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<document id="4BD1D1FC24BB4A75F1644845B7C588B7" ENCODING="UTF-8" ID-GBIF-Dataset="d8efa4ec-b5a3-457c-8492-6d60d7f0576f" ModsDocID="z01202p001" checkinTime="1247156166445" checkinUser="thomas" docAuthor="Yukio Iwatsuki &amp; Kent E. Carpenter" docDate="2006" docId="5932F797FEAE25243789CBA282E24C5B" docLanguage="en" docName="2006_Iwatsuki_Carpenter_gg1.xml" docOrigin="Zootaxa 1202" docSource="http://www.zoobank.org/urn:lsid:zoobank.org:pub:E4C0CD38-51A6-4B95-B54E-056ECE859536" docTitle="Acanthopagrus taiwanensis Iwatsuki &amp; Carpenter, 2006, n. sp." docType="treatment" docVersion="15" lastPageNumber="15" masterDocId="6042678B25E3536A72E43E7BE14ED41A" masterDocTitle="Acanthopagrus taiwanensis, a new sparid fish (Perciformes), with comparisons to Acanthopagrus berda (Forsskål, 1775) and other nominal species of Acanthopagrus." masterLastPageNumber="19" masterPageNumber="1" pageNumber="4" updateTime="1701326208839" updateUser="ExternalLinkService">
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<mods:title id="5292B8C1F37FF4A1D64B8A5DFDF860F3">Acanthopagrus taiwanensis, a new sparid fish (Perciformes), with comparisons to Acanthopagrus berda (Forsskål, 1775) and other nominal species of Acanthopagrus.</mods:title>
</mods:titleInfo>
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<mods:role id="26D405749924150C53772A96BC61B1F1">
<mods:roleTerm id="3F68522B5FA70F079B60499FCD969335">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6FEDCBB473C455E0778EE48B6D303DA3">Yukio Iwatsuki</mods:namePart>
</mods:name>
<mods:name id="58632AF092C038538FEE8A21F71E863B" type="personal">
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<mods:roleTerm id="C2F6D0E895B8890F126B00CB20651C46">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5A7DA3ADD5C360CAFB7EF9C1771BF924">Kent E. Carpenter</mods:namePart>
</mods:name>
<mods:typeOfResource id="A95427F0E85497575871DE80DB707926">text</mods:typeOfResource>
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<mods:date id="4CBCE64A68F43F8A15E6BBD135E77799">2006</mods:date>
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<mods:number id="7FFA0209F62297CAA3F78EFDC1442DB6">1202</mods:number>
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</mods:location>
<mods:classification id="21A1917BCB2DD9914D43DF6C427BB9F4">journal article</mods:classification>
<mods:identifier id="D19B9B9E3294E8A2C70DAC07815D3F66" type="Plazi-Custom">z01202p001</mods:identifier>
</mods:mods>
<treatment id="5932F797FEAE25243789CBA282E24C5B" ID-DOI="http://doi.org/10.5281/zenodo.6492271" ID-GBIF-Taxon="100118926" ID-Zenodo-Dep="6492271" LSID="urn:lsid:zoobank.org:act:65CDB643-2754-4FE4-92C5-E4EB68DF0DA2" httpUri="http://treatment.plazi.org/id/5932F797FEAE25243789CBA282E24C5B" lastPageNumber="15" pageNumber="4">
<subSubSection id="7B9F29AD6BEAF37ABFEE9A0AE46975D4" type="nomenclature">
<paragraph id="DBF95AA4C4EB223DACCB7D1F6FA268DD" pageNumber="4">
<taxonomicName id="4AA531650D774785063DC95D7C044B6C" ID-CoL="8YVV" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="4" phylum="Chordata" rank="species" species="taiwanensis">Acanthopagrus taiwanensis</taxonomicName>
<taxonomicNameLabel id="A44118D4DC44B972B21A72D61A12B04B" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
<paragraph id="7B1DABF4F4C0623B198E37470963D639" pageNumber="4">New English Name: Taiwan Picnic Seabream</paragraph>
</subSubSection>
<subSubSection id="5B42AA241F9AFAA6B5BAB0CB81EE35F2" type="description">
<paragraph id="51F4C48E7B100BDC4C62C2392999E01A" pageNumber="4">Figs. 1A, 2D -E, 3A -4A and 5A -B, Table 1</paragraph>
</subSubSection>
<subSubSection id="A1BCD5B1BC0459AAE5970E1ECF126C0C" type="nomenclature">
<paragraph id="CBF8540D18D19622C3616961C2E4788C" pageNumber="4">
<materialsCitation id="AE80070C594C4416CC492805B2C1F031" ID-GBIF-Occurrence="923906749" collectingDate="2005-05-22" collectingMethod="hook-and-line (according to sellers in market)" collectionCode="MUFS" collectorName="Y. Iwatsuki" country="Taiwan" location="estuary basin of Tung-kang River (purchased in Tung-kang Fish Market), southwestern Taiwan" specimenCode="MUFS 22854" typeStatus="Holotype">
<typeStatus id="DFD9791C55378CBF9C58B0037087416C">Holotype</typeStatus>
:
<specimenCode id="CFBF4EF41A0041490BF00418AAA3EF31">
<collectionCode id="BBCC4FB45E4879EC032772AEB3B8DAFF">MUFS</collectionCode>
22854
</specimenCode>
, male, 167 mm SL,
<location id="E5D2781B0375C8BC3E523B0340597C5A" LSID="urn:lsid:plazi:treatment:5932F797FEAE25243789CBA282E24C5B:E5D2781B0375C8BC3E523B0340597C5A" country="Taiwan" name="estuary basin of Tung-kang River (purchased in Tung-kang Fish Market), southwestern Taiwan">estuary basin of Tung-kang River (purchased in Tung-kang Fish Market), southwestern Taiwan</location>
,
<collectorName id="DC74A36B2956C8759AE301AA213AE2A8">Y. Iwatsuki</collectorName>
,
<collectingMethod id="8EA87D9E67FA2F78D7EA530E4647041B">hook-and-line (according to sellers in market)</collectingMethod>
,
<collectingDate id="40EEE5E91F2C05216ABB54D173C27631">22 May, 2005</collectingDate>
</materialsCitation>
.
</paragraph>
<paragraph id="58E910FED6E37F9FE47B7A8AE7D4FF61" pageNumber="4">
<materialsCitation id="40AC78466423EABBF19CA861E143839E" ID-GBIF-Occurrence="923906759" collectingDate="1973-02-25" collectionCode="MUFS" collectorName="M. Akazaki" country="Taiwan" location="Tung-kang, southwestern Taiwan" specimenCode="MUFS 11870" typeStatus="Paratype">
<typeStatus id="1353EFD0D33CF99F2DA39F7ED06DDDC0">Paratypes</typeStatus>
: (5 specimens)
<specimenCode id="AEC1042EC6ABB9B0C5E99AB2F6E16150">
<collectionCode id="5DB1F6EB0C08A3A294C152C6749EB11D">MUFS</collectionCode>
11870
</specimenCode>
, sex not determined, 110 mm SL,
<location id="98AA98FA4559AA6D3AAF8EFB9B3C433E" LSID="urn:lsid:plazi:treatment:5932F797FEAE25243789CBA282E24C5B:98AA98FA4559AA6D3AAF8EFB9B3C433E" country="Taiwan" name="Tung-kang, southwestern Taiwan">Tung-kang, southwestern Taiwan</location>
,
<collectorName id="234287A35D09198C56E51F4FE91E6592">M. Akazaki</collectorName>
,
<date id="1EB77A93A893F4F6F283E52A7BE8C372" day="25" month="02" value="1973-02-25" year="1973">
<collectingDate id="2DAD26703467480C425F45BA5A2E1EF1" day="25" month="02" value="1973-02-25" year="1973">25 February 1973</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="61A58E6AB636E14DFA672099F6E30DBE" ID-GBIF-Occurrence="923906756" collectingDate="2002-12-27" collectionCode="MUFS" collectorName="Y. Iwatsuki" country="Taiwan" location="Tung-kang, southwestern Taiwan" specimenCode="MUFS 22165" typeStatus="Paratype">
<specimenCode id="045840A74CD9DD3FC760E51F0D36C19A">
<collectionCode id="C777D59F1FF6213C8E72C26EDC72032F">MUFS</collectionCode>
22165
</specimenCode>
, sex not determined, 184 mm SL,
<location id="B30CB7B33D6313C1E50DDB416C1D29AE" LSID="urn:lsid:plazi:treatment:5932F797FEAE25243789CBA282E24C5B:B30CB7B33D6313C1E50DDB416C1D29AE" country="Taiwan" name="Tung-kang, southwestern Taiwan">Tung-kang, southwestern Taiwan</location>
,
<collectorName id="30D54E19626CA49BD5D0AD0F069F49AE">Y. Iwatsuki</collectorName>
,
<date id="94D570FD4291207780BF7DBCDDC8E3D3" day="27" month="12" value="2002-12-27" year="2002">
<collectingDate id="B960FED303280B1B3B251B3F995AD2A4" day="27" month="12" value="2002-12-27" year="2002">27 December 2002</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="1127FF2DAC21B40582861785E429AF9D" ID-GBIF-Occurrence="923906758" collectingDate="2002-12-27" collectionCode="MUFS" collectorName="Y. Iwatsuki" country="Taiwan" location="Tung-kang, southwestern Taiwan" specimenCode="MUFS 22166" typeStatus="Paratype">
<specimenCode id="474620937224A2546C07D63BE130A39C">
<collectionCode id="CCFE3519643BF94F4AF17A8872995CB8">MUFS</collectionCode>
22166
</specimenCode>
, female, 216 mm,
<location id="4F52A97AFBDE110A0E56DC2A4E5B7E32" LSID="urn:lsid:plazi:treatment:5932F797FEAE25243789CBA282E24C5B:4F52A97AFBDE110A0E56DC2A4E5B7E32" country="Taiwan" name="Tung-kang, southwestern Taiwan">same data as MUFS 22165</location>
</materialsCitation>
;
<materialsCitation id="CA52C947D78738976AFFED262A0470BB" ID-GBIF-Occurrence="923906761" collectingDate="2005-05-22" collectingMethod="hook-and-line (according to sellers in market)" collectionCode="MUFS" collectorName="Y. Iwatsuki" country="Taiwan" location="estuary basin of Tung-kang River (purchased in Tung-kang Fish Market), southwestern Taiwan" specimenCode="MUFS 22855" typeStatus="Paratype">
<specimenCode id="E5B67667959062C99C38C58FF0200A4F">
<collectionCode id="5F89D42E5477E6D43606A6A64B612242">MUFS</collectionCode>
22855
</specimenCode>
, female, 175 mm SL,
<location id="193746833D9806014C1F7C6EC4255188" LSID="urn:lsid:plazi:treatment:5932F797FEAE25243789CBA282E24C5B:193746833D9806014C1F7C6EC4255188" country="Taiwan" name="estuary basin of Tung-kang River (purchased in Tung-kang Fish Market), southwestern Taiwan">data same as holotype</location>
</materialsCitation>
;
<materialsCitation id="68B2EDBA128064C4BA71DB1E1CA4191A" ID-GBIF-Occurrence="923906772" collectingDate="2005-05-22" collectingMethod="shrimp set nets" collectionCode="MUFS" collectorName="Y. Iwatsuki" country="Taiwan" location="mouth of Tungkang River, southwestern Taiwan" specimenCode="MUFS 22857" typeStatus="Paratype">
<specimenCode id="0EF651FEDAC66E9DD5AC378CC593EC41">
<collectionCode id="31D74F5551354236D6F402AA4F114189">MUFS</collectionCode>
22857
</specimenCode>
, sex not determined, 106 mm SL,
<location id="B6865F782204DF552CAF9E0B4E4D3000" LSID="urn:lsid:plazi:treatment:5932F797FEAE25243789CBA282E24C5B:B6865F782204DF552CAF9E0B4E4D3000" country="Taiwan" name="mouth of Tungkang River, southwestern Taiwan">mouth of Tungkang River, southwestern Taiwan</location>
,
<collectorName id="A54C0B0A28B1D97BBDC757BC384A5638">Y. Iwatsuki</collectorName>
,
<collectingMethod id="80452FE0770A7947E8331A887B823F90">shrimp set nets</collectingMethod>
,
<collectingDate id="8C1B351148371CC9051B6FB3358E292F">22 May, 2005</collectingDate>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="074B94AEB41ECA2CDB06F6ABD383FCCD" type="diagnosis">
<paragraph id="4C6DA836A4F4755500432CED98502BED" pageNumber="4">Diagnosis</paragraph>
<paragraph id="4DD0DED8A0586F574E57D1C41183D8FE" lastPageNumber="5" pageNumber="4">
Dorsal fin with 11 or 12 spines and 10 to 12 soft rays; anal fin with 3 spines and 8 or 9 soft rays; 3 ½ scale rows between the fifth dorsal-fin spine base and lateral line; 3 or 4 cheek scale rows; body rounded; ventral infraorbital series above posterior part of upper jaw changing from almost straight to weakly concave with growth (Figs. 2B, 3A); second anal-fin spine robust and dull pointed (Fig. 2C); upper and lower jaws with thin lips, especially anteriorly; broadly rounded rostral-most head squamation with around 10 somewhat smaller scales anteriorly (Fig. 4A); prominent ridge developing just before eye with growth, dorsal head profile becoming slightly convex from snout to just above eye with growth (Figs. 1A, 2B and 3A); upper and lower molar teeth strongly developed and flattened on both sides, gradually more molariform posteriorly (Fig. 5A -B); upper molars
<pageStartToken id="BD482F6B7261A6B2690B3AC3CABA0A7F" pageNumber="5">in</pageStartToken>
3 or 4 rows anteriorly and 4 or 5 posteriorly, rows generally irregular, outer third row markedly larger; lower molars in 3 or 4 rows anteriorly and 2 or 3 rows posteriorly, rows generally irregular, the innermost row largest posteriorly (outer second row gently curved outward)(Fig. 5C -D); head and body mostly black, sharply demarcated from whitish belly and chin (Figs. 1A and 3A); a dark spot at top of pectoral-fin base (Figs. 1A and 3A).
</paragraph>
</subSubSection>
<subSubSection id="40A2EEDBCEB8EB279E957C1ADB988242" type="multiple">
<caption id="4DCDBCDB1BD23F0FE96318AE50B75759">
<paragraph id="A48633B3AFDD7A233BEC35E1FAE729E7" pageNumber="5">
FIGURE 1. A
<taxonomicName id="CE8C6C3278835E9C9F8283F4BDA631D9" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="5" phylum="Chordata" rank="species" species="taiwanensis">Acanthopagrus taiwanensis</taxonomicName>
<taxonomicNameLabel id="4F57A6C9466281BEEF294C13B5E0A981" rank="species">n. sp.</taxonomicNameLabel>
, 22854, holotype, 167 mm SL, Tung-kang, southwestern Taiwan and B
<taxonomicName id="6C4CAAC83553A8C437745502D1F75513" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="5" phylum="Chordata" rank="species" species="berda">
A. berda (
<normalizedToken id="B680ACB9EA6D7B3204678F71B408280E" originalValue="Forrskål">Forrskal</normalizedToken>
, 1775)
</taxonomicName>
, MUFS 22938, 183 mm SL, Tung-kang, Taiwan.
</paragraph>
</caption>
<caption id="BEF7CCE8A0D5AB8B2653BAF0F77C497A">
<paragraph id="9E697F0C3E45F767B6C7748932D3F81A" pageNumber="6">
<pageStartToken id="398E6C0FE8659FF366725FFF4EBC0F7A" pageNumber="6">FIGURE</pageStartToken>
2. A,
<taxonomicName id="6AEA9F03D3DF3BD25D1C5CB20B6E2EE0" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="6" phylum="Chordata" rank="species" species="berda">
Acanthopagrus berda (
<normalizedToken id="7B1E0E314639035E3492F720CD42C324" originalValue="Forrskâl">Forrskal</normalizedToken>
, 1775)
</taxonomicName>
, ZMUC-P 50555 (left side dried skin), holotype, presumably ca. 213 mm SL, Luhaiya, Yemen, Red Sea, B-C,
<taxonomicName id="90BED1540374F858D7A188150998746B" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="6" phylum="Chordata" rank="species" species="taiwanensis">A. taiwanensis</taxonomicName>
, MUFS 22166, paratype, 212 mm SL, Tung-kang, southwestern Taiwan and D-E, ventral edge of first two infraorbitals above posterior part of upper jaw and anal-fin rays of
<taxonomicName id="00AC95F5A07089EF4E13B272ECF4B8C6" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="6" phylum="Chordata" rank="species" species="berda">A. berda</taxonomicName>
, ZRC 38094, 249 mm SL, Kisap River, Langkawi Island, Malaysia. Holotype of
<taxonomicName id="A7F322EB73119633BEAB30A813708CB7" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="6" phylum="Chordata" rank="species" species="berda">
Acanthopagrus berda (
<normalizedToken id="B82CFBFE1929E9EA302922C66C5F9295" originalValue="Forsskål">Forsskal</normalizedToken>
, 1775)
</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="9F5F76B0CF299FCCD2574E37A24819D8" type="description">
<paragraph id="F470D1290F91001BD80B001FA00ECF4A" pageNumber="6">Description</paragraph>
<paragraph id="910E1C473429C1A72B40835F0C761717" pageNumber="6">Counts and measurements of the holotype and 5 paratypes are given in Table 1. In the following description, data for the holotype are presented first, followed by the 5 paratype specimens in parentheses where different. Characters stated in the diagnosis are not repeated.</paragraph>
<caption id="421E09D011F8CD3384245B37D4D951C6">
<paragraph id="B7B9EFD0C907E43EAC31B395202A8D14" pageNumber="7">
<pageStartToken id="9D9D920C40E0CC2EF649B7FF433961B3" pageNumber="7">FIGURE</pageStartToken>
3. Head of: A,
<taxonomicName id="053382AB0AD7DEE534D5C2278F3AFEEC" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="7" phylum="Chordata" rank="species" species="taiwanensis">Acanthopagrus taiwanensis</taxonomicName>
, MUFS 22166, paratype, 212 mm SL, Tungkang, southwestern Taiwan and B,
<taxonomicName id="8A092AF6A31221B1B6E891201397A528" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="7" phylum="Chordata" rank="species" species="berda">
A. berda (
<normalizedToken id="D9DECD3D495943AC1568FA3BABF91058" originalValue="Forrskål">Forrskal</normalizedToken>
, 1775)
</taxonomicName>
, RMNH 15683, 241 mm SL, Red Sea.
</paragraph>
</caption>
<caption id="892610EAEAAC2470D93AC481F3A29348">
<paragraph id="5D1DE9ABD492C441436D9FD7F705E4CF" pageNumber="8">
<pageStartToken id="7452EACB2B39BE8DC07668C4BC9FDFE7" pageNumber="8">TABLE</pageStartToken>
1. Counts and morphometric data for the holotype of
<taxonomicName id="F6C457A35B54AFD5CF5E29DCFE1CB7A9" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="8" phylum="Chordata" rank="species" species="taiwanensis">Acanthopagrus taiwanensis</taxonomicName>
<taxonomicNameLabel id="3675823F00BC27FC66048BCCAAB9602A" rank="species">n. sp.</taxonomicNameLabel>
and
<taxonomicName id="698D4918FAD909F71B5BA50E0BEE2D41" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="8" phylum="Chordata" rank="species" species="berda">
A. berda (
<normalizedToken id="66F5C4FAA7C979E3A0AD5A3CA558E82A" originalValue="Forsskål">Forsskal</normalizedToken>
, 1775)
</taxonomicName>
. Proprtional measurements expressed as percentage of standard length except where noted.
</paragraph>
</caption>
<paragraph id="0E86780A7F51BA7D52E2C01028EDDD16">
<table id="45A78EF686909AFFECCD2FFC186DA196" border="0" frame="box" pageNumber="8" rules="all">
<tr id="B187054255745661A37B006EED2094EF" pageNumber="8">
<td id="97C2BDBDA6F60501358711B8DECA5B53" colspan="2" pageNumber="8">
<taxonomicName id="8EA1E018AD1501A8B670A097A2F5D4FA" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="taiwanensis">Acanthopagrus taiwanensis</taxonomicName>
<taxonomicNameLabel id="E5CE0CA6A1A0E2F71EF50F63BD6E5221" rank="species">n. sp.</taxonomicNameLabel>
</td>
<td id="F09337354CB4EA3DEB7896E23B1304F0" colspan="2" pageNumber="8">
<taxonomicName id="9A9B3A1A7A0C08A8515C0FBCEFDCF23C" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="berda">Acanthopagrus berda</taxonomicName>
</td>
</tr>
<tr id="F0397253960DDBDCD46F4594912DE237" pageNumber="8">
<td id="832FF5D6EB9C42F7455A84CB456B11FC" pageNumber="8">Holotype</td>
<td id="D4FF34FF3BB06CEC720653DB81D2A704" pageNumber="8">Paratypes</td>
<td id="2FB91828A7AF8149111D89654F97257B" pageNumber="8">Holotype</td>
<td id="9654EA5E278C4464B1742FD551416A7C" pageNumber="8">Non-type specimens</td>
</tr>
<tr id="B584A23A04AC57E3B693E7B12154EB31" pageNumber="8">
<td id="FE1B59EA1289F8AFAB29B98FE33739D6" pageNumber="8">MUFS 22854</td>
<td id="51E8074B9E1C5B4B729A88A51F61119E" pageNumber="8">MUFS 11870, 22165, 22854, 22855, 22857 n = 5</td>
<td id="EC1EC5394D10078A227F970943C96A80" pageNumber="8">ZMUC P50555 n = 1</td>
<td id="6E3C3EF665B2D4AF8D36CC8C619A55E5" pageNumber="8">n = 39</td>
</tr>
<tr id="3A7E0EBD38A766EBC66EB13A012CA53C" pageNumber="8">
<td id="84C92074A5674F432ABF4C8525459B71" colspan="5" pageNumber="8">Counts:</td>
</tr>
<tr id="C3266985F27D2EDDB33D5A0BD6902A47" pageNumber="8">
<td id="66BA86DB72867EA59E158506D06C7C97" pageNumber="8">Dorsal fin rays</td>
<td id="06401200AEFCDC67B5319B6027ECCE76" pageNumber="8">XI, 11</td>
<td id="630FBE6311BEFE3CBBD3E9620629D437" pageNumber="8">XI, 11 (XII, 10 and XI, 12 at each)</td>
<td id="F1A08FBF4660AD6BCC452F16CED2315C" pageNumber="8">XI, -</td>
<td id="CE93BAA7CC39699FAE7FE0434BFCC060" pageNumber="8">usually XI, 11 (XII, 10 or XI, 12)</td>
</tr>
<tr id="493F00387241958B70D8A009B841AC6C" pageNumber="8">
<td id="6CC595E8B5A74DF1CBBB2932C0189DEF" pageNumber="8">Anal fin rays</td>
<td id="30A4EA843C2E4C08195E1411D3787B18" pageNumber="8">III, 9</td>
<td id="39D3D94AE3557449EA7B2B1860BA7D34" pageNumber="8">III, 8-9</td>
<td id="555E1C271D37F2A481153F07D1C69BF4" pageNumber="8">III, 9</td>
<td id="15870E1E943D0510BA36ABC0647A8F91" pageNumber="8">III, 8-9 (usually 9)</td>
</tr>
<tr id="3A6E0ECE4E86D5872FB7FEC5375EEADA" pageNumber="8">
<td id="CF3A181FCA1B04639B12705623990E20" pageNumber="8">Pectoral-fin rays</td>
<td id="24C25711CAE9C41EC22A202D3D27F40A" pageNumber="8">15</td>
<td id="12C6D8B2D378A778772046ED12581950" pageNumber="8">15</td>
<td id="289F9C36816C82FF7430CD24038F004E" pageNumber="8">15</td>
<td id="B67A9B2F46C502FE1C86F8FF4C48118C" pageNumber="8">usually 15 (rarely 14)</td>
</tr>
<tr id="D81798F338E7D6CA6B202399965EDA19" pageNumber="8">
<td id="EA09CA1DEE66927D98B4A6F1309CE8D9" pageNumber="8">Pored lateral-line scales</td>
<td id="127BA532A490A80A419275854187CBAB" pageNumber="8">43</td>
<td id="C67C6D7425C618BD14950C908F7AED7B" pageNumber="8">43 or 44</td>
<td id="74CB48CE298DC0195411B1EDB6F95633" pageNumber="8">-</td>
<td id="BF29F1E5276C5922B1C42E9AE8508D94" pageNumber="8">42-44</td>
</tr>
<tr id="EB2B5758D5889F024ED276E5857321F0" pageNumber="8">
<td id="739CB915CC082A15E8BBF139392293C9" pageNumber="8">Scales above/below lateral line</td>
<td id="E35214DB063DEA607C604C906053A755" pageNumber="8">4 / 11</td>
<td id="6294F2E13A896060E804178BDEDE5CE2" pageNumber="8">3 ½-4 / 11</td>
<td id="9FFDE39B4211F54555538927B86D1127" pageNumber="8">-</td>
<td id="FDEA3FA5092E1390746F2C2106B0866D" pageNumber="8">3 ½-4 ½ / 11-12</td>
</tr>
<tr id="2D62FAD7C1A1E115D0FE5DF325D8D7A5" pageNumber="8">
<td id="9F8C9B764F95BB5ECA40D28030509660" pageNumber="8">Scales from 5th dorsal spine base to lateral line</td>
<td id="76A2A68D2A54D772F7238D5622C294A5" pageNumber="8">3 ½</td>
<td id="DF6B126929D2A607FF6401A541581BB8" pageNumber="8">3 ½</td>
<td id="FC38903AF9EDDD1232A749CA974490B7" pageNumber="8">3 ½</td>
<td id="CA6ABDD42C7D47B866A1A788A991C822" pageNumber="8">3 ½</td>
</tr>
<tr id="AB70909D231FBDABE53EBDE3B6308948" pageNumber="8">
<td id="7A568A9149C8E2E773CA359A184C0295" pageNumber="8">Gill rakers</td>
<td id="56D38A7A5B1C90D0BC24BD50AAE52FC5" pageNumber="8">4+9 = 13</td>
<td id="8ED6483EA78A09BBA0EE964B42AC72FA" pageNumber="8">4-5+9-10 = 13-15</td>
<td id="4FA0CE5A982957D545CA1EE29823E1BA" pageNumber="8">-</td>
<td id="075A817EE65B786ED48E0B8A3AFF06CA" pageNumber="8">5-7 (usually 5 or 6) + 8-12 (usually 9 or 10) = 13-17 (usually 14 or 15)</td>
</tr>
<tr id="A037874E413134AE488A3F8767D819F6" pageNumber="8">
<td id="FE119D41599BC8A9C8CA7646D90CEEB3" pageNumber="8">Standard length (mm)</td>
<td id="6791B0B08E53B9D55A5A2E52EBF71AAD" pageNumber="8">167</td>
<td id="C3D8C8FEB08E7DEDF39786F4B3651536" pageNumber="8">110-212</td>
<td id="D5AD14E89A78D32B397B0177D0495099" pageNumber="8">ca. 213*</td>
<td id="DD2E2BFB9068EAF26FF1570C29BC8E50" pageNumber="8">33-288</td>
</tr>
<tr id="4906D52AA5E3C7D5662ED454DD3CEF1F" pageNumber="8">
<td id="6025E38C9F5F7007680BEB8F00318014" colspan="5" pageNumber="8">Proportions:</td>
</tr>
<tr id="0D01FA3B09C5D5287D65EE6E0BB7456B" pageNumber="8">
<td id="3355485780C0927A808020870BA70452" pageNumber="8">Body depth</td>
<td id="39648E315CCBE280D4729373E9ED4C97" pageNumber="8">46</td>
<td id="ADCBFC9442006CF3C48A7D886F168337" pageNumber="8">45-49</td>
<td id="32C4BF3648B4CE2AD5C0FC3FD12EEECE" pageNumber="8">-</td>
<td id="1801434FB8CDF8DA932D7336E9DA06F1" pageNumber="8">45-53 (48)</td>
</tr>
<tr id="4BF88B4B90E2D3429E7EF8DD6CB98768" pageNumber="8">
<td id="8DC820AD521FB0B80D7081C2B709084F" pageNumber="8">Head length</td>
<td id="088C7508C9EB88975D54BB86118E1C13" pageNumber="8">33</td>
<td id="C142C32AD24E0B99858BC8707DC13462" pageNumber="8">32-35</td>
<td id="8031C15F422AF88BF2164CA8673FCEEE" pageNumber="8">32</td>
<td id="1ECEB5DF8620516F6F197E16D27EAB23" pageNumber="8">31-41 (35)</td>
</tr>
<tr id="6331BE5E9B5451F84E7327E4E1C409B2" pageNumber="8">
<td id="D1FE5BBF64DD4872504AD2A1C9683711" pageNumber="8">Body width at pectoral fin base</td>
<td id="1FE192F0950AB20D17AA82118C4DE1CF" pageNumber="8">19</td>
<td id="2B8433DFA3603B853418B99419A9FDD7" pageNumber="8">17-21</td>
<td id="A91117A188668736093AB83FA50F3590" pageNumber="8">-</td>
<td id="E82C66B2382BD9931D761D8C51BB29E8" pageNumber="8">13-23 (18)</td>
</tr>
<tr id="5D80385F14940CFB989767B88A20023C" pageNumber="8">
<td id="D8C51C50D0756DF8FB80FCC19CBF0D28" pageNumber="8">Snout length</td>
<td id="904E5F3B7C5E60E159D7AF04773E733A" pageNumber="8">11</td>
<td id="6B1A1BB6B640F1DC9EB5EE8998C83C89" pageNumber="8">9-12</td>
<td id="326B682D34C2B04CA56717E894A92A30" pageNumber="8">13</td>
<td id="93E03305276A769A2F2091D9BE3524AE" pageNumber="8">11-15 (12)</td>
</tr>
<tr id="1D0E9DD42481FF66FEBEAA68D6F04028" pageNumber="8">
<td id="99A9E8CA9DBF691EA38D64C8D7FB9790" pageNumber="8">Orbit diameter</td>
<td id="7D223F3914B508F8BC0EA8CE2FC67FB0" pageNumber="8">9</td>
<td id="40D7F63D88072E4CB9C0850F0D00D31B" pageNumber="8">9-12</td>
<td id="DFBC0E90D25EB7576EF474A48FD357FD" pageNumber="8">10</td>
<td id="D88E0A80F420985EEF9FD98EFA6280D4" pageNumber="8">8-16 (12)</td>
</tr>
<tr id="5F8968FE80F7B63A8E2E1CBF3987090B" pageNumber="8">
<td id="A8A356E6AA0AB545FD6824CB5ED52E7A" pageNumber="8">Dermal eye opening</td>
<td id="734667559E099DA8E0491438E4E888C4" pageNumber="8">8</td>
<td id="4093643237F8761150E68076D6EBBFA6" pageNumber="8">8-11</td>
<td id="DD2A87D835BA35CF27559276346BA11C" pageNumber="8">9</td>
<td id="2770B0DC6F67AF4C099BC1EB2A6B5871" pageNumber="8">6-14 (10)</td>
</tr>
<tr id="3B5E6102C86153EF8D981C2BFABEC735" pageNumber="8">
<td id="6CD6984FD32EF260F5FB47B3AAA34112" pageNumber="8">Bony interorbital width</td>
<td id="B970DD416118D035533DB556D1A22B85" pageNumber="8">9</td>
<td id="9ADC032C768FBFFA1198B00621B74282" pageNumber="8">9-10</td>
<td id="62BD7FBA5D4D14A43D093C4881B682F2" pageNumber="8">-</td>
<td id="8908608ECD4793670455F84DD2F8FC1D" pageNumber="8">7-11 (10)</td>
</tr>
<tr id="95ED320E6555C4EF440C6DA5BB4D08F6" pageNumber="8">
<td id="903D684CECA746FB229A85A50C30B246" pageNumber="8">Upper jaw length</td>
<td id="195D1B1A15FCFF6206511164A03F3AC4" pageNumber="8">13</td>
<td id="9FB12639BD9D51726E839661C7F91607" pageNumber="8">12-14</td>
<td id="5097FEB7DB1AB6BF7894D0D29EBCCB66" pageNumber="8">13</td>
<td id="B2DCCE669FCE5C4F1A189C8BE10C85B4" pageNumber="8">13-17 (15)</td>
</tr>
<tr id="12BEA67F7B24A52558F135961D1A50E2" pageNumber="8">
<td id="286BDA65FDA3B3400A7501655694468B" pageNumber="8">Caudal peduncle depth</td>
<td id="A207D38ED3BC57EDE29551B076AF2A59" pageNumber="8">13</td>
<td id="7A34EF8D515C0C84DC06D1EF21813759" pageNumber="8">13-14</td>
<td id="442439444745D48E4275907EC9991F2C" pageNumber="8">-</td>
<td id="D6250E381B242A44E7B4D4641AD87C1C" pageNumber="8">10-15 (13)</td>
</tr>
<tr id="FBBB01065B75567F4CF930BB1597371B" pageNumber="8">
<td id="18388A064820D4FEC91686BF7307A2A1" pageNumber="8">Caudal peduncle length</td>
<td id="E2D7570308E6A61CF4B6E7822BDB4B6B" pageNumber="8">18</td>
<td id="A5634B238A84AEE1E5A6863E3A34C029" pageNumber="8">18-20</td>
<td id="A6577FF03BA9888102182C52EFD6D104" pageNumber="8">-</td>
<td id="A78CD88322A2603274F2DFE07995080D" pageNumber="8">16-23 (18)</td>
</tr>
<tr id="A263F7D204D7C8738C1AAE0D69125B94" pageNumber="8">
<td id="2A7AAB2898FF9CAD89CDC9EA5DB85B5C" pageNumber="8">Predorsal length</td>
<td id="1FDF19BAFD24583FB1F570FD25FD59B4" pageNumber="8">42</td>
<td id="99C94A64E96281E79C83466BF1E2517D" pageNumber="8">41-43</td>
<td id="32D9D089586F0C9A15D215D26FC08A06" pageNumber="8">-</td>
<td id="1FD8B1179EE8CEB116092EB8EC5339D7" pageNumber="8">42-52 (46)</td>
</tr>
<tr id="B9FFD0B12A597087987FC9A8CF0DA959" pageNumber="8">
<td id="65BE7BF7611773775C2F27F20BA69669" pageNumber="8">Preanal length</td>
<td id="FF0EDAFA5BD03238D61D5F0861F6D9C9" pageNumber="8">65</td>
<td id="343C60CD71ECFBA6DED4008985168F77" pageNumber="8">64-73</td>
<td id="78468840D1A7A8ECE3D40FE882B942E7" pageNumber="8">-</td>
<td id="75AD6EB99CF3500DA616F6BC03D9E023" pageNumber="8">67-74 (71)</td>
</tr>
</table>
</paragraph>
<footnote id="2ADA1E384422F3D7CE8468AF3F976AD1">
<paragraph id="18FA97751750523BE185681AB96FBC15" pageNumber="8">to be continued.</paragraph>
</footnote>
<caption id="2E66ED30479032005A7BDC3723A3C85E">
<paragraph id="558C0196D0DDD0E7F8556A06FEF32E18" pageNumber="9">
<pageStartToken id="D15CB6D305D55325841F80CCD0950E3A" pageNumber="9">TABLE</pageStartToken>
1 (continued).
</paragraph>
</caption>
<paragraph id="D03BA3CBB631554D9A90CB39076F6FC2">
<table id="DE799DEA2902019A41A87632E99CAF0A" border="0" frame="box" pageNumber="9" rules="all">
<tr id="2145A92563B849955347FE43232DBA0F" pageNumber="9">
<td id="83351F1B76E3252F05CE74A8CF311A8D" colspan="2" pageNumber="9">
<taxonomicName id="3B3C1E56C9C3F82F1BC56D0D14070F0A" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="taiwanensis">Acanthopagrus taiwanensis</taxonomicName>
<taxonomicNameLabel id="573892B29D927F10837D04E2ABE833BF" rank="species">n. sp.</taxonomicNameLabel>
</td>
<td id="32F867F0419BE43A6FC0B55DF2BF7C69" colspan="2" pageNumber="9">
<taxonomicName id="04CAB75CB10CDEA73282E11652417FFA" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="berda">Acanthopagrus berda</taxonomicName>
</td>
</tr>
<tr id="E88C01AC9393FAE4628F4E5425495733" pageNumber="9">
<td id="1245C5EED9BE6F7E1546FEF36B7A8F1B" pageNumber="9">Holotype</td>
<td id="113706AA6E191DCB355416F4E622F061" pageNumber="9">Paratypes</td>
<td id="4C868E22B41AA0203F76C2AC77651C9F" pageNumber="9">Holotype</td>
<td id="225302B3CDC88662EE374DA0DCA4427F" pageNumber="9">Non-type specimens</td>
</tr>
<tr id="52621C5FE12A21FF9DBCDAF71D3469C9" pageNumber="9">
<td id="9D4A8C79D67B824F7090709E4650184F" pageNumber="9">MUFS 22854</td>
<td id="C1176876481F82507E70460A4D5150C1" pageNumber="9">MUFS 11870, 22165, 22854, 22855, 22857 n = 5</td>
<td id="3C8728DE9D5F08D09621879B312A2C68" pageNumber="9">ZMUC P50555 n = 1</td>
<td id="81BA8D6688E6DBF35B46FFA6B42B90BB" pageNumber="9">n = 39</td>
</tr>
<tr id="816BF72F6404A91EC93F56284702C1E0" pageNumber="9">
<td id="B7B00A201FB5224F6405D813FE09D312" pageNumber="9">Prepelvic length</td>
<td id="2339A66EC46E51CAA00EF744A9CD35E4" pageNumber="9">38</td>
<td id="D0198B43B9A685394A35F4216BE1E96F" pageNumber="9">35-40</td>
<td id="DF0C338C4E2C7125F8A861D560F20A60" pageNumber="9">-</td>
<td id="3247BDE53A3F6DD6CDD476303922C4EB" pageNumber="9">35-45 (40)</td>
</tr>
<tr id="28C9F0B21464EE9C165A5631D79FE9A1" pageNumber="9">
<td id="112FAB52F20A94778B07D02D528BA62D" pageNumber="9">Dorsal fin base</td>
<td id="3D10557EACE77D203FCE349DEC1A22F4" pageNumber="9">58</td>
<td id="DDD81DCB8625AF8188CA3E94DDFD0692" pageNumber="9">56-60</td>
<td id="142D18492047C6845E403CCF10122049" pageNumber="9">-</td>
<td id="DAD28E4E08D73B4F08A6EC75134CBA48" pageNumber="9">52-64 (58)</td>
</tr>
<tr id="B60F10634AE0EFE11DB0B670F9F354D9" pageNumber="9">
<td id="50A21B53C31DDB10175F4074E611CB76" pageNumber="9">Anal fin base</td>
<td id="6F31615E2EF5F6F50E1482A0673724C1" pageNumber="9">19</td>
<td id="E33B08922F3ED2AC5EA9D2CB007A8E5B" pageNumber="9">18-20</td>
<td id="66956F46B7B96888458A96746FB845A9" pageNumber="9">-</td>
<td id="886947BFA45F19A3B7D78C4D8F216E71" pageNumber="9">16-22 (19)</td>
</tr>
<tr id="14B21BEC6D66EFD00388DE7BEA021865" pageNumber="9">
<td id="9F488A54331486F5FF926E685DD9F48E" pageNumber="9">Caudal fin length</td>
<td id="1B65CD219A4E3ECCD5130B4360A57B97" pageNumber="9">26</td>
<td id="A8B45C4D32F095FEAA848424C5838607" pageNumber="9">28-30</td>
<td id="FEE4BBFC60EACFACB9EAC6D218869750" pageNumber="9">-</td>
<td id="20BBC7EC011A44589B107C29EADE5CD1" pageNumber="9">12-39 (30)</td>
</tr>
<tr id="3FB20D1A42F1FF32CAC0D330A83421B1" pageNumber="9">
<td id="F910027B6FDB88C4B8026BC1CE5A38A5" pageNumber="9">Pelvic fin spine</td>
<td id="558D31E399670011E06591E7A0369C93" pageNumber="9">17</td>
<td id="347AA20E6736E303C4403AD5ACED4920" pageNumber="9">15-18</td>
<td id="0B6612C97DAF2753BC017483C4600C1F" pageNumber="9">18</td>
<td id="234CDC44367484BFF77C38F431DED0E4" pageNumber="9">16-20 (17)</td>
</tr>
<tr id="BC4A8288A39E11B8303ED6175624A67A" pageNumber="9">
<td id="C0D04E0C16BA9367D7E75B54F99D01BF" pageNumber="9">First pelvic fin ray</td>
<td id="F15E7A70A48C221ACB82BDBBEFE4FC62" pageNumber="9">26 in right</td>
<td id="B1BECC625E970317D4E5A3D3E59291C7" pageNumber="9">26-29</td>
<td id="E0CA9F9E3B67D8CE83190A1371451ACE" pageNumber="9">20</td>
<td id="63ACBE17E69C8A7622BB30F9C0F2897F" pageNumber="9">17-31 (26)</td>
</tr>
<tr id="4DAD9144F1C38406DF6D85CD7C7D2597" pageNumber="9">
<td id="CD7F7E8D1EE82F99574276FBB68DCAFA" pageNumber="9">Longest pectoral fin ray</td>
<td id="49107FE6BDF1C9322934652304ACEB10" pageNumber="9">37</td>
<td id="50E0497F55BEEDE6479B21BC17A4D1FA" pageNumber="9">34-40</td>
<td id="53D88DA3120A876517A6AC626D27F0B3" pageNumber="9">41</td>
<td id="3CAB2FBCAF14F4F3A6DF1FCEF3B62D15" pageNumber="9">25-43 (39)</td>
</tr>
<tr id="61B57770DC0692B57FABFF66B9CE78BB" pageNumber="9">
<td id="706394F04D27006DE5D2567901572FD4" pageNumber="9">First dorsal fin spine</td>
<td id="3D19075EF6CCC89B840F5A1E505D00C9" pageNumber="9">7</td>
<td id="A0A4B6622A0E48F6D3977EB558015DD8" pageNumber="9">7</td>
<td id="CC26CFDA8C46FD59C6B4320FCABEB985" pageNumber="9">-</td>
<td id="BC92BAB7224A8E077F5BAB347AE30EFA" pageNumber="9">5-10 (8)</td>
</tr>
<tr id="B6A0AFF2E86F425A1AC580D65A5C7743" pageNumber="9">
<td id="F44FCAB3B055B8D4DB10F75F34B6CC6F" pageNumber="9">Second dorsal fin spine</td>
<td id="ECE3A0913D6B059337E537A93C729B9F" pageNumber="9">12</td>
<td id="E06BB2FDD93E982D9F19DF59C48EE6E7" pageNumber="9">11-12</td>
<td id="8FCF4AC9D89189E9EE629BDABF607826" pageNumber="9">-</td>
<td id="4A578EDFCA2E3562EEFEC101DBACC5B1" pageNumber="9">9-18 (14)</td>
</tr>
<tr id="B9C0E70F4885388D5C6B0C5658FEA9F2" pageNumber="9">
<td id="15D3FC0AA70542203F1E05F7EC4F6EDD" pageNumber="9">Third dorsal fin spine</td>
<td id="A87DE57EF4454B62A43DB02C4DE8EE51" pageNumber="9">15</td>
<td id="CB69575609E61F58FB6076974989C85D" pageNumber="9">15-16</td>
<td id="C828DF1CA33E505EA3558E0B69D7F063" pageNumber="9">-</td>
<td id="43D38285B3C9FD1F6EE73604980D1228" pageNumber="9">13-21 (18)</td>
</tr>
<tr id="1F345019765B05EE2AA78CC748EF8FBF" pageNumber="9">
<td id="F477B305EE12938F6CC1E7231004195D" pageNumber="9">Fourth dorsal fin spine</td>
<td id="9B579B11D218105437CADCD7B2132012" pageNumber="9">18</td>
<td id="5CD6C9B31D1CF4AC399D7A9F0FDFA461" pageNumber="9">16-19</td>
<td id="BF76D550B997080CAF2F4ADC823421C3" pageNumber="9">-</td>
<td id="14F9B39903D1F743367825EC80269806" pageNumber="9">14-22 (19)</td>
</tr>
<tr id="329FDC121AAE612D174FFB1EA8A7F397" pageNumber="9">
<td id="FCED53B63838BACECC550349884E55AA" pageNumber="9">Fifth dorsal fin spine</td>
<td id="AFF532C78AF9ABCBD4E00E11B473FD4D" pageNumber="9">17</td>
<td id="0D2A75BB490AC45218EADAC11486D258" pageNumber="9">16-19</td>
<td id="D7A62A52F5F0AD596805ADD872A9D345" pageNumber="9">-</td>
<td id="480D49CF143F344ADC53942DAC90E6ED" pageNumber="9">14-21 (18)</td>
</tr>
<tr id="60DAD150BBF6D30D2A8104C31DACD323" pageNumber="9">
<td id="4C47E953CEA9957D27C7D2371B8D593A" pageNumber="9">Sixth dorsal fin spine</td>
<td id="2CA72D55C26B7D3C0EE99CD69D8FB96B" pageNumber="9">16</td>
<td id="0850649F3C7CC4EA4D45365A68CF1E22" pageNumber="9">16-19</td>
<td id="BA013D2EF4BEA3C4E0D6A846595CB029" pageNumber="9">-</td>
<td id="0ED5D9C89AE772A5904BCDE8A1B154C0" pageNumber="9">15-20 (17)</td>
</tr>
<tr id="7AAF4429B680536A8615B2FF19705690" pageNumber="9">
<td id="0C321461BAA2BBBA17842807F4637A88" pageNumber="9">Last dorsal fin spine</td>
<td id="893BDE4FC6778439C0DD3D237AC2419C" pageNumber="9">12</td>
<td id="DCDC363FCB825D109F7E94B59BBD5868" pageNumber="9">11-14</td>
<td id="3FE25DAD12ACC8EAF2FEBE8F734175B6" pageNumber="9">-</td>
<td id="C4D6E934590CFF496E580CCE1C1328EB" pageNumber="9">11-16 (13)</td>
</tr>
<tr id="308BF091FD8057426213935A671D5954" pageNumber="9">
<td id="482699DAF8021BA5087643065108775A" pageNumber="9">Longest dorsal fin ray</td>
<td id="FDB610CE93C0E9777CB651594F1BA09A" pageNumber="9">15</td>
<td id="24423B5024FCF9E526335DF3CC5A37DF" pageNumber="9">14-17</td>
<td id="946CBBFF33B1F88CA0B006A06FEAB201" pageNumber="9">-</td>
<td id="EA8513FBD3F6472C6A94DB0805BB98A7" pageNumber="9">14-19 (16)</td>
</tr>
<tr id="4736F4C72529CFBBE22CE106227303F2" pageNumber="9">
<td id="BBA0134D30E0C5E15AF61A9EDBA00C04" pageNumber="9">First anal fin spine</td>
<td id="4A0A0D13A29AF8D253E8BEA22D2BF36F" pageNumber="9">7</td>
<td id="A40D30821135969217C414F0E5110B8D" pageNumber="9">6-9</td>
<td id="0F423960D32A8196506927552E6293B0" pageNumber="9">-</td>
<td id="197E5264968A99D3562AB81F147F1443" pageNumber="9">5-10 (8)</td>
</tr>
<tr id="0588ECF635C29A25442C2F43979DF6C5" pageNumber="9">
<td id="1955400C82206B241AC26F362B925BB1" pageNumber="9">Second anal fin spine</td>
<td id="36CD7F31FD200C5878D5C25CB9DF6F3D" pageNumber="9">21</td>
<td id="680F1AA8B97774D0B549522A8000BE0F" pageNumber="9">18-23</td>
<td id="483A99BDDCEA0C256168D8DB63014AC3" pageNumber="9">19</td>
<td id="71BCE0367BDF9CD27D527EDB91596564" pageNumber="9">15-26 (22)</td>
</tr>
<tr id="5B20EFDFC7BFE102E77314C88C0285C1" pageNumber="9">
<td id="4B6B774CC90B0AB3CFF61B1AEFDDA9A7" pageNumber="9">Third anal fin spine</td>
<td id="4EECBEAF1E49E846394548799857F803" pageNumber="9">15</td>
<td id="55A1C3D878A3866FCE877FA7F44EA9E2" pageNumber="9">15-17</td>
<td id="E417965FCF53708B99216619A6427A59" pageNumber="9">15</td>
<td id="FFCB44E282D3F0565E410C33C8C11E53" pageNumber="9">11-18 (16)</td>
</tr>
<tr id="401AEC7CE8D2EE0FAD6A5823701F853A" pageNumber="9">
<td id="2CACBB4DF9E451EFE8FF2465426EE38F" pageNumber="9">First anal fin ray</td>
<td id="5261F7321A010818D1DB42E83EB9C861" pageNumber="9">19</td>
<td id="B7D0599B34C407D85EE8D5DA0C85BEA5" pageNumber="9">14-18</td>
<td id="9A395188A66CFF57E9B7AF136C993B76" pageNumber="9">-</td>
<td id="9C04FCABAAA284A9F925A729A401DA3C" pageNumber="9">15-20 (17)</td>
</tr>
<tr id="CDC4E8A94A06ED4432F91D25FA5E27AC" pageNumber="9">
<td id="3ABEED82FF7B2CD73D920FE0A4757CBA" pageNumber="9">Suborbital depth</td>
<td id="E1EEB2218E395C899A8A81FCB407AA62" pageNumber="9">5</td>
<td id="4498AC07C1A9CB96C36F198BE5E37762" pageNumber="9">4</td>
<td id="F359AB220FF8BA5A116F2C53674B30C9" pageNumber="9">4</td>
<td id="C2BC42C6587C20B73914A9B7EB8CCE79" pageNumber="9">3-5 (4)</td>
</tr>
<tr id="9055A0C536D220E162FE9345AAE86C81" pageNumber="9">
<td id="C15AA3B3A3C807912AA1F4ED23BF1A65" pageNumber="9">2nd anal-fin spine / 3rd anal-fin spine</td>
<td id="E3CC414ADD04FA980C0D8DCFB6F84140" pageNumber="9">1.39</td>
<td id="1B178DEFCC76B6DC09B19ABB78B321D3" pageNumber="9">1.32-1.40</td>
<td id="413CF65F688AFEAF9328734ECE3F2B70" pageNumber="9">1.27</td>
<td id="E5817C83C61A9997F60BEAB3B2512CD4" pageNumber="9">1.23-1.50 (1.36)</td>
</tr>
</table>
</paragraph>
<footnote id="91A0085DD530066EE1BFA508B8809FD4">
<paragraph id="FE7F1E27017F40EBC755D26C29089103" pageNumber="9">*Standard length estimated from head length using the following regression parameters based on non-type measurements: y = 0.314x, y = head length, x = standard length, R2 = 0.9863.</paragraph>
</footnote>
<paragraph id="21688A8B2FA93B941E5C00AACDCDE428" lastPageNumber="10" pageNumber="9">
Body compressed. Mouth somewhat oblique. Maxillary reaching to below middle of pupil and larger than eye diameter. Upper jaw protruding slightly in front of lower jaw. Teeth in jaws in 3 to 5 crowded rows, about 6 (6 or 7) curved canines anteriorly in the upper jaw and 6 in the lower jaw. Suborbital depth greater than dermal eye opening. Anterodorsal profile from above eye gently curved. Profile of occipital ridge distinct above eye. Dorsal-fin spines strong, length of first slightly more than half length of second; second slightly shorter than third; fourth or fifth spine longest. Longest soft
<pageStartToken id="3AC6D604D040EF09D7C7F48949A3F6A6" pageNumber="10">dorsal-fin</pageStartToken>
ray shorter than longest dorsal-fin spine. First anal-fin spine short, much less than orbit diameter; second spine robust, its length clearly less than length of head without snout and slightly longer than snout; third anal-fin spine shorter than second spine. First anal-fin ray somewhat shorter than second and third anal-fin spines. Pectoral-fin tip not reaching to vertical through first anal-fin spine base, its length clearly greater than head length. Pelvic fin with first ray somewhat produced, its length shorter than head; pelvic-fin spine longer than snout.
</paragraph>
<paragraph id="4F699063019A1FBE87CFBBC812DBE0C3" pageNumber="10">Color (from fresh specimens)</paragraph>
<paragraph id="B549C575F9553B0F0FDA68B36A15D6D0" pageNumber="10">Head and body black (Fig. 1A). Both chin and belly markedly whitish in large specimens, especially over about 180 mm SL. Dorsal, caudal, anal, and pectoral fins black; ventral fins whitish. Color does not change much with preservation.</paragraph>
</subSubSection>
<subSubSection id="9CD02BD12EB2ECC3D1672863E6536B3A" type="distribution">
<paragraph id="BFAADEC6FEA64DE9B2FEA4A26F51B280" pageNumber="10">Distribution</paragraph>
<paragraph id="9E12B474C2B8CE1D8020E26BBF248A03" pageNumber="10">
<taxonomicName id="F3AE80FD939E38336D470E1BC41782C8" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="10" phylum="Chordata" rank="species" species="taiwanensis">Acanthopagrus taiwanensis</taxonomicName>
is currently known only from Tung-kang, southwestern Taiwan. According to local fishermen and buyers, the species is common around southwestern Taiwan. As the first author did not observe the species in extensive field sampling and examination of museum specimens from Korea, Japan, China (including around Hong Kong and Hainan Island), Vietnam, and elsewhere around Southeast Asia, it may be endemic to Taiwan.
</paragraph>
</subSubSection>
<subSubSection id="4CE1D7E630A07343C8D6E63C7BD43114" type="etymology">
<paragraph id="E5AE1C95C4BDE11A2223B5F620C8E1ED" pageNumber="10">Etymology</paragraph>
<paragraph id="3B19F2F782997B116F751026C2D4F787" pageNumber="10">The species name, “taiwanensis” reflects the type locality, Taiwan.</paragraph>
</subSubSection>
<subSubSection id="F80B614450290E43FC0DBC1AA4396D87" type="discussion">
<paragraph id="26BA33F1FB24382A272F84EBF3D1463B" pageNumber="10">Discussion</paragraph>
<paragraph id="CF9B74BBAF4004A43D38DC84BD84E1A6" pageNumber="10">
<taxonomicName id="6ED105DEA7244E55F3AB83432A067F79" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="10" phylum="Chordata" rank="species" species="taiwanensis">Acanthopagrus taiwanensis</taxonomicName>
and
<taxonomicName id="E921B26A512C426F514BA54866D86CF1" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="10" phylum="Chordata" rank="species" species="berda">A. berda</taxonomicName>
:
<taxonomicName id="E97044C460EBA8E40B371E5ECF8D9F92" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="10" phylum="Chordata" rank="species" species="taiwanensis">Acanthopagrus taiwanensis</taxonomicName>
and
<taxonomicName id="CFBB74418672D1EFA009D5913150BDCB" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="10" phylum="Chordata" rank="species" species="berda">A. berda</taxonomicName>
appear to be closely related and are perhaps sister species based on shared specializations. A cladistic analysis of species within this genus is premature as confusion remains about the status of some species and the relation of this genus to
<taxonomicName id="9A9FBAECC4574A13B9FB616441B4D681" LSID-ZBK="urn:lsid:zoobank.org:act:499A51BC-ABBD-4D81-8990-D60A67430880" class="Actinopterygii" family="Sparidae" genus="Sparidentex" kingdom="Animalia" lsidName-ZBK="Sparidentex Munro 1948:276" order="Perciformes" pageNumber="10" phylum="Chordata" rank="genus">Sparidentex</taxonomicName>
(Orrell &amp; Carpenter, 2004). The combination of body shape, fin coloration and typical number of scale rows above the lateral line is unique for these species within
<taxonomicName id="745764709D87B56AA10B65A6A1C14874" LSID-ZBK="urn:lsid:zoobank.org:act:8F9E30E9-3A84-43EA-8E93-F852D3E82151" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" lsidName-ZBK="Acanthopagrus Chrysophrys Peters 1855:242" order="Perciformes" pageNumber="10" phylum="Chordata" rank="genus">Acanthopagrus</taxonomicName>
, supporting a close relationship. In addition, the shape of the ventral edge of the first 2 infraorbitals is concave in larger individuals of both species, while it is straight in all other described species of
<taxonomicName id="CC4C672193D8736CC62C5BF8064B9A92" LSID-ZBK="urn:lsid:zoobank.org:act:8F9E30E9-3A84-43EA-8E93-F852D3E82151" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" lsidName-ZBK="Acanthopagrus Chrysophrys Peters 1855:242" order="Perciformes" pageNumber="10" phylum="Chordata" rank="genus">Acanthopagrus</taxonomicName>
. The concavity is much more pronounced in
<taxonomicName id="2620D82EB05FEC06B06EB7E95E8B4629" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="10" phylum="Chordata" rank="species" species="berda">A. berda</taxonomicName>
than
<taxonomicName id="073F0BF30E8AA728D53B97BE11FAB3FF" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="10" phylum="Chordata" rank="species" species="taiwanensis">A. taiwanensis</taxonomicName>
and represents a clear difference between the 2 species.
</paragraph>
<paragraph id="ABF6B374A3018FA8248672DBAE70ED4A" lastPageNumber="11" pageNumber="10">
Synonyms of
<taxonomicName id="6D0B87F8B7450291520B76D49F8EC193" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="10" phylum="Chordata" rank="species" species="berda">Acanthopagrus berda</taxonomicName>
:
<taxonomicName id="FEDC55C7D3198BD7248E841DC8616276" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="10" phylum="Chordata" rank="species" species="berda">
Acanthopagrus berda (
<normalizedToken id="F30077100CAD7165541EFB5E1852B64D" originalValue="Forsskål">Forsskal</normalizedToken>
, 1775)
</taxonomicName>
was described on the basis of a dried skin of the holotype (see Table 1; Fig. 2A) from Luhaiya, Yemen, Red Sea. Despite the condition of the holotype, the 4 most important diagnostic characters of this species are evident: a strong concavity of ventral edge of first 2
<pageStartToken id="256C0FF3F8896D9EC05DCC737453E2B5" pageNumber="11">infraorbitals</pageStartToken>
above posterior part of upper jaw (Figs. 1B, 2A and 3B); 3 ½ scale rows between fifth spinous dorsal-fin spine base and lateral line; anal, ventral, and caudal fins uniformly blackish; and second anal-fin spine longer than third spine. We have examined numerous specimens from throughout the Indo-Pacific, including the Red Sea, that correspond to these key characters. The concavity of the ventral edge of the first 2 infraorbitals is observed in specimens larger than around 130-150 mm SL. The holotype of
<taxonomicName id="FE1E9CA6B2463A3507CC78028D59D002" LSID-ZBK="urn:lsid:zoobank.org:act:F406CB69-0B66-4AB0-A5D2-C433112AEB20" class="Actinopterygii" family="Sparidae" genus="Sparus" kingdom="Animalia" lsidName-ZBK="Sparus hasta Bloch &amp; Schneider 1801:275" order="Perciformes" pageNumber="10" phylum="Chordata" rank="species" species="hasta">Sparus hasta Bloch and Schneider</taxonomicName>
(1801) has the same diagnostic characters as
<taxonomicName id="B048A2D0B6FDCC70C918F4C47D10A692" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="10" phylum="Chordata" rank="species" species="berda">A. berda</taxonomicName>
. Bauchot and Skelton (1986) determined this species to be a junior synonym of
<taxonomicName id="E00596F28A8763CC1772A96941D88700" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="10" phylum="Chordata" rank="species" species="berda">A. berda</taxonomicName>
and we concur.
</paragraph>
<caption id="578D47056079CC06EAE1EC207307FE43">
<paragraph id="B4EB5B3841E8738B95A4B8C7C670F6AC" pageNumber="11">
FIGURE 4. Anteriormost head squamation type of A,
<taxonomicName id="3941BDCA68EF3744878C8EBD402426C7" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="11" phylum="Chordata" rank="species" species="taiwanensis">Acanthopagrus taiwanensis</taxonomicName>
<taxonomicNameLabel id="2287FB12CAF804D0E861A75825873F27" rank="species">n. sp.</taxonomicNameLabel>
, MUFS 22166, paratype, 212 mm SL, Tung-kang, southwestern Taiwan; and B,
<taxonomicName id="38CEEAA60ACFB41BD24DF5FD0C9A580C" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="11" phylum="Chordata" rank="species" species="berda">
A. berda (
<normalizedToken id="780B32A94B6EB9573544A2E2A0955FF3" originalValue="Forrskål">Forrskal</normalizedToken>
, 1775)
</taxonomicName>
, ZRC 38094, 249 mm SL, Kisap River, Langkawi Island, Malaysia.
</paragraph>
</caption>
<caption id="FF6876369C9A9067AC72F16DF4BC8C0B">
<paragraph id="AF113FFBFA39029ACEB3C33A7E7B0B8D" pageNumber="12">
<pageStartToken id="1D8BF9EC1774A7D9C30A28C90FCA7EF1" pageNumber="12">FIGURE</pageStartToken>
5. Molar teeth of upper and lower jaws of two sparid species: A -B,
<taxonomicName id="CD8D83206E9544332218B0C56ACEF416" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="12" phylum="Chordata" rank="species" species="taiwanensis">Acanthopagrus taiwanensis</taxonomicName>
<taxonomicNameLabel id="CD6234D24EB4E2CF6DA166FD47C5683C" rank="species">n. sp.</taxonomicNameLabel>
, MUFS 22166, paratype, 212 mm SL, Tung-kang, southwestern Taiwan and C -D,
<taxonomicName id="28C36785E01BCF41A70BB3F6E63010F3" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="12" phylum="Chordata" rank="species" species="berda">
A. berda (
<normalizedToken id="D642C94C5878CE40CDEB8E288244ECE0" originalValue="Forrskål">Forrskal</normalizedToken>
, 1775)
</taxonomicName>
, SAIAB, 203 mm SL, South Africa.
</paragraph>
</caption>
<paragraph id="09331635D7D236C74F8BADD8479E5D6E" pageNumber="12">
<taxonomicName id="111803DAA739CC95B07BB49E9A3FCA64" LSID-ZBK="urn:lsid:zoobank.org:act:135BF887-0394-4434-90B2-1CB9FE6603B8" class="Actinopterygii" family="Sparidae" genus="Sparus" kingdom="Animalia" lsidName-ZBK="Sparus calamara Cuvier 1829:182" order="Perciformes" pageNumber="12" phylum="Chordata" rank="species" species="calamara">Sparus calamara Cuvier, 1829</taxonomicName>
(based on a drawing of Russell, 1803: pl. 92) is clearly a junior synonym of
<taxonomicName id="2F917115D2457D10F9297989CEC39059" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="12" phylum="Chordata" rank="species" species="berda">A. berda</taxonomicName>
, with the following diagnostic characters evident on the drawing: a relatively deep body (52 % of SL, see Table 1); 3 ½ scale rows between the fifth spinous dorsal-fin spine base and lateral line; ventral edge of the first 2 infraorbitals above the posterior part of the upper jaw weakly concave; and black ventral, anal, and caudal fins (Table 1, Figs. 1i -5).
<taxonomicName id="0DD82AD954565F92AE71EAE8AAE7BB0D" LSID-ZBK="urn:lsid:zoobank.org:act:26DDEF88-4F61-46BD-9A14-6598DA40733F" family="Sparidae" genus="Chrysophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Chrysophrys calamara Valenciennes 1830:117" order="Perciformes" pageNumber="12" phylum="Chordata" rank="species" species="calamara">Chrysophrys calamara Valenciennes in Cuvier &amp; Valenciennes, 1830</taxonomicName>
(preoccupied by Cuvier, 1829), based on 3 syntypes has long been identified as a junior synonym of
<taxonomicName id="BAB4D72C52919A105F4136BD474A4CAE" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="12" phylum="Chordata" rank="species" species="berda">A. berda</taxonomicName>
by subsequent researchers (Bauchot &amp; Daget, 1972; Bauchot &amp; Skelton, 1986). Day (1875) also described “
<taxonomicName id="59F7D7491BAE0B33102F2AC82B7712F7" LSID-ZBK="urn:lsid:zoobank.org:act:9D0B43D0-C3E7-4711-B7F5-47363CF76137" family="Sparidae" genus="Chrysophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Chrysophrys berda calamara Day 1876:140" order="Perciformes" pageNumber="12" phylum="Chordata" rank="variety" species="berda" variety="calamara">Chrysophrys berda var. calamara</taxonomicName>
” on the basis of a ZSI F1785s specimen. However, since the name “
<taxonomicName id="EE8531E1E67ADAC60E27FD44BD304B36" LSID-ZBK="urn:lsid:zoobank.org:act:135BF887-0394-4434-90B2-1CB9FE6603B8" class="Actinopterygii" family="Sparidae" genus="Sparus" kingdom="Animalia" lsidName-ZBK="Sparus calamara Cuvier 1829:182" order="Perciformes" pageNumber="12" phylum="Chordata" rank="species" species="calamara">calamara</taxonomicName>
”, is preoccupied by Cuvier, 1829, Days “
<taxonomicName id="E7D5D687882D5192F109EF3A5C02963B" LSID-ZBK="urn:lsid:zoobank.org:act:9D0B43D0-C3E7-4711-B7F5-47363CF76137" family="Sparidae" genus="Chrysophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Chrysophrys berda calamara Day 1876:140" order="Perciformes" pageNumber="12" phylum="Chordata" rank="variety" species="berda" variety="calamara">calamara</taxonomicName>
” has no taxonomic significance in the context of
<taxonomicName id="5BF9CB3B9B78999DD75C7988C217FD24" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="12" phylum="Chordata" rank="species" species="berda">A. berda</taxonomicName>
.
</paragraph>
<paragraph id="17321518D16047C623F8F578834C0811" lastPageNumber="13" pageNumber="12">
Castelnau (1861) described
<taxonomicName id="E9FB6D2A64D2CFD046BEF210C39427BC" LSID-ZBK="urn:lsid:zoobank.org:act:2FBDAE0C-8BA2-4EC3-98FD-0CD596EBC622" class="Actinopterygii" family="Sparidae" genus="Pagrus" kingdom="Animalia" lsidName-ZBK="Pagrus caffer Castelnau 1861:30" order="Perciformes" pageNumber="12" phylum="Chordata" rank="species" species="caffer">Pagrus caffer</taxonomicName>
(type probably lost, P. Pruvost, pers. comm.) from Durban (Port Natal), South Africa and Smith &amp; Smith (1986) later considered it to be a synonym of
<taxonomicName id="DEA31265EA0743C7594CA6F77D94F2F3" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="12" phylum="Chordata" rank="species" species="berda">A. berda</taxonomicName>
; we tentatively concur. We examined the holotype of Gilchrist &amp; Thompsons (1908)
<taxonomicName id="3F6349C2F2A5FFCD98BB3605F289F8F6" LSID-ZBK="urn:lsid:zoobank.org:act:AC70590A-A323-436D-A22F-88789CB706D3" family="Sparidae" genus="Chrysophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Chrysophrys robinsoni Gilchrist &amp; Thompson 1908:170" order="Perciformes" pageNumber="12" phylum="Chordata" rank="species" species="robinsoni">Chrysophrys robinsoni</taxonomicName>
and its characters are consistent with
<taxonomicName id="95FF5D7E3FA5234E055DAC711C6E29BC" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="12" phylum="Chordata" rank="species" species="berda">A. berda</taxonomicName>
, including the broadly rounded rostral-most head squamation with small scales anteriorly
<pageStartToken id="0DE52C8179FFF893B182F4AD3B6C1519" pageNumber="13">(</pageStartToken>
as in Fig. 4B).
</paragraph>
<paragraph id="E1D49AE0AB15ABBA883F29DED78C81CB" pageNumber="13">
Comments on other nominal species assigned to
<taxonomicName id="BE047491EDECF2159538FDF6CFA51D66" LSID-ZBK="urn:lsid:zoobank.org:act:8F9E30E9-3A84-43EA-8E93-F852D3E82151" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" lsidName-ZBK="Acanthopagrus Chrysophrys Peters 1855:242" order="Perciformes" pageNumber="13" phylum="Chordata" rank="genus">Acanthopagrus</taxonomicName>
: No type specimen is known for
<taxonomicName id="3FD89332121B82ED1671237D4E9B4629" LSID-ZBK="urn:lsid:zoobank.org:act:B2DA4958-A70A-42BB-AECA-254BD654C7C6" family="Sparidae" genus="Coius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Coius datnia Hamilton 1822:88, 369" order="Perciformes" pageNumber="13" phylum="Chordata" rank="species" species="datnia">Coius datnia</taxonomicName>
, described by Hamilton (1822) from the Ganges River mouth, India, but a fine plate (fig. 29 of pl. 9) accompanied the description. This species has long been considered a junior synonym of
<taxonomicName id="FC075806298A82A0E124EBF1BFD62FB3" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="13" phylum="Chordata" rank="species" species="latus">A. latus (Houttuyn, 1782)</taxonomicName>
because of its yellow pelvic and anal fins (Kottelat, 1986, 2000). However, the absence of molar teeth given in the original description and plate strongly support its placement in the genus
<taxonomicName id="B45BCB0739F34134E2938BD31C09E3A3" LSID-ZBK="urn:lsid:zoobank.org:act:499A51BC-ABBD-4D81-8990-D60A67430880" class="Actinopterygii" family="Sparidae" genus="Sparidentex" kingdom="Animalia" lsidName-ZBK="Sparidentex Munro 1948:276" order="Perciformes" pageNumber="13" phylum="Chordata" rank="genus">Sparidentex</taxonomicName>
(Bauchot &amp; Smith, 1983) and clearly distinguish it from
<taxonomicName id="A5E5F0268B1CED39C5E9B5552732DF11" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="13" phylum="Chordata" rank="species" species="berda">A. berda</taxonomicName>
and
<taxonomicName id="18BB23A64AE71C2508157AAC78E34DF5" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="13" phylum="Chordata" rank="species" species="taiwanensis">A. taiwanensis</taxonomicName>
. Hamiltons species needs further study.
</paragraph>
<paragraph id="9A3C8EA68017490002B0A90667CF256C" pageNumber="13">
Gilchrist &amp; Thompson (1908) described
<taxonomicName id="8AD204C484E0AA83B6E70D21319B9DE4" LSID-ZBK="urn:lsid:zoobank.org:act:ACD72605-1F75-48B7-8AF1-5B269FB4DEEE" family="Sparidae" genus="Chrysophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Chrysophrys estuarius Gilchrist &amp; Thompson 1908:170" order="Perciformes" pageNumber="13" phylum="Chordata" rank="species" species="estuarius">Chrysophrys estuarius</taxonomicName>
from South Africa. The three syntypes have a relatively slender body and a straight ventral edge of the first 2 infraorbitals above the posterior upper jaw, characters that distinguish it from
<taxonomicName id="E840DBEE823B019161615290F7F5CE2D" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="13" phylum="Chordata" rank="species" species="berda">A. berda</taxonomicName>
and
<taxonomicName id="7D6C8F4DA4F8411D20336E956BE66867" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="13" phylum="Chordata" rank="species" species="taiwanensis">A. taiwanensis</taxonomicName>
.
</paragraph>
<paragraph id="E0B13E5C059F306A33D0C7AE4074D9DE" pageNumber="13">
<taxonomicName id="E9B659207C0968B6A4048ABBBD824EC3" LSID-ZBK="urn:lsid:zoobank.org:act:AF2146E3-8A2B-470F-A4F6-041F4B7AE21F" family="Sparidae" genus="Chrysophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Chrysophrys schlegelii Bleeker 1854:400" order="Perciformes" pageNumber="13" phylum="Chordata" rank="species" species="schlegelii">Chrysophrys schlegelii Bleeker, 1854</taxonomicName>
and
<taxonomicName id="87C60A83D7C74F4751F0DECB0BF35ACA" LSID-ZBK="urn:lsid:zoobank.org:act:F536E6AC-3C2B-451A-9021-0543E15B0D5F" family="Sparidae" genus="Mylio" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Mylio butcheri Munro 1949:191" order="Perciformes" pageNumber="13" phylum="Chordata" rank="species" species="butcheri">Mylio butcheri Munro, 1949</taxonomicName>
are valid species in the genus
<taxonomicName id="21B131D63644ECDEC004E894141347B0" LSID-ZBK="urn:lsid:zoobank.org:act:8F9E30E9-3A84-43EA-8E93-F852D3E82151" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" lsidName-ZBK="Acanthopagrus Chrysophrys Peters 1855:242" order="Perciformes" pageNumber="13" phylum="Chordata" rank="genus">Acanthopagrus</taxonomicName>
from the East Asian Shelf and southern Australia, respectively (Akazaki, 1984; Allen et al., 2002; Cadwallader and Backhouse, 1983; Gomon et al., 1994). They are clearly distinct from both
<taxonomicName id="E828FC726FB4C3891907CE8307444AA0" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="13" phylum="Chordata" rank="species" species="berda">A. berda</taxonomicName>
and
<taxonomicName id="C3A77082D3C003DE3F212E77EDEE217C" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="13" phylum="Chordata" rank="species" species="taiwanensis">A. taiwanensis</taxonomicName>
with slender bodies and 4 ½-5 ½ scale rows between the fifth dorsal-fin spine base and lateral line (vs. 3 ½ for
<taxonomicName id="C6DC79EFAEF7C296EBE0B623D2B4D913" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="13" phylum="Chordata" rank="species" species="berda">A. berda</taxonomicName>
and
<taxonomicName id="4E974A71520B0F922A4CCF0C63ECBD2B" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="13" phylum="Chordata" rank="species" species="taiwanensis">A. taiwanensis</taxonomicName>
). Akazaki (1962) considered
<taxonomicName id="38391DC193C20F3BB3F98F31245EB380" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="13" phylum="Chordata" rank="subSpecies" species="swinhonis" subSpecies="czerskii">Acanthopagrus swinhonis czerskii Berg, 1914</taxonomicName>
as a junior synonym of
<taxonomicName id="D7ABB2F006BF774FFA205F53843F829A" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="13" phylum="Chordata" rank="species" species="schlegelii">A. schlegelii</taxonomicName>
. We did not see the type specimens but followed his idea based on the detailed figure included in the description.
</paragraph>
<paragraph id="95D90BE95BFB100DE3BA07E87B7DC58D" pageNumber="13">
<taxonomicName id="60AB440BAB121AC5899C15A57E418D9E" LSID-ZBK="urn:lsid:zoobank.org:act:9E3FE3F2-D878-4801-826C-29371D028E4D" family="Sparidae" genus="Chrysophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Chrysophrys australis Günther 1859:494" order="Perciformes" pageNumber="13" phylum="Chordata" rank="species" species="australis">
Chrysophrys australis
<normalizedToken id="558D8EEE994CBECE79A2C6BDE4BFF2CA" originalValue="Günther">Guenther</normalizedToken>
, 1859
</taxonomicName>
,
<taxonomicName id="4F6CB04E9869DDDF49D0F1B51F46142E" LSID-ZBK="urn:lsid:zoobank.org:act:14D5083D-963A-415C-A8A1-FABBE8DF68C4" family="Sparidae" genus="Chrysophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Chrysophrys swinhonis Günther 1874:155" order="Perciformes" pageNumber="13" phylum="Chordata" rank="species" species="swinhonis">
C. swinhonis
<normalizedToken id="414B7E16B638F0CC848096E24F954957" originalValue="Günther">Guenther</normalizedToken>
, 1874
</taxonomicName>
,
<taxonomicName id="86443D8DB5E9CDA736910D9020728579" LSID-ZBK="urn:lsid:zoobank.org:act:AEEF3B1B-CD90-4A88-93C7-FE85B92A181E" family="Sparidae" genus="Chrysophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Chrysophrys rubroptera Tirant 1883:[14]" order="Perciformes" pageNumber="13" phylum="Chordata" rank="species" species="rubroptera">C. rubroptera Tirant, 1883</taxonomicName>
,
<taxonomicName id="9023ECAE6C2C7844A89DFDC4F4872D82" LSID-ZBK="urn:lsid:zoobank.org:act:1CAA8327-86CC-4EDD-B157-98E13DFCAB41" class="Actinopterygii" family="Sparidae" genus="Petrus" kingdom="Animalia" lsidName-ZBK="Petrus belayewi Hora &amp; Misra 1943:12" order="Perciformes" pageNumber="13" phylum="Chordata" rank="species" species="belayewi">Petrus belayewi Hora &amp; Misra, 1943</taxonomicName>
,
<taxonomicName id="9D79C89F970B5E260C2C97A5C585E215" LSID-ZBK="urn:lsid:zoobank.org:act:3F6383DE-915F-443E-96F8-64B900E0121F" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" lsidName-ZBK="Acanthopagrus sivicolus Akazaki 1962:187, 355" order="Perciformes" pageNumber="13" phylum="Chordata" rank="species" species="sivicolus">Acanthopagrus sivicolus Akazaki, 1962</taxonomicName>
, and
<taxonomicName id="6032A24AAA8F9EC625F3B9904C84FE6B" LSID-ZBK="urn:lsid:zoobank.org:act:F674D9D5-E140-40F4-8081-2AD2E086F838" family="Sparidae" genus="Chrysophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Chrysophrys novaecaledoniae Castelnau 1873:110" order="Perciformes" pageNumber="13" phylum="Chordata" rank="species" species="novaecaledoniae">Chrysophrys novaecaledoniae Castelnau, 1873</taxonomicName>
show 4 ½ scale rows between the fifth dorsal-fin spine base and lateral line (rarely 3 ½ in
<taxonomicName id="BDB8D2224B2941483E14F5C24CB0CD16" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="13" phylum="Chordata" rank="species" species="australis">A. australis</taxonomicName>
and yellowish pelvic- and anal-fin rays, as described in Carpenter, 2001, and 2 of 4 syntypes of
<taxonomicName id="118FDC056ED77892DF5FA8B51935B3A6" LSID-ZBK="urn:lsid:zoobank.org:act:14D5083D-963A-415C-A8A1-FABBE8DF68C4" family="Sparidae" genus="Chrysophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Chrysophrys swinhonis Günther 1874:155" order="Perciformes" pageNumber="13" phylum="Chordata" rank="species" species="swinhonis">C. swinhonis</taxonomicName>
with 6 ½, as discussed below) on the basis of their type specimens. But
<taxonomicName id="9B8214415E1756BE2E9B9116AF6BD637" LSID-ZBK="urn:lsid:zoobank.org:act:AEEF3B1B-CD90-4A88-93C7-FE85B92A181E" family="Sparidae" genus="Chrysophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Chrysophrys rubroptera Tirant 1883:[14]" order="Perciformes" pageNumber="13" phylum="Chordata" rank="species" species="rubroptera">C. rubroptera</taxonomicName>
and
<taxonomicName id="0244D6C60D9890FFE2D09A65FE50F3DC" LSID-ZBK="urn:lsid:zoobank.org:act:3F6383DE-915F-443E-96F8-64B900E0121F" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" lsidName-ZBK="Acanthopagrus sivicolus Akazaki 1962:187, 355" order="Perciformes" pageNumber="13" phylum="Chordata" rank="species" species="sivicolus">A. sivicolus</taxonomicName>
were confirmed by a fine photograph (Kottelat, 1986) and nontype specimens examined, respectively. In addition, these species have a straight ventral edge of the first 2 infraorbitals above the posterior part of the upper jaw, in contrast to strongly or weakly concave in
<taxonomicName id="13A3217189B034DE279BCB36343E7CF8" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="13" phylum="Chordata" rank="species" species="berda">A. berda</taxonomicName>
and
<taxonomicName id="2B39A2826C986FDDF27C88172BE8246F" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="13" phylum="Chordata" rank="species" species="taiwanensis">A. taiwanensis</taxonomicName>
.
</paragraph>
<paragraph id="E89F8C359869CA8594309C1D519A6CC4" pageNumber="13">
The taxonomic status of
<taxonomicName id="1E2BBE7CADE177D08F0FB80C99C0F219" LSID-ZBK="urn:lsid:zoobank.org:act:14D5083D-963A-415C-A8A1-FABBE8DF68C4" family="Sparidae" genus="Chrysophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Chrysophrys swinhonis Günther 1874:155" order="Perciformes" pageNumber="13" phylum="Chordata" rank="species" species="swinhonis">C. swinhonis</taxonomicName>
has not been examined in detail since the original description by
<normalizedToken id="212717E0F8E699FCFB67592E61AB720F" originalValue="Günther">Guenther</normalizedToken>
(1859). In 4 syntypes (87-279 mm SL), the pored lateralline scales range from 51 to 56 (vs. 42 to 44 in
<taxonomicName id="E49BFE87CBD39DBD02E148CA539FD291" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="13" phylum="Chordata" rank="species" species="berda">A. berda</taxonomicName>
and 43 or 44 in
<taxonomicName id="878B725782870CD83BEC2538A725A87A" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="13" phylum="Chordata" rank="species" species="taiwanensis">A. taiwanensis</taxonomicName>
), and scale rows between the fifth dorsal-fin spine base and lateral line are 4 ½ or 6 ½. Although a detailed review of
<taxonomicName id="3FC33949A6347922EE25A5A5569EFF2A" LSID-ZBK="urn:lsid:zoobank.org:act:14D5083D-963A-415C-A8A1-FABBE8DF68C4" family="Sparidae" genus="Chrysophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Chrysophrys swinhonis Günther 1874:155" order="Perciformes" pageNumber="13" phylum="Chordata" rank="species" species="swinhonis">C. swinhonis</taxonomicName>
is still required, it is clearly distinct from both
<taxonomicName id="CA29F6D3EAFC60DC88C58974231B61F4" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="13" phylum="Chordata" rank="species" species="berda">A. berda</taxonomicName>
and
<taxonomicName id="C9863F7F00452D7DD41A643F7F7B6878" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="13" phylum="Chordata" rank="species" species="taiwanensis">A. taiwanensis</taxonomicName>
.
</paragraph>
<paragraph id="BCE3F9A26CDBADDAA6FBB78ABBCD46DD" pageNumber="13">
The holotype of
<taxonomicName id="76609C3AEFD1ACAF29420B0909F022F0" LSID-ZBK="urn:lsid:zoobank.org:act:CAC1ADE9-A5D0-421A-85FE-94AE8C01972E" family="Sparidae" genus="Roughleyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Roughleyia palmaris Whitley 1935:234" order="Perciformes" pageNumber="13" phylum="Chordata" rank="species" species="palmaris">Roughleyia palmaris Whitley, 1935</taxonomicName>
has 4 scale rows between the fifth dorsal-fin spine base and lateral line, and the upper head profile gibbous (not gibbous in
<taxonomicName id="3B901CEB1F2B8497EC50F495347E2F29" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="13" phylum="Chordata" rank="species" species="berda">A. berda</taxonomicName>
and
<taxonomicName id="F85D9E273D42D25B628E40C320FCCC8C" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="13" phylum="Chordata" rank="species" species="taiwanensis">A. taiwanensis</taxonomicName>
). Hutchins (2001) and Allen et al. (2002) regard it as a valid species of
<taxonomicName id="2D6C2E707DFAA6AD7EF4AD1E7644347E" LSID-ZBK="urn:lsid:zoobank.org:act:8F9E30E9-3A84-43EA-8E93-F852D3E82151" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" lsidName-ZBK="Acanthopagrus Chrysophrys Peters 1855:242" order="Perciformes" pageNumber="13" phylum="Chordata" rank="genus">Acanthopagrus</taxonomicName>
confined to southwestern Australia.
</paragraph>
<caption id="830829CA34B1D8478595E503003E6B6A">
<paragraph id="298C0E4BA3F2576B562FB226A9D87D5B" pageNumber="14">
<pageStartToken id="705AD4C967F7A3C4B9C191B426424570" pageNumber="14">TABLE</pageStartToken>
2. Comparison of selected characters of
<taxonomicName id="3C4E1EB7B4E3C5B87C1C1F22A26C4137" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="14" phylum="Chordata" rank="species" species="taiwanensis">Acanthopagrus taiwanensis</taxonomicName>
<taxonomicNameLabel id="46B4CA23E1C36C0A5A86059817D230EE" rank="species">n. sp.</taxonomicNameLabel>
and
<taxonomicName id="4DCDBCCFF95B0EF510546902197B85A3" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="14" phylum="Chordata" rank="species" species="berda">
A. berda (
<normalizedToken id="9FC9F77D49A9399B0A91F7CA344DDE3F" originalValue="Forsskål">Forsskal</normalizedToken>
, 1775)
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="A953424F2B385AE18BA4162E3B99146D">
<table id="7095035B7933BEA947FCD4411AF49A6D" border="0" frame="box" pageNumber="14" rules="all">
<tr id="25C9105673389077C4BE9F8B06492AC2" pageNumber="14">
<td id="324B8C6A01F50874017CE151213E0AD0" pageNumber="14">
<taxonomicName id="6F2C232B978923D435EAFA7DF6CD6150" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="taiwanensis">Acanthopagrus taiwanensis</taxonomicName>
<taxonomicNameLabel id="6C4C31B924A089ABAC984E2D5A2C61B3" rank="species">n. sp.</taxonomicNameLabel>
110-212 mm SL (n = 3)
</td>
<td id="F0FAAEFBDB617D1503B84BE889F74D65" pageNumber="14">
<taxonomicName id="1F15E404B912FDD25CD9AA6CAE978BF3" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" phylum="Chordata" rank="species" species="berda">Acanthopagrus berda</taxonomicName>
33-288 mm SL (n = 39)
</td>
</tr>
<tr id="E1AC459E0F8A783D44B544320CFB5B17" pageNumber="14">
<td id="9031FFF08766A5A45BFA46858685D7A5" pageNumber="14">dorsal-fin rays</td>
<td id="EF20055753F87E130A19A4473F466061" pageNumber="14">XI-XII, 10-12</td>
<td id="F730B150920B5C70C31DFD55837993B3" pageNumber="14">usually XI, 11 (XII, 10 or XI, 12)</td>
</tr>
<tr id="0C0D50980A5AB60AA97C6B9645476947" pageNumber="14">
<td id="8B911F63D0D5068094FBE6A3CBC72884" pageNumber="14">anal-fin rays</td>
<td id="DE7B8EC179A9D11AC7862F1B6C4A3CF7" pageNumber="14">III, 8-9</td>
<td id="73BAE21DDD09D878D918729C66E52996" pageNumber="14">usually III, 9 (rarely 10)</td>
</tr>
<tr id="4F2C75E8A2BEEDACA3A119593F441BED" pageNumber="14">
<td id="14E940620E77F7B937F26E5B2ABDE029" pageNumber="14">gill rakers</td>
<td id="BA135D91C5E3147324AD13606CA83712" pageNumber="14">4-5+9-11 = 13-15</td>
<td id="4425B74C8989F3C598377166B9FECEDA" pageNumber="14">5-7 (usually 5 or 6) + 8-12 (usually 9 or 10) = 13-17 (usually 14 or 15)</td>
</tr>
<tr id="3FC81A7B1965CE5626B196D0FD365FE1" pageNumber="14">
<td id="D82F4A6CBE50989D3CF33435136FAE4A" pageNumber="14">pored lateral-line scales</td>
<td id="B7F4E35A3123534C898CA8D65BD37187" pageNumber="14">43 or 44</td>
<td id="1C771D0BD8A19A27E4C44FB0324462BA" pageNumber="14">42-44</td>
</tr>
<tr id="C4C8BE9D9726D23F80A77E36AE52F305" pageNumber="14">
<td id="A50CD46EA11A988C6A0CE5B13F6A6AB4" pageNumber="14">cheek scale rows</td>
<td id="876CF92E152B51F1A3AA4844FF4F7255" pageNumber="14">3 or 4</td>
<td id="038B43154F40FD9E6424B28BD3DC46A9" pageNumber="14">5 or 6</td>
</tr>
<tr id="757526372F3EAADD4029AAEDB8FC1886" pageNumber="14">
<td id="DD1ADD02CC883338498A0785CBF65992" pageNumber="14">upper and lower lips</td>
<td id="CF5027DF4F461A156C5C5BB6F0F2F6E6" pageNumber="14">thin</td>
<td id="E15074F5E86EB29F93192E612BC6C295" pageNumber="14">thick</td>
</tr>
<tr id="37BCC1122197ED2A5ACA69BE05088D1B" pageNumber="14">
<td id="222CC2B3FD856CE66034B5DB945EAFA6" pageNumber="14">ventral edge of the first two infraorbitals above posterior part of upper jaw</td>
<td id="EC37303CA287C70A6F03AEC0091F084D" pageNumber="14">almost straight but weakly concave in specimens of 210 mm SL</td>
<td id="4D073244878A0C4F1C00272FCA27A927" pageNumber="14">strongly concave with growth, in specimens over 130-150 mm SL</td>
</tr>
<tr id="1E7053125CFEFDE3934AE91685FBBB44" pageNumber="14">
<td id="44867C382A758EADA38EA497CC2F568E" pageNumber="14">anteriormost head scallation from top view</td>
<td id="EEBCD3515A0A92054F4CFEBB090C4F8A" pageNumber="14">rounded with ca. 10 somewhat smaller scales anteriorly</td>
<td id="9B8986E8AA8BB8C66D14F0CDFC803C01" pageNumber="14">rounded with over 20 distinct small scales anteriorly</td>
</tr>
<tr id="8786DC7E416C3DC51FD2E94C054A2F8D" pageNumber="14">
<td id="712110B75573E2B530195F8FD972F101" pageNumber="14">ridge just before or above eye</td>
<td id="D9C1DAAB2B321E5E8F71CB0884FFA9C3" pageNumber="14">obvious ridge just before eye in specimen over ca. 180 mm SL</td>
<td id="2D80A3565C429D4D8EAB2F23C7F75001" pageNumber="14">concave outline from snout to just above eye with growth and noticeably in specimen over ca. 130-150 mm SL</td>
</tr>
<tr id="7A5AC237A24075D3903CACD82C9C4CCB" pageNumber="14">
<td id="9FEB50059E92B0793F91CAD46EF31F11" pageNumber="14">upper head profile</td>
<td id="20AB609257EB05CF9677654C0E8CFC8C" pageNumber="14">gently convex even with growth</td>
<td id="FDADB4D26695F2A139797676238DB17D" pageNumber="14">concave in specimens over 130-150 mm SL with growth</td>
</tr>
<tr id="45EA35FE0883838EF449289ED005405D" pageNumber="14">
<td id="4206CAE3962179085F2EE2E9E67E026C" pageNumber="14">molar teeth</td>
<td id="65A1585010EAA4B4F67041B49ED94A21" pageNumber="14">flat</td>
<td id="6F23FDAEC4C6A1312CCB2B9B7267C886" pageNumber="14">somewhat pointed</td>
</tr>
<tr id="5F19DEB33524453B41EC7423784B366E" pageNumber="14">
<td id="30090B42CEE17340E5E7BD09DB200296" pageNumber="14">3rd and 2nd anal-fin spine</td>
<td id="77D1566A17EB264BF2BA873240DD05AA" pageNumber="14">robust and dull pointed</td>
<td id="5B8B2D8855ADDD5DC4CDA4423E097D29" pageNumber="14">sharply pointed</td>
</tr>
<tr id="7EEEC092E33DC623FB7F79D72D25CE32" pageNumber="14">
<td id="D83C0C82B0524814AAACC7660363D132" pageNumber="14">body color</td>
<td id="91AD98E5A1F3D0F5463FDFBFFB9EA778" pageNumber="14">Dense black with clear whitish part of ventral head and abdomen</td>
<td id="6160F990CDFB03B9D0C7872E30985BAB" pageNumber="14">dusky black in whole body</td>
</tr>
<tr id="00D46578CA4FAACE160CE74C7F79CE27" pageNumber="14">
<td id="ACDFF3B088E60EC8FAAB49684B95AE3A" pageNumber="14">black spot of pectoral-fin base</td>
<td id="D7294F83798F927429B7889AFC728146" pageNumber="14">present</td>
<td id="7234F4C1EA78B54AA95D6C7B4F17F8CD" pageNumber="14">absent</td>
</tr>
<tr id="80DD451EE081DD491F875D7DD39E1857" pageNumber="14">
<td id="D26407C6E04C8933CF8E23EB3002EC43" pageNumber="14">pelvic-fin color</td>
<td id="140124066114CC2D99E4A2880140D27A" pageNumber="14">whitish</td>
<td id="A0ECA826E24733C786CDBDD873AD9608" pageNumber="14">dusky</td>
</tr>
</table>
</paragraph>
<paragraph id="B9FA6D5DDCE77962A511E10AEB9ADE93" lastPageNumber="15" pageNumber="14">
Although no types of
<taxonomicName id="0DB3E9EF09A99FFD45BAD1845737B4BC" LSID-ZBK="urn:lsid:zoobank.org:act:A2A91B6B-88CF-4D9F-B9BC-0C78FE2A2CAD" class="Actinopterygii" family="Sparidae" genus="Pagrus" kingdom="Animalia" lsidName-ZBK="Pagrus macrocephalus Basilewsky 1855:222" order="Perciformes" pageNumber="14" phylum="Chordata" rank="species" species="macrocephalus">Pagrus macrocephalus Basilewsky, 1855</taxonomicName>
are known at ZIN was considered to be a junior synonym of
<taxonomicName id="B9F1026DB4DF5F69DDD5C21502BF1C65" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="14" phylum="Chordata" rank="species" species="schlegelii">A. schlegelii (Bleeker, 1854)</taxonomicName>
by Akazaki (1962) and Sadovy &amp; Cornish (2000). However, Basilewskys (1855), figure 3, tab. I (referred to erroneously as tab. III, fig. 1, which is a sciaenid) is a sparid that is clearly a species of
<taxonomicName id="3000EB181921FDDB1735D6CE07DD05D2" LSID-ZBK="urn:lsid:zoobank.org:act:882A86DA-054A-4083-B084-A53AE774D481" class="Actinopterygii" family="Sparidae" genus="Pagrus" kingdom="Animalia" lsidName-ZBK="Pagrus Cuvier 1816:272" order="Perciformes" pageNumber="14" phylum="Chordata" rank="genus">Pagrus</taxonomicName>
, recognizable by its scaly interorbital area, red body, and 7 or 8 scale rows between the fifth dorsal-fin spine base and lateral line (Tables 1-2)(Akazaki, 1962).
<taxonomicName id="A4456A6BF9B9F8E9D9378A7BB16416B9" LSID-ZBK="urn:lsid:zoobank.org:act:882A86DA-054A-4083-B084-A53AE774D481" class="Actinopterygii" family="Sparidae" genus="Pagrus" kingdom="Animalia" lsidName-ZBK="Pagrus Cuvier 1816:272" order="Perciformes" pageNumber="14" phylum="Chordata" rank="genus">Pagrus</taxonomicName>
<taxonomicName id="F2670DC3F5DB1EE15292A6BD6C420C30" LSID-ZBK="urn:lsid:zoobank.org:act:A2A91B6B-88CF-4D9F-B9BC-0C78FE2A2CAD" class="Actinopterygii" family="Sparidae" genus="Pagrus" kingdom="Animalia" lsidName-ZBK="Pagrus macrocephalus Basilewsky 1855:222" order="Perciformes" pageNumber="14" phylum="Chordata" rank="species" species="macrocephalus">
<pageStartToken id="1EFD21364D97F98F4D26BFD31EB6B680" pageNumber="15">macrocephalus</pageStartToken>
</taxonomicName>
needs further examination and may be a synonym of
<taxonomicName id="DFBE7B7F1197AB6254CD64FAFECDCFD8" class="Actinopterygii" family="Sparidae" genus="Pagrus" kingdom="Animalia" order="Perciformes" pageNumber="14" phylum="Chordata" rank="species" species="major">Pagrus major (Temminck &amp; Schlegel, 1844)</taxonomicName>
.
</paragraph>
<paragraph id="D33A6F1CF6999FF17A33767B41252EF1" pageNumber="15">
Richardsons (1846)
<taxonomicName id="510F718845B7B0ECBAB5525738F01683" LSID-ZBK="urn:lsid:zoobank.org:act:371A4EC5-C0CA-4715-B9A6-CD1802B03F9F" family="Sparidae" genus="Chrysophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Chrysophrys xanthopoda Richardson 1846:241" order="Perciformes" pageNumber="15" phylum="Chordata" rank="species" species="xanthopoda">Chrysophrys xanthopoda</taxonomicName>
and
<taxonomicName id="FBAB499BA26AA06D563EC899C23DA2B0" LSID-ZBK="urn:lsid:zoobank.org:act:14BEE51E-67F6-4852-AFC8-31039E0395A8" family="Sparidae" genus="Chrysophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Chrysophrys auripes Richardson 1846:241" order="Perciformes" pageNumber="15" phylum="Chordata" rank="species" species="auripes">C. auripes</taxonomicName>
from Canton, Chinese Seas are based on BMNH stuffed syntypes that have not been located (J. Maclaine, pers. comm.). However, detailed unpublished color plates of both species in the BMNH library have yellowish pelvic, anal, and caudal fins, especially along with lower margins. These nominal species may, therefore, be junior synonyms of
<taxonomicName id="7C88D60E9F5409A99C29084F0F839545" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="15" phylum="Chordata" rank="species" species="latus">A. latus</taxonomicName>
, which has the same fin coloration (see description of Akazaki, 1962, 1984 and Carpenter, 2001).
</paragraph>
<paragraph id="13FABD235AD2290EF82BF61E337401EC" pageNumber="15">
<taxonomicName id="E57096729C4C1CBE483E9D88000F9B32" LSID-ZBK="urn:lsid:zoobank.org:act:0CC2B529-9327-4928-8B49-758CC0351A34" family="Sparidae" genus="Chrysophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Chrysophrys longispinnis Valenciennes 1830:116" order="Perciformes" pageNumber="15" phylum="Chordata" rank="species" species="longispinnis">Chrysophrys longispinnis Valenciennes in Cuvier &amp; Valenciennes, 1830</taxonomicName>
, considered by Bauchot &amp; Daget (1972) to be a junior synonym of
<taxonomicName id="29455FD82B4C7E6FC491ADC713FCA184" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="15" phylum="Chordata" rank="species" species="berda">A. berda</taxonomicName>
, is unlikely to be that species. Two of 3 syntypes of
<taxonomicName id="AD144E8B595977B2027B33572F25B4FC" LSID-ZBK="urn:lsid:zoobank.org:act:0CC2B529-9327-4928-8B49-758CC0351A34" family="Sparidae" genus="Chrysophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Chrysophrys longispinnis Valenciennes 1830:116" order="Perciformes" pageNumber="15" phylum="Chordata" rank="species" species="longispinnis">C. longispinnis</taxonomicName>
from Bengal have an extremely long second anal-fin spine (1.59-1.60 ratio of second anal-fin spine to third anal-fin spine vs. 1.23-1.50, mean 1.36 in
<taxonomicName id="4B8A4D52F795B199DB70FF1E27980B26" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="15" phylum="Chordata" rank="species" species="berda">A. berda</taxonomicName>
, and 1.32-1.40, mean 1.37 in
<taxonomicName id="44C0B8E53120C0C941036C3E17ACC997" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="15" phylum="Chordata" rank="species" species="taiwanensis">A. taiwanensis</taxonomicName>
; see Table 1). The third syntype from Japan has a relatively deep body and fine minute posteroventral serration of the preopercular flange characteristic of Japanese
<taxonomicName id="A01B2B64033EEEE746DF75809E6A8675" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="15" phylum="Chordata" rank="species" species="latus">A. latus (Houttuyn, 1782)</taxonomicName>
(Akazaki, 1962). The type series of
<taxonomicName id="00AB68064C43826EEBCF4B724B382FDA" LSID-ZBK="urn:lsid:zoobank.org:act:0CC2B529-9327-4928-8B49-758CC0351A34" family="Sparidae" genus="Chrysophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Chrysophrys longispinnis Valenciennes 1830:116" order="Perciformes" pageNumber="15" phylum="Chordata" rank="species" species="longispinnis">C. longispinnis</taxonomicName>
, therefore, includes 2 species, neither of which is
<taxonomicName id="9512B7D8AA3EE5EA6AC1DC1A8DCADB25" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="15" phylum="Chordata" rank="species" species="berda">A. berda</taxonomicName>
or
<taxonomicName id="0BED48B0A2724F901CD8FD5BC3563D24" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="15" phylum="Chordata" rank="species" species="taiwanensis">A. taiwanensis</taxonomicName>
.
</paragraph>
<paragraph id="B5C15F36CA86CA4735752AEF3A11CC26" pageNumber="15">
<taxonomicName id="3D6D6D20EC82C105E14F091643E3CCF8" LSID-ZBK="urn:lsid:zoobank.org:act:48EDB4F6-3F5B-477D-B703-9840D6709A3C" family="Sparidae" genus="Chrysophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Chrysophrys cuvieri Day 1875:141" order="Perciformes" pageNumber="15" phylum="Chordata" rank="species" species="cuvieri">Chrysophrys cuvieri</taxonomicName>
, described (Day, 1875) from Mangalore, India, has been synonymized with
<taxonomicName id="A0225C835E906D9D91891CD8B172CBBB" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="15" phylum="Chordata" rank="species" species="berda">A. berda</taxonomicName>
(Whitehead &amp; Talwar, 1976; Randall, 1995; Ferraris et al, 2000), although the type designation is unclear. Whitehead &amp; Talwar (1976) identified the following potential Day specimens of
<taxonomicName id="4DD7A1530779CBEA7BCD6AF94E5C41A5" LSID-ZBK="urn:lsid:zoobank.org:act:48EDB4F6-3F5B-477D-B703-9840D6709A3C" family="Sparidae" genus="Chrysophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Chrysophrys cuvieri Day 1875:141" order="Perciformes" pageNumber="15" phylum="Chordata" rank="species" species="cuvieri">C. cuvieri</taxonomicName>
: AMS B.8225, 129 mm SL, Madras; BMNH 1975.9.30.21, 109 mm SL, Cochin; RMNH 1060, not seen by us. The original description and type locality of
<taxonomicName id="9B4DE203C8F2F263986CEE8637C768A5" LSID-ZBK="urn:lsid:zoobank.org:act:48EDB4F6-3F5B-477D-B703-9840D6709A3C" family="Sparidae" genus="Chrysophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Chrysophrys cuvieri Day 1875:141" order="Perciformes" pageNumber="15" phylum="Chordata" rank="species" species="cuvieri">C. cuvieri</taxonomicName>
was apparently based on ZSI 1782 although the type was not seen by us. Bauchot &amp; Smith (1983) synonymized
<taxonomicName id="EE8C484748A3F10CA7821811262D7DE6" LSID-ZBK="urn:lsid:zoobank.org:act:48EDB4F6-3F5B-477D-B703-9840D6709A3C" family="Sparidae" genus="Chrysophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Chrysophrys cuvieri Day 1875:141" order="Perciformes" pageNumber="15" phylum="Chordata" rank="species" species="cuvieri">C. cuvieri</taxonomicName>
with
<taxonomicName id="4EE9CAFE50F497CF061EC6850368F441" class="Actinopterygii" family="Sparidae" genus="Sparidentex" kingdom="Animalia" order="Perciformes" pageNumber="15" phylum="Chordata" rank="species" species="hasta">Sparidentex hasta (Valenciennes in Cuvier &amp; Valenciennes, 1830)</taxonomicName>
using dentition inconsistent with
<taxonomicName id="4193023C42A4C5405C6C6AA8660B5783" LSID-ZBK="urn:lsid:zoobank.org:act:8F9E30E9-3A84-43EA-8E93-F852D3E82151" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" lsidName-ZBK="Acanthopagrus Chrysophrys Peters 1855:242" order="Perciformes" pageNumber="15" phylum="Chordata" rank="genus">Acanthopagrus</taxonomicName>
as a key character. Days original description of
<taxonomicName id="1B9C567E501B1EB36DE7463CCAF7E1AA" LSID-ZBK="urn:lsid:zoobank.org:act:48EDB4F6-3F5B-477D-B703-9840D6709A3C" family="Sparidae" genus="Chrysophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Chrysophrys cuvieri Day 1875:141" order="Perciformes" pageNumber="15" phylum="Chordata" rank="species" species="cuvieri">C. cuvieri</taxonomicName>
supports this contention since species of
<taxonomicName id="328E9BF87B8852CD9488F7C4A63D02F3" LSID-ZBK="urn:lsid:zoobank.org:act:499A51BC-ABBD-4D81-8990-D60A67430880" class="Actinopterygii" family="Sparidae" genus="Sparidentex" kingdom="Animalia" lsidName-ZBK="Sparidentex Munro 1948:276" order="Perciformes" pageNumber="15" phylum="Chordata" rank="genus">Sparidentex</taxonomicName>
lack the large molariform teeth typically found in
<taxonomicName id="947F6B1490BF4762B003C3F2A4375DDC" LSID-ZBK="urn:lsid:zoobank.org:act:8F9E30E9-3A84-43EA-8E93-F852D3E82151" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" lsidName-ZBK="Acanthopagrus Chrysophrys Peters 1855:242" order="Perciformes" pageNumber="15" phylum="Chordata" rank="genus">Acanthopagrus</taxonomicName>
. Days (1875) description states: “Teeth -four to six sharp, pointed and rather conical incisors in front of either jaw, with villiform teeth behind them: a pointed and compressed row along the outer side of either jaw, the last few of which are small and with rounded crowns; internal to these are two rounded small molars in the lower and three in the upper jaw.” One of Days specimens of AMS B.8225 is clearly an example of
<taxonomicName id="30AAC70793DAD3CB92004DADB3F3F7FF" LSID-ZBK="urn:lsid:zoobank.org:act:499A51BC-ABBD-4D81-8990-D60A67430880" class="Actinopterygii" family="Sparidae" genus="Sparidentex" kingdom="Animalia" lsidName-ZBK="Sparidentex Munro 1948:276" order="Perciformes" pageNumber="15" phylum="Chordata" rank="genus">Sparidentex</taxonomicName>
without the typical molariform teeth found in
<taxonomicName id="998EE3985D9FE8913058ADCDB3DAA0D6" LSID-ZBK="urn:lsid:zoobank.org:act:8F9E30E9-3A84-43EA-8E93-F852D3E82151" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" lsidName-ZBK="Acanthopagrus Chrysophrys Peters 1855:242" order="Perciformes" pageNumber="15" phylum="Chordata" rank="genus">Acanthopagrus</taxonomicName>
. Although its status as a synonym of
<taxonomicName id="C98F1AFE5EAE458426601B6AD999CF64" class="Actinopterygii" family="Sparidae" genus="Sparidentex" kingdom="Animalia" order="Perciformes" pageNumber="15" phylum="Chordata" rank="species" species="hasta">S. hasta</taxonomicName>
needs to be resolved,
<taxonomicName id="931D64ECC26D1F384E2895D82E5677BC" LSID-ZBK="urn:lsid:zoobank.org:act:48EDB4F6-3F5B-477D-B703-9840D6709A3C" family="Sparidae" genus="Chrysophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Chrysophrys cuvieri Day 1875:141" order="Perciformes" pageNumber="15" phylum="Chordata" rank="species" species="cuvieri">C. cuvieri</taxonomicName>
is clearly distinct from
<taxonomicName id="A558DFBD6ABB54D97ACA4146B8249974" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="15" phylum="Chordata" rank="species" species="berda">A. berda</taxonomicName>
and
<taxonomicName id="4E473CC07D6617769CBE66FF29B787F7" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="15" phylum="Chordata" rank="species" species="taiwanensis">A. taiwanensis</taxonomicName>
.
</paragraph>
<paragraph id="637F20B17C6823A0D9254A5231682DBA" pageNumber="15">
Akazaki (1962) synonymized
<taxonomicName id="FA62C94F21E1EE745D274317C070CB5A" LSID-ZBK="urn:lsid:zoobank.org:act:4EDB16E8-A3F3-4CEA-B5E1-78C96B102347" class="Actinopterygii" family="Sparidae" genus="Sparus" kingdom="Animalia" lsidName-ZBK="Sparus chrysopterus Kishinouye 1907:327" order="Perciformes" pageNumber="15" phylum="Chordata" rank="species" species="chrysopterus">Sparus chrysopterus Kishinouye, 1907</taxonomicName>
(type locality: Kyushu, Shikoku, Inland Sea, Pacific coast of Hondo, Japan) with
<taxonomicName id="D8D1891678DF6709F64DCEE4E36CF399" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="15" phylum="Chordata" rank="species" species="latus">A. latus</taxonomicName>
. Although the type appears to be lost at ZUMT, Kishinouye (1907) noted that the pelvic and anal fins of
<taxonomicName id="C76FEA25EBE12D42B8E089DB052AC542" LSID-ZBK="urn:lsid:zoobank.org:act:4EDB16E8-A3F3-4CEA-B5E1-78C96B102347" class="Actinopterygii" family="Sparidae" genus="Sparus" kingdom="Animalia" lsidName-ZBK="Sparus chrysopterus Kishinouye 1907:327" order="Perciformes" pageNumber="15" phylum="Chordata" rank="species" species="chrysopterus">S. chrysopterus</taxonomicName>
are yellow, a character typical of
<taxonomicName id="2E61893942225F003D4C4020B4B4A3F0" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="15" phylum="Chordata" rank="species" species="latus">A. latus</taxonomicName>
. The description of the molariform teeth of this species is similar to those of
<taxonomicName id="A590F9E0C4A85CBE3D8988F3479DF0B9" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="15" phylum="Chordata" rank="species" species="schlegelii">A. schlegelii</taxonomicName>
and clearly places it in the genus
<taxonomicName id="617944BA0417268D9B859C71F19B83BD" LSID-ZBK="urn:lsid:zoobank.org:act:8F9E30E9-3A84-43EA-8E93-F852D3E82151" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" lsidName-ZBK="Acanthopagrus Chrysophrys Peters 1855:242" order="Perciformes" pageNumber="15" phylum="Chordata" rank="genus">Acanthopagrus</taxonomicName>
, although distinct from
<taxonomicName id="A575D0F1F8473274199654F22A5CE798" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="15" phylum="Chordata" rank="species" species="berda">A. berda</taxonomicName>
and
<taxonomicName id="A48DC9CC72A0DD384DEA8D117AA297BC" class="Actinopterygii" family="Sparidae" genus="Acanthopagrus" kingdom="Animalia" order="Perciformes" pageNumber="15" phylum="Chordata" rank="species" species="taiwanensis">A. taiwanensis</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>