298 lines
35 KiB
XML
298 lines
35 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.975.55502" ID-GBIF-Dataset="005c19e6-92bc-4fa3-88d7-59f3abb88ce5" ID-PMC="PMC7572540" ID-Pensoft-Pub="1313-2970-975-87" ID-Pensoft-UUID="782B7D099D2E5F2BAEF492BD4AF41337" ID-PubMed="33117066" ID-ZooBank="68B311A14AA5408F95090A52245290AA" ModsDocID="1313-2970-975-87" checkinTime="1602606909261" checkinUser="pensoft" docAuthor="Sorokina, Vera S. & Ovtshinnikova, Olga G." docDate="2020" docId="1FCD2321A12A5327ACF19253F92FEF69" docLanguage="en" docName="ZooKeys 975: 87-110" docOrigin="ZooKeys 975" docSource="http://dx.doi.org/10.3897/zookeys.975.55502" docTitle="Drymeia firthiana" docType="treatment" docVersion="4" id="782B7D099D2E5F2BAEF492BD4AF41337" lastPageNumber="87" masterDocId="782B7D099D2E5F2BAEF492BD4AF41337" masterDocTitle="The position of the Azeliinae in the Muscidae (Diptera) based on musculature of the male terminalia" masterLastPageNumber="110" masterPageNumber="87" pageNumber="87" updateTime="1668169611172" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The position of the Azeliinae in the Muscidae (Diptera) based on musculature of the male terminalia</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sorokina, Vera S.</mods:namePart>
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<mods:affiliation>Institute of Systematics and Ecology of Animals, Siberian Branch of the Russian Academy of Sciences, Novosibirsk, 630091, Russia</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0003-3679-9005</mods:nameIdentifier>
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<mods:nameIdentifier type="email">sorokinavs@mail.ru</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ovtshinnikova, Olga G.</mods:namePart>
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<mods:affiliation>Zoological Institute, Russian Academy of Sciences, St. Petersburg, 199034, Russia</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>975</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>87</mods:start>
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<mods:end>110</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.975.55502</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.975.55502</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-975-87</mods:identifier>
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<mods:identifier type="ZooBank">68B311A14AA5408F95090A52245290AA</mods:identifier>
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<mods:identifier type="Pensoft-UUID">782B7D099D2E5F2BAEF492BD4AF41337</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="168338821" LSID="urn:lsid:plazi:treatment:1FCD2321A12A5327ACF19253F92FEF69" httpUri="http://treatment.plazi.org/id/1FCD2321A12A5327ACF19253F92FEF69" lastPageNumber="87" pageId="0" pageNumber="87">
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<subSubSection pageId="0" pageNumber="87" type="nomenclature">
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<paragraph pageId="0" pageNumber="87">
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<taxonomicName LSID="1FCD2321-A12A-5327-ACF1-9253F92FEF69" authority="(Huckett, 1965)" baseAuthorityName="Huckett" baseAuthorityYear="1965" class="Insecta" family="Muscidae" genus="Drymeia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Drymeia firthiana" order="Diptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="firthiana">Drymeia firthiana (Huckett, 1965)</taxonomicName>
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Drymeia firthiana (Huckett, 1965). Male pregenital segments, inner view. Upper muscles ISM 5 removed left and lower muscles ISM 5 removed right." figureDoi="10.3897/zookeys.975.55502.figure1" httpUri="https://binary.pensoft.net/fig/462754" pageId="0" pageNumber="87">Figures 1</figureCitation>
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male genitalia of Drymeia firthiana (Huckett, 1965) A hypandrium, inner view B aedeagal complex, lateral view." figureDoi="10.3897/zookeys.975.55502.figure2" httpUri="https://binary.pensoft.net/fig/462755" pageId="0" pageNumber="87">, 2</figureCitation>
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male genitalia of Drymeia firthiana (Huckett, 1965) A genitalia, lateral view B epandrial complex, dorsal view." figureDoi="10.3897/zookeys.975.55502.figure3" httpUri="https://binary.pensoft.net/fig/462756" pageId="0" pageNumber="87">, 3</figureCitation>
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male genitalia of Drymeia firthiana (Huckett, 1965). Epandrial complex, inner view." figureDoi="10.3897/zookeys.975.55502.figure4" httpUri="https://binary.pensoft.net/fig/462757" pageId="0" pageNumber="87">, 4</figureCitation>
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Male genitalia of Drymeia firthiana (Huckett, 1965) A aedeagal complex, lateral view, with muscles M 1, M 21, M 22, M 23, M 23 and part of hypandrium, epiphallus removed B aedeagal complex, lateral view, sclerites C genitalia, lateral view, with muscles M 18 r, M 18 l and M 5 D genitalia, lateral view, sclerites E surstyli and subepandrial sclerite, inner view, with muscles M 4, M 3 F epandrial complex, inner view, sclerites." figureDoi="10.3897/zookeys.975.55502.figure9" httpUri="https://binary.pensoft.net/fig/462762" pageId="0" pageNumber="87">, 9</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="87" type="materials_examined">
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<paragraph pageId="0" pageNumber="87">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="87">
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10 males, Russia, Altai Republic, Kosh-Agach district, 8 km NE Maitobe Mt., 2420 m,
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<geoCoordinate degrees="49" direction="north" minutes="34" orientation="latitude" precision="925" value="49.566666">49°34'N</geoCoordinate>
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,
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<geoCoordinate degrees="87" direction="east" minutes="43" orientation="longitude" precision="925" value="87.71667">87°43'E</geoCoordinate>
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, pan traps, 7-10.vii.2006, leg. V. Sorokina.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="87" type="description">
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<paragraph pageId="0" pageNumber="87">Description.</paragraph>
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<paragraph pageId="0" pageNumber="87">
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<emphasis bold="true" pageId="0" pageNumber="87">
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<emphasis italics="true" pageId="0" pageNumber="87">Abdominal segments</emphasis>
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.
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</emphasis>
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Sternite I reduced to narrow band, tergites I and II fused. Segments III and IV and tergite V not modified; sternite V enlarged, with wide median notch.
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</paragraph>
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<paragraph pageId="0" pageNumber="87">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="87">Pregenital segments</emphasis>
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(Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Drymeia firthiana (Huckett, 1965). Male pregenital segments, inner view. Upper muscles ISM 5 removed left and lower muscles ISM 5 removed right." figureDoi="10.3897/zookeys.975.55502.figure1" httpUri="https://binary.pensoft.net/fig/462754" pageId="0" pageNumber="87">1</figureCitation>
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). Tergite VI reduced to long narrow sclerotized band. Sternite VI positioned under sternite V, reduced in size, represented by short, narrow, wavy sclerite; remainder of sternite VI desclerotized. Sternite VII long, narrow, positioned on left side of body, dilated at articulation with syntergosternite VII + VIII; ventrally connected to desclerotized left margins of sternite VI; laterally connected to syntergosternite VII + VIII. Syntergosternite relatively wide, positioned dorsally; left end wider than right end and connected to sternite VII, right end free; posterior margin extended to epandrium.
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</paragraph>
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<caption doi="10.3897/zookeys.975.55502.figure1" httpUri="https://binary.pensoft.net/fig/462754" pageId="0" pageNumber="87" start="Figure 1" startId="F1">
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<paragraph pageId="0" pageNumber="87">
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<emphasis bold="true" pageId="0" pageNumber="87">Figure 1.</emphasis>
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<taxonomicName baseAuthorityName="Huckett" baseAuthorityYear="1965" class="Insecta" family="Muscidae" genus="Drymeia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Drymeia firthiana" order="Diptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="firthiana">
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<emphasis italics="true" pageId="0" pageNumber="87">Drymeia firthiana</emphasis>
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</taxonomicName>
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(Huckett, 1965). Male pregenital segments, inner view. Upper
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<taxonomicName class="Insecta" family="Muscidae" genus="Drymeia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="muscles" order="Diptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="muscles">muscles</taxonomicName>
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ISM 5 removed left and lower
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<taxonomicName class="Insecta" family="Muscidae" genus="Drymeia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="muscles" order="Diptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="muscles">muscles</taxonomicName>
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ISM 5 removed right.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="87">
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<emphasis bold="true" pageId="0" pageNumber="87">
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<emphasis italics="true" pageId="0" pageNumber="87">Genitalia</emphasis>
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.
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</emphasis>
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Hypandrium in form of concave plate, V-shaped (Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male genitalia of Drymeia firthiana (Huckett, 1965) A hypandrium, inner view B aedeagal complex, lateral view." figureDoi="10.3897/zookeys.975.55502.figure2" httpUri="https://binary.pensoft.net/fig/462755" pageId="0" pageNumber="87">2A</figureCitation>
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); lateral arms of hypandrium articulated with surstyli and epandrium. Pregonites and postgonites present; pregonites larger than postgonites, tapered apically, and longer phallapodeme (Figs
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male genitalia of Drymeia firthiana (Huckett, 1965) A hypandrium, inner view B aedeagal complex, lateral view." figureDoi="10.3897/zookeys.975.55502.figure2" httpUri="https://binary.pensoft.net/fig/462755" pageId="0" pageNumber="87">2B</figureCitation>
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,
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Male genitalia of Drymeia firthiana (Huckett, 1965) A aedeagal complex, lateral view, with muscles M 1, M 21, M 22, M 23, M 23 and part of hypandrium, epiphallus removed B aedeagal complex, lateral view, sclerites C genitalia, lateral view, with muscles M 18 r, M 18 l and M 5 D genitalia, lateral view, sclerites E surstyli and subepandrial sclerite, inner view, with muscles M 4, M 3 F epandrial complex, inner view, sclerites." figureDoi="10.3897/zookeys.975.55502.figure9" httpUri="https://binary.pensoft.net/fig/462762" pageId="0" pageNumber="87">9B</figureCitation>
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). Phallus comprises epiphallus and distiphallus, basiphallus inconspicuous, either absent or fused with distiphallus. Phallapodeme long, articulated with phallus. Epiphallus well-developed, shaped as long and distally rounded plate. Distiphallus large, as long as pregonite, expanded distally, broadly articulated with epiphallus. Ejaculatory apodeme concave-shaped plate. Epandrium hemispherical, with large posteromedian notch (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male genitalia of Drymeia firthiana (Huckett, 1965) A genitalia, lateral view B epandrial complex, dorsal view." figureDoi="10.3897/zookeys.975.55502.figure3" httpUri="https://binary.pensoft.net/fig/462756" pageId="0" pageNumber="87">3</figureCitation>
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). Cerci large, wide, fused for a considerable length; each cercus with distal semicircular apical notch and well-formed process (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male genitalia of Drymeia firthiana (Huckett, 1965) A genitalia, lateral view B epandrial complex, dorsal view." figureDoi="10.3897/zookeys.975.55502.figure3" httpUri="https://binary.pensoft.net/fig/462756" pageId="0" pageNumber="87">3B</figureCitation>
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). Surstylus well-developed, wide, expanded and rounded apically, bent inward, with small process. Cercus approximately as long as surstylus. Subepandrial sclerite present as two long, quite wide, medially not connected plates, as long as length of surstyli and merging with them (Figs
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male genitalia of Drymeia firthiana (Huckett, 1965). Epandrial complex, inner view." figureDoi="10.3897/zookeys.975.55502.figure4" httpUri="https://binary.pensoft.net/fig/462757" pageId="0" pageNumber="87">4</figureCitation>
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,
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Male genitalia of Drymeia firthiana (Huckett, 1965) A aedeagal complex, lateral view, with muscles M 1, M 21, M 22, M 23, M 23 and part of hypandrium, epiphallus removed B aedeagal complex, lateral view, sclerites C genitalia, lateral view, with muscles M 18 r, M 18 l and M 5 D genitalia, lateral view, sclerites E surstyli and subepandrial sclerite, inner view, with muscles M 4, M 3 F epandrial complex, inner view, sclerites." figureDoi="10.3897/zookeys.975.55502.figure9" httpUri="https://binary.pensoft.net/fig/462762" pageId="0" pageNumber="87">9F</figureCitation>
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).
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</paragraph>
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<caption doi="10.3897/zookeys.975.55502.figure2" httpUri="https://binary.pensoft.net/fig/462755" pageId="0" pageNumber="87" start="Figure 2" startId="F2">
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<paragraph pageId="0" pageNumber="87">
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<emphasis bold="true" pageId="0" pageNumber="87">Figure 2.</emphasis>
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Male genitalia of
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<taxonomicName baseAuthorityName="Huckett" baseAuthorityYear="1965" class="Insecta" family="Muscidae" genus="Drymeia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Drymeia firthiana" order="Diptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="firthiana">
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<emphasis italics="true" pageId="0" pageNumber="87">Drymeia firthiana</emphasis>
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</taxonomicName>
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(Huckett, 1965)
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<emphasis bold="true" pageId="0" pageNumber="87">A</emphasis>
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hypandrium, inner view
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<emphasis bold="true" pageId="0" pageNumber="87">B</emphasis>
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aedeagal complex, lateral view.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="87">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="87">
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Thoracic
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<taxonomicName class="Insecta" family="Muscidae" genus="Drymeia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="muscles" order="Diptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="muscles">muscles</taxonomicName>
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</emphasis>
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. Paired symmetrical conical
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<taxonomicName class="Insecta" family="Muscidae" genus="Drymeia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="muscles" order="Diptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="muscles">muscles</taxonomicName>
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extend from thorax to lateromedian parts of tergite I + II, and also straight
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<taxonomicName class="Insecta" family="Muscidae" genus="Drymeia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="muscles" order="Diptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="muscles">muscles</taxonomicName>
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extend from thorax to basal parts of sternite II.
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</paragraph>
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<caption doi="10.3897/zookeys.975.55502.figure3" httpUri="https://binary.pensoft.net/fig/462756" pageId="0" pageNumber="87" start="Figure 3" startId="F3">
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<paragraph pageId="0" pageNumber="87">
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<emphasis bold="true" pageId="0" pageNumber="87">Figure 3.</emphasis>
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Male genitalia of
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<taxonomicName baseAuthorityName="Huckett" baseAuthorityYear="1965" class="Insecta" family="Muscidae" genus="Drymeia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Drymeia firthiana" order="Diptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="firthiana">
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<emphasis italics="true" pageId="0" pageNumber="87">Drymeia firthiana</emphasis>
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</taxonomicName>
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(Huckett, 1965)
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<emphasis bold="true" pageId="0" pageNumber="87">A</emphasis>
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genitalia, lateral view
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<emphasis bold="true" pageId="0" pageNumber="87">B</emphasis>
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epandrial complex, dorsal view.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="87">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="87">
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Abdominal
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<taxonomicName class="Insecta" family="Muscidae" genus="Drymeia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="muscles" order="Diptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="muscles">muscles</taxonomicName>
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</emphasis>
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(Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Drymeia firthiana (Huckett, 1965). Male pregenital segments, inner view. Upper muscles ISM 5 removed left and lower muscles ISM 5 removed right." figureDoi="10.3897/zookeys.975.55502.figure1" httpUri="https://binary.pensoft.net/fig/462754" pageId="0" pageNumber="87">1</figureCitation>
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): ITM 2-ITM 4, ITM 5a, ITM 5b, ISM 2-ISM 5, TSM 1-TSM 5. Flat, very short
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<taxonomicName class="Insecta" family="Muscidae" genus="Drymeia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="muscles" order="Diptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="muscles">muscles</taxonomicName>
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ITM 2-ITM 4 extend from distal parts of tergites II-IV along their entire width to basal margins of tergites III-V. Paired symmetrical
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<taxonomicName class="Insecta" family="Muscidae" genus="Drymeia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="muscles" order="Diptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="muscles">muscles</taxonomicName>
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ITM 5a extend from median parts of tergite V to median parts of basal margin of tergite VI. Long, paired, almost symmetrical conical
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<taxonomicName class="Insecta" family="Muscidae" genus="Drymeia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="muscles" order="Diptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="muscles">muscles</taxonomicName>
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ITM 5b extend from laterobasal parts of tergite V to membrane at lateral parts of tergite VI.
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</paragraph>
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<caption doi="10.3897/zookeys.975.55502.figure4" httpUri="https://binary.pensoft.net/fig/462757" pageId="0" pageNumber="87" start="Figure 4" startId="F4">
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<paragraph pageId="0" pageNumber="87">
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<emphasis bold="true" pageId="0" pageNumber="87">Figure 4.</emphasis>
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Male genitalia of
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<taxonomicName baseAuthorityName="Huckett" baseAuthorityYear="1965" class="Insecta" family="Muscidae" genus="Drymeia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Drymeia firthiana" order="Diptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="firthiana">
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<emphasis italics="true" pageId="0" pageNumber="87">Drymeia firthiana</emphasis>
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</taxonomicName>
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(Huckett, 1965). Epandrial complex, inner view.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="87">
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Paired symmetrical
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<taxonomicName class="Insecta" family="Muscidae" genus="Drymeia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="muscles" order="Diptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="muscles">muscles</taxonomicName>
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ISM 2-ISM 4 extend along the entire basal margin of sternites II-IV to basal margins of sternites III-V, respectively. Very powerful, paired, symmetrical, fan-shaped
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<taxonomicName class="Insecta" family="Muscidae" genus="Drymeia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="muscles" order="Diptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="muscles">muscles</taxonomicName>
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ISM 5 extend in two layers from basal margin of sternite V to sclerotized plate of sternite VI and to sternite VII at connection with membrane of sternite VI. Muscles extending to sternite VI connected with middle part of sternite V, but
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<taxonomicName class="Insecta" family="Muscidae" genus="Drymeia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="muscles" order="Diptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="muscles">muscles</taxonomicName>
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extending to sternite VII connected with basal part of sternite V. Wide and flat pleural abdominal
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<taxonomicName class="Insecta" family="Muscidae" genus="Drymeia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="muscles" order="Diptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="muscles">muscles</taxonomicName>
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TSM 1-TSM 5 easily discernible on corresponding segments.
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</paragraph>
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<paragraph pageId="0" pageNumber="87">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="87">
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Pregenital
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<taxonomicName class="Insecta" family="Muscidae" genus="Drymeia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="muscles" order="Diptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="muscles">muscles</taxonomicName>
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</emphasis>
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(Figs
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Drymeia firthiana (Huckett, 1965). Male pregenital segments, inner view. Upper muscles ISM 5 removed left and lower muscles ISM 5 removed right." figureDoi="10.3897/zookeys.975.55502.figure1" httpUri="https://binary.pensoft.net/fig/462754" pageId="0" pageNumber="87">1</figureCitation>
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,
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male genitalia of Drymeia firthiana (Huckett, 1965) A hypandrium, inner view B aedeagal complex, lateral view." figureDoi="10.3897/zookeys.975.55502.figure2" httpUri="https://binary.pensoft.net/fig/462755" pageId="0" pageNumber="87">2A</figureCitation>
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,
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male genitalia of Drymeia firthiana (Huckett, 1965) A genitalia, lateral view B epandrial complex, dorsal view." figureDoi="10.3897/zookeys.975.55502.figure3" httpUri="https://binary.pensoft.net/fig/462756" pageId="0" pageNumber="87">3A</figureCitation>
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,
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male genitalia of Drymeia firthiana (Huckett, 1965). Epandrial complex, inner view." figureDoi="10.3897/zookeys.975.55502.figure4" httpUri="https://binary.pensoft.net/fig/462757" pageId="0" pageNumber="87">4</figureCitation>
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|
): ITM 6, ISM 6, ISM 7, TSM 7, M 18, M 191, M 192. Small and short, paired, slightly asymmetrical
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<taxonomicName class="Insecta" family="Muscidae" genus="Drymeia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="muscles" order="Diptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="muscles">muscles</taxonomicName>
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ITM 6 extend from lateral parts of tergite VI to lateral parts of syntergosternite VII + VIII.
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</paragraph>
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<paragraph pageId="0" pageNumber="87">
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Paired
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<taxonomicName class="Insecta" family="Muscidae" genus="Drymeia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="muscles" order="Diptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="muscles">muscles</taxonomicName>
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|
ISM 6: left ISM 6 extends from left membranous parts of sternite VI to lateral margin of inner surface of sternite VII, articulated with syntergosternite VII + VIII; right muscle ISM 6 extends from right membranous parts of sternite VI to membrane near right laterobasal margin of syntergosternite VII + VIII; left ISM 6 larger than right muscle ISM 6. Unpaired left muscle ISM 7 short but powerful, extending from lateral margin of outer surface of sternite VII to outgrowth on lateral part of basal margin of syntergosternite VII + VIII. Paired asymmetrical
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<taxonomicName class="Insecta" family="Muscidae" genus="Drymeia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="muscles" order="Diptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="muscles">muscles</taxonomicName>
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|
TSM 7: left muscle TSM 7 wide, short, fan-shaped, extending from lateral part of inner surface of sternite VII to small outgrowth on lateral margin of syntergosternite VII + VIII; right muscle TSM 7 long, fan-shaped, extending from right basal margin of sternite VII to small, sclerotized part of membrane adjacent to syntergosternite VII + VIII.
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</paragraph>
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<paragraph pageId="0" pageNumber="87">
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Paired asymmetrical
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<taxonomicName class="Insecta" family="Muscidae" genus="Drymeia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="muscles" order="Diptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="muscles">muscles</taxonomicName>
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M 18: right muscle M 18 r wide and flat (homologous with left M 18 in
|
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<taxonomicName authorityName="Verrall" authorityYear="1888" lsidName="" pageId="0" pageNumber="87" rank="subFamily" subFamily="Mydaeinae">Mydaeinae</taxonomicName>
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), extending from membrane covering genital cavity near syntergosternite VII + VIII to middle of basal margin of hypandrium (Figs
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male genitalia of Drymeia firthiana (Huckett, 1965) A hypandrium, inner view B aedeagal complex, lateral view." figureDoi="10.3897/zookeys.975.55502.figure2" httpUri="https://binary.pensoft.net/fig/462755" pageId="0" pageNumber="87">2A</figureCitation>
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,
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male genitalia of Drymeia firthiana (Huckett, 1965) A genitalia, lateral view B epandrial complex, dorsal view." figureDoi="10.3897/zookeys.975.55502.figure3" httpUri="https://binary.pensoft.net/fig/462756" pageId="0" pageNumber="87">3A</figureCitation>
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,
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|
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Male genitalia of Drymeia firthiana (Huckett, 1965) A aedeagal complex, lateral view, with muscles M 1, M 21, M 22, M 23, M 23 and part of hypandrium, epiphallus removed B aedeagal complex, lateral view, sclerites C genitalia, lateral view, with muscles M 18 r, M 18 l and M 5 D genitalia, lateral view, sclerites E surstyli and subepandrial sclerite, inner view, with muscles M 4, M 3 F epandrial complex, inner view, sclerites." figureDoi="10.3897/zookeys.975.55502.figure9" httpUri="https://binary.pensoft.net/fig/462762" pageId="0" pageNumber="87">9C</figureCitation>
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); left muscle M 18 l long (homologous with right M 18 in
|
|
<taxonomicName authorityName="Verrall" authorityYear="1888" lsidName="" pageId="0" pageNumber="87" rank="subFamily" subFamily="Mydaeinae">Mydaeinae</taxonomicName>
|
|
), extending from left part of syntergosternite VII + VIII to inner surface of left laterobasal part of hypandrium. Paired asymmetrical
|
|
<taxonomicName class="Insecta" family="Muscidae" genus="Drymeia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="muscles" order="Diptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="muscles">muscles</taxonomicName>
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|
M 191 (Fig.
|
|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male genitalia of Drymeia firthiana (Huckett, 1965). Epandrial complex, inner view." figureDoi="10.3897/zookeys.975.55502.figure4" httpUri="https://binary.pensoft.net/fig/462757" pageId="0" pageNumber="87">4</figureCitation>
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|
): left muscle M 191 l short but powerful, extending from inner surface of small area on left lateral part of syntergosternite VII + VIII (close to connection with sternite VII) to small area on left laterobasal margin of epandrium; right muscle M 191 r longer but weaker than M 191 l, extending from right part of syntergosternite VII + VIII to right laterobasal margin of epandrium. Unpaired asymmetrical muscle M 192 powerful, fan-shaped, and oblique, extending from right lateral part of syntergosternite VII + VIII to slightly to right from middle of basal margin of epandrium.
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|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="87">
|
|
<emphasis bold="true" pageId="0" pageNumber="87">
|
|
<emphasis italics="true" pageId="0" pageNumber="87">
|
|
Genital
|
|
<taxonomicName class="Insecta" family="Muscidae" genus="Drymeia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="muscles" order="Diptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="muscles">muscles</taxonomicName>
|
|
</emphasis>
|
|
.
|
|
</emphasis>
|
|
<taxonomicName genus="Tergosternal" lsidName="Tergosternal muscles" pageId="0" pageNumber="87" rank="species" species="muscles">
|
|
<emphasis italics="true" pageId="0" pageNumber="87">Tergosternal muscles</emphasis>
|
|
</taxonomicName>
|
|
(Figs
|
|
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male genitalia of Drymeia firthiana (Huckett, 1965) A genitalia, lateral view B epandrial complex, dorsal view." figureDoi="10.3897/zookeys.975.55502.figure3" httpUri="https://binary.pensoft.net/fig/462756" pageId="0" pageNumber="87">3A</figureCitation>
|
|
,
|
|
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Male genitalia of Drymeia firthiana (Huckett, 1965) A aedeagal complex, lateral view, with muscles M 1, M 21, M 22, M 23, M 23 and part of hypandrium, epiphallus removed B aedeagal complex, lateral view, sclerites C genitalia, lateral view, with muscles M 18 r, M 18 l and M 5 D genitalia, lateral view, sclerites E surstyli and subepandrial sclerite, inner view, with muscles M 4, M 3 F epandrial complex, inner view, sclerites." figureDoi="10.3897/zookeys.975.55502.figure9" httpUri="https://binary.pensoft.net/fig/462762" pageId="0" pageNumber="87">9C</figureCitation>
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|
): M 5. Paired, symmetrical powerful
|
|
<taxonomicName genus="Tergosternal" lsidName="muscles" pageId="0" pageNumber="87" rank="species" species="muscles">muscles</taxonomicName>
|
|
M 5 extend from lateral parts of basal margin of hypandrium to median parts of basal margin of epandrium.
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="87">
|
|
<emphasis italics="true" pageId="0" pageNumber="87">Muscles of hypandrial complex</emphasis>
|
|
(Figs
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male genitalia of Drymeia firthiana (Huckett, 1965) A hypandrium, inner view B aedeagal complex, lateral view." figureDoi="10.3897/zookeys.975.55502.figure2" httpUri="https://binary.pensoft.net/fig/462755" pageId="0" pageNumber="87">2</figureCitation>
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,
|
|
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Male genitalia of Drymeia firthiana (Huckett, 1965) A aedeagal complex, lateral view, with muscles M 1, M 21, M 22, M 23, M 23 and part of hypandrium, epiphallus removed B aedeagal complex, lateral view, sclerites C genitalia, lateral view, with muscles M 18 r, M 18 l and M 5 D genitalia, lateral view, sclerites E surstyli and subepandrial sclerite, inner view, with muscles M 4, M 3 F epandrial complex, inner view, sclerites." figureDoi="10.3897/zookeys.975.55502.figure9" httpUri="https://binary.pensoft.net/fig/462762" pageId="0" pageNumber="87">9A</figureCitation>
|
|
): M 1, M 21, M 22, M 23, M 23. Wide and powerful, paired, symmetrical
|
|
<taxonomicName genus="Tergosternal" lsidName="muscles" pageId="0" pageNumber="87" rank="species" species="muscles">muscles</taxonomicName>
|
|
M 1 extend from hypandrium, occupying considerable part of inner surface, to curve of mediobasal part of phallapodeme in front of pregonites. Long paired symmetrical
|
|
<taxonomicName genus="Tergosternal" lsidName="muscles" pageId="0" pageNumber="87" rank="species" species="muscles">muscles</taxonomicName>
|
|
M 21 extend from arms of hypandrium to laterodistal parts of phallapodeme. Powerful paired symmetrical
|
|
<taxonomicName genus="Tergosternal" lsidName="muscles" pageId="0" pageNumber="87" rank="species" species="muscles">muscles</taxonomicName>
|
|
M 22 extend from entire laterobasal part of pregonites to distal half of phallapodeme. Symmetrical
|
|
<taxonomicName genus="Tergosternal" lsidName="muscles" pageId="0" pageNumber="87" rank="species" species="muscles">muscles</taxonomicName>
|
|
M 23 long and close to each other, extending from membrane of basal margin of epiphallus between lateral ends of hypandrial arms to distal part of phallapodeme, opposite epandrium;
|
|
<taxonomicName genus="Tergosternal" lsidName="muscles" pageId="0" pageNumber="87" rank="species" species="muscles">muscles</taxonomicName>
|
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M 23 very close to
|
|
<taxonomicName genus="Tergosternal" lsidName="muscles" pageId="0" pageNumber="87" rank="species" species="muscles">muscles</taxonomicName>
|
|
M 21 and both look like one muscle. Constrictors of ejaculatory apodeme small;
|
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<taxonomicName genus="Tergosternal" lsidName="muscles" pageId="0" pageNumber="87" rank="species" species="muscles">muscles</taxonomicName>
|
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M 23 surrounding ejaculatory apodeme, contraction pumps seminal fluid into phallus.
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|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="87">
|
|
<emphasis italics="true" pageId="0" pageNumber="87">Muscles of epandrial complex</emphasis>
|
|
(Figs
|
|
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male genitalia of Drymeia firthiana (Huckett, 1965) A genitalia, lateral view B epandrial complex, dorsal view." figureDoi="10.3897/zookeys.975.55502.figure3" httpUri="https://binary.pensoft.net/fig/462756" pageId="0" pageNumber="87">3B</figureCitation>
|
|
,
|
|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male genitalia of Drymeia firthiana (Huckett, 1965). Epandrial complex, inner view." figureDoi="10.3897/zookeys.975.55502.figure4" httpUri="https://binary.pensoft.net/fig/462757" pageId="0" pageNumber="87">4</figureCitation>
|
|
,
|
|
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Male genitalia of Drymeia firthiana (Huckett, 1965) A aedeagal complex, lateral view, with muscles M 1, M 21, M 22, M 23, M 23 and part of hypandrium, epiphallus removed B aedeagal complex, lateral view, sclerites C genitalia, lateral view, with muscles M 18 r, M 18 l and M 5 D genitalia, lateral view, sclerites E surstyli and subepandrial sclerite, inner view, with muscles M 4, M 3 F epandrial complex, inner view, sclerites." figureDoi="10.3897/zookeys.975.55502.figure9" httpUri="https://binary.pensoft.net/fig/462762" pageId="0" pageNumber="87">9E</figureCitation>
|
|
): M 3, M 4, M 7, M 24-M 26. Powerful paired symmetrical
|
|
<taxonomicName genus="Tergosternal" lsidName="muscles" pageId="0" pageNumber="87" rank="species" species="muscles">muscles</taxonomicName>
|
|
M 3 extend from inner surface of basal parts of epandrium to inner surface of basal parts of subepandrial sclerite. Powerful paired symmetrical
|
|
<taxonomicName genus="Tergosternal" lsidName="muscles" pageId="0" pageNumber="87" rank="species" species="muscles">muscles</taxonomicName>
|
|
M 4 extend from lateral parts of inner surface of epandrium to inner surface of basal parts of surstyli. Paired symmetrical thin cercal
|
|
<taxonomicName genus="Tergosternal" lsidName="muscles" pageId="0" pageNumber="87" rank="species" species="muscles">muscles</taxonomicName>
|
|
M 7 extend from inner part of subepandrial sclerite to laterobasal parts of cerci. Broad powerful muscle M 24 passes inside cerci connecting lateral parts of two halves of cerci. Broad paired
|
|
<taxonomicName genus="Tergosternal" lsidName="muscles" pageId="0" pageNumber="87" rank="species" species="muscles">muscles</taxonomicName>
|
|
M 25 extend from median parts of distal margin of epandrium to integument of anus. Powerful paired symmetrical, fan-shaped
|
|
<taxonomicName genus="Tergosternal" lsidName="muscles" pageId="0" pageNumber="87" rank="species" species="muscles">muscles</taxonomicName>
|
|
M 26 extend from distolateral parts of epandrium to laterobasal margins of small cercal outgrowths.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |