treatments-xml/data/1D/36/87/1D3687F8A761A668FF62FAAFFBCC75B8.xml

<document id="5CBCA61FA432FA81490D925B490539A2" ID-CLB-Dataset="22701" ID-DOI="10.3897/zookeys.7.117" ID-GBIF-Dataset="7a12aaad-a568-4920-91cf-af3e9407eadc" ID-ISSN="1313–2970" ID-Zenodo-Dep="576436" ID-ZooBank="67B4D2EC-2C6D-4226-847D-0BA2B2777AE3" IM.bibliography_approvedBy="admin" IM.illustrations_approvedBy="admin" IM.materialsCitations_approvedBy="admin" IM.metadata_approvedBy="admin" IM.taxonomicNames_approvedBy="admin" IM.treatments_approvedBy="admin" checkinTime="1587986763134" checkinUser="plazi" docAuthor="Golovatch, Sergei, Geoffroy, Jean-Jacques, Mauriès, Jean-Paul &amp; VandenSpiegel, Didier" docDate="2009" docId="1D3687F8A761A668FF62FAAFFBCC75B8" docLanguage="en" docName="ZK_article_1970.pdf" docOrigin="ZooKeys 7 (7)" docStyle="DocumentStyle:BEF2356135E79670CA200BCAD40C4E3A:zookeys.2008.journal_article" docStyleId="BEF2356135E79670CA200BCAD40C4E3A" docStyleName="zookeys.2008.journal_article" docTitle="Eutrichodesmus basalis Golovatch, Geoffroy, Mauries &amp; VandenSpiegel 2009, sp. n." docType="treatment" docUuid="49B81FBA-709E-40C4-BA47-0E36E1E39664" docUuidSource="ZooBank" docVersion="7" lastPageNumber="13" masterDocId="E10FFF80A766A664FFEEFF9FFF8E742C" masterDocTitle="Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida)" masterLastPageNumber="53" masterPageNumber="1" pageNumber="8" updateTime="1693525343643" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:title id="E771CFF26D75C236678DC08ED21B3FA8">Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida)</mods:title>
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<mods:namePart id="8A5509B3C44E9ABDF58C15151F8F0553">Golovatch, Sergei</mods:namePart>
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<mods:affiliation id="5F3FBFF9EFB9947694FCFE51D3DC4DE5">Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071, Russia</mods:affiliation>
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<mods:namePart id="31C552EDF174B95E3FCA7C378CB8FE7E">Geoffroy, Jean-Jacques</mods:namePart>
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<mods:affiliation id="55A572C35A11A12F06DDDE27A1EAD487">Muséum national d’Histoire naturelle, Département Ecologie &amp; Gestion de la Biodiversité, UMR 7179 du CNRS, Equipe EVOLTRAIT, 4, avenue du Petit Château, F- 91800 Brunoy, France</mods:affiliation>
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<mods:affiliation id="9B28F50DB88211E56C606A1C5590BB7F">Muséum national d’Histoire naturelle, Département Systématique et Evolution, USM 602, Section Arthropodes Case Postale n ° 53, 61 rue Buffon F- 75231 Paris, France</mods:affiliation>
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<taxonomicName id="529F4D6DA761A663FF62FAAFFC507166" ID-CoL="6HKQM" authority="Golovatch, Geoffroy, Mauries &amp; VandenSpiegel" authorityName="Golovatch, Geoffroy, Mauries &amp; VandenSpiegel" authorityYear="2009" box="[140,990,1327,1355]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="basalis" status="sp. n.">
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<emphasis id="A7EBEAFCA761A663FF62FAAFFEC67167" box="[140,328,1328,1355]" inLineHeading="true" pageId="7" pageNumber="8" reason="1">Eutrichodesmus</emphasis>
basalis
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Golovatch, Geoffroy, Mauriès &amp; VandenSpiegel
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, 
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<paragraph id="952036EEA761A663FF62FAE8FF6071BD" blockId="7.[140,1067,1327,1425]" box="[140,238,1399,1425]" pageId="7" pageNumber="8">
<figureCitation id="0DA42A6BA761A663FF62FAE8FF6771BD" box="[140,233,1399,1425]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStartId-0="8.[140,212,1558,1580]" captionStartId-1="9.[140,212,1590,1612]" captionStartId-2="10.[140,212,1590,1612]" captionStartId-3="11.[140,212,1622,1644]" captionTargetBox-0="[140,1108,506,1539]" captionTargetBox-1="[140,1108,407,1570]" captionTargetBox-2="[140,1108,826,1570]" captionTargetBox-3="[200,1045,807,1593]" captionTargetId-0="figure@8.[140,1108,505,1539]" captionTargetId-1="figure@9.[140,1108,407,1570]" captionTargetId-2="figure@10.[140,1108,825,1570]" captionTargetId-3="figure@11.[194,1054,799,1602]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionTargetPageId-3="11" captionText-0="Figure 1. Eutrichodesmus basalis sp. n., ♀ paratype; A, habitus, lateral view; B, anterior part of body, lateral view; C, midbody segments, lateral view; D, posterior part of body, lateral view; E, midbody segments, dorsal view; F, posterior part of body, dorsal view. − Scale bars: A, 0.5 mm, B, C, D &amp; F, 0.1 mm &amp; E, 0.2 mm." captionText-1="Figure 2. Eutrichodesmus basalis sp. n., ♀ paratype; A, anterior part of body, frontodorsal view; B, head, front view; C, antenna; D, cross-section of a midbody segment, caudal view; E, posterior part of body, ventral view; F, legs, ventral view. − Scale bars: A, B, D, E, 0.1 mm &amp; C, F, 0.05 mm." captionText-2="Figure 3. Eutrichodesmus basalis sp. n., ♀ paratype; A, enlarged paraterga with ozopores, lateral view; B, bisegmented seta; C, tegument texture; D, limbus and adjacent parts; E, claw. − Scale bars: A, 0.05 mm, B, D, E, 0.01 mm, C, 0.02 mm." captionText-3="Figure 4. Eutrichodesmus basalis sp. n., ♂ paratype; A, leg 9, caudal view; B &amp; C, left gonopod, mesal and lateral views, respectively (dp = distofemoral process). − Scale bar: A, 0.2 mm, B, C, 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3787336" figureDoi-1="http://doi.org/10.5281/zenodo.3769071" figureDoi-2="http://doi.org/10.5281/zenodo.3769073" figureDoi-3="http://doi.org/10.5281/zenodo.3769075" httpUri-0="https://zenodo.org/record/3787336/files/figure.png" httpUri-1="https://zenodo.org/record/3769071/files/figure.png" httpUri-2="https://zenodo.org/record/3769073/files/figure.png" httpUri-3="https://zenodo.org/record/3769075/files/figure.png" pageId="7" pageNumber="8">Figs 1-4</figureCitation>
.
</paragraph>
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<emphasis id="A7EBEAFCA761A663FF62FA22FEB571FB" bold="true" box="[140,315,1469,1495]" pageId="7" pageNumber="8">
<emphasis id="A7EBEAFCA761A663FF62FA22FF4871FB" box="[140,198,1469,1495]" inLineHeading="true" pageId="7" pageNumber="8" reason="1">Type</emphasis>
material.
</emphasis>
<materialsCitation id="25F73CB3A761A663FEAAFA22FC5971D6" ID-GBIF-Occurrence="2610433329" collectingDate="2003-06-12" collectionCode="MNHN" collectorName="P. Trontelj &amp; B. Sket" country="Vietnam" location="Hang Bo Nau Cave" municipality="Dao Bo Hon" pageId="7" pageNumber="8" specimenCode="JC 309" specimenCount="1" specimenCount-male="1" stateProvince="Vinh Ha Long Prov" typeStatus="holotype">
<collectingCountry id="ED88767EA761A663FEAAFA22FE2771FB" box="[324,425,1469,1495]" name="Vietnam" pageId="7" pageNumber="8" value="Vietnam">Vietnam</collectingCountry>
, 
<collectingRegion id="575BF80CA761A663FE56FA22FD2A71F4" box="[440,676,1469,1496]" pageId="7" pageNumber="8">Vinh Ha Long Prov</collectingRegion>
. (SW), 
<collectingMunicipality id="7544AC94A761A663FCEFFA21FC1071F4" box="[769,926,1470,1496]" pageId="7" pageNumber="8">Dao Bo Hon</collectingMunicipality>
, 
<location id="90406035A761A663FC40FA21FF4971D6" LSID="urn:lsid:plazi:treatment:1D3687F8A761A668FF62FAAFFBCC75B8:90406035A761A663FC40FA21FF4971D6" country="Vietnam" municipality="Dao Bo Hon" name="Hang Bo Nau Cave" pageId="7" pageNumber="8" stateProvince="Vinh Ha Long Prov">Hang Bo Nau Cave</location>
, 
<date id="E121102EA761A663FF3CFA7FFED671D6" box="[210,344,1504,1530]" pageId="7" pageNumber="8" value="2003-06-12">
<collectingDate id="F165E9C6A761A663FF3CFA7FFED671D6" box="[210,344,1504,1530]" pageId="7" pageNumber="8" value="2003-06-12">12.VI.2003</collectingDate>
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, leg. 
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&amp; 
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, 
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(
<collectionCode id="F38EAE2BA761A663FCFDFA7FFCFA71D6" box="[787,884,1504,1530]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="7" pageNumber="8">MNHN</collectionCode>
<specimenCode id="C5399E95A761A663FC94FA7FFC4071D6" box="[890,974,1504,1530]" pageId="7" pageNumber="8">JC 309</specimenCode>
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, 
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<typeStatus id="4A24884CA761A663FC0CFA7EFBDE71D7" box="[994,1104,1505,1531]" pageId="7" pageNumber="8">paratypes</typeStatus>
: 
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(
<collectionCode id="F38EAE2BA761A663FF29F99BFEA67232" box="[199,296,1540,1566]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="7" pageNumber="8">MNHN</collectionCode>
<specimenCode id="C5399E95A761A663FEDEF99BFE0B7232" box="[304,389,1540,1566]" pageId="7" pageNumber="8">JC 309</specimenCode>
), 
<specimenCount id="8399FD67A761A663FE74F99BFE317232" box="[410,447,1540,1566]" pageId="7" pageNumber="8" type="female">1 ♀</specimenCount>
(
<collectionCode id="F38EAE2BA761A663FE38F99BFD987232" box="[470,534,1540,1566]" pageId="7" pageNumber="8">SEM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DD856565A761A663FF57F9B8FBDA726D" box="[185,1108,1575,1601]" pageId="7" pageNumber="8" type="etymology">
<paragraph id="952036EEA761A663FF57F9B8FBDA726D" blockId="7.[140,1108,1469,1672]" box="[185,1108,1575,1601]" pageId="7" pageNumber="8">
<emphasis id="A7EBEAFCA761A663FF57F9B8FE8D726D" bold="true" box="[185,259,1575,1601]" pageId="7" pageNumber="8">
<emphasis id="A7EBEAFCA761A663FF57F9B8FF73726D" box="[185,253,1575,1601]" inLineHeading="true" pageId="7" pageNumber="8" reason="1">Name</emphasis>
.
</emphasis>
To emphasize the obvious basal position amongst the volvatory “doratodesmids”.
</paragraph>
</subSubSection>
<subSubSection id="DD856565A761A663FF57F9D6FBAB72A4" pageId="7" pageNumber="8" type="diagnosis">
<paragraph id="952036EEA761A663FF57F9D6FBAB72A4" blockId="7.[140,1108,1469,1672]" pageId="7" pageNumber="8">
<emphasis id="A7EBEAFCA761A663FF57F9D6FEB6724F" bold="true" box="[185,312,1609,1635]" pageId="7" pageNumber="8">
<emphasis id="A7EBEAFCA761A663FF57F9D6FEBD724F" box="[185,307,1609,1635]" inLineHeading="true" pageId="7" pageNumber="8" reason="1">Diagnosis</emphasis>
.
</emphasis>
Differs from congeners by the apparently imperfect volvation, due to particularly short paraterga, coupled with an especially simple gonopod structure.
</paragraph>
</subSubSection>
<subSubSection id="DD856565A76EA66FFF57FF38FBC075D1" lastPageId="11" lastPageNumber="12" pageId="8" pageNumber="9" type="description">
<paragraph id="952036EEA76EA66CFF57FF38FC24752B" blockId="8.[140,1108,166,440]" pageId="8" pageNumber="9">
<emphasis id="A7EBEAFCA76EA66CFF57FF38FEB274EC" box="[185,316,167,192]" italics="true" pageId="8" pageNumber="9">Description:</emphasis>
Length of adults of both sexes 
<emphasis id="A7EBEAFCA76EA66CFD42FF38FD4D74EC" box="[684,707,167,192]" italics="true" pageId="8" pageNumber="9">ca</emphasis>
4.8-5.0 mm, width 
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, body broadest at segment 3 or 4; 
<specimenCount id="8399FD67A76EA66CFDE1FF54FDBA74CA" box="[527,564,203,230]" pageId="8" pageNumber="9">♂♂</specimenCount>
a little smaller than 
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. 
<typeStatus id="4A24884CA76EA66CFC8BFF55FC5E74C8" box="[869,976,202,228]" pageId="8" pageNumber="9">Holotype</typeStatus>
<emphasis id="A7EBEAFCA76EA66CFC36FF54FC6174C8" box="[984,1007,203,228]" italics="true" pageId="8" pageNumber="9">ca</emphasis>
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long and 
<quantity id="52679B0BA76EA66CFF19FF72FEDF7524" box="[247,337,237,264]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="8" pageNumber="9" unit="mm" value="0.6">0.6 mm</quantity>
wide. Coloration uniformly pallid or light yellowish.
</paragraph>
<paragraph id="952036EEA76EA66CFF57FE8FFBDD759B" blockId="8.[140,1108,166,440]" pageId="8" pageNumber="9">
Adults with 20 segments, body subcylindrical (
<figureCitation id="0DA42A6BA76EA66CFD3BFE8FFCA37506" box="[725,813,272,298]" captionStart="Figure 1" captionStartId="8.[140,212,1558,1580]" captionTargetBox="[140,1108,506,1539]" captionTargetId="figure@8.[140,1108,505,1539]" captionTargetPageId="8" captionText="Figure 1. Eutrichodesmus basalis sp. n., ♀ paratype; A, habitus, lateral view; B, anterior part of body, lateral view; C, midbody segments, lateral view; D, posterior part of body, lateral view; E, midbody segments, dorsal view; F, posterior part of body, dorsal view. − Scale bars: A, 0.5 mm, B, C, D &amp; F, 0.1 mm &amp; E, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3787336" httpUri="https://zenodo.org/record/3787336/files/figure.png" pageId="8" pageNumber="9">Fig. 1A</figureCitation>
), pattern of conglobation typical of “ 
<taxonomicName id="529F4D6DA76EA66CFEE5FEACFE407561" authorityName="Cook" authorityYear="1896" box="[267,462,307,333]" class="Diplopoda" family="Doratodesmidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="family">Doratodesmidae</taxonomicName>
” as described by 
<bibRefCitation id="F10E4B1FA76EA66CFD77FEACFCE27562" author="Golovatch" box="[665,876,307,334]" firstAuthor="Golovatch" pageId="8" pageNumber="9" pagination="39 - 60" refId="ref19093" refString="Golovatch SI (2003) A review of the volvatory Polydesmida, with special reference to the patterns of volvation (Diplopoda). African Invertebrates 44 (1): 39 - 60." type="journal article" year="2003">Golovatch (2003)</bibRefCitation>
, but volvation itself obviously somewhat imperfect in barely concealing all (especially hind) legs (
<figureCitation id="0DA42A6BA76EA66CFBCAFEC8FF7075B8" captionStart="Figure 1" captionStartId="8.[140,212,1558,1580]" captionTargetBox="[140,1108,506,1539]" captionTargetId="figure@8.[140,1108,505,1539]" captionTargetPageId="8" captionText="Figure 1. Eutrichodesmus basalis sp. n., ♀ paratype; A, habitus, lateral view; B, anterior part of body, lateral view; C, midbody segments, lateral view; D, posterior part of body, lateral view; E, midbody segments, dorsal view; F, posterior part of body, dorsal view. − Scale bars: A, 0.5 mm, B, C, D &amp; F, 0.1 mm &amp; E, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3787336" httpUri="https://zenodo.org/record/3787336/files/figure.png" pageId="8" pageNumber="9">Figs 1A, C, D</figureCitation>
). Head (
<figureCitation id="0DA42A6BA76EA66CFE88FEE5FE3075B8" box="[358,446,378,404]" captionStart="Figure 2" captionStartId="9.[140,212,1590,1612]" captionTargetBox="[140,1108,407,1570]" captionTargetId="figure@9.[140,1108,407,1570]" captionTargetPageId="9" captionText="Figure 2. Eutrichodesmus basalis sp. n., ♀ paratype; A, anterior part of body, frontodorsal view; B, head, front view; C, antenna; D, cross-section of a midbody segment, caudal view; E, posterior part of body, ventral view; F, legs, ventral view. − Scale bars: A, B, D, E, 0.1 mm &amp; C, F, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3769071" httpUri="https://zenodo.org/record/3769071/files/figure.png" pageId="8" pageNumber="9">Fig. 2B</figureCitation>
) slightly transverse, rather densely pilose, microgranular amd microvillose just below antennae and on vertex, with a pair of paramedian,
</paragraph>
<caption id="C1E06666A76EA66CFF62F989FE9E72A0" ID-DOI="http://doi.org/10.5281/zenodo.3787336" ID-Zenodo-Dep="3787336" httpUri="https://zenodo.org/record/3787336/files/figure.png" pageId="8" pageNumber="9" startId="8.[140,212,1558,1580]" targetBox="[140,1108,506,1539]" targetPageId="8">
<paragraph id="952036EEA76EA66CFF62F989FE9E72A0" blockId="8.[140,1108,1558,1676]" pageId="8" pageNumber="9">
<emphasis id="A7EBEAFCA76EA66CFF62F989FF7E7200" bold="true" box="[140,240,1558,1580]" pageId="8" pageNumber="9">Figure 1.</emphasis>
<taxonomicName id="529F4D6DA76EA66CFF16F989FE4D7207" authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel, 2009" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel" authorityYear="2009" box="[248,451,1558,1579]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="basalis" status="sp. n.">
<emphasis id="A7EBEAFCA76EA66CFF16F989FE4D7207" box="[248,451,1558,1579]" italics="true" pageId="8" pageNumber="9">Eutrichodesmus basalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BCD85787A76EA66CFE25F988FD8F7200" box="[459,513,1559,1580]" pageId="8" pageNumber="9" rank="species">sp. n.</taxonomicNameLabel>
, 
<specimenCount id="8399FD67A76EA66CFDE3F988FD977200" box="[525,537,1559,1580]" pageId="8" pageNumber="9" type="female">♀</specimenCount>
<typeStatus id="4A24884CA76EA66CFDC9F988FDF77200" box="[551,633,1559,1580]" pageId="8" pageNumber="9">paratype</typeStatus>
; 
<emphasis id="A7EBEAFCA76EA66CFD6AF989FD1D7207" bold="true" box="[644,659,1558,1579]" pageId="8" pageNumber="9">A</emphasis>
, habitus, lateral view; 
<emphasis id="A7EBEAFCA76EA66CFC85F989FCF47207" bold="true" box="[875,890,1558,1579]" pageId="8" pageNumber="9">B</emphasis>
, anterior part of body, lateral view; 
<emphasis id="A7EBEAFCA76EA66CFEEFF9A9FE9C7260" bold="true" box="[257,274,1590,1612]" pageId="8" pageNumber="9">C</emphasis>
, midbody segments, lateral view; 
<emphasis id="A7EBEAFCA76EA66CFDA1F9A9FDEC7267" bold="true" box="[591,610,1590,1611]" pageId="8" pageNumber="9">D</emphasis>
, posterior part of body, lateral view; 
<emphasis id="A7EBEAFCA76EA66CFC55F9A9FC477267" bold="true" box="[955,969,1590,1611]" pageId="8" pageNumber="9">E</emphasis>
, midbody segments, dorsal view; 
<emphasis id="A7EBEAFCA76EA66CFEADF9C9FEDE7247" bold="true" box="[323,336,1622,1643]" pageId="8" pageNumber="9">F</emphasis>
, posterior part of body, dorsal view. − Scale bars: A, 
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, B, C, D &amp; F, 
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&amp; E, 
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.
</paragraph>
</caption>
<paragraph id="952036EEA76FA66EFF62FF38FCC676D8" blockId="9.[140,1108,167,334]" lastBlockId="10.[140,1108,167,756]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
almost contiguous knobs; isthmus between antennal sockets very narrow. Antennae (
<figureCitation id="0DA42A6BA76FA66DFF7AFF55FF7E74C8" box="[148,240,202,228]" captionStart="Figure 2" captionStartId="9.[140,212,1590,1612]" captionTargetBox="[140,1108,407,1570]" captionTargetId="figure@9.[140,1108,407,1570]" captionTargetPageId="9" captionText="Figure 2. Eutrichodesmus basalis sp. n., ♀ paratype; A, anterior part of body, frontodorsal view; B, head, front view; C, antenna; D, cross-section of a midbody segment, caudal view; E, posterior part of body, ventral view; F, legs, ventral view. − Scale bars: A, B, D, E, 0.1 mm &amp; C, F, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3769071" httpUri="https://zenodo.org/record/3769071/files/figure.png" pageId="9" pageNumber="10">Fig. 2C</figureCitation>
) very short and stout; antennomere 6 longer than 5, dorso-apically with an evident pit containing a tight group of minute bacilliform sensilla; antennomere 8 with the usual four sensory cones apically. Collum rather large, regularly convex, not covering the head from above, entire surface microvillose, with four transverse rows of round tubercles with a pit on top (apparently representing the former place of insertion of bisegmented, tactile setae, which are mostly lost) (
<figureCitation id="0DA42A6BA76CA66EFC6BFF55FC6F74C8" box="[901,993,202,228]" captionStart="Figure 2" captionStartId="9.[140,212,1590,1612]" captionTargetBox="[140,1108,407,1570]" captionTargetId="figure@9.[140,1108,407,1570]" captionTargetPageId="9" captionText="Figure 2. Eutrichodesmus basalis sp. n., ♀ paratype; A, anterior part of body, frontodorsal view; B, head, front view; C, antenna; D, cross-section of a midbody segment, caudal view; E, posterior part of body, ventral view; F, legs, ventral view. − Scale bars: A, B, D, E, 0.1 mm &amp; C, F, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3769071" httpUri="https://zenodo.org/record/3769071/files/figure.png" pageId="10" pageNumber="11">Fig. 2A</figureCitation>
). Prozona very finely alveolate, collum and metaterga covered with a cerotegumental crust held by abundant microvilli; stricture between pro- and metazona broad and shallow, more finely alveolate-microgranular than prozona; limbus microcrenulate, nearly fully hidden by nearby abundant microvilli (
<figureCitation id="0DA42A6BA76CA66EFD7FFEC8FC99755D" box="[657,791,343,369]" captionStart="Figure 3" captionStartId="10.[140,212,1590,1612]" captionTargetBox="[140,1108,826,1570]" captionTargetId="figure@10.[140,1108,825,1570]" captionTargetPageId="10" captionText="Figure 3. Eutrichodesmus basalis sp. n., ♀ paratype; A, enlarged paraterga with ozopores, lateral view; B, bisegmented seta; C, tegument texture; D, limbus and adjacent parts; E, claw. − Scale bars: A, 0.05 mm, B, D, E, 0.01 mm, C, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.3769073" httpUri="https://zenodo.org/record/3769073/files/figure.png" pageId="10" pageNumber="11">Figs 3C, D</figureCitation>
). Metaterga behind collum with three transverse rows of tubercles (
<figureCitation id="0DA42A6BA76CA66EFDB4FEE5FD1375B8" box="[602,669,378,404]" captionStart="Figure 1" captionStartId="8.[140,212,1558,1580]" captionTargetBox="[140,1108,506,1539]" captionTargetId="figure@8.[140,1108,505,1539]" captionTargetPageId="8" captionText="Figure 1. Eutrichodesmus basalis sp. n., ♀ paratype; A, habitus, lateral view; B, anterior part of body, lateral view; C, midbody segments, lateral view; D, posterior part of body, lateral view; E, midbody segments, dorsal view; F, posterior part of body, dorsal view. − Scale bars: A, 0.5 mm, B, C, D &amp; F, 0.1 mm &amp; E, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3787336" httpUri="https://zenodo.org/record/3787336/files/figure.png" pageId="10" pageNumber="11">Figs 1</figureCitation>
A-F), some of which still retain long, bisegmented setae (
<figureCitation id="0DA42A6BA76CA66EFE9EFE02FE42759B" box="[368,460,413,439]" captionStart="Figure 3" captionStartId="10.[140,212,1590,1612]" captionTargetBox="[140,1108,826,1570]" captionTargetId="figure@10.[140,1108,825,1570]" captionTargetPageId="10" captionText="Figure 3. Eutrichodesmus basalis sp. n., ♀ paratype; A, enlarged paraterga with ozopores, lateral view; B, bisegmented seta; C, tegument texture; D, limbus and adjacent parts; E, claw. − Scale bars: A, 0.05 mm, B, D, E, 0.01 mm, C, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.3769073" httpUri="https://zenodo.org/record/3769073/files/figure.png" pageId="10" pageNumber="11">Fig. 3B</figureCitation>
). Paraterga subvertical, very narrow, downwards barely reaching level of venter, clearly trisinuate caudolaterally at base due to two lobulations (
<figureCitation id="0DA42A6BA76CA66EFF3DFE7CFEDA75D2" box="[211,340,483,510]" captionStart="Figure 1" captionStartId="8.[140,212,1558,1580]" captionTargetBox="[140,1108,506,1539]" captionTargetId="figure@8.[140,1108,505,1539]" captionTargetPageId="8" captionText="Figure 1. Eutrichodesmus basalis sp. n., ♀ paratype; A, habitus, lateral view; B, anterior part of body, lateral view; C, midbody segments, lateral view; D, posterior part of body, lateral view; E, midbody segments, dorsal view; F, posterior part of body, dorsal view. − Scale bars: A, 0.5 mm, B, C, D &amp; F, 0.1 mm &amp; E, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3787336" httpUri="https://zenodo.org/record/3787336/files/figure.png" pageId="10" pageNumber="11">Figs 1C, D</figureCitation>
; 2D; 3A); paraterga 2 strongly enlarged, with a series of lobulations anterolaterally, both schism and hyposchism very small; paraterga 3 and 4 slightly shorter than others (
<figureCitation id="0DA42A6BA76CA66EFE93FDB5FE587668" box="[381,470,554,580]" captionStart="Figure 1" captionStartId="8.[140,212,1558,1580]" captionTargetBox="[140,1108,506,1539]" captionTargetId="figure@8.[140,1108,505,1539]" captionTargetPageId="8" captionText="Figure 1. Eutrichodesmus basalis sp. n., ♀ paratype; A, habitus, lateral view; B, anterior part of body, lateral view; C, midbody segments, lateral view; D, posterior part of body, lateral view; E, midbody segments, dorsal view; F, posterior part of body, dorsal view. − Scale bars: A, 0.5 mm, B, C, D &amp; F, 0.1 mm &amp; E, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3787336" httpUri="https://zenodo.org/record/3787336/files/figure.png" pageId="10" pageNumber="11">Fig. 1B</figureCitation>
), overlap of following paraterga typical, latter broadly rounded and evidently trilobate. Pore formula normal, ozopores located on top of a small porostele-like tubercle on ventrocaudal lobulation (
<figureCitation id="0DA42A6BA76CA66EFCBFFDEFFC2176A6" box="[849,943,624,650]" captionStart="Figure 3" captionStartId="10.[140,212,1590,1612]" captionTargetBox="[140,1108,826,1570]" captionTargetId="figure@10.[140,1108,825,1570]" captionTargetPageId="10" captionText="Figure 3. Eutrichodesmus basalis sp. n., ♀ paratype; A, enlarged paraterga with ozopores, lateral view; B, bisegmented seta; C, tegument texture; D, limbus and adjacent parts; E, claw. − Scale bars: A, 0.05 mm, B, D, E, 0.01 mm, C, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.3769073" httpUri="https://zenodo.org/record/3769073/files/figure.png" pageId="10" pageNumber="11">Fig. 3A</figureCitation>
). Pleurotergal ridges very small, alveolate-microgranular like entire ventral surface. Epiproct short, also with tubercles, directed ventrocaudad, with the usual four cones (= spinneret setae) just below tip; para- and hypoprocts as in 
<figureCitation id="0DA42A6BA76CA66EFD52FD45FC9D76D8" box="[700,787,730,756]" captionStart="Figure 1" captionStartId="8.[140,212,1558,1580]" captionTargetBox="[140,1108,506,1539]" captionTargetId="figure@8.[140,1108,505,1539]" captionTargetPageId="8" captionText="Figure 1. Eutrichodesmus basalis sp. n., ♀ paratype; A, habitus, lateral view; B, anterior part of body, lateral view; C, midbody segments, lateral view; D, posterior part of body, lateral view; E, midbody segments, dorsal view; F, posterior part of body, dorsal view. − Scale bars: A, 0.5 mm, B, C, D &amp; F, 0.1 mm &amp; E, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3787336" httpUri="https://zenodo.org/record/3787336/files/figure.png" pageId="10" pageNumber="11">Figs 1F</figureCitation>
; 2E.
</paragraph>
<caption id="C1E06666A76FA66DFF62F9A9FC2872A0" ID-DOI="http://doi.org/10.5281/zenodo.3769071" ID-Zenodo-Dep="3769071" httpUri="https://zenodo.org/record/3769071/files/figure.png" pageId="9" pageNumber="10" startId="9.[140,212,1590,1612]" targetBox="[140,1108,407,1570]" targetPageId="9">
<paragraph id="952036EEA76FA66DFF62F9A9FC2872A0" blockId="9.[140,1108,1590,1676]" pageId="9" pageNumber="10">
<emphasis id="A7EBEAFCA76FA66DFF62F9A9FF607260" bold="true" box="[140,238,1590,1612]" pageId="9" pageNumber="10">Figure 2.</emphasis>
<taxonomicName id="529F4D6DA76FA66DFF1AF9A9FE337267" authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel, 2009" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel" authorityYear="2009" box="[244,445,1590,1611]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="basalis" status="sp. n.">
<emphasis id="A7EBEAFCA76FA66DFF1AF9A9FE337267" box="[244,445,1590,1611]" italics="true" pageId="9" pageNumber="10">Eutrichodesmus basalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BCD85787A76FA66DFE2DF9A8FE787260" box="[451,502,1591,1612]" pageId="9" pageNumber="10" rank="species">sp. n.</taxonomicNameLabel>
, 
<specimenCount id="8399FD67A76FA66DFDEFF9A8FD837260" box="[513,525,1591,1612]" pageId="9" pageNumber="10">♀</specimenCount>
paratype; 
<emphasis id="A7EBEAFCA76FA66DFD9DF9A9FD0C7267" bold="true" box="[627,642,1590,1611]" pageId="9" pageNumber="10">A</emphasis>
, anterior part of body, frontodorsal view; 
<emphasis id="A7EBEAFCA76FA66DFBE8F9A9FB9B7267" bold="true" box="[1030,1045,1590,1611]" pageId="9" pageNumber="10">B</emphasis>
, head, front view; 
<emphasis id="A7EBEAFCA76FA66DFF17F9C9FE847240" bold="true" box="[249,266,1622,1644]" pageId="9" pageNumber="10">C</emphasis>
, antenna; 
<emphasis id="A7EBEAFCA76FA66DFE80F9C9FE0F7247" bold="true" box="[366,385,1622,1643]" pageId="9" pageNumber="10">D</emphasis>
, cross-section of a midbody segment, caudal view; 
<emphasis id="A7EBEAFCA76FA66DFC8BF9C9FCFD7247" bold="true" box="[869,883,1622,1643]" pageId="9" pageNumber="10">E</emphasis>
, posterior part of body, ventral view; 
<emphasis id="A7EBEAFCA76FA66DFEE9F9E9FE9A72A7" bold="true" box="[263,276,1654,1675]" pageId="9" pageNumber="10">F</emphasis>
, legs, ventral view. − Scale bars: A, B, D, E, 0.1 mm &amp; C, F, 0.05 mm.
</paragraph>
</caption>
<caption id="C1E06666A76CA66EFF62F9A9FE3472A0" ID-DOI="http://doi.org/10.5281/zenodo.3769073" ID-Zenodo-Dep="3769073" httpUri="https://zenodo.org/record/3769073/files/figure.png" pageId="10" pageNumber="11" startId="10.[140,212,1590,1612]" targetBox="[140,1108,826,1570]" targetPageId="10">
<paragraph id="952036EEA76CA66EFF62F9A9FE3472A0" blockId="10.[140,1108,1590,1676]" pageId="10" pageNumber="11">
<emphasis id="A7EBEAFCA76CA66EFF62F9A9FF607260" bold="true" box="[140,238,1590,1612]" pageId="10" pageNumber="11">Figure 3.</emphasis>
<taxonomicName id="529F4D6DA76CA66EFF1AF9A9FE337267" authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel, 2009" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel" authorityYear="2009" box="[244,445,1590,1611]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="basalis" status="sp. n.">
<emphasis id="A7EBEAFCA76CA66EFF1AF9A9FE337267" box="[244,445,1590,1611]" italics="true" pageId="10" pageNumber="11">Eutrichodesmus basalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BCD85787A76CA66EFE2DF9A8FE787260" box="[451,502,1591,1612]" pageId="10" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
, 
<specimenCount id="8399FD67A76CA66EFDEFF9A8FD837260" box="[513,525,1591,1612]" pageId="10" pageNumber="11">♀</specimenCount>
paratype; 
<emphasis id="A7EBEAFCA76CA66EFD9AF9A9FD0D7267" bold="true" box="[628,643,1590,1611]" pageId="10" pageNumber="11">A</emphasis>
, enlarged paraterga with ozopores, lateral view; 
<emphasis id="A7EBEAFCA76CA66EFBD0F9A9FBC37267" bold="true" box="[1086,1101,1590,1611]" pageId="10" pageNumber="11">B</emphasis>
, bisegmented seta; 
<emphasis id="A7EBEAFCA76CA66EFEDBF9C9FEC87240" bold="true" box="[309,326,1622,1644]" pageId="10" pageNumber="11">C</emphasis>
, tegument texture; 
<emphasis id="A7EBEAFCA76CA66EFE12F9C9FD817247" bold="true" box="[508,527,1622,1643]" pageId="10" pageNumber="11">D</emphasis>
, limbus and adjacent parts; 
<emphasis id="A7EBEAFCA76CA66EFCF8F9C9FCAA7247" bold="true" box="[790,804,1622,1643]" pageId="10" pageNumber="11">E</emphasis>
, claw. − Scale bars: A, 0.05 mm, B, D, E, 0.01 mm, C, 0.02 mm.
</paragraph>
</caption>
<paragraph id="952036EEA76DA66FFF57FF38FD077562" blockId="11.[140,1108,167,756]" pageId="11" pageNumber="12">
Sterna usually with a deep, narrow, longitudinal depression between coxae (
<figureCitation id="0DA42A6BA76DA66FFBC9FF38FF2374C8" captionStart="Figure 2" captionStartId="9.[140,212,1590,1612]" captionTargetBox="[140,1108,407,1570]" captionTargetId="figure@9.[140,1108,407,1570]" captionTargetPageId="9" captionText="Figure 2. Eutrichodesmus basalis sp. n., ♀ paratype; A, anterior part of body, frontodorsal view; B, head, front view; C, antenna; D, cross-section of a midbody segment, caudal view; E, posterior part of body, ventral view; F, legs, ventral view. − Scale bars: A, B, D, E, 0.1 mm &amp; C, F, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3769071" httpUri="https://zenodo.org/record/3769071/files/figure.png" pageId="11" pageNumber="12">Fig. 2F</figureCitation>
), but sterna between coxae 6, 7 and 9 much wider (
<figureCitation id="0DA42A6BA76DA66FFCE0FF55FCE474C8" box="[782,874,202,228]" captionStart="Figure 4" captionStartId="11.[140,212,1622,1644]" captionTargetBox="[200,1045,807,1593]" captionTargetId="figure@11.[194,1054,799,1602]" captionTargetPageId="11" captionText="Figure 4. Eutrichodesmus basalis sp. n., ♂ paratype; A, leg 9, caudal view; B &amp; C, left gonopod, mesal and lateral views, respectively (dp = distofemoral process). − Scale bar: A, 0.2 mm, B, C, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3769075" httpUri="https://zenodo.org/record/3769075/files/figure.png" pageId="11" pageNumber="12">Fig. 4A</figureCitation>
). Gonopod aperture transverse-oval, relatively small, far from reaching lateral sides of segment 7. Legs very short; tarsal segment longest; claw usual, simple, very slightly curved ventrad; some setae sparsely microdenticulate (
<figureCitation id="0DA42A6BA76DA66FFE1BFEACFDC27562" box="[501,588,307,334]" captionStart="Figure 2" captionStartId="9.[140,212,1590,1612]" captionTargetBox="[140,1108,407,1570]" captionTargetId="figure@9.[140,1108,407,1570]" captionTargetPageId="9" captionText="Figure 2. Eutrichodesmus basalis sp. n., ♀ paratype; A, anterior part of body, frontodorsal view; B, head, front view; C, antenna; D, cross-section of a midbody segment, caudal view; E, posterior part of body, ventral view; F, legs, ventral view. − Scale bars: A, B, D, E, 0.1 mm &amp; C, F, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3769071" httpUri="https://zenodo.org/record/3769071/files/figure.png" pageId="11" pageNumber="12">Figs 2F</figureCitation>
; 
<figureCitation id="0DA42A6BA76DA66FFDB6FEABFDF57562" box="[600,635,308,334]" captionStart="Figure 3" captionStartId="10.[140,212,1590,1612]" captionTargetBox="[140,1108,826,1570]" captionTargetId="figure@10.[140,1108,825,1570]" captionTargetPageId="10" captionText="Figure 3. Eutrichodesmus basalis sp. n., ♀ paratype; A, enlarged paraterga with ozopores, lateral view; B, bisegmented seta; C, tegument texture; D, limbus and adjacent parts; E, claw. − Scale bars: A, 0.05 mm, B, D, E, 0.01 mm, C, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.3769073" httpUri="https://zenodo.org/record/3769073/files/figure.png" pageId="11" pageNumber="12">3E</figureCitation>
).
</paragraph>
<paragraph id="952036EEA76DA66FFF57FEC8FBC075D1" blockId="11.[140,1108,167,756]" pageId="11" pageNumber="12">
Gonopods (
<figureCitation id="0DA42A6BA76DA66FFED5FEC8FE3F755D" box="[315,433,343,369]" captionStart="Figure 4" captionStartId="11.[140,212,1622,1644]" captionTargetBox="[200,1045,807,1593]" captionTargetId="figure@11.[194,1054,799,1602]" captionTargetPageId="11" captionText="Figure 4. Eutrichodesmus basalis sp. n., ♂ paratype; A, leg 9, caudal view; B &amp; C, left gonopod, mesal and lateral views, respectively (dp = distofemoral process). − Scale bar: A, 0.2 mm, B, C, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3769075" httpUri="https://zenodo.org/record/3769075/files/figure.png" pageId="11" pageNumber="12">Figs 4B, C</figureCitation>
) very simple. Coxae subquadrate, large, microtuberculate and abundantly setose ventrolaterally, with a conspicuous triangular lobe frontolaterally. Telopodite much longer than coxite, slender throughout, setose in its basal half, with a conspicuous, only apically denticulate, lateral, distofemoral process (dp) at about midway, with a hairpad in distal third, but seminal groove unexpectedly terminating subapically.
</paragraph>
</subSubSection>
<subSubSection id="DD856565A76DA668FF57FD98FBCC75B8" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="discussion">
<paragraph id="952036EEA76DA668FF57FD98FD217506" blockId="11.[140,1108,167,756]" lastBlockId="12.[140,1108,167,404]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<emphasis id="A7EBEAFCA76DA66FFF57FD98FEA4760D" bold="true" box="[185,298,519,545]" pageId="11" pageNumber="12">
<emphasis id="A7EBEAFCA76DA66FFF57FD98FEAB760D" box="[185,293,519,545]" inLineHeading="true" pageId="11" pageNumber="12" reason="1">Remarks</emphasis>
.
</emphasis>
This remarkable species somewhat bridges the former 
<taxonomicName id="529F4D6DA76DA66FFC4DFD98FBDA760D" authorityName="Cook" authorityYear="1895" box="[931,1108,519,545]" class="Diplopoda" family="Haplodesmidae" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="family">Haplodesmidae</taxonomicName>
and 
<taxonomicName id="529F4D6DA76DA66FFF50FDB5FEF37668" authorityName="Cook" authorityYear="1896" box="[190,381,554,580]" class="Diplopoda" family="Doratodesmidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="family">Doratodesmidae</taxonomicName>
. Hence the quotation marks which are used for “haplodesmids” and “doratodesmids” above and hereafter. In addition to showing the tergal trichome of peculiar, bisegmented setae and the unusually simple gonopods so characteristic of 
<taxonomicName id="529F4D6DA76DA66FFF62FD0CFEB87681" authorityName="Pocock" authorityYear="1889" box="[140,310,659,685]" class="Diplopoda" family="Haplodesmidae" genus="Cylindrodesmus" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="A7EBEAFCA76DA66FFF62FD0CFEB87681" box="[140,310,659,685]" italics="true" pageId="11" pageNumber="12">Cylindrodesmus</emphasis>
</taxonomicName>
(both features partly shared also with several true “doratodesmids”), it also, most importantly, has the body only capable of incomplete volvation, due to the relatively very short paraterga. Moreover, the ozopores are borne on small porostele-like tubercles, a feature characteristic of most “haplodesmids”, but only rarely encountered among “doratodesmids”. At least the incomplete volvation and the simple biramous gonopods seem to be relatively primitive characters amongst the more advanced “doratodesmids” in which volvation is already perfect.
</paragraph>
<caption id="C1E06666A76DA66FFF62F9C9FC6672A0" ID-DOI="http://doi.org/10.5281/zenodo.3769075" ID-Zenodo-Dep="3769075" httpUri="https://zenodo.org/record/3769075/files/figure.png" pageId="11" pageNumber="12" startId="11.[140,212,1622,1644]" targetBox="[200,1045,807,1593]" targetPageId="11">
<paragraph id="952036EEA76DA66FFF62F9C9FC6672A0" blockId="11.[140,1108,1532,1676]" pageId="11" pageNumber="12">
<emphasis id="A7EBEAFCA76DA66FFF62F9C9FF637240" bold="true" box="[140,237,1622,1644]" pageId="11" pageNumber="12">Figure 4.</emphasis>
<taxonomicName id="529F4D6DA76DA66FFF1FF9C9FE347247" authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel, 2009" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; VandenSpiegel" authorityYear="2009" box="[241,442,1622,1643]" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="basalis" status="sp. n.">
<emphasis id="A7EBEAFCA76DA66FFF1FF9C9FE347247" box="[241,442,1622,1643]" italics="true" pageId="11" pageNumber="12">Eutrichodesmus basalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BCD85787A76DA66FFE50F9C8FE7F7240" box="[446,497,1623,1644]" pageId="11" pageNumber="12" rank="species">sp. n.</taxonomicNameLabel>
, 
<specimenCount id="8399FD67A76DA66FFE14F9C8FD877241" box="[506,521,1623,1645]" pageId="11" pageNumber="12">♂</specimenCount>
paratype; 
<emphasis id="A7EBEAFCA76DA66FFD85F9C9FDF47247" bold="true" box="[619,634,1622,1643]" pageId="11" pageNumber="12">A</emphasis>
, leg 9, caudal view; 
<emphasis id="A7EBEAFCA76DA66FFCDEF9C9FCE07240" bold="true" box="[816,878,1622,1644]" pageId="11" pageNumber="12">B &amp; C</emphasis>
, left gonopod, mesal and lateral views, respectively (dp = distofemoral process). − Scale bar: A, 0.2 mm, B, C, 0.1 mm.
</paragraph>
</caption>
<paragraph id="952036EEA76AA668FF57FEACFBCC75B8" blockId="12.[140,1108,167,404]" pageId="12" pageNumber="13">
The new species especially resembles the similarly tuberculate-setose 
<taxonomicName id="529F4D6DA76AA668FC53FEACFE2F755D" authority="Hoffman, 1982" authorityName="Hoffman" authorityYear="1982" class="Diplopoda" family="Haplodesmidae" genus="Dyomerothrix" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="gremialis">
<emphasis id="A7EBEAFCA76AA668FC53FEACFF62755D" italics="true" pageId="12" pageNumber="13">Dyomerothrix gremialis</emphasis>
<bibRefCitation id="F10E4B1FA76AA668FF1CFEC8FE2F755D" author="Hoffman RL" box="[242,417,343,369]" pageId="12" pageNumber="13" pagination="87 - 93" refId="ref19444" refString="Hoffman RL (1982 a) A new genus and species of doratodesmid millipede from Thailand. Archives des Sciences 35 (1): 87 - 93." type="journal article" year="1982">Hoffman, 1982</bibRefCitation>
</taxonomicName>
or 
<taxonomicName id="529F4D6DA76AA668FE2AFEC8FBC0755D" authority="Zhang in Zhang &amp; Wang, 1993" authorityName="Zhang in Zhang &amp; Wang" authorityYear="1993" box="[452,1102,343,369]" class="Diplopoda" family="Haplodesmidae" genus="Parapauroplus" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="monodentus">
<emphasis id="A7EBEAFCA76AA668FE2AFEC8FD6B755D" box="[452,741,343,369]" italics="true" pageId="12" pageNumber="13">Parapauroplus monodentus</emphasis>
Zhang 
<emphasis id="A7EBEAFCA76AA668FCD4FEC8FCDF755C" box="[826,849,343,368]" italics="true" pageId="12" pageNumber="13">in</emphasis>
<bibRefCitation id="F10E4B1FA76AA668FCB8FEC8FBC0755D" author="Zhang &amp; Wang" box="[854,1102,343,369]" firstAuthor="Zhang" pageId="12" pageNumber="13" pagination="205 - 220" refId="ref20260" refString="Zhang Chunzhou, Wang Daqing (1993) Diplopoda in caves of Yunnan - 1. A study of new genera and species of the millipede family Doratodesmidae (Diplopoda: Polydesmida). In: Song Linhua, Ting Huaiyuan (eds) Karst Landscape and Cave Tourism. China Environmental Science Press, Beijing: 205 - 220." type="book chapter" year="1993">Zhang &amp; Wang, 1993</bibRefCitation>
</taxonomicName>
, which also show rather simple gonopods (
<bibRefCitation id="F10E4B1FA76AA668FD8AFEE5FC9175B8" author="Hoffman" box="[612,799,378,404]" firstAuthor="Hoffman" pageId="12" pageNumber="13" pagination="87 - 93" refId="ref19444" refString="Hoffman RL (1982 a) A new genus and species of doratodesmid millipede from Thailand. Archives des Sciences 35 (1): 87 - 93." type="journal article" year="1982" yearSuffix="a">Hoffman 1982a</bibRefCitation>
; 
<bibRefCitation id="F10E4B1FA76AA668FCC5FEE5FBB875B8" author="Zhang and Wang" box="[811,1078,378,404]" firstAuthor="Zhang" pageId="12" pageNumber="13" pagination="205 - 220" refId="ref20260" refString="Zhang Chunzhou, Wang Daqing (1993) Diplopoda in caves of Yunnan - 1. A study of new genera and species of the millipede family Doratodesmidae (Diplopoda: Polydesmida). In: Song Linhua, Ting Huaiyuan (eds) Karst Landscape and Cave Tourism. China Environmental Science Press, Beijing: 205 - 220." type="book chapter" year="1993">Zhang and Wang 1993</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>