394 lines
58 KiB
XML
394 lines
58 KiB
XML
<document id="04AE5F0950929FDA12B2A70E972E60D3" ID-DOI="10.11646/zootaxa.5375.3.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="10199873" ID-ZooBank="537770D1-40D2-40A7-A07C-4C2A9CB6C4E2" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1700727168078" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2023" docId="731687CCFFC54C25FF37EF87FA37F805" docLanguage="en" docName="zootaxa.5375.3.6.pdf" docOrigin="Zootaxa 5375 (3)" docSource="https://mapress.com/zt/article/download/zootaxa.5375.3.6/52337" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sinacrulia Shavrin 2023, gen. n." docType="treatment" docVersion="3" lastPageNumber="434" masterDocId="8F2FFFB4FFC44C20FFA0EE79FFA3FFDC" masterDocTitle="A new genus of Omaliini from China (Coleoptera: Staphylinidae: Omaliinae)" masterLastPageNumber="438" masterPageNumber="429" pageNumber="430" updateTime="1700833506740" updateUser="valdenar" zenodo-license-document="CLOSED">
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<mods:title id="91FF8ED4237EC8A61F64FFCA9B560D69">A new genus of Omaliini from China (Coleoptera: Staphylinidae: Omaliinae)</mods:title>
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<mods:namePart id="3B7FF633FD96ABA81E8DA74B356780D8">Shavrin, Alexey V.</mods:namePart>
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<mods:date id="0FC5C0D9E91B18F08E9A4DCFB369847B">2023</mods:date>
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<mods:number id="794E3C5F6D7857B8F1399FDEFB6D01C1">2023-11-23</mods:number>
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<paragraph id="FB0036DAFFC54C21FF37EF87FED3FDC5" blockId="1.[151,368,510,573]" box="[151,368,510,537]" pageId="1" pageNumber="430">
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<emphasis id="C9CBEAC8FFC54C21FF37EF87FED3FDC5" bold="true" box="[151,368,510,537]" pageId="1" pageNumber="430">
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<taxonomicName id="3CBF4D59FFC54C21FF37EF87FEB5FDC5" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[151,278,510,537]" class="Insecta" family="Staphylinidae" genus="Sinacrulia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="430" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="C9CBEAC8FFC54C21FF37EF87FEB5FDC5" bold="true" box="[151,278,510,537]" italics="true" pageId="1" pageNumber="430">Sinacrulia</emphasis>
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<taxonomicNameLabel id="D2F857B3FFC54C21FEBEEF86FED3FDC5" box="[286,368,511,537]" pageId="1" pageNumber="430" rank="genus">gen. n.</taxonomicNameLabel>
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<subSubSection id="B3A56551FFC54C21FF37EC5AFEBAFDE1" box="[151,281,547,573]" pageId="1" pageNumber="430" type="description">
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<paragraph id="FB0036DAFFC54C21FF37EC5AFEBAFDE1" blockId="1.[151,368,510,573]" box="[151,281,547,573]" pageId="1" pageNumber="430">
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(
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<figureCitation id="63842A5FFFC54C21FF3FEC5AFEB3FDE1" box="[159,272,547,573]" captionStart-0="FIGURES 1–2" captionStart-1="FIGURES 3–8" captionStart-2="FIGURES 9–10" captionStart-3="FIGURES 11–15" captionStart-4="FIGURE 16" captionStart-5="FIGURE 17" captionStartId-0="2.[151,264,1563,1587]" captionStartId-1="3.[151,264,1435,1459]" captionStartId-2="4.[151,264,1964,1988]" captionStartId-3="5.[151,264,1481,1505]" captionStartId-4="7.[151,250,2003,2027]" captionStartId-5="8.[151,250,916,940]" captionTargetBox-0="[212,1371,391,1520]" captionTargetBox-1="[154,1433,193,1408]" captionTargetBox-2="[155,1432,637,1937]" captionTargetBox-3="[155,1433,185,1453]" captionTargetBox-4="[167,1419,309,1980]" captionTargetBox-5="[151,1436,181,893]" captionTargetId-0="figure-97@2.[200,1388,378,1537]" captionTargetId-1="figure-15@3.[151,1436,181,1411]" captionTargetId-2="figure-227@4.[151,1436,633,1941]" captionTargetId-3="figure-15@5.[151,1436,181,1457]" captionTargetId-4="figure-76@7.[167,1419,309,1980]" captionTargetId-5="figure-18@8.[151,1436,181,893]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="7" captionTargetPageId-5="8" captionText-0="FIGURES 1–2. Habitus of Sinacrulia: 1–S. fortiterpunctata (holotype), 2–S. sp. Scale bars: 1.0 mm." captionText-1="FIGURES 3–8. Mouthparts of Sinacrulia fortiterpunctata (holotype): 3–labrum (dorsal view), 4–left mandible (dorsal view), 5–right mandible (dorsal view), 6–labium (ventral view), 7–mentum (ventral view), 8–right maxilla (dorsal view). Scale bars: 0.1 mm." captionText-2="FIGURES 9–10. Aedeagus of Sinacrulia fortiterpunctata (holotype): 9–parameral view, 10–lateral view. Scale bar: 0.1 mm." captionText-3="FIGURES 11–15. Abdominal segments of Sinacrulia (Figs 11–12: S. fortiterpunctata (holotype), Figs. 13–15: S. sp.): 11–male tergite VIII (dorsal view), 12–male sternite VIII (ventral view), 13–female tergite VIII (dorsal view), 14–female sternite VIII (ventral view), 15–female genital segment (dorsal view). Scale bars: 0.1 mm." captionText-4="FIGURE 16. Type locality of Sinacrulia fortiterpunctata in the Diancang Shan, Yunnan, China (photograph: M. Schülke)." captionText-5="FIGURE 17. Distribution of Sinacrulia in China (black circle: S. fortiterpunctata, white circle: S. sp.)." figureDoi-0="http://doi.org/10.5281/zenodo.10199875" figureDoi-1="http://doi.org/10.5281/zenodo.10199877" figureDoi-2="http://doi.org/10.5281/zenodo.10199879" figureDoi-3="http://doi.org/10.5281/zenodo.10199883" figureDoi-4="http://doi.org/10.5281/zenodo.10199885" figureDoi-5="http://doi.org/10.5281/zenodo.10199887" httpUri-0="https://zenodo.org/record/10199875/files/figure.png" httpUri-1="https://zenodo.org/record/10199877/files/figure.png" httpUri-2="https://zenodo.org/record/10199879/files/figure.png" httpUri-3="https://zenodo.org/record/10199883/files/figure.png" httpUri-4="https://zenodo.org/record/10199885/files/figure.png" httpUri-5="https://zenodo.org/record/10199887/files/figure.png" pageId="1" pageNumber="430">Figs 1–17</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="B3A56551FFC54C21FF37EC12FD62FD59" box="[151,705,618,645]" pageId="1" pageNumber="430" type="type_taxon">
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<paragraph id="FB0036DAFFC54C21FF37EC12FD62FD59" blockId="1.[151,705,618,645]" box="[151,705,618,645]" pageId="1" pageNumber="430">
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<emphasis id="C9CBEAC8FFC54C21FF37EC12FE9FFD58" bold="true" box="[151,316,618,645]" pageId="1" pageNumber="430">
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<typeStatus id="24048878FFC54C21FF37EC12FF77FD59" box="[151,212,619,645]" pageId="1" pageNumber="430">Type</typeStatus>
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species:
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</emphasis>
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<taxonomicName id="3CBF4D59FFC54C21FEE3EC12FDDCFD59" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[323,639,619,645]" class="Insecta" family="Staphylinidae" genus="Sinacrulia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="430" phylum="Arthropoda" rank="species" species="fortiterpunctata" status="sp. nov.">
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<emphasis id="C9CBEAC8FFC54C21FEE3EC12FDDCFD59" box="[323,639,619,645]" italics="true" pageId="1" pageNumber="430">Sinacrulia fortiterpunctata</emphasis>
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</taxonomicName>
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<emphasis id="C9CBEAC8FFC54C21FD26EC12FD62FD59" bold="true" box="[646,705,619,645]" pageId="1" pageNumber="430">
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<taxonomicNameLabel id="D2F857B3FFC54C21FD26EC12FD62FD59" box="[646,705,619,645]" pageId="1" pageNumber="430" rank="species">sp.n.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="B3A56551FFC54C23FF37ECCBFCA5F96C" lastPageId="3" lastPageNumber="432" pageId="1" pageNumber="430" type="description">
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<paragraph id="FB0036DAFFC54C21FF37ECCBFF69FC79" blockId="1.[151,1437,690,2013]" pageId="1" pageNumber="430">
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<emphasis id="C9CBEAC8FFC54C21FF37ECCBFE8AFD10" bold="true" box="[151,297,690,716]" pageId="1" pageNumber="430">Description.</emphasis>
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Body moderately convex and slightly elongate, brown or reddish-brown (
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<figureCitation id="63842A5FFFC54C21FB23ECCAFB4FFD11" box="[1155,1260,691,717]" captionStart="FIGURES 1–2" captionStartId="2.[151,264,1563,1587]" captionTargetBox="[212,1371,391,1520]" captionTargetId="figure-97@2.[200,1388,378,1537]" captionTargetPageId="2" captionText="FIGURES 1–2. Habitus of Sinacrulia: 1–S. fortiterpunctata (holotype), 2–S. sp. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10199875" httpUri="https://zenodo.org/record/10199875/files/figure.png" pageId="1" pageNumber="430">Figs 1–2</figureCitation>
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). Punctation of forebody very dense, large and coarse, markedly larger and deeper on pronotum and elytra, with distinct elevations between punctures in pronotum and elytra. Dorsal surface of body glossy; head with dense microsculpture in apical and lateral portions; pronotum and elytra without microreticulation; abdomen with very dense and coarse isodiametric microreticulation. Body without pubescence except of frontal portion of head with very sparse moderately long setae on apical part of clypeus and abdomen with sparse and very short recumbent setation. Body length:
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<quantity id="3C479B3FFFC54C21FA8CED1EFF65FC79" metricMagnitude="-3" metricUnit="m" metricValue="2.115" metricValueMax="2.22" metricValueMin="2.01" pageId="1" pageNumber="430" unit="mm" value="2.115" valueMax="2.22" valueMin="2.01">2.01–2.22 mm</quantity>
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.
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</paragraph>
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<paragraph id="FB0036DAFFC54C21FF67EDD6FC57F8D9" blockId="1.[151,1437,690,2013]" pageId="1" pageNumber="430">
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Head markedly narrower than pronotum, distinctly transverse, slightly convex in middle, with wide frontoclypeal portion and distinctly convex supra-antennal elevations, widely rounded and more convex at level of antennal insertions; each laterobasal part of supra-antennal elevation with narrow elongate wrinkle diagonally or directly stretching posteriad to level of anterior margin of eyes; nuchal constriction sharp, head strongly separated from neck. Head with a pair of ocelli located near nuchal constriction at level of hind angles of temples, with indistinct and narrow anteocellar foveae slightly curved lateroapicad in front of anterior margins of each ocellus. Postocular ridges missing. Anteriomedian depressions at level of antennal insertions fine and relatively wide. Eyes large, strongly convex, about twice longer than temple (if seen dorsally). Temples moderately long, subparallel at level slightly behind eyes and then widely rounded toward neck. Antennal insertion concealed under supra-antennal elevation; antenna slightly longer than head and pronotum combined, reaching basal margin of elytra when reclined; antennomeres 1–3 strongly elongate, 4–6 small, about as broad as long or slightly elongate, 7–10 distinctly transverse. Labrum (
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<figureCitation id="63842A5FFFC54C21FEEAEB42FE2EFA89" box="[330,397,1339,1365]" captionStart="FIGURES 3–8" captionStartId="3.[151,264,1435,1459]" captionTargetBox="[154,1433,193,1408]" captionTargetId="figure-15@3.[151,1436,181,1411]" captionTargetPageId="3" captionText="FIGURES 3–8. Mouthparts of Sinacrulia fortiterpunctata (holotype): 3–labrum (dorsal view), 4–left mandible (dorsal view), 5–right mandible (dorsal view), 6–labium (ventral view), 7–mentum (ventral view), 8–right maxilla (dorsal view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10199877" httpUri="https://zenodo.org/record/10199877/files/figure.png" pageId="1" pageNumber="430">Fig. 3</figureCitation>
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) small, with widely rounded latero-apical portions, widely concaved anteriorly, with four to six pairs of long anterolateral setae and two rows of of shorter setae in middle of apical portion. Mandibles (
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<figureCitation id="63842A5FFFC54C21FAC4EB26FF60FA41" captionStart="FIGURES 3–8" captionStartId="3.[151,264,1435,1459]" captionTargetBox="[154,1433,193,1408]" captionTargetId="figure-15@3.[151,1436,181,1411]" captionTargetPageId="3" captionText="FIGURES 3–8. Mouthparts of Sinacrulia fortiterpunctata (holotype): 3–labrum (dorsal view), 4–left mandible (dorsal view), 5–right mandible (dorsal view), 6–labium (ventral view), 7–mentum (ventral view), 8–right maxilla (dorsal view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10199877" httpUri="https://zenodo.org/record/10199877/files/figure.png" pageId="1" pageNumber="430">Figs. 4–5</figureCitation>
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) broad, each with concave latero-basal margin and subacute apex; right mandible (
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<figureCitation id="63842A5FFFC54C21FBD6EBFAFB19FA41" box="[1142,1210,1410,1437]" captionStart="FIGURES 3–8" captionStartId="3.[151,264,1435,1459]" captionTargetBox="[154,1433,193,1408]" captionTargetId="figure-15@3.[151,1436,181,1411]" captionTargetPageId="3" captionText="FIGURES 3–8. Mouthparts of Sinacrulia fortiterpunctata (holotype): 3–labrum (dorsal view), 4–left mandible (dorsal view), 5–right mandible (dorsal view), 6–labium (ventral view), 7–mentum (ventral view), 8–right maxilla (dorsal view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10199877" httpUri="https://zenodo.org/record/10199877/files/figure.png" pageId="1" pageNumber="430">Fig. 5</figureCitation>
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) with slightly more curved apex and small obtuse tooth at distal third of cutting edge; articulations of molar lobes with strong acute dents almost in entire half of latero-apical margin. Maxilla (
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<figureCitation id="63842A5FFFC54C21FC96EBB2FCDBFA39" box="[822,888,1483,1509]" captionStart="FIGURES 3–8" captionStartId="3.[151,264,1435,1459]" captionTargetBox="[154,1433,193,1408]" captionTargetId="figure-15@3.[151,1436,181,1411]" captionTargetPageId="3" captionText="FIGURES 3–8. Mouthparts of Sinacrulia fortiterpunctata (holotype): 3–labrum (dorsal view), 4–left mandible (dorsal view), 5–right mandible (dorsal view), 6–labium (ventral view), 7–mentum (ventral view), 8–right maxilla (dorsal view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10199877" httpUri="https://zenodo.org/record/10199877/files/figure.png" pageId="1" pageNumber="430">Fig. 8</figureCitation>
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) narrow, long; galea oblong, moderately narrow, with relatively dense long apical setae; lacinia very narrow, slightly curved on outer margin of apical half, with row of small thorns on apical third of inner margin; maxillary palpi moderately long, second palpomere distinctly longer than broad, third palpomere slightly transverse and slightly broader than preapical segment, apical palpomere moderately narrow, slightly less than three times as long as preapical segment, from basal part gradually narrowing apicad toward rounded apex. Labium (
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<figureCitation id="63842A5FFFC54C21FDDAE806FD1DF945" box="[634,702,1663,1689]" captionStart="FIGURES 3–8" captionStartId="3.[151,264,1435,1459]" captionTargetBox="[154,1433,193,1408]" captionTargetId="figure-15@3.[151,1436,181,1411]" captionTargetPageId="3" captionText="FIGURES 3–8. Mouthparts of Sinacrulia fortiterpunctata (holotype): 3–labrum (dorsal view), 4–left mandible (dorsal view), 5–right mandible (dorsal view), 6–labium (ventral view), 7–mentum (ventral view), 8–right maxilla (dorsal view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10199877" httpUri="https://zenodo.org/record/10199877/files/figure.png" pageId="1" pageNumber="430">Fig. 6</figureCitation>
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) distinctly bilobed, two preapical segments distinctly transverse, apical segment about twice longer than preapical segment, with rounded apex. Mentum (
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<figureCitation id="63842A5FFFC54C21FB24E8DAFB6BF961" box="[1156,1224,1699,1725]" captionStart="FIGURES 3–8" captionStartId="3.[151,264,1435,1459]" captionTargetBox="[154,1433,193,1408]" captionTargetId="figure-15@3.[151,1436,181,1411]" captionTargetPageId="3" captionText="FIGURES 3–8. Mouthparts of Sinacrulia fortiterpunctata (holotype): 3–labrum (dorsal view), 4–left mandible (dorsal view), 5–right mandible (dorsal view), 6–labium (ventral view), 7–mentum (ventral view), 8–right maxilla (dorsal view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10199877" httpUri="https://zenodo.org/record/10199877/files/figure.png" pageId="1" pageNumber="430">Fig. 7</figureCitation>
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) moderately large, transverse, trapezoidal, with three short setae on lateral margins. Gular sutures relatively narrowly separated from each other at level of posterior margins of eyes, strongly divergent posteriad.
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</paragraph>
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Pronotum transverse, convex, without (
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<taxonomicName id="3CBF4D59FFC54C21FD2EE976FCFCF8F5" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[654,863,1807,1833]" class="Insecta" family="Staphylinidae" genus="Sinacrulia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="430" phylum="Arthropoda" rank="species" species="fortiterpunctata" status="sp. nov.">
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<emphasis id="C9CBEAC8FFC54C21FD2EE976FD3FF8F5" box="[654,668,1807,1833]" italics="true" pageId="1" pageNumber="430">S</emphasis>
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.
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<emphasis id="C9CBEAC8FFC54C21FD0EE976FCFCF8F5" box="[686,863,1807,1833]" italics="true" pageId="1" pageNumber="430">fortiterpunctata</emphasis>
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</taxonomicName>
|
||
<emphasis id="C9CBEAC8FFC54C21FCCAE976FC02F8F5" bold="true" box="[874,929,1807,1833]" pageId="1" pageNumber="430">
|
||
<taxonomicNameLabel id="D2F857B3FFC54C21FCCAE976FC02F8F5" box="[874,929,1807,1833]" pageId="1" pageNumber="430" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
) or with (
|
||
<emphasis id="C9CBEAC8FFC54C21FBBCE976FB89F8F5" box="[1052,1066,1807,1833]" italics="true" pageId="1" pageNumber="430">S</emphasis>
|
||
. sp.) indistinct longitudinal and semioval mediobasal impressions, with slightly and widely impressed laterobasal portions, with elongate curved relatively short narrow elevation in latero-apical part; lateral margins distinctly crenulate, with indistinct and irregular (
|
||
<emphasis id="C9CBEAC8FFC54C21FF56E902FEA7F849" box="[246,260,1915,1941]" italics="true" pageId="1" pageNumber="430">S</emphasis>
|
||
. sp.) or with relatively regular small acute thorn-shaped crenulation. Prothorax short, strongly transverse, convex in middle, with short acute intercoxal process reaching about middle of front coxae. Mesoventrite very short, without carina, with acute elongate process, reaching apex of long and narrow intercoxal process extending to about middle of mesocoxae. Scutellum wide, triangular. Metaventrite markedly transverse, convex, with deep mesocoxal cavities and wide moderately short intercoxal process, obtuse apically and extending to mesosternal process.
|
||
</paragraph>
|
||
<paragraph id="FB0036DAFFC64C22FF67EEA6FDA7FEB9" blockId="2.[151,1436,151,357]" pageId="2" pageNumber="431">Elytra convex, indistinctly depressed in mediobasal portion, moderately short and wide, extending to apical margin of abdominal tergite III or basal margin of abdominal tergite IV, slightly less than twice as long as pronotum, slightly broadened posteriad, with broad epipleura and well-developed rounded humeri, with slightly serrate lateral margins. Wings fully developed.</paragraph>
|
||
<caption id="AFC06652FFC64C22FF37E862FB34F9EF" ID-DOI="http://doi.org/10.5281/zenodo.10199875" ID-Zenodo-Dep="10199875" box="[151,1175,1563,1588]" httpUri="https://zenodo.org/record/10199875/files/figure.png" pageId="2" pageNumber="431" startId="2.[151,264,1563,1587]" targetBox="[212,1371,391,1520]" targetPageId="2" targetType="figure">
|
||
<paragraph id="FB0036DAFFC64C22FF37E862FB34F9EF" blockId="2.[151,1175,1563,1588]" box="[151,1175,1563,1588]" pageId="2" pageNumber="431">
|
||
<emphasis id="C9CBEAC8FFC64C22FF37E862FE99F9E8" bold="true" box="[151,314,1563,1588]" pageId="2" pageNumber="431">FIGURES 1–2.</emphasis>
|
||
Habitus of
|
||
<taxonomicName id="3CBF4D59FFC64C22FE11E865FDB8F9E8" authorityName="Shavrin" authorityYear="2023" box="[433,539,1564,1588]" class="Insecta" family="Staphylinidae" genus="Sinacrulia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="431" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C9CBEAC8FFC64C22FE11E865FDB8F9E8" box="[433,539,1564,1588]" italics="true" pageId="2" pageNumber="431">Sinacrulia</emphasis>
|
||
</taxonomicName>
|
||
: 1–
|
||
<taxonomicName id="3CBF4D59FFC64C22FDE2E865FD58F9E8" authorityName="Shavrin" authorityYear="2023" box="[578,763,1564,1588]" class="Insecta" family="Staphylinidae" genus="Sinacrulia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="431" phylum="Arthropoda" rank="species" species="fortiterpunctata">
|
||
<emphasis id="C9CBEAC8FFC64C22FDE2E865FD58F9E8" box="[578,763,1564,1588]" italics="true" pageId="2" pageNumber="431">S. fortiterpunctata</emphasis>
|
||
</taxonomicName>
|
||
(holotype), 2–
|
||
<emphasis id="C9CBEAC8FFC64C22FC31E865FC06F9EF" box="[913,933,1564,1587]" italics="true" pageId="2" pageNumber="431">S.</emphasis>
|
||
sp. Scale bars: 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="FB0036DAFFC64C22FF67E81BFE1EF8D0" blockId="2.[151,1436,1634,2020]" pageId="2" pageNumber="431">
|
||
Legs moderately short; mesocoxae oval; metacoxae large and subtriangular; tibiae slightly widened apicad; apical half of pro- and mesotibia with elongate spines on inner and outer margins, slightly denser on protibiae; metatibiae without modification and spines, covered with long setae, denser and longer on inner margin; tarsi short, apical tarsomere long, twice longer than previous four tarsomeres combined; protarsi with very long lateral setae (
|
||
<taxonomicName id="3CBF4D59FFC64C22FF01E88BFECDF8D0" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[161,366,1778,1804]" class="Insecta" family="Staphylinidae" genus="Sinacrulia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="431" phylum="Arthropoda" rank="species" species="fortiterpunctata" status="sp. nov.">
|
||
<emphasis id="C9CBEAC8FFC64C22FF01E88BFF0CF8D0" box="[161,175,1778,1804]" italics="true" pageId="2" pageNumber="431">S</emphasis>
|
||
.
|
||
<emphasis id="C9CBEAC8FFC64C22FF1DE88BFECDF8D0" box="[189,366,1778,1804]" italics="true" pageId="2" pageNumber="431">fortiterpunctata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C9CBEAC8FFC64C22FED5E88BFE0FF8D0" bold="true" box="[373,428,1778,1804]" pageId="2" pageNumber="431">
|
||
<taxonomicNameLabel id="D2F857B3FFC64C22FED5E88BFE0FF8D0" box="[373,428,1778,1804]" pageId="2" pageNumber="431" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="FB0036DAFFC64C22FF67E96FFD2BF8A4" blockId="2.[151,1436,1634,2020]" pageId="2" pageNumber="431">Abdomen distinctly convex, slightly narrower or broader than elytra, with a pair of small oval and sometimes indistinct tomentose spots (wing-folding patches) in middle of abdominal tergite IV; apical margin of abdominal tergite VII with very narrow palisade fringe.</paragraph>
|
||
<paragraph id="FB0036DAFFC64C22FF67E9FBFEE9F838" blockId="2.[151,1436,1634,2020]" pageId="2" pageNumber="431">
|
||
Male. Aedeagus short and wide, with very wide median lobe, strongly narrowed apically, with elongate accessory plates, short and moderately broad cylindrical parameres and very long flagellum spirally folded in middle portion (
|
||
<figureCitation id="63842A5FFFC64C22FF57E9B3FE9FF838" box="[247,316,1994,2020]" captionStart="FIGURES 9–10" captionStartId="4.[151,264,1964,1988]" captionTargetBox="[155,1432,637,1937]" captionTargetId="figure-227@4.[151,1436,633,1941]" captionTargetPageId="4" captionText="FIGURES 9–10. Aedeagus of Sinacrulia fortiterpunctata (holotype): 9–parameral view, 10–lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10199879" httpUri="https://zenodo.org/record/10199879/files/figure.png" pageId="2" pageNumber="431">Fig. 9</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="AFC06652FFC74C23FF37EBE2FF4FFA27" ID-DOI="http://doi.org/10.5281/zenodo.10199877" ID-Zenodo-Dep="10199877" httpUri="https://zenodo.org/record/10199877/files/figure.png" pageId="3" pageNumber="432" startId="3.[151,264,1435,1459]" targetBox="[154,1433,193,1408]" targetPageId="3" targetType="figure">
|
||
<paragraph id="FB0036DAFFC74C23FF37EBE2FF4FFA27" blockId="3.[151,1437,1435,1531]" pageId="3" pageNumber="432">
|
||
<emphasis id="C9CBEAC8FFC74C23FF37EBE2FE98FA6F" bold="true" box="[151,315,1435,1459]" pageId="3" pageNumber="432">FIGURES 3–8.</emphasis>
|
||
Mouthparts of
|
||
<taxonomicName id="3CBF4D59FFC74C23FE7AEBE2FD48FA6F" authorityName="Shavrin" authorityYear="2023" box="[474,747,1435,1459]" class="Insecta" family="Staphylinidae" genus="Sinacrulia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="432" phylum="Arthropoda" rank="species" species="fortiterpunctata">
|
||
<emphasis id="C9CBEAC8FFC74C23FE7AEBE2FD48FA6F" box="[474,747,1435,1459]" italics="true" pageId="3" pageNumber="432">Sinacrulia fortiterpunctata</emphasis>
|
||
</taxonomicName>
|
||
(holotype): 3–labrum (dorsal view), 4–left mandible (dorsal view), 5–right mandible (dorsal view), 6–labium (ventral view), 7–mentum (ventral view), 8–right maxilla (dorsal view). Scale bars: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="FB0036DAFFC74C23FF67E850FB3EF9BB" blockId="3.[151,1437,1577,2036]" pageId="3" pageNumber="432">
|
||
Female. Accessory abdominal sclerite weakly sclerotized, transverse and wide (
|
||
<figureCitation id="63842A5FFFC74C23FB9DE850FB33F998" box="[1085,1168,1577,1604]" captionStart="FIGURES 11–15" captionStartId="5.[151,264,1481,1505]" captionTargetBox="[155,1433,185,1453]" captionTargetId="figure-15@5.[151,1436,181,1457]" captionTargetPageId="5" captionText="FIGURES 11–15. Abdominal segments of Sinacrulia (Figs 11–12: S. fortiterpunctata (holotype), Figs. 13–15: S. sp.): 11–male tergite VIII (dorsal view), 12–male sternite VIII (ventral view), 13–female tergite VIII (dorsal view), 14–female sternite VIII (ventral view), 15–female genital segment (dorsal view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10199883" httpUri="https://zenodo.org/record/10199883/files/figure.png" pageId="3" pageNumber="432">Fig. 15</figureCitation>
|
||
). Gonocoxites wide and long, with relatively short styli and long apical setae (
|
||
<figureCitation id="63842A5FFFC74C23FD50E834FCE0F9B4" box="[752,835,1613,1640]" captionStart="FIGURES 11–15" captionStartId="5.[151,264,1481,1505]" captionTargetBox="[155,1433,185,1453]" captionTargetId="figure-15@5.[151,1436,181,1457]" captionTargetPageId="5" captionText="FIGURES 11–15. Abdominal segments of Sinacrulia (Figs 11–12: S. fortiterpunctata (holotype), Figs. 13–15: S. sp.): 11–male tergite VIII (dorsal view), 12–male sternite VIII (ventral view), 13–female tergite VIII (dorsal view), 14–female sternite VIII (ventral view), 15–female genital segment (dorsal view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10199883" httpUri="https://zenodo.org/record/10199883/files/figure.png" pageId="3" pageNumber="432">Fig. 15</figureCitation>
|
||
). Spermatheca not recognized.
|
||
</paragraph>
|
||
<paragraph id="FB0036DAFFC74C23FF67E80BFE55F957" blockId="3.[151,1437,1577,2036]" box="[199,502,1649,1676]" pageId="3" pageNumber="432">Immature stages unknown.</paragraph>
|
||
<paragraph id="FB0036DAFFC74C23FF67E8ECFCA5F96C" blockId="3.[151,1437,1577,2036]" box="[199,774,1685,1712]" pageId="3" pageNumber="432">
|
||
<emphasis id="C9CBEAC8FFC74C23FF67E8ECFE30F973" bold="true" box="[199,403,1685,1711]" pageId="3" pageNumber="432">Species included.</emphasis>
|
||
<taxonomicName id="3CBF4D59FFC74C23FE3AE8ECFD64F96C" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[410,711,1685,1712]" class="Insecta" family="Staphylinidae" genus="Sinacrulia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="432" phylum="Arthropoda" rank="species" species="fortiterpunctata" status="sp. nov.">
|
||
<emphasis id="C9CBEAC8FFC74C23FE3AE8ECFD64F96C" box="[410,711,1685,1712]" italics="true" pageId="3" pageNumber="432">Sinacrulia fortiterpunctata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C9CBEAC8FFC74C23FD6EE8EFFCA5F96C" bold="true" box="[718,774,1686,1712]" pageId="3" pageNumber="432">
|
||
<taxonomicNameLabel id="D2F857B3FFC74C23FD6EE8EFFCA5F96C" box="[718,774,1686,1712]" pageId="3" pageNumber="432" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B3A56551FFC74C24FF67E8C0FE07FDE1" lastPageId="4" lastPageNumber="433" pageId="3" pageNumber="432" type="diagnosis">
|
||
<paragraph id="FB0036DAFFC74C24FF67E8C0FCFCFE29" blockId="3.[151,1437,1577,2036]" lastBlockId="4.[151,1437,151,609]" lastPageId="4" lastPageNumber="433" pageId="3" pageNumber="432">
|
||
<emphasis id="C9CBEAC8FFC74C23FF67E8C0FE73F90F" bold="true" box="[199,464,1721,1747]" pageId="3" pageNumber="432">Differential diagnosis.</emphasis>
|
||
Based on the general shapes of the body and the aedeagus, the hind tarsomeres 1–4 together distinctly shorter than apical tarsomere and transverse preapical antennomeres,
|
||
<taxonomicName id="3CBF4D59FFC74C23FBDDE8A4FB44F92B" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[1149,1255,1757,1783]" class="Insecta" family="Staphylinidae" genus="Sinaculia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="432" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="C9CBEAC8FFC74C23FBDDE8A4FB44F92B" box="[1149,1255,1757,1783]" italics="true" pageId="3" pageNumber="432">Sinaculia</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C9CBEAC8FFC74C23FB4FE8A7FA93F924" bold="true" box="[1263,1328,1758,1784]" pageId="3" pageNumber="432">
|
||
<taxonomicNameLabel id="D2F857B3FFC74C23FB4FE8A7FA93F924" box="[1263,1328,1758,1784]" pageId="3" pageNumber="432" rank="genus">gen.n</taxonomicNameLabel>
|
||
</emphasis>
|
||
undoubtedly belongs to the tribe
|
||
<taxonomicName id="3CBF4D59FFC74C23FE0DE978FDB7F8C7" authorityName="Thomson" authorityYear="1858" box="[429,532,1793,1819]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="432" phylum="Arthropoda" rank="tribe" tribe="Omaliini">Omaliini</taxonomicName>
|
||
. Regarding the general shape of small and convex body, with the presence of the distinctly crenulate lateral margins of the pronotum,
|
||
<taxonomicName id="3CBF4D59FFC74C23FD46E95CFCF8F8E3" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[742,859,1829,1855]" class="Insecta" family="Staphylinidae" genus="Sinacrulia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="432" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="C9CBEAC8FFC74C23FD46E95CFCF8F8E3" box="[742,859,1829,1855]" italics="true" pageId="3" pageNumber="432">Sinacrulia</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C9CBEAC8FFC74C23FCC2E95FFC0AF89C" bold="true" box="[866,937,1830,1856]" pageId="3" pageNumber="432">
|
||
<taxonomicNameLabel id="D2F857B3FFC74C23FCC2E95FFC0AF89C" box="[866,937,1830,1856]" pageId="3" pageNumber="432" rank="genus">gen.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be compared with several genera which have recently been reviewed by
|
||
<bibRefCitation id="9F2E4B2BFFC74C23FE5CE930FCBBF8B8" author="Shavrin, A. V. & Zanetti, A." box="[508,792,1865,1892]" pageId="3" pageNumber="432" pagination="1 - 63" refId="ref4812" refString="Shavrin, A. V. & Zanetti, A. (2020) Revision of Acrulia Thomson, 1858 and Dialycera Ganglbauer, 1895, with taxonomic notes on related genera and description of a new genus from the Caucasus (Coleoptera: Staphylinidae: Omaliinae: Omaliini). Zootaxa, 4800 (1), 1 - 63. https: // doi. org / 10.11646 / zootaxa. 4800.1.1" type="journal article" year="2020">Shavrin & Zanetti (2020)</bibRefCitation>
|
||
: Palaearctic
|
||
<taxonomicName id="3CBF4D59FFC74C23FC03E930FB08F8B8" authority="Thomson, 1858" authorityName="Thomson" authorityYear="1858" box="[931,1195,1865,1892]" class="Insecta" family="Staphylinidae" genus="Acrulia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="432" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C9CBEAC8FFC74C23FC03E930FC54F8BF" box="[931,1015,1865,1891]" italics="true" pageId="3" pageNumber="432">Acrulia</emphasis>
|
||
Thomson, 1858
|
||
</taxonomicName>
|
||
(three species), western palaearctic
|
||
<taxonomicName id="3CBF4D59FFC74C23FEE3E914FD20F854" authority="Ganglbauer, 1895" authorityName="Ganglbauer" authorityYear="1895" box="[323,643,1901,1928]" class="Insecta" family="Staphylinidae" genus="Dialycera" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="432" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C9CBEAC8FFC74C23FEE3E914FE10F85B" box="[323,435,1901,1927]" italics="true" pageId="3" pageNumber="432">Dialycera</emphasis>
|
||
Ganglbauer, 1895
|
||
</taxonomicName>
|
||
(three species), Holarctic
|
||
<taxonomicName id="3CBF4D59FFC74C23FC0CE914FB64F854" authority="Zerche, 2003" authorityName="Zerche" authorityYear="2003" box="[940,1223,1901,1928]" class="Insecta" family="Staphylinidae" genus="Acruliopsis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="432" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C9CBEAC8FFC74C23FC0CE914FB8FF85B" box="[940,1068,1901,1927]" italics="true" pageId="3" pageNumber="432">Acruliopsis</emphasis>
|
||
<bibRefCitation id="9F2E4B2BFFC74C23FB93E914FB64F854" author="Zerche, L." box="[1075,1223,1901,1928]" pageId="3" pageNumber="432" pagination="297 - 316" refId="ref4966" refString="Zerche, L. (2003) Acruliopsis gen. n. aus der Ostpalaarktis (Coleoptera, Staphylinidae, Omaliinae, Omaliini). Beitrage zur Entomologie, 53 (2), 297 - 316. https: // doi. org / 10.21248 / contrib. entomol. 53.2.297 - 316" type="journal article" year="2003">Zerche, 2003</bibRefCitation>
|
||
</taxonomicName>
|
||
(five species), and Holarctic
|
||
<taxonomicName id="3CBF4D59FFC74C23FEA8E9E8FDE4F870" authority="Thomson, 1858" authorityName="Thomson" authorityYear="1858" box="[264,583,1937,1964]" class="Insecta" family="Staphylinidae" genus="Hapalaraea" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="432" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C9CBEAC8FFC74C23FEA8E9E8FE2CF877" box="[264,399,1937,1963]" italics="true" pageId="3" pageNumber="432">Hapalaraea</emphasis>
|
||
Thomson, 1858
|
||
</taxonomicName>
|
||
(six species). From all these genera
|
||
<taxonomicName id="3CBF4D59FFC74C23FC49E9E8FBFDF877" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[1001,1118,1937,1963]" class="Insecta" family="Staphylinidae" genus="Sinacrulia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="432" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="C9CBEAC8FFC74C23FC49E9E8FBFDF877" box="[1001,1118,1937,1963]" italics="true" pageId="3" pageNumber="432">Sinacrulia</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C9CBEAC8FFC74C23FBC6E9EBFB0EF870" bold="true" box="[1126,1197,1938,1964]" pageId="3" pageNumber="432">
|
||
<taxonomicNameLabel id="D2F857B3FFC74C23FBC6E9EBFB0EF870" box="[1126,1197,1938,1964]" pageId="3" pageNumber="432" rank="genus">gen.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be distinguished by the following morphological features: 1) apical segment of the maxillary palpi very long and narrow, slightly less than three times as long as moderately broad penultimate segment, 2) the presence of indistinct pronotal impressions (
|
||
<emphasis id="C9CBEAC8FFC04C24FF01EEEEFF0CFF6D" box="[161,175,151,177]" italics="true" pageId="4" pageNumber="433">S</emphasis>
|
||
. sp.), 3) the shape of very broad median lobe, from widest middle strongly narrowed apically, 4) the shape of the very short parameres, reaching middle of the median lobe, 5) the presence of elongate accessory plates (“accessory lamella” according
|
||
<bibRefCitation id="9F2E4B2BFFC04C24FED5EEA6FD82FF25" author="Zanetti, A." box="[373,545,223,249]" pageId="4" pageNumber="433" pagination="1 - 472" refId="ref4886" refString="Zanetti, A. (1987) Coleoptera. Staphylinidae. Omaliinae. In: Fauna d'Italia. Vol. 25. Calderini, Bologna, pp. i - xii + 1 - 472." type="book chapter" year="1987">Zanetti (1987))</bibRefCitation>
|
||
rounded apically at level of middle part of the median lobe, 6) the missing of sclerotized spines of internal sac, 7) the presence of very long and strong flagellum. Based on the general shape of the transverse head evenly convex in middle, without shield-like median elevation and with moderately long temples, the shape of the slightly elongate antennomere 5, and the strong punctation of the forebody,
|
||
<taxonomicName id="3CBF4D59FFC04C24FB51EF32FAC5FEB9" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[1265,1382,331,357]" class="Insecta" family="Staphylinidae" genus="Sinacrulia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="433" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="C9CBEAC8FFC04C24FB51EF32FAC5FEB9" box="[1265,1382,331,357]" italics="true" pageId="4" pageNumber="433">Sinacrulia</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C9CBEAC8FFC04C24FACBEF32FF0DFE55" bold="true" pageId="4" pageNumber="433">
|
||
<taxonomicNameLabel id="D2F857B3FFC04C24FACBEF32FF0DFE55" pageId="4" pageNumber="433" rank="genus">gen. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is somewhat similar to the genus
|
||
<taxonomicName id="3CBF4D59FFC04C24FD8BEF16FD08FE55" authorityName="Zerche" authorityYear="2003" box="[555,683,367,393]" class="Insecta" family="Staphylinidae" genus="Acruliopsis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="433" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C9CBEAC8FFC04C24FD8BEF16FD08FE55" box="[555,683,367,393]" italics="true" pageId="4" pageNumber="433">Acruliopsis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="9F2E4B2BFFC04C24FD19EF16FCE9FE55" author="Zerche, L." box="[697,842,367,393]" pageId="4" pageNumber="433" pagination="297 - 316" refId="ref4966" refString="Zerche, L. (2003) Acruliopsis gen. n. aus der Ostpalaarktis (Coleoptera, Staphylinidae, Omaliinae, Omaliini). Beitrage zur Entomologie, 53 (2), 297 - 316. https: // doi. org / 10.21248 / contrib. entomol. 53.2.297 - 316" type="journal article" year="2003">Zerche 2003</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="9F2E4B2BFFC04C24FCF5EF16FC4DFE55" author="Shavrin, A. V." box="[853,1006,367,393]" pageId="4" pageNumber="433" pagination="125 - 130" refId="ref4560" refString="Shavrin, A. V. (2013) Redescription of Acruliopsis tumidula (Maklin, 1853) comb. n., with new data on A. ussuriensis Zerche, 2003 (Coleoptera: Staphylinidae: Omaliinae). Baltic Journal of Coleopteroogy, 13 (2), 125 - 130." type="journal article" year="2013">Shavrin 2013</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="9F2E4B2BFFC04C24FC59EF16FAA4FE55" author="Shavrin, A. V. & Zanetti, A." box="[1017,1287,367,393]" pageId="4" pageNumber="433" pagination="1 - 63" refId="ref4812" refString="Shavrin, A. V. & Zanetti, A. (2020) Revision of Acrulia Thomson, 1858 and Dialycera Ganglbauer, 1895, with taxonomic notes on related genera and description of a new genus from the Caucasus (Coleoptera: Staphylinidae: Omaliinae: Omaliini). Zootaxa, 4800 (1), 1 - 63. https: // doi. org / 10.11646 / zootaxa. 4800.1.1" type="journal article" year="2020">Shavrin & Zanetti 2020</bibRefCitation>
|
||
), from which it differs by the missing of pubescence of the forebody, finer crenulation of lateral margins of the pronotum, longer elytra and different morphology of the aedeagus. From all genera of
|
||
<taxonomicName id="3CBF4D59FFC04C24FC30EFCEFC57FE0D" authorityName="Thomson" authorityYear="1858" box="[912,1012,439,465]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="433" phylum="Arthropoda" rank="tribe" tribe="Omaliini">Omaliini</taxonomicName>
|
||
it can be distinguished by the unusual shape of the aedeagus, and the presence of very long flagellum.
|
||
</paragraph>
|
||
<paragraph id="FB0036DAFFC04C24FF67EF86FE07FDE1" blockId="4.[151,1437,151,609]" pageId="4" pageNumber="433">
|
||
<emphasis id="C9CBEAC8FFC04C24FF67EF86FE26FDC4" bold="true" box="[199,389,510,537]" pageId="4" pageNumber="433">Natural history.</emphasis>
|
||
The species of
|
||
<taxonomicName id="3CBF4D59FFC04C24FD92EF86FD04FDC5" authorityName="Shavrin" authorityYear="2023" box="[562,679,511,537]" class="Insecta" family="Staphylinidae" genus="Sinacrulia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="433" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C9CBEAC8FFC04C24FD92EF86FD04FDC5" box="[562,679,511,537]" italics="true" pageId="4" pageNumber="433">Sinacrulia</emphasis>
|
||
</taxonomicName>
|
||
were collected at high elevations from
|
||
<quantity id="3C479B3FFFC04C24FBFDEF86FAAFFDC5" box="[1117,1292,510,537]" metricMagnitude="3" metricUnit="m" metricValue="3.01" metricValueMax="3.05" metricValueMin="2.9699999999999998" pageId="4" pageNumber="433" unit="m" value="3010.0" valueMax="3050.0" valueMin="2970.0">2970 to 3050 m</quantity>
|
||
a.s.l. by sifting of litter and mosses.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B3A56551FFC04C24FF67EC3FFDDDFDBD" box="[199,638,582,609]" pageId="4" pageNumber="433" type="distribution">
|
||
<paragraph id="FB0036DAFFC04C24FF67EC3FFDDDFDBD" blockId="4.[151,1437,151,609]" box="[199,638,582,609]" pageId="4" pageNumber="433">
|
||
<emphasis id="C9CBEAC8FFC04C24FF67EC3FFEC2FDBC" bold="true" box="[199,353,582,608]" pageId="4" pageNumber="433">Distribution.</emphasis>
|
||
<collectingCountry id="83A8764AFFC04C24FEC7EC3EFE0DFDBD" box="[359,430,583,609]" name="China" pageId="4" pageNumber="433">China</collectingCountry>
|
||
:
|
||
<collectingRegion id="397BF838FFC04C24FE18EC3EFDADFDBD" box="[440,526,583,609]" country="China" name="Yunnan" pageId="4" pageNumber="433">Yunnan</collectingRegion>
|
||
(
|
||
<figureCitation id="63842A5FFFC04C24FDBDEC3EFDD3FDBD" box="[541,624,583,609]" captionStart="FIGURE 17" captionStartId="8.[151,250,916,940]" captionTargetBox="[151,1436,181,893]" captionTargetId="figure-18@8.[151,1436,181,893]" captionTargetPageId="8" captionText="FIGURE 17. Distribution of Sinacrulia in China (black circle: S. fortiterpunctata, white circle: S. sp.)." figureDoi="http://doi.org/10.5281/zenodo.10199887" httpUri="https://zenodo.org/record/10199887/files/figure.png" pageId="4" pageNumber="433">Fig. 17</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="AFC06652FFC04C24FF37E9D5FA2AF818" ID-DOI="http://doi.org/10.5281/zenodo.10199879" ID-Zenodo-Dep="10199879" box="[151,1417,1964,1989]" httpUri="https://zenodo.org/record/10199879/files/figure.png" pageId="4" pageNumber="433" startId="4.[151,264,1964,1988]" targetBox="[155,1432,637,1937]" targetPageId="4" targetType="figure">
|
||
<paragraph id="FB0036DAFFC04C24FF37E9D5FA2AF818" blockId="4.[151,1417,1964,1989]" box="[151,1417,1964,1989]" pageId="4" pageNumber="433">
|
||
<emphasis id="C9CBEAC8FFC04C24FF37E9D5FEE4F818" bold="true" box="[151,327,1964,1988]" pageId="4" pageNumber="433">FIGURES 9–10.</emphasis>
|
||
Aedeagus of
|
||
<taxonomicName id="3CBF4D59FFC04C24FE73E9D5FD40F819" authorityName="Shavrin" authorityYear="2023" box="[467,739,1964,1989]" class="Insecta" family="Staphylinidae" genus="Sinacrulia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="433" phylum="Arthropoda" rank="species" species="fortiterpunctata">
|
||
<emphasis id="C9CBEAC8FFC04C24FE73E9D5FD40F819" box="[467,739,1964,1989]" italics="true" pageId="4" pageNumber="433">Sinacrulia fortiterpunctata</emphasis>
|
||
</taxonomicName>
|
||
(holotype): 9–parameral view, 10–lateral view. Scale bar: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="AFC06652FFC14C25FF37EBB0FC05F9F5" ID-DOI="http://doi.org/10.5281/zenodo.10199883" ID-Zenodo-Dep="10199883" httpUri="https://zenodo.org/record/10199883/files/figure.png" pageId="5" pageNumber="434" startId="5.[151,264,1481,1505]" targetBox="[155,1433,185,1453]" targetPageId="5" targetType="figure">
|
||
<paragraph id="FB0036DAFFC14C25FF37EBB0FC05F9F5" blockId="5.[151,1437,1481,1578]" pageId="5" pageNumber="434">
|
||
<emphasis id="C9CBEAC8FFC14C25FF37EBB0FEF1FA3D" bold="true" box="[151,338,1481,1505]" pageId="5" pageNumber="434">FIGURES 11–15.</emphasis>
|
||
Abdominal segments of
|
||
<taxonomicName id="3CBF4D59FFC14C25FDEEEBB0FD1BFA3D" authorityName="Shavrin" authorityYear="2023" box="[590,696,1481,1505]" class="Insecta" family="Staphylinidae" genus="Sinacrulia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="434" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C9CBEAC8FFC14C25FDEEEBB0FD1BFA3D" box="[590,696,1481,1505]" italics="true" pageId="5" pageNumber="434">Sinacrulia</emphasis>
|
||
</taxonomicName>
|
||
(Figs 11–12:
|
||
<taxonomicName id="3CBF4D59FFC14C25FCE3EBB0FC58FA3D" authorityName="Shavrin" authorityYear="2023" box="[835,1019,1481,1505]" class="Insecta" family="Staphylinidae" genus="Sinacrulia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="434" phylum="Arthropoda" rank="species" species="fortiterpunctata">
|
||
<emphasis id="C9CBEAC8FFC14C25FCE3EBB0FC58FA3D" box="[835,1019,1481,1505]" italics="true" pageId="5" pageNumber="434">S. fortiterpunctata</emphasis>
|
||
</taxonomicName>
|
||
(holotype), Figs. 13–15:
|
||
<emphasis id="C9CBEAC8FFC14C25FB5BEBB0FAACFA3C" box="[1275,1295,1481,1504]" italics="true" pageId="5" pageNumber="434">S.</emphasis>
|
||
sp.): 11–male tergite VIII (dorsal view), 12–male sternite VIII (ventral view), 13–female tergite VIII (dorsal view), 14–female sternite VIII (ventral view), 15–female genital segment (dorsal view). Scale bars: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="B3A56551FFC14C25FF67E82EFD4DF949" pageId="5" pageNumber="434" type="etymology">
|
||
<paragraph id="FB0036DAFFC14C25FF67E82EFD4DF949" blockId="5.[151,1437,1623,2010]" pageId="5" pageNumber="434">
|
||
<emphasis id="C9CBEAC8FFC14C25FF67E82EFEEDF9AD" bold="true" box="[199,334,1623,1649]" pageId="5" pageNumber="434">Etymology.</emphasis>
|
||
The name of the new genus is derived from the prefix
|
||
<emphasis id="C9CBEAC8FFC14C25FC07E821FC7AF9AE" box="[935,985,1624,1650]" italics="true" pageId="5" pageNumber="434">Sino</emphasis>
|
||
-, referring to
|
||
<collectingCountry id="83A8764AFFC14C25FBD2E82EFB1BF9AD" box="[1138,1208,1623,1649]" name="China" pageId="5" pageNumber="434">China</collectingCountry>
|
||
, and the genus name
|
||
<taxonomicName id="3CBF4D59FFC14C25FF37E802FF48F949" authorityName="Thomson" authorityYear="1858" box="[151,235,1659,1685]" class="Insecta" family="Staphylinidae" genus="Acrulia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="434" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C9CBEAC8FFC14C25FF37E802FF48F949" box="[151,235,1659,1685]" italics="true" pageId="5" pageNumber="434">Acrulia</emphasis>
|
||
</taxonomicName>
|
||
, one of the relative taxa of the tribe
|
||
<taxonomicName id="3CBF4D59FFC14C25FD23E802FD49F949" authorityName="Thomson" authorityYear="1858" box="[643,746,1659,1685]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="434" phylum="Arthropoda" rank="tribe" tribe="Omaliini">Omaliini</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B3A56551FFC14C25FF67E8E6FA37F805" pageId="5" pageNumber="434" type="discussion">
|
||
<paragraph id="FB0036DAFFC14C25FF67E8E6FA37F805" blockId="5.[151,1437,1623,2010]" pageId="5" pageNumber="434">
|
||
<emphasis id="C9CBEAC8FFC14C25FF67E8E6FE98F965" bold="true" box="[199,315,1695,1721]" pageId="5" pageNumber="434">Remarks.</emphasis>
|
||
The general structure of the aedeagus of
|
||
<taxonomicName id="3CBF4D59FFC14C25FCACE8E6FC22F965" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[780,897,1695,1721]" class="Insecta" family="Staphylinidae" genus="Sinacrulia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="434" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="C9CBEAC8FFC14C25FCACE8E6FC22F965" box="[780,897,1695,1721]" italics="true" pageId="5" pageNumber="434">Sinacrulia</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C9CBEAC8FFC14C25FC28E8E6FC6CF965" bold="true" box="[904,975,1695,1721]" pageId="5" pageNumber="434">
|
||
<taxonomicNameLabel id="D2F857B3FFC14C25FC28E8E6FC6CF965" box="[904,975,1695,1721]" pageId="5" pageNumber="434" rank="genus">gen.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is somewhat similar to that of some species of
|
||
<taxonomicName id="3CBF4D59FFC14C25FF49E8BAFD89F901" authority="Gravenhorst, 1802" authorityName="Gravenhorst" authorityYear="1802" box="[233,554,1731,1757]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="434" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C9CBEAC8FFC14C25FF49E8BAFEF2F901" box="[233,337,1731,1757]" italics="true" pageId="5" pageNumber="434">Omalium</emphasis>
|
||
Gravenhorst, 1802
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="3CBF4D59FFC14C25FDC1E8BAFC2BF902" authority="Thomson 1858" authorityName="Thomson" authorityYear="1858" box="[609,904,1731,1758]" class="Insecta" family="Staphylinidae" genus="Acrolocha" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="434" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C9CBEAC8FFC14C25FDC1E8BAFD76F901" box="[609,725,1731,1757]" italics="true" pageId="5" pageNumber="434">Acrolocha</emphasis>
|
||
Thomson 1858
|
||
</taxonomicName>
|
||
. Almost all species of these genera have developed accessory plates, sometimes strongly sclerotized and protruded apicad between elongate median lobe, and also have similar shapes of the parameres (usually more complicated in many species
|
||
<taxonomicName id="3CBF4D59FFC14C25FB8BE972FB30F8F9" authorityName="Gravenhorst" authorityYear="1802" box="[1067,1171,1803,1829]" class="Insecta" family="Staphylinidae" genus="Omalium" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="434" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C9CBEAC8FFC14C25FB8BE972FB30F8F9" box="[1067,1171,1803,1829]" italics="true" pageId="5" pageNumber="434">Omalium</emphasis>
|
||
</taxonomicName>
|
||
). The presence of large and long flagellum in
|
||
<taxonomicName id="3CBF4D59FFC14C25FE36E956FDA8F895" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[406,523,1839,1865]" class="Insecta" family="Staphylinidae" genus="Sinacrulia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="434" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="C9CBEAC8FFC14C25FE36E956FDA8F895" box="[406,523,1839,1865]" italics="true" pageId="5" pageNumber="434">Sinacrulia</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C9CBEAC8FFC14C25FDB3E956FDF8F895" bold="true" box="[531,603,1839,1865]" pageId="5" pageNumber="434">
|
||
<taxonomicNameLabel id="D2F857B3FFC14C25FDB3E956FDF8F895" box="[531,603,1839,1865]" pageId="5" pageNumber="434" rank="genus">gen.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is unique. Only species of some genera have short narrow flagellum not spirally folded in basal portion of the aedeagus (e.g.
|
||
<taxonomicName id="3CBF4D59FFC14C25FD54E92AFBC8F8B1" authority="Berhauer, 1903" authorityName="Berhauer" authorityYear="1903" box="[756,1131,1875,1901]" class="Insecta" family="Staphylinidae" genus="Carcinocephalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="434" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C9CBEAC8FFC14C25FD54E92AFC11F8B1" box="[756,946,1875,1901]" italics="true" pageId="5" pageNumber="434">Carcinocephalus</emphasis>
|
||
Berhauer, 1903
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="3CBF4D59FFC14C25FBD8E92AFA3FF8B1" authority="Jeannel, 1940" authorityName="Jeannel" authorityYear="1940" box="[1144,1436,1875,1901]" class="Insecta" family="Staphylinidae" genus="Omaliopsis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="434" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C9CBEAC8FFC14C25FBD8E92AFB5BF8B1" box="[1144,1272,1875,1901]" italics="true" pageId="5" pageNumber="434">Omaliopsis</emphasis>
|
||
Jeannel, 1940
|
||
</taxonomicName>
|
||
(some species),
|
||
<taxonomicName id="3CBF4D59FFC14C25FEE9E90EFDC6F84D" authority="Heer, 1839" authorityName="Heer" authorityYear="1839" box="[329,613,1911,1938]" class="Insecta" family="Staphylinidae" genus="Phloeonomus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="434" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C9CBEAC8FFC14C25FEE9E90EFE42F84D" box="[329,481,1911,1937]" italics="true" pageId="5" pageNumber="434">Phloeonomus</emphasis>
|
||
Heer, 1839
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="3CBF4D59FFC14C25FDCFE90EFC23F84D" authority="Fauvel, 1904" authorityName="Fauvel" authorityYear="1904" box="[623,896,1911,1937]" class="Insecta" family="Staphylinidae" genus="Tetradelus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="434" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C9CBEAC8FFC14C25FDCFE90EFD47F84D" box="[623,740,1911,1937]" italics="true" pageId="5" pageNumber="434">Tetradelus</emphasis>
|
||
Fauvel, 1904
|
||
</taxonomicName>
|
||
). Usually, the very long flagellum with the spiral in basal portion of the aedeagus can be observed in some genera of the tribe
|
||
<taxonomicName id="3CBF4D59FFC14C25FC4DE9E2FAE1F86A" authority="Thomson, 1859" authorityName="Thomson" authorityYear="1859" box="[1005,1346,1947,1974]" class="Insecta" family="Staphylinidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="434" phylum="Arthropoda" rank="tribe" tribe="Anthophagini">Anthophagini Thomson, 1859</taxonomicName>
|
||
:
|
||
<taxonomicName id="3CBF4D59FFC14C25FAECE9E5FE12F805" authority="Tronquet, 1982" authorityName="Tronquet" authorityYear="1982" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="434" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C9CBEAC8FFC14C25FAECE9E5FF5DF805" italics="true" pageId="5" pageNumber="434">Hygrodromicus</emphasis>
|
||
Tronquet, 1982
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="9F2E4B2BFFC14C25FE1FE9C6FDC4F805" author="Shavrin, A. V." box="[447,615,1983,2009]" pageId="5" pageNumber="434" pagination="335 - 348" refId="ref4672" refString="Shavrin, A. V. (2021 b) New taxonomic and faunistic data of the genus Hygrodromicus, 1981 of the Himalayan Region (Coleoptera: Staphylinidaae: Omaliinae: Anthophagini). In: Hartmann, M., Barclay, M. & Weipert, J. (Eds.), Biodiversitat und Naturausstattung im Himalaya VII. Verein der Freunde und Forderer des Naturkundmuseums Erfurt e. V., Erfurt, pp. 335 - 348." type="book chapter" year="2021">Shavrin 2021b</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="9F2E4B2BFFC14C25FDD3E9C6FD0CF805" author="Shavrin, A. V." box="[627,687,1983,2009]" pageId="5" pageNumber="434" pagination="1 - 15" refId="ref4756" refString="Shavrin, A. V. (2022) New species and records of Hygrodromicus Tronquet, 1981 from China (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Journal of Insect Biodiversity, 31 (1), 1 - 15. https: // doi. org / 10.12976 / jib / 2022.31.1.1" type="journal article" year="2022">2022</bibRefCitation>
|
||
),
|
||
<taxonomicName id="3CBF4D59FFC14C25FD62E9C6FC53F805" authority="Luze, 1903" authorityName="Luze" authorityYear="1903" box="[706,1008,1983,2010]" class="Insecta" family="Staphylinidae" genus="Trichodromeus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="434" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C9CBEAC8FFC14C25FD62E9C6FCC8F805" box="[706,875,1983,2009]" italics="true" pageId="5" pageNumber="434">Trichodromeus</emphasis>
|
||
Luze, 1903
|
||
</taxonomicName>
|
||
(
|
||
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |