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<document ID-DOI="http://dx.doi.org/10.3897/jhr.55.11482" ID-GBIF-Dataset="310c95ad-8bf1-4955-acfb-c1368c0222a7" ID-GBIF-Taxon="128834706" ID-Pensoft-Pub="1314-2607-55-139" ID-Pensoft-UUID="FFF5667EAD5E7A6FFFEC2B6CFFB7D800" ID-ZBK="0E1B0A14F8714CD89F2910B255677621" ID-Zenodo-Dep="575140" ID-ZooBank="0E1B0A14F8714CD89F2910B255677621" LSID="urn:lsid:zoobank.org:act:8C723EBE-7AFA-4B38-8671-285CF7E2B443" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1314-2607-55-139" ModsDocOrigin="Journal of Hymenoptera Research 55" ModsDocTitle="Biting the bullet: revisionary notes on the Oraseminae of the Old World (Hymenoptera, Chalcidoidea, Eucharitidae)" checkinTime="1493464749952" checkinUser="pensoft" docAuthor="Burks, Roger A., Heraty, John M., Mottern, Jason, Dominguez, Chrysalyn &amp; Heacox, Scott" docDate="2017" docId="598DCAC562107936D3BA8D9E8729D9B1" docLanguage="en" docName="JourHymenoptRes 55: 139-188" docOrigin="Journal of Hymenoptera Research 55" docPubDate="2017-04-28" docSource="http://dx.doi.org/10.3897/jhr.55.11482" docTitle="Cymosema waterworthae Burks &amp; Mottern 2017, sp. n." docType="treatment" docUuid="8C723EBE-7AFA-4B38-8671-285CF7E2B443" docUuidSource="ZooBank" docVersion="4" id="FFF5667EAD5E7A6FFFEC2B6CFFB7D800" lastPageId="17" lastPageNumber="156" masterDocId="FFF5667EAD5E7A6FFFEC2B6CFFB7D800" masterDocTitle="Biting the bullet: revisionary notes on the Oraseminae of the Old World (Hymenoptera, Chalcidoidea, Eucharitidae)" masterLastPageNumber="188" masterPageNumber="139" pageId="13" pageNumber="152" updateTime="1643484763995" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Biting the bullet: revisionary notes on the Oraseminae of the Old World (Hymenoptera, Chalcidoidea, Eucharitidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Burks, Roger A.</mods:namePart>
<mods:affiliation>Department of Entomology, University of California, Riverside, CA 92521</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Heraty, John M.</mods:namePart>
<mods:affiliation>Department of Entomology, University of California, Riverside, CA 92521</mods:affiliation>
<mods:nameIdentifier type="email">john.heraty@ucr.edu</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mottern, Jason</mods:namePart>
<mods:affiliation>Department of Entomology, University of California, Riverside, CA 92521 &amp; USDA Systematic Entomology Laboratory, Washington, DC 20013 - 7012</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dominguez, Chrysalyn</mods:namePart>
<mods:affiliation>Department of Entomology, University of California, Riverside, CA 92521</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Heacox, Scott</mods:namePart>
<mods:affiliation>Department of Entomology, University of California, Riverside, CA 92521</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Journal of Hymenoptera Research</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2017</mods:date>
<mods:detail type="pubDate">
<mods:number>2017-04-28</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>55</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>139</mods:start>
<mods:end>188</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/jhr.55.11482</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/jhr.55.11482</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2607-55-139</mods:identifier>
<mods:identifier type="ZooBank">0E1B0A14F8714CD89F2910B255677621</mods:identifier>
<mods:identifier type="Pensoft-UUID">FFF5667EAD5E7A6FFFEC2B6CFFB7D800</mods:identifier>
<mods:identifier type="Zenodo-Dep">575140</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="128834706" LSID="urn:lsid:zoobank.org:act:8C723EBE-7AFA-4B38-8671-285CF7E2B443" httpUri="http://treatment.plazi.org/id/598DCAC562107936D3BA8D9E8729D9B1" lastPageId="17" lastPageNumber="156" pageId="13" pageNumber="152">
<subSubSection pageId="13" pageNumber="152" type="nomenclature">
<paragraph pageId="13" pageNumber="152">
<taxonomicName LSID="http://zoobank.org/8C723EBE-7AFA-4B38-8671-285CF7E2B443" authority="Burks &amp; Mottern" authorityName="Burks &amp; Mottern" authorityYear="2017" class="Insecta" family="Eucharitidae" genus="Cymosema" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cymosema waterworthae" order="Hymenoptera" pageId="13" pageNumber="152" phylum="Arthropoda" rank="species" species="waterworthae" status="sp. n.">Cymosema waterworthae Burks &amp; Mottern</taxonomicName>
<taxonomicNameLabel pageId="13" pageNumber="152">sp. n.</taxonomicNameLabel>
<figureCitation captionStart="Figures 2126" captionStartId="F5" captionText="Figures 21 - 26. Cymosema waterworthae. Holotype female: 21 head, anterior 22 antennae, lateral 23 mesosoma, lateral 24 mesosoma, dorsal 25 petiole, lateral 26 ovipositor. pet = petiole." figureDoi="10.3897/jhr.55.11482.figures21-26" httpUri="https://binary.pensoft.net/fig/134104" pageId="13" pageNumber="152">Figs 21-26</figureCitation>
<figureCitation captionStart="Figure 27" captionStartId="F6" captionText="Figure 27. Cymosema waterworthae. Holotype female: fore wing. arrow points to end of postmarginal vein." figureDoi="10.3897/jhr.55.11482.figure27" httpUri="https://binary.pensoft.net/fig/134105" pageId="13" pageNumber="152">, 27</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="153" type="etymology">
<paragraph pageId="14" pageNumber="153">
<pageBreakToken pageId="14" pageNumber="153" start="start">Etymology</pageBreakToken>
.
</paragraph>
<paragraph pageId="14" pageNumber="153">Named in honor of the collector, Rebeccah Waterworth.</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="153" type="diagnosis">
<paragraph pageId="14" pageNumber="153">Diagnosis.</paragraph>
<paragraph pageId="14" pageNumber="153">Mandibles bidentate and overlapping, teeth of equal length and projecting medially. Funiculars cylindrical. Frons weakly reticulate. Submarginal vein bare, fore wing without marginal fringe.</paragraph>
</subSubSection>
<subSubSection lastPageId="16" lastPageNumber="155" pageId="15" pageNumber="154" type="female">
<paragraph pageId="15" pageNumber="154">
<pageBreakToken pageId="15" pageNumber="154" start="start">Female</pageBreakToken>
.
</paragraph>
<paragraph pageId="15" pageNumber="154">Length 1.4 mm. Head, mesosoma, and metasoma brown with metallic luster. Scape yellowish-brown, pedicel and flagellum brown. Legs brown, with tibial apices, and first four tarsomeres pale brown. Fore wing hyaline; venation pale brown, setae dark. Gaster brown with faint metallic luster.</paragraph>
<paragraph lastPageId="16" lastPageNumber="155" pageId="15" pageNumber="154">
<emphasis italics="true" pageId="15" pageNumber="154">Head</emphasis>
(Figs
<figureCitation captionStart="Figures 2126" captionStartId="F5" captionText="Figures 21 - 26. Cymosema waterworthae. Holotype female: 21 head, anterior 22 antennae, lateral 23 mesosoma, lateral 24 mesosoma, dorsal 25 petiole, lateral 26 ovipositor. pet = petiole." figureDoi="10.3897/jhr.55.11482.figures21-26" httpUri="https://binary.pensoft.net/fig/134104" pageId="15" pageNumber="154">21-22</figureCitation>
) transverse, 1.6
<normalizedToken originalValue="×">x</normalizedToken>
as broad as high. Head receding ventral to toruli; face reticulate, including supraclypeal area; clypeus shallowly transversely rugose-reticulate, anteclypeus present; vertex not carinate. Mandibles bidentate, both teeth projecting medially, narrow and only weakly curved (Fig.
<figureCitation captionStart="Figures 2126" captionStartId="F5" captionText="Figures 21 - 26. Cymosema waterworthae. Holotype female: 21 head, anterior 22 antennae, lateral 23 mesosoma, lateral 24 mesosoma, dorsal 25 petiole, lateral 26 ovipositor. pet = petiole." figureDoi="10.3897/jhr.55.11482.figures21-26" httpUri="https://binary.pensoft.net/fig/134104" pageId="15" pageNumber="154">21</figureCitation>
). Palpal formula 2:1, palpi minute. Pedicel broader than F2; scape and pedicel reticulate; flagellum 1.5
<normalizedToken originalValue="×">x</normalizedToken>
<pageBreakToken pageId="16" pageNumber="155" start="start">head</pageBreakToken>
height, with 7 parallel-sided funiculars; anellus minute and hardly visible; F2 1.6
<normalizedToken originalValue="×">x</normalizedToken>
as long as broad, F2 1.1
<normalizedToken originalValue="×">x</normalizedToken>
as long as F3.
</paragraph>
<paragraph pageId="16" pageNumber="155">
<emphasis italics="true" pageId="16" pageNumber="155">Mesosoma</emphasis>
(Figs
<figureCitation captionStart="Figures 2126" captionStartId="F5" captionText="Figures 21 - 26. Cymosema waterworthae. Holotype female: 21 head, anterior 22 antennae, lateral 23 mesosoma, lateral 24 mesosoma, dorsal 25 petiole, lateral 26 ovipositor. pet = petiole." figureDoi="10.3897/jhr.55.11482.figures21-26" httpUri="https://binary.pensoft.net/fig/134104" pageId="16" pageNumber="155">23-24</figureCitation>
,
<figureCitation captionStart="Figure 27" captionStartId="F6" captionText="Figure 27. Cymosema waterworthae. Holotype female: fore wing. arrow points to end of postmarginal vein." figureDoi="10.3897/jhr.55.11482.figure27" httpUri="https://binary.pensoft.net/fig/134105" pageId="16" pageNumber="155">27</figureCitation>
). Mesoscutal midlobe coriaceous anteriorly, becoming smooth posteriorly, lateral lobe mostly smooth but with shallow irregular sculpture anteriorly and laterally; notauli deep anteriorly but very shallow posteriorly. Axilla strongly advanced, smooth anteriorly, coriaceous posteriorly; mesoscutellar disc smooth to very shallowly coriaceous; frenal line indicated by narrow groove, frenum smooth; axillular sulcus narrow, axillula coriaceous. Mesepisternum reticulate anterodorsally, becoming smooth ventrally and forming a rounded horizontal shelf anterior to mesocoxae; mesepimeron shallowly reticulate and without a transepimeral sulcus. Propodeal disc finely reticulate, with a weak median carina, sublaterally reticulate and with an oblique elevation anterior to a steep descent towards the broad petiolar foramen; callus bare, with a flange-like elevation posterolateral to spiracle (Fig.
<figureCitation captionStart="Figures 2126" captionStartId="F5" captionText="Figures 21 - 26. Cymosema waterworthae. Holotype female: 21 head, anterior 22 antennae, lateral 23 mesosoma, lateral 24 mesosoma, dorsal 25 petiole, lateral 26 ovipositor. pet = petiole." figureDoi="10.3897/jhr.55.11482.figures21-26" httpUri="https://binary.pensoft.net/fig/134104" pageId="16" pageNumber="155">24</figureCitation>
, flange). Metacoxa shallowly reticulate dorsolaterally. Fore wing spatulate (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F6" captionText="Figure 27. Cymosema waterworthae. Holotype female: fore wing. arrow points to end of postmarginal vein." figureDoi="10.3897/jhr.55.11482.figure27" httpUri="https://binary.pensoft.net/fig/134105" pageId="16" pageNumber="155">27</figureCitation>
), 2.7
<normalizedToken originalValue="×">x</normalizedToken>
as long as broad; costal cell and submarginal vein bare; basal cell ventrally setose, also with a few dorsal setae
<taxonomicName authority="Heraty &amp; Burks, 2017" authorityName="Heraty &amp; Burks" authorityYear="2017" class="Insecta" family="Eucharitidae" genus="Cymosema" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="near" order="Hymenoptera" pageId="16" pageNumber="155" phylum="Arthropoda" rank="species" species="near">near</taxonomicName>
basal fold; speculum absent; cubital and subcubital folds setose; marginal fringe absent; stigmal vein 2.0
<normalizedToken originalValue="×">x</normalizedToken>
as long as broad; postmarginal vein about 1.5
<normalizedToken originalValue="×">x</normalizedToken>
as long as stigmal vein. Hind wing costal cell bare.
</paragraph>
<paragraph pageId="16" pageNumber="155">
<emphasis italics="true" pageId="16" pageNumber="155">Metasoma</emphasis>
(Figs
<figureCitation captionStart="Figures 2126" captionStartId="F5" captionText="Figures 21 - 26. Cymosema waterworthae. Holotype female: 21 head, anterior 22 antennae, lateral 23 mesosoma, lateral 24 mesosoma, dorsal 25 petiole, lateral 26 ovipositor. pet = petiole." figureDoi="10.3897/jhr.55.11482.figures21-26" httpUri="https://binary.pensoft.net/fig/134104" pageId="16" pageNumber="155">25-26</figureCitation>
). Petiole 1.6
<normalizedToken originalValue="×">x</normalizedToken>
as long as broad, 1.0
<normalizedToken originalValue="×">x</normalizedToken>
as long as metacoxa, finely longitudinally rugose-reticulate, with a pair of complete lateral longitudinal carinae and some additional incomplete longitudinal carinae anteriorly, narrowing anteriorly to abruptly expand anteriorly at the condyle, without anterior transverse carina, ventrally with a longitudinal groove that expands at apices. Acrosternite with irregular raised sculpture, swollen and differentiated across indistinct antecostal sulcus from remaining smooth Gs1. Cercus with all setae subequal in length. Hypopygium bare. Ovipositor elongate and slightly recurved, dorsal valve with 7 medially interrupted annuli; ventral valve with 5-6 teeth, and with oblique carina ending in a notch immediately basal to toothed apex.
</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="155" type="male">
<paragraph pageId="16" pageNumber="155">Male.</paragraph>
<paragraph pageId="16" pageNumber="155">Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="17" pageNumber="156" type="materials_examined">
<paragraph pageId="17" pageNumber="156">
<pageBreakToken pageId="17" pageNumber="156" start="start">Material</pageBreakToken>
examined.
</paragraph>
<paragraph pageId="17" pageNumber="156">
<materialsCitation collectingDate="2009-12-08" collectorName="D'Entrecasteaux N. P., R. Waterworth" country="Australia" elevation="105" latitude="-34.83861" location="Coastal Survivors Walk" longLatPrecision="19" longitude="116.00472" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia" typeStatus="Holotype">
<emphasis bold="true" pageId="17" pageNumber="156">
<typeStatus>Holotype</typeStatus>
:
<collectingCountry name="Australia">Australia</collectingCountry>
:
<collectingRegion country="Australia" name="Western Australia">Western Australia</collectingRegion>
</emphasis>
:
<collectorName>
<normalizedToken originalValue="DEntrecasteaux">D'Entrecasteaux</normalizedToken>
N.P.
</collectorName>
,
<location LSID="urn:lsid:plazi:treatment:598DCAC562107936D3BA8D9E8729D9B1:947FCA34059489D297DB73F5439EB182" country="Australia" latitude="-34.83861" longLatPrecision="19" longitude="116.00472" name="Coastal Survivors Walk" stateProvince="Western Australia">Coastal Survivors Walk</location>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.05" unit="m" value="105.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.05" unit="m" value="105.0">105m</elevation>
</quantity>
,
<geoCoordinate degrees="34" direction="south" minutes="50" orientation="latitude" precision="15" seconds="19" value="-34.83861">34°50'19&quot;S</geoCoordinate>
,
<geoCoordinate degrees="116" direction="east" minutes="00" orientation="longitude" precision="15" seconds="17" value="116.00472">116°00'17&quot;E</geoCoordinate>
,
<collectingDate value="2009-12-08">8.xii.2009</collectingDate>
,
<collectorName>R. Waterworth</collectorName>
, sweep [
<specimenCount type="female">1♀</specimenCount>
, UCRC: UCRCENT00364784], deposited in UCRC. Gaster mounted separately on card.
</materialsCitation>
</paragraph>
<caption ID-Zenodo-Dep="989725" doi="10.3897/jhr.55.11482.figures21-26" httpUri="https://binary.pensoft.net/fig/134104" pageId="17" pageNumber="156" start="Figures 2126" startId="F5">
<paragraph pageId="17" pageNumber="156">
<emphasis bold="true" pageId="17" pageNumber="156">Figures 21-26.</emphasis>
<taxonomicName authority="Heraty &amp; Burks, 2017" authorityName="Burks &amp; Mottern" authorityYear="2017" class="Insecta" family="Eucharitidae" genus="Cymosema" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cymosema waterworthae" order="Hymenoptera" pageId="17" pageNumber="156" phylum="Arthropoda" rank="species" species="waterworthae">
<emphasis italics="true" pageId="17" pageNumber="156">Cymosema waterworthae</emphasis>
</taxonomicName>
.
<typeStatus>Holotype</typeStatus>
female:
<emphasis bold="true" pageId="17" pageNumber="156">21</emphasis>
head, anterior
<emphasis bold="true" pageId="17" pageNumber="156">22</emphasis>
antennae, lateral
<emphasis bold="true" pageId="17" pageNumber="156">23</emphasis>
mesosoma, lateral
<emphasis bold="true" pageId="17" pageNumber="156">24</emphasis>
mesosoma, dorsal
<emphasis bold="true" pageId="17" pageNumber="156">25</emphasis>
petiole, lateral
<emphasis bold="true" pageId="17" pageNumber="156">26</emphasis>
ovipositor. pet = petiole.
</paragraph>
</caption>
<caption ID-Zenodo-Dep="989727" doi="10.3897/jhr.55.11482.figure27" httpUri="https://binary.pensoft.net/fig/134105" pageId="17" pageNumber="156" start="Figure 27" startId="F6">
<paragraph pageId="17" pageNumber="156">
<emphasis bold="true" pageId="17" pageNumber="156">Figure 27.</emphasis>
<taxonomicName authority="Heraty &amp; Burks, 2017" authorityName="Burks &amp; Mottern" authorityYear="2017" class="Insecta" family="Eucharitidae" genus="Cymosema" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cymosema waterworthae" order="Hymenoptera" pageId="17" pageNumber="156" phylum="Arthropoda" rank="species" species="waterworthae">
<emphasis italics="true" pageId="17" pageNumber="156">Cymosema waterworthae</emphasis>
</taxonomicName>
.
<typeStatus>Holotype</typeStatus>
female: fore wing. arrow points to end of postmarginal vein.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>