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<document id="9E9F7CB36CB0E08FBC6DD13228706C1E" ID-DOI="10.5252/geodiversitas2019v41a19" ID-GBIF-Dataset="92402f3e-075a-440f-87df-a545fd221c0c" ID-ISSN="1638-9395" ID-Zenodo-Dep="3695594" ID-ZooBank="urn:lsid:zoobank.org:pub:C34F475B-6315-400C-ACE3-B67B1667C033" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1583245730904" checkinUser="valdenar" docAuthor="Downs, Jason P., Daeschler, Edward B., Lo, Nathanael, Carey, Emily N. &amp; Shubin, Neil H." docDate="2019" docId="594A8781BF1DE569FC52CC50DEDA3F76" docLanguage="en" docName="GEODIVERSITAS.41.19.679-698.pdf.imf" docOrigin="Geodiversitas 41 (19)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Asterolepis alticristata Downs &amp; Daeschler &amp; Lo &amp; Carey &amp; Shubin 2019, n. sp." docType="treatment" docUuid="D6EAF712-9D00-4A37-9612-93CF17333564" docUuidSource="ZooBank" docVersion="8" lastPageNumber="688" masterDocId="A573FFF9BF18E562FF84C865DA193D45" masterDocTitle="Asterolepis alticristata n. sp. (Antiarchi) from the Upper Devonian (Frasnian) of Nunavut, Canada, and a report on the antiarch diversity of the Fram Formation" masterLastPageNumber="698" masterPageNumber="679" pageNumber="682" updateTime="1698724279455" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:titleInfo id="6207398BC58D83A4C3A2EECECDC29A09">
<mods:title id="D1E8B4F689B3E4D986C3AF814A8B5FF7">Asterolepis alticristata n. sp. (Antiarchi) from the Upper Devonian (Frasnian) of Nunavut, Canada, and a report on the antiarch diversity of the Fram Formation</mods:title>
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<mods:namePart id="DA384E1B7D816636C4AC3C89B828F706">Downs, Jason P.</mods:namePart>
</mods:name>
<mods:name id="A2C86C52F2E9FCE506164C83FECF187B" type="personal">
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<mods:namePart id="5875E674C0FFC3D3785EFBCA5D1A0C40">Daeschler, Edward B.</mods:namePart>
</mods:name>
<mods:name id="DA963348353C7CFCC2AAFD99D63304DC" type="personal">
<mods:role id="16FC90CF06647EFA42B361B2B329A7C6">
<mods:roleTerm id="C5DC6EBBED607F8CB6E0D7608F2487DA">Author</mods:roleTerm>
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<mods:namePart id="87C4F11013FC2EC3866A52C8A56BF614">Lo, Nathanael</mods:namePart>
</mods:name>
<mods:name id="B588D2FB5C76C990228D3F90CF29DE0B" type="personal">
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<mods:namePart id="F12F5B85A287D14E2D6E91C1EDFC28EB">Carey, Emily N.</mods:namePart>
</mods:name>
<mods:name id="3315E64A3ED156F9D06AAD210CA7D5DC" type="personal">
<mods:role id="3DF01DE6DC862634099E25F038C5C1C3">
<mods:roleTerm id="B613EC021DCC3AF4D880322CD96721F6">Author</mods:roleTerm>
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<mods:namePart id="F48F159050E4C468F39F13735621D1B8">Shubin, Neil H.</mods:namePart>
</mods:name>
<mods:typeOfResource id="0845A20A6187F1BCB6F15FDF80C403F8">text</mods:typeOfResource>
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<mods:title id="0E7911B78C8EA621215465DA6639218A">Geodiversitas</mods:title>
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<mods:part id="F8DD87EC2F0AF140829BAF310C90D1B2">
<mods:date id="D11314673CD7F92CA677C8D4BC340B5F">2019</mods:date>
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<mods:number id="5B253EED222A5117488EFBF60B4ACA1E">2019-10-17</mods:number>
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<mods:detail id="0D88325CD242AB7ED078985C94F0B6D8" type="volume">
<mods:number id="3BC6C50A2B0657D0B2DC1FAC8C2F1031">41</mods:number>
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<mods:number id="D657FB32988434E95984E889679B2B72">19</mods:number>
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<mods:identifier id="6E3E3092AE001E1AEDB07999724892F7" type="DOI">10.5252/geodiversitas2019v41a19</mods:identifier>
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<treatment id="594A8781BF1DE569FC52CC50DEDA3F76" ID-DOI="http://doi.org/10.5281/zenodo.3703489" ID-GBIF-Taxon="162381842" ID-Zenodo-Dep="3703489" LSID="urn:lsid:zoobank.org:act:D6EAF712-9D00-4A37-9612-93CF17333564" httpUri="http://treatment.plazi.org/id/594A8781BF1DE569FC52CC50DEDA3F76" lastPageId="11" lastPageNumber="688" pageId="5" pageNumber="682">
<subSubSection id="99F9651CBF1DE567FC52CC50DF1F3914" box="[982,1286,1077,1105]" pageId="5" pageNumber="682" type="nomenclature">
<paragraph id="D15C3697BF1DE567FC52CC50DF1F3914" blockId="5.[982,1286,1077,1105]" box="[982,1286,1077,1105]" pageId="5" pageNumber="682">
<heading id="8A1481FBBF1DE567FC52CC50DF1F3914" box="[982,1286,1077,1105]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="682" reason="2">
<taxonomicName id="16E34D14BF1DE567FC52CC50DEDD3915" authority="Downs &amp; Daeschler &amp; Lo &amp; Carey &amp; Shubin, 2019" authorityName="Downs &amp; Daeschler &amp; Lo &amp; Carey &amp; Shubin" authorityYear="2019" box="[982,1220,1077,1104]" class="Insecta" family="Tortricidae" genus="Asterolepis" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="682" phylum="Arthropoda" rank="species" species="alticristata" status="sp. nov.">
<emphasis id="E397EA85BF1DE567FC52CC50DEDD3915" bold="true" box="[982,1220,1077,1104]" italics="true" pageId="5" pageNumber="682">Asterolepis alticristata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F8A457FEBF1DE567FB4FCC52DF1F3914" box="[1227,1286,1079,1105]" pageId="5" pageNumber="682" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="99F9651CBF1DE567FCA9CCD1DF1339BA" pageId="5" pageNumber="682" type="reference_group">
<paragraph id="D15C3697BF1DE567FCA9CCD1DED63989" blockId="5.[813,1231,1204,1228]" box="[813,1231,1204,1228]" pageId="5" pageNumber="682">
<treatmentCitationGroup id="F1F311B9BF1DE567FCA9CCD1DED63989" box="[813,1231,1204,1228]" pageId="5" pageNumber="682">
<taxonomicName id="16E34D14BF1DE567FCA9CCD1D9973989" ID-CoL="9235F" authorityName="Eichwald, 1840 sensu Agassiz" authorityYear="1844" box="[813,910,1204,1228]" class="Insecta" family="Tortricidae" genus="Asterolepis" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="688" phylum="Arthropoda" rank="genus">
<emphasis id="E397EA85BF1DE567FCA9CCD1D9973989" box="[813,910,1204,1228]" italics="true" pageId="5" pageNumber="682">Asterolepis</emphasis>
</taxonomicName>
sp. –
<treatmentCitation id="50421086BF1DE567FC4CCCD1DED23989" author="DAESCHLER E. B. &amp; SHUBIN N. H. &amp; JENKINS JR F. A." box="[968,1227,1204,1228]" page="759" pageId="5" pageNumber="682" year="2006">
<bibRefCitation id="B5724B66BF1DE567FC4CCCD1DED23989" author="DAESCHLER E. B. &amp; SHUBIN N. H. &amp; JENKINS JR F. A." box="[968,1227,1204,1228]" pageId="5" pageNumber="682" pagination="757 - 763" refId="ref16243" refString="DAESCHLER E. B., SHUBIN N. H. &amp; JENKINS JR F. A. 2006. - A Devonian tetrapod-like fish and the evolution of the tetrapod body plan. Nature 440: 757 - 763. https: // doi. org / 10.1038 / nature 04639" type="journal article" year="2006">
Daeschler
<emphasis id="E397EA85BF1DE567FBAACCD0DE443989" box="[1070,1117,1204,1228]" italics="true" pageId="5" pageNumber="682">et al.</emphasis>
2006: 759
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="D15C3697BF1DE567FCA9CC82DF1339BA" blockId="5.[813,1290,1254,1279]" box="[813,1290,1254,1279]" pageId="5" pageNumber="682">
“Large asterolepidid” –
<bibRefCitation id="B5724B66BF1DE567FB97CC82DF1F39BA" author="DOWNS J. P. &amp; DAESCHLER E. B. &amp; JENKINS JR. F. A. &amp; SHUBIN N. H." box="[1043,1286,1254,1279]" pageId="5" pageNumber="682" pagination="981 - 996" refId="ref16348" refString="DOWNS J. P., DAESCHLER E. B., JENKINS JR. F. A. &amp; SHUBIN N. H. 2011 a. - A new species of Laccognathus (Sarcopterygii, Porolepiformes) from the Late Devonian of Ellesmere Island, Nunavut, Canada. Journal of Vertebrate Paleontology 31 (5): 981 - 996. https: // doi. org / 10.1080 / 02724634.2011.599462" type="journal article" year="2011">
Downs
<emphasis id="E397EA85BF1DE567FBDBCC82DE9739BB" box="[1119,1166,1254,1278]" italics="true" pageId="5" pageNumber="682">et al.</emphasis>
2011a: 994
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="99F9651CBF1DE567FCA9CD7CDEF43826" pageId="5" pageNumber="682" type="description">
<paragraph id="D15C3697BF1DE567FCA9CD7CDF0A3874" blockId="5.[813,1299,1305,1329]" box="[813,1299,1305,1329]" pageId="5" pageNumber="682">
“Asterolepid placoderm” –
<bibRefCitation id="B5724B66BF1DE567FBB7CD7CDF173874" author="DOWNS J. P. &amp; DAESCHLER E. B. &amp; JENKINS JR. F. A. &amp; SHUBIN N. H." box="[1075,1294,1305,1329]" pageId="5" pageNumber="682" pagination="47 - 59" refId="ref16495" refString="DOWNS J. P., DAESCHLER E. B., JENKINS JR. F. A. &amp; SHUBIN N. H. 2013. - Holoptychius bergmanni sp. nov. (Sarcopterygii, Porolepiformes) from the Upper Devonian of Nunavut, Canada, and a review of Holoptychius taxonomy. Proceedings of the Academy of Natural Sciences of Philadelphia 162: 47 - 59. https: // doi. org / 10.1635 / 053.162.0104" type="journal article" year="2013">
Downs
<emphasis id="E397EA85BF1DE567FBFBCD7CDEB73874" box="[1151,1198,1305,1329]" italics="true" pageId="5" pageNumber="682">et al.</emphasis>
2013: 48
</bibRefCitation>
.
</paragraph>
<paragraph id="D15C3697BF1DE567FCA9CD2EDEF43826" blockId="5.[813,1261,1355,1379]" box="[813,1261,1355,1379]" pageId="5" pageNumber="682">
“Asterolepid antiarch” –
<bibRefCitation id="B5724B66BF1DE567FB9FCD2EDEF03826" author="DOWNS J. P. &amp; DAESCHLER E. B. &amp; LONG A. M. &amp; SHUBIN N. H." box="[1051,1257,1355,1379]" pageId="5" pageNumber="682" pagination="1 - 24" refId="ref16644" refString="DOWNS J. P., DAESCHLER E. B., LONG A. M. &amp; SHUBIN N. H. 2018. - Eusthenopteron jenkinsi sp. nov. (Sarcopterygii, Tristichopteridae) from the Upper Devonian of Nunavut, Canada, and a review of Eusthenopteron taxonomy. Breviora 562: 1 - 24. https: // doi. org / 10.3099 / MCZ 44.1" type="journal article" year="2018">
Downs
<emphasis id="E397EA85BF1DE567FBE3CD2EDE8F3826" box="[1127,1174,1355,1379]" italics="true" pageId="5" pageNumber="682">et al.</emphasis>
2018: 1
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="99F9651CBF1DE567FCA9CD19D9C838EA" pageId="5" pageNumber="682" type="materials_examined">
<paragraph id="D15C3697BF1DE567FCA9CD19D9C838EA" blockId="5.[813,1455,1404,1455]" pageId="5" pageNumber="682">
<materialsCitation id="618B3CCABF1DE567FCA9CD19D9C838EA" ID-GBIF-Occurrence="2573220419" collectionCode="NUFV" pageId="5" pageNumber="682" specimenCode="NUFV 897" specimenCount="1" typeStatus="holotype">
<typeStatus id="0E588835BF1DE567FCA9CD19D98C38D7" box="[813,917,1404,1428]" pageId="5" pageNumber="682" type="holotype">HOLOTYPE</typeStatus>
. —
<specimenCode id="81459EECBF1DE567FC3ACD18DE2938D0" box="[958,1072,1405,1429]" collectionCode="NUFV" pageId="5" pageNumber="682">NUFV 897</specimenCode>
, anterior median dorsal plate of thoracic skeleton (
<figureCitation id="49D82A12BF1DE567FC0FCDF2D9DA38EA" box="[907,963,1431,1455]" captionStart="FIG" captionStartId="6.[132,143,893,911]" captionTargetBox="[137,1449,215,847]" captionTargetId="figure@6.[132,1455,214,867]" captionTargetPageId="6" captionText="FIG. 2. — Asterolepis alticristata n. sp., NUFV 897, holotype, anterior median dorsal plate in visceral (A, photograph; A’, labeled illustration), cranial (B), left lateral (C), and dermal (D) views. Abbreviations:cd.PMD, area overlapped by the posterior median dorsal plate; cv.ADL, area overlapping the anterior dorsolateral plate; cv.Mxl, area overlapping the mixilateral plate; lc, lateral corner; op.r oblique posterolateral ridge. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.3695598" httpUri="https://zenodo.org/record/3695598/files/figure.png" pageId="5" pageNumber="682">Fig. 2</figureCitation>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="99F9651CBF1DE567FCA9CDACDE6E3B53" pageId="5" pageNumber="682" type="etymology">
<paragraph id="D15C3697BF1DE567FCA9CDACDE6E3B53" blockId="5.[813,1455,1481,1558]" pageId="5" pageNumber="682">ETYMOLOGY. — From the Latin ‘altus,’ high, and ‘crista,’ crest, in reference to the tall midline crest of the anterior median dorsal plate that helps to diagnose the species.</paragraph>
</subSubSection>
<subSubSection id="99F9651CBF1DE567FCA9CE55DED13AA8" pageId="5" pageNumber="682" type="materials_examined">
<paragraph id="D15C3697BF1DE567FCA9CE55DE263BF7" blockId="5.[813,1455,1584,1714]" pageId="5" pageNumber="682">
<collectionCode id="B7F2AE52BF1DE567FCA9CE55DE5F3B03" box="[813,1094,1584,1608]" pageId="5" pageNumber="682">TYPE LOCALITY AND HORIZON</collectionCode>
. —
<collectionCode id="B7F2AE52BF1DE567FBEACE55DE8D3B0D" box="[1134,1172,1584,1608]" pageId="5" pageNumber="682">NV</collectionCode>
2K17 locality (77˚09’59.1”N, 86˚16’9.42”W), Fram Formation near the eastern arm of Bird Fiord on southern Ellesmere Island,
<collectingRegion id="1327F875BF1DE567FBD6CE03DF1C3B38" box="[1106,1285,1637,1661]" country="Canada" name="Nunavut" pageId="5" pageNumber="682">Nunavut Territory</collectingRegion>
,
<collectingCountry id="A9F47607BF1DE567FA8ACE00DF453B38" box="[1294,1372,1637,1661]" name="Canada" pageId="5" pageNumber="682">Canada</collectingCountry>
. Palynological data indicate an early Frasnian age (
<emphasis id="E397EA85BF1DE567FB53CE1ADF013BD2" box="[1239,1304,1663,1687]" italics="true" pageId="5" pageNumber="682">medius</emphasis>
and
<emphasis id="E397EA85BF1DE567FAC8CE1ADFB63BD2" box="[1356,1455,1663,1687]" italics="true" pageId="5" pageNumber="682">maclarenii</emphasis>
zones of
<bibRefCitation id="B5724B66BF1DE567FC06CEFFDE283BF7" author="CHI B. I. &amp; HILLS L. V." box="[898,1073,1690,1714]" pageId="5" pageNumber="682" pagination="640 - 818" refId="ref16174" refString="CHI B. I. &amp; HILLS L. V. 1976. - Biostratigraphy and taxonomy of Devonian megaspores, Arctic Canada. Bulletin of Canadian Petroleum Geology 24: 640 - 818." type="journal article" year="1976">Chi &amp; Hills 1976</bibRefCitation>
).
</paragraph>
<paragraph id="D15C3697BF1DE567FCA9CEA9DED13AA8" blockId="5.[813,1456,1740,2029]" pageId="5" pageNumber="682">
REFERRED MATERIAL. —
<materialsCitation id="618B3CCABF1DE567FBA6CEA8DEDC3AA8" ID-GBIF-Occurrence="2573220418" pageId="5" pageNumber="682">
<collectionCode id="B7F2AE52BF1DE567FBA6CEA8DE7E3BA1" box="[1058,1127,1741,1764]" pageId="5" pageNumber="682">NUFV</collectionCode>
1265, nuchal plate;
<collectionCode id="B7F2AE52BF1DE567FAB2CEA8DF623BA1" box="[1334,1403,1741,1764]" pageId="5" pageNumber="682">NUFV</collectionCode>
812, 813, premedian plates;
<collectionCode id="B7F2AE52BF1DE567FB95CE82DE4F3BBB" box="[1041,1110,1767,1790]" pageId="5" pageNumber="682">NUFV</collectionCode>
822, partial anterior median dorsal plate;
<collectionCode id="B7F2AE52BF1DE567FCEECF64D9B63A5D" box="[874,943,1793,1816]" pageId="5" pageNumber="682">NUFV</collectionCode>
814, 815, 823, 828, 834, 835, 842, 845, 849-851, 854, 856, 860-861, 864, 867, 870-873, 876-878, 884-886, 889, 890, 896,
<date id="A55D1057BF1DE567FC1FCF53DE093A0B" box="[923,1040,1846,1870]" pageId="5" pageNumber="682" value="1270" valueMax="1277">1270-1277</date>
, crests of anterior median dorsal plates;
<collectionCode id="B7F2AE52BF1DE567FCA9CF34D96A3A2D" box="[813,883,1873,1896]" pageId="5" pageNumber="682">NUFV</collectionCode>
796, 806, 874, 879, 898, posterior median dorsal plates;
<collectionCode id="B7F2AE52BF1DE567FCA9CF0ED96B3AC7" box="[813,882,1899,1922]" pageId="5" pageNumber="682">NUFV</collectionCode>
802.1, 802.2, 821, 848, posterior median dorsal fragments;
<collectionCode id="B7F2AE52BF1DE567FCA9CFE3D96B3AD8" box="[813,882,1926,1949]" pageId="5" pageNumber="682">NUFV</collectionCode>
892, articulated partial thoracic and pectoral skeleton including median ventral, right and left anterior dorsolateral plates, anterior ventrolateral plates, dorsal central 1 and ventral central 1 plates;
<collectionCode id="B7F2AE52BF1DE567FCF5CFB0D9AF3AA9" box="[881,950,2005,2028]" pageId="5" pageNumber="682">NUFV</collectionCode>
895, right mixilateral plate
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="859C661FBF1EE564FF00CB18DE2F3EF9" ID-DOI="http://doi.org/10.5281/zenodo.3695598" ID-Zenodo-Dep="3695598" httpUri="https://zenodo.org/record/3695598/files/figure.png" pageId="6" pageNumber="683" startId="6.[132,143,893,911]" targetBox="[137,1449,215,847]" targetPageId="6">
<paragraph id="D15C3697BF1EE564FF00CB18DE2F3EF9" blockId="6.[132,1457,893,956]" pageId="6" pageNumber="683">
FIG. 2. —
<taxonomicName id="16E34D14BF1EE564FF57CB18DB913ECA" authority="Downs &amp; Daeschler &amp; Lo &amp; Carey &amp; Shubin, 2019" authorityName="Downs &amp; Daeschler &amp; Lo &amp; Carey &amp; Shubin" authorityYear="2019" box="[211,392,893,911]" class="Insecta" family="Tortricidae" genus="Asterolepis" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="683" phylum="Arthropoda" rank="species" species="alticristata" status="sp. nov.">
<emphasis id="E397EA85BF1EE564FF57CB18DB913ECA" box="[211,392,893,911]" italics="true" pageId="6" pageNumber="683">Asterolepis alticristata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F8A457FEBF1EE564FE0ACB18DBA23ECA" box="[398,443,893,911]" pageId="6" pageNumber="683" rank="species">n. sp.</taxonomicNameLabel>
, NUFV 897, holotype, anterior median dorsal plate in visceral (
<emphasis id="E397EA85BF1EE564FC44CB18D9D43ECA" bold="true" box="[960,973,893,911]" pageId="6" pageNumber="683">A</emphasis>
, photograph;
<emphasis id="E397EA85BF1EE564FBC6CB18DE483ECA" bold="true" box="[1090,1105,893,911]" pageId="6" pageNumber="683">A’</emphasis>
, labeled illustration), cranial (
<emphasis id="E397EA85BF1EE564FAC6CB18DF563ECA" bold="true" box="[1346,1359,893,911]" pageId="6" pageNumber="683">B</emphasis>
), left lateral (
<emphasis id="E397EA85BF1EE564FF0DCBF1DA8E3EE3" bold="true" box="[137,151,916,934]" pageId="6" pageNumber="683">C</emphasis>
), and dermal (
<emphasis id="E397EA85BF1EE564FE8FCBF1DB003EE3" bold="true" box="[267,281,916,934]" pageId="6" pageNumber="683">D</emphasis>
) views. Abbreviations:
<emphasis id="E397EA85BF1EE564FE52CBF1D8043EE3" bold="true" box="[470,541,916,934]" pageId="6" pageNumber="683">cd.PMD</emphasis>
, area overlapped by the posterior median dorsal plate;
<emphasis id="E397EA85BF1EE564FC66CBF1DE393EE3" bold="true" box="[994,1056,916,934]" pageId="6" pageNumber="683">cv.ADL</emphasis>
, area overlapping the anterior dorsolateral plate;
<emphasis id="E397EA85BF1EE564FF00CBCFDAA53EF9" bold="true" box="[132,188,938,956]" pageId="6" pageNumber="683">cv.Mxl</emphasis>
, area overlapping the mixilateral plate;
<emphasis id="E397EA85BF1EE564FD87CBCFD80A3EF9" bold="true" box="[515,531,938,956]" pageId="6" pageNumber="683">lc</emphasis>
, lateral corner;
<emphasis id="E397EA85BF1EE564FD11CBCFD8AB3EF9" bold="true" box="[661,690,938,956]" pageId="6" pageNumber="683">op.</emphasis>
<taxonomicName id="16E34D14BF1EE564FD36CBCFD8E53EF9" box="[690,764,938,956]" pageId="6" pageNumber="683" rank="subSpecies" subSpecies="oblique">
<emphasis id="E397EA85BF1EE564FD36CBCFD8A03EF9" bold="true" box="[690,697,938,956]" pageId="6" pageNumber="683">r</emphasis>
oblique
</taxonomicName>
posterolateral ridge. Scale bar: 5 cm.
</paragraph>
</caption>
<subSubSection id="99F9651CBF1EE564FF00CB93DB6939A7" pageId="6" pageNumber="683" type="diagnosis">
<paragraph id="D15C3697BF1EE564FF00CB93DB6939A7" blockId="6.[132,775,1014,1250]" pageId="6" pageNumber="683">
DIAGNOSIS. — Antiarch referred to
<taxonomicName id="16E34D14BF1EE564FE5ECB93D822394B" authorityName="Eichwald, 1840 sensu Agassiz" authorityYear="1844" box="[474,571,1014,1038]" class="Insecta" family="Tortricidae" genus="Asterolepis" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="688" phylum="Arthropoda" rank="genus">
<emphasis id="E397EA85BF1EE564FE5ECB93D822394B" box="[474,571,1014,1038]" italics="true" pageId="6" pageNumber="683">Asterolepis</emphasis>
</taxonomicName>
by the presence of an anterior median dorsal plate that broadly overlaps the mixilateral plates (
<bibRefCitation id="B5724B66BF1EE564FF42CC4EDB603906" author="MOLOSHNIKOV S." box="[198,377,1067,1091]" pageId="6" pageNumber="683" pagination="691 - 773" refId="ref18033" refString="MOLOSHNIKOV S. 2008. - Devonian antiarchs (Pisces, Antiarchi) from Central and Southern European Russia. Paleontological Journal 42 (7): 691 - 773. https: // doi. org / 10.1134 / S 0031030108070010" type="journal article" year="2008">Moloshnikov 2008</bibRefCitation>
). Distinguished from previously described species by the following combination of anatomical features: 1) a tall, rounded midline crest that rises from a position near the tergal angle of the anterior median dorsal plate and loses most of its height before reaching the posterior median dorsal plate and (2) a midline crest of the posterior median dorsal plate that decreases in height cranial to caudal.
</paragraph>
</subSubSection>
<subSubSection id="99F9651CBF1EE564FF00CD70D8F73B6B" pageId="6" pageNumber="683" type="discussion">
<paragraph id="D15C3697BF1EE564FF00CD70DAF13868" blockId="6.[132,775,1301,1583]" box="[132,232,1301,1327]" pageId="6" pageNumber="683">REMARKS</paragraph>
<paragraph id="D15C3697BF1EE564FF00CD53D8F73B6B" blockId="6.[132,775,1301,1583]" pageId="6" pageNumber="683">
Because the new species is diagnosed by a crest of the anterior and posterior median dorsal plates, we use spatial association with diagnostic plates or plate fragments to refer additional material to the species (including isolated premedian, mixilateral and nuchal plates and an articulated partial skeleton [NUFV 892] that lacks the median dorsal plates). All of these referred materials also share a size category and dermal ornament with the
<taxonomicName id="16E34D14BF1EE564FEA0CE71D81D3B6B" authority="Downs &amp; Daeschler &amp; Lo &amp; Carey &amp; Shubin, 2019" authorityName="Downs &amp; Daeschler &amp; Lo &amp; Carey &amp; Shubin" authorityYear="2019" box="[292,516,1556,1582]" class="Insecta" family="Tortricidae" genus="Asterolepis" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="683" phylum="Arthropoda" rank="species" species="alticristata" status="sp. nov.">
<emphasis id="E397EA85BF1EE564FEA0CE71D81D3B6B" box="[292,516,1556,1582]" italics="true" pageId="6" pageNumber="683">Asterolepis alticristata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F8A457FEBF1EE564FD8FCE70D85F3B6A" box="[523,582,1557,1583]" pageId="6" pageNumber="683" rank="species">n. sp.</taxonomicNameLabel>
<typeStatus id="0E588835BF1EE564FDC8CE70D8633B6A" box="[588,634,1557,1583]" pageId="6" pageNumber="683">type</typeStatus>
specimen.
</paragraph>
</subSubSection>
<subSubSection id="99F9651CBF1EE569FF00CE31DEDA3F76" lastPageId="11" lastPageNumber="688" pageId="6" pageNumber="683" type="description">
<paragraph id="D15C3697BF1EE564FF00CE31D80F3B29" blockId="6.[132,774,1620,2030]" box="[132,534,1620,1646]" pageId="6" pageNumber="683">DESCRIPTION OF CEPHALIC SKELETON</paragraph>
<paragraph id="D15C3697BF1EE564FF00CE11DB903BCB" blockId="6.[132,774,1620,2030]" box="[132,393,1652,1678]" pageId="6" pageNumber="683">
<emphasis id="E397EA85BF1EE564FF00CE11DB903BCB" box="[132,393,1652,1678]" italics="true" pageId="6" pageNumber="683">
Nuchal plate (
<figureCitation id="49D82A12BF1EE564FE9CCE10DB4C3BCB" box="[280,341,1652,1678]" captionStart="FIG" captionStartId="7.[132,143,1408,1426]" captionTargetBox="[137,1452,218,1369]" captionTargetId="figure@7.[132,1455,214,1380]" captionTargetPageId="7" captionText="FIG. 3. — Asterolepis alticristata n. sp.: A-C, NUFV 1265, nuchal plate in dermal (A, photograph; A’, labeled illustration), visceral (B, photograph; B’, labeled illustration), and caudal (C) views; D-F, NUFV 813, premedian plate in dermal (D), visceral (E), and caudal (F) views; G-I, NUFV 812, premedian plate in dermal (G), visceral (H), and caudal (I) views. Abbreviations: cpl, central pit-line line groove; cv.L, area overlapping the lateral plate; cv.Pn, area overlapping the paranuchal plate; d.end, endolymphatic duct; ifc1, principal section of infraorbital pit-line groove; ml.p, midline process; mo.c, median occipital crista; nm, obtected area; pr.p, prerostral process; ps.f, parasagittal foramina; sop, supraoccipital pit; sot, supraotic thickening. Scale bar: 2 cm." figureDoi="http://doi.org/10.5281/zenodo.3695600" httpUri="https://zenodo.org/record/3695600/files/figure.png" pageId="6" pageNumber="683">Fig. 3</figureCitation>
A-C)
</emphasis>
</paragraph>
<paragraph id="D15C3697BF1EE564FF00CEF1DEF53875" blockId="6.[132,774,1620,2030]" lastBlockId="6.[813,1456,1014,2030]" pageId="6" pageNumber="683">
The nuchal plate (NUFV 1265:
<figureCitation id="49D82A12BF1EE564FE53CEF1D80B3BEB" box="[471,530,1684,1710]" captionStart="FIG" captionStartId="7.[132,143,1408,1426]" captionTargetBox="[137,1452,218,1369]" captionTargetId="figure@7.[132,1455,214,1380]" captionTargetPageId="7" captionText="FIG. 3. — Asterolepis alticristata n. sp.: A-C, NUFV 1265, nuchal plate in dermal (A, photograph; A’, labeled illustration), visceral (B, photograph; B’, labeled illustration), and caudal (C) views; D-F, NUFV 813, premedian plate in dermal (D), visceral (E), and caudal (F) views; G-I, NUFV 812, premedian plate in dermal (G), visceral (H), and caudal (I) views. Abbreviations: cpl, central pit-line line groove; cv.L, area overlapping the lateral plate; cv.Pn, area overlapping the paranuchal plate; d.end, endolymphatic duct; ifc1, principal section of infraorbital pit-line groove; ml.p, midline process; mo.c, median occipital crista; nm, obtected area; pr.p, prerostral process; ps.f, parasagittal foramina; sop, supraoccipital pit; sot, supraotic thickening. Scale bar: 2 cm." figureDoi="http://doi.org/10.5281/zenodo.3695600" httpUri="https://zenodo.org/record/3695600/files/figure.png" pageId="6" pageNumber="683">Fig. 3</figureCitation>
A-C) is short and wide; the width/length index (maximum transverse width × 100 divided by maximum sagittal length) is 195. The maximum reported index for
<taxonomicName id="16E34D14BF1EE564FEC7CE91DBB33A4B" authorityName="Eichwald, 1840 sensu Agassiz" authorityYear="1844" box="[323,426,1780,1806]" class="Insecta" family="Tortricidae" genus="Asterolepis" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="688" phylum="Arthropoda" rank="genus">
<emphasis id="E397EA85BF1EE564FEC7CE91DBB33A4B" box="[323,426,1780,1806]" italics="true" pageId="6" pageNumber="683">Asterolepis</emphasis>
</taxonomicName>
is
<quantity id="161B9B72BF1EE564FE40CE91D81F3A4B" box="[452,518,1780,1806]" metricMagnitude="0" metricUnit="m" metricValue="5.334" pageId="6" pageNumber="683" unit="in" value="210.0">210 in</quantity>
<emphasis id="E397EA85BF1EE564FD88CE91D8AC3A4B" box="[524,693,1780,1806]" italics="true" pageId="6" pageNumber="683">
<taxonomicName id="16E34D14BF1EE564FD88CE91D86A3A4B" authorityName="Eichwald, 1840 sensu Agassiz" authorityYear="1844" box="[524,627,1780,1806]" class="Insecta" family="Tortricidae" genus="Asterolepis" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="688" phylum="Arthropoda" rank="genus">Asterolepis</taxonomicName>
scabra
</emphasis>
(
<bibRefCitation id="B5724B66BF1EE564FD47CE91DAA53A6B" author="YOUNG G. C." pageId="6" pageNumber="683" pagination="179 - 192" refId="ref19166" refString="YOUNG G. C. 1987. - Devonian fish remains from Billiluna, eastern Canning Basin, Western Australia. BMR Journal of Australian Geology and Geophysics 10 (2): 179 - 192." type="journal article" year="1987">Young 1987</bibRefCitation>
). Several smaller, unassignable nuchals from NV2K17 (NUFV 1247, 1248, 1252, 1254, 1263) have an index that is comparable with, though none higher than, this reported maximum (See Description of Other Asterolepidoid Specimens from NV2K17). As in
<emphasis id="E397EA85BF1EE564FEF0CFF6D8313AE8" box="[372,552,1939,1965]" italics="true" pageId="6" pageNumber="683">
<taxonomicName id="16E34D14BF1EE564FEF0CFF6DBC63AE8" authorityName="Eichwald, 1840 sensu Agassiz" authorityYear="1844" box="[372,479,1939,1965]" class="Insecta" family="Tortricidae" genus="Asterolepis" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="688" phylum="Arthropoda" rank="genus">Asterolepis</taxonomicName>
ornata
</emphasis>
, the width across the lateral corners is slightly greater than the width across the caudolateral corners; this is despite the presence of pronounced caudolateral processes. As is true of all known species of
<taxonomicName id="16E34D14BF1EE564FAF7CB92D97C3975" authorityName="Eichwald, 1840 sensu Agassiz" authorityYear="1844" class="Insecta" family="Tortricidae" genus="Asterolepis" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="688" phylum="Arthropoda" rank="genus">
<emphasis id="E397EA85BF1EE564FAF7CB92D97C3975" italics="true" pageId="6" pageNumber="683">Asterolepis</emphasis>
</taxonomicName>
, the notch for the postpineal plate is shallow, reaching no further caudal than the lateral corners of the nuchal plate. The central pit-line groove (
<figureCitation id="49D82A12BF1EE564FBD2CC33DE8A3935" box="[1110,1171,1110,1136]" captionStart="FIG" captionStartId="7.[132,143,1408,1426]" captionTargetBox="[137,1452,218,1369]" captionTargetId="figure@7.[132,1455,214,1380]" captionTargetPageId="7" captionText="FIG. 3. — Asterolepis alticristata n. sp.: A-C, NUFV 1265, nuchal plate in dermal (A, photograph; A’, labeled illustration), visceral (B, photograph; B’, labeled illustration), and caudal (C) views; D-F, NUFV 813, premedian plate in dermal (D), visceral (E), and caudal (F) views; G-I, NUFV 812, premedian plate in dermal (G), visceral (H), and caudal (I) views. Abbreviations: cpl, central pit-line line groove; cv.L, area overlapping the lateral plate; cv.Pn, area overlapping the paranuchal plate; d.end, endolymphatic duct; ifc1, principal section of infraorbital pit-line groove; ml.p, midline process; mo.c, median occipital crista; nm, obtected area; pr.p, prerostral process; ps.f, parasagittal foramina; sop, supraoccipital pit; sot, supraotic thickening. Scale bar: 2 cm." figureDoi="http://doi.org/10.5281/zenodo.3695600" httpUri="https://zenodo.org/record/3695600/files/figure.png" pageId="6" pageNumber="683">Fig. 3</figureCitation>
A’[cp1]) extends across the width of the nuchal; it exhibits a pronounced median dip and meets the lateral margins of the plate well caudal to the lateral corners. Tiny, parasagittal foramina just caudal to the pit-line are weakly visible among the tubercles of the ornament.
<bibRefCitation id="B5724B66BF1EE564FCA9CC90D9D03855" author="STENSIO E. A." box="[813,969,1269,1296]" pageId="6" pageNumber="683" pagination="1 - 212" refId="ref18490" refString="STENSIO E. A. 1931. - Upper Devonian vertebrates from East Greenland collected by the Danish Greenland expeditions in 1929 and 1930. Meddelelser om Gronland 86 (1): 1 - 212." type="journal article" year="1931">Stensiö (1931)</bibRefCitation>
presumed these to be the external openings of the endolympatic ducts (
<figureCitation id="49D82A12BF1EE564FBBDCD70DE6E3875" box="[1081,1143,1301,1328]" captionStart="FIG" captionStartId="7.[132,143,1408,1426]" captionTargetBox="[137,1452,218,1369]" captionTargetId="figure@7.[132,1455,214,1380]" captionTargetPageId="7" captionText="FIG. 3. — Asterolepis alticristata n. sp.: A-C, NUFV 1265, nuchal plate in dermal (A, photograph; A’, labeled illustration), visceral (B, photograph; B’, labeled illustration), and caudal (C) views; D-F, NUFV 813, premedian plate in dermal (D), visceral (E), and caudal (F) views; G-I, NUFV 812, premedian plate in dermal (G), visceral (H), and caudal (I) views. Abbreviations: cpl, central pit-line line groove; cv.L, area overlapping the lateral plate; cv.Pn, area overlapping the paranuchal plate; d.end, endolymphatic duct; ifc1, principal section of infraorbital pit-line groove; ml.p, midline process; mo.c, median occipital crista; nm, obtected area; pr.p, prerostral process; ps.f, parasagittal foramina; sop, supraoccipital pit; sot, supraotic thickening. Scale bar: 2 cm." figureDoi="http://doi.org/10.5281/zenodo.3695600" httpUri="https://zenodo.org/record/3695600/files/figure.png" pageId="6" pageNumber="683">Fig. 3</figureCitation>
A’[d.end]).
</paragraph>
<paragraph id="D15C3697BF1EE564FCC7CD50DFB63B6A" blockId="6.[813,1456,1014,2030]" pageId="6" pageNumber="683">
The nuchal is a thick plate with broad and deep zones of overlap onto lateral (
<figureCitation id="49D82A12BF1EE564FC79CD30DE2F382A" box="[1021,1078,1365,1391]" captionStart="FIG" captionStartId="7.[132,143,1408,1426]" captionTargetBox="[137,1452,218,1369]" captionTargetId="figure@7.[132,1455,214,1380]" captionTargetPageId="7" captionText="FIG. 3. — Asterolepis alticristata n. sp.: A-C, NUFV 1265, nuchal plate in dermal (A, photograph; A’, labeled illustration), visceral (B, photograph; B’, labeled illustration), and caudal (C) views; D-F, NUFV 813, premedian plate in dermal (D), visceral (E), and caudal (F) views; G-I, NUFV 812, premedian plate in dermal (G), visceral (H), and caudal (I) views. Abbreviations: cpl, central pit-line line groove; cv.L, area overlapping the lateral plate; cv.Pn, area overlapping the paranuchal plate; d.end, endolymphatic duct; ifc1, principal section of infraorbital pit-line groove; ml.p, midline process; mo.c, median occipital crista; nm, obtected area; pr.p, prerostral process; ps.f, parasagittal foramina; sop, supraoccipital pit; sot, supraotic thickening. Scale bar: 2 cm." figureDoi="http://doi.org/10.5281/zenodo.3695600" httpUri="https://zenodo.org/record/3695600/files/figure.png" pageId="6" pageNumber="683">Fig. 3</figureCitation>
A’[cv.L]) and paranuchal (
<figureCitation id="49D82A12BF1EE564FABACD30DF6F382A" box="[1342,1398,1365,1391]" captionStart="FIG" captionStartId="7.[132,143,1408,1426]" captionTargetBox="[137,1452,218,1369]" captionTargetId="figure@7.[132,1455,214,1380]" captionTargetPageId="7" captionText="FIG. 3. — Asterolepis alticristata n. sp.: A-C, NUFV 1265, nuchal plate in dermal (A, photograph; A’, labeled illustration), visceral (B, photograph; B’, labeled illustration), and caudal (C) views; D-F, NUFV 813, premedian plate in dermal (D), visceral (E), and caudal (F) views; G-I, NUFV 812, premedian plate in dermal (G), visceral (H), and caudal (I) views. Abbreviations: cpl, central pit-line line groove; cv.L, area overlapping the lateral plate; cv.Pn, area overlapping the paranuchal plate; d.end, endolymphatic duct; ifc1, principal section of infraorbital pit-line groove; ml.p, midline process; mo.c, median occipital crista; nm, obtected area; pr.p, prerostral process; ps.f, parasagittal foramina; sop, supraoccipital pit; sot, supraotic thickening. Scale bar: 2 cm." figureDoi="http://doi.org/10.5281/zenodo.3695600" httpUri="https://zenodo.org/record/3695600/files/figure.png" pageId="6" pageNumber="683">Fig.3</figureCitation>
A’[cv. Pn]) plates. There is a very short obtected area (
<figureCitation id="49D82A12BF1EE564FA9ACD10DF4338CA" box="[1310,1370,1397,1423]" captionStart="FIG" captionStartId="7.[132,143,1408,1426]" captionTargetBox="[137,1452,218,1369]" captionTargetId="figure@7.[132,1455,214,1380]" captionTargetPageId="7" captionText="FIG. 3. — Asterolepis alticristata n. sp.: A-C, NUFV 1265, nuchal plate in dermal (A, photograph; A’, labeled illustration), visceral (B, photograph; B’, labeled illustration), and caudal (C) views; D-F, NUFV 813, premedian plate in dermal (D), visceral (E), and caudal (F) views; G-I, NUFV 812, premedian plate in dermal (G), visceral (H), and caudal (I) views. Abbreviations: cpl, central pit-line line groove; cv.L, area overlapping the lateral plate; cv.Pn, area overlapping the paranuchal plate; d.end, endolymphatic duct; ifc1, principal section of infraorbital pit-line groove; ml.p, midline process; mo.c, median occipital crista; nm, obtected area; pr.p, prerostral process; ps.f, parasagittal foramina; sop, supraoccipital pit; sot, supraotic thickening. Scale bar: 2 cm." figureDoi="http://doi.org/10.5281/zenodo.3695600" httpUri="https://zenodo.org/record/3695600/files/figure.png" pageId="6" pageNumber="683">Fig. 3</figureCitation>
A’[nm]) along the nuchal’s caudal margin that is unornamented though not depressed. The caudal margin of the plate is notched in parasagittal positions close to the plate’s midline. These notches border a short caudal midline process (
<figureCitation id="49D82A12BF1EE564FB4FCD90DF113B4A" box="[1227,1288,1525,1551]" captionStart="FIG" captionStartId="7.[132,143,1408,1426]" captionTargetBox="[137,1452,218,1369]" captionTargetId="figure@7.[132,1455,214,1380]" captionTargetPageId="7" captionText="FIG. 3. — Asterolepis alticristata n. sp.: A-C, NUFV 1265, nuchal plate in dermal (A, photograph; A’, labeled illustration), visceral (B, photograph; B’, labeled illustration), and caudal (C) views; D-F, NUFV 813, premedian plate in dermal (D), visceral (E), and caudal (F) views; G-I, NUFV 812, premedian plate in dermal (G), visceral (H), and caudal (I) views. Abbreviations: cpl, central pit-line line groove; cv.L, area overlapping the lateral plate; cv.Pn, area overlapping the paranuchal plate; d.end, endolymphatic duct; ifc1, principal section of infraorbital pit-line groove; ml.p, midline process; mo.c, median occipital crista; nm, obtected area; pr.p, prerostral process; ps.f, parasagittal foramina; sop, supraoccipital pit; sot, supraotic thickening. Scale bar: 2 cm." figureDoi="http://doi.org/10.5281/zenodo.3695600" httpUri="https://zenodo.org/record/3695600/files/figure.png" pageId="6" pageNumber="683">Fig. 3</figureCitation>
A’[ml.p]) of the plate similar to the one described in
<emphasis id="E397EA85BF1EE564FB1DCE70DEEF3B6B" box="[1177,1270,1557,1583]" italics="true" pageId="6" pageNumber="683">A. ornata</emphasis>
(
<bibRefCitation id="B5724B66BF1EE564FA80CE71DFB93B6A" author="UPENIECE I." box="[1284,1440,1556,1583]" pageId="6" pageNumber="683" refId="ref18774" refString="UPENIECE I. 2011. - Palaeoecology and juvenile individuals of the Devonian placoderm and acanthodian fishes from Lode site, Latvia. (Doctoral dissertation). Retrieved from https: // dspace. lu. lv / dspace / handle / 7 / 4659 (last access on 3 rd October 2019)." type="url" year="2011">Upeniece 2011</bibRefCitation>
).
</paragraph>
<paragraph id="D15C3697BF1EE565FCC7CE51DB8D3B2B" blockId="6.[813,1456,1014,2030]" lastBlockId="7.[132,776,1588,1743]" lastPageId="7" lastPageNumber="684" pageId="6" pageNumber="683">
The visceral surface of the nuchal exhibits a supraotic thickening (
<figureCitation id="49D82A12BF1EE564FC34CE31D9F63B2B" box="[944,1007,1620,1646]" captionStart="FIG" captionStartId="7.[132,143,1408,1426]" captionTargetBox="[137,1452,218,1369]" captionTargetId="figure@7.[132,1455,214,1380]" captionTargetPageId="7" captionText="FIG. 3. — Asterolepis alticristata n. sp.: A-C, NUFV 1265, nuchal plate in dermal (A, photograph; A’, labeled illustration), visceral (B, photograph; B’, labeled illustration), and caudal (C) views; D-F, NUFV 813, premedian plate in dermal (D), visceral (E), and caudal (F) views; G-I, NUFV 812, premedian plate in dermal (G), visceral (H), and caudal (I) views. Abbreviations: cpl, central pit-line line groove; cv.L, area overlapping the lateral plate; cv.Pn, area overlapping the paranuchal plate; d.end, endolymphatic duct; ifc1, principal section of infraorbital pit-line groove; ml.p, midline process; mo.c, median occipital crista; nm, obtected area; pr.p, prerostral process; ps.f, parasagittal foramina; sop, supraoccipital pit; sot, supraotic thickening. Scale bar: 2 cm." figureDoi="http://doi.org/10.5281/zenodo.3695600" httpUri="https://zenodo.org/record/3695600/files/figure.png" pageId="6" pageNumber="683">Fig. 3</figureCitation>
B’[sot]) that surrounds the large supraoccipital pits (containing the internal openings of the endolymphatic ducts;
<bibRefCitation id="B5724B66BF1EE564FC69CEF1DE673BEB" author="STENSIO E. A." box="[1005,1150,1684,1710]" pageId="6" pageNumber="683" pagination="1 - 212" refId="ref18490" refString="STENSIO E. A. 1931. - Upper Devonian vertebrates from East Greenland collected by the Danish Greenland expeditions in 1929 and 1930. Meddelelser om Gronland 86 (1): 1 - 212." type="journal article" year="1931">Stensiö 1931</bibRefCitation>
;
<figureCitation id="49D82A12BF1EE564FB0ACEF1DED73BEB" box="[1166,1230,1684,1710]" captionStart="FIG" captionStartId="7.[132,143,1408,1426]" captionTargetBox="[137,1452,218,1369]" captionTargetId="figure@7.[132,1455,214,1380]" captionTargetPageId="7" captionText="FIG. 3. — Asterolepis alticristata n. sp.: A-C, NUFV 1265, nuchal plate in dermal (A, photograph; A’, labeled illustration), visceral (B, photograph; B’, labeled illustration), and caudal (C) views; D-F, NUFV 813, premedian plate in dermal (D), visceral (E), and caudal (F) views; G-I, NUFV 812, premedian plate in dermal (G), visceral (H), and caudal (I) views. Abbreviations: cpl, central pit-line line groove; cv.L, area overlapping the lateral plate; cv.Pn, area overlapping the paranuchal plate; d.end, endolymphatic duct; ifc1, principal section of infraorbital pit-line groove; ml.p, midline process; mo.c, median occipital crista; nm, obtected area; pr.p, prerostral process; ps.f, parasagittal foramina; sop, supraoccipital pit; sot, supraotic thickening. Scale bar: 2 cm." figureDoi="http://doi.org/10.5281/zenodo.3695600" httpUri="https://zenodo.org/record/3695600/files/figure.png" pageId="6" pageNumber="683">Fig. 3</figureCitation>
B’[sop]). The caudal surface of the nuchal plate exhibits a median occipital crista (
<figureCitation id="49D82A12BF1EE564FCB2CEB1D9913BAB" box="[822,904,1747,1774]" captionStart="FIG" captionStartId="7.[132,143,1408,1426]" captionTargetBox="[137,1452,218,1369]" captionTargetId="figure@7.[132,1455,214,1380]" captionTargetPageId="7" captionText="FIG. 3. — Asterolepis alticristata n. sp.: A-C, NUFV 1265, nuchal plate in dermal (A, photograph; A’, labeled illustration), visceral (B, photograph; B’, labeled illustration), and caudal (C) views; D-F, NUFV 813, premedian plate in dermal (D), visceral (E), and caudal (F) views; G-I, NUFV 812, premedian plate in dermal (G), visceral (H), and caudal (I) views. Abbreviations: cpl, central pit-line line groove; cv.L, area overlapping the lateral plate; cv.Pn, area overlapping the paranuchal plate; d.end, endolymphatic duct; ifc1, principal section of infraorbital pit-line groove; ml.p, midline process; mo.c, median occipital crista; nm, obtected area; pr.p, prerostral process; ps.f, parasagittal foramina; sop, supraoccipital pit; sot, supraotic thickening. Scale bar: 2 cm." figureDoi="http://doi.org/10.5281/zenodo.3695600" httpUri="https://zenodo.org/record/3695600/files/figure.png" pageId="6" pageNumber="683">Fig. 3C</figureCitation>
[mo.c]) and, on either side of it, parasagittal fossae that were interpreted by
<bibRefCitation id="B5724B66BF1EE564FBC0CE91DEFE3A4B" author="STENSIO E. A." box="[1092,1255,1780,1806]" pageId="6" pageNumber="683" pagination="1 - 212" refId="ref18490" refString="STENSIO E. A. 1931. - Upper Devonian vertebrates from East Greenland collected by the Danish Greenland expeditions in 1929 and 1930. Meddelelser om Gronland 86 (1): 1 - 212." type="journal article" year="1931">Stensiö (1931)</bibRefCitation>
to be attachment areas for cranial levator muscles. In caudal view, the visceral surface of the plate has a tight, nearly semicircular arch to it. The areas lateral to the visceral arch are depressed relative to the area external to it such that a pronounced horizontal ledge runs across the plate at the most external reach of the visceral arch. The caudal surface of the nuchal is the thickest part of the plate and there is a dramatic decrease in nuchal thickness rostrally from the depressed visceral extensions of the plate’s caudal surface.
</paragraph>
<caption id="859C661FBF1FE565FF00CDE5DE4638A9" ID-DOI="http://doi.org/10.5281/zenodo.3695600" ID-Zenodo-Dep="3695600" httpUri="https://zenodo.org/record/3695600/files/figure.png" pageId="7" pageNumber="684" startId="7.[132,143,1408,1426]" targetBox="[137,1452,218,1369]" targetPageId="7">
<paragraph id="D15C3697BF1FE565FF00CDE5DE4638A9" blockId="7.[132,1457,1408,1517]" pageId="7" pageNumber="684">
FIG. 3. —
<taxonomicName id="16E34D14BF1FE565FF52CDE5DB9538D7" authority="Downs &amp; Daeschler &amp; Lo &amp; Carey &amp; Shubin, 2019" authorityName="Downs &amp; Daeschler &amp; Lo &amp; Carey &amp; Shubin" authorityYear="2019" box="[214,396,1408,1426]" class="Insecta" family="Tortricidae" genus="Asterolepis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="684" phylum="Arthropoda" rank="species" species="alticristata" status="sp. nov.">
<emphasis id="E397EA85BF1FE565FF52CDE5DB9538D7" box="[214,396,1408,1426]" italics="true" pageId="7" pageNumber="684">Asterolepis alticristata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F8A457FEBF1FE565FE16CDE5DBDC38D7" box="[402,453,1408,1426]" pageId="7" pageNumber="684" rank="species">n. sp.</taxonomicNameLabel>
:
<emphasis id="E397EA85BF1FE565FE48CDE5DBF738D7" bold="true" box="[460,494,1408,1426]" pageId="7" pageNumber="684">A -C</emphasis>
, NUFV 1265, nuchal plate in dermal (
<emphasis id="E397EA85BF1FE565FCAECDE5D92E38D7" bold="true" box="[810,823,1408,1426]" pageId="7" pageNumber="684">A</emphasis>
, photograph;
<emphasis id="E397EA85BF1FE565FC2ACDE5D9A438D7" bold="true" box="[942,957,1408,1426]" pageId="7" pageNumber="684">A’</emphasis>
, labeled illustration), visceral (
<emphasis id="E397EA85BF1FE565FB3FCDE5DED138D7" bold="true" box="[1211,1224,1408,1426]" pageId="7" pageNumber="684">B</emphasis>
, photograph;
<emphasis id="E397EA85BF1FE565FAC4CDE5DF4B38D7" bold="true" box="[1344,1362,1408,1426]" pageId="7" pageNumber="684">B’</emphasis>
, labeled illustration), and caudal (
<emphasis id="E397EA85BF1FE565FEC1CDF3DB4A38ED" bold="true" box="[325,339,1430,1448]" pageId="7" pageNumber="684">C</emphasis>
) views;
<emphasis id="E397EA85BF1FE565FE12CDF3DBAF38ED" bold="true" box="[406,438,1430,1448]" pageId="7" pageNumber="684">D -F</emphasis>
, NUFV 813, premedian plate in dermal (
<emphasis id="E397EA85BF1FE565FD7BCDF3D91438ED" bold="true" box="[767,781,1430,1448]" pageId="7" pageNumber="684">D</emphasis>
), visceral (
<emphasis id="E397EA85BF1FE565FCE1CDF3D96838ED" bold="true" box="[869,881,1430,1448]" pageId="7" pageNumber="684">E</emphasis>
), and caudal (
<emphasis id="E397EA85BF1FE565FC61CDF3D9E938ED" bold="true" box="[997,1008,1430,1448]" pageId="7" pageNumber="684">F</emphasis>
) views;
<emphasis id="E397EA85BF1FE565FBB6CDF3DE5738ED" bold="true" box="[1074,1102,1430,1448]" pageId="7" pageNumber="684">G -I</emphasis>
, NUFV 812, premedian plate in dermal (
<emphasis id="E397EA85BF1FE565FA13CDF3DFBC38ED" bold="true" box="[1431,1445,1430,1448]" pageId="7" pageNumber="684">G</emphasis>
), visceral (
<emphasis id="E397EA85BF1FE565FF4ACDC8DAC538FA" bold="true" box="[206,220,1453,1471]" pageId="7" pageNumber="684">H</emphasis>
), and caudal (
<emphasis id="E397EA85BF1FE565FED5CDC8DB4E38FA" bold="true" box="[337,343,1453,1471]" pageId="7" pageNumber="684">I</emphasis>
) views. Abbreviations:
<emphasis id="E397EA85BF1FE565FD93CDC8D82B38FA" bold="true" box="[535,562,1453,1471]" pageId="7" pageNumber="684">cpl</emphasis>
, central pit-line line groove;
<emphasis id="E397EA85BF1FE565FC9BCDC8D95B38FA" bold="true" box="[799,834,1453,1471]" pageId="7" pageNumber="684">cv.L</emphasis>
, area overlapping the lateral plate;
<emphasis id="E397EA85BF1FE565FBECCDC8DE8138FA" bold="true" box="[1128,1176,1453,1471]" pageId="7" pageNumber="684">cv.Pn</emphasis>
, area overlapping the paranuchal plate;
<emphasis id="E397EA85BF1FE565FF3CCDA1DAF33893" bold="true" box="[184,234,1476,1494]" pageId="7" pageNumber="684">d.end</emphasis>
, endolymphatic duct;
<emphasis id="E397EA85BF1FE565FE21CDA1DBDC3893" bold="true" box="[421,453,1476,1494]" pageId="7" pageNumber="684">ifc1</emphasis>
, principal section of infraorbital pit-line groove;
<emphasis id="E397EA85BF1FE565FCD7CDA1D9633893" bold="true" box="[851,890,1476,1494]" pageId="7" pageNumber="684">ml.p</emphasis>
, midline process;
<emphasis id="E397EA85BF1FE565FB96CDA1DE263893" bold="true" box="[1042,1087,1476,1494]" pageId="7" pageNumber="684">mo.c</emphasis>
, median occipital crista;
<emphasis id="E397EA85BF1FE565FA8BCDA1DF323893" bold="true" box="[1295,1323,1476,1494]" pageId="7" pageNumber="684">nm</emphasis>
, obtected area;
<emphasis id="E397EA85BF1FE565FF00CDBEDABF38A8" bold="true" box="[132,166,1499,1517]" pageId="7" pageNumber="684">pr.p</emphasis>
, prerostral process;
<emphasis id="E397EA85BF1FE565FED5CDBEDB6B38A8" bold="true" box="[337,370,1499,1517]" pageId="7" pageNumber="684">ps.f</emphasis>
, parasagittal foramina;
<emphasis id="E397EA85BF1FE565FDB3CDBED84138A8" bold="true" box="[567,600,1499,1517]" pageId="7" pageNumber="684">sop</emphasis>
, supraoccipital pit;
<emphasis id="E397EA85BF1FE565FD78CDBED90138A8" bold="true" box="[764,792,1499,1517]" pageId="7" pageNumber="684">sot</emphasis>
, supraotic thickening. Scale bar: 2 cm.
</paragraph>
</caption>
<paragraph id="D15C3697BF1FE565FF1FCE11D8F33B8A" blockId="7.[132,776,1588,1743]" pageId="7" pageNumber="684">The ornament of the nuchal plate is coarsely tuberculate. Tubercles are fused into ridges close to the rostral and lateral margins of the plate; these ridges radiate from the center.</paragraph>
<paragraph id="D15C3697BF1FE565FF00CE91DBB03A4B" blockId="7.[132,775,1780,2030]" box="[132,425,1780,1806]" pageId="7" pageNumber="684">
<emphasis id="E397EA85BF1FE565FF00CE91DBB03A4B" box="[132,425,1780,1806]" italics="true" pageId="7" pageNumber="684">
Premedian plate (
<figureCitation id="49D82A12BF1FE565FEB8CE90DB603A4B" box="[316,377,1780,1806]" captionStart="FIG" captionStartId="7.[132,143,1408,1426]" captionTargetBox="[137,1452,218,1369]" captionTargetId="figure@7.[132,1455,214,1380]" captionTargetPageId="7" captionText="FIG. 3. — Asterolepis alticristata n. sp.: A-C, NUFV 1265, nuchal plate in dermal (A, photograph; A’, labeled illustration), visceral (B, photograph; B’, labeled illustration), and caudal (C) views; D-F, NUFV 813, premedian plate in dermal (D), visceral (E), and caudal (F) views; G-I, NUFV 812, premedian plate in dermal (G), visceral (H), and caudal (I) views. Abbreviations: cpl, central pit-line line groove; cv.L, area overlapping the lateral plate; cv.Pn, area overlapping the paranuchal plate; d.end, endolymphatic duct; ifc1, principal section of infraorbital pit-line groove; ml.p, midline process; mo.c, median occipital crista; nm, obtected area; pr.p, prerostral process; ps.f, parasagittal foramina; sop, supraoccipital pit; sot, supraotic thickening. Scale bar: 2 cm." figureDoi="http://doi.org/10.5281/zenodo.3695600" httpUri="https://zenodo.org/record/3695600/files/figure.png" pageId="7" pageNumber="684">Fig. 3</figureCitation>
D-I)
</emphasis>
</paragraph>
<paragraph id="D15C3697BF1FE565FF00CF71DECF3AAB" blockId="7.[132,775,1780,2030]" lastBlockId="7.[813,1456,1588,2030]" pageId="7" pageNumber="684">
The two premedian plate specimens referred to
<taxonomicName id="16E34D14BF1FE565FD1ECF71DAEB3A08" authority="Downs &amp; Daeschler &amp; Lo &amp; Carey &amp; Shubin, 2019" authorityName="Downs &amp; Daeschler &amp; Lo &amp; Carey &amp; Shubin" authorityYear="2019" class="Insecta" family="Tortricidae" genus="Asterolepis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="684" phylum="Arthropoda" rank="species" species="alticristata" status="sp. nov.">
<emphasis id="E397EA85BF1FE565FD1ECF71DAEB3A08" italics="true" pageId="7" pageNumber="684">Asterolepis alticristata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F8A457FEBF1FE565FF7FCF51DB2E3A0B" box="[251,311,1844,1870]" pageId="7" pageNumber="684" rank="species">n. sp.</taxonomicNameLabel>
(NUFV 812 [maximum sagittal length of
<quantity id="161B9B72BF1FE565FF00CF36DAE33A2B" box="[132,250,1875,1902]" metricMagnitude="-2" metricUnit="m" metricValue="2.985" pageId="7" pageNumber="684" unit="mm" value="29.85">29.85 mm</quantity>
] and 813 [same length =
<quantity id="161B9B72BF1FE565FD8ACF31D89F3A2B" box="[526,646,1875,1902]" metricMagnitude="-2" metricUnit="m" metricValue="2.202" pageId="7" pageNumber="684" unit="mm" value="22.02">22.02 mm</quantity>
]:
<figureCitation id="49D82A12BF1FE565FD1CCF36D8CF3A2B" box="[664,726,1875,1902]" captionStart="FIG" captionStartId="7.[132,143,1408,1426]" captionTargetBox="[137,1452,218,1369]" captionTargetId="figure@7.[132,1455,214,1380]" captionTargetPageId="7" captionText="FIG. 3. — Asterolepis alticristata n. sp.: A-C, NUFV 1265, nuchal plate in dermal (A, photograph; A’, labeled illustration), visceral (B, photograph; B’, labeled illustration), and caudal (C) views; D-F, NUFV 813, premedian plate in dermal (D), visceral (E), and caudal (F) views; G-I, NUFV 812, premedian plate in dermal (G), visceral (H), and caudal (I) views. Abbreviations: cpl, central pit-line line groove; cv.L, area overlapping the lateral plate; cv.Pn, area overlapping the paranuchal plate; d.end, endolymphatic duct; ifc1, principal section of infraorbital pit-line groove; ml.p, midline process; mo.c, median occipital crista; nm, obtected area; pr.p, prerostral process; ps.f, parasagittal foramina; sop, supraoccipital pit; sot, supraotic thickening. Scale bar: 2 cm." figureDoi="http://doi.org/10.5281/zenodo.3695600" httpUri="https://zenodo.org/record/3695600/files/figure.png" pageId="7" pageNumber="684">Fig. 3</figureCitation>
D-I) share a size category and an ornament
<typeStatus id="0E588835BF1FE565FDA8CF11D8433ACB" box="[556,602,1908,1934]" pageId="7" pageNumber="684">type</typeStatus>
but differ from one another in some specific details. The premedian plate has been recognized as a source of intraspecific variation throughout
<taxonomicName id="16E34D14BF1FE565FE8ECFB6DB683AA8" authorityName="Cope" authorityYear="1885" box="[266,369,2003,2029]" class="Leptocardii" higherTaxonomySource="GBIF" kingdom="Animalia" order="Antiarchi" pageId="7" pageNumber="684" phylum="Chordata" rank="order">Antiarchi</taxonomicName>
(
<bibRefCitation id="B5724B66BF1FE565FE07CFB1D8123AA8" author="YOUNG G. C." box="[387,523,2003,2030]" pageId="7" pageNumber="684" pagination="635 - 660" refId="ref19128" refString="YOUNG G. C. 1984. - Reconstruction of the jaws and braincase in the Devonian placoderm fish Bothriolepis. Palaeontology 27: 635 - 660. https: // biodiversitylibrary. org / page / 49742174" type="journal article" year="1984">Young 1984</bibRefCitation>
;
<bibRefCitation id="B5724B66BF1FE565FD9ECFB1D8EF3AA8" author="DOWNS J. P. &amp; CRISWELL K. E. &amp; DAESCHLER E. B." box="[538,758,2003,2030]" pageId="7" pageNumber="684" pagination="191 - 203" refId="ref16422" refString="DOWNS J. P., CRISWELL K. E. &amp; DAESCHLER E. B. 2011 b. - Mass mortality of juvenile antiarchs (Bothriolepis sp.) from the Catskill Formation (Upper Devonian, Famennian Stage), Tioga County, Pennsylvania. Proceedings of the Academy of Natural Sciences of Philadelphia 161: 191 - 203. https: // doi. org / 10.1635 / 053.161.0111" type="journal article" year="2011">
Downs
<emphasis id="E397EA85BF1FE565FDF5CFB1D8BE3AA8" box="[625,679,2003,2029]" italics="true" pageId="7" pageNumber="684">et al.</emphasis>
2011b
</bibRefCitation>
). Both specimens exhibit the generalized premedian plate shape that has been well described in
<emphasis id="E397EA85BF1FE565FB05CE31DF213B2B" box="[1153,1336,1620,1646]" italics="true" pageId="7" pageNumber="684">
<taxonomicName id="16E34D14BF1FE565FB05CE31DEF53B2B" authorityName="Eichwald, 1840 sensu Agassiz" authorityYear="1844" box="[1153,1260,1620,1646]" class="Insecta" family="Tortricidae" genus="Asterolepis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="688" phylum="Arthropoda" rank="genus">Asterolepis</taxonomicName>
ornata
</emphasis>
(
<bibRefCitation id="B5724B66BF1FE565FACCCE31D97F3BCB" author="LUKsEVICS E." pageId="7" pageNumber="684" pagination="687 - 692" refId="ref17738" refString="LUKsEVICS E. 2001. - The orbito-nasal area of Asterolepis ornata, a Middle Devonian placoderm fish. Journal of Vertebrate Paleontology 21 (4): 687 - 692. https: // doi. org / 10.1671 / 0272 - 4634 (2001) 021 [0687: TON AOA] 2.0. CO; 2" type="journal article" year="2001">Lukševičs 2001</bibRefCitation>
). Both are thick plates, especially at the caudal end, and are arched across the midline. Both have unornamented zones in lateral positions along the depressed rostral margin of the plate; the rostral margin exhibits a midline notch. In both, the tall caudal surface is depressed rostrally (analogue of the bothriolepidid preorbital recess;
<bibRefCitation id="B5724B66BF1FE565FB55CF71DF6E3A6B" author="LUKsEVICS E." box="[1233,1399,1812,1838]" pageId="7" pageNumber="684" pagination="687 - 692" refId="ref17738" refString="LUKsEVICS E. 2001. - The orbito-nasal area of Asterolepis ornata, a Middle Devonian placoderm fish. Journal of Vertebrate Paleontology 21 (4): 687 - 692. https: // doi. org / 10.1671 / 0272 - 4634 (2001) 021 [0687: TON AOA] 2.0. CO; 2" type="journal article" year="2001">Lukševičs 2001</bibRefCitation>
) and this surface has a prerostral process (
<figureCitation id="49D82A12BF1FE565FB2BCF56DEE43A0B" box="[1199,1277,1843,1870]" captionStart="FIG" captionStartId="7.[132,143,1408,1426]" captionTargetBox="[137,1452,218,1369]" captionTargetId="figure@7.[132,1455,214,1380]" captionTargetPageId="7" captionText="FIG. 3. — Asterolepis alticristata n. sp.: A-C, NUFV 1265, nuchal plate in dermal (A, photograph; A’, labeled illustration), visceral (B, photograph; B’, labeled illustration), and caudal (C) views; D-F, NUFV 813, premedian plate in dermal (D), visceral (E), and caudal (F) views; G-I, NUFV 812, premedian plate in dermal (G), visceral (H), and caudal (I) views. Abbreviations: cpl, central pit-line line groove; cv.L, area overlapping the lateral plate; cv.Pn, area overlapping the paranuchal plate; d.end, endolymphatic duct; ifc1, principal section of infraorbital pit-line groove; ml.p, midline process; mo.c, median occipital crista; nm, obtected area; pr.p, prerostral process; ps.f, parasagittal foramina; sop, supraoccipital pit; sot, supraotic thickening. Scale bar: 2 cm." figureDoi="http://doi.org/10.5281/zenodo.3695600" httpUri="https://zenodo.org/record/3695600/files/figure.png" pageId="7" pageNumber="684">Fig. 3F</figureCitation>
[pr.p]) across the midline with a small depression on the midline (the nasal notch of
<bibRefCitation id="B5724B66BF1FE565FC17CF16DE243AC8" author="LUKsEVICS E." box="[915,1085,1907,1933]" pageId="7" pageNumber="684" pagination="687 - 692" refId="ref17738" refString="LUKsEVICS E. 2001. - The orbito-nasal area of Asterolepis ornata, a Middle Devonian placoderm fish. Journal of Vertebrate Paleontology 21 (4): 687 - 692. https: // doi. org / 10.1671 / 0272 - 4634 (2001) 021 [0687: TON AOA] 2.0. CO; 2" type="journal article" year="2001">Lukševičs 2001</bibRefCitation>
). The prerostral process separates the orbital cavities. Both premedian specimens are widest at the lateral corners and these corners are closer to the rostral margin than they are to the caudal one.
</paragraph>
<caption id="859C661FBF10E56AFF00CB0ADE973EDD" ID-DOI="http://doi.org/10.5281/zenodo.3695602" ID-Zenodo-Dep="3695602" httpUri="https://zenodo.org/record/3695602/files/figure.png" pageId="8" pageNumber="685" startId="8.[132,143,879,897]" targetBox="[137,1451,215,835]" targetPageId="8">
<paragraph id="D15C3697BF10E56AFF00CB0ADE973EDD" blockId="8.[132,1455,879,920]" pageId="8" pageNumber="685">
FIG. 4. —
<taxonomicName id="16E34D14BF10E56AFF52CB0ADB953EC4" authority="Downs &amp; Daeschler &amp; Lo &amp; Carey &amp; Shubin, 2019" authorityName="Downs &amp; Daeschler &amp; Lo &amp; Carey &amp; Shubin" authorityYear="2019" box="[214,396,879,897]" class="Insecta" family="Tortricidae" genus="Asterolepis" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="685" phylum="Arthropoda" rank="species" species="alticristata" status="sp. nov.">
<emphasis id="E397EA85BF10E56AFF52CB0ADB953EC4" box="[214,396,879,897]" italics="true" pageId="8" pageNumber="685">Asterolepis alticristata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F8A457FEBF10E56AFE16CB0ADBD83EC4" box="[402,449,879,897]" pageId="8" pageNumber="685" rank="species">n. sp.</taxonomicNameLabel>
, NUFV 898, posterior median dorsal plate in dermal (
<emphasis id="E397EA85BF10E56AFC06CB0AD9963EC4" bold="true" box="[898,911,879,897]" pageId="8" pageNumber="685">A</emphasis>
), visceral (
<emphasis id="E397EA85BF10E56AFC6CCB0AD9EC3EC4" bold="true" box="[1000,1013,879,897]" pageId="8" pageNumber="685">B</emphasis>
), and cranial (
<emphasis id="E397EA85BF10E56AFBE9CB0ADE623EC4" bold="true" box="[1133,1147,879,897]" pageId="8" pageNumber="685">C</emphasis>
) views. Abbreviations:
<emphasis id="E397EA85BF10E56AFAB8CB0ADF993EC4" bold="true" box="[1340,1408,879,897]" pageId="8" pageNumber="685">cv.AMD</emphasis>
, area overlapping the anterior median dorsal plate;
<emphasis id="E397EA85BF10E56AFE7BCBE3D82F3EDD" bold="true" box="[511,566,902,920]" pageId="8" pageNumber="685">cv.Mxl</emphasis>
, area overlapping the mixilateral plate;
<emphasis id="E397EA85BF10E56AFCFACBE3D98C3EDD" bold="true" box="[894,917,902,920]" pageId="8" pageNumber="685">v.p</emphasis>
, visceral pit. Scale bar: 2 cm.
</paragraph>
</caption>
<paragraph id="D15C3697BF10E56AFF1FCBB3D85B39F5" blockId="8.[132,776,982,1806]" pageId="8" pageNumber="685">
The two premedian specimens referred to
<taxonomicName id="16E34D14BF10E56AFDC2CBB2D8D53EB5" authority="Downs &amp; Daeschler &amp; Lo &amp; Carey &amp; Shubin, 2019" authorityName="Downs &amp; Daeschler &amp; Lo &amp; Carey &amp; Shubin" authorityYear="2019" box="[582,716,982,1009]" class="Insecta" family="Tortricidae" genus="Asterolepis" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="685" phylum="Arthropoda" rank="species" species="alticristata" status="sp. nov.">
<emphasis id="E397EA85BF10E56AFDC2CBB2D8D53EB5" box="[582,716,982,1009]" italics="true" pageId="8" pageNumber="685">A. alticristata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F8A457FEBF10E56AFD54CBB2D91E3EB4" box="[720,775,983,1009]" pageId="8" pageNumber="685" rank="species">n. sp.</taxonomicNameLabel>
have a width/length index (as calculated for the nuchal above) of 134 (NUFV 812) and 157 (NUFV 813). These values are within the range of variation that
<bibRefCitation id="B5724B66BF10E56AFE74CC53D8BC3914" author="UPENIECE I." box="[496,677,1078,1105]" pageId="8" pageNumber="685" refId="ref18774" refString="UPENIECE I. 2011. - Palaeoecology and juvenile individuals of the Devonian placoderm and acanthodian fishes from Lode site, Latvia. (Doctoral dissertation). Retrieved from https: // dspace. lu. lv / dspace / handle / 7 / 4659 (last access on 3 rd October 2019)." type="url" year="2011">Upeniece (2011)</bibRefCitation>
reported for adult
<emphasis id="E397EA85BF10E56AFF69CC33DBBC3935" box="[237,421,1110,1136]" italics="true" pageId="8" pageNumber="685">
<taxonomicName id="16E34D14BF10E56AFF69CC33DB403935" authorityName="Eichwald, 1840 sensu Agassiz" authorityYear="1844" box="[237,345,1110,1136]" class="Insecta" family="Tortricidae" genus="Asterolepis" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="688" phylum="Arthropoda" rank="genus">Asterolepis</taxonomicName>
ornata
</emphasis>
(120-233). This broad range of values for adult
<emphasis id="E397EA85BF10E56AFEB0CC13DB8E39D5" box="[308,407,1142,1168]" italics="true" pageId="8" pageNumber="685">A. ornata</emphasis>
again highlights the variability of this plate within a single antiarch species.
</paragraph>
<paragraph id="D15C3697BF10E56AFF1FCCD3D89B3B6B" blockId="8.[132,776,982,1806]" pageId="8" pageNumber="685">
The principal section of the infraorbital pit-line groove (
<figureCitation id="49D82A12BF10E56AFF09CCB3DAC439B5" box="[141,221,1237,1264]" captionStart="FIG" captionStartId="7.[132,143,1408,1426]" captionTargetBox="[137,1452,218,1369]" captionTargetId="figure@7.[132,1455,214,1380]" captionTargetPageId="7" captionText="FIG. 3. — Asterolepis alticristata n. sp.: A-C, NUFV 1265, nuchal plate in dermal (A, photograph; A’, labeled illustration), visceral (B, photograph; B’, labeled illustration), and caudal (C) views; D-F, NUFV 813, premedian plate in dermal (D), visceral (E), and caudal (F) views; G-I, NUFV 812, premedian plate in dermal (G), visceral (H), and caudal (I) views. Abbreviations: cpl, central pit-line line groove; cv.L, area overlapping the lateral plate; cv.Pn, area overlapping the paranuchal plate; d.end, endolymphatic duct; ifc1, principal section of infraorbital pit-line groove; ml.p, midline process; mo.c, median occipital crista; nm, obtected area; pr.p, prerostral process; ps.f, parasagittal foramina; sop, supraoccipital pit; sot, supraotic thickening. Scale bar: 2 cm." figureDoi="http://doi.org/10.5281/zenodo.3695600" httpUri="https://zenodo.org/record/3695600/files/figure.png" pageId="8" pageNumber="685">Fig. 3D</figureCitation>
[ifc1], G[ifc1]) extends across the width of the premedian but follows a different path on each of the two specimens. In NUFV 812 (
<figureCitation id="49D82A12BF10E56AFEAECD70DB653875" box="[298,380,1301,1328]" captionStart="FIG" captionStartId="7.[132,143,1408,1426]" captionTargetBox="[137,1452,218,1369]" captionTargetId="figure@7.[132,1455,214,1380]" captionTargetPageId="7" captionText="FIG. 3. — Asterolepis alticristata n. sp.: A-C, NUFV 1265, nuchal plate in dermal (A, photograph; A’, labeled illustration), visceral (B, photograph; B’, labeled illustration), and caudal (C) views; D-F, NUFV 813, premedian plate in dermal (D), visceral (E), and caudal (F) views; G-I, NUFV 812, premedian plate in dermal (G), visceral (H), and caudal (I) views. Abbreviations: cpl, central pit-line line groove; cv.L, area overlapping the lateral plate; cv.Pn, area overlapping the paranuchal plate; d.end, endolymphatic duct; ifc1, principal section of infraorbital pit-line groove; ml.p, midline process; mo.c, median occipital crista; nm, obtected area; pr.p, prerostral process; ps.f, parasagittal foramina; sop, supraoccipital pit; sot, supraotic thickening. Scale bar: 2 cm." figureDoi="http://doi.org/10.5281/zenodo.3695600" httpUri="https://zenodo.org/record/3695600/files/figure.png" pageId="8" pageNumber="685">Fig. 3G</figureCitation>
), the groove forms a caudally directed point at the midline and strongly arches rostrally on either side of the midline. It is positioned at about mid-length along the plate, such that the midline dip of the groove reaches into the caudal half of the plate. In contrast, the groove on NUFV 813 (
<figureCitation id="49D82A12BF10E56AFF3ACDD0DB08388A" box="[190,273,1461,1487]" captionStart="FIG" captionStartId="7.[132,143,1408,1426]" captionTargetBox="[137,1452,218,1369]" captionTargetId="figure@7.[132,1455,214,1380]" captionTargetPageId="7" captionText="FIG. 3. — Asterolepis alticristata n. sp.: A-C, NUFV 1265, nuchal plate in dermal (A, photograph; A’, labeled illustration), visceral (B, photograph; B’, labeled illustration), and caudal (C) views; D-F, NUFV 813, premedian plate in dermal (D), visceral (E), and caudal (F) views; G-I, NUFV 812, premedian plate in dermal (G), visceral (H), and caudal (I) views. Abbreviations: cpl, central pit-line line groove; cv.L, area overlapping the lateral plate; cv.Pn, area overlapping the paranuchal plate; d.end, endolymphatic duct; ifc1, principal section of infraorbital pit-line groove; ml.p, midline process; mo.c, median occipital crista; nm, obtected area; pr.p, prerostral process; ps.f, parasagittal foramina; sop, supraoccipital pit; sot, supraotic thickening. Scale bar: 2 cm." figureDoi="http://doi.org/10.5281/zenodo.3695600" httpUri="https://zenodo.org/record/3695600/files/figure.png" pageId="8" pageNumber="685">Fig. 3D</figureCitation>
) has a shallow, rounded dip at the midline and is much straighter across its entire width. The groove is situated close to the rostral margin of the premedian such that no part of it reaches into the plate’s caudal half.
</paragraph>
<paragraph id="D15C3697BF10E56AFF1FCE51D9113A4B" blockId="8.[132,776,982,1806]" pageId="8" pageNumber="685">
Tiny foramina (
<figureCitation id="49D82A12BF10E56AFECFCE51DB853B0B" box="[331,412,1588,1615]" captionStart="FIG" captionStartId="7.[132,143,1408,1426]" captionTargetBox="[137,1452,218,1369]" captionTargetId="figure@7.[132,1455,214,1380]" captionTargetPageId="7" captionText="FIG. 3. — Asterolepis alticristata n. sp.: A-C, NUFV 1265, nuchal plate in dermal (A, photograph; A’, labeled illustration), visceral (B, photograph; B’, labeled illustration), and caudal (C) views; D-F, NUFV 813, premedian plate in dermal (D), visceral (E), and caudal (F) views; G-I, NUFV 812, premedian plate in dermal (G), visceral (H), and caudal (I) views. Abbreviations: cpl, central pit-line line groove; cv.L, area overlapping the lateral plate; cv.Pn, area overlapping the paranuchal plate; d.end, endolymphatic duct; ifc1, principal section of infraorbital pit-line groove; ml.p, midline process; mo.c, median occipital crista; nm, obtected area; pr.p, prerostral process; ps.f, parasagittal foramina; sop, supraoccipital pit; sot, supraotic thickening. Scale bar: 2 cm." figureDoi="http://doi.org/10.5281/zenodo.3695600" httpUri="https://zenodo.org/record/3695600/files/figure.png" pageId="8" pageNumber="685">Fig. 3E</figureCitation>
[ps.f], H[ps.f]) penetrate the visceral surface of the premedian; the two most prominent are parasagittally located in the rostral half of the plate. The external ornament of the premedian is similar to that of adult
<emphasis id="E397EA85BF10E56AFF00CED1DB223B8B" box="[132,315,1716,1742]" italics="true" pageId="8" pageNumber="685">
<taxonomicName id="16E34D14BF10E56AFF00CED1DAF63B8B" authorityName="Eichwald, 1840 sensu Agassiz" authorityYear="1844" box="[132,239,1716,1742]" class="Insecta" family="Tortricidae" genus="Asterolepis" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="688" phylum="Arthropoda" rank="genus">Asterolepis</taxonomicName>
ornata
</emphasis>
in that rows of tubercles radiate out from the center of ossification. Close to the lateral margins of the plate, the tubercles within a row fuse together to
<taxonomicName id="16E34D14BF10E56AFD02CE91D91A3A4B" box="[646,771,1780,1806]" form="ridges" pageId="8" pageNumber="685" rank="form">form ridges</taxonomicName>
.
</paragraph>
<paragraph id="D15C3697BF10E56AFF00CF56D8183A2B" blockId="8.[132,775,1843,2030]" pageId="8" pageNumber="685">
DESCRIPTION OF THORACIC AND PECTORAL SKELETONS
<emphasis id="E397EA85BF10E56AFF00CF31D8183A2B" box="[132,513,1875,1902]" italics="true" pageId="8" pageNumber="685">
Anterior median dorsal plate (
<figureCitation id="49D82A12BF10E56AFE3FCF31DBE13A2B" box="[443,504,1876,1902]" captionStart="FIG" captionStartId="6.[132,143,893,911]" captionTargetBox="[137,1449,215,847]" captionTargetId="figure@6.[132,1455,214,867]" captionTargetPageId="6" captionText="FIG. 2. — Asterolepis alticristata n. sp., NUFV 897, holotype, anterior median dorsal plate in visceral (A, photograph; A’, labeled illustration), cranial (B), left lateral (C), and dermal (D) views. Abbreviations:cd.PMD, area overlapped by the posterior median dorsal plate; cv.ADL, area overlapping the anterior dorsolateral plate; cv.Mxl, area overlapping the mixilateral plate; lc, lateral corner; op.r oblique posterolateral ridge. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.3695598" httpUri="https://zenodo.org/record/3695598/files/figure.png" pageId="8" pageNumber="685">Fig. 2</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="D15C3697BF10E56AFF00CF16D91E3AA8" blockId="8.[132,775,1843,2030]" pageId="8" pageNumber="685">
The anterior median dorsal plate of
<taxonomicName id="16E34D14BF10E56AFE77CF16D8D23AC8" authority="Downs &amp; Daeschler &amp; Lo &amp; Carey &amp; Shubin, 2019" authorityName="Downs &amp; Daeschler &amp; Lo &amp; Carey &amp; Shubin" authorityYear="2019" box="[499,715,1907,1933]" class="Insecta" family="Tortricidae" genus="Asterolepis" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="685" phylum="Arthropoda" rank="species" species="alticristata" status="sp. nov.">
<emphasis id="E397EA85BF10E56AFE77CF16D8D23AC8" box="[499,715,1907,1933]" italics="true" pageId="8" pageNumber="685">Asterolepis alticristata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F8A457FEBF10E56AFD4BCF11D91E3ACB" box="[719,775,1908,1934]" pageId="8" pageNumber="685" rank="species">n. sp.</taxonomicNameLabel>
features a tall rounded crest on the midline. The remarkable height (
<emphasis id="E397EA85BF10E56AFF53CFD1DAF13A88" box="[215,232,1972,1997]" italics="true" pageId="8" pageNumber="685">c.</emphasis>
<quantity id="161B9B72BF10E56AFF69CFD6DB223A8B" box="[237,315,1971,1998]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="8" pageNumber="685" unit="mm" value="60.0">60 mm</quantity>
in the largest specimen, NUFV 897, maximum anterior median dorsal plate width =
<quantity id="161B9B72BF10E56AFDD4CFB6D8C93AAB" box="[592,720,2003,2030]" metricMagnitude="-1" metricUnit="m" metricValue="1.1255" pageId="8" pageNumber="685" unit="mm" value="112.55">112.55 mm</quantity>
) and
</paragraph>
<paragraph id="D15C3697BF10E56AFCA9CBB3DFB63A6B" blockId="8.[813,1456,982,2030]" pageId="8" pageNumber="685">
thickness (
<emphasis id="E397EA85BF10E56AFC1ACBB2D9B63EB5" box="[926,943,983,1008]" italics="true" pageId="8" pageNumber="685">c.</emphasis>
<quantity id="161B9B72BF10E56AFC30CBB2DE1B3EB4" box="[948,1026,983,1009]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="8" pageNumber="685" unit="mm" value="18.0">18 mm</quantity>
in NUFV 897) of the crest is unlike any previously described for
<taxonomicName id="16E34D14BF10E56AFBB9CB93DEB03955" authorityName="Eichwald, 1840 sensu Agassiz" authorityYear="1844" box="[1085,1193,1014,1040]" class="Insecta" family="Tortricidae" genus="Asterolepis" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="688" phylum="Arthropoda" rank="genus">
<emphasis id="E397EA85BF10E56AFBB9CB93DEB03955" box="[1085,1193,1014,1040]" italics="true" pageId="8" pageNumber="685">Asterolepis</emphasis>
</taxonomicName>
(
<figureCitation id="49D82A12BF10E56AFB3FCB93DEE23954" box="[1211,1275,1014,1041]" captionStart="FIG" captionStartId="6.[132,143,893,911]" captionTargetBox="[137,1449,215,847]" captionTargetId="figure@6.[132,1455,214,867]" captionTargetPageId="6" captionText="FIG. 2. — Asterolepis alticristata n. sp., NUFV 897, holotype, anterior median dorsal plate in visceral (A, photograph; A’, labeled illustration), cranial (B), left lateral (C), and dermal (D) views. Abbreviations:cd.PMD, area overlapped by the posterior median dorsal plate; cv.ADL, area overlapping the anterior dorsolateral plate; cv.Mxl, area overlapping the mixilateral plate; lc, lateral corner; op.r oblique posterolateral ridge. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.3695598" httpUri="https://zenodo.org/record/3695598/files/figure.png" pageId="8" pageNumber="685">Fig. 2</figureCitation>
). The crest rises sharply from a position just cranial to the tergal angle and reaches its maximum height just caudal to the center of the anterior median dorsal plate, nearly in line with the plate’s greatest width. The long dorsal margin of the crest caudal to the maximum height is slightly convex to deeply concave. The closest point of comparison among antiarchs, for a crest in this position and of this height, is in
<emphasis id="E397EA85BF10E56AFB66CCB3DFB639AA" box="[1250,1455,1238,1264]" italics="true" pageId="8" pageNumber="685">Bothriolepis cristata</emphasis>
<bibRefCitation id="B5724B66BF10E56AFCA9CC90D9D53855" author="TRAQUAIR R. H." box="[813,972,1269,1296]" pageId="8" pageNumber="685" pagination="235 - 285" refId="ref18633" refString="TRAQUAIR R. H. 1895. - The extinct vertebrate animals of the Moray Firth area in HARVIE- BROWN J. A. &amp; BUCKLEY T. A. (eds), A Vertebrate Fauna of the Moray Firth Basin. Volume 2. T. and A. Constable, Edinburgh: 235 - 285. https: // doi. org / 10.5962 / bhl. title. 69239" type="book chapter" year="1895">Traquair, 1895</bibRefCitation>
(
<collectingCountry id="A9F47607BF10E56AFC5FCC90DE593855" box="[987,1088,1269,1296]" name="United Kingdom" pageId="8" pageNumber="685">Scotland</collectingCountry>
). A tall dorsal crest also appears in
<emphasis id="E397EA85BF10E56AFCA9CD70DE5C386A" box="[813,1093,1301,1327]" italics="true" pageId="8" pageNumber="685">Bothriolepis gippslandiensis</emphasis>
<bibRefCitation id="B5724B66BF10E56AFBCACD70DED03875" author="HILLS E. S." box="[1102,1225,1301,1328]" pageId="8" pageNumber="685" pagination="176 - 201" refId="ref17351" refString="HILLS E. S. 1929. - The geology and palaeontography of the Cathedral Range and Blue Hills in north-western Gippsland. Proceedings of the Royal Society of Victoria 41: 176 - 201. https: // biodiversitylibrary. org / page / 54367530" type="journal article" year="1929">Hills, 1929</bibRefCitation>
(
<collectingRegion id="1327F875BF10E56AFB5CCD70DF2D386A" box="[1240,1332,1301,1327]" country="Australia" name="Victoria" pageId="8" pageNumber="685">Victoria</collectingRegion>
,
<collectingCountry id="A9F47607BF10E56AFAC4CD70DFBD3875" box="[1344,1444,1301,1328]" name="Australia" pageId="8" pageNumber="685">Australia</collectingCountry>
);
<emphasis id="E397EA85BF10E56AFCA9CD50DE35380A" box="[813,1068,1333,1359]" italics="true" pageId="8" pageNumber="685">Bothriolepis cullodenensis</emphasis>
<bibRefCitation id="B5724B66BF10E56AFBB5CD53DEB2380A" author="LONG J. A." box="[1073,1195,1333,1360]" pageId="8" pageNumber="685" pagination="295 - 320" refId="ref17698" refString="LONG J. A. 1983. - New bothriolepid fish from the Late Devonian of Victoria, Australia. Palaeontology 26 (2): 295 - 320. https: // biodiversitylibrary. org / page / 49741248" type="journal article" year="1983">Long, 1983</bibRefCitation>
(
<collectingRegion id="1327F875BF10E56AFB3CCD50DF09380A" box="[1208,1296,1333,1359]" country="Australia" name="Victoria" pageId="8" pageNumber="685">Victoria</collectingRegion>
<collectingCountry id="A9F47607BF10E56AFA91CD50DF6E3815" box="[1301,1399,1333,1360]" name="Australia" pageId="8" pageNumber="685">Australia</collectingCountry>
); and
<emphasis id="E397EA85BF10E56AFCA9CD30DE11382A" box="[813,1032,1365,1391]" italics="true" pageId="8" pageNumber="685">Bothriolepis zadonica</emphasis>
<bibRefCitation id="B5724B66BF10E56AFB8BCD30DEF8382A" author="MOLOSHNIKOV S." box="[1039,1249,1365,1392]" pageId="8" pageNumber="685" pagination="135 - 146" refId="ref17997" refString="MOLOSHNIKOV S. 2004. - Crested antiarch Bothriolepis zadonica H. D. Obrucheva from the Lower Famennian of Central European Russia. Acta Palaeontologica Polonica 49 (1): 135 - 146." type="journal article" year="2004">Moloshnikov, 2004</bibRefCitation>
(Central European
<collectingCountry id="A9F47607BF10E56AFCA9CD10D96E38D5" box="[813,887,1397,1424]" name="Russia" pageId="8" pageNumber="685">Russia</collectingCountry>
). In each of these three examples, however, the crest is formed nearly equally by anterior median dorsal and posterior median dorsal plates; the crest rises from near the anterior median dorsal plate’s tergal angle and reaches its maximum height only at the contact between anterior median dorsal and posterior median dorsal plates (
<bibRefCitation id="B5724B66BF10E56AFB35CE71DF993B6A" author="MOLOSHNIKOV S." box="[1201,1408,1556,1583]" pageId="8" pageNumber="685" pagination="135 - 146" refId="ref17997" refString="MOLOSHNIKOV S. 2004. - Crested antiarch Bothriolepis zadonica H. D. Obrucheva from the Lower Famennian of Central European Russia. Acta Palaeontologica Polonica 49 (1): 135 - 146." type="journal article" year="2004">Moloshnikov 2004</bibRefCitation>
). In addition to the three partial anterior median dorsal plates with crests (NUFV 822, 896, 897 [
<typeStatus id="0E588835BF10E56AFB30CE31DF053B2A" box="[1204,1308,1620,1647]" pageId="8" pageNumber="685" type="holotype">holotype</typeStatus>
]), the
<taxonomicName id="16E34D14BF10E56AFAE7CE30D9613BCB" authority="Downs &amp; Daeschler &amp; Lo &amp; Carey &amp; Shubin, 2019" authorityName="Downs &amp; Daeschler &amp; Lo &amp; Carey &amp; Shubin" authorityYear="2019" class="Insecta" family="Tortricidae" genus="Asterolepis" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="685" phylum="Arthropoda" rank="species" species="alticristata" status="sp. nov.">
<emphasis id="E397EA85BF10E56AFAE7CE30D9613BCB" italics="true" pageId="8" pageNumber="685">A. alticristata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F8A457FEBF10E56AFCFACE10D9A03BCA" box="[894,953,1653,1679]" pageId="8" pageNumber="685" rank="species">n. sp.</taxonomicNameLabel>
referred material includes thirty-seven isolated crests from anterior median dorsal plates. No other part of the
<taxonomicName id="16E34D14BF10E56AFCD2CED1D9FA3B8B" authority="Downs &amp; Daeschler &amp; Lo &amp; Carey &amp; Shubin, 2019" authorityName="Downs &amp; Daeschler &amp; Lo &amp; Carey &amp; Shubin" authorityYear="2019" box="[854,995,1716,1742]" class="Insecta" family="Tortricidae" genus="Asterolepis" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="685" phylum="Arthropoda" rank="species" species="alticristata" status="sp. nov.">
<emphasis id="E397EA85BF10E56AFCD2CED1D9FA3B8B" box="[854,995,1716,1742]" italics="true" pageId="8" pageNumber="685">A. alticristata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F8A457FEBF10E56AFC6ECED0DE3C3B8A" box="[1002,1061,1717,1743]" pageId="8" pageNumber="685" rank="species">n. sp.</taxonomicNameLabel>
anatomy is better represented in the sample. The size and shape variation among these crests does not fit a single growth trend (See Discussion: Variation in anterior median dorsal plate crests of
<taxonomicName id="16E34D14BF10E56AFB02CF71DF7B3A6B" authority="Downs &amp; Daeschler &amp; Lo &amp; Carey &amp; Shubin, 2019" authorityName="Downs &amp; Daeschler &amp; Lo &amp; Carey &amp; Shubin" authorityYear="2019" box="[1158,1378,1812,1838]" class="Insecta" family="Tortricidae" genus="Asterolepis" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="685" phylum="Arthropoda" rank="species" species="alticristata" status="sp. nov.">
<emphasis id="E397EA85BF10E56AFB02CF71DF7B3A6B" box="[1158,1378,1812,1838]" italics="true" pageId="8" pageNumber="685">Asterolepis alticristata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F8A457FEBF10E56AFAE3CF71DFBB3A6B" box="[1383,1442,1812,1838]" pageId="8" pageNumber="685" rank="species">n. sp.</taxonomicNameLabel>
).
</paragraph>
<paragraph id="D15C3697BF10E56BFCC7CF51DB7D3F37" blockId="8.[813,1456,982,2030]" lastBlockId="9.[132,776,217,1328]" lastPageId="9" lastPageNumber="686" pageId="8" pageNumber="685">
At the caudal end of the anterior median dorsal plate, the midline crest continues onto the posterior median dorsal plate. In NUFV 897 (maximum crest height
<emphasis id="E397EA85BF10E56AFAB5CF11DF5A3AC8" box="[1329,1347,1908,1933]" italics="true" pageId="8" pageNumber="685">c.</emphasis>
<quantity id="161B9B72BF10E56AFAC8CF16DFBA3ACB" box="[1356,1443,1907,1934]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="8" pageNumber="685" unit="mm" value="60.0">60 mm</quantity>
), the height of the anterior median dorsal plate crest at its caudal terminus is
<emphasis id="E397EA85BF10E56AFC7ACFD1DE163A88" box="[1022,1039,1972,1997]" italics="true" pageId="8" pageNumber="685">c.</emphasis>
<quantity id="161B9B72BF10E56AFB93CFD6DE753A8B" box="[1047,1132,1971,1998]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="8" pageNumber="685" unit="mm" value="14.0">14 mm</quantity>
. The total length of the anterior median dorsal plate crest on NUFV 897, from where it first rises to its contact with the crest of the posterior median dorsal plate is
<quantity id="161B9B72BF11E56BFE0CC89CD80B3C51" box="[392,530,249,276]" metricMagnitude="-1" metricUnit="m" metricValue="1.081" pageId="9" pageNumber="686" unit="mm" value="108.1">108.10 mm</quantity>
. The anterior median dorsal plate of
<taxonomicName id="16E34D14BF11E56BFEACC97CDBA33C76" authority="Downs &amp; Daeschler &amp; Lo &amp; Carey &amp; Shubin, 2019" authorityName="Downs &amp; Daeschler &amp; Lo &amp; Carey &amp; Shubin" authorityYear="2019" box="[296,442,281,307]" class="Insecta" family="Tortricidae" genus="Asterolepis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="686" phylum="Arthropoda" rank="species" species="alticristata" status="sp. nov.">
<emphasis id="E397EA85BF11E56BFEACC97CDBA33C76" box="[296,442,281,307]" italics="true" pageId="9" pageNumber="686">A. alticristata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F8A457FEBF11E56BFE45C97FDBE43C71" box="[449,509,282,308]" pageId="9" pageNumber="686" rank="species">n. sp.</taxonomicNameLabel>
is steeply vaulted in the transverse plane such that the dorsal surfaces lateral to the crest face distinctly craniolaterally. The posterior median dorsal plate overlaps an unornamented, depressed zone along the caudal end of the anterior median dorsal plate’s dorsal surface (
<figureCitation id="49D82A12BF11E56BFEABC9DDDB913C96" box="[303,392,440,467]" captionStart="FIG" captionStartId="8.[132,143,879,897]" captionTargetBox="[137,1451,215,835]" captionTargetId="figure@8.[132,1455,214,845]" captionTargetPageId="8" captionText="FIG. 4. — Asterolepis alticristata n. sp., NUFV 898, posterior median dorsal plate in dermal (A), visceral (B), and cranial (C) views. Abbreviations: cv.AMD, area overlapping the anterior median dorsal plate; cv.Mxl, area overlapping the mixilateral plate; v.p, visceral pit. Scale bar: 2 cm." figureDoi="http://doi.org/10.5281/zenodo.3695602" httpUri="https://zenodo.org/record/3695602/files/figure.png" pageId="9" pageNumber="686">Fig. 4D</figureCitation>
[cd.PMD]). None of the available material allows description of the anterior median dorsal plate cranial to the crest. In NUFV 897, the tergal angle is cranial to the lateral corners (
<figureCitation id="49D82A12BF11E56BFE50CA7DD80C3F77" box="[468,533,536,562]" captionStart="FIG" captionStartId="6.[132,143,893,911]" captionTargetBox="[137,1449,215,847]" captionTargetId="figure@6.[132,1455,214,867]" captionTargetPageId="6" captionText="FIG. 2. — Asterolepis alticristata n. sp., NUFV 897, holotype, anterior median dorsal plate in visceral (A, photograph; A’, labeled illustration), cranial (B), left lateral (C), and dermal (D) views. Abbreviations:cd.PMD, area overlapped by the posterior median dorsal plate; cv.ADL, area overlapping the anterior dorsolateral plate; cv.Mxl, area overlapping the mixilateral plate; lc, lateral corner; op.r oblique posterolateral ridge. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.3695598" httpUri="https://zenodo.org/record/3695598/files/figure.png" pageId="9" pageNumber="686">Fig. 2</figureCitation>
A’[lc]) of the anterior median dorsal plate and caudal to the most cranial position of the midline crest.
</paragraph>
<paragraph id="D15C3697BF11E56BFF1FCA1DD86C3E94" blockId="9.[132,776,217,1328]" pageId="9" pageNumber="686">
The visceral surface of the anterior median dorsal plate shows the depressed surfaces where the plate overlaps the anterior dorsolateral (
<figureCitation id="49D82A12BF11E56BFE05CAD2DBDC3F97" box="[385,453,695,722]" captionStart="FIG" captionStartId="6.[132,143,893,911]" captionTargetBox="[137,1449,215,847]" captionTargetId="figure@6.[132,1455,214,867]" captionTargetPageId="6" captionText="FIG. 2. — Asterolepis alticristata n. sp., NUFV 897, holotype, anterior median dorsal plate in visceral (A, photograph; A’, labeled illustration), cranial (B), left lateral (C), and dermal (D) views. Abbreviations:cd.PMD, area overlapped by the posterior median dorsal plate; cv.ADL, area overlapping the anterior dorsolateral plate; cv.Mxl, area overlapping the mixilateral plate; lc, lateral corner; op.r oblique posterolateral ridge. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.3695598" httpUri="https://zenodo.org/record/3695598/files/figure.png" pageId="9" pageNumber="686">Fig. 2</figureCitation>
A’[cv.ADL]) and mixilateral (
<figureCitation id="49D82A12BF11E56BFF0ACAB2DAD43FB7" box="[142,205,727,754]" captionStart="FIG" captionStartId="6.[132,143,893,911]" captionTargetBox="[137,1449,215,847]" captionTargetId="figure@6.[132,1455,214,867]" captionTargetPageId="6" captionText="FIG. 2. — Asterolepis alticristata n. sp., NUFV 897, holotype, anterior median dorsal plate in visceral (A, photograph; A’, labeled illustration), cranial (B), left lateral (C), and dermal (D) views. Abbreviations:cd.PMD, area overlapped by the posterior median dorsal plate; cv.ADL, area overlapping the anterior dorsolateral plate; cv.Mxl, area overlapping the mixilateral plate; lc, lateral corner; op.r oblique posterolateral ridge. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.3695598" httpUri="https://zenodo.org/record/3695598/files/figure.png" pageId="9" pageNumber="686">Fig. 2</figureCitation>
A’[cv.Mxl]) plates cranial and caudal to the anterior median dorsal lateral corners. On each side, these overlap surfaces are continuous with one another across the lateral corner. Obliquely oriented, low, wide ridges (oblique posterolateral ridge of Karatajūte-Talimaa 1963;
<figureCitation id="49D82A12BF11E56BFDE8CB32D8B03E34" box="[620,681,855,881]" captionStart="FIG" captionStartId="8.[132,143,879,897]" captionTargetBox="[137,1451,215,835]" captionTargetId="figure@8.[132,1455,214,845]" captionTargetPageId="8" captionText="FIG. 4. — Asterolepis alticristata n. sp., NUFV 898, posterior median dorsal plate in dermal (A), visceral (B), and cranial (C) views. Abbreviations: cv.AMD, area overlapping the anterior median dorsal plate; cv.Mxl, area overlapping the mixilateral plate; v.p, visceral pit. Scale bar: 2 cm." figureDoi="http://doi.org/10.5281/zenodo.3695602" httpUri="https://zenodo.org/record/3695602/files/figure.png" pageId="9" pageNumber="686">Fig. 4</figureCitation>
A’[op.r]) extend and widen from the center of the anterior median dorsal plate’s visceral surface to the mixilateral underlap close to the caudolateral corners of the plate.
</paragraph>
<paragraph id="D15C3697BF11E56BFF1FCBB3D9113875" blockId="9.[132,776,217,1328]" pageId="9" pageNumber="686">
The ornament of the dorsolateral faces of the anterior median dorsal plate is densely tuberculate centrally and ridged closer to the lateral and caudal margins of the plate. The ridges retain the appearance of fused tubercles and radiate from the center. The ornament of the crest is densely tuberculate especially at the base. The tubercles are disorganized at the base, continuing the ornament
<typeStatus id="0E588835BF11E56BFD09CCF3D8A539F5" box="[653,700,1174,1200]" pageId="9" pageNumber="686">type</typeStatus>
of the plate’s dorsolateral surfaces, and are organized into rows more distally. In one specimen (NUFV 872), these organized tubercles are fused into high ridges. The rows of tubercles or ridges radiate from a cranioproximal position on the crest.
</paragraph>
<paragraph id="D15C3697BF11E56BFF00CD33D808382A" blockId="9.[132,776,1365,2030]" box="[132,529,1365,1391]" pageId="9" pageNumber="686">
<emphasis id="E397EA85BF11E56BFF00CD33D808382A" box="[132,529,1365,1391]" italics="true" pageId="9" pageNumber="686">
Posterior median dorsal plate (
<figureCitation id="49D82A12BF11E56BFE4CCD33D81E382A" box="[456,519,1365,1391]" captionStart="FIG" captionStartId="8.[132,143,879,897]" captionTargetBox="[137,1451,215,835]" captionTargetId="figure@8.[132,1455,214,845]" captionTargetPageId="8" captionText="FIG. 4. — Asterolepis alticristata n. sp., NUFV 898, posterior median dorsal plate in dermal (A), visceral (B), and cranial (C) views. Abbreviations: cv.AMD, area overlapping the anterior median dorsal plate; cv.Mxl, area overlapping the mixilateral plate; v.p, visceral pit. Scale bar: 2 cm." figureDoi="http://doi.org/10.5281/zenodo.3695602" httpUri="https://zenodo.org/record/3695602/files/figure.png" pageId="9" pageNumber="686">Fig. 4</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="D15C3697BF11E56BFF00CD10D8F73ACB" blockId="9.[132,776,1365,2030]" pageId="9" pageNumber="686">
The posterior median dorsal plate of
<taxonomicName id="16E34D14BF11E56BFD9BCD10D91E38CA" authority="Downs &amp; Daeschler &amp; Lo &amp; Carey &amp; Shubin, 2019" authorityName="Downs &amp; Daeschler &amp; Lo &amp; Carey &amp; Shubin" authorityYear="2019" box="[543,775,1397,1423]" class="Insecta" family="Tortricidae" genus="Asterolepis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="686" phylum="Arthropoda" rank="species" species="alticristata" status="sp. nov.">
<emphasis id="E397EA85BF11E56BFD9BCD10D91E38CA" box="[543,775,1397,1423]" italics="true" pageId="9" pageNumber="686">Asterolepis alticristata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F8A457FEBF11E56BFF00CDF3DAA638F5" box="[132,191,1430,1456]" pageId="9" pageNumber="686" rank="species">n. sp.</taxonomicNameLabel>
is elongate and narrow. The greatest width of the plate is across the lateral corners; these corners are located close to the caudal margin of the plate. There is little increase in width from craniolateral to caudolateral corners and much of that increase occurs abruptly just cranial to the caudolateral corner. With nearly parallel lateral margins and craniolateral margins that angle toward the midline, the posterior median dorsal plate of
<taxonomicName id="16E34D14BF11E56BFEAFCE10DBA43BCB" authority="Downs &amp; Daeschler &amp; Lo &amp; Carey &amp; Shubin, 2019" authorityName="Downs &amp; Daeschler &amp; Lo &amp; Carey &amp; Shubin" authorityYear="2019" box="[299,445,1652,1679]" class="Insecta" family="Tortricidae" genus="Asterolepis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="686" phylum="Arthropoda" rank="species" species="alticristata" status="sp. nov.">
<emphasis id="E397EA85BF11E56BFEAFCE10DBA43BCB" box="[299,445,1652,1679]" italics="true" pageId="9" pageNumber="686">A. alticristata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F8A457FEBF11E56BFE41CE10D81B3BCA" box="[453,514,1653,1679]" pageId="9" pageNumber="686" rank="species">n. sp.</taxonomicNameLabel>
is more obelisk-shaped in dorsal view than the bulging diamond-shaped posterior median dorsal plate of
<emphasis id="E397EA85BF11E56BFE0CCED1D85C3B8B" box="[392,581,1716,1742]" italics="true" pageId="9" pageNumber="686">
<taxonomicName id="16E34D14BF11E56BFE0CCED1DBEE3B8B" authorityName="Eichwald, 1840 sensu Agassiz" authorityYear="1844" box="[392,503,1716,1742]" class="Insecta" family="Tortricidae" genus="Asterolepis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="688" phylum="Arthropoda" rank="genus">Asterolepis</taxonomicName>
ornata
</emphasis>
(
<bibRefCitation id="B5724B66BF11E56BFDD3CED1D8E63B8B" author="LYARSKAJA L. A." box="[599,767,1716,1742]" pageId="9" pageNumber="686" refId="ref17797" refString="LYARSKAJA L. A. 1981. - Baltic Devonian Placodermi: Asterolepididae. Zinatne, Riga, Latvia, 152 p. [in Russian with English summary]." type="book" year="1981">Lyarskaja 1981</bibRefCitation>
: pl. 30, figs 1, 3). As is typical for
<taxonomicName id="16E34D14BF11E56BFE78CEB1D8723BAB" authorityName="Eichwald, 1840 sensu Agassiz" authorityYear="1844" box="[508,619,1748,1774]" class="Insecta" family="Tortricidae" genus="Asterolepis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="688" phylum="Arthropoda" rank="genus">
<emphasis id="E397EA85BF11E56BFE78CEB1D8723BAB" box="[508,619,1748,1774]" italics="true" pageId="9" pageNumber="686">Asterolepis</emphasis>
</taxonomicName>
, the posterior median dorsal plate broadly overlaps both anterior median dorsal (
<figureCitation id="49D82A12BF11E56BFF51CF71DB3D3A6B" box="[213,292,1811,1838]" captionStart="FIG" captionStartId="8.[132,143,879,897]" captionTargetBox="[137,1451,215,835]" captionTargetId="figure@8.[132,1455,214,845]" captionTargetPageId="8" captionText="FIG. 4. — Asterolepis alticristata n. sp., NUFV 898, posterior median dorsal plate in dermal (A), visceral (B), and cranial (C) views. Abbreviations: cv.AMD, area overlapping the anterior median dorsal plate; cv.Mxl, area overlapping the mixilateral plate; v.p, visceral pit. Scale bar: 2 cm." figureDoi="http://doi.org/10.5281/zenodo.3695602" httpUri="https://zenodo.org/record/3695602/files/figure.png" pageId="9" pageNumber="686">Fig. 4B</figureCitation>
[cv.AMD]) and mixilateral (
<figureCitation id="49D82A12BF11E56BFDD7CF71D8BB3A6B" box="[595,674,1811,1838]" captionStart="FIG" captionStartId="8.[132,143,879,897]" captionTargetBox="[137,1451,215,835]" captionTargetId="figure@8.[132,1455,214,845]" captionTargetPageId="8" captionText="FIG. 4. — Asterolepis alticristata n. sp., NUFV 898, posterior median dorsal plate in dermal (A), visceral (B), and cranial (C) views. Abbreviations: cv.AMD, area overlapping the anterior median dorsal plate; cv.Mxl, area overlapping the mixilateral plate; v.p, visceral pit. Scale bar: 2 cm." figureDoi="http://doi.org/10.5281/zenodo.3695602" httpUri="https://zenodo.org/record/3695602/files/figure.png" pageId="9" pageNumber="686">Fig. 4B</figureCitation>
[cv.Mxl]) plates with the deepest zones of overlap at the rostral midline (onto the posterior median dorsal plate) and in the positions of the caudolateral corners (onto the mixilateral plates).
</paragraph>
<paragraph id="D15C3697BF11E56BFF1FCFF6DF0C3F17" blockId="9.[132,776,1365,2030]" lastBlockId="9.[813,1456,217,850]" pageId="9" pageNumber="686">
A low midline crest extends along the entire length of the plate that shallows from its cranial to its caudal end. There is a sharp division between the narrow peak of the posterior median dorsal plate’s external tissue layers and the more shallow vaulting of the plate’s visceral surface. Between the two there is a mass of bone tissue that fills the vaulted core of the peak (
<figureCitation id="49D82A12BF11E56BFC3AC95CDE0E3C16" box="[958,1047,313,339]" captionStart="FIG" captionStartId="8.[132,143,879,897]" captionTargetBox="[137,1451,215,835]" captionTargetId="figure@8.[132,1455,214,845]" captionTargetPageId="8" captionText="FIG. 4. — Asterolepis alticristata n. sp., NUFV 898, posterior median dorsal plate in dermal (A), visceral (B), and cranial (C) views. Abbreviations: cv.AMD, area overlapping the anterior median dorsal plate; cv.Mxl, area overlapping the mixilateral plate; v.p, visceral pit. Scale bar: 2 cm." figureDoi="http://doi.org/10.5281/zenodo.3695602" httpUri="https://zenodo.org/record/3695602/files/figure.png" pageId="9" pageNumber="686">Fig. 4C</figureCitation>
). The division between external and internal tissue layers results in a dramatic thickening of the plate along the midline, from the peak of the midline crest to the smooth visceral surface. The non-overlapping portion of the visceral surface (
<figureCitation id="49D82A12BF11E56BFBA3C9DDDE613C96" box="[1063,1144,440,467]" captionStart="FIG" captionStartId="8.[132,143,879,897]" captionTargetBox="[137,1451,215,835]" captionTargetId="figure@8.[132,1455,214,845]" captionTargetPageId="8" captionText="FIG. 4. — Asterolepis alticristata n. sp., NUFV 898, posterior median dorsal plate in dermal (A), visceral (B), and cranial (C) views. Abbreviations: cv.AMD, area overlapping the anterior median dorsal plate; cv.Mxl, area overlapping the mixilateral plate; v.p, visceral pit. Scale bar: 2 cm." figureDoi="http://doi.org/10.5281/zenodo.3695602" httpUri="https://zenodo.org/record/3695602/files/figure.png" pageId="9" pageNumber="686">Fig. 4B</figureCitation>
) is generally rectangular but for a broadening at its caudal end. On the visceral midline, near the caudal end, there is a small circular pit (
<figureCitation id="49D82A12BF11E56BFAACC99DDF6E3F56" box="[1320,1399,504,531]" captionStart="FIG" captionStartId="8.[132,143,879,897]" captionTargetBox="[137,1451,215,835]" captionTargetId="figure@8.[132,1455,214,845]" captionTargetPageId="8" captionText="FIG. 4. — Asterolepis alticristata n. sp., NUFV 898, posterior median dorsal plate in dermal (A), visceral (B), and cranial (C) views. Abbreviations: cv.AMD, area overlapping the anterior median dorsal plate; cv.Mxl, area overlapping the mixilateral plate; v.p, visceral pit. Scale bar: 2 cm." figureDoi="http://doi.org/10.5281/zenodo.3695602" httpUri="https://zenodo.org/record/3695602/files/figure.png" pageId="9" pageNumber="686">Fig. 4B</figureCitation>
[v.p]) that is surrounded by parasagittal depressions cranial to it and a single midline depression caudal to it.
</paragraph>
<paragraph id="D15C3697BF11E56BFCC7CA3DDFA93E14" blockId="9.[813,1456,217,850]" pageId="9" pageNumber="686">
The ornament is densely tuberculate. The tubercles are tightly-spaced and arranged into rows in locations close to the plate’s margin; these rows are parallel to the plate’s lateral edges in a small specimen (NUFV 796) and perpendicular to the cranial and lateral margins in large specimens (NUFV 874, 898;
<figureCitation id="49D82A12BF11E56BFC25CA92D9F83E54" box="[929,993,759,785]" captionStart="FIG" captionStartId="8.[132,143,879,897]" captionTargetBox="[137,1451,215,835]" captionTargetId="figure@8.[132,1455,214,845]" captionTargetPageId="8" captionText="FIG. 4. — Asterolepis alticristata n. sp., NUFV 898, posterior median dorsal plate in dermal (A), visceral (B), and cranial (C) views. Abbreviations: cv.AMD, area overlapping the anterior median dorsal plate; cv.Mxl, area overlapping the mixilateral plate; v.p, visceral pit. Scale bar: 2 cm." figureDoi="http://doi.org/10.5281/zenodo.3695602" httpUri="https://zenodo.org/record/3695602/files/figure.png" pageId="9" pageNumber="686">Fig. 4</figureCitation>
). In positions along the lateral margin of the largest known posterior median dorsal plate (NUFV 898;
<figureCitation id="49D82A12BF11E56BFCE0CB52D9BB3E14" box="[868,930,823,849]" captionStart="FIG" captionStartId="8.[132,143,879,897]" captionTargetBox="[137,1451,215,835]" captionTargetId="figure@8.[132,1455,214,845]" captionTargetPageId="8" captionText="FIG. 4. — Asterolepis alticristata n. sp., NUFV 898, posterior median dorsal plate in dermal (A), visceral (B), and cranial (C) views. Abbreviations: cv.AMD, area overlapping the anterior median dorsal plate; cv.Mxl, area overlapping the mixilateral plate; v.p, visceral pit. Scale bar: 2 cm." figureDoi="http://doi.org/10.5281/zenodo.3695602" httpUri="https://zenodo.org/record/3695602/files/figure.png" pageId="9" pageNumber="686">Fig. 4</figureCitation>
), the tubercles are fused together to
<taxonomicName id="16E34D14BF11E56BFAAFCB52DFB53E14" box="[1323,1452,823,849]" form="ridges" pageId="9" pageNumber="686" rank="form">form ridges</taxonomicName>
.
</paragraph>
<paragraph id="D15C3697BF11E56BFCA9CB12DE833ED4" blockId="9.[813,1456,887,1679]" box="[813,1178,887,913]" pageId="9" pageNumber="686">
<emphasis id="E397EA85BF11E56BFCA9CB12DE833ED4" box="[813,1178,887,913]" italics="true" pageId="9" pageNumber="686">
Anterior dorsolateral plate (
<figureCitation id="49D82A12BF11E56BFBD6CB1DDE883ED4" box="[1106,1169,887,913]" captionStart="FIG" captionStartId="10.[132,143,1337,1355]" captionTargetBox="[138,1452,219,1294]" captionTargetId="figure@10.[132,1455,214,1316]" captionTargetPageId="10" captionText="FIG. 5. — Asterolepis alticristata n. sp.: A-C, NUFV 892,left side of articulated partial thoracic skeleton in visceral (A, photograph;A’, labeled illustration),left lateral (B, photograph; B’, labeled illustration), and cranial (C, photograph; C’, labeled illustration) views; D, NUFV 895, right mixilateral plate in dermal view. Abbreviations:ADL, anterior dorsolateral plate;AVL, anterior ventrolateral;c1, central 1 plate; cd, area overlapped by ventral thoracic plates;cd.AMD, area overlapped by the anterior median dorsal plate; cr.d, crescent-shaped depression; cv.Mxl, area overlapping the mixilateral plate; dlr, dorsolateral ridge; f.ax, foramen axillaris; lcg, main lateral line groove; MV, median ventral plate; pr.b, processus brachialis; pr.o, processus obstans. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.3695604" httpUri="https://zenodo.org/record/3695604/files/figure.png" pageId="9" pageNumber="686">Fig. 5</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="D15C3697BF11E56BFCA9CBF2DFA93BCA" blockId="9.[813,1456,887,1679]" pageId="9" pageNumber="686">
The anterior dorsolateral plate of
<taxonomicName id="16E34D14BF11E56BFB3ECBF2DFB63EF4" authority="Downs &amp; Daeschler &amp; Lo &amp; Carey &amp; Shubin, 2019" authorityName="Downs &amp; Daeschler &amp; Lo &amp; Carey &amp; Shubin" authorityYear="2019" box="[1210,1455,919,945]" class="Insecta" family="Tortricidae" genus="Asterolepis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="686" phylum="Arthropoda" rank="species" species="alticristata" status="sp. nov.">
<emphasis id="E397EA85BF11E56BFB3ECBF2DFB63EF4" box="[1210,1455,919,945]" italics="true" pageId="9" pageNumber="686">Asterolepis alticristata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F8A457FEBF11E56BFCA9CBD2D9713E94" box="[813,872,951,977]" pageId="9" pageNumber="686" rank="species">n. sp.</taxonomicNameLabel>
expectedly overlaps the mixilateral plate (
<figureCitation id="49D82A12BF11E56BFAB0CBD2DF6B3E94" box="[1332,1394,950,977]" captionStart="FIG" captionStartId="10.[132,143,1337,1355]" captionTargetBox="[138,1452,219,1294]" captionTargetId="figure@10.[132,1455,214,1316]" captionTargetPageId="10" captionText="FIG. 5. — Asterolepis alticristata n. sp.: A-C, NUFV 892,left side of articulated partial thoracic skeleton in visceral (A, photograph;A’, labeled illustration),left lateral (B, photograph; B’, labeled illustration), and cranial (C, photograph; C’, labeled illustration) views; D, NUFV 895, right mixilateral plate in dermal view. Abbreviations:ADL, anterior dorsolateral plate;AVL, anterior ventrolateral;c1, central 1 plate; cd, area overlapped by ventral thoracic plates;cd.AMD, area overlapped by the anterior median dorsal plate; cr.d, crescent-shaped depression; cv.Mxl, area overlapping the mixilateral plate; dlr, dorsolateral ridge; f.ax, foramen axillaris; lcg, main lateral line groove; MV, median ventral plate; pr.b, processus brachialis; pr.o, processus obstans. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.3695604" httpUri="https://zenodo.org/record/3695604/files/figure.png" pageId="9" pageNumber="686">Fig. 5</figureCitation>
A’[cv. Mxl]) and is overlapped by the anterior median dorsal plate (
<figureCitation id="49D82A12BF11E56BFCB2CB93D96C3954" box="[822,885,1014,1041]" captionStart="FIG" captionStartId="10.[132,143,1337,1355]" captionTargetBox="[138,1452,219,1294]" captionTargetId="figure@10.[132,1455,214,1316]" captionTargetPageId="10" captionText="FIG. 5. — Asterolepis alticristata n. sp.: A-C, NUFV 892,left side of articulated partial thoracic skeleton in visceral (A, photograph;A’, labeled illustration),left lateral (B, photograph; B’, labeled illustration), and cranial (C, photograph; C’, labeled illustration) views; D, NUFV 895, right mixilateral plate in dermal view. Abbreviations:ADL, anterior dorsolateral plate;AVL, anterior ventrolateral;c1, central 1 plate; cd, area overlapped by ventral thoracic plates;cd.AMD, area overlapped by the anterior median dorsal plate; cr.d, crescent-shaped depression; cv.Mxl, area overlapping the mixilateral plate; dlr, dorsolateral ridge; f.ax, foramen axillaris; lcg, main lateral line groove; MV, median ventral plate; pr.b, processus brachialis; pr.o, processus obstans. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.3695604" httpUri="https://zenodo.org/record/3695604/files/figure.png" pageId="9" pageNumber="686">Fig. 5</figureCitation>
B’[cd.AMD]). The dorsolateral ridge (
<figureCitation id="49D82A12BF11E56BFA98CB93DF453954" box="[1308,1372,1014,1041]" captionStart="FIG" captionStartId="10.[132,143,1337,1355]" captionTargetBox="[138,1452,219,1294]" captionTargetId="figure@10.[132,1455,214,1316]" captionTargetPageId="10" captionText="FIG. 5. — Asterolepis alticristata n. sp.: A-C, NUFV 892,left side of articulated partial thoracic skeleton in visceral (A, photograph;A’, labeled illustration),left lateral (B, photograph; B’, labeled illustration), and cranial (C, photograph; C’, labeled illustration) views; D, NUFV 895, right mixilateral plate in dermal view. Abbreviations:ADL, anterior dorsolateral plate;AVL, anterior ventrolateral;c1, central 1 plate; cd, area overlapped by ventral thoracic plates;cd.AMD, area overlapped by the anterior median dorsal plate; cr.d, crescent-shaped depression; cv.Mxl, area overlapping the mixilateral plate; dlr, dorsolateral ridge; f.ax, foramen axillaris; lcg, main lateral line groove; MV, median ventral plate; pr.b, processus brachialis; pr.o, processus obstans. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.3695604" httpUri="https://zenodo.org/record/3695604/files/figure.png" pageId="9" pageNumber="686">Fig. 5</figureCitation>
B’[dlr]) separates the dorsal lamina (maximum transverse width of
<quantity id="161B9B72BF11E56BFCA9CC53D9873914" box="[813,926,1078,1105]" metricMagnitude="-2" metricUnit="m" metricValue="4.76" pageId="9" pageNumber="686" unit="mm" value="47.6">47.60 mm</quantity>
in NUFV 892 [left side]) from the lateral lamina (maximum transverse height of
<quantity id="161B9B72BF11E56BFB15CC33DF1E3934" box="[1169,1287,1110,1137]" metricMagnitude="-2" metricUnit="m" metricValue="3.8689999999999998" pageId="9" pageNumber="686" unit="mm" value="38.69">38.69 mm</quantity>
in NUFV 892 [left side]). The angle between dorsal and lateral laminae is impossible to measure in NUFV 892 due to preservational flattening of the specimen that has deformed those laminae into nearly the same plane. The thickest part of the plate is in the position of the processus obstans (
<figureCitation id="49D82A12BF11E56BFA89CC90DF543855" box="[1293,1357,1269,1296]" captionStart="FIG" captionStartId="10.[132,143,1337,1355]" captionTargetBox="[138,1452,219,1294]" captionTargetId="figure@10.[132,1455,214,1316]" captionTargetPageId="10" captionText="FIG. 5. — Asterolepis alticristata n. sp.: A-C, NUFV 892,left side of articulated partial thoracic skeleton in visceral (A, photograph;A’, labeled illustration),left lateral (B, photograph; B’, labeled illustration), and cranial (C, photograph; C’, labeled illustration) views; D, NUFV 895, right mixilateral plate in dermal view. Abbreviations:ADL, anterior dorsolateral plate;AVL, anterior ventrolateral;c1, central 1 plate; cd, area overlapped by ventral thoracic plates;cd.AMD, area overlapped by the anterior median dorsal plate; cr.d, crescent-shaped depression; cv.Mxl, area overlapping the mixilateral plate; dlr, dorsolateral ridge; f.ax, foramen axillaris; lcg, main lateral line groove; MV, median ventral plate; pr.b, processus brachialis; pr.o, processus obstans. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.3695604" httpUri="https://zenodo.org/record/3695604/files/figure.png" pageId="9" pageNumber="686">Fig. 5</figureCitation>
C’[pr.o]) which consists of a short, medial, cranially-pointing process and a cranially-facing facet that likely accommodated the postmarginal plate of the cephalic skeleton. The ornament is densely tuberculate with those tubercles close to the dorsolateral ridge, on both dorsal and lateral laminae, appearing in rows parallel to that linear feature. Along the ventral margin of the lateral lamina, tubercles are conjoined into ridges oriented perpendicular to the contact between anterior median dorsal and anterior ventrolateral plates. The main lateral line groove (
<figureCitation id="49D82A12BF11E56BFB8CCE51DE533B0A" box="[1032,1098,1588,1615]" captionStart="FIG" captionStartId="10.[132,143,1337,1355]" captionTargetBox="[138,1452,219,1294]" captionTargetId="figure@10.[132,1455,214,1316]" captionTargetPageId="10" captionText="FIG. 5. — Asterolepis alticristata n. sp.: A-C, NUFV 892,left side of articulated partial thoracic skeleton in visceral (A, photograph;A’, labeled illustration),left lateral (B, photograph; B’, labeled illustration), and cranial (C, photograph; C’, labeled illustration) views; D, NUFV 895, right mixilateral plate in dermal view. Abbreviations:ADL, anterior dorsolateral plate;AVL, anterior ventrolateral;c1, central 1 plate; cd, area overlapped by ventral thoracic plates;cd.AMD, area overlapped by the anterior median dorsal plate; cr.d, crescent-shaped depression; cv.Mxl, area overlapping the mixilateral plate; dlr, dorsolateral ridge; f.ax, foramen axillaris; lcg, main lateral line groove; MV, median ventral plate; pr.b, processus brachialis; pr.o, processus obstans. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.3695604" httpUri="https://zenodo.org/record/3695604/files/figure.png" pageId="9" pageNumber="686">Fig. 5</figureCitation>
C’[lcg]) is on the lateral lamina of the anterior dorsolateral plate and follows a path that is roughly parallel with the dorsolateral ridge (
<figureCitation id="49D82A12BF11E56BFA97CE11DF4B3BCA" box="[1299,1362,1652,1679]" captionStart="FIG" captionStartId="10.[132,143,1337,1355]" captionTargetBox="[138,1452,219,1294]" captionTargetId="figure@10.[132,1455,214,1316]" captionTargetPageId="10" captionText="FIG. 5. — Asterolepis alticristata n. sp.: A-C, NUFV 892,left side of articulated partial thoracic skeleton in visceral (A, photograph;A’, labeled illustration),left lateral (B, photograph; B’, labeled illustration), and cranial (C, photograph; C’, labeled illustration) views; D, NUFV 895, right mixilateral plate in dermal view. Abbreviations:ADL, anterior dorsolateral plate;AVL, anterior ventrolateral;c1, central 1 plate; cd, area overlapped by ventral thoracic plates;cd.AMD, area overlapped by the anterior median dorsal plate; cr.d, crescent-shaped depression; cv.Mxl, area overlapping the mixilateral plate; dlr, dorsolateral ridge; f.ax, foramen axillaris; lcg, main lateral line groove; MV, median ventral plate; pr.b, processus brachialis; pr.o, processus obstans. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.3695604" httpUri="https://zenodo.org/record/3695604/files/figure.png" pageId="9" pageNumber="686">Fig. 5</figureCitation>
C’[dlr]).
</paragraph>
<paragraph id="D15C3697BF11E56BFCA9CED1DEBC3B8B" blockId="9.[813,1456,1716,2030]" box="[813,1189,1716,1742]" pageId="9" pageNumber="686">
<emphasis id="E397EA85BF11E56BFCA9CED1DEBC3B8B" box="[813,1189,1716,1742]" italics="true" pageId="9" pageNumber="686">
Anterior ventrolateral plate (
<figureCitation id="49D82A12BF11E56BFBD9CED0DE853B8B" box="[1117,1180,1716,1742]" captionStart="FIG" captionStartId="10.[132,143,1337,1355]" captionTargetBox="[138,1452,219,1294]" captionTargetId="figure@10.[132,1455,214,1316]" captionTargetPageId="10" captionText="FIG. 5. — Asterolepis alticristata n. sp.: A-C, NUFV 892,left side of articulated partial thoracic skeleton in visceral (A, photograph;A’, labeled illustration),left lateral (B, photograph; B’, labeled illustration), and cranial (C, photograph; C’, labeled illustration) views; D, NUFV 895, right mixilateral plate in dermal view. Abbreviations:ADL, anterior dorsolateral plate;AVL, anterior ventrolateral;c1, central 1 plate; cd, area overlapped by ventral thoracic plates;cd.AMD, area overlapped by the anterior median dorsal plate; cr.d, crescent-shaped depression; cv.Mxl, area overlapping the mixilateral plate; dlr, dorsolateral ridge; f.ax, foramen axillaris; lcg, main lateral line groove; MV, median ventral plate; pr.b, processus brachialis; pr.o, processus obstans. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.3695604" httpUri="https://zenodo.org/record/3695604/files/figure.png" pageId="9" pageNumber="686">Fig. 5</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="D15C3697BF11E568FCA9CEB1DBF23AAB" blockId="9.[813,1456,1716,2030]" lastBlockId="10.[132,776,1525,2030]" lastPageId="10" lastPageNumber="687" pageId="9" pageNumber="686">
The anterior ventrolateral plate forms a broad thick platform for the submarginal plates of the
<taxonomicName id="16E34D14BF11E56BFB01CE91DEF63A4B" authorityName="Eichwald, 1840 sensu Agassiz" authorityYear="1844" box="[1157,1263,1780,1806]" class="Insecta" family="Tortricidae" genus="Asterolepis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="688" phylum="Arthropoda" rank="genus">
<emphasis id="E397EA85BF11E56BFB01CE91DEF63A4B" box="[1157,1263,1780,1806]" italics="true" pageId="9" pageNumber="686">Asterolepis</emphasis>
</taxonomicName>
cephalic skeleton. Along with the anterior dorsolateral plate, the cranial end of the anterior ventrolateral plate’s lateral lamina forms a wide, crested backwall for the cephalic skeleton. One specimen (NUFV 892;
<figureCitation id="49D82A12BF11E56BFC43CF16DE113ACB" box="[967,1032,1907,1934]" captionStart="FIG" captionStartId="10.[132,143,1337,1355]" captionTargetBox="[138,1452,219,1294]" captionTargetId="figure@10.[132,1455,214,1316]" captionTargetPageId="10" captionText="FIG. 5. — Asterolepis alticristata n. sp.: A-C, NUFV 892,left side of articulated partial thoracic skeleton in visceral (A, photograph;A’, labeled illustration),left lateral (B, photograph; B’, labeled illustration), and cranial (C, photograph; C’, labeled illustration) views; D, NUFV 895, right mixilateral plate in dermal view. Abbreviations:ADL, anterior dorsolateral plate;AVL, anterior ventrolateral;c1, central 1 plate; cd, area overlapped by ventral thoracic plates;cd.AMD, area overlapped by the anterior median dorsal plate; cr.d, crescent-shaped depression; cv.Mxl, area overlapping the mixilateral plate; dlr, dorsolateral ridge; f.ax, foramen axillaris; lcg, main lateral line groove; MV, median ventral plate; pr.b, processus brachialis; pr.o, processus obstans. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.3695604" httpUri="https://zenodo.org/record/3695604/files/figure.png" pageId="9" pageNumber="686">Fig. 5</figureCitation>
) preserves the lateral lamina, pectoral articulation, and only a short section of the ventral lamina of both anterior ventrolateral plates of
<taxonomicName id="16E34D14BF11E56BFB4ACFD6DFA93A88" authority="Downs &amp; Daeschler &amp; Lo &amp; Carey &amp; Shubin, 2019" authorityName="Downs &amp; Daeschler &amp; Lo &amp; Carey &amp; Shubin" authorityYear="2019" box="[1230,1456,1971,1997]" class="Insecta" family="Tortricidae" genus="Asterolepis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="686" phylum="Arthropoda" rank="species" species="alticristata" status="sp. nov.">
<emphasis id="E397EA85BF11E56BFB4ACFD6DFA93A88" box="[1230,1456,1971,1997]" italics="true" pageId="9" pageNumber="686">Asterolepis alticristata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F8A457FEBF11E56BFCA9CFB1D97E3AAB" box="[813,871,2004,2030]" pageId="9" pageNumber="686" rank="species">n. sp.</taxonomicNameLabel>
As in adult
<emphasis id="E397EA85BF11E56BFC6ECFB6DEB93AA8" box="[1002,1184,2003,2029]" italics="true" pageId="9" pageNumber="686">
<taxonomicName id="16E34D14BF11E56BFC6ECFB6DE4C3AA8" authorityName="Eichwald, 1840 sensu Agassiz" authorityYear="1844" box="[1002,1109,2003,2029]" class="Insecta" family="Tortricidae" genus="Asterolepis" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="688" phylum="Arthropoda" rank="genus">Asterolepis</taxonomicName>
ornata
</emphasis>
, the lateral lamina of the anterior ventrolateral plate is tall across its entire length and reaches its maximum height close to the caudal end. The anterior ventrolateral plate is remarkably thick at the pectoral articulation and, at
<quantity id="161B9B72BF12E568FED9CE31DBC83B2A" box="[349,465,1620,1647]" metricMagnitude="-2" metricUnit="m" metricValue="3.8869999999999996" pageId="10" pageNumber="687" unit="mm" value="38.87">38.87 mm</quantity>
in NUFV 892, presents the position of greatest skeletal thickness known for the species. The processus brachialis (
<figureCitation id="49D82A12BF12E568FE26CEF1DBFA3BEA" box="[418,483,1684,1711]" captionStart="FIG" captionStartId="10.[132,143,1337,1355]" captionTargetBox="[138,1452,219,1294]" captionTargetId="figure@10.[132,1455,214,1316]" captionTargetPageId="10" captionText="FIG. 5. — Asterolepis alticristata n. sp.: A-C, NUFV 892,left side of articulated partial thoracic skeleton in visceral (A, photograph;A’, labeled illustration),left lateral (B, photograph; B’, labeled illustration), and cranial (C, photograph; C’, labeled illustration) views; D, NUFV 895, right mixilateral plate in dermal view. Abbreviations:ADL, anterior dorsolateral plate;AVL, anterior ventrolateral;c1, central 1 plate; cd, area overlapped by ventral thoracic plates;cd.AMD, area overlapped by the anterior median dorsal plate; cr.d, crescent-shaped depression; cv.Mxl, area overlapping the mixilateral plate; dlr, dorsolateral ridge; f.ax, foramen axillaris; lcg, main lateral line groove; MV, median ventral plate; pr.b, processus brachialis; pr.o, processus obstans. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.3695604" httpUri="https://zenodo.org/record/3695604/files/figure.png" pageId="10" pageNumber="687">Fig. 5</figureCitation>
B’[pr.b]) resides in a fossa that is deeply excavated from the ornamented surface. This fossa narrows to a point cranially. The processus brachialis is penetrated by an oval-shaped (long axis dorsal-ventral) foramen axillaris (
<figureCitation id="49D82A12BF12E568FE9FCF71DB413A6B" box="[283,344,1811,1838]" captionStart="FIG" captionStartId="10.[132,143,1337,1355]" captionTargetBox="[138,1452,219,1294]" captionTargetId="figure@10.[132,1455,214,1316]" captionTargetPageId="10" captionText="FIG. 5. — Asterolepis alticristata n. sp.: A-C, NUFV 892,left side of articulated partial thoracic skeleton in visceral (A, photograph;A’, labeled illustration),left lateral (B, photograph; B’, labeled illustration), and cranial (C, photograph; C’, labeled illustration) views; D, NUFV 895, right mixilateral plate in dermal view. Abbreviations:ADL, anterior dorsolateral plate;AVL, anterior ventrolateral;c1, central 1 plate; cd, area overlapped by ventral thoracic plates;cd.AMD, area overlapped by the anterior median dorsal plate; cr.d, crescent-shaped depression; cv.Mxl, area overlapping the mixilateral plate; dlr, dorsolateral ridge; f.ax, foramen axillaris; lcg, main lateral line groove; MV, median ventral plate; pr.b, processus brachialis; pr.o, processus obstans. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.3695604" httpUri="https://zenodo.org/record/3695604/files/figure.png" pageId="10" pageNumber="687">Fig. 5</figureCitation>
B’[f.ax]). The most cranial margin of the plate’s ventral platform exhibits a tall concave, cranial-facing facet that likely accommodated the semilunar plate. As in
<emphasis id="E397EA85BF12E568FF00CF11DAFF3AC8" box="[132,230,1908,1934]" italics="true" pageId="10" pageNumber="687">A. ornata</emphasis>
, this facet terminates laterally in a cranial-pointing process (the antero-mesial process of
<bibRefCitation id="B5724B66BF12E568FD9BCFF6D8DC3AE8" author="UPENIECE I." box="[543,709,1939,1965]" pageId="10" pageNumber="687" refId="ref18774" refString="UPENIECE I. 2011. - Palaeoecology and juvenile individuals of the Devonian placoderm and acanthodian fishes from Lode site, Latvia. (Doctoral dissertation). Retrieved from https: // dspace. lu. lv / dspace / handle / 7 / 4659 (last access on 3 rd October 2019)." type="url" year="2011">Upeniece 2011</bibRefCitation>
); further lateral is the most cranial point of the plate (the anterior lateral corner of
<bibRefCitation id="B5724B66BF12E568FEB3CFB6DBC23AA8" author="UPENIECE I." box="[311,475,2003,2029]" pageId="10" pageNumber="687" refId="ref18774" refString="UPENIECE I. 2011. - Palaeoecology and juvenile individuals of the Devonian placoderm and acanthodian fishes from Lode site, Latvia. (Doctoral dissertation). Retrieved from https: // dspace. lu. lv / dspace / handle / 7 / 4659 (last access on 3 rd October 2019)." type="url" year="2011">Upeniece 2011</bibRefCitation>
).
</paragraph>
<caption id="859C661FBF12E568FF00CD5CDE9738E3" ID-DOI="http://doi.org/10.5281/zenodo.3695604" ID-Zenodo-Dep="3695604" httpUri="https://zenodo.org/record/3695604/files/figure.png" pageId="10" pageNumber="687" startId="10.[132,143,1337,1355]" targetBox="[138,1452,219,1294]" targetPageId="10">
<paragraph id="D15C3697BF12E568FF00CD5CDE9738E3" blockId="10.[132,1457,1337,1446]" pageId="10" pageNumber="687">
FIG. 5. —
<taxonomicName id="16E34D14BF12E568FF56CD5CDB9F380E" authority="Downs &amp; Daeschler &amp; Lo &amp; Carey &amp; Shubin, 2019" authorityName="Downs &amp; Daeschler &amp; Lo &amp; Carey &amp; Shubin" authorityYear="2019" box="[210,390,1337,1355]" class="Insecta" family="Tortricidae" genus="Asterolepis" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="687" phylum="Arthropoda" rank="species" species="alticristata" status="sp. nov.">
<emphasis id="E397EA85BF12E568FF56CD5CDB9F380E" box="[210,390,1337,1355]" italics="true" pageId="10" pageNumber="687">Asterolepis alticristata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F8A457FEBF12E568FE0ECD5CDBA5380E" box="[394,444,1337,1355]" pageId="10" pageNumber="687" rank="species">n. sp.</taxonomicNameLabel>
:
<emphasis id="E397EA85BF12E568FE45CD5CDBFA380E" bold="true" box="[449,483,1337,1355]" pageId="10" pageNumber="687">A -C</emphasis>
, NUFV 892,left side of articulated partial thoracic skeleton in visceral (
<emphasis id="E397EA85BF12E568FBA5CD5CDE37380E" bold="true" box="[1057,1070,1337,1355]" pageId="10" pageNumber="687">A</emphasis>
, photograph;
<emphasis id="E397EA85BF12E568FB25CD5CDEA9380E" bold="true" box="[1185,1200,1337,1355]" pageId="10" pageNumber="687">A’</emphasis>
, labeled illustration),left lateral (
<emphasis id="E397EA85BF12E568FF0DCD35DA8F3827" bold="true" box="[137,150,1360,1378]" pageId="10" pageNumber="687">B</emphasis>
, photograph;
<emphasis id="E397EA85BF12E568FE88CD35DB073827" bold="true" box="[268,286,1360,1378]" pageId="10" pageNumber="687">B’</emphasis>
, labeled illustration), and cranial (
<emphasis id="E397EA85BF12E568FDB1CD35D85A3827" bold="true" box="[565,579,1360,1378]" pageId="10" pageNumber="687">C</emphasis>
, photograph;
<emphasis id="E397EA85BF12E568FD3DCD35D8D53827" bold="true" box="[697,716,1360,1378]" pageId="10" pageNumber="687">C’</emphasis>
, labeled illustration) views;
<emphasis id="E397EA85BF12E568FC36CD35D9D93827" bold="true" box="[946,960,1360,1378]" pageId="10" pageNumber="687">D</emphasis>
, NUFV 895, right mixilateral plate in dermal view. Abbreviations:
<emphasis id="E397EA85BF12E568FF32CD02DAC5383C" bold="true" box="[182,220,1383,1401]" pageId="10" pageNumber="687">ADL</emphasis>
, anterior dorsolateral plate;
<emphasis id="E397EA85BF12E568FE46CD02DBFD383C" bold="true" box="[450,484,1383,1401]" pageId="10" pageNumber="687">AVL</emphasis>
, anterior ventrolateral;
<emphasis id="E397EA85BF12E568FD26CD02D8AE383C" bold="true" box="[674,695,1383,1401]" pageId="10" pageNumber="687">c1</emphasis>
, central 1 plate;
<emphasis id="E397EA85BF12E568FCBBCD02D94C383C" bold="true" box="[831,853,1383,1401]" pageId="10" pageNumber="687">cd</emphasis>
, area overlapped by ventral thoracic plates;
<emphasis id="E397EA85BF12E568FB44CD02DF1E383C" bold="true" box="[1216,1287,1383,1401]" pageId="10" pageNumber="687">cd.AMD</emphasis>
, area overlapped by the anterior median dorsal plate;
<emphasis id="E397EA85BF12E568FE13CD18DBA138CA" bold="true" box="[407,440,1405,1423]" pageId="10" pageNumber="687">cr.d</emphasis>
, crescent-shaped depression;
<emphasis id="E397EA85BF12E568FD3FCD18D8EB38CA" bold="true" box="[699,754,1405,1423]" pageId="10" pageNumber="687">cv.Mxl</emphasis>
, area overlapping the mixilateral plate;
<emphasis id="E397EA85BF12E568FBBECD18DE4838CA" bold="true" box="[1082,1105,1405,1423]" pageId="10" pageNumber="687">dlr</emphasis>
, dorsolateral ridge;
<emphasis id="E397EA85BF12E568FB7CCD18DF0138CA" bold="true" box="[1272,1304,1405,1423]" pageId="10" pageNumber="687">f.ax</emphasis>
, foramen axillaris;
<emphasis id="E397EA85BF12E568FF00CDF1DA8638E3" bold="true" box="[132,159,1428,1446]" pageId="10" pageNumber="687">lcg</emphasis>
, main lateral line groove;
<emphasis id="E397EA85BF12E568FEF1CDF1DB8B38E3" bold="true" box="[373,402,1428,1446]" pageId="10" pageNumber="687">MV</emphasis>
, median ventral plate;
<emphasis id="E397EA85BF12E568FDCBCDF1D86838E3" bold="true" box="[591,625,1428,1446]" pageId="10" pageNumber="687">pr.b</emphasis>
, processus brachialis;
<emphasis id="E397EA85BF12E568FCB4CDF1D94B38E3" bold="true" box="[816,850,1428,1446]" pageId="10" pageNumber="687">pr.o</emphasis>
, processus obstans. Scale bar: 5 cm.
</paragraph>
</caption>
<paragraph id="D15C3697BF12E568FCC7CD90DE1F3B8B" blockId="10.[813,1455,1525,1743]" pageId="10" pageNumber="687">The ornament of the anterior ventrolateral plate’s lateral lamina is densely tuberculate; the tubercles are arranged in rows that radiate from a position just caudal to the pectoral articulation. The preserved portion of the ventral lamina, just ventral to the pectoral articulation, is unornamented as is a depressed zone just dorsal to the fossa that surrounds the processus brachialis.</paragraph>
<paragraph id="D15C3697BF12E568FCA9CE96DF253AA8" blockId="10.[813,1456,1779,2030]" pageId="10" pageNumber="687">
<emphasis id="E397EA85BF12E568FCA9CE96D9723A48" bold="true" box="[813,875,1779,1805]" pageId="10" pageNumber="687">Note.</emphasis>
On the lateral lamina of the left anterior ventrolateral plate of NUFV 892, there is a crescent-shaped depression (
<figureCitation id="49D82A12BF12E568FCB1CF56D9693A0B" box="[821,880,1843,1870]" captionStart="FIG" captionStartId="10.[132,143,1337,1355]" captionTargetBox="[138,1452,219,1294]" captionTargetId="figure@10.[132,1455,214,1316]" captionTargetPageId="10" captionText="FIG. 5. — Asterolepis alticristata n. sp.: A-C, NUFV 892,left side of articulated partial thoracic skeleton in visceral (A, photograph;A’, labeled illustration),left lateral (B, photograph; B’, labeled illustration), and cranial (C, photograph; C’, labeled illustration) views; D, NUFV 895, right mixilateral plate in dermal view. Abbreviations:ADL, anterior dorsolateral plate;AVL, anterior ventrolateral;c1, central 1 plate; cd, area overlapped by ventral thoracic plates;cd.AMD, area overlapped by the anterior median dorsal plate; cr.d, crescent-shaped depression; cv.Mxl, area overlapping the mixilateral plate; dlr, dorsolateral ridge; f.ax, foramen axillaris; lcg, main lateral line groove; MV, median ventral plate; pr.b, processus brachialis; pr.o, processus obstans. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.3695604" httpUri="https://zenodo.org/record/3695604/files/figure.png" pageId="10" pageNumber="687">Fig. 5</figureCitation>
B’[cr.d]) that interrupts the surrounding ornament pattern. The ornament within the depression consists of larger, isolated and fused tubercles that are more widely spaced and chaotically-organized than those of the surrounding lateral lamina. This depression may represent bone repair of puncturetype damage or an instance of skeletal pathology.
</paragraph>
<caption id="859C661FBF13E569FF00C8B8DBEF3C40" ID-Table-UUID="859C661FBF13E569FF00C8B8DBEF3C40" httpUri="http://table.plazi.org/id/859C661FBF13E569FF00C8B8DBEF3C40" pageId="11" pageNumber="688" targetBox="[134,775,320,762]" targetIsTable="true" targetPageId="11">
<paragraph id="D15C3697BF13E569FF00C8B8DBEF3C40" blockId="11.[132,775,221,262]" pageId="11" pageNumber="688">TABLE 1. — Antiarch-producing sites within the Fram Formation on southern Ellesmere Island, Nunavut Territory, Canada.</paragraph>
</caption>
<paragraph id="D15C3697BF13E569FF02C925D8F93FBF" pageId="11" pageNumber="688">
<table id="A3E3C437BF131A9DFF02C925D91E3FBF" box="[134,775,320,762]" gridcols="4" gridrows="18" pageId="11" pageNumber="688">
<tr id="6FD334D5BF131A9DFF02C925D91E3C11" box="[134,775,320,340]" gridrow="0" pageId="11" pageNumber="688" rowspan-1="1" rowspan-3="1">
<th id="2C025DA9BF131A9DFF02C925DB193C11" box="[134,256,320,340]" gridcol="0" gridrow="0" pageId="11" pageNumber="688">
<emphasis id="E397EA85BF13E569FF02C925DAB43C11" bold="true" box="[134,173,320,340]" pageId="11" pageNumber="688">Site</emphasis>
</th>
<th id="2C025DA9BF131A9DFDC1C925D8FA3C11" box="[581,739,320,340]" gridcol="2" gridrow="0" pageId="11" pageNumber="688">
<emphasis id="E397EA85BF13E569FDC1C925D8D03C11" bold="true" box="[581,713,320,340]" pageId="11" pageNumber="688">Stratigraphic</emphasis>
</th>
</tr>
<tr id="6FD334D5BF131A9DFF02C93CD91E3C28" box="[134,775,345,365]" gridrow="1" pageId="11" pageNumber="688" rowspan-3="1">
<th id="2C025DA9BF131A9DFF02C93CDB193C28" box="[134,256,345,365]" gridcol="0" gridrow="1" pageId="11" pageNumber="688">
<emphasis id="E397EA85BF13E569FF02C93CDB193C28" bold="true" box="[134,256,345,365]" pageId="11" pageNumber="688">Designation</emphasis>
</th>
<td id="2C025DA9BF131A9DFE8EC93CD82B3C28" box="[266,562,345,365]" gridcol="1" gridrow="1" pageId="11" pageNumber="688">
<emphasis id="E397EA85BF13E569FE8EC93CDB9F3C28" bold="true" box="[266,390,345,365]" pageId="11" pageNumber="688">Coordinates</emphasis>
</td>
<td id="2C025DA9BF131A9DFDC1C93CD8FA3C28" box="[581,739,345,365]" gridcol="2" gridrow="1" pageId="11" pageNumber="688">
<emphasis id="E397EA85BF13E569FDC1C93CD8813C28" bold="true" box="[581,664,345,365]" pageId="11" pageNumber="688">Position</emphasis>
</td>
</tr>
<tr id="6FD334D5BF131A9DFF02C919D91E3CD5" box="[134,775,380,400]" gridrow="2" pageId="11" pageNumber="688">
<th id="2C025DA9BF131A9DFF02C919DB193CD5" box="[134,256,380,400]" gridcol="0" gridrow="2" pageId="11" pageNumber="688">NV0207</th>
<td id="2C025DA9BF131A9DFE8EC919D82B3CD5" box="[266,562,380,400]" gridcol="1" gridrow="2" pageId="11" pageNumber="688">77°39’8.34”N, 83°24’18.9”W</td>
<td id="2C025DA9BF131A9DFDC1C919D8FA3CD5" box="[581,739,380,400]" gridcol="2" gridrow="2" pageId="11" pageNumber="688">lower-most Fram</td>
<td id="2C025DA9BF131A9DFD6DC919D91E3CD5" box="[745,775,380,400]" gridcol="3" gridrow="2" pageId="11" pageNumber="688">Fm</td>
</tr>
<tr id="6FD334D5BF131A9DFF02C9F0D91E3CEC" box="[134,775,405,425]" gridrow="3" pageId="11" pageNumber="688" rowspan-3="1">
<th id="2C025DA9BF131A9DFF02C9F0DB193CEC" box="[134,256,405,425]" gridcol="0" gridrow="3" pageId="11" pageNumber="688">NV0202</th>
<td id="2C025DA9BF131A9DFE8EC9F0D82B3CEC" box="[266,562,405,425]" gridcol="1" gridrow="3" pageId="11" pageNumber="688">77°13’14.94”N, 84°22’6.72”W</td>
<td id="2C025DA9BF131A9DFDC1C9F0D8FA3CEC" box="[581,739,405,425]" gridcol="2" gridrow="3" pageId="11" pageNumber="688">upper Fram Fm</td>
</tr>
<tr id="6FD334D5BF131A9DFF02C9C8D91E3C84" box="[134,775,429,449]" gridrow="4" pageId="11" pageNumber="688" rowspan-3="1">
<th id="2C025DA9BF131A9DFF02C9C8DB193C84" box="[134,256,429,449]" gridcol="0" gridrow="4" pageId="11" pageNumber="688">NV0203</th>
<td id="2C025DA9BF131A9DFE8EC9C8D82B3C84" box="[266,562,429,449]" gridcol="1" gridrow="4" pageId="11" pageNumber="688">77°11’32.05”N, 84°19’17.28”W</td>
<td id="2C025DA9BF131A9DFDC1C9C8D8FA3C84" box="[581,739,429,449]" gridcol="2" gridrow="4" pageId="11" pageNumber="688">middle Fram Fm</td>
</tr>
<tr id="6FD334D5BF131A9DFF02C9A0D91E3C9C" box="[134,775,453,473]" gridrow="5" pageId="11" pageNumber="688" rowspan-3="1">
<th id="2C025DA9BF131A9DFF02C9A0DB193C9C" box="[134,256,453,473]" gridcol="0" gridrow="5" pageId="11" pageNumber="688">NV0205</th>
<td id="2C025DA9BF131A9DFE8EC9A0D82B3C9C" box="[266,562,453,473]" gridcol="1" gridrow="5" pageId="11" pageNumber="688">77°10’19.02”N, 84°16’14.1”W</td>
<td id="2C025DA9BF131A9DFDC1C9A0D8FA3C9C" box="[581,739,453,473]" gridcol="2" gridrow="5" pageId="11" pageNumber="688">lower Fram Fm</td>
</tr>
<tr id="6FD334D5BF131A9DFF02C9B8D91E3CB4" box="[134,775,477,497]" gridrow="6" pageId="11" pageNumber="688">
<th id="2C025DA9BF131A9DFF02C9B8DB193CB4" box="[134,256,477,497]" gridcol="0" gridrow="6" pageId="11" pageNumber="688">NV0403</th>
<td id="2C025DA9BF131A9DFE8EC9B8D82B3CB4" box="[266,562,477,497]" gridcol="1" gridrow="6" pageId="11" pageNumber="688">77°10’35.82”N, 86°16’47.7”W</td>
<td id="2C025DA9BF131A9DFDC1C9B8D8FA3CB4" box="[581,739,477,497]" gridcol="2" gridrow="6" pageId="11" pageNumber="688">lower-most Fram</td>
<td id="2C025DA9BF131A9DFD6DC9B8D91E3CB4" box="[745,775,477,497]" gridcol="3" gridrow="6" pageId="11" pageNumber="688">Fm</td>
</tr>
<tr id="6FD334D5BF131A9DFF02C990D91E3F4C" box="[134,775,501,521]" gridrow="7" pageId="11" pageNumber="688" rowspan-3="1">
<th id="2C025DA9BF131A9DFF02C990DB193F4C" box="[134,256,501,521]" gridcol="0" gridrow="7" pageId="11" pageNumber="688">NV0404</th>
<td id="2C025DA9BF131A9DFE8EC990D82B3F4C" box="[266,562,501,521]" gridcol="1" gridrow="7" pageId="11" pageNumber="688">77°10’41.64”N, 86°15’42.96”W</td>
<td id="2C025DA9BF131A9DFDC1C990D8FA3F4C" box="[581,739,501,521]" gridcol="2" gridrow="7" pageId="11" pageNumber="688">lower Fram Fm</td>
</tr>
<tr id="6FD334D5BF131A9DFF02CA68D91E3F64" box="[134,775,525,545]" gridrow="8" pageId="11" pageNumber="688" rowspan-3="1">
<th id="2C025DA9BF131A9DFF02CA68DB193F64" box="[134,256,525,545]" gridcol="0" gridrow="8" pageId="11" pageNumber="688">NV2K16</th>
<td id="2C025DA9BF131A9DFE8ECA68D82B3F64" box="[266,562,525,545]" gridcol="1" gridrow="8" pageId="11" pageNumber="688">77°9’54.42”N, 86°8’5.94”W</td>
<td id="2C025DA9BF131A9DFDC1CA68D8FA3F64" box="[581,739,525,545]" gridcol="2" gridrow="8" pageId="11" pageNumber="688">middle Fram Fm</td>
</tr>
<tr id="6FD334D5BF131A9DFF02CA40D91E3F7C" box="[134,775,549,569]" gridrow="9" pageId="11" pageNumber="688">
<th id="2C025DA9BF131A9DFF02CA40DB193F7C" box="[134,256,549,569]" gridcol="0" gridrow="9" pageId="11" pageNumber="688">NV0802</th>
<td id="2C025DA9BF131A9DFE8ECA40D82B3F7C" box="[266,562,549,569]" gridcol="1" gridrow="9" pageId="11" pageNumber="688">77°10’36.24”N, 86°14’54.6”W</td>
<td id="2C025DA9BF131A9DFDC1CA40D8FA3F7C" box="[581,739,549,569]" gridcol="2" gridrow="9" pageId="11" pageNumber="688">lower-most Fram</td>
<td id="2C025DA9BF131A9DFD6DCA40D91E3F7C" box="[745,775,549,569]" gridcol="3" gridrow="9" pageId="11" pageNumber="688">Fm</td>
</tr>
<tr id="6FD334D5BF131A9DFF02CA58D91E3F14" box="[134,775,573,593]" gridrow="10" pageId="11" pageNumber="688" rowspan-3="1">
<th id="2C025DA9BF131A9DFF02CA58DB193F14" box="[134,256,573,593]" gridcol="0" gridrow="10" pageId="11" pageNumber="688">NV2K17</th>
<td id="2C025DA9BF131A9DFE8ECA58D82B3F14" box="[266,562,573,593]" gridcol="1" gridrow="10" pageId="11" pageNumber="688">77°9’53.7”N, 86°16’9.42”W</td>
<td id="2C025DA9BF131A9DFDC1CA58D8FA3F14" box="[581,739,573,593]" gridcol="2" gridrow="10" pageId="11" pageNumber="688">middle Fram Fm</td>
</tr>
<tr id="6FD334D5BF131A9DFF02CA30D91E3F2C" box="[134,775,597,617]" gridrow="11" pageId="11" pageNumber="688" rowspan-3="1">
<th id="2C025DA9BF131A9DFF02CA30DB193F2C" box="[134,256,597,617]" gridcol="0" gridrow="11" pageId="11" pageNumber="688">NV0402</th>
<td id="2C025DA9BF131A9DFE8ECA30D82B3F2C" box="[266,562,597,617]" gridcol="1" gridrow="11" pageId="11" pageNumber="688">77°10’8.34”N, 86°10’58.26”W</td>
<td id="2C025DA9BF131A9DFDC1CA30D8FA3F2C" box="[581,739,597,617]" gridcol="2" gridrow="11" pageId="11" pageNumber="688">middle Fram Fm</td>
</tr>
<tr id="6FD334D5BF131A9DFF02CA08D91E3FC4" box="[134,775,621,641]" gridrow="12" pageId="11" pageNumber="688" rowspan-3="1">
<th id="2C025DA9BF131A9DFF02CA08DB193FC4" box="[134,256,621,641]" gridcol="0" gridrow="12" pageId="11" pageNumber="688">NV2K15</th>
<td id="2C025DA9BF131A9DFE8ECA08D82B3FC4" box="[266,562,621,641]" gridcol="1" gridrow="12" pageId="11" pageNumber="688">77°9’37.14”N, 86°9’7.74”W</td>
<td id="2C025DA9BF131A9DFDC1CA08D8FA3FC4" box="[581,739,621,641]" gridcol="2" gridrow="12" pageId="11" pageNumber="688">upper Fram Fm</td>
</tr>
<tr id="6FD334D5BF131A9DFF02CAE0D91E3FDC" box="[134,775,645,665]" gridrow="13" pageId="11" pageNumber="688" rowspan-3="1">
<th id="2C025DA9BF131A9DFF02CAE0DB193FDC" box="[134,256,645,665]" gridcol="0" gridrow="13" pageId="11" pageNumber="688">NV2K01</th>
<td id="2C025DA9BF131A9DFE8ECAE0D82B3FDC" box="[266,562,645,665]" gridcol="1" gridrow="13" pageId="11" pageNumber="688">76°45’39.06”N, 88°28’33.42”W</td>
<td id="2C025DA9BF131A9DFDC1CAE0D8FA3FDC" box="[581,739,645,665]" gridcol="2" gridrow="13" pageId="11" pageNumber="688">middle Fram Fm</td>
</tr>
<tr id="6FD334D5BF131A9DFF02CAFBD91E3FF7" box="[134,775,670,690]" gridrow="14" pageId="11" pageNumber="688" rowspan-3="1">
<th id="2C025DA9BF131A9DFF02CAFBDB193FF7" box="[134,256,670,690]" gridcol="0" gridrow="14" pageId="11" pageNumber="688">NV2K02</th>
<td id="2C025DA9BF131A9DFE8ECAFBD82B3FF7" box="[266,562,670,690]" gridcol="1" gridrow="14" pageId="11" pageNumber="688">76°45’42”N, 88°28’32.58”W</td>
<td id="2C025DA9BF131A9DFDC1CAFBD8FA3FF7" box="[581,739,670,690]" gridcol="2" gridrow="14" pageId="11" pageNumber="688">middle Fram Fm</td>
</tr>
<tr id="6FD334D5BF131A9DFF02CAD3D91E3F8F" box="[134,775,694,714]" gridrow="15" pageId="11" pageNumber="688" rowspan-3="1">
<th id="2C025DA9BF131A9DFF02CAD3DB193F8F" box="[134,256,694,714]" gridcol="0" gridrow="15" pageId="11" pageNumber="688">NV2K03</th>
<td id="2C025DA9BF131A9DFE8ECAD3D82B3F8F" box="[266,562,694,714]" gridcol="1" gridrow="15" pageId="11" pageNumber="688">76°45’50.7”N, 88°28’26.94”W</td>
<td id="2C025DA9BF131A9DFDC1CAD3D8FA3F8F" box="[581,739,694,714]" gridcol="2" gridrow="15" pageId="11" pageNumber="688">middle Fram Fm</td>
</tr>
<tr id="6FD334D5BF131A9DFF02CAABD91E3FA7" box="[134,775,718,738]" gridrow="16" pageId="11" pageNumber="688" rowspan-3="1">
<th id="2C025DA9BF131A9DFF02CAABDB193FA7" box="[134,256,718,738]" gridcol="0" gridrow="16" pageId="11" pageNumber="688">NV2K05</th>
<td id="2C025DA9BF131A9DFE8ECAABD82B3FA7" box="[266,562,718,738]" gridcol="1" gridrow="16" pageId="11" pageNumber="688">76°45’53.1”N, 88°28’24.18”W</td>
<td id="2C025DA9BF131A9DFDC1CAABD8FA3FA7" box="[581,739,718,738]" gridcol="2" gridrow="16" pageId="11" pageNumber="688">middle Fram Fm</td>
</tr>
<tr id="6FD334D5BF131A9DFF02CA83D91E3FBF" box="[134,775,742,762]" gridrow="17" pageId="11" pageNumber="688" rowspan-3="1">
<th id="2C025DA9BF131A9DFF02CA83DB193FBF" box="[134,256,742,762]" gridcol="0" gridrow="17" pageId="11" pageNumber="688">NV2K07</th>
<td id="2C025DA9BF131A9DFE8ECA83D82B3FBF" box="[266,562,742,762]" gridcol="1" gridrow="17" pageId="11" pageNumber="688">76°45’36.54”N, 88°28’43.14”W</td>
<td id="2C025DA9BF131A9DFDC1CA83D8FA3FBF" box="[581,739,742,762]" gridcol="2" gridrow="17" pageId="11" pageNumber="688">middle Fram Fm</td>
</tr>
</table>
</paragraph>
<paragraph id="D15C3697BF13E569FF00CB52DBF83E14" blockId="11.[132,776,823,1232]" box="[132,481,823,849]" pageId="11" pageNumber="688">
<emphasis id="E397EA85BF13E569FF00CB52DBF83E14" box="[132,481,823,849]" italics="true" pageId="11" pageNumber="688">
Median ventral plate (
<figureCitation id="49D82A12BF13E569FEE8CB5DDBC03E14" box="[364,473,823,849]" captionStart="FIG" captionStartId="10.[132,143,1337,1355]" captionTargetBox="[138,1452,219,1294]" captionTargetId="figure@10.[132,1455,214,1316]" captionTargetPageId="10" captionText="FIG. 5. — Asterolepis alticristata n. sp.: A-C, NUFV 892,left side of articulated partial thoracic skeleton in visceral (A, photograph;A’, labeled illustration),left lateral (B, photograph; B’, labeled illustration), and cranial (C, photograph; C’, labeled illustration) views; D, NUFV 895, right mixilateral plate in dermal view. Abbreviations:ADL, anterior dorsolateral plate;AVL, anterior ventrolateral;c1, central 1 plate; cd, area overlapped by ventral thoracic plates;cd.AMD, area overlapped by the anterior median dorsal plate; cr.d, crescent-shaped depression; cv.Mxl, area overlapping the mixilateral plate; dlr, dorsolateral ridge; f.ax, foramen axillaris; lcg, main lateral line groove; MV, median ventral plate; pr.b, processus brachialis; pr.o, processus obstans. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.3695604" httpUri="https://zenodo.org/record/3695604/files/figure.png" pageId="11" pageNumber="688">Fig. 5A, B</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="D15C3697BF13E569FF00CB32D91E3995" blockId="11.[132,776,823,1232]" pageId="11" pageNumber="688">
The one median ventral plate (NUFV 892;
<figureCitation id="49D82A12BF13E569FDCACB32D8A33E37" box="[590,698,855,882]" captionStart="FIG" captionStartId="10.[132,143,1337,1355]" captionTargetBox="[138,1452,219,1294]" captionTargetId="figure@10.[132,1455,214,1316]" captionTargetPageId="10" captionText="FIG. 5. — Asterolepis alticristata n. sp.: A-C, NUFV 892,left side of articulated partial thoracic skeleton in visceral (A, photograph;A’, labeled illustration),left lateral (B, photograph; B’, labeled illustration), and cranial (C, photograph; C’, labeled illustration) views; D, NUFV 895, right mixilateral plate in dermal view. Abbreviations:ADL, anterior dorsolateral plate;AVL, anterior ventrolateral;c1, central 1 plate; cd, area overlapped by ventral thoracic plates;cd.AMD, area overlapped by the anterior median dorsal plate; cr.d, crescent-shaped depression; cv.Mxl, area overlapping the mixilateral plate; dlr, dorsolateral ridge; f.ax, foramen axillaris; lcg, main lateral line groove; MV, median ventral plate; pr.b, processus brachialis; pr.o, processus obstans. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.3695604" httpUri="https://zenodo.org/record/3695604/files/figure.png" pageId="11" pageNumber="688">Fig. 5A, B</figureCitation>
) is best preserved in visceral view. The outline of the median ventral plate is generally circular although it appears to pinch to
<taxonomicName id="16E34D14BF13E569FD56CBF2DAA63E94" form="blunt" pageId="11" pageNumber="688" rank="form">form blunt</taxonomicName>
points in at least three corners. What is exposed of the external surface (
<figureCitation id="49D82A12BF13E569FEB8CBB3DB953EB4" box="[316,396,982,1009]" captionStart="FIG" captionStartId="10.[132,143,1337,1355]" captionTargetBox="[138,1452,219,1294]" captionTargetId="figure@10.[132,1455,214,1316]" captionTargetPageId="10" captionText="FIG. 5. — Asterolepis alticristata n. sp.: A-C, NUFV 892,left side of articulated partial thoracic skeleton in visceral (A, photograph;A’, labeled illustration),left lateral (B, photograph; B’, labeled illustration), and cranial (C, photograph; C’, labeled illustration) views; D, NUFV 895, right mixilateral plate in dermal view. Abbreviations:ADL, anterior dorsolateral plate;AVL, anterior ventrolateral;c1, central 1 plate; cd, area overlapped by ventral thoracic plates;cd.AMD, area overlapped by the anterior median dorsal plate; cr.d, crescent-shaped depression; cv.Mxl, area overlapping the mixilateral plate; dlr, dorsolateral ridge; f.ax, foramen axillaris; lcg, main lateral line groove; MV, median ventral plate; pr.b, processus brachialis; pr.o, processus obstans. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.3695604" httpUri="https://zenodo.org/record/3695604/files/figure.png" pageId="11" pageNumber="688">Fig. 5B</figureCitation>
) shows a densely tuberculate ornament with evidence for rows of fused tubercles in some areas. The median ventral plate is overlapped by anterior ventral lateral and posterior ventral lateral plates. Two such overlap surfaces are continuous with one another and visible on the dermal surface of the median ventral plate of NUFV 892. It is impossible to determine which plates overlap these visible surfaces because the plate is out of articulation in NUFV 892.
</paragraph>
<paragraph id="D15C3697BF13E569FF00CC90DB853855" blockId="11.[132,775,1269,1743]" box="[132,412,1269,1296]" pageId="11" pageNumber="688">
<emphasis id="E397EA85BF13E569FF00CC90DB853855" box="[132,412,1269,1296]" italics="true" pageId="11" pageNumber="688">
Mixilateral plate (
<figureCitation id="49D82A12BF13E569FEC5CC93DB8A3855" box="[321,403,1270,1296]" captionStart="FIG" captionStartId="10.[132,143,1337,1355]" captionTargetBox="[138,1452,219,1294]" captionTargetId="figure@10.[132,1455,214,1316]" captionTargetPageId="10" captionText="FIG. 5. — Asterolepis alticristata n. sp.: A-C, NUFV 892,left side of articulated partial thoracic skeleton in visceral (A, photograph;A’, labeled illustration),left lateral (B, photograph; B’, labeled illustration), and cranial (C, photograph; C’, labeled illustration) views; D, NUFV 895, right mixilateral plate in dermal view. Abbreviations:ADL, anterior dorsolateral plate;AVL, anterior ventrolateral;c1, central 1 plate; cd, area overlapped by ventral thoracic plates;cd.AMD, area overlapped by the anterior median dorsal plate; cr.d, crescent-shaped depression; cv.Mxl, area overlapping the mixilateral plate; dlr, dorsolateral ridge; f.ax, foramen axillaris; lcg, main lateral line groove; MV, median ventral plate; pr.b, processus brachialis; pr.o, processus obstans. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.3695604" httpUri="https://zenodo.org/record/3695604/files/figure.png" pageId="11" pageNumber="688">Fig. 5D</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="D15C3697BF13E569FF00CD70D8713B8B" blockId="11.[132,775,1269,1743]" pageId="11" pageNumber="688">
Only one specimen (NUFV 895;
<figureCitation id="49D82A12BF13E569FE64CD70D8283875" box="[480,561,1301,1328]" captionStart="FIG" captionStartId="10.[132,143,1337,1355]" captionTargetBox="[138,1452,219,1294]" captionTargetId="figure@10.[132,1455,214,1316]" captionTargetPageId="10" captionText="FIG. 5. — Asterolepis alticristata n. sp.: A-C, NUFV 892,left side of articulated partial thoracic skeleton in visceral (A, photograph;A’, labeled illustration),left lateral (B, photograph; B’, labeled illustration), and cranial (C, photograph; C’, labeled illustration) views; D, NUFV 895, right mixilateral plate in dermal view. Abbreviations:ADL, anterior dorsolateral plate;AVL, anterior ventrolateral;c1, central 1 plate; cd, area overlapped by ventral thoracic plates;cd.AMD, area overlapped by the anterior median dorsal plate; cr.d, crescent-shaped depression; cv.Mxl, area overlapping the mixilateral plate; dlr, dorsolateral ridge; f.ax, foramen axillaris; lcg, main lateral line groove; MV, median ventral plate; pr.b, processus brachialis; pr.o, processus obstans. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.3695604" httpUri="https://zenodo.org/record/3695604/files/figure.png" pageId="11" pageNumber="688">Fig. 5D</figureCitation>
) is complete enough to both recognize it as a mixilateral plate and assign it to
<taxonomicName id="16E34D14BF13E569FD45CD53DB06382A" authority="Downs &amp; Daeschler &amp; Lo &amp; Carey &amp; Shubin, 2019" authorityName="Downs &amp; Daeschler &amp; Lo &amp; Carey &amp; Shubin" authorityYear="2019" class="Insecta" family="Tortricidae" genus="Asterolepis" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="688" phylum="Arthropoda" rank="species" species="alticristata" status="sp. nov.">
<emphasis id="E397EA85BF13E569FD45CD53DB06382A" italics="true" pageId="11" pageNumber="688">Asterolepis alticristata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F8A457FEBF13E569FEA0CD33DB453835" box="[292,348,1366,1392]" pageId="11" pageNumber="688" rank="species">n. sp.</taxonomicNameLabel>
NUFV 895 is incomplete and lacks most of the zones of overlap that accommodate the anterior median dorsal and posterior median dorsal plates. But for a short segment, the specimen also lacks the plate’s lateral lamina. The ornament is densely tuberculate with the tubercles arranged in rows close to cranial and caudomedial edges. These rows of tubercles meet each of these two ornamented margins at steeply oblique angles. On the visceral surface, oblique ridges extend caudolaterally from the caudal end of the mixilateral’s contact with the anterior median dorsal plate. These continue the path of the ridges on the visceral surface of the anterior median dorsal plate that are described above.
</paragraph>
<paragraph id="D15C3697BF13E569FF00CE90DB873A4B" blockId="11.[132,775,1780,2030]" box="[132,414,1780,1806]" pageId="11" pageNumber="688">
<emphasis id="E397EA85BF13E569FF00CE90DB873A4B" box="[132,414,1780,1806]" italics="true" pageId="11" pageNumber="688">Posterior ventrolateral plate</emphasis>
</paragraph>
<paragraph id="D15C3697BF13E569FF00CF71DBB03AA8" blockId="11.[132,775,1780,2030]" pageId="11" pageNumber="688">
Several fragments (NUFV 802.1, 802.2, 821, 848) are identifiable as the caudoventral corner of the posterior ventrolateral plates and are here attributed to
<taxonomicName id="16E34D14BF13E569FE60CF36D8DC3A28" authority="Downs &amp; Daeschler &amp; Lo &amp; Carey &amp; Shubin, 2019" authorityName="Downs &amp; Daeschler &amp; Lo &amp; Carey &amp; Shubin" authorityYear="2019" box="[484,709,1875,1901]" class="Insecta" family="Tortricidae" genus="Asterolepis" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="688" phylum="Arthropoda" rank="species" species="alticristata" status="sp. nov.">
<emphasis id="E397EA85BF13E569FE60CF36D8DC3A28" box="[484,709,1875,1901]" italics="true" pageId="11" pageNumber="688">Asterolepis alticristata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F8A457FEBF13E569FD48CF31D91E3A2B" box="[716,775,1876,1902]" pageId="11" pageNumber="688" rank="species">n. sp.</taxonomicNameLabel>
They are not complete enough to support an anatomical description but for the ornament pattern of rows of isolated or conjoined tubercles that is consistent with much of the rest of the known skeleton.
</paragraph>
<paragraph id="D15C3697BF13E569FCA9C8BCDE6C3DB6" blockId="11.[813,1455,217,563]" box="[813,1141,217,244]" pageId="11" pageNumber="688">
<emphasis id="E397EA85BF13E569FCA9C8BCDE6C3DB6" box="[813,1141,217,244]" italics="true" pageId="11" pageNumber="688">
Pectoral appendage (
<figureCitation id="49D82A12BF13E569FB84C8BFDE753DB6" box="[1024,1132,217,244]" captionStart="FIG" captionStartId="10.[132,143,1337,1355]" captionTargetBox="[138,1452,219,1294]" captionTargetId="figure@10.[132,1455,214,1316]" captionTargetPageId="10" captionText="FIG. 5. — Asterolepis alticristata n. sp.: A-C, NUFV 892,left side of articulated partial thoracic skeleton in visceral (A, photograph;A’, labeled illustration),left lateral (B, photograph; B’, labeled illustration), and cranial (C, photograph; C’, labeled illustration) views; D, NUFV 895, right mixilateral plate in dermal view. Abbreviations:ADL, anterior dorsolateral plate;AVL, anterior ventrolateral;c1, central 1 plate; cd, area overlapped by ventral thoracic plates;cd.AMD, area overlapped by the anterior median dorsal plate; cr.d, crescent-shaped depression; cv.Mxl, area overlapping the mixilateral plate; dlr, dorsolateral ridge; f.ax, foramen axillaris; lcg, main lateral line groove; MV, median ventral plate; pr.b, processus brachialis; pr.o, processus obstans. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.3695604" httpUri="https://zenodo.org/record/3695604/files/figure.png" pageId="11" pageNumber="688">Fig. 5A, B</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="D15C3697BF13E569FCA9C89CDEDA3F76" blockId="11.[813,1455,217,563]" pageId="11" pageNumber="688">
The identifiable plates of the
<taxonomicName id="16E34D14BF13E569FBDBC89CDF273C56" authority="Downs &amp; Daeschler &amp; Lo &amp; Carey &amp; Shubin, 2019" authorityName="Downs &amp; Daeschler &amp; Lo &amp; Carey &amp; Shubin" authorityYear="2019" box="[1119,1342,249,275]" class="Insecta" family="Tortricidae" genus="Asterolepis" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="688" phylum="Arthropoda" rank="species" species="alticristata" status="sp. nov.">
<emphasis id="E397EA85BF13E569FBDBC89CDF273C56" box="[1119,1342,249,275]" italics="true" pageId="11" pageNumber="688">Asterolepis alticristata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F8A457FEBF13E569FAC7C89FDF643C51" box="[1347,1405,250,276]" pageId="11" pageNumber="688" rank="species">n. sp.</taxonomicNameLabel>
pectoral appendage include dorsal and ventral central 1 plates (
<figureCitation id="49D82A12BF13E569FCB2C95CD9943C16" box="[822,909,312,340]" captionStart="FIG" captionStartId="10.[132,143,1337,1355]" captionTargetBox="[138,1452,219,1294]" captionTargetId="figure@10.[132,1455,214,1316]" captionTargetPageId="10" captionText="FIG. 5. — Asterolepis alticristata n. sp.: A-C, NUFV 892,left side of articulated partial thoracic skeleton in visceral (A, photograph;A’, labeled illustration),left lateral (B, photograph; B’, labeled illustration), and cranial (C, photograph; C’, labeled illustration) views; D, NUFV 895, right mixilateral plate in dermal view. Abbreviations:ADL, anterior dorsolateral plate;AVL, anterior ventrolateral;c1, central 1 plate; cd, area overlapped by ventral thoracic plates;cd.AMD, area overlapped by the anterior median dorsal plate; cr.d, crescent-shaped depression; cv.Mxl, area overlapping the mixilateral plate; dlr, dorsolateral ridge; f.ax, foramen axillaris; lcg, main lateral line groove; MV, median ventral plate; pr.b, processus brachialis; pr.o, processus obstans. Scale bar: 5 cm." figureDoi="http://doi.org/10.5281/zenodo.3695604" httpUri="https://zenodo.org/record/3695604/files/figure.png" pageId="11" pageNumber="688">Fig. 5A</figureCitation>
[c1], B[c1]) and a lateral marginal 2 plate, but the material also includes additional unidentified central and marginal plates. The dorsal and ventral central 1 and lateral marginal 2 plates appear as they do in the
<typeStatus id="0E588835BF13E569FB79C9FCDF323CF6" box="[1277,1323,409,435]" pageId="11" pageNumber="688">type</typeStatus>
species. On the central plates, the ornament is tuberculate with larger, more widely spaced tubercles at the proximal end. On the marginal plate, the ornament is densely tuberculate with no obvious pattern to their arrangement.
</paragraph>
</subSubSection>
</treatment>
</document>
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