treatments-xml/data/AF/14/E9/AF14E937FFB5E455FF05FF67FD871779.xml

<document id="B147AD77033C8ABFD82D919FCAEB06A2" ID="10.11646/zootaxa.4168.3.3" ID-DOI="10.11646/zootaxa.4168.3.3" ID-GBIF-Dataset="e637580d-a728-4232-be4e-4965c8f4d9b0" ID-ISSN="1175-5326" ID-Zenodo-Dep="155271" ID-ZooBank="CBAB62EC-4FB6-49BD-94DB-7954E0CF5AD0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1474008133758" checkinUser="plazi" docAuthor="Arias, Andrés" docDate="2016" docId="AF14E937FFB5E455FF05FF67FD871779" docLanguage="en" docName="zootaxa.4168.3.3.pdf" docOrigin="Zootaxa 4168 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Onuphis hanneloreae Arias, 2016, sp. nov." docType="treatment" docVersion="12" lastPageNumber="501" masterDocId="532D914FFFB9E441FF92FFF0FFC2136A" masterDocTitle="Onuphi s and Mooreonuphis (Annelida: Onuphidae) from West Africa with the description of three new species and the reinstatement of O. landanaensis Augener, 1918" masterLastPageNumber="511" masterPageNumber="481" pageNumber="493" updateTime="1698386055454" updateUser="plazi">
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<mods:title id="7CF74744EB8433B39B2A1F0BD743309D">Onuphi s and Mooreonuphis (Annelida: Onuphidae) from West Africa with the description of three new species and the reinstatement of O. landanaensis Augener, 1918</mods:title>
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<mods:namePart id="C1C00A4E0822D817B73FCF45C6E62447">Arias, Andrés</mods:namePart>
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<mods:date id="D04783AB2BFF058398283F73EC185A37">2016</mods:date>
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<paragraph id="27025821FFB5E44DFF05FF67FDC013D8" blockId="12.[151,514,151,211]" box="[151,514,151,178]" pageId="12" pageNumber="493">
<heading id="7C4AEF4DFFB5E44DFF05FF67FDC013D8" bold="true" box="[151,514,151,178]" fontSize="11" level="1" pageId="12" pageNumber="493" reason="1">
<emphasis id="15C98433FFB5E44DFF05FF67FDC013D8" bold="true" box="[151,514,151,178]" pageId="12" pageNumber="493">
<taxonomicName id="E0BD23A2FFB5E44DFF05FF67FE5F13DB" ID-CoL="49QJP" box="[151,413,151,177]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="12" pageNumber="493" phylum="Annelida" rank="species" species="hanneloreae" status="sp. nov.">
<emphasis id="15C98433FFB5E44DFF05FF67FE5F13DB" bold="true" box="[151,413,151,177]" italics="true" pageId="12" pageNumber="493">Onuphis hanneloreae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0EFA3948FFB5E44DFE37FF68FDC013D8" box="[421,514,152,178]" pageId="12" pageNumber="493" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
<paragraph id="27025821FFB5E44DFF05FF4AFE7013B9" blockId="12.[151,514,151,211]" box="[151,434,186,211]" pageId="12" pageNumber="493">
<figureCitation id="BF8644A4FFB5E44DFF05FF4AFEEF13B9" box="[151,301,186,211]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStart-2="FIGURE 9" captionStart-3="FIGURE 10" captionStart-4="FIGURE 11" captionStart-5="FIGURE 12" captionStartId-0="13.[151,250,1953,1975]" captionStartId-1="14.[151,250,1933,1955]" captionStartId-2="15.[151,250,1905,1927]" captionStartId-3="16.[151,250,1917,1939]" captionStartId-4="17.[151,250,2005,2027]" captionStartId-5="18.[151,250,1953,1975]" captionTargetBox-0="[193,1393,193,1930]" captionTargetBox-1="[151,1436,193,1910]" captionTargetBox-2="[235,1385,260,1865]" captionTargetBox-3="[184,1388,193,1874]" captionTargetBox-4="[183,1401,1124,1933]" captionTargetBox-5="[202,1384,193,1930]" captionTargetId-0="figure@13.[193,1393,193,1932]" captionTargetId-1="figure@14.[151,1436,193,1911]" captionTargetId-2="figure@15.[180,1406,163,1902]" captionTargetId-3="figure@16.[184,1388,193,1876]" captionTargetId-4="figure@17.[151,1436,1093,1984]" captionTargetId-5="figure@18.[202,1385,193,1932]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionTargetPageId-2="15" captionTargetPageId-3="16" captionTargetPageId-4="17" captionTargetPageId-5="18" captionText-0="FIGURE 7. Onuphis hanneloreae sp. nov. Holotype A, B (MNCN 16.01 / 17458), paratype C, D (MNCN 16.01 / 17459). Photomicrographs. A, anterior end, dorsal view; B, detailed view of anterior end; C, anterior end, dorsal view; D, anterior end, ventral view." captionText-1="FIGURE 8. Onuphis hanneloreae sp. nov. juvenile. Scanning electron micrographs. A, anterior end, dorsal view; B, detailed view of prostomium; C, detailed view of nuchal Groove; D, anterior modified parapodia, dorsal view. b, branchia; ch, chaetiger; dc, dorsal cirrus; pol, postchaetal lobe; prl, prechaetal lobe; vc, ventral cirrus." captionText-2="FIGURE 9. Onuphis hanneloreae sp. nov. Paratype A-E (MNCN 16.01 / 17459). Line drawings. A, parapodium chaetiger 1, anterior view; B, parapodium median chaetiger, anterior view; C slighly shorter tridentate pseudocompound hook from chaetiger 1; D, bidentate pseudocompound hook from chaetiger 1; E, tridentate pseudocompound hook from chaetiger 1; F, maxillae (from largest juvenile)." captionText-3="FIGURE 10. Onuphis hanneloreae sp. nov. Scanning electron micrographs. A, glandular pad from median chaetiger; B, median chaetigers, antero-dorsal view; C, pseudocompound hooks from chaetiger 1; D, detailed view of a bidentate pseudocompound hook of chaetiger 2; E, detailed view of a quatridentate pseudocompound hook of chaetiger 2; F, pectinate chaetae from chaetiger 10; G, subacicular hooks from chaetiger 12." captionText-4="FIGURE 11. Comparison of the progression of hooks in Onuphis hanneloreae sp. nov. juveniles." captionText-5="FIGURE 12. Onuphis hanneloreae sp. nov. juvenile. Scanning electron micrographs. A, bidentate compound hook; B, median anterior provisional subacicular hook; C, ventral anterior provisional subacicular hook; D, posterior provisional subacicular hook; E, posterior parapodium showing both permanent subacicular hook (top) and provisional subacicular hook (bottom)." httpUri-0="http://zenodo.org/record/155278/files/figure.png" httpUri-1="http://zenodo.org/record/155279/files/figure.png" httpUri-2="http://zenodo.org/record/155280/files/figure.png" httpUri-3="http://zenodo.org/record/155281/files/figure.png" httpUri-4="http://zenodo.org/record/155282/files/figure.png" httpUri-5="http://zenodo.org/record/155283/files/figure.png" pageId="12" pageNumber="493">Figures 7–12</figureCitation>
; 
<tableCitation id="6A3F6D9AFFB5E44DFEABFF4AFE5413B9" box="[313,406,186,211]" captionStart="TABLE 1" captionStartId="5.[125,202,650,670]" captionTargetBox="[125,1883,695,1111]" captionText="TABLE 1. Comparison of key features and habitats of the eastern Atlantic species of the Onuphis eremita complex." httpUri="http://table.plazi.org/id/73C208A9FFBCE444FFEFFD7AFB6011F5" pageId="12" pageNumber="493" tableUuid="73C208A9FFBCE444FFEFFD7AFB6011F5">Tables 1</tableCitation>
, 
<tableCitation id="6A3F6D9AFFB5E44DFE36FF4BFE7013B9" box="[420,434,187,211]" captionStart="TABLE 3" captionStartId="19.[125,202,666,689]" captionTargetBox="[147,1878,732,1088]" captionTargetPageId="19" captionText="TABLE 3. Ontogeny of Onuphis hanneloreae sp. nov. juveniles" httpUri="http://table.plazi.org/id/73C208A9FFAAE452FFEFFD6AFD0711DB" pageId="12" pageNumber="493" tableUuid="73C208A9FFAAE452FFEFFD6AFD0711DB">3</tableCitation>
</paragraph>
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<subSubSection id="6FA70BAAFFB5E44DFF05FEF2FC9B1299" pageId="12" pageNumber="493" type="materials_examined">
<paragraph id="27025821FFB5E44DFF05FEF2FCCB1209" blockId="12.[151,1437,258,2011]" pageId="12" pageNumber="493">
<materialsCitation id="97D5527CFFB5E44DFF05FEF2FE761255" ID-GBIF-Occurrence="1313442834" collectionCode="MNCN" latitude="16.6" location="Sal" longLatPrecision="1281" longitude="-22.883333" pageId="12" pageNumber="493" specimenCode="MNCN 16.01" specimenCount="2" stateProvince="Sal" typeStatus="holotype">
<emphasis id="15C98433FFB5E44DFF05FEF2FE031271" bold="true" box="[151,449,258,283]" pageId="12" pageNumber="493">
<typeStatus id="F806E683FFB5E44DFF05FEF2FF101271" box="[151,210,258,283]" pageId="12" pageNumber="493">Type</typeStatus>
material. 
<typeStatus id="F806E683FFB5E44DFEC6FEF2FE031271" box="[340,449,258,283]" pageId="12" pageNumber="493" type="holotype">Holotype</typeStatus>
</emphasis>
(
<specimenCode id="771BF05AFFB5E44DFE46FEF3FDB61271" box="[468,628,258,283]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="12" pageNumber="493">
<collectionCode id="41ACC0E4FFB5E44DFE46FEF3FDE81271" box="[468,554,259,283]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="12" pageNumber="493">MNCN</collectionCode>
16.01
</specimenCode>
/17458), Igrejinha beach (
<geoCoordinate id="42893EE6FFB5E44DFC36FEF2FBC81271" box="[932,1034,258,283]" direction="north" orientation="latitude" pageId="12" pageNumber="493" precision="925" value="16.6">16º 36’N</geoCoordinate>
– 
<geoCoordinate id="42893EE6FFB5E44DFB88FEF2FB591271" box="[1050,1179,258,283]" direction="west" orientation="longitude" pageId="12" pageNumber="493" precision="925" value="-22.883333">22º 53’ W</geoCoordinate>
), intertidal to shallow subtidal, SE of 
<collectingRegion id="E57996C3FFB5E44DFED1FED6FEA51255" box="[323,359,294,319]" country="Cape Verde" name="Sal" pageId="12" pageNumber="493">Sal</collectingRegion>
island
</materialsCitation>
, Cape Verde archipelago, 
<materialsCitation id="97D5527CFFB5E44DFD4BFED7FCCB1209" ID-GBIF-Occurrence="1313442844" collectionCode="MNCN" collectorName="A. Arias July" latitude="16.6" location="eastern Atlantic" longLatPrecision="1281" longitude="-22.883333" pageId="12" pageNumber="493" specimenCode="MNCN 16.01" specimenCount="1" stateProvince="West" typeStatus="paratype">
<collectingRegion id="E57996C3FFB5E44DFD4BFED7FCD21254" box="[729,784,295,318]" country="Cameroon" name="West" pageId="12" pageNumber="493">West</collectingRegion>
Africa, tropical 
<location id="22620EFAFFB5E44DFC58FED7FBBF1255" LSID="urn:lsid:plazi:treatment:AF14E937FFB5E455FF05FF67FD871779:22620EFAFFB5E44DFC58FED7FBBF1255" box="[970,1149,294,319]" name="eastern Atlantic" pageId="12" pageNumber="493" stateProvince="West">eastern Atlantic</location>
, coll. 
<collectorName id="8A483DF7FFB5E44DFB53FED7FA941255" box="[1217,1366,294,319]" pageId="12" pageNumber="493">
A. Arias 
<date id="53037EE1FFB5E44DFAB5FED6FA941255" box="[1319,1366,294,319]" pageId="12" pageNumber="493" value="2012-07">July</date>
</collectorName>
2012; 
<emphasis id="15C98433FFB5E44DFF05FEBAFEC11209" bold="true" box="[151,259,330,355]" pageId="12" pageNumber="493">
<typeStatus id="F806E683FFB5E44DFF05FEBAFEC11209" box="[151,259,330,355]" pageId="12" pageNumber="493" type="paratype">paratype</typeStatus>
</emphasis>
(
<specimenCode id="771BF05AFFB5E44DFE82FEBBFE691209" box="[272,427,330,355]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="12" pageNumber="493">
<collectionCode id="41ACC0E4FFB5E44DFE82FEBBFEA41209" box="[272,358,331,355]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="12" pageNumber="493">MNCN</collectionCode>
16.01
</specimenCode>
/17459) same data as holotype.
</materialsCitation>
</paragraph>
<paragraph id="27025821FFB5E44DFF55FE9EFE2C12C1" blockId="12.[151,1437,258,2011]" pageId="12" pageNumber="493">
<emphasis id="15C98433FFB5E44DFF55FE9EFE6812ED" bold="true" box="[199,426,366,391]" pageId="12" pageNumber="493">Non-type material.</emphasis>
Juvenile specimens of different sizes, same data as holotype, coated with gold for SEM studies (MNCN 16.01/17460).
</paragraph>
<paragraph id="27025821FFB5E44DFF55FE46FC9B1299" blockId="12.[151,1437,258,2011]" pageId="12" pageNumber="493">
<emphasis id="15C98433FFB5E44DFF55FE46FEA512A5" bold="true" box="[199,359,438,463]" pageId="12" pageNumber="493">Type locality.</emphasis>
Intertidal to shallow subtidal sands of Igrejinha beach 16º 36’N– 
<materialsCitation id="97D5527CFFB5E44DFBC7FE46FA5A12A5" ID-GBIF-Occurrence="1313442831" box="[1109,1432,438,463]" location="Sal" longitude="-22.883333" pageId="12" pageNumber="493" specimenCount="1" stateProvince="Sal">
<geoCoordinate id="42893EE6FFB5E44DFBC7FE46FB0C12A4" box="[1109,1230,438,463]" direction="west" orientation="longitude" pageId="12" pageNumber="493" precision="925" value="-22.883333">22º 53’ W</geoCoordinate>
, SE of 
<collectingRegion id="E57996C3FFB5E44DFAB7FE46FA8B12A5" box="[1317,1353,438,463]" country="Cape Verde" name="Sal" pageId="12" pageNumber="493">Sal</collectingRegion>
island
</materialsCitation>
, Cape Verde archipelago, 
<materialsCitation id="97D5527CFFB5E44DFE22FE2BFC971299" ID-GBIF-Occurrence="1313442843" box="[432,853,474,499]" location="eastern Atlantic" pageId="12" pageNumber="493" specimenCount="1" stateProvince="West">
<collectingRegion id="E57996C3FFB5E44DFE22FE2BFE251298" box="[432,487,475,498]" country="Cameroon" name="West" pageId="12" pageNumber="493">West</collectingRegion>
Africa, tropical 
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</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="6FA70BAAFFB5E44DFF55FE0EFC9F1079" pageId="12" pageNumber="493" type="diagnosis">
<paragraph id="27025821FFB5E44DFF55FE0EFC9F1079" blockId="12.[151,1437,258,2011]" pageId="12" pageNumber="493">
<emphasis id="15C98433FFB5E44DFF55FE0EFEF8117D" bold="true" box="[199,314,510,535]" pageId="12" pageNumber="493">Diagnosis</emphasis>
. Prostomium anteriorly extended. Eyespots absent. Palps reaching chaetiger 2, lateral antennae reaching chaetiger 5–6 and median antenna chaetiger 4. Ceratophores long and strongly ringed, palpophores with 10–11 rings, lateral antennophores with 15–16 and median antennophore with 7–10 rings. Subulate ventral cirri in first six chaetigers; distinct subulate postchaetal lobes in first 10–15 chaetigers. Interramal papillae at base of dorsal cirrus absent. Bi-, tri- and quatridentate pseudocompound hooks in first three chaetigers; slender long-appendaged hooks absent. Subacicular hooks from chaetiger 10. Flat pectinate chaetae with 10–11 teeth. Single branchial filaments from chaetiger 1 to 16–21, thereafter number increasing to maximum of three to four. Tube cylindrical in shape, parchment-like and externally covered with sand-grains.
</paragraph>
</subSubSection>
<subSubSection id="6FA70BAAFFB5E44DFF55FCEEFC3414B1" pageId="12" pageNumber="493" type="description">
<paragraph id="27025821FFB5E44DFF55FCEEFBF71031" blockId="12.[151,1437,258,2011]" pageId="12" pageNumber="493">
<emphasis id="15C98433FFB5E44DFF55FCEEFE9B105D" bold="true" box="[199,345,798,823]" pageId="12" pageNumber="493">Description.</emphasis>
Small and slender species. Both 
<typeStatus id="F806E683FFB5E44DFD5DFCEFFD3D105D" box="[719,767,799,823]" pageId="12" pageNumber="493">type</typeStatus>
specimens incomplete. Length of 
<typeStatus id="F806E683FFB5E44DFB16FCEEFB24105D" box="[1156,1254,798,823]" pageId="12" pageNumber="493" type="holotype">holotype</typeStatus>
<quantity id="E045F5C4FFB5E44DFB7FFCEEFA93105C" box="[1261,1361,798,823]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="12" pageNumber="493" unit="mm" value="11.5">11.5 mm</quantity>
for 30 chaetigers, width 
<quantity id="E045F5C4FFB5E44DFECFFCB3FE741030" box="[349,438,835,859]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="12" pageNumber="493" unit="mm" value="0.8">0.8 mm</quantity>
; 
<typeStatus id="F806E683FFB5E44DFE53FCB3FDE01031" box="[449,546,835,859]" pageId="12" pageNumber="493" type="paratype">paratype</typeStatus>
<quantity id="E045F5C4FFB5E44DFDBBFCB2FDBD1030" box="[553,639,834,859]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="12" pageNumber="493" unit="mm" value="8.5">8.5 mm</quantity>
long, 
<quantity id="E045F5C4FFB5E44DFD57FCB3FCD91030" box="[709,795,835,859]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="12" pageNumber="493" unit="mm" value="0.9">0.9 mm</quantity>
wide with 28 chaetigers.
</paragraph>
<paragraph id="27025821FFB5E44DFF55FC96FDA7178D" blockId="12.[151,1437,258,2011]" pageId="12" pageNumber="493">
Colour pattern consisting of: 1–2 circular brown subcutaneous spots on underside of ceratophores of antennae and palps; frontal lips with brown spots at base; three large, sometimes fused, dark brown spots on peristomium, occupying almost entire surface, first six chaetigers almost uniform cream to light brown coloured, subsequent chaetigers with a brown coloured band at posterior margin of segment forming transversal banding pattern (
<figureCitation id="BF8644A4FFB5E44DFAD8FC22FA4C1081" box="[1354,1422,978,1003]" captionStart="FIGURE 7" captionStartId="13.[151,250,1953,1975]" captionTargetBox="[193,1393,193,1930]" captionTargetId="figure@13.[193,1393,193,1932]" captionTargetPageId="13" captionText="FIGURE 7. Onuphis hanneloreae sp. nov. Holotype A, B (MNCN 16.01 / 17458), paratype C, D (MNCN 16.01 / 17459). Photomicrographs. A, anterior end, dorsal view; B, detailed view of anterior end; C, anterior end, dorsal view; D, anterior end, ventral view." httpUri="http://zenodo.org/record/155278/files/figure.png" pageId="12" pageNumber="493">Fig. 7</figureCitation>
). Prostomium subtriangular with pair of subulate frontal lips (
<figureCitation id="BF8644A4FFB5E44DFCD3FC06FC461765" box="[833,900,1014,1039]" captionStart="FIGURE 8" captionStartId="14.[151,250,1933,1955]" captionTargetBox="[151,1436,193,1910]" captionTargetId="figure@14.[151,1436,193,1911]" captionTargetPageId="14" captionText="FIGURE 8. Onuphis hanneloreae sp. nov. juvenile. Scanning electron micrographs. A, anterior end, dorsal view; B, detailed view of prostomium; C, detailed view of nuchal Groove; D, anterior modified parapodia, dorsal view. b, branchia; ch, chaetiger; dc, dorsal cirrus; pol, postchaetal lobe; prl, prechaetal lobe; vc, ventral cirrus." httpUri="http://zenodo.org/record/155279/files/figure.png" pageId="12" pageNumber="493">Fig. 8</figureCitation>
B). Eyespots absent. Palps reaching chaetiger 2 with 9–10 basal rings and longer distal ring in palpophores. Lateral antennae reaching chaetiger 5–6 with 14–15 basal rings in antennophores, median antenna reaching chaetiger 4 with 6–9 basal rings, all three antennophores with slightly longer distal ring (
<figureCitation id="BF8644A4FFB5E44DFE69FB92FD861710" box="[507,580,1122,1147]" captionStart="FIGURE 7" captionStartId="13.[151,250,1953,1975]" captionTargetBox="[193,1393,193,1930]" captionTargetId="figure@13.[193,1393,193,1932]" captionTargetPageId="13" captionText="FIGURE 7. Onuphis hanneloreae sp. nov. Holotype A, B (MNCN 16.01 / 17458), paratype C, D (MNCN 16.01 / 17459). Photomicrographs. A, anterior end, dorsal view; B, detailed view of anterior end; C, anterior end, dorsal view; D, anterior end, ventral view." httpUri="http://zenodo.org/record/155278/files/figure.png" pageId="12" pageNumber="493">Figs 7</figureCitation>
B, 8 
<collectingRegion id="E57996C3FFB5E44DFDE3FB93FD4A1711" box="[625,648,1123,1147]" country="Nigeria" name="Rivers" pageId="12" pageNumber="493">A</collectingRegion>
, B). Palpostyles and antennostyles with irregular longitudinal rows of sensory buds. Nuchal grooves straight with narrow middorsal separation (
<figureCitation id="BF8644A4FFB5E44DFB96FB76FB8E17F5" box="[1028,1100,1158,1183]" captionStart="FIGURE 8" captionStartId="14.[151,250,1933,1955]" captionTargetBox="[151,1436,193,1910]" captionTargetId="figure@14.[151,1436,193,1911]" captionTargetPageId="14" captionText="FIGURE 8. Onuphis hanneloreae sp. nov. juvenile. Scanning electron micrographs. A, anterior end, dorsal view; B, detailed view of prostomium; C, detailed view of nuchal Groove; D, anterior modified parapodia, dorsal view. b, branchia; ch, chaetiger; dc, dorsal cirrus; pol, postchaetal lobe; prl, prechaetal lobe; vc, ventral cirrus." httpUri="http://zenodo.org/record/155279/files/figure.png" pageId="12" pageNumber="493">Fig. 8</figureCitation>
C). Peristomial cirri inserted distally on peristomium slightly lateral to lateral antennae, shorter than length of peristomium not exceeding anterior margin of prostomium (
<figureCitation id="BF8644A4FFB5E44DFD92FB3EFD81178D" box="[512,579,1230,1255]" captionStart="FIGURE 7" captionStartId="13.[151,250,1953,1975]" captionTargetBox="[193,1393,193,1930]" captionTargetId="figure@13.[193,1393,193,1932]" captionTargetPageId="13" captionText="FIGURE 7. Onuphis hanneloreae sp. nov. Holotype A, B (MNCN 16.01 / 17458), paratype C, D (MNCN 16.01 / 17459). Photomicrographs. A, anterior end, dorsal view; B, detailed view of anterior end; C, anterior end, dorsal view; D, anterior end, ventral view." httpUri="http://zenodo.org/record/155278/files/figure.png" pageId="12" pageNumber="493">Fig. 7</figureCitation>
B).
</paragraph>
<paragraph id="27025821FFB5E44DFF55FB02FD881689" blockId="12.[151,1437,258,2011]" pageId="12" pageNumber="493">
Anterior chaetigers (1–3) slightly longer than those following (
<figureCitation id="BF8644A4FFB5E44DFC39FB02FC3A1660" box="[939,1016,1266,1291]" captionStart="FIGURE 7" captionStartId="13.[151,250,1953,1975]" captionTargetBox="[193,1393,193,1930]" captionTargetId="figure@13.[193,1393,193,1932]" captionTargetPageId="13" captionText="FIGURE 7. Onuphis hanneloreae sp. nov. Holotype A, B (MNCN 16.01 / 17458), paratype C, D (MNCN 16.01 / 17459). Photomicrographs. A, anterior end, dorsal view; B, detailed view of anterior end; C, anterior end, dorsal view; D, anterior end, ventral view." httpUri="http://zenodo.org/record/155278/files/figure.png" pageId="12" pageNumber="493">Figs 7</figureCitation>
, 
<figureCitation id="BF8644A4FFB5E44DFB9BFB03FBD51661" box="[1033,1047,1267,1291]" captionStart="FIGURE 8" captionStartId="14.[151,250,1933,1955]" captionTargetBox="[151,1436,193,1910]" captionTargetId="figure@14.[151,1436,193,1911]" captionTargetPageId="14" captionText="FIGURE 8. Onuphis hanneloreae sp. nov. juvenile. Scanning electron micrographs. A, anterior end, dorsal view; B, detailed view of prostomium; C, detailed view of nuchal Groove; D, anterior modified parapodia, dorsal view. b, branchia; ch, chaetiger; dc, dorsal cirrus; pol, postchaetal lobe; prl, prechaetal lobe; vc, ventral cirrus." httpUri="http://zenodo.org/record/155279/files/figure.png" pageId="12" pageNumber="493">8</figureCitation>
<collectingRegion id="E57996C3FFB5E44DFB84FB03FBEC1661" box="[1046,1070,1267,1291]" country="Nigeria" name="Rivers" pageId="12" pageNumber="493">A</collectingRegion>
). First three pairs of parapodia modified, not enlarged, directed slightly anterolaterally, with low prechaetal fold, rounded prechaetal lobe and spindle-shaped postchaetal lobe, latter longer than base of parapodium; digitate dorsal cirrus as long as postchaetal lobe, ventral cirrus shorter than postchaetal lobe (
<figureCitation id="BF8644A4FFB5E44DFD79FAAEFCF6161D" box="[747,820,1374,1399]" captionStart="FIGURE 9" captionStartId="15.[151,250,1905,1927]" captionTargetBox="[235,1385,260,1865]" captionTargetId="figure@15.[180,1406,163,1902]" captionTargetPageId="15" captionText="FIGURE 9. Onuphis hanneloreae sp. nov. Paratype A-E (MNCN 16.01 / 17459). Line drawings. A, parapodium chaetiger 1, anterior view; B, parapodium median chaetiger, anterior view; C slighly shorter tridentate pseudocompound hook from chaetiger 1; D, bidentate pseudocompound hook from chaetiger 1; E, tridentate pseudocompound hook from chaetiger 1; F, maxillae (from largest juvenile)." httpUri="http://zenodo.org/record/155280/files/figure.png" pageId="12" pageNumber="493">Fig. 9</figureCitation>
<collectingRegion id="E57996C3FFB5E44DFCA6FAAFFC89161D" box="[820,843,1375,1399]" country="Nigeria" name="Rivers" pageId="12" pageNumber="493">A</collectingRegion>
). Subulate ventral cirri in six chaetigers, pad-like thereafter (
<figureCitation id="BF8644A4FFB5E44DFE86FA72FE9816F1" box="[276,346,1410,1435]" captionStart="FIGURE 7" captionStartId="13.[151,250,1953,1975]" captionTargetBox="[193,1393,193,1930]" captionTargetId="figure@13.[193,1393,193,1932]" captionTargetPageId="13" captionText="FIGURE 7. Onuphis hanneloreae sp. nov. Holotype A, B (MNCN 16.01 / 17458), paratype C, D (MNCN 16.01 / 17459). Photomicrographs. A, anterior end, dorsal view; B, detailed view of anterior end; C, anterior end, dorsal view; D, anterior end, ventral view." httpUri="http://zenodo.org/record/155278/files/figure.png" pageId="12" pageNumber="493">Fig. 7</figureCitation>
D). Ventral glandular pads with irregular cuticular pore pattern (
<figureCitation id="BF8644A4FFB5E44DFBD2FA72FB5616F1" box="[1088,1172,1410,1435]" captionStart="FIGURE 10" captionStartId="16.[151,250,1917,1939]" captionTargetBox="[184,1388,193,1874]" captionTargetId="figure@16.[184,1388,193,1876]" captionTargetPageId="16" captionText="FIGURE 10. Onuphis hanneloreae sp. nov. Scanning electron micrographs. A, glandular pad from median chaetiger; B, median chaetigers, antero-dorsal view; C, pseudocompound hooks from chaetiger 1; D, detailed view of a bidentate pseudocompound hook of chaetiger 2; E, detailed view of a quatridentate pseudocompound hook of chaetiger 2; F, pectinate chaetae from chaetiger 10; G, subacicular hooks from chaetiger 12." httpUri="http://zenodo.org/record/155281/files/figure.png" pageId="12" pageNumber="493">Fig. 10</figureCitation>
<collectingRegion id="E57996C3FFB5E44DFB06FA73FB6916F1" box="[1172,1195,1411,1435]" country="Nigeria" name="Rivers" pageId="12" pageNumber="493">A</collectingRegion>
). Postchaetal lobe as distinct subulate lobe in first 10–15 chaetigers, becoming smaller and conical knobs in shape thereafter. Interramal papillae at base of dorsal cirrus absent.
</paragraph>
<paragraph id="27025821FFB5E44DFF55FA1EFCDC1525" blockId="12.[151,1437,258,2011]" pageId="12" pageNumber="493">
Branchiae as single filament from first chaetiger to chaetiger 
<quantity id="E045F5C4FFB5E44DFCE2FA1FFC6A156D" box="[880,936,1518,1543]" metricMagnitude="-1" metricUnit="m" metricValue="5.334" pageId="12" pageNumber="493" unit="in" value="21.0">21 in</quantity>
<typeStatus id="F806E683FFB5E44DFC3DFA1EFBD6156D" box="[943,1044,1518,1543]" pageId="12" pageNumber="493" type="holotype">holotype</typeStatus>
, to chaetiger 16 (right) –17 (left) in 
<typeStatus id="F806E683FFB5E44DFF05F9E3FF351541" box="[151,247,1555,1579]" pageId="12" pageNumber="493" type="paratype">paratype</typeStatus>
(
<figureCitation id="BF8644A4FFB5E44DFE94F9E2FE8B1541" box="[262,329,1554,1579]" captionStart="FIGURE 9" captionStartId="15.[151,250,1905,1927]" captionTargetBox="[235,1385,260,1865]" captionTargetId="figure@15.[180,1406,163,1902]" captionTargetPageId="15" captionText="FIGURE 9. Onuphis hanneloreae sp. nov. Paratype A-E (MNCN 16.01 / 17459). Line drawings. A, parapodium chaetiger 1, anterior view; B, parapodium median chaetiger, anterior view; C slighly shorter tridentate pseudocompound hook from chaetiger 1; D, bidentate pseudocompound hook from chaetiger 1; E, tridentate pseudocompound hook from chaetiger 1; F, maxillae (from largest juvenile)." httpUri="http://zenodo.org/record/155280/files/figure.png" pageId="12" pageNumber="493">Fig. 9</figureCitation>
<collectingRegion id="E57996C3FFB5E44DFEDBF9E3FEA21541" box="[329,352,1555,1579]" country="Nigeria" name="Rivers" pageId="12" pageNumber="493">A</collectingRegion>
), usually longer than dorsal cirri, thereafter number of filaments increasing to maximum of three to four (
<figureCitation id="BF8644A4FFB5E44DFF63F9C6FEFB1525" box="[241,313,1590,1615]" captionStart="FIGURE 9" captionStartId="15.[151,250,1905,1927]" captionTargetBox="[235,1385,260,1865]" captionTargetId="figure@15.[180,1406,163,1902]" captionTargetPageId="15" captionText="FIGURE 9. Onuphis hanneloreae sp. nov. Paratype A-E (MNCN 16.01 / 17459). Line drawings. A, parapodium chaetiger 1, anterior view; B, parapodium median chaetiger, anterior view; C slighly shorter tridentate pseudocompound hook from chaetiger 1; D, bidentate pseudocompound hook from chaetiger 1; E, tridentate pseudocompound hook from chaetiger 1; F, maxillae (from largest juvenile)." httpUri="http://zenodo.org/record/155280/files/figure.png" pageId="12" pageNumber="493">Figs 9</figureCitation>
B, 10B), being present to end of fragments.
</paragraph>
<paragraph id="27025821FFB5E44DFF55F9AAFC3414B1" blockId="12.[151,1437,258,2011]" pageId="12" pageNumber="493">
Aciculae yellowish with pointed tips, generally three per parapodium. Hooded pseudocompound hooks in first three chaetigers. First three parapodia with following chaetal complement going from superior to inferior part of chaetal fan: one to two simple chaetae, three protruding distal tips of aciculae and four to five pseudocompound hooks with bi-, tri- (
<figureCitation id="BF8644A4FFB5E44DFE16F936FE0A15B5" box="[388,456,1734,1759]" captionStart="FIGURE 9" captionStartId="15.[151,250,1905,1927]" captionTargetBox="[235,1385,260,1865]" captionTargetId="figure@15.[180,1406,163,1902]" captionTargetPageId="15" captionText="FIGURE 9. Onuphis hanneloreae sp. nov. Paratype A-E (MNCN 16.01 / 17459). Line drawings. A, parapodium chaetiger 1, anterior view; B, parapodium median chaetiger, anterior view; C slighly shorter tridentate pseudocompound hook from chaetiger 1; D, bidentate pseudocompound hook from chaetiger 1; E, tridentate pseudocompound hook from chaetiger 1; F, maxillae (from largest juvenile)." httpUri="http://zenodo.org/record/155280/files/figure.png" pageId="12" pageNumber="493">Fig. 9</figureCitation>
C–E) and quatridentate tips (with two little denticles protuding directly from base of second distal tooth) (
<figureCitation id="BF8644A4FFB5E44DFE12F91AFE131469" box="[384,465,1770,1795]" captionStart="FIGURE 10" captionStartId="16.[151,250,1917,1939]" captionTargetBox="[184,1388,193,1874]" captionTargetId="figure@16.[184,1388,193,1876]" captionTargetPageId="16" captionText="FIGURE 10. Onuphis hanneloreae sp. nov. Scanning electron micrographs. A, glandular pad from median chaetiger; B, median chaetigers, antero-dorsal view; C, pseudocompound hooks from chaetiger 1; D, detailed view of a bidentate pseudocompound hook of chaetiger 2; E, detailed view of a quatridentate pseudocompound hook of chaetiger 2; F, pectinate chaetae from chaetiger 10; G, subacicular hooks from chaetiger 12." httpUri="http://zenodo.org/record/155281/files/figure.png" pageId="12" pageNumber="493">Fig. 10</figureCitation>
E). All hooks with falcate hoods (
<figureCitation id="BF8644A4FFB5E44DFCD4F91AFC541469" box="[838,918,1770,1795]" captionStart="FIGURE 10" captionStartId="16.[151,250,1917,1939]" captionTargetBox="[184,1388,193,1874]" captionTargetId="figure@16.[184,1388,193,1876]" captionTargetPageId="16" captionText="FIGURE 10. Onuphis hanneloreae sp. nov. Scanning electron micrographs. A, glandular pad from median chaetiger; B, median chaetigers, antero-dorsal view; C, pseudocompound hooks from chaetiger 1; D, detailed view of a bidentate pseudocompound hook of chaetiger 2; E, detailed view of a quatridentate pseudocompound hook of chaetiger 2; F, pectinate chaetae from chaetiger 10; G, subacicular hooks from chaetiger 12." httpUri="http://zenodo.org/record/155281/files/figure.png" pageId="12" pageNumber="493">Fig. 10</figureCitation>
D) and of almost equal thickness and length of appendages (
<figureCitation id="BF8644A4FFB5E44DFEBDF8FEFEBE144D" box="[303,380,1806,1831]" captionStart="FIGURE 9" captionStartId="15.[151,250,1905,1927]" captionTargetBox="[235,1385,260,1865]" captionTargetId="figure@15.[180,1406,163,1902]" captionTargetPageId="15" captionText="FIGURE 9. Onuphis hanneloreae sp. nov. Paratype A-E (MNCN 16.01 / 17459). Line drawings. A, parapodium chaetiger 1, anterior view; B, parapodium median chaetiger, anterior view; C slighly shorter tridentate pseudocompound hook from chaetiger 1; D, bidentate pseudocompound hook from chaetiger 1; E, tridentate pseudocompound hook from chaetiger 1; F, maxillae (from largest juvenile)." httpUri="http://zenodo.org/record/155280/files/figure.png" pageId="12" pageNumber="493">Figs 9</figureCitation>
C–E; 10C). Slender long-appendaged hooks absent. Pseudocompound hooks replaced by limbate chaetae from chaetiger 4. Flat pectinate chaetae with 10–11 teeth, two per parapodium (
<figureCitation id="BF8644A4FFB5E44DFB6EF8C2FA921421" box="[1276,1360,1842,1867]" captionStart="FIGURE 10" captionStartId="16.[151,250,1917,1939]" captionTargetBox="[184,1388,193,1874]" captionTargetId="figure@16.[184,1388,193,1876]" captionTargetPageId="16" captionText="FIGURE 10. Onuphis hanneloreae sp. nov. Scanning electron micrographs. A, glandular pad from median chaetiger; B, median chaetigers, antero-dorsal view; C, pseudocompound hooks from chaetiger 1; D, detailed view of a bidentate pseudocompound hook of chaetiger 2; E, detailed view of a quatridentate pseudocompound hook of chaetiger 2; F, pectinate chaetae from chaetiger 10; G, subacicular hooks from chaetiger 12." httpUri="http://zenodo.org/record/155281/files/figure.png" pageId="12" pageNumber="493">Fig. 10</figureCitation>
F) and hooded bidentate subacicular hooks from chaetiger 10 (
<figureCitation id="BF8644A4FFB5E44DFC8DF8A6FCB61405" box="[799,884,1878,1903]" captionStart="FIGURE 10" captionStartId="16.[151,250,1917,1939]" captionTargetBox="[184,1388,193,1874]" captionTargetId="figure@16.[184,1388,193,1876]" captionTargetPageId="16" captionText="FIGURE 10. Onuphis hanneloreae sp. nov. Scanning electron micrographs. A, glandular pad from median chaetiger; B, median chaetigers, antero-dorsal view; C, pseudocompound hooks from chaetiger 1; D, detailed view of a bidentate pseudocompound hook of chaetiger 2; E, detailed view of a quatridentate pseudocompound hook of chaetiger 2; F, pectinate chaetae from chaetiger 10; G, subacicular hooks from chaetiger 12." httpUri="http://zenodo.org/record/155281/files/figure.png" pageId="12" pageNumber="493">Fig. 10</figureCitation>
G). Mandibles with transparent poorly calcified cutting plates and slender shafts. Maxillae (
<figureCitation id="BF8644A4FFB5E44DFD0DF88AFD2514F9" box="[671,743,1914,1939]" captionStart="FIGURE 9" captionStartId="15.[151,250,1905,1927]" captionTargetBox="[235,1385,260,1865]" captionTargetId="figure@15.[180,1406,163,1902]" captionTargetPageId="15" captionText="FIGURE 9. Onuphis hanneloreae sp. nov. Paratype A-E (MNCN 16.01 / 17459). Line drawings. A, parapodium chaetiger 1, anterior view; B, parapodium median chaetiger, anterior view; C slighly shorter tridentate pseudocompound hook from chaetiger 1; D, bidentate pseudocompound hook from chaetiger 1; E, tridentate pseudocompound hook from chaetiger 1; F, maxillae (from largest juvenile)." httpUri="http://zenodo.org/record/155280/files/figure.png" pageId="12" pageNumber="493">Fig. 9</figureCitation>
F) weakly sclerotised; maxillary formula (based on largest juvenile): 
<collectingCountry id="5FAA18B1FFB5E44DFE9EF86FFEF114DC" box="[268,307,1951,1974]" name="Mexico" pageId="12" pageNumber="493">Mx</collectingCountry>
I = 1 + 1, 
<collectingCountry id="5FAA18B1FFB5E44DFE28F86FFE2314DC" box="[442,481,1951,1974]" name="Mexico" pageId="12" pageNumber="493">Mx</collectingCountry>
II = 7 + 7, 
<collectingCountry id="5FAA18B1FFB5E44DFDE0F86FFD5B14DC" box="[626,665,1951,1974]" name="Mexico" pageId="12" pageNumber="493">Mx</collectingCountry>
III = 8 + 0, 
<collectingCountry id="5FAA18B1FFB5E44DFCA1F86FFC9814DC" box="[819,858,1951,1974]" name="Mexico" pageId="12" pageNumber="493">Mx</collectingCountry>
IV = 6 + 10, 
<collectingCountry id="5FAA18B1FFB5E44DFB91F86FFBE814DC" box="[1027,1066,1951,1974]" name="Mexico" pageId="12" pageNumber="493">Mx</collectingCountry>
V = 1 +1, 
<collectingCountry id="5FAA18B1FFB5E44DFB21F86FFB1814DC" box="[1203,1242,1951,1974]" name="Mexico" pageId="12" pageNumber="493">Mx</collectingCountry>
VI absent. Tube cylindrical in shape, parchment-like and externally covered with sand grains.
</paragraph>
</subSubSection>
<caption id="73C208A9FFB4E44CFF05F851FED8149E" httpUri="http://zenodo.org/record/155278/files/figure.png" pageId="13" pageNumber="494" targetBox="[193,1393,193,1930]" targetPageId="13">
<paragraph id="27025821FFB4E44CFF05F851FED8149E" blockId="13.[151,1437,1953,2036]" pageId="13" pageNumber="494">
<emphasis id="15C98433FFB4E44CFF05F851FED514DD" bold="true" box="[151,279,1953,1975]" pageId="13" pageNumber="494">FIGURE 7.</emphasis>
<taxonomicName id="E0BD23A2FFB4E44CFEB0F851FDC214DD" box="[290,512,1953,1975]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="13" pageNumber="494" phylum="Annelida" rank="species" species="hanneloreae" status="sp. nov.">
<emphasis id="15C98433FFB4E44CFEB0F851FDC214DD" box="[290,512,1953,1975]" italics="true" pageId="13" pageNumber="494">Onuphis hanneloreae</emphasis>
</taxonomicName>
<emphasis id="15C98433FFB4E44CFD9EF851FDA314DD" bold="true" box="[524,609,1953,1975]" pageId="13" pageNumber="494">
<taxonomicNameLabel id="0EFA3948FFB4E44CFD9EF851FDA314DD" box="[524,609,1953,1975]" pageId="13" pageNumber="494" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Holotype A, B (MNCN 16.01/17458), paratype C, D (MNCN 16.01/17459). Photomicrographs. A, anterior end, dorsal view; B, detailed view of anterior end; C, anterior end, dorsal view; D, anterior end, ventral view.
</paragraph>
</caption>
<caption id="73C208A9FFB7E44FFF05F87DFBD3148A" httpUri="http://zenodo.org/record/155279/files/figure.png" pageId="14" pageNumber="495" targetBox="[151,1436,193,1910]" targetPageId="14">
<paragraph id="27025821FFB7E44FFF05F87DFBD3148A" blockId="14.[151,1436,1933,2016]" pageId="14" pageNumber="495">
<emphasis id="15C98433FFB7E44FFF05F87DFED614C9" bold="true" box="[151,276,1933,1955]" pageId="14" pageNumber="495">FIGURE 8.</emphasis>
<taxonomicName id="E0BD23A2FFB7E44FFE8EF87DFE3514C9" box="[284,503,1933,1955]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="14" pageNumber="495" phylum="Annelida" rank="species" species="hanneloreae" status="sp. nov.">
<emphasis id="15C98433FFB7E44FFE8EF87DFE3514C9" box="[284,503,1933,1955]" italics="true" pageId="14" pageNumber="495">Onuphis hanneloreae</emphasis>
</taxonomicName>
<emphasis id="15C98433FFB7E44FFE6DF87DFD8814C9" bold="true" box="[511,586,1933,1955]" pageId="14" pageNumber="495">
<taxonomicNameLabel id="0EFA3948FFB7E44FFE6DF87DFD8814C9" box="[511,586,1933,1955]" pageId="14" pageNumber="495" rank="species">sp.nov.</taxonomicNameLabel>
</emphasis>
juvenile. Scanning electron micrographs. A, anterior end, dorsal view; B, detailed view of prostomium; C, detailed view of nuchal Groove; D, anterior modified parapodia, dorsal view. b, branchia; ch, chaetiger; dc, dorsal cirrus; pol, postchaetal lobe; prl, prechaetal lobe; vc, ventral cirrus.
</paragraph>
</caption>
<caption id="73C208A9FFB6E44EFF05F881FE1D1489" httpUri="http://zenodo.org/record/155280/files/figure.png" pageId="15" pageNumber="496" targetBox="[235,1385,260,1865]" targetPageId="15">
<paragraph id="27025821FFB6E44EFF05F881FE1D1489" blockId="15.[151,1436,1905,2019]" pageId="15" pageNumber="496">
<emphasis id="15C98433FFB6E44EFF05F881FED614ED" bold="true" box="[151,276,1905,1927]" pageId="15" pageNumber="496">FIGURE 9.</emphasis>
<taxonomicName id="E0BD23A2FFB6E44EFE8EF881FE3A14ED" box="[284,504,1905,1927]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="15" pageNumber="496" phylum="Annelida" rank="species" species="hanneloreae" status="sp. nov.">
<emphasis id="15C98433FFB6E44EFE8EF881FE3A14ED" box="[284,504,1905,1927]" italics="true" pageId="15" pageNumber="496">Onuphis hanneloreae</emphasis>
</taxonomicName>
<emphasis id="15C98433FFB6E44EFD92F881FD9014ED" bold="true" box="[512,594,1905,1927]" pageId="15" pageNumber="496">
<taxonomicNameLabel id="0EFA3948FFB6E44EFD92F881FD9014ED" box="[512,594,1905,1927]" pageId="15" pageNumber="496" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Paratype A-E (MNCN 16.01/17459). Line drawings. A, parapodium chaetiger 1, anterior view; B, parapodium median chaetiger, anterior view; C slighly shorter tridentate pseudocompound hook from chaetiger 1; D, bidentate pseudocompound hook from chaetiger 1; E, tridentate pseudocompound hook from chaetiger 1; F, maxillae (from largest juvenile).
</paragraph>
</caption>
<caption id="73C208A9FFA9E451FF05F88DFCFF1485" httpUri="http://zenodo.org/record/155281/files/figure.png" pageId="16" pageNumber="497" targetBox="[184,1388,193,1874]" targetPageId="16">
<paragraph id="27025821FFA9E451FF05F88DFCFF1485" blockId="16.[151,1436,1917,2031]" pageId="16" pageNumber="497">
<emphasis id="15C98433FFA9E451FF05F88DFEE614F9" bold="true" box="[151,292,1917,1939]" pageId="16" pageNumber="497">FIGURE 10.</emphasis>
<taxonomicName id="E0BD23A2FFA9E451FEBCF88DFDC914F9" box="[302,523,1917,1939]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="16" pageNumber="497" phylum="Annelida" rank="species" species="hanneloreae" status="sp. nov.">
<emphasis id="15C98433FFA9E451FEBCF88DFDC914F9" box="[302,523,1917,1939]" italics="true" pageId="16" pageNumber="497">Onuphis hanneloreae</emphasis>
</taxonomicName>
<emphasis id="15C98433FFA9E451FD84F88DFDA914F9" bold="true" box="[534,619,1917,1939]" pageId="16" pageNumber="497">
<taxonomicNameLabel id="0EFA3948FFA9E451FD84F88DFDA914F9" box="[534,619,1917,1939]" pageId="16" pageNumber="497" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Scanning electron micrographs. A, glandular pad from median chaetiger; B, median chaetigers, antero-dorsal view; C, pseudocompound hooks from chaetiger 1; D, detailed view of a bidentate pseudocompound hook of chaetiger 2; E, detailed view of a quatridentate pseudocompound hook of chaetiger 2; F, pectinate chaetae from chaetiger 10; G, subacicular hooks from chaetiger 12.
</paragraph>
</caption>
<subSubSection id="6FA70BAAFFA8E450FF55FF67FE741392" pageId="17" pageNumber="498" type="etymology">
<paragraph id="27025821FFA8E450FF55FF67FE741392" blockId="17.[151,1437,151,1040]" pageId="17" pageNumber="498">
<emphasis id="15C98433FFA8E450FF55FF67FE8C13DA" bold="true" box="[199,334,151,176]" pageId="17" pageNumber="498">Etymology.</emphasis>
It is a great pleasure to dedicate this new species to Dr Hannelore Paxton, whose significant contributions have greatly increased and improved the knowledge on this family and have brought about a modern view of their systematics.
</paragraph>
</subSubSection>
<subSubSection id="6FA70BAAFFA8E450FF55FEF4FD58120E" pageId="17" pageNumber="498" type="distribution">
<paragraph id="27025821FFA8E450FF55FEF4FD58120E" blockId="17.[151,1437,151,1040]" pageId="17" pageNumber="498">
<emphasis id="15C98433FFA8E450FF55FEF4FE3A1277" bold="true" box="[199,504,260,285]" pageId="17" pageNumber="498">Distribution and ecology.</emphasis>
<taxonomicName id="E0BD23A2FFA8E450FD93FEF4FD371276" box="[513,757,260,284]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="498" phylum="Annelida" rank="species" species="hanneloreae" status="sp. nov.">
<emphasis id="15C98433FFA8E450FD93FEF4FD371276" box="[513,757,260,284]" italics="true" pageId="17" pageNumber="498">Onuphis hanneloreae</emphasis>
</taxonomicName>
<emphasis id="15C98433FFA8E450FD6CFEF4FC981277" bold="true" box="[766,858,260,285]" pageId="17" pageNumber="498">
<taxonomicNameLabel id="0EFA3948FFA8E450FD6CFEF4FC981277" box="[766,858,260,285]" pageId="17" pageNumber="498" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is known from the intertidal and shallow subtidal sandy flats and patchy sandy areas between rocks of 
<collectingRegion id="E57996C3FFA8E450FD74FED7FCCE122A" box="[742,780,295,320]" country="Cape Verde" name="Sal" pageId="17" pageNumber="498">Sal</collectingRegion>
, the second easternmost island of 
<collectingCountry id="5FAA18B1FFA8E450FB18FED8FACE122A" box="[1162,1292,295,320]" name="Cape Verde" pageId="17" pageNumber="498">
<collectingRegion id="E57996C3FFA8E450FB18FED8FB06122A" box="[1162,1220,296,320]" country="South Africa" name="Western Cape" pageId="17" pageNumber="498">Cape</collectingRegion>
Verde
</collectingCountry>
archipelago, 
<collectingRegion id="E57996C3FFA8E450FF05FEBDFF0C120E" box="[151,206,333,356]" country="Cameroon" name="West" pageId="17" pageNumber="498">West</collectingRegion>
Africa, tropical eastern Atlantic (
<figureCitation id="BF8644A4FFA8E450FDD5FEBCFD4A120E" box="[583,648,332,357]" captionStart="FIGURE 1" captionStartId="1.[151,250,1904,1926]" captionTargetBox="[151,1436,493,1869]" captionTargetId="figure@1.[151,1436,493,1871]" captionTargetPageId="1" captionText="FIGURE 1. Distribution map of the type localities of Onuphis augeneri sp. nov., O. hanneloreae sp. nov. and Mooreonuphis nunezi sp. nov. Satellite images modified from Google Earth." httpUri="http://zenodo.org/record/155272/files/figure.png" pageId="17" pageNumber="498">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="6FA70BAAFFA8E455FF55FE9FFD871779" lastPageId="20" lastPageNumber="501" pageId="17" pageNumber="498" type="discussion">
<paragraph id="27025821FFA8E450FF55FE9FFDFF119A" blockId="17.[151,1437,151,1040]" pageId="17" pageNumber="498">
<emphasis id="15C98433FFA8E450FF55FE9FFEF912E2" bold="true" box="[199,315,367,392]" pageId="17" pageNumber="498">Remarks.</emphasis>
<taxonomicName id="E0BD23A2FFA8E450FED7FE80FDFB12E2" box="[325,569,368,392]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="498" phylum="Annelida" rank="species" species="hanneloreae" status="sp. nov.">
<emphasis id="15C98433FFA8E450FED7FE80FDFB12E2" box="[325,569,368,392]" italics="true" pageId="17" pageNumber="498">Onuphis hanneloreae</emphasis>
</taxonomicName>
<emphasis id="15C98433FFA8E450FDD1FE9FFD6212E2" bold="true" box="[579,672,367,392]" pageId="17" pageNumber="498">
<taxonomicNameLabel id="0EFA3948FFA8E450FDD1FE9FFD6212E2" box="[579,672,367,392]" pageId="17" pageNumber="498" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
belongs to the 
<taxonomicName id="E0BD23A2FFA8E450FCCAFE80FC1112ED" box="[856,979,368,391]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="498" phylum="Annelida" rank="species" species="eremita">
<emphasis id="15C98433FFA8E450FCCAFE80FC1112ED" box="[856,979,368,391]" italics="true" pageId="17" pageNumber="498">O. eremita</emphasis>
</taxonomicName>
species complex (
<tableCitation id="6A3F6D9AFFA8E450FB38FE9FFAC312E2" box="[1194,1281,367,392]" captionStart="TABLE 1" captionStartId="5.[125,202,650,670]" captionTargetBox="[125,1883,695,1111]" captionText="TABLE 1. Comparison of key features and habitats of the eastern Atlantic species of the Onuphis eremita complex." httpUri="http://table.plazi.org/id/73C208A9FFBCE444FFEFFD7AFB6011F5" pageId="17" pageNumber="498" tableUuid="73C208A9FFBCE444FFEFFD7AFB6011F5">Table 1</tableCitation>
). Within this group the new species resembles the European 
<taxonomicName id="E0BD23A2FFA8E450FD22FE64FCF212C6" box="[688,816,404,428]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="498" phylum="Annelida" rank="species" species="pancerii">
<emphasis id="15C98433FFA8E450FD22FE64FCF212C6" box="[688,816,404,428]" italics="true" pageId="17" pageNumber="498">O. pancerii</emphasis>
</taxonomicName>
and the Japanese 
<taxonomicName id="E0BD23A2FFA8E450FC6DFE64FF0D12BA" authority="Maekawa &amp; Hayashi, 1999" authorityName="Maekawa &amp; Hayashi" authorityYear="1999" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="498" phylum="Annelida" rank="species" species="shijikiensis">
<emphasis id="15C98433FFA8E450FC6DFE64FB5D12C6" box="[1023,1183,404,428]" italics="true" pageId="17" pageNumber="498">O. shijikiensis</emphasis>
<bibRefCitation id="432C25D0FFA8E450FB34FE64FF0D12BA" author="Maekawa" pageId="17" pageNumber="498" refString="Maekawa, N. &amp; Hayashi, I. (1999) Taxonomic study on the genus Onuphis (Polychaeta, Onuphidae) from Japan and adjacent seas, with descriptions of six new species. Bulletin of the National Science Museum of Tokyo, 25 (3), 163 - 214." type="journal article" year="1999">
Maekawa &amp; Hayashi, 
<quantity id="E045F5C4FFA8E450FF05FE48FF0D12BA" box="[151,207,440,464]" metricMagnitude="1" metricUnit="m" metricValue="5.07746" pageId="17" pageNumber="498" unit="in" value="1999.0">1999</quantity>
</bibRefCitation>
</taxonomicName>
in having both bi- and tridentate pseudocompound hooks in the first three chaetigers and subacicular hooks from chaetiger 10. However 
<taxonomicName id="E0BD23A2FFA8E450FE78FE2CFD59129E" box="[490,667,476,500]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="498" phylum="Annelida" rank="species" species="hanneloreae" status="sp. nov.">
<emphasis id="15C98433FFA8E450FE78FE2CFD59129E" box="[490,667,476,500]" italics="true" pageId="17" pageNumber="498">O. hanneloreae</emphasis>
</taxonomicName>
<emphasis id="15C98433FFA8E450FD34FE2CFCC6129F" bold="true" box="[678,772,476,501]" pageId="17" pageNumber="498">
<taxonomicNameLabel id="0EFA3948FFA8E450FD34FE2CFCC6129F" box="[678,772,476,501]" pageId="17" pageNumber="498" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="E0BD23A2FFA8E450FCD0FE2CFC07129E" box="[834,965,476,500]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="498" phylum="Annelida" rank="species" species="pancerii">
<emphasis id="15C98433FFA8E450FCD0FE2CFC07129E" box="[834,965,476,500]" italics="true" pageId="17" pageNumber="498">O. pancerii</emphasis>
</taxonomicName>
differ from the Japanese species by the presence of quatridentate pseudocompound hooks in anterior modified chaetigers and in the number of hooks per fascicle, bearing five hooks, in contrast 
<taxonomicName id="E0BD23A2FFA8E450FDCBFDD4FD3A1156" box="[601,760,548,572]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="498" phylum="Annelida" rank="species" species="shijikiensis">
<emphasis id="15C98433FFA8E450FDCBFDD4FD3A1156" box="[601,760,548,572]" italics="true" pageId="17" pageNumber="498">O. shijikiensis</emphasis>
</taxonomicName>
only has three hooks in each fascicle (
<bibRefCitation id="432C25D0FFA8E450FB3FFDD4FF11110A" author="Maekawa" pageId="17" pageNumber="498" refString="Maekawa, N. &amp; Hayashi, I. (1999) Taxonomic study on the genus Onuphis (Polychaeta, Onuphidae) from Japan and adjacent seas, with descriptions of six new species. Bulletin of the National Science Museum of Tokyo, 25 (3), 163 - 214." type="journal article" year="1999">Maekawa &amp; Hayashi 1999</bibRefCitation>
). 
<taxonomicName id="E0BD23A2FFA8E450FF74FDB8FE15110A" box="[230,471,584,608]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="498" phylum="Annelida" rank="species" species="hanneloreae" status="sp. nov.">
<emphasis id="15C98433FFA8E450FF74FDB8FE15110A" box="[230,471,584,608]" italics="true" pageId="17" pageNumber="498">Onuphis hanneloreae</emphasis>
</taxonomicName>
<emphasis id="15C98433FFA8E450FE4CFDB7FDFA110A" bold="true" box="[478,568,583,608]" pageId="17" pageNumber="498">
<taxonomicNameLabel id="0EFA3948FFA8E450FE4CFDB7FDFA110A" box="[478,568,583,608]" pageId="17" pageNumber="498" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
clearly differs from 
<taxonomicName id="E0BD23A2FFA8E450FCB2FDB8FC7D110A" box="[800,959,584,608]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="498" phylum="Annelida" rank="species" species="shijikiensis">
<emphasis id="15C98433FFA8E450FCB2FDB8FC7D110A" box="[800,959,584,608]" italics="true" pageId="17" pageNumber="498">O. shijikiensis</emphasis>
</taxonomicName>
in its colour pattern and 
<taxonomicName id="E0BD23A2FFA8E450FB45FDB8FA951135" box="[1239,1367,584,607]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="498" phylum="Annelida" rank="species" species="pancerii">
<emphasis id="15C98433FFA8E450FB45FDB8FA951135" box="[1239,1367,584,607]" italics="true" pageId="17" pageNumber="498">O. pancerii</emphasis>
</taxonomicName>
by its smaller size and the absence of the interramal papillae at the base of dorsal cirri on anterior non-modified chaetigers (present in 
<taxonomicName id="E0BD23A2FFA8E450FE1CFD60FDCF11CD" box="[398,525,656,679]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="498" phylum="Annelida" rank="species" species="pancerii">
<emphasis id="15C98433FFA8E450FE1CFD60FDCF11CD" box="[398,525,656,679]" italics="true" pageId="17" pageNumber="498">O. pancerii</emphasis>
</taxonomicName>
). Furthermore, their reproductive biology is different, 
<taxonomicName id="E0BD23A2FFA8E450FBE6FD60FAE311C2" box="[1140,1313,656,680]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="498" phylum="Annelida" rank="species" species="hanneloreae" status="sp. nov.">
<emphasis id="15C98433FFA8E450FBE6FD60FAE311C2" box="[1140,1313,656,680]" italics="true" pageId="17" pageNumber="498">O. hanneloreae</emphasis>
</taxonomicName>
<emphasis id="15C98433FFA8E450FABAFD7FFA4011C2" bold="true" box="[1320,1410,655,680]" pageId="17" pageNumber="498">
<taxonomicNameLabel id="0EFA3948FFA8E450FABAFD7FFA4011C2" box="[1320,1410,655,680]" pageId="17" pageNumber="498" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is a brooder species that undergoes direct development; by contrast 
<taxonomicName id="E0BD23A2FFA8E450FCEEFD44FC3F11A6" box="[892,1021,692,716]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="498" phylum="Annelida" rank="species" species="pancerii">
<emphasis id="15C98433FFA8E450FCEEFD44FC3F11A6" box="[892,1021,692,716]" italics="true" pageId="17" pageNumber="498">O. pancerii</emphasis>
</taxonomicName>
is a broadcast spawner with indirect development (Arias &amp; Paxton, 2014).
</paragraph>
<paragraph id="27025821FFA8E450FF55FD0CFDEB177A" blockId="17.[151,1437,151,1040]" pageId="17" pageNumber="498">
<emphasis id="15C98433FFA8E450FF55FD0CFDBB107F" bold="true" box="[199,633,764,789]" pageId="17" pageNumber="498">Reproductive biology and ontogeny.</emphasis>
Among the 
<typeStatus id="F806E683FFA8E450FC9AFD0DFCFA107F" box="[776,824,765,789]" pageId="17" pageNumber="498">type</typeStatus>
material, the 
<typeStatus id="F806E683FFA8E450FC45FD0DFBF5107F" box="[983,1079,765,789]" pageId="17" pageNumber="498" type="paratype">paratype</typeStatus>
specimen was collected within its tube together with three vermiform juveniles. The brooded juveniles range from 
<quantity id="E045F5C4FFA8E450FBA8FCD0FB02105D" box="[1082,1216,800,824]" metricMagnitude="-3" metricUnit="m" metricValue="4.05" metricValueMax="4.2" metricValueMin="3.9" pageId="17" pageNumber="498" unit="mm" value="4.05" valueMax="4.2" valueMin="3.9">3.9–4.2 mm</quantity>
in length for 28–39 chaetigers and measure 
<quantity id="E045F5C4FFA8E450FE23FCB4FD9A1036" box="[433,600,836,861]" metricMagnitude="-4" metricUnit="m" metricValue="3.4499999999999997" metricValueMax="3.5" metricValueMin="3.4" pageId="17" pageNumber="498" unit="mm" value="0.345" valueMax="0.35" valueMin="0.34">0.34–0.35 mm</quantity>
in width (
<tableCitation id="6A3F6D9AFFA8E450FD4AFCB4FCF01036" box="[728,818,836,861]" captionStart="TABLE 3" captionStartId="19.[125,202,666,689]" captionTargetBox="[147,1878,732,1088]" captionTargetPageId="19" captionText="TABLE 3. Ontogeny of Onuphis hanneloreae sp. nov. juveniles" httpUri="http://table.plazi.org/id/73C208A9FFAAE452FFEFFD6AFD0711DB" pageId="17" pageNumber="498" tableUuid="73C208A9FFAAE452FFEFFD6AFD0711DB">Table 3</tableCitation>
), with the largest presenting a colour pattern on the dorsum of the anterior chaetigers similar to the adults. All juveniles have a pair of small eyespots on prostomium, small peristomial cirri, well developed frontal lips, palps and antennae and the ceratophores of the largest two with three to four rings. The parapodia are well developed with dorsal cirri on the first 20 to 30 chaetigers and subulate ventral cirri on the first two to three chaetigers. Single branchiae are present on chaetigers 6 and 7; the pygidium bears two pairs of slender anal cirri.
</paragraph>
<caption id="73C208A9FFA8E450FF05F825FBB71481" box="[151,1141,2005,2027]" httpUri="http://zenodo.org/record/155282/files/figure.png" pageId="17" pageNumber="498" targetBox="[183,1401,1124,1933]" targetPageId="17">
<paragraph id="27025821FFA8E450FF05F825FBB71481" blockId="17.[151,1141,2005,2027]" box="[151,1141,2005,2027]" pageId="17" pageNumber="498">
<emphasis id="15C98433FFA8E450FF05F825FEDC1481" bold="true" box="[151,286,2005,2027]" pageId="17" pageNumber="498">FIGURE 11.</emphasis>
Comparison of the progression of hooks in 
<taxonomicName id="E0BD23A2FFA8E450FD4EF825FC771481" box="[732,949,2005,2027]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="498" phylum="Annelida" rank="species" species="hanneloreae" status="sp. nov.">
<emphasis id="15C98433FFA8E450FD4EF825FC771481" box="[732,949,2005,2027]" italics="true" pageId="17" pageNumber="498">Onuphis hanneloreae</emphasis>
</taxonomicName>
<emphasis id="15C98433FFA8E450FC2EF825FBCE1481" bold="true" box="[956,1036,2005,2027]" pageId="17" pageNumber="498">
<taxonomicNameLabel id="0EFA3948FFA8E450FC2EF825FBCE1481" box="[956,1036,2005,2027]" pageId="17" pageNumber="498" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
juveniles.
</paragraph>
</caption>
<caption id="73C208A9FFABE453FF05F851FAB0149E" httpUri="http://zenodo.org/record/155283/files/figure.png" pageId="18" pageNumber="499" targetBox="[202,1384,193,1930]" targetPageId="18">
<paragraph id="27025821FFABE453FF05F851FAB0149E" blockId="18.[151,1436,1953,2036]" pageId="18" pageNumber="499">
<emphasis id="15C98433FFABE453FF05F851FEDD14DD" bold="true" box="[151,287,1953,1975]" pageId="18" pageNumber="499">FIGURE 12.</emphasis>
<taxonomicName id="E0BD23A2FFABE453FEB7F851FE3C14DD" box="[293,510,1953,1975]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="18" pageNumber="499" phylum="Annelida" rank="species" species="hanneloreae" status="sp. nov.">
<emphasis id="15C98433FFABE453FEB7F851FE3C14DD" box="[293,510,1953,1975]" italics="true" pageId="18" pageNumber="499">Onuphis hanneloreae</emphasis>
</taxonomicName>
<emphasis id="15C98433FFABE453FD97F851FD9714DD" bold="true" box="[517,597,1953,1975]" pageId="18" pageNumber="499">
<taxonomicNameLabel id="0EFA3948FFABE453FD97F851FD9714DD" box="[517,597,1953,1975]" pageId="18" pageNumber="499" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
juvenile. Scanning electron micrographs. A, bidentate compound hook; B, median anterior provisional subacicular hook; C, ventral anterior provisional subacicular hook; D, posterior provisional subacicular hook; E, posterior parapodium showing both permanent subacicular hook (top) and provisional subacicular hook (bottom).
</paragraph>
</caption>
<caption id="73C208A9FFAAE452FFEFFD6AFD0711DB" ID-Table-UUID="73C208A9FFAAE452FFEFFD6AFD0711DB" box="[125,709,666,689]" httpUri="http://table.plazi.org/id/73C208A9FFAAE452FFEFFD6AFD0711DB" pageId="19" pageNumber="500" targetBox="[147,1878,732,1088]" targetIsTable="true" targetPageId="19">
<paragraph id="27025821FFAAE452FFEFFD6AFD0711DB" blockId="19.[125,709,666,689]" box="[125,709,666,689]" pageId="19" pageNumber="500">
<emphasis id="15C98433FFAAE452FFEFFD6AFF2211DB" bold="true" box="[125,224,666,689]" pageId="19" pageNumber="500">TABLE 3.</emphasis>
Ontogeny of 
<taxonomicName id="E0BD23A2FFAAE452FECEFD6AFDE211DB" box="[348,544,666,689]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="19" pageNumber="500" phylum="Annelida" rank="species" species="hanneloreae" status="sp. nov.">
<emphasis id="15C98433FFAAE452FECEFD6AFDE211DB" box="[348,544,666,689]" italics="true" pageId="19" pageNumber="500">Onuphis hanneloreae</emphasis>
</taxonomicName>
<emphasis id="15C98433FFAAE452FDB7FD6AFDAC11DB" bold="true" box="[549,622,666,689]" pageId="19" pageNumber="500">
<taxonomicNameLabel id="0EFA3948FFAAE452FDB7FD6AFDAC11DB" box="[549,622,666,689]" pageId="19" pageNumber="500" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
juveniles
</paragraph>
</caption>
<paragraph id="27025821FFAAE452FF08FD2CF97B1755" pageId="19" pageNumber="500">
<table id="55BDAA81FFAA1BBEFF01FD2CF894172A" box="[147,1878,732,1088]" gridcols="6" gridrows="7" pageId="19" pageNumber="500">
<tr id="998D5A63FFAA1BBEFF01FD2CF8941027" box="[147,1878,732,845]" gridrow="0" pageId="19" pageNumber="500">
<th id="DA5C331FFFAA1BBEFF01FD2CFEBC1027" box="[147,382,732,845]" gridcol="0" gridrow="0" pageId="19" pageNumber="500">
. 
<emphasis id="15C98433FFAAE452FF31FD2CFEBC1199" bold="true" box="[163,382,732,755]" pageId="19" pageNumber="500">of ch. Width (mm)</emphasis>
0.34
</th>
<th id="DA5C331FFFAA1BBEFE54FD2CFD871027" box="[454,581,732,845]" gridcol="1" gridrow="0" pageId="19" pageNumber="500">
<emphasis id="15C98433FFAAE452FE54FD2CFE211027" bold="true" pageId="19" pageNumber="500">Length (mm) 3.9</emphasis>
</th>
<th id="DA5C331FFFAA1BBEFDF7FD2CFCE21027" box="[613,800,732,845]" gridcol="2" gridrow="0" pageId="19" pageNumber="500">
<emphasis id="15C98433FFAAE452FDF7FD2CFD1A107A" bold="true" pageId="19" pageNumber="500">No. of ceratophoral rings on LA</emphasis>
6
</th>
<th id="DA5C331FFFAA1BBEFCCCFD2CFC161027" box="[862,980,732,845]" gridcol="3" gridrow="0" pageId="19" pageNumber="500">
<emphasis id="15C98433FFAAE452FCCCFD2CFC16107A" bold="true" pageId="19" pageNumber="500">Eyespots on prostomium</emphasis>
present
</th>
<th id="DA5C331FFFAA1BBEFBB5FD2CFA511027" box="[1063,1427,732,845]" gridcol="4" gridrow="0" pageId="19" pageNumber="500">
<emphasis id="15C98433FFAAE452FBB5FD2CFA81107A" bold="true" pageId="19" pageNumber="500">No. of chaetigers Single branchiae with subulate VC from ch.</emphasis>
3 6
</th>
<th id="DA5C331FFFAA1BBEFA46FD2CF8941027" box="[1492,1878,732,845]" gridcol="5" gridrow="0" pageId="19" pageNumber="500">
<emphasis id="15C98433FFAAE452FA46FD2CF8C51044" bold="true" pageId="19" pageNumber="500">Two or more Maximal number branchial filaments of branchial from ch. filaments</emphasis>
- 1
</th>
</tr>
<tr id="998D5A63FFAA1BBEFF01FCAEF894101F" box="[147,1878,862,885]" gridrow="1" pageId="19" pageNumber="500">
<th id="DA5C331FFFAA1BBEFF01FCAEFEBC101F" box="[147,382,862,885]" gridcol="0" gridrow="1" pageId="19" pageNumber="500">0.34</th>
<td id="DA5C331FFFAA1BBEFE54FCAEFD87101F" box="[454,581,862,885]" gridcol="1" gridrow="1" pageId="19" pageNumber="500">4.1</td>
<td id="DA5C331FFFAA1BBEFDF7FCAEFCE2101F" box="[613,800,862,885]" gridcol="2" gridrow="1" pageId="19" pageNumber="500">5</td>
<td id="DA5C331FFFAA1BBEFCCCFCAEFC16101F" box="[862,980,862,885]" gridcol="3" gridrow="1" pageId="19" pageNumber="500">present</td>
<td id="DA5C331FFFAA1BBEFBB5FCAEFA51101F" box="[1063,1427,862,885]" gridcol="4" gridrow="1" pageId="19" pageNumber="500">2 6</td>
<td id="DA5C331FFFAA1BBEFA46FCAEF894101F" box="[1492,1878,862,885]" gridcol="5" gridrow="1" pageId="19" pageNumber="500">- 1</td>
</tr>
<tr id="998D5A63FFAA1BBEFF01FC77F89410F4" box="[147,1878,903,926]" gridrow="2" pageId="19" pageNumber="500">
<th id="DA5C331FFFAA1BBEFF01FC77FEBC10F4" box="[147,382,903,926]" gridcol="0" gridrow="2" pageId="19" pageNumber="500">0.35</th>
<td id="DA5C331FFFAA1BBEFE54FC77FD8710F4" box="[454,581,903,926]" gridcol="1" gridrow="2" pageId="19" pageNumber="500">4.2</td>
<td id="DA5C331FFFAA1BBEFDF7FC77FCE210F4" box="[613,800,903,926]" gridcol="2" gridrow="2" pageId="19" pageNumber="500">7</td>
<td id="DA5C331FFFAA1BBEFCCCFC77FC1610F4" box="[862,980,903,926]" gridcol="3" gridrow="2" pageId="19" pageNumber="500">present</td>
<td id="DA5C331FFFAA1BBEFBB5FC77FA5110F4" box="[1063,1427,903,926]" gridcol="4" gridrow="2" pageId="19" pageNumber="500">3 6</td>
<td id="DA5C331FFFAA1BBEFA46FC77F89410F4" box="[1492,1878,903,926]" gridcol="5" gridrow="2" pageId="19" pageNumber="500">- 1</td>
</tr>
<tr id="998D5A63FFAA1BBEFF01FC5FF89410AC" box="[147,1878,943,966]" gridrow="3" pageId="19" pageNumber="500">
<th id="DA5C331FFFAA1BBEFF01FC5FFEBC10AC" box="[147,382,943,966]" gridcol="0" gridrow="3" pageId="19" pageNumber="500">0.40</th>
<td id="DA5C331FFFAA1BBEFE54FC5FFD8710AC" box="[454,581,943,966]" gridcol="1" gridrow="3" pageId="19" pageNumber="500">5.3</td>
<td id="DA5C331FFFAA1BBEFDF7FC5FFCE210AC" box="[613,800,943,966]" gridcol="2" gridrow="3" pageId="19" pageNumber="500">9</td>
<td id="DA5C331FFFAA1BBEFCCCFC5FFC1610AC" box="[862,980,943,966]" gridcol="3" gridrow="3" pageId="19" pageNumber="500">present</td>
<td id="DA5C331FFFAA1BBEFBB5FC5FFA5110AC" box="[1063,1427,943,966]" gridcol="4" gridrow="3" pageId="19" pageNumber="500">3 5</td>
<td id="DA5C331FFFAA1BBEFA46FC5FF89410AC" box="[1492,1878,943,966]" gridcol="5" gridrow="3" pageId="19" pageNumber="500">- 1</td>
</tr>
<tr id="998D5A63FFAA1BBEFF01FC27F8941084" box="[147,1878,983,1006]" gridrow="4" pageId="19" pageNumber="500">
<th id="DA5C331FFFAA1BBEFF01FC27FEBC1084" box="[147,382,983,1006]" gridcol="0" gridrow="4" pageId="19" pageNumber="500">0.42</th>
<td id="DA5C331FFFAA1BBEFE54FC27FD871084" box="[454,581,983,1006]" gridcol="1" gridrow="4" pageId="19" pageNumber="500">5.5</td>
<td id="DA5C331FFFAA1BBEFDF7FC27FCE21084" box="[613,800,983,1006]" gridcol="2" gridrow="4" pageId="19" pageNumber="500">10</td>
<td id="DA5C331FFFAA1BBEFCCCFC27FC161084" box="[862,980,983,1006]" gridcol="3" gridrow="4" pageId="19" pageNumber="500">present</td>
<td id="DA5C331FFFAA1BBEFBB5FC27FA511084" box="[1063,1427,983,1006]" gridcol="4" gridrow="4" pageId="19" pageNumber="500">3 3</td>
<td id="DA5C331FFFAA1BBEFA46FC27F8941084" box="[1492,1878,983,1006]" gridcol="5" gridrow="4" pageId="19" pageNumber="500">- 1</td>
</tr>
<tr id="998D5A63FFAA1BBEFF01FC0FF894177C" box="[147,1878,1023,1046]" gridrow="5" pageId="19" pageNumber="500">
<th id="DA5C331FFFAA1BBEFF01FC0FFEBC177C" box="[147,382,1023,1046]" gridcol="0" gridrow="5" pageId="19" pageNumber="500">0.49</th>
<td id="DA5C331FFFAA1BBEFE54FC0FFD87177C" box="[454,581,1023,1046]" gridcol="1" gridrow="5" pageId="19" pageNumber="500">6.2</td>
<td id="DA5C331FFFAA1BBEFDF7FC0FFCE2177C" box="[613,800,1023,1046]" gridcol="2" gridrow="5" pageId="19" pageNumber="500">8 9</td>
<td id="DA5C331FFFAA1BBEFCCCFC0FFC16177C" box="[862,980,1023,1046]" gridcol="3" gridrow="5" pageId="19" pageNumber="500">absent</td>
<td id="DA5C331FFFAA1BBEFBB5FC0FFA51177C" box="[1063,1427,1023,1046]" gridcol="4" gridrow="5" pageId="19" pageNumber="500">3 2</td>
<td id="DA5C331FFFAA1BBEFA46FC0FF894177C" box="[1492,1878,1023,1046]" gridcol="5" gridrow="5" pageId="19" pageNumber="500">- 1</td>
</tr>
<tr id="998D5A63FFAA1BBEFF01FBD8F894172A" box="[147,1878,1064,1088]" gridrow="6" pageId="19" pageNumber="500">
<th id="DA5C331FFFAA1BBEFF01FBD8FEBC172A" box="[147,382,1064,1088]" gridcol="0" gridrow="6" pageId="19" pageNumber="500">
<emphasis id="15C98433FFAAE452FF01FBD9FF62172A" bold="true" box="[147,160,1065,1088]" pageId="19" pageNumber="500">+</emphasis>
0.52
</th>
<td id="DA5C331FFFAA1BBEFE54FBD8FD87172A" box="[454,581,1064,1088]" gridcol="1" gridrow="6" pageId="19" pageNumber="500">6.4+</td>
<td id="DA5C331FFFAA1BBEFDF7FBD8FCE2172A" box="[613,800,1064,1088]" gridcol="2" gridrow="6" pageId="19" pageNumber="500">10</td>
<td id="DA5C331FFFAA1BBEFCCCFBD8FC16172A" box="[862,980,1064,1088]" gridcol="3" gridrow="6" pageId="19" pageNumber="500">present</td>
<td id="DA5C331FFFAA1BBEFBB5FBD8FA51172A" box="[1063,1427,1064,1088]" gridcol="4" gridrow="6" pageId="19" pageNumber="500">4 2</td>
<td id="DA5C331FFFAA1BBEFA46FBD8F894172A" box="[1492,1878,1064,1088]" gridcol="5" gridrow="6" pageId="19" pageNumber="500">16 2</td>
</tr>
</table>
</paragraph>
<paragraph id="27025821FFAAE452FFEFFB94F91A16A5" pageId="19" pageNumber="500">
<table id="55BDAA81FFAA1BBEFFEFFB94F8F716BA" box="[125,1845,1124,1488]" gridcols="8" gridrows="7" pageId="19" pageNumber="500">
<tr id="998D5A63FFAA1BBEFFEFFB94F8F717B4" box="[125,1845,1124,1246]" gridrow="0" pageId="19" pageNumber="500">
<th id="DA5C331FFFAA1BBEFFEFFB94FF2017B4" box="[125,226,1124,1246]" gridcol="0" gridrow="0" pageId="19" pageNumber="500">
Continueđ. 
<emphasis id="15C98433FFAAE452FFEFFB79FF5617B4" bold="true" pageId="19" pageNumber="500">No. of ch. 28</emphasis>
</th>
<th id="DA5C331FFFAA1BBEFE89FB94FE6517B4" box="[283,423,1124,1246]" gridcol="1" gridrow="0" pageId="19" pageNumber="500">
<emphasis id="15C98433FFAAE452FE89FB79FE9117D4" bold="true" pageId="19" pageNumber="500">No. of ch. with hooks</emphasis>
3
</th>
<th id="DA5C331FFFAA1BBEFE48FB94FD6317B4" box="[474,673,1124,1246]" gridcol="2" gridrow="0" pageId="19" pageNumber="500">
<emphasis id="15C98433FFAAE452FE48FB79FD8817D4" bold="true" pageId="19" pageNumber="500">Compound hooks on anterior ch.</emphasis>
present
</th>
<th id="DA5C331FFFAA1BBEFD48FB94FBF717B4" box="[730,1077,1124,1246]" gridcol="3" gridrow="0" pageId="19" pageNumber="500">
<emphasis id="15C98433FFAAE452FD48FB79FD3417D4" bold="true" pageId="19" pageNumber="500">Pseudocompound hooks on anterior ch.</emphasis>
absent
</th>
<th id="DA5C331FFFAA1BBEFBC3FB94FB3017B4" box="[1105,1266,1124,1246]" gridcol="4" gridrow="0" pageId="19" pageNumber="500">Ant. Prov. hooks present</th>
<th id="DA5C331FFFAA1BBEFA88FB94FA7D17B4" box="[1306,1471,1124,1246]" gridcol="5" gridrow="0" pageId="19" pageNumber="500">
<emphasis id="15C98433FFAAE452FA88FB79FA7D17CA" bold="true" box="[1306,1471,1161,1184]" pageId="19" pageNumber="500">Post. prov. hooks</emphasis>
present
</th>
<th id="DA5C331FFFAA1BBEFA76FB94F9AB17B4" box="[1508,1641,1124,1246]" gridcol="6" gridrow="0" pageId="19" pageNumber="500">
<emphasis id="15C98433FFAAE452FA76FB79FA2E17B7" bold="true" pageId="19" pageNumber="500">SAH from ch. no. -</emphasis>
</th>
<th id="DA5C331FFFAA1BBEF905FB94F8F717B4" box="[1687,1845,1124,1246]" gridcol="7" gridrow="0" pageId="19" pageNumber="500">
<emphasis id="15C98433FFAAE452F905FB79F8F717CA" bold="true" box="[1687,1845,1161,1184]" pageId="19" pageNumber="500">Pectinate chaeta</emphasis>
absent
</th>
</tr>
<tr id="998D5A63FFAA1BBEFFEFFB1EF8F7166C" box="[125,1845,1262,1286]" gridrow="1" pageId="19" pageNumber="500">
<th id="DA5C331FFFAA1BBEFFEFFB1EFF20166C" box="[125,226,1262,1286]" gridcol="0" gridrow="1" pageId="19" pageNumber="500">
<emphasis id="15C98433FFAAE452FFEFFB1FFF56166C" bold="true" box="[125,148,1263,1286]" pageId="19" pageNumber="500">33</emphasis>
</th>
<td id="DA5C331FFFAA1BBEFE89FB1EFE65166C" box="[283,423,1262,1286]" gridcol="1" gridrow="1" pageId="19" pageNumber="500">3</td>
<td id="DA5C331FFFAA1BBEFE48FB1EFD63166C" box="[474,673,1262,1286]" gridcol="2" gridrow="1" pageId="19" pageNumber="500">present</td>
<td id="DA5C331FFFAA1BBEFD48FB1EFBF7166C" box="[730,1077,1262,1286]" gridcol="3" gridrow="1" pageId="19" pageNumber="500">absent</td>
<td id="DA5C331FFFAA1BBEFBC3FB1EFB30166C" box="[1105,1266,1262,1286]" gridcol="4" gridrow="1" pageId="19" pageNumber="500">present</td>
<td id="DA5C331FFFAA1BBEFA88FB1EFA7D166C" box="[1306,1471,1262,1286]" gridcol="5" gridrow="1" pageId="19" pageNumber="500">present</td>
<td id="DA5C331FFFAA1BBEFA76FB1EF9AB166C" box="[1508,1641,1262,1286]" gridcol="6" gridrow="1" pageId="19" pageNumber="500">-</td>
<td id="DA5C331FFFAA1BBEF905FB1EF8F7166C" box="[1687,1845,1262,1286]" gridcol="7" gridrow="1" pageId="19" pageNumber="500">absent</td>
</tr>
<tr id="998D5A63FFAA1BBEFFEFFAE7F8F71644" box="[125,1845,1303,1326]" gridrow="2" pageId="19" pageNumber="500">
<th id="DA5C331FFFAA1BBEFFEFFAE7FF201644" box="[125,226,1303,1326]" gridcol="0" gridrow="2" pageId="19" pageNumber="500">
<emphasis id="15C98433FFAAE452FFEFFAE7FF561644" bold="true" box="[125,148,1303,1326]" pageId="19" pageNumber="500">39</emphasis>
</th>
<td id="DA5C331FFFAA1BBEFE89FAE7FE651644" box="[283,423,1303,1326]" gridcol="1" gridrow="2" pageId="19" pageNumber="500">3</td>
<td id="DA5C331FFFAA1BBEFE48FAE7FD631644" box="[474,673,1303,1326]" gridcol="2" gridrow="2" pageId="19" pageNumber="500">present</td>
<td id="DA5C331FFFAA1BBEFD48FAE7FBF71644" box="[730,1077,1303,1326]" gridcol="3" gridrow="2" pageId="19" pageNumber="500">absent</td>
<td id="DA5C331FFFAA1BBEFBC3FAE7FB301644" box="[1105,1266,1303,1326]" gridcol="4" gridrow="2" pageId="19" pageNumber="500">present</td>
<td id="DA5C331FFFAA1BBEFA88FAE7FA7D1644" box="[1306,1471,1303,1326]" gridcol="5" gridrow="2" pageId="19" pageNumber="500">present</td>
<td id="DA5C331FFFAA1BBEFA76FAE7F9AB1644" box="[1508,1641,1303,1326]" gridcol="6" gridrow="2" pageId="19" pageNumber="500">-</td>
<td id="DA5C331FFFAA1BBEF905FAE7F8F71644" box="[1687,1845,1303,1326]" gridcol="7" gridrow="2" pageId="19" pageNumber="500">absent</td>
</tr>
<tr id="998D5A63FFAA1BBEFFEFFACFF8F7163D" box="[125,1845,1343,1367]" gridrow="3" pageId="19" pageNumber="500">
<th id="DA5C331FFFAA1BBEFFEFFACFFF20163D" box="[125,226,1343,1367]" gridcol="0" gridrow="3" pageId="19" pageNumber="500">
<emphasis id="15C98433FFAAE452FFEFFAB0FF56163D" bold="true" box="[125,148,1344,1367]" pageId="19" pageNumber="500">46</emphasis>
</th>
<td id="DA5C331FFFAA1BBEFE89FACFFE65163D" box="[283,423,1343,1367]" gridcol="1" gridrow="3" pageId="19" pageNumber="500">3</td>
<td id="DA5C331FFFAA1BBEFE48FACFFD63163D" box="[474,673,1343,1367]" gridcol="2" gridrow="3" pageId="19" pageNumber="500">present</td>
<td id="DA5C331FFFAA1BBEFD48FACFFBF7163D" box="[730,1077,1343,1367]" gridcol="3" gridrow="3" pageId="19" pageNumber="500">absent</td>
<td id="DA5C331FFFAA1BBEFBC3FACFFB30163D" box="[1105,1266,1343,1367]" gridcol="4" gridrow="3" pageId="19" pageNumber="500">present</td>
<td id="DA5C331FFFAA1BBEFA88FACFFA7D163D" box="[1306,1471,1343,1367]" gridcol="5" gridrow="3" pageId="19" pageNumber="500">present</td>
<td id="DA5C331FFFAA1BBEFA76FACFF9AB163D" box="[1508,1641,1343,1367]" gridcol="6" gridrow="3" pageId="19" pageNumber="500">-</td>
<td id="DA5C331FFFAA1BBEF905FACFF8F7163D" box="[1687,1845,1343,1367]" gridcol="7" gridrow="3" pageId="19" pageNumber="500">absent</td>
</tr>
<tr id="998D5A63FFAA1BBEFFEFFA97F8F71615" box="[125,1845,1383,1407]" gridrow="4" pageId="19" pageNumber="500">
<th id="DA5C331FFFAA1BBEFFEFFA97FF201615" box="[125,226,1383,1407]" gridcol="0" gridrow="4" pageId="19" pageNumber="500">
<emphasis id="15C98433FFAAE452FFEFFA98FF561615" bold="true" box="[125,148,1384,1407]" pageId="19" pageNumber="500">57</emphasis>
</th>
<td id="DA5C331FFFAA1BBEFE89FA97FE651615" box="[283,423,1383,1407]" gridcol="1" gridrow="4" pageId="19" pageNumber="500">3</td>
<td id="DA5C331FFFAA1BBEFE48FA97FD631615" box="[474,673,1383,1407]" gridcol="2" gridrow="4" pageId="19" pageNumber="500">present</td>
<td id="DA5C331FFFAA1BBEFD48FA97FBF71615" box="[730,1077,1383,1407]" gridcol="3" gridrow="4" pageId="19" pageNumber="500">absent</td>
<td id="DA5C331FFFAA1BBEFBC3FA97FB301615" box="[1105,1266,1383,1407]" gridcol="4" gridrow="4" pageId="19" pageNumber="500">present</td>
<td id="DA5C331FFFAA1BBEFA88FA97FA7D1615" box="[1306,1471,1383,1407]" gridcol="5" gridrow="4" pageId="19" pageNumber="500">present</td>
<td id="DA5C331FFFAA1BBEFA76FA97F9AB1615" box="[1508,1641,1383,1407]" gridcol="6" gridrow="4" pageId="19" pageNumber="500">10</td>
<td id="DA5C331FFFAA1BBEF905FA97F8F71615" box="[1687,1845,1383,1407]" gridcol="7" gridrow="4" pageId="19" pageNumber="500">absent</td>
</tr>
<tr id="998D5A63FFAA1BBEFFEFFA7FF8F716CD" box="[125,1845,1423,1447]" gridrow="5" pageId="19" pageNumber="500">
<th id="DA5C331FFFAA1BBEFFEFFA7FFF2016CD" box="[125,226,1423,1447]" gridcol="0" gridrow="5" pageId="19" pageNumber="500">
<emphasis id="15C98433FFAAE452FFEFFA60FF5616CD" bold="true" box="[125,148,1424,1447]" pageId="19" pageNumber="500">64</emphasis>
</th>
<td id="DA5C331FFFAA1BBEFE89FA7FFE6516CD" box="[283,423,1423,1447]" gridcol="1" gridrow="5" pageId="19" pageNumber="500">3</td>
<td id="DA5C331FFFAA1BBEFE48FA7FFD6316CD" box="[474,673,1423,1447]" gridcol="2" gridrow="5" pageId="19" pageNumber="500">present</td>
<td id="DA5C331FFFAA1BBEFD48FA7FFBF716CD" box="[730,1077,1423,1447]" gridcol="3" gridrow="5" pageId="19" pageNumber="500">present</td>
<td id="DA5C331FFFAA1BBEFBC3FA7FFB3016CD" box="[1105,1266,1423,1447]" gridcol="4" gridrow="5" pageId="19" pageNumber="500">absent</td>
<td id="DA5C331FFFAA1BBEFA88FA7FFA7D16CD" box="[1306,1471,1423,1447]" gridcol="5" gridrow="5" pageId="19" pageNumber="500">present</td>
<td id="DA5C331FFFAA1BBEFA76FA7FF9AB16CD" box="[1508,1641,1423,1447]" gridcol="6" gridrow="5" pageId="19" pageNumber="500">10</td>
<td id="DA5C331FFFAA1BBEF905FA7FF8F716CD" box="[1687,1845,1423,1447]" gridcol="7" gridrow="5" pageId="19" pageNumber="500">present</td>
</tr>
<tr id="998D5A63FFAA1BBEFFEFFA48F8F716BA" box="[125,1845,1464,1488]" gridrow="6" pageId="19" pageNumber="500">
<th id="DA5C331FFFAA1BBEFFEFFA48FF2016BA" box="[125,226,1464,1488]" gridcol="0" gridrow="6" pageId="19" pageNumber="500">
<emphasis id="15C98433FFAAE452FFEFFA49FF6216BA" bold="true" box="[125,160,1465,1488]" pageId="19" pageNumber="500">66+</emphasis>
</th>
<td id="DA5C331FFFAA1BBEFE89FA48FE6516BA" box="[283,423,1464,1488]" gridcol="1" gridrow="6" pageId="19" pageNumber="500">3</td>
<td id="DA5C331FFFAA1BBEFE48FA48FD6316BA" box="[474,673,1464,1488]" gridcol="2" gridrow="6" pageId="19" pageNumber="500">present</td>
<td id="DA5C331FFFAA1BBEFD48FA48FBF716BA" box="[730,1077,1464,1488]" gridcol="3" gridrow="6" pageId="19" pageNumber="500">present</td>
<td id="DA5C331FFFAA1BBEFBC3FA48FB3016BA" box="[1105,1266,1464,1488]" gridcol="4" gridrow="6" pageId="19" pageNumber="500">absent</td>
<td id="DA5C331FFFAA1BBEFA88FA48FA7D16BA" box="[1306,1471,1464,1488]" gridcol="5" gridrow="6" pageId="19" pageNumber="500">present</td>
<td id="DA5C331FFFAA1BBEFA76FA48F9AB16BA" box="[1508,1641,1464,1488]" gridcol="6" gridrow="6" pageId="19" pageNumber="500">10</td>
<td id="DA5C331FFFAA1BBEF905FA48F8F716BA" box="[1687,1845,1464,1488]" gridcol="7" gridrow="6" pageId="19" pageNumber="500">present</td>
</tr>
</table>
</paragraph>
<paragraph id="27025821FFAAE452FF07FA04FB6B1561" blockId="19.[149,1193,1524,1547]" box="[149,1193,1524,1547]" pageId="19" pageNumber="500">. chaetiger; LA. lateral antennae; VC. ventral cirri; Ant. Prov. anterior provisional; Post. prov. posterior provisional.</paragraph>
<paragraph id="27025821FFADE455FF55FF68FF6E122A" blockId="20.[151,1436,151,1043]" pageId="20" pageNumber="501">
In addition, four recently settled juveniles were collected from the same location. The specimens range from 5.3–6.4+ mm in length for 46–66+ chaetigers and measured 
<quantity id="E045F5C4FFADE455FCADFF4CFC2013BE" box="[831,994,188,213]" metricMagnitude="-4" metricUnit="m" metricValue="4.6" metricValueMax="5.2" metricValueMin="4.0" pageId="20" pageNumber="501" unit="mm" value="0.46" valueMax="0.52" valueMin="0.4">0.40–0.52 mm</quantity>
in width (
<tableCitation id="6A3F6D9AFFADE455FBC6FF4CFB6913BE" box="[1108,1195,188,213]" captionStart="TABLE 3" captionStartId="19.[125,202,666,689]" captionTargetBox="[147,1878,732,1088]" captionTargetPageId="19" captionText="TABLE 3. Ontogeny of Onuphis hanneloreae sp. nov. juveniles" httpUri="http://table.plazi.org/id/73C208A9FFAAE452FFEFFD6AFD0711DB" pageId="20" pageNumber="501" tableUuid="73C208A9FFAAE452FFEFFD6AFD0711DB">Table 3</tableCitation>
). Three of them have a brownish pigmented colour pattern. All have branchiae, the two larger ones have single branchiae from chaetiger 2, the smaller one has single branchiae from chaetiger 5 and the 57 chaetiger specimen has branchia from chaetiger 3.
</paragraph>
<paragraph id="27025821FFADE455FF55FEBCFDBC1051" blockId="20.[151,1436,151,1043]" pageId="20" pageNumber="501">
The chaetal composition of juveniles differs from that of adults and has been detailed for these 
<taxonomicName id="E0BD23A2FFADE455FB7DFEBCFA5E120E" box="[1263,1436,332,356]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="20" pageNumber="501" phylum="Annelida" rank="species" species="hanneloreae">
<emphasis id="15C98433FFADE455FB7DFEBCFA5E120E" box="[1263,1436,332,356]" italics="true" pageId="20" pageNumber="501">O. hanneloreae</emphasis>
</taxonomicName>
specimens in 
<figureCitation id="BF8644A4FFADE455FEA2FE9FFE4012E2" box="[304,386,367,392]" captionStart="FIGURE 11" captionStartId="17.[151,250,2005,2027]" captionTargetBox="[183,1401,1124,1933]" captionTargetId="figure@17.[151,1436,1093,1984]" captionTargetPageId="17" captionText="FIGURE 11. Comparison of the progression of hooks in Onuphis hanneloreae sp. nov. juveniles." httpUri="http://zenodo.org/record/155282/files/figure.png" pageId="20" pageNumber="501">Fig. 11</figureCitation>
. The smallest examined juveniles have two to four compound hooks in the first three pairs of parapodia. In the 28- and 33- chaetiger specimens they have only bidentate and the 39-chaetiger specimen has bi- and tridentate hooks. Parapodia 4–9 have two limbate chaetae and two kinds of anterior provisional subacicular hooks (SAHs): one with a relatively large appendage in the medial position of the parapodium (
<figureCitation id="BF8644A4FFADE455FB5EFE2CFADE129E" box="[1228,1308,476,501]" captionStart="FIGURE 12" captionStartId="18.[151,250,1953,1975]" captionTargetBox="[202,1384,193,1930]" captionTargetId="figure@18.[202,1385,193,1932]" captionTargetPageId="18" captionText="FIGURE 12. Onuphis hanneloreae sp. nov. juvenile. Scanning electron micrographs. A, bidentate compound hook; B, median anterior provisional subacicular hook; C, ventral anterior provisional subacicular hook; D, posterior provisional subacicular hook; E, posterior parapodium showing both permanent subacicular hook (top) and provisional subacicular hook (bottom)." httpUri="http://zenodo.org/record/155283/files/figure.png" pageId="20" pageNumber="501">Fig. 12</figureCitation>
<collectingRegion id="E57996C3FFADE455FA8FFE2CFAF1129E" box="[1309,1331,476,500]" country="Nigeria" name="Rivers" pageId="20" pageNumber="501">A</collectingRegion>
) and two ventral hooks with smaller appendages (
<figureCitation id="BF8644A4FFADE455FDF2FE0FFD711172" box="[608,691,511,536]" captionStart="FIGURE 12" captionStartId="18.[151,250,1953,1975]" captionTargetBox="[202,1384,193,1930]" captionTargetId="figure@18.[202,1385,193,1932]" captionTargetPageId="18" captionText="FIGURE 12. Onuphis hanneloreae sp. nov. juvenile. Scanning electron micrographs. A, bidentate compound hook; B, median anterior provisional subacicular hook; C, ventral anterior provisional subacicular hook; D, posterior provisional subacicular hook; E, posterior parapodium showing both permanent subacicular hook (top) and provisional subacicular hook (bottom)." httpUri="http://zenodo.org/record/155283/files/figure.png" pageId="20" pageNumber="501">Fig. 12</figureCitation>
B, C). Two posterior provisional SAHs start on chaetiger 10 (
<figureCitation id="BF8644A4FFADE455FAFDFE0FFF761156" captionStart="FIGURE 12" captionStartId="18.[151,250,1953,1975]" captionTargetBox="[202,1384,193,1930]" captionTargetId="figure@18.[202,1385,193,1932]" captionTargetPageId="18" captionText="FIGURE 12. Onuphis hanneloreae sp. nov. juvenile. Scanning electron micrographs. A, bidentate compound hook; B, median anterior provisional subacicular hook; C, ventral anterior provisional subacicular hook; D, posterior provisional subacicular hook; E, posterior parapodium showing both permanent subacicular hook (top) and provisional subacicular hook (bottom)." httpUri="http://zenodo.org/record/155283/files/figure.png" pageId="20" pageNumber="501">Fig. 12</figureCitation>
D), being present to almost the end of the body. The brooded juveniles lack permanent subacicular hooks and pectinate chaetae. The recently settled juveniles have bi- and tridentate compound to pseudocompound hooks on the first three parapodia. They have four to five hooks (compound and pseudocompound) and one limbate chaeta in the first two parapodia and two or three compound hooks and two or three limbates in the 3rd parapodium. The anterior provisional SAHs are now restricted to chaetiger 7–8 to 9, with chaetigers 4–6 or 7 lacking any hooks, having only limbate chaetae. Two permanent SAHs are present from chaetiger 10–37 or 38 when first one (
<figureCitation id="BF8644A4FFADE455FAFDFD2AFF76107D" captionStart="FIGURE 12" captionStartId="18.[151,250,1953,1975]" captionTargetBox="[202,1384,193,1930]" captionTargetId="figure@18.[202,1385,193,1932]" captionTargetPageId="18" captionText="FIGURE 12. Onuphis hanneloreae sp. nov. juvenile. Scanning electron micrographs. A, bidentate compound hook; B, median anterior provisional subacicular hook; C, ventral anterior provisional subacicular hook; D, posterior provisional subacicular hook; E, posterior parapodium showing both permanent subacicular hook (top) and provisional subacicular hook (bottom)." httpUri="http://zenodo.org/record/155283/files/figure.png" pageId="20" pageNumber="501">Fig. 12</figureCitation>
E), and then both permanent SAHs are replaced by posterior provisional SAHs. 
<collectingRegion id="E57996C3FFADE455FBBEFD0FFB82107D" box="[1068,1088,767,791]" country="Nigeria" name="Rivers" pageId="20" pageNumber="501">A</collectingRegion>
flat pectinate chaeta with 9–10 teeth is present from chaetiger 10 to 20–30.
</paragraph>
<paragraph id="27025821FFADE455FF55FCB6FD871779" blockId="20.[151,1436,151,1043]" pageId="20" pageNumber="501">
The branchial development commences when juveniles consist of about 30 chaetigers. The branchiae start on chaetiger 6, from there they spread more posteriorly but also anteriorly, passing through intermediate forms with branchiae starting on chaetiger 5, 4, 3 and 2. When the worm has attained a width of about 
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, or about twothirds of its maximum size, the branchiae begin on chaetiger 1 which is considered here as representing the adult condition of the species. Similar patterns in branchial progression have been observed in species of 
<taxonomicName id="E0BD23A2FFADE455FAB3FC26FA5E1085" box="[1313,1436,982,1007]" class="Polychaeta" family="Onuphidae" kingdom="Animalia" order="Eunicida" pageId="20" pageNumber="501" phylum="Annelida" rank="family">Onuphidae</taxonomicName>
(
<bibRefCitation id="432C25D0FFADE455FF0CFC0BFEF21779" author="Paxton" box="[158,304,1018,1043]" pageId="20" pageNumber="501" refString="Paxton, H. (1996) Hirsutonuphis (Polychaeta: Onuphidae) from Australia, with a discussion of setal progression in juveniles. Invertebrate Taxonomy, 10, 77 - 96. http: // dx. doi. org / 10.1071 / IT 9960077" type="journal article" year="1996">Paxton 1996</bibRefCitation>
, 
<bibRefCitation id="432C25D0FFADE455FEA9FC0AFDFB1779" author="Arias" box="[315,569,1018,1043]" pageId="20" pageNumber="501" refString="Arias, A. &amp; Paxton, H. (2015 a) Onuphis and Aponuphis (Annelida: Onuphidae) from southwestern Europe, with the description of a new species. Zootaxa, 3949 (3), 345 - 369. http: // dx. doi. org / 10.11646 / zootaxa. 3949.3.3" type="journal article" year="2015" yearSuffix="a">Arias &amp; Paxton 2015a</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>