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<document id="C7E5245861068564DA7FE5B7A77F06B7" ID-CLB-Dataset="31806" ID-DOI="10.5281/zenodo.893555" ID-GBIF-Dataset="4a2efe36-c48b-456b-9223-7b5793025a72" ID-ISSN="0035-418" ID-Zenodo-Dep="893555" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1508361162557" checkinUser="plazi" docAuthor="Peter J. Schwendinger" docDate="2017" docId="4C30A452FFE6FFF3B99DFE733975FB0E" docLanguage="en" docName="rsz_124_2_11_Schwendinger.pdf" docOrigin="Revue suisse de Zoologie 124 (2)" docStyle="DocumentStyle:225056F0A23C51DDC06AC27EAEA15D5B.2:RevSuisseZool.2015-2019.journal_article" docStyleId="225056F0A23C51DDC06AC27EAEA15D5B" docStyleName="RevSuisseZool.2015-2019.journal_article" docStyleVersion="2" docTitle="Lipistius malayanus" docType="treatment" docVersion="9" lastPageNumber="395" masterDocId="B009DC2AFFE4FFF7BA6FFFF13818FFE3" masterDocTitle="A revision of the trapdoor spider genus Liphistius (Mesothelae: Liphistiidae) in peninsular Malaysia; part 1" masterLastPageNumber="445" masterPageNumber="391" pageNumber="393" updateTime="1698469770161" updateUser="ExternalLinkService">
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<mods:title id="5FCBB25F5D4BE84500E5136C72796906">A revision of the trapdoor spider genus Liphistius (Mesothelae: Liphistiidae) in peninsular Malaysia; part 1</mods:title>
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<mods:namePart id="3A4BAA6BA617A756E5D2DD4E20D2130D">Peter J. Schwendinger</mods:namePart>
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<mods:date id="A5BA322E348AE8FD5B20952D82B7D9D5">2017</mods:date>
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<mods:number id="175507968D2E663EC13B9FFAE9A7F3A6">2017-09-30</mods:number>
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<treatment id="4C30A452FFE6FFF3B99DFE733975FB0E" ID-DOI="http://doi.org/10.5281/zenodo.6042347" ID-GBIF-Taxon="135450778" ID-Zenodo-Dep="6042347" LSID="urn:lsid:plazi:treatment:4C30A452FFE6FFF3B99DFE733975FB0E" httpUri="http://treatment.plazi.org/id/4C30A452FFE6FFF3B99DFE733975FB0E" lastPageId="4" lastPageNumber="395" pageId="2" pageNumber="393">
<subSection id="AD160CAFFFE6FFF4B99DFE733BA3F7FE" lastPageId="3" lastPageNumber="394" pageId="2" pageNumber="393" type="key">
<subSection id="AD160CAFFFE6FFF5B99DFE733CF2FE79" box="[1010,1258,385,410]" pageId="2" pageNumber="393" type="multiple">
<subSubSection id="8C8346CFFFE6FFF5B99DFE733CF2FE79" box="[1010,1258,385,410]" pageId="2" pageNumber="393" type="nomenclature">
<paragraph id="C4261544FFE6FFF5B99DFE733CF2FE79" blockId="2.[1010,1258,385,410]" box="[1010,1258,385,410]" pageId="2" pageNumber="393">
<heading id="9F6EA228FFE6FFF5B99DFE733CF2FE79" bold="true" box="[1010,1258,385,410]" fontSize="10" level="2" pageId="2" pageNumber="393" reason="2">
<emphasis id="F6EDC956FFE6FFF5B99DFE733CF2FE79" bold="true" box="[1010,1258,385,410]" pageId="2" pageNumber="393">
The
<taxonomicName id="03996EC7FFE6FFF5BE4AFE703C85FE7B" box="[1061,1181,385,408]" class="Arachnida" family="Liphistiidae" genus="Lipistius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="393" phylum="Arthropoda" rank="species" species="malayanus">
<emphasis id="F6EDC956FFE6FFF5BE4AFE703C85FE7B" bold="true" box="[1061,1181,385,408]" italics="true" pageId="2" pageNumber="393">malayanus</emphasis>
</taxonomicName>
-group
</emphasis>
</heading>
</paragraph>
</subSubSection>
</subSection>
<subSubSection id="8C8346CFFFE6FFF5B950FE403B44FD2A" pageId="2" pageNumber="393" type="diagnosis">
<paragraph id="C4261544FFE6FFF5B950FE403B44FD2A" blockId="2.[831,1437,433,713]" pageId="2" pageNumber="393">
<emphasis id="F6EDC956FFE6FFF5B950FE403BA0FE2B" bold="true" box="[831,952,433,456]" pageId="2" pageNumber="393">Diagnosis:</emphasis>
Distinguished from the
<taxonomicName id="03996EC7FFE6FFF5BE83FE433D3FFE2A" authorityName="Platnick &amp; Sedgwick" authorityYear="1984" box="[1260,1319,434,457]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="393" phylum="Arthropoda" rank="species" species="trang">
<emphasis id="F6EDC956FFE6FFF5BE83FE433D3FFE2A" box="[1260,1319,434,457]" italics="true" pageId="2" pageNumber="393">trang</emphasis>
</taxonomicName>
-group by an indistinctly split embolus proper (
<figureCitation id="5CA209C1FFE6FFF5BE9DFE203B4BFDEA" captionStart="Fig. 3" captionStartId="5.[151,186,1940,1961]" captionTargetBox="[261,1329,261,1882]" captionTargetId="figure@5.[255,1334,257,1886]" captionTargetPageId="5" captionText="Fig. 3. SEM-micrographs of embolus proper of left palpal organ of two males: Liphistius desultor from Penang Hill (A, C, E, G) and Liphistius malayanus from Fraser’s Hill (B, D, F, H). (A) Retroventral-distal view. (B) Dorsal-distal view. (C) Distal view (dorsal side to the right). (D) Distal view (prolateral side to the right). (E) Retrodorsal view. (F) Prodorsal view. (G) Proventral view. (H) Prolateral view. Abbreviations: f - major embolic fold; m - membranous part of embolus proper; s - sclerotised part of embolus proper." httpUri="https://zenodo.org/record/1021943/files/figure.png" pageId="2" pageNumber="393">Fig. 3B, D, F, H</figureCitation>
) and from all other species groups by distal edge of contrategulum proventrally ending in a V-shaped or U-shaped row of denticles (the clearest synapomorphy of this group;
<figureCitation id="5CA209C1FFE6FFF5B986FDA03C5EFD8A" box="[1001,1094,593,617]" captionStart="Fig. 4" captionStartId="6.[151,186,1713,1734]" captionTargetBox="[156,1432,265,1656]" captionTargetId="figure@6.[148,1439,254,1693]" captionTargetPageId="6" captionText="Fig. 4. Liphistius malayanus, details of palp of four males: Templer Park (A, C-D, H-I); holotype of L. malayanus cameroni (B, E, G); Cameron Highlands, ZRC (F, J), Fraser’s Hill, MHNG (K-L). (A) Paracymbium and tibial apophysis of left palp, retroventral view. (B) Tibial apophysis of right palp, ventral view. (C) Distal part of left palpal organ, proventral view (arrow indicating V-shaped row of denticles at proventral end of distal edge of contrategulum). (D, F) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (G-H) Left palpal organ, distal view (dorsal side up). (I, K) Distal edge of contrategulum of right palp, distal view (dorsal side to the left). (J) Same, distal and slighly prolateral view. (L) Same of left palp, distal and slighly prolateral view (dorsal side to the right). Abbreviations: a - dorsal apex of contrategulum; de - distal edge of contrategulum. Scale lines: 1.0 mm (A; B; C-D; E, G; F; H-L)." httpUri="https://zenodo.org/record/1021945/files/figure.png" pageId="2" pageNumber="393">Figs 4C</figureCitation>
; 6A, C; 8A, D). Vulval plate strongly sclerotised, with a large, undivided CDO and with a large, undivided receptacular cluster (
<figureCitation id="5CA209C1FFE6FFF5BF5FFD603D61FD4B" box="[1328,1401,657,680]" captionStart="Fig. 5" captionStartId="8.[151,186,1809,1830]" captionTargetBox="[154,1439,260,1762]" captionTargetId="figure@8.[148,1439,254,1779]" captionTargetPageId="8" captionText="Fig. 5. Liphistius malayanus, vulval plate of eight females: paratype of L. m. cameroni from Berinchang, Cameron Highlands (A- B); exuvia of female from Ringlet, Cameron Highlands (C); female (SMF 64093) from Fraser’s Hill (D); female (moulted 24. VI. 2002) from Fraser’s Hill; poreplate slightly deformed by brief immersion in cold KOH (E); female (SMF 56206) from Fraser’s Hill (F); female (SMF 60037) from Genting Highlands (G); female (moulted 8. XII. 2005) from Genting Highlands (H); exuvia of female from Templer Park (I-J). (A, C-I) Dorsal view. (B, J) Ventral view. Scale lines: 1.0 mm (A-B; C, F; D; E, G; H; I-J)." httpUri="https://zenodo.org/record/1021947/files/figure.png" pageId="2" pageNumber="393">Figs 5</figureCitation>
,
<figureCitation id="5CA209C1FFE6FFF5BFE7FD633D8FFD4A" box="[1416,1431,658,681]" captionStart="Fig. 7" captionStartId="12.[151,186,1940,1961]" captionTargetBox="[159,1415,617,1889]" captionTargetId="figure@12.[151,1436,609,1911]" captionTargetPageId="12" captionText="Fig. 7. Liphistius endau, vulval plate of six females: adult from unknown locality (A-B); largest female from Kota Tinggi (C); exuvia of reproductive female (moulted 27. IV. 2002) from type locality (D-E); exuvia of reproductive female (moulted 28. IV. 2002) from type locality (F-G); medium-sized juvenile from type locality (H); small juvenile from type locality (I). (A, C-D, F, H-I) Entire vulval plate, dorsal view. (B) Poreplate, ventral view. (E, G) Entire vulval plate, ventral view. Scale lines: 1.0 mm (A-B; C; D-E; F-G; H-I)." httpUri="https://zenodo.org/record/1021951/files/figure.png" pageId="2" pageNumber="393">7</figureCitation>
,
<figureCitation id="5CA209C1FFE6FFF5B950FD403B57FD2A" box="[831,847,689,713]" captionStart="Fig. 9" captionStartId="16.[151,186,1607,1628]" captionTargetBox="[157,1426,257,1572]" captionTargetId="figure@16.[148,1439,254,1598]" captionTargetPageId="16" captionText="Fig. 9. Liphistius gracilis sp. nov., vulval plate of three females (all from exuviae): allotype (A-B); paratype, moult 31. VIII. 2001 (C- D); paratype, moult 27. IX. 2001 (E-G). (A, C, E) Vulval plate, dorsal view. (B, D, F) Same, ventral view. (G) Same, anterior view. Scale lines: 1.0 mm (A-B; C-G)." httpUri="https://zenodo.org/record/1021955/files/figure.png" pageId="2" pageNumber="393">9</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="90E645CCFFE7FFF4BAFAFC803954FC23" httpUri="https://zenodo.org/record/1021941/files/figure.png" pageId="3" pageNumber="394" targetBox="[166,1405,257,835]" targetPageId="3">
<paragraph id="C4261544FFE7FFF4BAFAFC803954FC23" blockId="3.[149,1437,881,961]" pageId="3" pageNumber="394">
Fig. 2. Habitus of two
<taxonomicName id="03996EC7FFE7FFF4BB14FC8039C2FC65" authorityName="Schiodte" authorityYear="1849" box="[379,474,881,902]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="394" phylum="Arthropoda" rank="genus">
<emphasis id="F6EDC956FFE7FFF4BB14FC8039C2FC65" box="[379,474,881,902]" italics="true" pageId="3" pageNumber="394">Liphistius</emphasis>
</taxonomicName>
species from peninsular Malaysia and southern Thailand. (A)
<taxonomicName id="03996EC7FFE7FFF4BE2FFC803CF9FC65" authorityName="Sedgwick &amp; Platnick" authorityYear="1987" box="[1088,1249,881,902]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="endau">
<emphasis id="F6EDC956FFE7FFF4BE2FFC803CF9FC65" box="[1088,1249,881,902]" italics="true" pageId="3" pageNumber="394">Liphistius endau</emphasis>
</taxonomicName>
, female from Kota Tinggi (Malaysia), dorsal view. (B)
<taxonomicName id="03996EC7FFE7FFF4B82CFC7F3AC6FC40" authority="Peter J. Schwendinger, 2017" authorityName="Peter J. Schwendinger" authorityYear="2017" box="[579,734,910,931]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="indra" status="sp. nov.">
<emphasis id="F6EDC956FFE7FFF4B82CFC7F3AC6FC40" box="[579,734,910,931]" italics="true" pageId="3" pageNumber="394">Liphistius indra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EDDE742DFFE7FFF4B889FC7E3B2CFC47" box="[742,820,911,932]" pageId="3" pageNumber="394" rank="species">sp. nov.</taxonomicNameLabel>
, male paratype from the Sankalakhierie Mountains (Thailand), same view.
</paragraph>
</caption>
<subSubSection id="8C8346CFFFE7FFF4BAF8FBE73CF1F9F5" pageId="3" pageNumber="394" type="description">
<paragraph id="C4261544FFE7FFF4BAF8FBE73CF1F9F5" blockId="3.[151,757,1046,2079]" lastBlockId="3.[831,1437,1046,1559]" pageId="3" pageNumber="394">
<emphasis id="F6EDC956FFE7FFF4BAF8FBE7393EFBCE" bold="true" box="[151,294,1046,1069]" pageId="3" pageNumber="394">Description:</emphasis>
Small to large species (carapace length of males 4.36-12.04, carapace width 3.64-10.99); body colouration uniformly dark (in large species:
<taxonomicName id="03996EC7FFE7FFF4BAF8FB883936FB6D" box="[151,302,1143,1168]" class="Arachnida" family="Liphistiidae" genus="Lipistius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="malayanus">
<emphasis id="F6EDC956FFE7FFF4BAF8FB883936FB6D" box="[151,302,1143,1168]" italics="true" pageId="3" pageNumber="394">L. malayanus</emphasis>
</taxonomicName>
and
<taxonomicName id="03996EC7FFE7FFF4BB02FB8839C9FB6D" authorityName="Sedgwick &amp; Platnick" authorityYear="1987" box="[365,465,1143,1168]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="endau">
<emphasis id="F6EDC956FFE7FFF4BB02FB8839C9FB6D" box="[365,465,1143,1168]" italics="true" pageId="3" pageNumber="394">L. endau</emphasis>
</taxonomicName>
;
<figureCitation id="5CA209C1FFE7FFF4BB8BFB863A25FB73" box="[484,573,1143,1168]" captionStart="Fig. 2" captionStartId="3.[149,184,881,902]" captionTargetBox="[166,1405,257,835]" captionTargetId="figure@3.[149,1420,257,835]" captionTargetPageId="3" captionText="Fig. 2. Habitus of two Liphistius species from peninsular Malaysia and southern Thailand. (A) Liphistius endau, female from Kota Tinggi (Malaysia), dorsal view. (B) Liphistius indra sp. nov., male paratype from the Sankalakhierie Mountains (Thailand), same view." httpUri="https://zenodo.org/record/1021941/files/figure.png" pageId="3" pageNumber="394">Fig. 2A</figureCitation>
) or brown with annulated legs and palps (in smaller species:
<taxonomicName id="03996EC7FFE7FFF4B8FFFB683AECFB4C" authorityName="Platnick &amp; Sedgwick" authorityYear="1984" box="[656,756,1176,1200]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="johore">
<emphasis id="F6EDC956FFE7FFF4B8FFFB683AECFB4C" box="[656,756,1176,1200]" italics="true" pageId="3" pageNumber="394">L. johore</emphasis>
</taxonomicName>
and
<taxonomicName id="03996EC7FFE7FFF4BAA7FB4B3922FB2C" authority="Peter J. Schwendinger, 2017" authorityName="Peter J. Schwendinger" authorityYear="2017" box="[200,314,1208,1233]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="gracilis" status="sp. nov.">
<emphasis id="F6EDC956FFE7FFF4BAA7FB4B3922FB2C" box="[200,314,1208,1233]" italics="true" pageId="3" pageNumber="394">L. gracilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EDDE742DFFE7FFF4BB2AFB4B39B8FB32" box="[325,416,1209,1233]" pageId="3" pageNumber="394" rank="species">sp. nov.</taxonomicNameLabel>
). Male palp with membranous and sclerotised part of embolus proper in close contact with each other (both distinctly separated in
<taxonomicName id="03996EC7FFE7FFF4B8DFFB0A3AF3FAF1" authorityName="Platnick &amp; Sedgwick" authorityYear="1984" box="[688,747,1275,1298]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="trang">
<emphasis id="F6EDC956FFE7FFF4B8DFFB0A3AF3FAF1" box="[688,747,1275,1298]" italics="true" pageId="3" pageNumber="394">trang</emphasis>
</taxonomicName>
- group), together forming a closed tube; dorsal wall of sclerotized part of embolus proper straight and directed proventrad (
<taxonomicName id="03996EC7FFE7FFF4BB12FAAD3A08FA91" box="[381,528,1371,1395]" class="Arachnida" family="Liphistiidae" genus="Lipistius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="malayanus">
<emphasis id="F6EDC956FFE7FFF4BB12FAAD3A08FA91" box="[381,528,1371,1395]" italics="true" pageId="3" pageNumber="394">L. malayanus</emphasis>
</taxonomicName>
;
<figureCitation id="5CA209C1FFE7FFF4B84EFAAA3A7BFA97" box="[545,611,1371,1396]" captionStart="Fig. 4" captionStartId="6.[151,186,1713,1734]" captionTargetBox="[156,1432,265,1656]" captionTargetId="figure@6.[148,1439,254,1693]" captionTargetPageId="6" captionText="Fig. 4. Liphistius malayanus, details of palp of four males: Templer Park (A, C-D, H-I); holotype of L. malayanus cameroni (B, E, G); Cameron Highlands, ZRC (F, J), Fraser’s Hill, MHNG (K-L). (A) Paracymbium and tibial apophysis of left palp, retroventral view. (B) Tibial apophysis of right palp, ventral view. (C) Distal part of left palpal organ, proventral view (arrow indicating V-shaped row of denticles at proventral end of distal edge of contrategulum). (D, F) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (G-H) Left palpal organ, distal view (dorsal side up). (I, K) Distal edge of contrategulum of right palp, distal view (dorsal side to the left). (J) Same, distal and slighly prolateral view. (L) Same of left palp, distal and slighly prolateral view (dorsal side to the right). Abbreviations: a - dorsal apex of contrategulum; de - distal edge of contrategulum. Scale lines: 1.0 mm (A; B; C-D; E, G; F; H-L)." httpUri="https://zenodo.org/record/1021945/files/figure.png" pageId="3" pageNumber="394">Fig. 4</figureCitation>
G-H), curved prodorsad (
<taxonomicName id="03996EC7FFE7FFF4BB74FA8C399AFA71" authorityName="Sedgwick &amp; Platnick" authorityYear="1987" box="[283,386,1403,1428]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="endau">
<emphasis id="F6EDC956FFE7FFF4BB74FA8C399AFA71" box="[283,386,1403,1428]" italics="true" pageId="3" pageNumber="394">L. endau</emphasis>
</taxonomicName>
;
<figureCitation id="5CA209C1FFE7FFF4BBF6FA8A3A00FA77" box="[409,536,1403,1428]" captionStart="Fig. 6" captionStartId="11.[151,186,1824,1845]" captionTargetBox="[232,1351,260,1775]" captionTargetId="figure@11.[215,1371,254,1797]" captionTargetPageId="11" captionText="Fig. 6. Liphistius endau, details of left palp of eight males: from type locality, matured 27. II. 2002 (A-F); from type locality, matured 1. II. 2002 (G-I); from unknown locality, died 22. III. 2012 (J); from unknown locality, killed 30. IX. 2010 (K); from Gunung Belumut (L); from Kota Tinggi, matured 22. X. 2014 (M); same place, matured 20. VIII. 2014 (N); same place, matured 6. XI. 2014 (O). (A) Distal part of palp, ventral view. (B) Paracymbium and tibial apophysis, retroventral view. (C) Palpal organ, proventral view (arrow indicating U-shaped row of denticles at proventral end of distal edge of contrategulum). (D) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (F, H) Same, distal view (dorsal side up). (G) Embolus complex, proventral and slightly distal view. (I) Distal edge of contrategulum, distal and slighly prolateral view (dorsal side to the right). (J-O) Same, distal view. Abbreviations: dw - dorsal wall of sclerotized part of embolus proper; t - tooth on distal edge of contrategulum. Scale lines: 1.0 mm (A-B; C-E, G-H; F, I-O)." httpUri="https://zenodo.org/record/1021949/files/figure.png" pageId="3" pageNumber="394">Fig. 6F, H</figureCitation>
) or bent prodosad (
<taxonomicName id="03996EC7FFE7FFF4BACFFA6C390FFA50" authority="Peter J. Schwendinger, 2017" authorityName="Peter J. Schwendinger" authorityYear="2017" box="[160,279,1436,1460]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="gracilis" status="sp. nov.">
<emphasis id="F6EDC956FFE7FFF4BACFFA6C390FFA50" box="[160,279,1436,1460]" italics="true" pageId="3" pageNumber="394">L. gracilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EDDE742DFFE7FFF4BB4BFA6C399AFA57" box="[292,386,1437,1460]" pageId="3" pageNumber="394" rank="species">sp. nov.</taxonomicNameLabel>
;
<figureCitation id="5CA209C1FFE7FFF4BBFBFA6D39C0FA56" box="[404,472,1436,1461]" captionStart="Fig. 8" captionStartId="15.[151,186,1862,1883]" captionTargetBox="[159,1417,261,1808]" captionTargetId="figure@15.[148,1439,254,1832]" captionTargetPageId="15" captionText="Fig. 8. Liphistius gracilis sp. nov., details of palp of three males: paratype, matured 5. I. 2002 (A-B, H); holotype (C-F, I-J); paratype, matured 19. IX. 2001 (G). (A) Distal part of palp, ventral view. (B) Same, retrolateral view. (C) Subtegulum, cymbium and distal part of palpal tibia, ventral view. (D) Distal part of cymbium and palpal organ, proventral view (arrow indicating U-shaped row of denticles at proventral end of distal edge of contrategulum). (E) Same, retrodorsal view. (F-G) Same, distal view (dorsal side up). (H-I) Right tibial apophysis, retrolateral and slightly proximal view. (J) Left tibial apophysis, same view. Scale lines: 1.0 mm (A-C), 0.5 mm (D-G; H-J)." httpUri="https://zenodo.org/record/1021953/files/figure.png" pageId="3" pageNumber="394">Fig. 8</figureCitation>
F-G); para-embolic plate short or absent (
<figureCitation id="5CA209C1FFE7FFF4BB3CFA4D39A9FA36" box="[339,433,1468,1493]" captionStart="Fig. 4" captionStartId="6.[151,186,1713,1734]" captionTargetBox="[156,1432,265,1656]" captionTargetId="figure@6.[148,1439,254,1693]" captionTargetPageId="6" captionText="Fig. 4. Liphistius malayanus, details of palp of four males: Templer Park (A, C-D, H-I); holotype of L. malayanus cameroni (B, E, G); Cameron Highlands, ZRC (F, J), Fraser’s Hill, MHNG (K-L). (A) Paracymbium and tibial apophysis of left palp, retroventral view. (B) Tibial apophysis of right palp, ventral view. (C) Distal part of left palpal organ, proventral view (arrow indicating V-shaped row of denticles at proventral end of distal edge of contrategulum). (D, F) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (G-H) Left palpal organ, distal view (dorsal side up). (I, K) Distal edge of contrategulum of right palp, distal view (dorsal side to the left). (J) Same, distal and slighly prolateral view. (L) Same of left palp, distal and slighly prolateral view (dorsal side to the right). Abbreviations: a - dorsal apex of contrategulum; de - distal edge of contrategulum. Scale lines: 1.0 mm (A; B; C-D; E, G; F; H-L)." httpUri="https://zenodo.org/record/1021945/files/figure.png" pageId="3" pageNumber="394">Figs 4D</figureCitation>
,
<figureCitation id="5CA209C1FFE7FFF4BBAEFA4D39FAFA30" box="[449,482,1468,1491]" captionStart="Fig. 6" captionStartId="11.[151,186,1824,1845]" captionTargetBox="[232,1351,260,1775]" captionTargetId="figure@11.[215,1371,254,1797]" captionTargetPageId="11" captionText="Fig. 6. Liphistius endau, details of left palp of eight males: from type locality, matured 27. II. 2002 (A-F); from type locality, matured 1. II. 2002 (G-I); from unknown locality, died 22. III. 2012 (J); from unknown locality, killed 30. IX. 2010 (K); from Gunung Belumut (L); from Kota Tinggi, matured 22. X. 2014 (M); same place, matured 20. VIII. 2014 (N); same place, matured 6. XI. 2014 (O). (A) Distal part of palp, ventral view. (B) Paracymbium and tibial apophysis, retroventral view. (C) Palpal organ, proventral view (arrow indicating U-shaped row of denticles at proventral end of distal edge of contrategulum). (D) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (F, H) Same, distal view (dorsal side up). (G) Embolus complex, proventral and slightly distal view. (I) Distal edge of contrategulum, distal and slighly prolateral view (dorsal side to the right). (J-O) Same, distal view. Abbreviations: dw - dorsal wall of sclerotized part of embolus proper; t - tooth on distal edge of contrategulum. Scale lines: 1.0 mm (A-B; C-E, G-H; F, I-O)." httpUri="https://zenodo.org/record/1021949/files/figure.png" pageId="3" pageNumber="394">6E</figureCitation>
); tegulum with coarsely dentate proximal edge (corresponding to dorsal extension of terminal apophysis in Heptathelinae; see
<bibRefCitation id="A00868B5FFE7FFF4BABFF9EF3A2CF9D4" author="Schwendinger P. J. &amp; Ono H." box="[208,564,1566,1591]" journalOrPublisher="Revue suisse de Zoologie" pageId="3" pageNumber="394" pagination="599 - 637" part="118" refString="Schwendinger P. J., Ono H. 2011. On two Heptathela species from southern Vietnam, with a discussion of copulatory organs and systematics of the Liphistiidae (Araneae; Mesothelae). Revue suisse de Zoologie 118 (4): 599 - 637." title="On two Heptathela species from southern Vietnam, with a discussion of copulatory organs and systematics of the Liphistiidae (Araneae; Mesothelae)" type="journal article" year="2011">Schwendinger &amp; Ono, 2011</bibRefCitation>
), distal margin (corresponding to marginal apophysis) indistinct, not drawn into a prominent edge (
<figureCitation id="5CA209C1FFE7FFF4B844F9AE3A6DF99B" box="[555,629,1631,1656]" captionStart="Fig. 4" captionStartId="6.[151,186,1713,1734]" captionTargetBox="[156,1432,265,1656]" captionTargetId="figure@6.[148,1439,254,1693]" captionTargetPageId="6" captionText="Fig. 4. Liphistius malayanus, details of palp of four males: Templer Park (A, C-D, H-I); holotype of L. malayanus cameroni (B, E, G); Cameron Highlands, ZRC (F, J), Fraser’s Hill, MHNG (K-L). (A) Paracymbium and tibial apophysis of left palp, retroventral view. (B) Tibial apophysis of right palp, ventral view. (C) Distal part of left palpal organ, proventral view (arrow indicating V-shaped row of denticles at proventral end of distal edge of contrategulum). (D, F) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (G-H) Left palpal organ, distal view (dorsal side up). (I, K) Distal edge of contrategulum of right palp, distal view (dorsal side to the left). (J) Same, distal and slighly prolateral view. (L) Same of left palp, distal and slighly prolateral view (dorsal side to the right). Abbreviations: a - dorsal apex of contrategulum; de - distal edge of contrategulum. Scale lines: 1.0 mm (A; B; C-D; E, G; F; H-L)." httpUri="https://zenodo.org/record/1021945/files/figure.png" pageId="3" pageNumber="394">Figs 4</figureCitation>
D-F, 6D-E, 8E); contrategulum with indistinct ventral process or without, distal edge carrying several teeth, prolateral ones numerous, tiny and forming a serrate crest (
<taxonomicName id="03996EC7FFE7FFF4BACFF9133904F91B" authority="Peter J. Schwendinger, 2017" authorityName="Peter J. Schwendinger" authorityYear="2017" box="[160,284,1761,1785]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="gracilis" status="sp. nov.">
<emphasis id="F6EDC956FFE7FFF4BACFF9133904F91B" box="[160,284,1761,1785]" italics="true" pageId="3" pageNumber="394">L. gracilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EDDE742DFFE7FFF4BB41F9133989F91A" box="[302,401,1762,1785]" pageId="3" pageNumber="394" rank="species">sp. nov.</taxonomicNameLabel>
;
<figureCitation id="5CA209C1FFE7FFF4BBC8F91039E9F919" box="[423,497,1761,1786]" captionStart="Fig. 8" captionStartId="15.[151,186,1862,1883]" captionTargetBox="[159,1417,261,1808]" captionTargetId="figure@15.[148,1439,254,1832]" captionTargetPageId="15" captionText="Fig. 8. Liphistius gracilis sp. nov., details of palp of three males: paratype, matured 5. I. 2002 (A-B, H); holotype (C-F, I-J); paratype, matured 19. IX. 2001 (G). (A) Distal part of palp, ventral view. (B) Same, retrolateral view. (C) Subtegulum, cymbium and distal part of palpal tibia, ventral view. (D) Distal part of cymbium and palpal organ, proventral view (arrow indicating U-shaped row of denticles at proventral end of distal edge of contrategulum). (E) Same, retrodorsal view. (F-G) Same, distal view (dorsal side up). (H-I) Right tibial apophysis, retrolateral and slightly proximal view. (J) Left tibial apophysis, same view. Scale lines: 1.0 mm (A-C), 0.5 mm (D-G; H-J)." httpUri="https://zenodo.org/record/1021953/files/figure.png" pageId="3" pageNumber="394">Fig. 8</figureCitation>
D-G), few and small (
<taxonomicName id="03996EC7FFE7FFF4BACFF8F2392FF8FA" box="[160,311,1794,1818]" class="Arachnida" family="Liphistiidae" genus="Lipistius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="malayanus">
<emphasis id="F6EDC956FFE7FFF4BACFF8F2392FF8FA" box="[160,311,1794,1818]" italics="true" pageId="3" pageNumber="394">L. malayanus</emphasis>
</taxonomicName>
;
<figureCitation id="5CA209C1FFE7FFF4BB24F8F33988F8F9" box="[331,400,1794,1818]" captionStart="Fig. 4" captionStartId="6.[151,186,1713,1734]" captionTargetBox="[156,1432,265,1656]" captionTargetId="figure@6.[148,1439,254,1693]" captionTargetPageId="6" captionText="Fig. 4. Liphistius malayanus, details of palp of four males: Templer Park (A, C-D, H-I); holotype of L. malayanus cameroni (B, E, G); Cameron Highlands, ZRC (F, J), Fraser’s Hill, MHNG (K-L). (A) Paracymbium and tibial apophysis of left palp, retroventral view. (B) Tibial apophysis of right palp, ventral view. (C) Distal part of left palpal organ, proventral view (arrow indicating V-shaped row of denticles at proventral end of distal edge of contrategulum). (D, F) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (G-H) Left palpal organ, distal view (dorsal side up). (I, K) Distal edge of contrategulum of right palp, distal view (dorsal side to the left). (J) Same, distal and slighly prolateral view. (L) Same of left palp, distal and slighly prolateral view (dorsal side to the right). Abbreviations: a - dorsal apex of contrategulum; de - distal edge of contrategulum. Scale lines: 1.0 mm (A; B; C-D; E, G; F; H-L)." httpUri="https://zenodo.org/record/1021945/files/figure.png" pageId="3" pageNumber="394">Fig. 4</figureCitation>
G-L), or only represented by a single large triangular tooth (
<taxonomicName id="03996EC7FFE7FFF4BB8FF8D23A5AF8DA" authorityName="Sedgwick &amp; Platnick" authorityYear="1987" box="[480,578,1826,1850]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="endau">
<emphasis id="F6EDC956FFE7FFF4BB8FF8D23A5AF8DA" box="[480,578,1826,1850]" italics="true" pageId="3" pageNumber="394">L. endau</emphasis>
</taxonomicName>
;
<figureCitation id="5CA209C1FFE7FFF4B83BF8D33ABEF8DA" box="[596,678,1826,1849]" captionStart="Fig. 6" captionStartId="11.[151,186,1824,1845]" captionTargetBox="[232,1351,260,1775]" captionTargetId="figure@11.[215,1371,254,1797]" captionTargetPageId="11" captionText="Fig. 6. Liphistius endau, details of left palp of eight males: from type locality, matured 27. II. 2002 (A-F); from type locality, matured 1. II. 2002 (G-I); from unknown locality, died 22. III. 2012 (J); from unknown locality, killed 30. IX. 2010 (K); from Gunung Belumut (L); from Kota Tinggi, matured 22. X. 2014 (M); same place, matured 20. VIII. 2014 (N); same place, matured 6. XI. 2014 (O). (A) Distal part of palp, ventral view. (B) Paracymbium and tibial apophysis, retroventral view. (C) Palpal organ, proventral view (arrow indicating U-shaped row of denticles at proventral end of distal edge of contrategulum). (D) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (F, H) Same, distal view (dorsal side up). (G) Embolus complex, proventral and slightly distal view. (I) Distal edge of contrategulum, distal and slighly prolateral view (dorsal side to the right). (J-O) Same, distal view. Abbreviations: dw - dorsal wall of sclerotized part of embolus proper; t - tooth on distal edge of contrategulum. Scale lines: 1.0 mm (A-B; C-E, G-H; F, I-O)." httpUri="https://zenodo.org/record/1021949/files/figure.png" pageId="3" pageNumber="394">Fig. 6F</figureCitation>
, H-O), proventral denticles of distal edge forming a downwardcurved V-shaped (
<figureCitation id="5CA209C1FFE7FFF4BB01F89239D4F89F" box="[366,460,1891,1916]" captionStart="Fig. 4" captionStartId="6.[151,186,1713,1734]" captionTargetBox="[156,1432,265,1656]" captionTargetId="figure@6.[148,1439,254,1693]" captionTargetPageId="6" captionText="Fig. 4. Liphistius malayanus, details of palp of four males: Templer Park (A, C-D, H-I); holotype of L. malayanus cameroni (B, E, G); Cameron Highlands, ZRC (F, J), Fraser’s Hill, MHNG (K-L). (A) Paracymbium and tibial apophysis of left palp, retroventral view. (B) Tibial apophysis of right palp, ventral view. (C) Distal part of left palpal organ, proventral view (arrow indicating V-shaped row of denticles at proventral end of distal edge of contrategulum). (D, F) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (G-H) Left palpal organ, distal view (dorsal side up). (I, K) Distal edge of contrategulum of right palp, distal view (dorsal side to the left). (J) Same, distal and slighly prolateral view. (L) Same of left palp, distal and slighly prolateral view (dorsal side to the right). Abbreviations: a - dorsal apex of contrategulum; de - distal edge of contrategulum. Scale lines: 1.0 mm (A; B; C-D; E, G; F; H-L)." httpUri="https://zenodo.org/record/1021945/files/figure.png" pageId="3" pageNumber="394">Fig. 4C</figureCitation>
) or U-shaped row (
<figureCitation id="5CA209C1FFE7FFF4B8ABF89238F9F87F" captionStart="Fig. 6" captionStartId="11.[151,186,1824,1845]" captionTargetBox="[232,1351,260,1775]" captionTargetId="figure@11.[215,1371,254,1797]" captionTargetPageId="11" captionText="Fig. 6. Liphistius endau, details of left palp of eight males: from type locality, matured 27. II. 2002 (A-F); from type locality, matured 1. II. 2002 (G-I); from unknown locality, died 22. III. 2012 (J); from unknown locality, killed 30. IX. 2010 (K); from Gunung Belumut (L); from Kota Tinggi, matured 22. X. 2014 (M); same place, matured 20. VIII. 2014 (N); same place, matured 6. XI. 2014 (O). (A) Distal part of palp, ventral view. (B) Paracymbium and tibial apophysis, retroventral view. (C) Palpal organ, proventral view (arrow indicating U-shaped row of denticles at proventral end of distal edge of contrategulum). (D) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (F, H) Same, distal view (dorsal side up). (G) Embolus complex, proventral and slightly distal view. (I) Distal edge of contrategulum, distal and slighly prolateral view (dorsal side to the right). (J-O) Same, distal view. Abbreviations: dw - dorsal wall of sclerotized part of embolus proper; t - tooth on distal edge of contrategulum. Scale lines: 1.0 mm (A-B; C-E, G-H; F, I-O)." httpUri="https://zenodo.org/record/1021949/files/figure.png" pageId="3" pageNumber="394">Figs 6A, C</figureCitation>
; 8D), dorsal apex of contrategulum widely to narrowly tongue-shaped (
<figureCitation id="5CA209C1FFE7FFF4BBDCF85539E5F85E" box="[435,509,1956,1981]" captionStart="Fig. 4" captionStartId="6.[151,186,1713,1734]" captionTargetBox="[156,1432,265,1656]" captionTargetId="figure@6.[148,1439,254,1693]" captionTargetPageId="6" captionText="Fig. 4. Liphistius malayanus, details of palp of four males: Templer Park (A, C-D, H-I); holotype of L. malayanus cameroni (B, E, G); Cameron Highlands, ZRC (F, J), Fraser’s Hill, MHNG (K-L). (A) Paracymbium and tibial apophysis of left palp, retroventral view. (B) Tibial apophysis of right palp, ventral view. (C) Distal part of left palpal organ, proventral view (arrow indicating V-shaped row of denticles at proventral end of distal edge of contrategulum). (D, F) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (G-H) Left palpal organ, distal view (dorsal side up). (I, K) Distal edge of contrategulum of right palp, distal view (dorsal side to the left). (J) Same, distal and slighly prolateral view. (L) Same of left palp, distal and slighly prolateral view (dorsal side to the right). Abbreviations: a - dorsal apex of contrategulum; de - distal edge of contrategulum. Scale lines: 1.0 mm (A; B; C-D; E, G; F; H-L)." httpUri="https://zenodo.org/record/1021945/files/figure.png" pageId="3" pageNumber="394">Figs 4</figureCitation>
G-L; 6F, H-O; 8F-G); subtegulum always without apophysis; paracymbium quite small and shallow to large and deep (
<figureCitation id="5CA209C1FFE7FFF4B8FFF8143AE8F81D" box="[656,752,2021,2046]" captionStart="Fig. 4" captionStartId="6.[151,186,1713,1734]" captionTargetBox="[156,1432,265,1656]" captionTargetId="figure@6.[148,1439,254,1693]" captionTargetPageId="6" captionText="Fig. 4. Liphistius malayanus, details of palp of four males: Templer Park (A, C-D, H-I); holotype of L. malayanus cameroni (B, E, G); Cameron Highlands, ZRC (F, J), Fraser’s Hill, MHNG (K-L). (A) Paracymbium and tibial apophysis of left palp, retroventral view. (B) Tibial apophysis of right palp, ventral view. (C) Distal part of left palpal organ, proventral view (arrow indicating V-shaped row of denticles at proventral end of distal edge of contrategulum). (D, F) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (G-H) Left palpal organ, distal view (dorsal side up). (I, K) Distal edge of contrategulum of right palp, distal view (dorsal side to the left). (J) Same, distal and slighly prolateral view. (L) Same of left palp, distal and slighly prolateral view (dorsal side to the right). Abbreviations: a - dorsal apex of contrategulum; de - distal edge of contrategulum. Scale lines: 1.0 mm (A; B; C-D; E, G; F; H-L)." httpUri="https://zenodo.org/record/1021945/files/figure.png" pageId="3" pageNumber="394">Figs 4A</figureCitation>
,
<figureCitation id="5CA209C1FFE7FFF4BAF8F7F738A0F7FE" box="[151,184,2054,2077]" captionStart="Fig. 6" captionStartId="11.[151,186,1824,1845]" captionTargetBox="[232,1351,260,1775]" captionTargetId="figure@11.[215,1371,254,1797]" captionTargetPageId="11" captionText="Fig. 6. Liphistius endau, details of left palp of eight males: from type locality, matured 27. II. 2002 (A-F); from type locality, matured 1. II. 2002 (G-I); from unknown locality, died 22. III. 2012 (J); from unknown locality, killed 30. IX. 2010 (K); from Gunung Belumut (L); from Kota Tinggi, matured 22. X. 2014 (M); same place, matured 20. VIII. 2014 (N); same place, matured 6. XI. 2014 (O). (A) Distal part of palp, ventral view. (B) Paracymbium and tibial apophysis, retroventral view. (C) Palpal organ, proventral view (arrow indicating U-shaped row of denticles at proventral end of distal edge of contrategulum). (D) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (F, H) Same, distal view (dorsal side up). (G) Embolus complex, proventral and slightly distal view. (I) Distal edge of contrategulum, distal and slighly prolateral view (dorsal side to the right). (J-O) Same, distal view. Abbreviations: dw - dorsal wall of sclerotized part of embolus proper; t - tooth on distal edge of contrategulum. Scale lines: 1.0 mm (A-B; C-E, G-H; F, I-O)." httpUri="https://zenodo.org/record/1021949/files/figure.png" pageId="3" pageNumber="394">6B</figureCitation>
,
<figureCitation id="5CA209C1FFE7FFF4BAA9F7F738F0F7FD" box="[198,232,2054,2078]" captionStart="Fig. 8" captionStartId="15.[151,186,1862,1883]" captionTargetBox="[159,1417,261,1808]" captionTargetId="figure@15.[148,1439,254,1832]" captionTargetPageId="15" captionText="Fig. 8. Liphistius gracilis sp. nov., details of palp of three males: paratype, matured 5. I. 2002 (A-B, H); holotype (C-F, I-J); paratype, matured 19. IX. 2001 (G). (A) Distal part of palp, ventral view. (B) Same, retrolateral view. (C) Subtegulum, cymbium and distal part of palpal tibia, ventral view. (D) Distal part of cymbium and palpal organ, proventral view (arrow indicating U-shaped row of denticles at proventral end of distal edge of contrategulum). (E) Same, retrodorsal view. (F-G) Same, distal view (dorsal side up). (H-I) Right tibial apophysis, retrolateral and slightly proximal view. (J) Left tibial apophysis, same view. Scale lines: 1.0 mm (A-C), 0.5 mm (D-G; H-J)." httpUri="https://zenodo.org/record/1021953/files/figure.png" pageId="3" pageNumber="394">8B</figureCitation>
); cumulus very low, indistinct, carrying several long, thick bristles; retrolateral tibial apophysis relatively short and carrying long apical megaspines (
<taxonomicName id="03996EC7FFE7FFF4B926FBA93BFAFB8D" box="[841,994,1111,1135]" class="Arachnida" family="Liphistiidae" genus="Lipistius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="malayanus">
<emphasis id="F6EDC956FFE7FFF4B926FBA93BFAFB8D" box="[841,994,1111,1135]" italics="true" pageId="3" pageNumber="394">L. malayanus</emphasis>
</taxonomicName>
and
<taxonomicName id="03996EC7FFE7FFF4BE48FBA93C95FB8D" authorityName="Sedgwick &amp; Platnick" authorityYear="1987" box="[1063,1165,1111,1135]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="endau">
<emphasis id="F6EDC956FFE7FFF4BE48FBA93C95FB8D" box="[1063,1165,1111,1135]" italics="true" pageId="3" pageNumber="394">L. endau</emphasis>
</taxonomicName>
;
<figureCitation id="5CA209C1FFE7FFF4BECCFBA63CF7FB93" box="[1187,1263,1111,1136]" captionStart="Fig. 4" captionStartId="6.[151,186,1713,1734]" captionTargetBox="[156,1432,265,1656]" captionTargetId="figure@6.[148,1439,254,1693]" captionTargetPageId="6" captionText="Fig. 4. Liphistius malayanus, details of palp of four males: Templer Park (A, C-D, H-I); holotype of L. malayanus cameroni (B, E, G); Cameron Highlands, ZRC (F, J), Fraser’s Hill, MHNG (K-L). (A) Paracymbium and tibial apophysis of left palp, retroventral view. (B) Tibial apophysis of right palp, ventral view. (C) Distal part of left palpal organ, proventral view (arrow indicating V-shaped row of denticles at proventral end of distal edge of contrategulum). (D, F) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (G-H) Left palpal organ, distal view (dorsal side up). (I, K) Distal edge of contrategulum of right palp, distal view (dorsal side to the left). (J) Same, distal and slighly prolateral view. (L) Same of left palp, distal and slighly prolateral view (dorsal side to the right). Abbreviations: a - dorsal apex of contrategulum; de - distal edge of contrategulum. Scale lines: 1.0 mm (A; B; C-D; E, G; F; H-L)." httpUri="https://zenodo.org/record/1021945/files/figure.png" pageId="3" pageNumber="394">Figs 4</figureCitation>
A-B, 6A-B) or quite long and carrying distinctly shorter megaspines (
<taxonomicName id="03996EC7FFE7FFF4B927FB6B3BA2FB4C" authority="Peter J. Schwendinger, 2017" authorityName="Peter J. Schwendinger" authorityYear="2017" box="[840,954,1176,1201]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="gracilis" status="sp. nov.">
<emphasis id="F6EDC956FFE7FFF4B927FB6B3BA2FB4C" box="[840,954,1176,1201]" italics="true" pageId="3" pageNumber="394">L. gracilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EDDE742DFFE7FFF4B9ACFB683C04FB53" box="[963,1052,1177,1200]" pageId="3" pageNumber="394" rank="species">sp. nov.</taxonomicNameLabel>
;
<figureCitation id="5CA209C1FFE7FFF4BE45FB693C72FB52" box="[1066,1130,1176,1201]" captionStart="Fig. 8" captionStartId="15.[151,186,1862,1883]" captionTargetBox="[159,1417,261,1808]" captionTargetId="figure@15.[148,1439,254,1832]" captionTargetPageId="15" captionText="Fig. 8. Liphistius gracilis sp. nov., details of palp of three males: paratype, matured 5. I. 2002 (A-B, H); holotype (C-F, I-J); paratype, matured 19. IX. 2001 (G). (A) Distal part of palp, ventral view. (B) Same, retrolateral view. (C) Subtegulum, cymbium and distal part of palpal tibia, ventral view. (D) Distal part of cymbium and palpal organ, proventral view (arrow indicating U-shaped row of denticles at proventral end of distal edge of contrategulum). (E) Same, retrodorsal view. (F-G) Same, distal view (dorsal side up). (H-I) Right tibial apophysis, retrolateral and slightly proximal view. (J) Left tibial apophysis, same view. Scale lines: 1.0 mm (A-C), 0.5 mm (D-G; H-J)." httpUri="https://zenodo.org/record/1021953/files/figure.png" pageId="3" pageNumber="394">Fig. 8</figureCitation>
A-C, I-J); both apical lobes of cymbium weakly developed (
<figureCitation id="5CA209C1FFE7FFF4BEB9FB483D23FB31" box="[1238,1339,1209,1234]" captionStart="Fig. 8" captionStartId="15.[151,186,1862,1883]" captionTargetBox="[159,1417,261,1808]" captionTargetId="figure@15.[148,1439,254,1832]" captionTargetPageId="15" captionText="Fig. 8. Liphistius gracilis sp. nov., details of palp of three males: paratype, matured 5. I. 2002 (A-B, H); holotype (C-F, I-J); paratype, matured 19. IX. 2001 (G). (A) Distal part of palp, ventral view. (B) Same, retrolateral view. (C) Subtegulum, cymbium and distal part of palpal tibia, ventral view. (D) Distal part of cymbium and palpal organ, proventral view (arrow indicating U-shaped row of denticles at proventral end of distal edge of contrategulum). (E) Same, retrodorsal view. (F-G) Same, distal view (dorsal side up). (H-I) Right tibial apophysis, retrolateral and slightly proximal view. (J) Left tibial apophysis, same view. Scale lines: 1.0 mm (A-C), 0.5 mm (D-G; H-J)." httpUri="https://zenodo.org/record/1021953/files/figure.png" pageId="3" pageNumber="394">Fig. 8D</figureCitation>
). Vulva (
<figureCitation id="5CA209C1FFE7FFF4B928FB283B89FB13" box="[839,913,1241,1264]" captionStart="Fig. 5" captionStartId="8.[151,186,1809,1830]" captionTargetBox="[154,1439,260,1762]" captionTargetId="figure@8.[148,1439,254,1779]" captionTargetPageId="8" captionText="Fig. 5. Liphistius malayanus, vulval plate of eight females: paratype of L. m. cameroni from Berinchang, Cameron Highlands (A- B); exuvia of female from Ringlet, Cameron Highlands (C); female (SMF 64093) from Fraser’s Hill (D); female (moulted 24. VI. 2002) from Fraser’s Hill; poreplate slightly deformed by brief immersion in cold KOH (E); female (SMF 56206) from Fraser’s Hill (F); female (SMF 60037) from Genting Highlands (G); female (moulted 8. XII. 2005) from Genting Highlands (H); exuvia of female from Templer Park (I-J). (A, C-I) Dorsal view. (B, J) Ventral view. Scale lines: 1.0 mm (A-B; C, F; D; E, G; H; I-J)." httpUri="https://zenodo.org/record/1021947/files/figure.png" pageId="3" pageNumber="394">Figs 5</figureCitation>
,
<figureCitation id="5CA209C1FFE7FFF4B9CFFB2A3BB7FB11" box="[928,943,1243,1266]" captionStart="Fig. 7" captionStartId="12.[151,186,1940,1961]" captionTargetBox="[159,1415,617,1889]" captionTargetId="figure@12.[151,1436,609,1911]" captionTargetPageId="12" captionText="Fig. 7. Liphistius endau, vulval plate of six females: adult from unknown locality (A-B); largest female from Kota Tinggi (C); exuvia of reproductive female (moulted 27. IV. 2002) from type locality (D-E); exuvia of reproductive female (moulted 28. IV. 2002) from type locality (F-G); medium-sized juvenile from type locality (H); small juvenile from type locality (I). (A, C-D, F, H-I) Entire vulval plate, dorsal view. (B) Poreplate, ventral view. (E, G) Entire vulval plate, ventral view. Scale lines: 1.0 mm (A-B; C; D-E; F-G; H-I)." httpUri="https://zenodo.org/record/1021951/files/figure.png" pageId="3" pageNumber="394">7</figureCitation>
,
<figureCitation id="5CA209C1FFE7FFF4B9D0FB2B3BD6FB11" box="[959,974,1242,1266]" captionStart="Fig. 9" captionStartId="16.[151,186,1607,1628]" captionTargetBox="[157,1426,257,1572]" captionTargetId="figure@16.[148,1439,254,1598]" captionTargetPageId="16" captionText="Fig. 9. Liphistius gracilis sp. nov., vulval plate of three females (all from exuviae): allotype (A-B); paratype, moult 31. VIII. 2001 (C- D); paratype, moult 27. IX. 2001 (E-G). (A, C, E) Vulval plate, dorsal view. (B, D, F) Same, ventral view. (G) Same, anterior view. Scale lines: 1.0 mm (A-B; C-G)." httpUri="https://zenodo.org/record/1021955/files/figure.png" pageId="3" pageNumber="394">9</figureCitation>
,
<figureCitation id="5CA209C1FFE7FFF4B9B1FB2B3C09FB11" box="[990,1041,1242,1266]" captionStart="Fig. 10" captionStartId="18.[831,866,1881,1902]" captionTargetBox="[838,1435,1514,1833]" captionTargetId="figure@18.[829,1439,1504,1842]" captionTargetPageId="18" captionText="Fig. 10. Liphistius johore, female holotype (A); Liphistius panching, male from type locality (B). (A) Dorsal view of vulval plate (redrawn from Platnick &amp; Sedgwick, 1984: fig. 79) (arrows indicating anterolateral invaginations in margin of poreplate). (B) Distal view of left palpal organ (dorsal side up) (redrawn from Sedgwick &amp; Platnick, 1986: fig. 5)." httpUri="https://zenodo.org/record/1021957/files/figure.png" pageId="3" pageNumber="394">10A</figureCitation>
): poreplate strongly pigmented and sclerotised, wider than long, its anterior margin more or less distinctly recurved, with or without anterolateral lobes or processes, never with mediolateral processes; CDO and ventral receptacular cluster large to very large, both undivided; posterior stalk wide (in
<taxonomicName id="03996EC7FFE7FFF4BF52FA8F3D84FA70" authorityName="Sedgwick &amp; Platnick" authorityYear="1987" box="[1341,1436,1404,1429]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="endau">
<emphasis id="F6EDC956FFE7FFF4BF52FA8F3D84FA70" box="[1341,1436,1404,1429]" italics="true" pageId="3" pageNumber="394">L. endau</emphasis>
</taxonomicName>
even covering entire width of genital atrium;
<figureCitation id="5CA209C1FFE7FFF4BF26FA6C3D96FA55" box="[1353,1422,1437,1462]" captionStart="Fig. 7" captionStartId="12.[151,186,1940,1961]" captionTargetBox="[159,1415,617,1889]" captionTargetId="figure@12.[151,1436,609,1911]" captionTargetPageId="12" captionText="Fig. 7. Liphistius endau, vulval plate of six females: adult from unknown locality (A-B); largest female from Kota Tinggi (C); exuvia of reproductive female (moulted 27. IV. 2002) from type locality (D-E); exuvia of reproductive female (moulted 28. IV. 2002) from type locality (F-G); medium-sized juvenile from type locality (H); small juvenile from type locality (I). (A, C-D, F, H-I) Entire vulval plate, dorsal view. (B) Poreplate, ventral view. (E, G) Entire vulval plate, ventral view. Scale lines: 1.0 mm (A-B; C; D-E; F-G; H-I)." httpUri="https://zenodo.org/record/1021951/files/figure.png" pageId="3" pageNumber="394">Fig. 7</figureCitation>
), somewhat trapezium-shaped or almost triangular (
<figureCitation id="5CA209C1FFE7FFF4BF1EFA4C3B69FA14" captionStart="Fig. 10" captionStartId="18.[831,866,1881,1902]" captionTargetBox="[838,1435,1514,1833]" captionTargetId="figure@18.[829,1439,1504,1842]" captionTargetPageId="18" captionText="Fig. 10. Liphistius johore, female holotype (A); Liphistius panching, male from type locality (B). (A) Dorsal view of vulval plate (redrawn from Platnick &amp; Sedgwick, 1984: fig. 79) (arrows indicating anterolateral invaginations in margin of poreplate). (B) Distal view of left palpal organ (dorsal side up) (redrawn from Sedgwick &amp; Platnick, 1986: fig. 5)." httpUri="https://zenodo.org/record/1021957/files/figure.png" pageId="3" pageNumber="394">Fig. 10A</figureCitation>
); genital atrium with few to many lateral hairs and with no, few or numerous median hairs.
</paragraph>
</subSubSection>
<subSubSection id="8C8346CFFFE7FFF4B950F9DF3BF9F96B" pageId="3" pageNumber="394" type="synonymic_list">
<paragraph id="C4261544FFE7FFF4B950F9DF3BF9F96B" blockId="3.[831,1436,1582,1672]" pageId="3" pageNumber="394">
<emphasis id="F6EDC956FFE7FFF4B950F9DF3C0AF9A6" bold="true" box="[831,1042,1582,1605]" pageId="3" pageNumber="394">Species included:</emphasis>
<emphasis id="F6EDC956FFE7FFF4BE4CF9DF3BB6F985" italics="true" pageId="3" pageNumber="394">
<taxonomicName id="03996EC7FFE7FFF4BE4CF9DF3CFCF9A6" authorityName="Sedgwick &amp; Platnick" authorityYear="1987" box="[1059,1252,1582,1605]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="endau">Liphistius endau</taxonomicName>
,
<taxonomicName id="03996EC7FFE7FFF4BE96F9C13D80F9A6" box="[1273,1432,1582,1607]" class="Arachnida" family="Liphistiidae" genus="Lipistius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="malayanus">L. malayanus</taxonomicName>
,
<taxonomicName id="03996EC7FFE7FFF4B950F9A13BB6F985" authority="Peter J. Schwendinger, 2017" authorityName="Peter J. Schwendinger" authorityYear="2017" box="[831,942,1615,1639]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="gracilis" status="sp. nov.">L. gracilis</taxonomicName>
</emphasis>
<taxonomicNameLabel id="EDDE742DFFE7FFF4B9D9F9A13C12F984" box="[950,1034,1616,1639]" pageId="3" pageNumber="394" rank="species">sp. nov.</taxonomicNameLabel>
and presumably also
<taxonomicName id="03996EC7FFE7FFF4BE99F9A13D40F985" authorityName="Platnick &amp; Sedgwick" authorityYear="1984" box="[1270,1368,1615,1639]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="johore">
<emphasis id="F6EDC956FFE7FFF4BE99F9A13D40F985" box="[1270,1368,1615,1639]" italics="true" pageId="3" pageNumber="394">L. johore</emphasis>
</taxonomicName>
(male still unknown).
</paragraph>
</subSubSection>
<subSubSection id="8C8346CFFFE7FFF4B950F96E3BA3F7FE" pageId="3" pageNumber="394" type="discussion">
<paragraph id="C4261544FFE7FFF4B950F96E3BA3F7FE" blockId="3.[831,1437,1695,2077]" pageId="3" pageNumber="394">
<emphasis id="F6EDC956FFE7FFF4B950F96E3BFDF955" bold="true" box="[831,997,1695,1718]" pageId="3" pageNumber="394">Relationships:</emphasis>
Species of the
<taxonomicName id="03996EC7FFE7FFF4BEC1F96E3D3BF955" box="[1198,1315,1695,1718]" class="Arachnida" family="Liphistiidae" genus="Lipistius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="malayanus">
<emphasis id="F6EDC956FFE7FFF4BEC1F96E3D3BF955" box="[1198,1315,1695,1718]" italics="true" pageId="3" pageNumber="394">malayanus</emphasis>
</taxonomicName>
-group are most similar and probably closely related to species of the
<taxonomicName id="03996EC7FFE7FFF4B906F9133BABF91A" authorityName="Platnick &amp; Sedgwick" authorityYear="1984" box="[873,947,1762,1785]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="tioman">
<emphasis id="F6EDC956FFE7FFF4B906F9133BABF91A" box="[873,947,1762,1785]" italics="true" pageId="3" pageNumber="394">tioman</emphasis>
</taxonomicName>
-group. The two large species,
<taxonomicName id="03996EC7FFE7FFF4BF65F9133D85F914" box="[1290,1437,1760,1785]" class="Arachnida" family="Liphistiidae" genus="Lipistius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="malayanus">
<emphasis id="F6EDC956FFE7FFF4BF65F9133D85F914" box="[1290,1437,1760,1785]" italics="true" pageId="3" pageNumber="394">L. malayanus</emphasis>
</taxonomicName>
and
<emphasis id="F6EDC956FFE7FFF4B91AF8F33BFAF8FB" box="[885,994,1793,1817]" italics="true" pageId="3" pageNumber="394">
<taxonomicName id="03996EC7FFE7FFF4B91AF8F33BC7F8FB" authorityName="Sedgwick &amp; Platnick" authorityYear="1987" box="[885,991,1793,1817]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="endau">L. endau</taxonomicName>
,
</emphasis>
are closer to each other than to the much smaller
<taxonomicName id="03996EC7FFE7FFF4B986F8D23C78F8DA" authority="Peter J. Schwendinger, 2017" authorityName="Peter J. Schwendinger" authorityYear="2017" box="[1001,1120,1826,1850]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="gracilis" status="sp. nov.">
<emphasis id="F6EDC956FFE7FFF4B986F8D23C78F8DA" box="[1001,1120,1826,1850]" italics="true" pageId="3" pageNumber="394">L. gracilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EDDE742DFFE7FFF4BE00F8D23CD3F8D9" box="[1135,1227,1827,1850]" pageId="3" pageNumber="394" rank="species">sp. nov.</taxonomicNameLabel>
(distinguished by serrate distal edge of contrategulum and by sharp bend in wide dorsal wall of sclerotised part of embolus proper). Small size, details of the vulval plate and geographical proximity suggest a close relationship between
<taxonomicName id="03996EC7FFE7FFF4B9CDF8373C09F838" authority="Peter J. Schwendinger, 2017" authorityName="Peter J. Schwendinger" authorityYear="2017" box="[930,1041,1988,2013]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="gracilis" status="sp. nov.">
<emphasis id="F6EDC956FFE7FFF4B9CDF8373C09F838" box="[930,1041,1988,2013]" italics="true" pageId="3" pageNumber="394">L. gracilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EDDE742DFFE7FFF4BE74F8373C68F83E" box="[1051,1136,1990,2013]" pageId="3" pageNumber="394" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="03996EC7FFE7FFF4BEC6F8373D15F838" authorityName="Platnick &amp; Sedgwick" authorityYear="1984" box="[1193,1293,1988,2013]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="johore">
<emphasis id="F6EDC956FFE7FFF4BEC6F8373D15F838" box="[1193,1293,1988,2013]" italics="true" pageId="3" pageNumber="394">L. johore</emphasis>
</taxonomicName>
, but this can only be confirmed when the male of the latter species is discovered.
</paragraph>
</subSubSection>
</subSection>
<subSection id="AD160CAFFFE0FFF3BAF8FEF33975FB0E" pageId="4" pageNumber="395" type="key">
<subSubSection id="8C8346CFFFE0FFF3BAF8FEF33D8FFBEB" pageId="4" pageNumber="395" type="key">
<paragraph id="C4261544FFE0FFF3BAF8FEF33A65FEF9" blockId="4.[151,637,257,282]" box="[151,637,257,282]" pageId="4" pageNumber="395">
<heading id="9F6EA228FFE0FFF3BAF8FEF33A65FEF9" bold="true" box="[151,637,257,282]" fontSize="10" level="2" pageId="4" pageNumber="395" reason="2">
<emphasis id="F6EDC956FFE0FFF3BAF8FEF33A65FEF9" bold="true" box="[151,637,257,282]" pageId="4" pageNumber="395">
Key to the species of the
<taxonomicName id="03996EC7FFE0FFF3BBC0FEF03A3FFEFB" box="[431,551,257,280]" class="Arachnida" family="Liphistiidae" genus="Lipistius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="395" phylum="Arthropoda" rank="species" species="malayanus">
<emphasis id="F6EDC956FFE0FFF3BBC0FEF03A3FFEFB" bold="true" box="[431,551,257,280]" italics="true" pageId="4" pageNumber="395">malayanus</emphasis>
</taxonomicName>
-group:
</emphasis>
</heading>
</paragraph>
<paragraph id="C4261544FFE0FFF3BAF8FEC03D8FFE6A" blockId="4.[151,1437,305,1033]" pageId="4" pageNumber="395">
1 Small spiders (carapace length of mature males and females 4.36-5.34, carapace width 3.64-4.22) with brown body colouration and annulated limbs; lateral and posterolateral margins on ventral side of poreplate not bulging (
<figureCitation id="5CA209C1FFE0FFF3BF1EFEA0395AFE6A" captionStart="Fig. 9" captionStartId="16.[151,186,1607,1628]" captionTargetBox="[157,1426,257,1572]" captionTargetId="figure@16.[148,1439,254,1598]" captionTargetPageId="16" captionText="Fig. 9. Liphistius gracilis sp. nov., vulval plate of three females (all from exuviae): allotype (A-B); paratype, moult 31. VIII. 2001 (C- D); paratype, moult 27. IX. 2001 (E-G). (A, C, E) Vulval plate, dorsal view. (B, D, F) Same, ventral view. (G) Same, anterior view. Scale lines: 1.0 mm (A-B; C-G)." httpUri="https://zenodo.org/record/1021955/files/figure.png" pageId="4" pageNumber="395">Fig. 9B, D, F</figureCitation>
;
<bibRefCitation id="A00868B5FFE0FFF3BB22FE803A6EFE6A" author="Platnick N. &amp; Sedgwick W. C." box="[333,630,369,393]" journalOrPublisher="American Museum Novitates" pageId="4" pageNumber="395" pagination="1 - 31" part="2781" refString="Platnick N. I., Sedgwick W. C. 1984. A revision of the spider genus Liphistius (Araneae, Mesothelae). American Museum Novitates 2781: 1 - 31." title="A revision of the spider genus Liphistius (Araneae, Mesothelae)" type="journal article" year="1984">Platnick &amp; Sedgwick, 1984</bibRefCitation>
: fig. 80).........................................................................................................2
</paragraph>
<paragraph id="C4261544FFE0FFF3BAF8FE633D80FE2A" blockId="4.[151,1437,305,1033]" pageId="4" pageNumber="395">
- Large (carapace lengths in males 7.90-12.04, carapace width 7.0-10.99), uniformly dark-coloured spiders; lateral and posterolateral margins of poreplate on ventral side of vulval plate bulging (
<figureCitation id="5CA209C1FFE0FFF3BE40FE403C86FE2A" box="[1071,1182,433,457]" captionStart="Fig. 5" captionStartId="8.[151,186,1809,1830]" captionTargetBox="[154,1439,260,1762]" captionTargetId="figure@8.[148,1439,254,1779]" captionTargetPageId="8" captionText="Fig. 5. Liphistius malayanus, vulval plate of eight females: paratype of L. m. cameroni from Berinchang, Cameron Highlands (A- B); exuvia of female from Ringlet, Cameron Highlands (C); female (SMF 64093) from Fraser’s Hill (D); female (moulted 24. VI. 2002) from Fraser’s Hill; poreplate slightly deformed by brief immersion in cold KOH (E); female (SMF 56206) from Fraser’s Hill (F); female (SMF 60037) from Genting Highlands (G); female (moulted 8. XII. 2005) from Genting Highlands (H); exuvia of female from Templer Park (I-J). (A, C-I) Dorsal view. (B, J) Ventral view. Scale lines: 1.0 mm (A-B; C, F; D; E, G; H; I-J)." httpUri="https://zenodo.org/record/1021947/files/figure.png" pageId="4" pageNumber="395">Figs 5B, J</figureCitation>
; 7B, E, G)..................3
</paragraph>
<paragraph id="C4261544FFE0FFF3BAF8FE203D80FDCA" blockId="4.[151,1437,305,1033]" pageId="4" pageNumber="395">
2 Vulval plate with indistinct transition between poreplate and posterior stalk, the latter wide in transition zone; anterior margin of poreplate without or with only indistinct anterolateral invaginations; genital atrium with numerous lateral and median hairs (
<figureCitation id="5CA209C1FFE0FFF3B80EFDE03AE9FDCA" box="[609,753,529,553]" captionStart="Fig. 9" captionStartId="16.[151,186,1607,1628]" captionTargetBox="[157,1426,257,1572]" captionTargetId="figure@16.[148,1439,254,1598]" captionTargetPageId="16" captionText="Fig. 9. Liphistius gracilis sp. nov., vulval plate of three females (all from exuviae): allotype (A-B); paratype, moult 31. VIII. 2001 (C- D); paratype, moult 27. IX. 2001 (E-G). (A, C, E) Vulval plate, dorsal view. (B, D, F) Same, ventral view. (G) Same, anterior view. Scale lines: 1.0 mm (A-B; C-G)." httpUri="https://zenodo.org/record/1021955/files/figure.png" pageId="4" pageNumber="395">Fig. 9A, C, E</figureCitation>
) ......................................................................
<taxonomicName id="03996EC7FFE0FFF3BEBEFDE33D26FDCB" ID-CoL="72J5N" authority="Peter J. Schwendinger, 2017" authorityName="Peter J. Schwendinger" authorityYear="2017" box="[1233,1342,529,553]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="395" phylum="Arthropoda" rank="species" species="gracilis" status="sp. nov.">
<emphasis id="F6EDC956FFE0FFF3BEBEFDE33D26FDCB" box="[1233,1342,529,553]" italics="true" pageId="4" pageNumber="395">L. gracilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EDDE742DFFE0FFF3BF2AFDE33D80FDCA" box="[1349,1432,530,553]" pageId="4" pageNumber="395" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="C4261544FFE0FFF3BAF8FDC33D80FD6B" blockId="4.[151,1437,305,1033]" pageId="4" pageNumber="395">
- Vulval plate with distinct transition between poreplate and posterior stalk, the latter very narrow in transition zone; anterior margin of poreplate with pronounced anterolateral invaginations; genital atrium without hairs (
<figureCitation id="5CA209C1FFE0FFF3BF5CFDA03D8DFD8A" box="[1331,1429,593,617]" captionStart="Fig. 10" captionStartId="18.[831,866,1881,1902]" captionTargetBox="[838,1435,1514,1833]" captionTargetId="figure@18.[829,1439,1504,1842]" captionTargetPageId="18" captionText="Fig. 10. Liphistius johore, female holotype (A); Liphistius panching, male from type locality (B). (A) Dorsal view of vulval plate (redrawn from Platnick &amp; Sedgwick, 1984: fig. 79) (arrows indicating anterolateral invaginations in margin of poreplate). (B) Distal view of left palpal organ (dorsal side up) (redrawn from Sedgwick &amp; Platnick, 1986: fig. 5)." httpUri="https://zenodo.org/record/1021957/files/figure.png" pageId="4" pageNumber="395">Fig. 10A</figureCitation>
) (male unknown) ...........................................................................................................................................
<taxonomicName id="03996EC7FFE0FFF3BF59FD833D80FD6B" ID-CoL="72J5M" authorityName="Platnick &amp; Sedgwick" authorityYear="1984" box="[1334,1432,625,649]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="395" phylum="Arthropoda" rank="species" species="johore">
<emphasis id="F6EDC956FFE0FFF3BF59FD833D80FD6B" box="[1334,1432,625,649]" italics="true" pageId="4" pageNumber="395">L. johore</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="C4261544FFE0FFF3BAF8FD603D80FCAB" blockId="4.[151,1437,305,1033]" pageId="4" pageNumber="395">
3 Dorsal and ventral wall of sclerotised part of embolus proper equally wide, both ends facing proventrad (
<figureCitation id="5CA209C1FFE0FFF3BF2EFD603D98FD4A" box="[1345,1408,657,681]" captionStart="Fig. 4" captionStartId="6.[151,186,1713,1734]" captionTargetBox="[156,1432,265,1656]" captionTargetId="figure@6.[148,1439,254,1693]" captionTargetPageId="6" captionText="Fig. 4. Liphistius malayanus, details of palp of four males: Templer Park (A, C-D, H-I); holotype of L. malayanus cameroni (B, E, G); Cameron Highlands, ZRC (F, J), Fraser’s Hill, MHNG (K-L). (A) Paracymbium and tibial apophysis of left palp, retroventral view. (B) Tibial apophysis of right palp, ventral view. (C) Distal part of left palpal organ, proventral view (arrow indicating V-shaped row of denticles at proventral end of distal edge of contrategulum). (D, F) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (G-H) Left palpal organ, distal view (dorsal side up). (I, K) Distal edge of contrategulum of right palp, distal view (dorsal side to the left). (J) Same, distal and slighly prolateral view. (L) Same of left palp, distal and slighly prolateral view (dorsal side to the right). Abbreviations: a - dorsal apex of contrategulum; de - distal edge of contrategulum. Scale lines: 1.0 mm (A; B; C-D; E, G; F; H-L)." httpUri="https://zenodo.org/record/1021945/files/figure.png" pageId="4" pageNumber="395">Fig. 4</figureCitation>
G- H), dorsal wall ending in indistinct rounded lobe (
<figureCitation id="5CA209C1FFE0FFF3B89AFD403B2CFD2A" box="[757,820,689,713]" captionStart="Fig. 4" captionStartId="6.[151,186,1713,1734]" captionTargetBox="[156,1432,265,1656]" captionTargetId="figure@6.[148,1439,254,1693]" captionTargetPageId="6" captionText="Fig. 4. Liphistius malayanus, details of palp of four males: Templer Park (A, C-D, H-I); holotype of L. malayanus cameroni (B, E, G); Cameron Highlands, ZRC (F, J), Fraser’s Hill, MHNG (K-L). (A) Paracymbium and tibial apophysis of left palp, retroventral view. (B) Tibial apophysis of right palp, ventral view. (C) Distal part of left palpal organ, proventral view (arrow indicating V-shaped row of denticles at proventral end of distal edge of contrategulum). (D, F) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (G-H) Left palpal organ, distal view (dorsal side up). (I, K) Distal edge of contrategulum of right palp, distal view (dorsal side to the left). (J) Same, distal and slighly prolateral view. (L) Same of left palp, distal and slighly prolateral view (dorsal side to the right). Abbreviations: a - dorsal apex of contrategulum; de - distal edge of contrategulum. Scale lines: 1.0 mm (A; B; C-D; E, G; F; H-L)." httpUri="https://zenodo.org/record/1021945/files/figure.png" pageId="4" pageNumber="395">Fig. 4</figureCitation>
C-F); membranous part of embolus proper distally narrow (
<figureCitation id="5CA209C1FFE0FFF3BA85FD203978FD0A" box="[234,352,721,745]" captionStart="Fig. 3" captionStartId="5.[151,186,1940,1961]" captionTargetBox="[261,1329,261,1882]" captionTargetId="figure@5.[255,1334,257,1886]" captionTargetPageId="5" captionText="Fig. 3. SEM-micrographs of embolus proper of left palpal organ of two males: Liphistius desultor from Penang Hill (A, C, E, G) and Liphistius malayanus from Fraser’s Hill (B, D, F, H). (A) Retroventral-distal view. (B) Dorsal-distal view. (C) Distal view (dorsal side to the right). (D) Distal view (prolateral side to the right). (E) Retrodorsal view. (F) Prodorsal view. (G) Proventral view. (H) Prolateral view. Abbreviations: f - major embolic fold; m - membranous part of embolus proper; s - sclerotised part of embolus proper." httpUri="https://zenodo.org/record/1021943/files/figure.png" pageId="4" pageNumber="395">Figs 3F, H</figureCitation>
; 4C); distal edge of contrategulum with one to several small denticles (
<figureCitation id="5CA209C1FFE0FFF3BE0AFD203CA1FD0A" box="[1125,1209,721,745]" captionStart="Fig. 4" captionStartId="6.[151,186,1713,1734]" captionTargetBox="[156,1432,265,1656]" captionTargetId="figure@6.[148,1439,254,1693]" captionTargetPageId="6" captionText="Fig. 4. Liphistius malayanus, details of palp of four males: Templer Park (A, C-D, H-I); holotype of L. malayanus cameroni (B, E, G); Cameron Highlands, ZRC (F, J), Fraser’s Hill, MHNG (K-L). (A) Paracymbium and tibial apophysis of left palp, retroventral view. (B) Tibial apophysis of right palp, ventral view. (C) Distal part of left palpal organ, proventral view (arrow indicating V-shaped row of denticles at proventral end of distal edge of contrategulum). (D, F) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (G-H) Left palpal organ, distal view (dorsal side up). (I, K) Distal edge of contrategulum of right palp, distal view (dorsal side to the left). (J) Same, distal and slighly prolateral view. (L) Same of left palp, distal and slighly prolateral view (dorsal side to the right). Abbreviations: a - dorsal apex of contrategulum; de - distal edge of contrategulum. Scale lines: 1.0 mm (A; B; C-D; E, G; F; H-L)." httpUri="https://zenodo.org/record/1021945/files/figure.png" pageId="4" pageNumber="395">Fig. 4C</figureCitation>
, G-L); paracymbium about as deep as wide (
<figureCitation id="5CA209C1FFE0FFF3BBB7FD003A36FCEA" box="[472,558,753,777]" captionStart="Fig. 4" captionStartId="6.[151,186,1713,1734]" captionTargetBox="[156,1432,265,1656]" captionTargetId="figure@6.[148,1439,254,1693]" captionTargetPageId="6" captionText="Fig. 4. Liphistius malayanus, details of palp of four males: Templer Park (A, C-D, H-I); holotype of L. malayanus cameroni (B, E, G); Cameron Highlands, ZRC (F, J), Fraser’s Hill, MHNG (K-L). (A) Paracymbium and tibial apophysis of left palp, retroventral view. (B) Tibial apophysis of right palp, ventral view. (C) Distal part of left palpal organ, proventral view (arrow indicating V-shaped row of denticles at proventral end of distal edge of contrategulum). (D, F) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (G-H) Left palpal organ, distal view (dorsal side up). (I, K) Distal edge of contrategulum of right palp, distal view (dorsal side to the left). (J) Same, distal and slighly prolateral view. (L) Same of left palp, distal and slighly prolateral view (dorsal side to the right). Abbreviations: a - dorsal apex of contrategulum; de - distal edge of contrategulum. Scale lines: 1.0 mm (A; B; C-D; E, G; F; H-L)." httpUri="https://zenodo.org/record/1021945/files/figure.png" pageId="4" pageNumber="395">Fig. 4A</figureCitation>
). Dorsal side of vulval plate with anterolateral corners of genital atrium not bulged and elevated above level of poreplate; usually no median hairs in genital atrium (
<figureCitation id="5CA209C1FFE0FFF3BE3EFCE03CBFFCCB" box="[1105,1191,785,808]" captionStart="Fig. 5" captionStartId="8.[151,186,1809,1830]" captionTargetBox="[154,1439,260,1762]" captionTargetId="figure@8.[148,1439,254,1779]" captionTargetPageId="8" captionText="Fig. 5. Liphistius malayanus, vulval plate of eight females: paratype of L. m. cameroni from Berinchang, Cameron Highlands (A- B); exuvia of female from Ringlet, Cameron Highlands (C); female (SMF 64093) from Fraser’s Hill (D); female (moulted 24. VI. 2002) from Fraser’s Hill; poreplate slightly deformed by brief immersion in cold KOH (E); female (SMF 56206) from Fraser’s Hill (F); female (SMF 60037) from Genting Highlands (G); female (moulted 8. XII. 2005) from Genting Highlands (H); exuvia of female from Templer Park (I-J). (A, C-I) Dorsal view. (B, J) Ventral view. Scale lines: 1.0 mm (A-B; C, F; D; E, G; H; I-J)." httpUri="https://zenodo.org/record/1021947/files/figure.png" pageId="4" pageNumber="395">Fig. 5A</figureCitation>
, C-H, but see
<figureCitation id="5CA209C1FFE0FFF3BF2BFCE03D96FCCB" box="[1348,1422,785,808]" captionStart="Fig. 5" captionStartId="8.[151,186,1809,1830]" captionTargetBox="[154,1439,260,1762]" captionTargetId="figure@8.[148,1439,254,1779]" captionTargetPageId="8" captionText="Fig. 5. Liphistius malayanus, vulval plate of eight females: paratype of L. m. cameroni from Berinchang, Cameron Highlands (A- B); exuvia of female from Ringlet, Cameron Highlands (C); female (SMF 64093) from Fraser’s Hill (D); female (moulted 24. VI. 2002) from Fraser’s Hill; poreplate slightly deformed by brief immersion in cold KOH (E); female (SMF 56206) from Fraser’s Hill (F); female (SMF 60037) from Genting Highlands (G); female (moulted 8. XII. 2005) from Genting Highlands (H); exuvia of female from Templer Park (I-J). (A, C-I) Dorsal view. (B, J) Ventral view. Scale lines: 1.0 mm (A-B; C, F; D; E, G; H; I-J)." httpUri="https://zenodo.org/record/1021947/files/figure.png" pageId="4" pageNumber="395">
Fig.
<date id="B0273384FFE0FFF3BF19FCE03D96FCCB" box="[1398,1422,785,808]" pageId="4" pageNumber="395">5I</date>
</figureCitation>
); receptacular cluster not divided into subclusters (
<figureCitation id="5CA209C1FFE0FFF3B881FCC03B4FFCAA" box="[750,855,817,841]" captionStart="Fig. 5" captionStartId="8.[151,186,1809,1830]" captionTargetBox="[154,1439,260,1762]" captionTargetId="figure@8.[148,1439,254,1779]" captionTargetPageId="8" captionText="Fig. 5. Liphistius malayanus, vulval plate of eight females: paratype of L. m. cameroni from Berinchang, Cameron Highlands (A- B); exuvia of female from Ringlet, Cameron Highlands (C); female (SMF 64093) from Fraser’s Hill (D); female (moulted 24. VI. 2002) from Fraser’s Hill; poreplate slightly deformed by brief immersion in cold KOH (E); female (SMF 56206) from Fraser’s Hill (F); female (SMF 60037) from Genting Highlands (G); female (moulted 8. XII. 2005) from Genting Highlands (H); exuvia of female from Templer Park (I-J). (A, C-I) Dorsal view. (B, J) Ventral view. Scale lines: 1.0 mm (A-B; C, F; D; E, G; H; I-J)." httpUri="https://zenodo.org/record/1021947/files/figure.png" pageId="4" pageNumber="395">Fig. 5B, J</figureCitation>
)...............................................................
<taxonomicName id="03996EC7FFE0FFF3BF68FCC33D80FCAB" box="[1287,1432,817,841]" class="Arachnida" family="Liphistiidae" genus="Lipistius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="395" phylum="Arthropoda" rank="species" species="malayanus">
<emphasis id="F6EDC956FFE0FFF3BF68FCC33D80FCAB" box="[1287,1432,817,841]" italics="true" pageId="4" pageNumber="395">L. malayanus</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="C4261544FFE0FFF3BAF8FCA33D8FFBEB" blockId="4.[151,1437,305,1033]" pageId="4" pageNumber="395">
- Dorsal wall of sclerotised part of embolus proper wider than ventral wall, end of dorsal wall bent prodorsad (
<figureCitation id="5CA209C1FFE0FFF3BF1EFCA03939FC6A" captionStart="Fig. 6" captionStartId="11.[151,186,1824,1845]" captionTargetBox="[232,1351,260,1775]" captionTargetId="figure@11.[215,1371,254,1797]" captionTargetPageId="11" captionText="Fig. 6. Liphistius endau, details of left palp of eight males: from type locality, matured 27. II. 2002 (A-F); from type locality, matured 1. II. 2002 (G-I); from unknown locality, died 22. III. 2012 (J); from unknown locality, killed 30. IX. 2010 (K); from Gunung Belumut (L); from Kota Tinggi, matured 22. X. 2014 (M); same place, matured 20. VIII. 2014 (N); same place, matured 6. XI. 2014 (O). (A) Distal part of palp, ventral view. (B) Paracymbium and tibial apophysis, retroventral view. (C) Palpal organ, proventral view (arrow indicating U-shaped row of denticles at proventral end of distal edge of contrategulum). (D) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (F, H) Same, distal view (dorsal side up). (G) Embolus complex, proventral and slightly distal view. (I) Distal edge of contrategulum, distal and slighly prolateral view (dorsal side to the right). (J-O) Same, distal view. Abbreviations: dw - dorsal wall of sclerotized part of embolus proper; t - tooth on distal edge of contrategulum. Scale lines: 1.0 mm (A-B; C-E, G-H; F, I-O)." httpUri="https://zenodo.org/record/1021949/files/figure.png" pageId="4" pageNumber="395">Fig. 6F, H</figureCitation>
) and forming pronounced lobe (
<figureCitation id="5CA209C1FFE0FFF3B8E8FC803AD1FC6B" box="[647,713,881,904]" captionStart="Fig. 6" captionStartId="11.[151,186,1824,1845]" captionTargetBox="[232,1351,260,1775]" captionTargetId="figure@11.[215,1371,254,1797]" captionTargetPageId="11" captionText="Fig. 6. Liphistius endau, details of left palp of eight males: from type locality, matured 27. II. 2002 (A-F); from type locality, matured 1. II. 2002 (G-I); from unknown locality, died 22. III. 2012 (J); from unknown locality, killed 30. IX. 2010 (K); from Gunung Belumut (L); from Kota Tinggi, matured 22. X. 2014 (M); same place, matured 20. VIII. 2014 (N); same place, matured 6. XI. 2014 (O). (A) Distal part of palp, ventral view. (B) Paracymbium and tibial apophysis, retroventral view. (C) Palpal organ, proventral view (arrow indicating U-shaped row of denticles at proventral end of distal edge of contrategulum). (D) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (F, H) Same, distal view (dorsal side up). (G) Embolus complex, proventral and slightly distal view. (I) Distal edge of contrategulum, distal and slighly prolateral view (dorsal side to the right). (J-O) Same, distal view. Abbreviations: dw - dorsal wall of sclerotized part of embolus proper; t - tooth on distal edge of contrategulum. Scale lines: 1.0 mm (A-B; C-E, G-H; F, I-O)." httpUri="https://zenodo.org/record/1021949/files/figure.png" pageId="4" pageNumber="395">Fig. 6</figureCitation>
C-E); membranous part of embolus proper distally wide (
<figureCitation id="5CA209C1FFE0FFF3BF2DFC803D8FFC6B" box="[1346,1431,881,904]" captionStart="Fig. 6" captionStartId="11.[151,186,1824,1845]" captionTargetBox="[232,1351,260,1775]" captionTargetId="figure@11.[215,1371,254,1797]" captionTargetPageId="11" captionText="Fig. 6. Liphistius endau, details of left palp of eight males: from type locality, matured 27. II. 2002 (A-F); from type locality, matured 1. II. 2002 (G-I); from unknown locality, died 22. III. 2012 (J); from unknown locality, killed 30. IX. 2010 (K); from Gunung Belumut (L); from Kota Tinggi, matured 22. X. 2014 (M); same place, matured 20. VIII. 2014 (N); same place, matured 6. XI. 2014 (O). (A) Distal part of palp, ventral view. (B) Paracymbium and tibial apophysis, retroventral view. (C) Palpal organ, proventral view (arrow indicating U-shaped row of denticles at proventral end of distal edge of contrategulum). (D) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (F, H) Same, distal view (dorsal side up). (G) Embolus complex, proventral and slightly distal view. (I) Distal edge of contrategulum, distal and slighly prolateral view (dorsal side to the right). (J-O) Same, distal view. Abbreviations: dw - dorsal wall of sclerotized part of embolus proper; t - tooth on distal edge of contrategulum. Scale lines: 1.0 mm (A-B; C-E, G-H; F, I-O)." httpUri="https://zenodo.org/record/1021949/files/figure.png" pageId="4" pageNumber="395">Fig. 6C</figureCitation>
, F-H); distal edge of contrategulum with a single large triangular tooth (
<figureCitation id="5CA209C1FFE0FFF3B9B5FC603C31FC4B" box="[986,1065,913,936]" captionStart="Fig. 6" captionStartId="11.[151,186,1824,1845]" captionTargetBox="[232,1351,260,1775]" captionTargetId="figure@11.[215,1371,254,1797]" captionTargetPageId="11" captionText="Fig. 6. Liphistius endau, details of left palp of eight males: from type locality, matured 27. II. 2002 (A-F); from type locality, matured 1. II. 2002 (G-I); from unknown locality, died 22. III. 2012 (J); from unknown locality, killed 30. IX. 2010 (K); from Gunung Belumut (L); from Kota Tinggi, matured 22. X. 2014 (M); same place, matured 20. VIII. 2014 (N); same place, matured 6. XI. 2014 (O). (A) Distal part of palp, ventral view. (B) Paracymbium and tibial apophysis, retroventral view. (C) Palpal organ, proventral view (arrow indicating U-shaped row of denticles at proventral end of distal edge of contrategulum). (D) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (F, H) Same, distal view (dorsal side up). (G) Embolus complex, proventral and slightly distal view. (I) Distal edge of contrategulum, distal and slighly prolateral view (dorsal side to the right). (J-O) Same, distal view. Abbreviations: dw - dorsal wall of sclerotized part of embolus proper; t - tooth on distal edge of contrategulum. Scale lines: 1.0 mm (A-B; C-E, G-H; F, I-O)." httpUri="https://zenodo.org/record/1021949/files/figure.png" pageId="4" pageNumber="395">Fig. 6F</figureCitation>
, H-O); paracymbium much deeper than wide (
<figureCitation id="5CA209C1FFE0FFF3BB33FC4039A9FC2B" box="[348,433,945,968]" captionStart="Fig. 6" captionStartId="11.[151,186,1824,1845]" captionTargetBox="[232,1351,260,1775]" captionTargetId="figure@11.[215,1371,254,1797]" captionTargetPageId="11" captionText="Fig. 6. Liphistius endau, details of left palp of eight males: from type locality, matured 27. II. 2002 (A-F); from type locality, matured 1. II. 2002 (G-I); from unknown locality, died 22. III. 2012 (J); from unknown locality, killed 30. IX. 2010 (K); from Gunung Belumut (L); from Kota Tinggi, matured 22. X. 2014 (M); same place, matured 20. VIII. 2014 (N); same place, matured 6. XI. 2014 (O). (A) Distal part of palp, ventral view. (B) Paracymbium and tibial apophysis, retroventral view. (C) Palpal organ, proventral view (arrow indicating U-shaped row of denticles at proventral end of distal edge of contrategulum). (D) Same, retrodorsal and slightly proximal view. (E) Same, retrodorsal view. (F, H) Same, distal view (dorsal side up). (G) Embolus complex, proventral and slightly distal view. (I) Distal edge of contrategulum, distal and slighly prolateral view (dorsal side to the right). (J-O) Same, distal view. Abbreviations: dw - dorsal wall of sclerotized part of embolus proper; t - tooth on distal edge of contrategulum. Scale lines: 1.0 mm (A-B; C-E, G-H; F, I-O)." httpUri="https://zenodo.org/record/1021949/files/figure.png" pageId="4" pageNumber="395">Fig. 6B</figureCitation>
). Dorsal side of vulval plate with anterolateral corners of genital atrium bulged and elevated above level of poreplate; median hairs usually present in genital atrium (
<figureCitation id="5CA209C1FFE0FFF3BE6FFC203C40FC0A" box="[1024,1112,977,1001]" captionStart="Fig. 7" captionStartId="12.[151,186,1940,1961]" captionTargetBox="[159,1415,617,1889]" captionTargetId="figure@12.[151,1436,609,1911]" captionTargetPageId="12" captionText="Fig. 7. Liphistius endau, vulval plate of six females: adult from unknown locality (A-B); largest female from Kota Tinggi (C); exuvia of reproductive female (moulted 27. IV. 2002) from type locality (D-E); exuvia of reproductive female (moulted 28. IV. 2002) from type locality (F-G); medium-sized juvenile from type locality (H); small juvenile from type locality (I). (A, C-D, F, H-I) Entire vulval plate, dorsal view. (B) Poreplate, ventral view. (E, G) Entire vulval plate, ventral view. Scale lines: 1.0 mm (A-B; C; D-E; F-G; H-I)." httpUri="https://zenodo.org/record/1021951/files/figure.png" pageId="4" pageNumber="395">Fig. 7A</figureCitation>
, C-D, F); receptacular cluster divided into three more or less distinct subclusters (
<figureCitation id="5CA209C1FFE0FFF3B966FC003B81FBEA" box="[777,921,1009,1033]" captionStart="Fig. 7" captionStartId="12.[151,186,1940,1961]" captionTargetBox="[159,1415,617,1889]" captionTargetId="figure@12.[151,1436,609,1911]" captionTargetPageId="12" captionText="Fig. 7. Liphistius endau, vulval plate of six females: adult from unknown locality (A-B); largest female from Kota Tinggi (C); exuvia of reproductive female (moulted 27. IV. 2002) from type locality (D-E); exuvia of reproductive female (moulted 28. IV. 2002) from type locality (F-G); medium-sized juvenile from type locality (H); small juvenile from type locality (I). (A, C-D, F, H-I) Entire vulval plate, dorsal view. (B) Poreplate, ventral view. (E, G) Entire vulval plate, ventral view. Scale lines: 1.0 mm (A-B; C; D-E; F-G; H-I)." httpUri="https://zenodo.org/record/1021951/files/figure.png" pageId="4" pageNumber="395">Fig. 7B, E, G</figureCitation>
).............................................................
<taxonomicName id="03996EC7FFE0FFF3BF55FC033D8FFBEB" ID-CoL="72J5Q" authorityName="Sedgwick &amp; Platnick" authorityYear="1987" box="[1338,1431,1009,1033]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="395" phylum="Arthropoda" rank="species" species="endau">
<emphasis id="F6EDC956FFE0FFF3BF55FC033D8FFBEB" box="[1338,1431,1009,1033]" italics="true" pageId="4" pageNumber="395">L. endau</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="8C8346CFFFE0FFF3BAF8FBA53975FB0E" pageId="4" pageNumber="395" type="distribution">
<paragraph id="C4261544FFE0FFF3BAF8FBA53975FB0E" blockId="4.[151,756,1108,1261]" pageId="4" pageNumber="395">
<emphasis id="F6EDC956FFE0FFF3BAF8FBA53934FB88" bold="true" box="[151,300,1108,1131]" pageId="4" pageNumber="395">Distribution:</emphasis>
All described species of this group occur in the western and southern part of peninsular
<collectingCountry id="BC8E55D4FFE0FFF3B8FDFB853AECFB68" box="[658,756,1140,1163]" name="Malaysia" pageId="4" pageNumber="395">Malaysia</collectingCountry>
(
<figureCitation id="5CA209C1FFE0FFF3BAF0FB6538FAFB4E" box="[159,226,1172,1197]" captionStart="Fig. 1" captionStartId="1.[151,186,1392,1413]" captionTargetBox="[153,758,625,1349]" captionTargetId="figure@1.[149,499,895,1170]" captionTargetPageId="1" captionText="Fig. 1. Localities of Liphistius species of the malayanus - group, tioman - group, linang - group and batuenis - group in peninsular Malaysia and southern Thailand (coast of Sumatra omitted): 1 - Gunung Angsi (type locality of L. malayanus); 2 - Templer Park and Gua Anak Takun (L. malayanus, L. batuensis); 3 - Genting Highlands (L. malayanus); 4 - Fraser’s Hill (L. malayanus); 5 - Cameron Highlands (type locality of L. malayanus cameroni); 6 - Sungai Jasin in Endau Rompin National Park (type locality of L. endau); 7 - Gunung Belumut (L. endau); 8 - Gunung Muntahak (L. endau; type locality of L. gracilis sp. nov.); 9 - Sungai Rengit (type locality of L. johore); 10 - Gunung Kajang on Tioman Island (type locality of L. tioman); 11 - Gua Charas in Bukit Charas (type locality of L. panching); 12 - Nusa Camp in Taman Negara (type locality of L. negara sp. nov.); 13 - Jeram Linang Waterfall (type locality of L. linang sp. nov.); 14 - Sankalakhierie Mountains (type locality of L. indra); 15 - Batu Caves (type locality of L. batuensis); 16 - Gua Tempurung (type locality of L. tempurung); 17 - Gua Cicak (L. tempurung); 18 - Gua Keris (type locality of L. priceae sp. nov). Localities with conspecific populations are encircled. Colours distinguish species groups." httpUri="https://zenodo.org/record/1021937/files/figure.png" pageId="4" pageNumber="395">Fig. 1</figureCitation>
, localities 1-9); an undescribed species appears to be present in the northeastern corner of the country (
<figureCitation id="5CA209C1FFE0FFF3BAF0FB2538C6FB0E" box="[159,222,1236,1261]" captionStart="Fig. 1" captionStartId="1.[151,186,1392,1413]" captionTargetBox="[153,758,625,1349]" captionTargetId="figure@1.[149,499,895,1170]" captionTargetPageId="1" captionText="Fig. 1. Localities of Liphistius species of the malayanus - group, tioman - group, linang - group and batuenis - group in peninsular Malaysia and southern Thailand (coast of Sumatra omitted): 1 - Gunung Angsi (type locality of L. malayanus); 2 - Templer Park and Gua Anak Takun (L. malayanus, L. batuensis); 3 - Genting Highlands (L. malayanus); 4 - Fraser’s Hill (L. malayanus); 5 - Cameron Highlands (type locality of L. malayanus cameroni); 6 - Sungai Jasin in Endau Rompin National Park (type locality of L. endau); 7 - Gunung Belumut (L. endau); 8 - Gunung Muntahak (L. endau; type locality of L. gracilis sp. nov.); 9 - Sungai Rengit (type locality of L. johore); 10 - Gunung Kajang on Tioman Island (type locality of L. tioman); 11 - Gua Charas in Bukit Charas (type locality of L. panching); 12 - Nusa Camp in Taman Negara (type locality of L. negara sp. nov.); 13 - Jeram Linang Waterfall (type locality of L. linang sp. nov.); 14 - Sankalakhierie Mountains (type locality of L. indra); 15 - Batu Caves (type locality of L. batuensis); 16 - Gua Tempurung (type locality of L. tempurung); 17 - Gua Cicak (L. tempurung); 18 - Gua Keris (type locality of L. priceae sp. nov). Localities with conspecific populations are encircled. Colours distinguish species groups." httpUri="https://zenodo.org/record/1021937/files/figure.png" pageId="4" pageNumber="395">Fig. 1</figureCitation>
, locality 13).
</paragraph>
</subSubSection>
</subSection>
</treatment>
</document>
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