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<document id="DC4BECD80933E3FE123BDC98016F8CAE" ID-CLB-Dataset="28377" ID-DOI="10.11646/zootaxa.4565.4.2" ID-GBIF-Dataset="7603c4db-a658-4941-88b2-75bdc2d2b402" ID-ISSN="1175-5326" ID-Zenodo-Dep="2591307" ID-ZooBank="EC3B414E-9699-4178-AC58-9D640C6B64CD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1552367174595" checkinUser="plazi" docAuthor="Huerta, Heron, Dzul-Manzanilla, Felipe, Navarrete-Carballo, Juan C., Manrique-Saide, Pablo &amp; Hancock, E. Geoffrey" docDate="2019" docId="8B1E550EFF89C812FF0725E2FA4E74E1" docLanguage="en" docName="zootaxa.4565.4.2.pdf" docOrigin="Zootaxa 4565 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Olbiogaster Osten Sacken 1886" docType="key" docVersion="7" lastPageNumber="477" masterDocId="77272D76FF88C810FF902344FFD27511" masterDocTitle="Olbiogaster Osten Sacken (Diptera: Anisopodidae) from Mexico, with the description of three new species" masterLastPageNumber="492" masterPageNumber="475" pageNumber="476" updateTime="1698701625847" updateUser="ExternalLinkService">
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<mods:title id="711DDBA9B56289330E758AD751182BF2">Olbiogaster Osten Sacken (Diptera: Anisopodidae) from Mexico, with the description of three new species</mods:title>
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<mods:namePart id="C634DF61F588D00E7909E54D8390A2D6">Huerta, Heron</mods:namePart>
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<mods:namePart id="BD647EBF64386171EA9FCF2156CA6150">Dzul-Manzanilla, Felipe</mods:namePart>
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<mods:namePart id="7CA5ABDFD42F8C99E4E73F3855E6A5D4">Navarrete-Carballo, Juan C.</mods:namePart>
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<mods:namePart id="61FAA9910A013ED7EA23623E85D3218D">Manrique-Saide, Pablo</mods:namePart>
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<mods:namePart id="975E41FB8B0EC00825B3596EE9EBC9F0">Hancock, E. Geoffrey</mods:namePart>
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<mods:title id="E3A9F63B71829000219D824F5D86DB4A">Zootaxa</mods:title>
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<mods:date id="77AB0620C3BA4349B8A329FBAD8C9A8C">2019</mods:date>
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<mods:number id="387EA707C549E5010918C779429B8C57">2019-03-12</mods:number>
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<mods:number id="9872881CE4E0D93AB0BF9783EDB3BEC3">4565</mods:number>
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<treatment id="8B1E550EFF89C812FF0725E2FA4E74E1" ID-DOI="http://doi.org/10.5281/zenodo.5942455" ID-GBIF-Taxon="154402105" ID-Zenodo-Dep="5942455" LSID="urn:lsid:plazi:treatment:8B1E550EFF89C812FF0725E2FA4E74E1" httpUri="http://treatment.plazi.org/id/8B1E550EFF89C812FF0725E2FA4E74E1" lastPageId="2" lastPageNumber="477" pageId="1" pageNumber="476">
<subSubSection id="4BADB793FF89C811FF0725E2FC9F73D1" box="[151,845,1701,1728]" pageId="1" pageNumber="476" type="nomenclature">
<paragraph id="0308E418FF89C811FF0725E2FC9F73D1" blockId="1.[151,845,1701,1728]" box="[151,845,1701,1728]" pageId="1" pageNumber="476">
<heading id="58405374FF89C811FF0725E2FC9F73D1" bold="true" box="[151,845,1701,1728]" fontSize="11" level="1" pageId="1" pageNumber="476" reason="1">
<emphasis id="31C3380AFF89C811FF0725E2FC9F73D1" bold="true" box="[151,845,1701,1728]" pageId="1" pageNumber="476">
Key to the Mexican species of
<taxonomicName id="C4B79F9BFF89C811FD9F25E1FC9F73D1" ID-CoL="67JN" ID-ENA="560768" authority="Osten Sacken" authorityName="Osten Sacken" authorityYear="1886" box="[527,845,1701,1728]" class="Insecta" family="Anisopodidae" genus="Olbiogaster" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="476" phylum="Arthropoda" rank="genus">
<emphasis id="31C3380AFF89C811FD9F25E1FD4973AE" bold="true" box="[527,667,1701,1727]" italics="true" pageId="1" pageNumber="476">Olbiogaster</emphasis>
Osten Sacken
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="4BADB793FF89C811FF0725A9FE3A7260" pageId="1" pageNumber="476" type="discussion">
<paragraph id="0308E418FF89C811FF0725A9FE3A7260" blockId="1.[151,1436,1772,1905]" pageId="1" pageNumber="476">
<taxonomicName id="C4B79F9BFF89C811FF0725A9FDA17214" authority="Lane &amp; d'Andretta" authorityName="Lane &amp; d'Andretta" authorityYear="1958" box="[151,627,1772,1797]" class="Insecta" family="Anisopodidae" genus="Olbiogaster" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="476" phylum="Arthropoda" rank="species" species="mexicana">
<emphasis id="31C3380AFF89C811FF0725A9FE437215" box="[151,401,1773,1797]" italics="true" pageId="1" pageNumber="476">Olbiogaster mexicana</emphasis>
Lane &amp; d'Andretta
</taxonomicName>
is only known from females. Regarding
<taxonomicName id="C4B79F9BFF89C811FBDE25A9FA4E7214" authority="Edwards" authorityName="Edwards" box="[1102,1436,1772,1797]" class="Insecta" family="Anisopodidae" genus="Olbiogaster" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="476" phylum="Arthropoda" rank="species" species="sackeni">
<emphasis id="31C3380AFF89C811FBDE25A9FAE37214" box="[1102,1329,1773,1797]" italics="true" pageId="1" pageNumber="476">Olbiogaster sackeni</emphasis>
Edwards
</taxonomicName>
and
<taxonomicName id="C4B79F9BFF89C811FF572455FE697238" authority="(Bellardi)" authorityName="Bellardi" baseAuthorityName="Bellardi" baseAuthorityYear="1862" box="[199,443,1808,1833]" class="Insecta" family="Anisopodidae" genus="Olbiogaster" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="476" phylum="Arthropoda" rank="species" species="taeniata">
<emphasis id="31C3380AFF89C811FF572455FE947239" box="[199,326,1809,1832]" italics="true" pageId="1" pageNumber="476">O. taeniata</emphasis>
(Bellardi)
</taxonomicName>
the description of females included some characteristics indicated in the key of
<bibRefCitation id="672699E9FF89C811FAD72455FE75725C" author="Lane, J." pageId="1" pageNumber="476" pagination="497 - 528" refId="ref8777" refString="Lane, J. &amp; d'Andretta (1958) Neotropical Anisopodidae (Diptera, Nematocera). Studia Entomologica, 1 (3 - 4), 497 - 528." type="journal article" year="1958">Lane &amp; d'Andretta (1958: 514)</bibRefCitation>
.
<bibRefCitation id="672699E9FF89C811FE272471FDBF725C" author="Peterson" box="[439,621,1845,1869]" pageId="1" pageNumber="476" pagination="305 - 312" refId="ref9112" refString="Peterson, B. V. (1981) Chapter 19. Anisopodidae. In: McAlpine, J. F., Peterson, B. V., Shewell, G. E., Teskey, H. J., Vockeroth, J. R. &amp; Wood, D. M. (eds.), Manual of Nearctic Diptera: Folume 1. Research Branch Agriculture Canada, Monograph No. 27, Ottawa, Ontario, pp. 305 - 312." type="book chapter" year="1981">Peterson (1981)</bibRefCitation>
added some illustrations of the wing, thorax, tarsomere and claw of
<taxonomicName id="C4B79F9BFF89C811FF07241CFE3A7260" authority="Edwards." authorityName="Edwards." box="[151,488,1880,1905]" class="Insecta" family="Anisopodidae" genus="Olbiogaster" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="476" phylum="Arthropoda" rank="species" species="sackeni">
<emphasis id="31C3380AFF89C811FF07241CFEAA7261" box="[151,376,1880,1904]" italics="true" pageId="1" pageNumber="476">Olbiogaster sackeni</emphasis>
Edwards.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="4BADB793FF89C812FF0724DBFA4E74E1" lastPageId="2" lastPageNumber="477" pageId="1" pageNumber="476" type="key">
<keyStep id="B843FCBDFF89C812FF0724DBFA4975BC" lastPageId="2" lastPageNumber="477" pageId="1" pageNumber="476">
<paragraph id="0308E418FF89C811FF0724DBFA4972E1" blockId="1.[151,1435,1951,2032]" pageId="1" pageNumber="476">
<keyLead id="B846472DFF89C811FF0724DBFA4972E1" pageId="1" pageNumber="476">
1. Male................................................................................................ 2 - Female.............................................................................................. 6 2(1). Front coxa yellow......................................................................
<taxonomicName id="C4B79F9BFF89C811FB4C249EFA4972E1" ID-CoL="492PP" authority="Edwards" authorityName="Edwards" box="[1244,1435,2010,2032]" class="Insecta" family="Anisopodidae" genus="Olbiogaster" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="476" phylum="Arthropoda" rank="species" species="sackeni">
<emphasis id="31C3380AFF89C811FB4C249EFA9072E1" box="[1244,1346,2010,2032]" italics="true" pageId="1" pageNumber="476">O. sackeni</emphasis>
Edwards
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="0308E418FF8AC812FF0723D3FA4975BC" blockId="2.[151,1437,151,496]" box="[151,1435,151,173]" pageId="2" pageNumber="477">
<keyLead id="B846472DFF8AC812FF0723D3FA4975BC" box="[151,1435,151,173]" pageId="2" pageNumber="477">- Front coxa dark brown................................................................................. 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="B843FCBDFF8AC812FF0723F0FA4E75F9" pageId="2" pageNumber="477">
<paragraph id="0308E418FF8AC812FF0723F0FA4975DB" blockId="2.[151,1437,151,496]" box="[151,1435,180,202]" pageId="2" pageNumber="477">
<keyLead id="B846472DFF8AC812FF0723F0FA4975DB" box="[151,1435,180,202]" pageId="2" pageNumber="477">3(2). Hind femur dark brown with basal or distal yellow ring....................................................... 4</keyLead>
</paragraph>
<paragraph id="0308E418FF8AC812FF072396FA4E75F9" blockId="2.[151,1437,151,496]" box="[151,1436,209,232]" pageId="2" pageNumber="477">
<keyLead id="B846472DFF8AC812FF072396FA4E75F9" box="[151,1436,209,232]" pageId="2" pageNumber="477">
- Hind femur completely dark brown (
<figureCitation id="9B8CF89DFF8AC812FDD12395FD5F75F6" box="[577,653,209,231]" captionStart="FIGURE 9" captionStartId="12.[151,250,1971,1993]" captionTargetBox="[341,1307,208,1862]" captionTargetId="figure@12.[226,1351,193,1945]" captionTargetPageId="12" captionText="FIGURE 9. Olbiogaster puuc sp. nov. Male holotype, Legs, lateral view. A. foreleg, B. midleg, C. hind leg. Abbreviations. cx= coxa; fe= femur; tb= tibia; tr= trochanter; tsm= tarsomere 1 to 5. Scale bar= 0.1 mm." httpUri="https://zenodo.org/record/2591323/files/figure.png" pageId="2" pageNumber="477">Fig. 9C</figureCitation>
)....................................................
<taxonomicName id="C4B79F9BFF8AC812FB6D2395FA9B75F6" authority="Huerta &amp; Dzul-Manzanilla &amp; Navarrete-Carballo &amp; Manrique-Saide &amp; Hancock, 2019" authorityName="Huerta &amp; Dzul-Manzanilla &amp; Navarrete-Carballo &amp; Manrique-Saide &amp; Hancock" authorityYear="2019" box="[1277,1353,209,231]" class="Insecta" family="Anisopodidae" genus="Olbiogaster" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="477" phylum="Arthropoda" rank="species" species="puuc" status="sp. nov.">
<emphasis id="31C3380AFF8AC812FB6D2395FA9B75F6" box="[1277,1353,209,231]" italics="true" pageId="2" pageNumber="477">O. puuc</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2AF08571FF8AC812FADF2396FA4E75F9" box="[1359,1436,210,232]" pageId="2" pageNumber="477" rank="species">
<emphasis id="31C3380AFF8AC812FADF2396FA4E75F9" bold="true" box="[1359,1436,210,232]" pageId="2" pageNumber="477">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="B843FCBDFF8AC812FF0723ABFA4E742E" pageId="2" pageNumber="477">
<paragraph id="0308E418FF8AC812FF0723ABFA4E7433" blockId="2.[151,1437,151,496]" pageId="2" pageNumber="477">
<keyLead id="B846472DFF8AC812FF0723ABFA4E7433" pageId="2" pageNumber="477">
4(3). Mesonotum with a dark brown central stripe, surrounded U-shaped black stripe (
<figureCitation id="9B8CF89DFF8AC812FC6323ABFB927414" box="[1011,1088,239,261]" captionStart="FIGURE 2" captionStartId="5.[151,250,1977,1999]" captionTargetBox="[197,1348,358,1897]" captionTargetId="figure@5.[161,1427,345,1958]" captionTargetPageId="5" captionText="FIGURE 2. Olbiogaster nuxco sp. nov. Male paratype, A. Antenna; B. Head, frontal view; C. Thorax, dorsal view. Scale bar: A-B=0.5 mm, C=1.0 mm." httpUri="https://zenodo.org/record/2591309/files/figure.png" pageId="2" pageNumber="477">Fig. 2C</figureCitation>
); sternite 10 with elongated slender arms, apically with a long spine, curved (
<figureCitation id="9B8CF89DFF8AC812FDFC2248FD7D7433" box="[620,687,268,290]" captionStart="FIGURE 6" captionStartId="9.[151,250,1928,1950]" captionTargetBox="[276,1322,226,1869]" captionTargetId="figure@9.[229,1361,193,1911]" captionTargetPageId="9" captionText="FIGURE 6. Male genitalia, ventral view. A–D. Olbiogaster halffteri sp. nov.; E–G. Olbiogaster nuxco sp. nov.; H–J. Olbiogaster puuc sp. nov. Aedeagal guide: A, B, E, H; gonostylus: C, F, I; sternite X: D, G, J. Scale bar: A, C, D= 0.1 mm; E– J= 0.16 mm." httpUri="https://zenodo.org/record/2591317/files/figure.png" pageId="2" pageNumber="477">Fig. 6J</figureCitation>
)................................................
<taxonomicName id="C4B79F9BFF8AC812FB622248FA9B7433" authority="Huerta &amp; Dzul-Manzanilla &amp; Navarrete-Carballo &amp; Manrique-Saide &amp; Hancock, 2019" authorityName="Huerta &amp; Dzul-Manzanilla &amp; Navarrete-Carballo &amp; Manrique-Saide &amp; Hancock" authorityYear="2019" box="[1266,1353,268,290]" class="Insecta" family="Anisopodidae" genus="Olbiogaster" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="477" phylum="Arthropoda" rank="species" species="nuxco" status="sp. nov.">
<emphasis id="31C3380AFF8AC812FB622248FA9B7433" box="[1266,1353,268,290]" italics="true" pageId="2" pageNumber="477">O. nuxco</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2AF08571FF8AC812FADF2248FA4E7433" box="[1359,1436,268,290]" pageId="2" pageNumber="477" rank="species">
<emphasis id="31C3380AFF8AC812FADF2248FA4E7433" bold="true" box="[1359,1436,268,290]" pageId="2" pageNumber="477">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="0308E418FF8AC812FF07226EFA4E742E" blockId="2.[151,1437,151,496]" box="[151,1436,297,319]" pageId="2" pageNumber="477">
<keyLead id="B846472DFF8AC812FF07226EFA4E742E" box="[151,1436,297,319]" pageId="2" pageNumber="477">- Mesonotum with different pattern; sternite 10 with elongated slender arms, apically with spine, slightly curved or a few setae5</keyLead>
</paragraph>
</keyStep>
<keyStep id="B843FCBDFF8AC812FF072203FA4E7486" pageId="2" pageNumber="477">
<paragraph id="0308E418FF8AC812FF072203FA44746B" blockId="2.[151,1437,151,496]" pageId="2" pageNumber="477">
<keyLead id="B846472DFF8AC812FF072203FA44746B" pageId="2" pageNumber="477">
5(4). Hind femur basal and apical 1/3 dark, middle 1/3 yellow; sternite 10 with elongated slender arms, apically not widened (
<figureCitation id="9B8CF89DFF8AC812FAE52203FA4F744C" box="[1397,1437,327,349]" captionStart="FIGURE 6" captionStartId="9.[151,250,1928,1950]" captionTargetBox="[276,1322,226,1869]" captionTargetId="figure@9.[229,1361,193,1911]" captionTargetPageId="9" captionText="FIGURE 6. Male genitalia, ventral view. A–D. Olbiogaster halffteri sp. nov.; E–G. Olbiogaster nuxco sp. nov.; H–J. Olbiogaster puuc sp. nov. Aedeagal guide: A, B, E, H; gonostylus: C, F, I; sternite X: D, G, J. Scale bar: A, C, D= 0.1 mm; E– J= 0.16 mm." httpUri="https://zenodo.org/record/2591317/files/figure.png" pageId="2" pageNumber="477">Fig.</figureCitation>
6D)..................................................................................
<taxonomicName id="C4B79F9BFF8AC812FB452220FA90746B" authority="Huerta &amp; Dzul-Manzanilla &amp; Navarrete-Carballo &amp; Manrique-Saide &amp; Hancock, 2019" authorityName="Huerta &amp; Dzul-Manzanilla &amp; Navarrete-Carballo &amp; Manrique-Saide &amp; Hancock" authorityYear="2019" box="[1237,1346,356,378]" class="Insecta" family="Anisopodidae" genus="Olbiogaster" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="477" phylum="Arthropoda" rank="species" species="halffteri" status="sp. nov.">
<emphasis id="31C3380AFF8AC812FB452220FA90746B" box="[1237,1346,356,378]" italics="true" pageId="2" pageNumber="477">O. halffteri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2AF08571FF8AC812FAD92220FA44746B" box="[1353,1430,356,378]" pageId="2" pageNumber="477" rank="species">
<emphasis id="31C3380AFF8AC812FAD92220FA44746B" bold="true" box="[1353,1430,356,378]" pageId="2" pageNumber="477">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="0308E418FF8AC812FF0722C6FA4E7486" blockId="2.[151,1437,151,496]" box="[151,1436,385,407]" pageId="2" pageNumber="477">
<keyLead id="B846472DFF8AC812FF0722C6FA4E7486" box="[151,1436,385,407]" pageId="2" pageNumber="477">
- Hind femur dark, except for yellow apex; sternite 10 with elongated slender arms, apically enlarged....
<taxonomicName id="C4B79F9BFF8AC812FB5C22C5FA4E7486" authority="(Bellardi)" authorityName="Bellardi" baseAuthorityName="Bellardi" baseAuthorityYear="1862" box="[1228,1436,385,407]" class="Insecta" family="Anisopodidae" genus="Olbiogaster" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="477" phylum="Arthropoda" rank="species" species="taeniata">
<emphasis id="31C3380AFF8AC812FB5C22C5FAEB7486" box="[1228,1337,385,407]" italics="true" pageId="2" pageNumber="477">O. taeniata</emphasis>
(Bellardi)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="B843FCBDFF8AC812FF0722DBFA4E74E1" pageId="2" pageNumber="477">
<paragraph id="0308E418FF8AC812FF0722DBFA4974A4" blockId="2.[151,1437,151,496]" box="[151,1435,415,437]" pageId="2" pageNumber="477">
<keyLead id="B846472DFF8AC812FF0722DBFA4974A4" box="[151,1435,415,437]" pageId="2" pageNumber="477">
6(1). Coxae mostly blackish brown; hind femur entirely yellowish..........................
<taxonomicName id="C4B79F9BFF8AC812FBF622DBFA4974A4" ID-CoL="492PC" authority="Lane &amp; d'Andretta" authorityName="Lane &amp; d'Andretta" authorityYear="1958" box="[1126,1435,415,437]" class="Insecta" family="Anisopodidae" genus="Olbiogaster" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="477" phylum="Arthropoda" rank="species" species="mexicana">
<emphasis id="31C3380AFF8AC812FBF622DBFB0D74A4" box="[1126,1247,415,437]" italics="true" pageId="2" pageNumber="477">O. mexicana</emphasis>
Lane &amp; d´Andretta
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="0308E418FF8AC812FF0722F9FA4E74E1" blockId="2.[151,1437,151,496]" pageId="2" pageNumber="477">
<keyLead id="B846472DFF8AC812FF0722F9FA4E74E1" pageId="2" pageNumber="477">
- Fore and midcoxae yellow, with dark brown tinge; hind coxae dark brown; hind femur dark, except for basal 2/3 yellow ring (
<figureCitation id="9B8CF89DFF8AC812FF78229DFEE074FE" box="[232,306,473,495]" captionStart="FIGURE 7" captionStartId="10.[151,250,1590,1612]" captionTargetBox="[181,1377,211,1568]" captionTargetId="figure@10.[151,1436,193,1568]" captionTargetPageId="10" captionText="FIGURE 7. Olbiogaster nuxco sp. nov. Female paratype, A. head, thorax and first and second segment abdominal, dorsal view; B. head and thorax, lateral view; C. Legs, top: hind leg, middle: midleg, lower: foreleg, lateral view; D. Abdomen, dorsal view; E. genitalia, ventral view. Scale bar: A, B, D=1.0 mm; C=0.1 mm, E=0.05 mm." httpUri="https://zenodo.org/record/2591319/files/figure.png" pageId="2" pageNumber="477">Fig. 7C</figureCitation>
)...............................................................................
<taxonomicName id="C4B79F9BFF8AC812FB62229DFA9B74FE" authority="Huerta &amp; Dzul-Manzanilla &amp; Navarrete-Carballo &amp; Manrique-Saide &amp; Hancock, 2019" authorityName="Huerta &amp; Dzul-Manzanilla &amp; Navarrete-Carballo &amp; Manrique-Saide &amp; Hancock" authorityYear="2019" box="[1266,1353,473,495]" class="Insecta" family="Anisopodidae" genus="Olbiogaster" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="477" phylum="Arthropoda" rank="species" species="nuxco" status="sp. nov.">
<emphasis id="31C3380AFF8AC812FB62229DFA9B74FE" box="[1266,1353,473,495]" italics="true" pageId="2" pageNumber="477">O. nuxco</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2AF08571FF8AC812FADF229EFA4E74E1" box="[1359,1436,474,496]" pageId="2" pageNumber="477" rank="species">
<emphasis id="31C3380AFF8AC812FADF229EFA4E74E1" bold="true" box="[1359,1436,474,496]" pageId="2" pageNumber="477">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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