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<document id="21053160F6DCAFE91AEC62ECE172D5BA" ID-CLB-Dataset="6318" ID-DOI="10.11646/zootaxa.4970.2.1" ID-GBIF-Dataset="89d3c795-198b-4162-8903-9c6cf3f1e665" ID-ISSN="1175-5326" ID-Zenodo-Dep="4761544" ID-ZooBank="6DBA0C02-9E52-4FD0-8B48-8C7B405713A8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1620984897660" checkinUser="plazi" docAuthor="Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M. &amp; Brescovit, Antonio Domingos" docDate="2021" docId="052487CFFFD3E53846C6B7358F2AFE7E" docLanguage="en" docName="zootaxa.4970.2.1.pdf" docOrigin="Zootaxa 4970 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gangugia boitata Bouzan &amp; Iniesta &amp; Brescovit 2021, n. sp." docType="treatment" docUuid="47505136-DC93-40BB-A9A8-E0821B2F7B03" docUuidSource="ZooBank" docVersion="6" lastPageNumber="227" masterDocId="F91DFFB7FFCAE5224651B2518D76FFF7" masterDocTitle="Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae)" masterLastPageNumber="256" masterPageNumber="201" pageNumber="226" updateTime="1698939836712" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="091563BD9AC68855016183BC00BCA118">Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae)</mods:title>
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<mods:namePart id="8762A674DC58A8576E9F7550C1AAE479">Bouzan, Rodrigo Salvador</mods:namePart>
<mods:affiliation id="AB51BE71D1F82CAC3E05F3A293AF901D">Pós-graduação em Zoologia, Instituto de Biociências, Universidade de São Paulo, São Paulo, Brasil.</mods:affiliation>
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<mods:namePart id="F8D6FC813FD97EA86A89678E6172FB2D">Iniesta, Luiz Felipe M.</mods:namePart>
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<mods:affiliation id="5E484F71E12C50140723A4A83D3A1F56">Pós-graduação em Zoologia, Instituto de Biociências, Universidade de São Paulo, São Paulo, Brasil. &amp; luiz-moretti @ hotmail. com; https: // orcid. org / 0000 - 0002 - 0529 - 4162</mods:affiliation>
<mods:nameIdentifier id="88D544E5DC7EFE184BE9A4AB87D1A9BC" type="email">luiz-moretti@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="6E919D041FECFCE54113BCDF7B80DBBB">Brescovit, Antonio Domingos</mods:namePart>
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<mods:affiliation id="30B02C7BA51755F594396E8642EF09BC">antonio. brescovit @ butantan. gov. br; https: // orcid. org / 0000 - 0002 - 1511 - 5324</mods:affiliation>
<mods:nameIdentifier id="6A409C1690CD145CA9B758771A018A53" type="email">antonio.brescovit@butantan.gov.br</mods:nameIdentifier>
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<treatment id="052487CFFFD3E53846C6B7358F2AFE7E" ID-DOI="http://doi.org/10.5281/zenodo.4761596" ID-GBIF-Taxon="182486292" ID-Zenodo-Dep="4761596" LSID="urn:lsid:zoobank.org:act:47505136-DC93-40BB-A9A8-E0821B2F7B03" httpUri="http://treatment.plazi.org/id/052487CFFFD3E53846C6B7358F2AFE7E" lastPageId="26" lastPageNumber="227" pageId="25" pageNumber="226">
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<paragraph id="8D3236D9FFD3E53B46C6B7358CC2FA89" blockId="25.[151,886,1379,1472]" box="[151,436,1379,1406]" pageId="25" pageNumber="226">
<heading id="D67A81B5FFD3E53B46C6B7358CC2FA89" bold="true" box="[151,436,1379,1406]" fontSize="11" level="1" pageId="25" pageNumber="226" reason="1">
<emphasis id="BFF9EACBFFD3E53B46C6B7358CC2FA89" bold="true" box="[151,436,1379,1406]" pageId="25" pageNumber="226">
<taxonomicName id="4A8D4D5AFFD3E53B46C6B7358C1CFA89" ID-CoL="8H7S3" authority="Bouzan &amp; Iniesta &amp; Brescovit, 2021" authorityName="Bouzan &amp; Iniesta &amp; Brescovit" authorityYear="2021" box="[151,362,1379,1406]" class="Diplopoda" family="Chelodesmidae" genus="Gangugia" kingdom="Animalia" order="Polydesmida" pageId="25" pageNumber="226" phylum="Arthropoda" rank="species" species="boitata" status="sp. nov.">
<emphasis id="BFF9EACBFFD3E53B46C6B7358C1CFA89" bold="true" box="[151,362,1379,1406]" italics="true" pageId="25" pageNumber="226">Gangugia boitata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A4CA57B0FFD3E53B4723B7358CC2FA89" box="[370,436,1380,1406]" pageId="25" pageNumber="226" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C5976552FFD3E53B46C6B7D98E00FA37" pageId="25" pageNumber="226" type="description">
<paragraph id="8D3236D9FFD3E53B46C6B7D98C4EFA55" blockId="25.[151,886,1379,1472]" box="[151,312,1416,1442]" pageId="25" pageNumber="226">
<figureCitation id="15B62A5CFFD3E53B46C6B7D98C66FA55" box="[151,272,1416,1442]" captionStart="FIGURE 14" captionStartId="27.[151,250,1704,1728]" captionTargetBox="[161,1426,181,1680]" captionTargetId="figure-20@27.[161,1426,181,1680]" captionTargetPageId="27" captionText="FIGURE 14. Gangugia boitata n. sp. (holotype, IBSP 1061), dorsal views of anterior (A), midbody (B) and posterior (C) body rings. Scale bars: 2 mm. Left gonopod. (D), mesal view; (E), ventral view; (F), ectal view. Scale bars: 0,5 mm. Abbreviations: C, cingulum; PfP, prefemoral process; S, solenomere." figureDoi="http://doi.org/10.5281/zenodo.4761594" httpUri="https://zenodo.org/record/4761594/files/figure.png" pageId="25" pageNumber="226">Figures 14</figureCitation>
,
<figureCitation id="15B62A5CFFD3E53B474DB7D98C4EFA55" box="[284,312,1416,1442]" captionStart="FIGURE 18" captionStartId="32.[151,250,2002,2026]" captionTargetBox="[166,1421,849,1979]" captionTargetId="figure-319@32.[166,1421,849,1979]" captionTargetPageId="32" captionText="FIGURE 18. Distribution map of Gangugia Schubart, 1947 species." figureDoi="http://doi.org/10.5281/zenodo.4761611" httpUri="https://zenodo.org/record/4761611/files/figure.png" pageId="25" pageNumber="226">18</figureCitation>
</paragraph>
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<uri id="F91C3ADBFFD3E53B46C6B7F98E00FA37" box="[151,886,1448,1472]" pageId="25" pageNumber="226">
urn:lsid:zoobank.org:act:
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<subSubSection id="C5976552FFD3E53B46C6B7B98E56F9D1" pageId="25" pageNumber="226" type="materials_examined">
<paragraph id="8D3236D9FFD3E53B46C6B7B98E56F9D1" blockId="25.[151,1437,1512,2043]" pageId="25" pageNumber="226">
<emphasis id="BFF9EACBFFD3E53B46C6B7B98C3CF9F5" bold="true" box="[151,330,1512,1538]" pageId="25" pageNumber="226">Type material.</emphasis>
<materialsCitation id="3DE53C84FFD3E53B4704B7B98E6AF9D1" ID-GBIF-Occurrence="3125622330" collectingDate="1992-10" collectionCode="IBSP" collectorName="Parque Nacional das Emas &amp; M. Barroso" country="Brazil" latitude="-18.366945" location="Mineiros" longLatPrecision="21" longitude="-52.79972" pageId="25" pageNumber="226" specimenCode="IBSP 1061" specimenCount="1" stateProvince="Goias" typeStatus="holotype">
Male
<typeStatus id="5236887BFFD3E53B47CAB7B98C8BF9F5" box="[411,509,1512,1538]" pageId="25" pageNumber="226" type="holotype">holotype</typeStatus>
from
<collectorName id="2078530FFFD3E53B441AB7B88EF2F9F4" box="[587,900,1512,1539]" pageId="25" pageNumber="226">Parque Nacional das Emas</collectorName>
(
<geoCoordinate id="E8B9501EFFD3E53B45C8B7B98956F9F5" box="[921,1056,1512,1538]" degrees="18" direction="south" minutes="22" orientation="latitude" pageId="25" pageNumber="226" precision="15" seconds="01" value="-18.366945">18°2201”S</geoCoordinate>
,
<geoCoordinate id="E8B9501EFFD3E53B4260B7B989B5F9F4" box="[1073,1219,1512,1539]" degrees="52" direction="west" minutes="47" orientation="longitude" pageId="25" pageNumber="226" precision="15" seconds="59" value="-52.79972">52°4759”W</geoCoordinate>
),
<location id="88526002FFD3E53B428DB7B98832F9F5" LSID="urn:lsid:plazi:treatment:052487CFFFD3E53846C6B7358F2AFE7E:88526002FFD3E53B428DB7B98832F9F5" box="[1244,1348,1512,1538]" country="Brazil" latitude="-18.366945" longLatPrecision="21" longitude="-52.79972" name="Mineiros" pageId="25" pageNumber="226" stateProvince="Goias">Mineiros</location>
,
<collectingRegion id="4F49F83BFFD3E53B4305B7B988EEF9F5" box="[1364,1432,1512,1538]" country="Brazil" name="Goias" pageId="25" pageNumber="226">Goiás</collectingRegion>
,
<collectingCountry id="F59A7649FFD3E53B46C6B45D8DA8F9D1" box="[151,222,1548,1574]" name="Brazil" pageId="25" pageNumber="226">Brazil</collectingCountry>
,
<date id="F9331019FFD3E53B46BBB45D8C36F9D1" box="[234,320,1548,1574]" pageId="25" pageNumber="226" value="1992-10">
<collectingDate id="E977E9F1FFD3E53B46BBB45D8C36F9D1" box="[234,320,1548,1574]" pageId="25" pageNumber="226" value="1992-10">X.1992</collectingDate>
</date>
,
<collectorName id="2078530FFFD3E53B471AB45C8CBDF9D0" box="[331,459,1549,1575]" pageId="25" pageNumber="226">M. Barroso</collectorName>
coll., deposited in
<specimenCode id="DD2B9EA2FFD3E53B44CEB45D8E6AF9D1" box="[671,796,1548,1574]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="25" pageNumber="226">IBSP 1061</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C5976552FFD3E53B4696B4618FBCF941" pageId="25" pageNumber="226" type="diagnosis">
<paragraph id="8D3236D9FFD3E53B4696B4618FBCF941" blockId="25.[151,1437,1512,2043]" pageId="25" pageNumber="226">
<emphasis id="BFF9EACBFFD3E53B4696B4618C4CF9BD" bold="true" box="[199,314,1584,1610]" pageId="25" pageNumber="226">Diagnosis</emphasis>
. Adult males differ from all other species of the genus by the combination of the following characters: gonocoxae conspicuously greater than half the size of the telopodite (
<figureCitation id="15B62A5CFFD3E53B4591B405895DF999" box="[960,1067,1620,1646]" captionStart="FIGURE 14" captionStartId="27.[151,250,1704,1728]" captionTargetBox="[161,1426,181,1680]" captionTargetId="figure-20@27.[161,1426,181,1680]" captionTargetPageId="27" captionText="FIGURE 14. Gangugia boitata n. sp. (holotype, IBSP 1061), dorsal views of anterior (A), midbody (B) and posterior (C) body rings. Scale bars: 2 mm. Left gonopod. (D), mesal view; (E), ventral view; (F), ectal view. Scale bars: 0,5 mm. Abbreviations: C, cingulum; PfP, prefemoral process; S, solenomere." figureDoi="http://doi.org/10.5281/zenodo.4761594" httpUri="https://zenodo.org/record/4761594/files/figure.png" pageId="25" pageNumber="226">Fig. 14D</figureCitation>
); absence of indentations in the prefemoral process (
<figureCitation id="15B62A5CFFD3E53B472CB4298C93F965" box="[381,485,1656,1682]" captionStart="FIGURE 14" captionStartId="27.[151,250,1704,1728]" captionTargetBox="[161,1426,181,1680]" captionTargetId="figure-20@27.[161,1426,181,1680]" captionTargetPageId="27" captionText="FIGURE 14. Gangugia boitata n. sp. (holotype, IBSP 1061), dorsal views of anterior (A), midbody (B) and posterior (C) body rings. Scale bars: 2 mm. Left gonopod. (D), mesal view; (E), ventral view; (F), ectal view. Scale bars: 0,5 mm. Abbreviations: C, cingulum; PfP, prefemoral process; S, solenomere." figureDoi="http://doi.org/10.5281/zenodo.4761594" httpUri="https://zenodo.org/record/4761594/files/figure.png" pageId="25" pageNumber="226">Fig. 14D</figureCitation>
); absence of a secondary process of the prefemoral process and presence of a lobe in the solenomere, visible in ventral view (
<figureCitation id="15B62A5CFFD3E53B4409B4CD8FC8F941" box="[600,702,1692,1718]" captionStart="FIGURE 14" captionStartId="27.[151,250,1704,1728]" captionTargetBox="[161,1426,181,1680]" captionTargetId="figure-20@27.[161,1426,181,1680]" captionTargetPageId="27" captionText="FIGURE 14. Gangugia boitata n. sp. (holotype, IBSP 1061), dorsal views of anterior (A), midbody (B) and posterior (C) body rings. Scale bars: 2 mm. Left gonopod. (D), mesal view; (E), ventral view; (F), ectal view. Scale bars: 0,5 mm. Abbreviations: C, cingulum; PfP, prefemoral process; S, solenomere." figureDoi="http://doi.org/10.5281/zenodo.4761594" httpUri="https://zenodo.org/record/4761594/files/figure.png" pageId="25" pageNumber="226">Fig. 14E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C5976552FFD3E5384696B4918CE8FF0E" lastPageId="26" lastPageNumber="227" pageId="25" pageNumber="226" type="description">
<paragraph id="8D3236D9FFD3E5384696B4918993FF22" blockId="25.[151,1437,1512,2043]" lastBlockId="26.[151,1436,151,393]" lastPageId="26" lastPageNumber="227" pageId="25" pageNumber="226">
<emphasis id="BFF9EACBFFD3E53B4696B4918C2FF92D" bold="true" box="[199,345,1728,1754]" pageId="25" pageNumber="226">Description.</emphasis>
Male (
<typeStatus id="5236887BFFD3E53B47FCB4918F6CF92D" box="[429,538,1728,1754]" pageId="25" pageNumber="226" type="holotype">Holotype</typeStatus>
, IBSP 1061). Coloration (long-preserved in 70% ethanol): Head and antennae ochre. Body ocher and paranota tip with the same color as the body (
<figureCitation id="15B62A5CFFD3E53B45DBB4B58961F909" box="[906,1047,1764,1790]" captionStart="FIGURE 14" captionStartId="27.[151,250,1704,1728]" captionTargetBox="[161,1426,181,1680]" captionTargetId="figure-20@27.[161,1426,181,1680]" captionTargetPageId="27" captionText="FIGURE 14. Gangugia boitata n. sp. (holotype, IBSP 1061), dorsal views of anterior (A), midbody (B) and posterior (C) body rings. Scale bars: 2 mm. Left gonopod. (D), mesal view; (E), ventral view; (F), ectal view. Scale bars: 0,5 mm. Abbreviations: C, cingulum; PfP, prefemoral process; S, solenomere." figureDoi="http://doi.org/10.5281/zenodo.4761594" httpUri="https://zenodo.org/record/4761594/files/figure.png" pageId="25" pageNumber="226">Figs 14AC</figureCitation>
). Legs ocher whitish. Telson ocher. Total length: 39.07. Total width: 5.57. Collum, length 2.30, width 5.19. Antennomere lengths (1&gt;7): 0.48; 1.35; 1.10; 1.06; 0.99; 1.00; 0.24. Podomeres lengths (1&gt;7): 0.66; 0.88; 1.50; 0.70; 0.89; 1.10; 0.33. Gonopod aperture, length 1.33, width 2.19. Telson, length 1.05. Gonopod: length 1.58, width 1.68. Gonocoxae: length 0.96, width 1.06. Gonocoxae globose, about 1/2 the size of the telopodite. Telopodite: length 1.44, width 0.74. Prefemoral region short, 1/3 the size of telopodite. Prefemoral process long and falciform, presenting a dorso-basal lobe, starting from an angle of 225°, involving the solenomere in its initial portion, subsequently ascending parallel to the solenomere and partially covering it in ectal view (
<figureCitation id="15B62A5CFFD3E53B4401B5B18FAAF80D" box="[592,732,2016,2042]" captionStart="FIGURE 14" captionStartId="27.[151,250,1704,1728]" captionTargetBox="[161,1426,181,1680]" captionTargetId="figure-20@27.[161,1426,181,1680]" captionTargetPageId="27" captionText="FIGURE 14. Gangugia boitata n. sp. (holotype, IBSP 1061), dorsal views of anterior (A), midbody (B) and posterior (C) body rings. Scale bars: 2 mm. Left gonopod. (D), mesal view; (E), ventral view; (F), ectal view. Scale bars: 0,5 mm. Abbreviations: C, cingulum; PfP, prefemoral process; S, solenomere." figureDoi="http://doi.org/10.5281/zenodo.4761594" httpUri="https://zenodo.org/record/4761594/files/figure.png" pageId="25" pageNumber="226">Figs 14DF</figureCitation>
). Solenomere long and falciform, not reaching the apex of the prefemoral process (
<figureCitation id="15B62A5CFFD0E53847D0B2C68C9FFF46" box="[385,489,151,177]" captionStart="FIGURE 14" captionStartId="27.[151,250,1704,1728]" captionTargetBox="[161,1426,181,1680]" captionTargetId="figure-20@27.[161,1426,181,1680]" captionTargetPageId="27" captionText="FIGURE 14. Gangugia boitata n. sp. (holotype, IBSP 1061), dorsal views of anterior (A), midbody (B) and posterior (C) body rings. Scale bars: 2 mm. Left gonopod. (D), mesal view; (E), ventral view; (F), ectal view. Scale bars: 0,5 mm. Abbreviations: C, cingulum; PfP, prefemoral process; S, solenomere." figureDoi="http://doi.org/10.5281/zenodo.4761594" httpUri="https://zenodo.org/record/4761594/files/figure.png" pageId="26" pageNumber="227">Fig. 14E</figureCitation>
). Cingulum in basal position (
<figureCitation id="15B62A5CFFD0E5384516B2C68ED8FF46" box="[839,942,151,177]" captionStart="FIGURE 14" captionStartId="27.[151,250,1704,1728]" captionTargetBox="[161,1426,181,1680]" captionTargetId="figure-20@27.[161,1426,181,1680]" captionTargetPageId="27" captionText="FIGURE 14. Gangugia boitata n. sp. (holotype, IBSP 1061), dorsal views of anterior (A), midbody (B) and posterior (C) body rings. Scale bars: 2 mm. Left gonopod. (D), mesal view; (E), ventral view; (F), ectal view. Scale bars: 0,5 mm. Abbreviations: C, cingulum; PfP, prefemoral process; S, solenomere." figureDoi="http://doi.org/10.5281/zenodo.4761594" httpUri="https://zenodo.org/record/4761594/files/figure.png" pageId="26" pageNumber="227">Fig. 14F</figureCitation>
). Median portion of the solenomere with a rounded lobe, visible in ventral view (
<figureCitation id="15B62A5CFFD0E5384410B2EA8FD1FF22" box="[577,679,187,213]" captionStart="FIGURE 14" captionStartId="27.[151,250,1704,1728]" captionTargetBox="[161,1426,181,1680]" captionTargetId="figure-20@27.[161,1426,181,1680]" captionTargetPageId="27" captionText="FIGURE 14. Gangugia boitata n. sp. (holotype, IBSP 1061), dorsal views of anterior (A), midbody (B) and posterior (C) body rings. Scale bars: 2 mm. Left gonopod. (D), mesal view; (E), ventral view; (F), ectal view. Scale bars: 0,5 mm. Abbreviations: C, cingulum; PfP, prefemoral process; S, solenomere." figureDoi="http://doi.org/10.5281/zenodo.4761594" httpUri="https://zenodo.org/record/4761594/files/figure.png" pageId="26" pageNumber="227">Fig. 14E</figureCitation>
). Apex of the solenomere sickle-shaped (
<figureCitation id="15B62A5CFFD0E5384222B2EA89AFFF22" box="[1139,1241,187,213]" captionStart="FIGURE 14" captionStartId="27.[151,250,1704,1728]" captionTargetBox="[161,1426,181,1680]" captionTargetId="figure-20@27.[161,1426,181,1680]" captionTargetPageId="27" captionText="FIGURE 14. Gangugia boitata n. sp. (holotype, IBSP 1061), dorsal views of anterior (A), midbody (B) and posterior (C) body rings. Scale bars: 2 mm. Left gonopod. (D), mesal view; (E), ventral view; (F), ectal view. Scale bars: 0,5 mm. Abbreviations: C, cingulum; PfP, prefemoral process; S, solenomere." figureDoi="http://doi.org/10.5281/zenodo.4761594" httpUri="https://zenodo.org/record/4761594/files/figure.png" pageId="26" pageNumber="227">Fig. 14E</figureCitation>
).
</paragraph>
<paragraph id="8D3236D9FFD0E5384696B28E8CE8FF0E" blockId="26.[151,1436,151,393]" box="[199,414,223,249]" pageId="26" pageNumber="227">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C5976552FFD0E5384696B3538E2DFEEA" box="[199,859,258,285]" pageId="26" pageNumber="227" type="distribution">
<paragraph id="8D3236D9FFD0E5384696B3538E2DFEEA" blockId="26.[151,1436,151,393]" box="[199,859,258,285]" pageId="26" pageNumber="227">
<emphasis id="BFF9EACBFFD0E5384696B3538C17FEEB" bold="true" box="[199,353,258,284]" pageId="26" pageNumber="227">Distribution.</emphasis>
Known only from the
<typeStatus id="5236887BFFD0E5384431B3528FE6FEEA" box="[608,656,259,285]" pageId="26" pageNumber="227">type</typeStatus>
locality (
<figureCitation id="15B62A5CFFD0E53844AAB3528E38FEEA" box="[763,846,259,285]" captionStart="FIGURE 18" captionStartId="32.[151,250,2002,2026]" captionTargetBox="[166,1421,849,1979]" captionTargetId="figure-319@32.[166,1421,849,1979]" captionTargetPageId="32" captionText="FIGURE 18. Distribution map of Gangugia Schubart, 1947 species." figureDoi="http://doi.org/10.5281/zenodo.4761611" httpUri="https://zenodo.org/record/4761611/files/figure.png" pageId="26" pageNumber="227">Fig. 18</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C5976552FFD0E5384696B3768F2AFE7E" pageId="26" pageNumber="227" type="etymology">
<paragraph id="8D3236D9FFD0E5384696B3768F2AFE7E" blockId="26.[151,1436,151,393]" pageId="26" pageNumber="227">
<emphasis id="BFF9EACBFFD0E5384696B3768C38FEB6" bold="true" box="[199,334,295,321]" pageId="26" pageNumber="227">Etymology.</emphasis>
The species epithet is a reference of the Brazilian folkloric character “Boitatá”. According to the indigenous legend, Boitatá is a large fire serpent that protects the animals and the forests from people who try to burn the vegetation. Noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>