338 lines
47 KiB
XML
338 lines
47 KiB
XML
<document id="04A2B50E48780D652DD511D141DE0CCC" ID="10.11646/zootaxa.4168.3.3" ID-DOI="10.11646/zootaxa.4168.3.3" ID-GBIF-Dataset="e637580d-a728-4232-be4e-4965c8f4d9b0" ID-ISSN="1175-5326" ID-Zenodo-Dep="155271" ID-ZooBank="CBAB62EC-4FB6-49BD-94DB-7954E0CF5AD0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1474008133758" checkinUser="plazi" docAuthor="Arias, Andrés" docDate="2016" docId="AF14E937FFADE454FF05FB96FD42141A" docLanguage="en" docName="zootaxa.4168.3.3.pdf" docOrigin="Zootaxa 4168 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Onuphis landanaensis Augener 1918" docType="treatment" docVersion="11" lastPageNumber="502" masterDocId="532D914FFFB9E441FF92FFF0FFC2136A" masterDocTitle="Onuphi s and Mooreonuphis (Annelida: Onuphidae) from West Africa with the description of three new species and the reinstatement of O. landanaensis Augener, 1918" masterLastPageNumber="511" masterPageNumber="481" pageNumber="501" updateTime="1698386055454" updateUser="plazi">
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<mods:title id="8B35A67C0A911213883A5F1B0D0E63F0">Onuphi s and Mooreonuphis (Annelida: Onuphidae) from West Africa with the description of three new species and the reinstatement of O. landanaensis Augener, 1918</mods:title>
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</mods:titleInfo>
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<mods:name id="F88CE5BAACD235EA055055860121E20D" type="personal">
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<mods:role id="7FCCD273963DD19EF90004B5C7628FE5">
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<mods:roleTerm id="2CDD8318B45A83DE14079E354FD3DCEA">Author</mods:roleTerm>
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<mods:namePart id="EED55049F34EBF7FC3993978D8E08BC8">Arias, Andrés</mods:namePart>
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<mods:typeOfResource id="7E8D225AD951C6A99BE19F6611B97881">text</mods:typeOfResource>
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<mods:title id="148EAAF0508484CF5651710F3DB17A72">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="74690B2AE40EB3D4371AEE43A1A410AF">
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<mods:date id="72020EE6D905F9C556D70EA7A5FC7940">2016</mods:date>
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<mods:detail id="D9215A303F2825AFAC0675F4C4EE8E6F" type="volume">
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<mods:number id="125227B08F3E418E4380112C34287C5F">4168</mods:number>
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</mods:detail>
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<mods:detail id="340C88FAC8ECF6AED4C068F3C0F4F7B4" type="issue">
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<mods:number id="69BA4BE0396561470F30747C17E89BD7">3</mods:number>
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<mods:start id="2D59AB7ADEE3B06E58511C957D0D98CC">481</mods:start>
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<mods:end id="DF028C2022A752CA73DFDA60617F1EAA">511</mods:end>
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<mods:identifier id="F361FCAB74795DD46554629AB06D1ED3" type="DOI">10.11646/zootaxa.4168.3.3</mods:identifier>
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<mods:identifier id="FD62AA9EE7FDB74D08D3725BBAAD9FE0" type="GBIF-Dataset">e637580d-a728-4232-be4e-4965c8f4d9b0</mods:identifier>
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<mods:identifier id="EEF02C6830DAE44EF2E89E9C73840B08" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="25F6F80C9BF57D3E518B1A78ECABCE4C" type="Zenodo-Dep">155271</mods:identifier>
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<treatment id="AF14E937FFADE454FF05FB96FD42141A" ID-DOI="http://doi.org/10.5281/zenodo.5632336" ID-GBIF-Taxon="124551878" ID-Zenodo-Dep="5632336" LSID="urn:lsid:plazi:treatment:AF14E937FFADE454FF05FB96FD42141A" httpUri="http://treatment.plazi.org/id/AF14E937FFADE454FF05FB96FD42141A" lastPageId="21" lastPageNumber="502" pageId="20" pageNumber="501">
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<subSubSection id="6FA70BAAFFADE455FF05FB96FDC117C8" pageId="20" pageNumber="501" type="nomenclature">
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<paragraph id="27025821FFADE455FF05FB96FD1417EA" blockId="20.[151,726,1126,1186]" box="[151,726,1126,1152]" pageId="20" pageNumber="501">
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<heading id="7C4AEF4DFFADE455FF05FB96FD1417EA" bold="true" box="[151,726,1126,1152]" fontSize="11" level="1" pageId="20" pageNumber="501" reason="1">
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<emphasis id="15C98433FFADE455FF05FB96FD1417EA" bold="true" box="[151,726,1126,1152]" pageId="20" pageNumber="501">
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<taxonomicName id="E0BD23A2FFADE455FF05FB96FDA217EA" ID-CoL="49QK6" authority="Augener, 1918" authorityName="Augener" authorityYear="1918" box="[151,608,1126,1152]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="20" pageNumber="501" phylum="Annelida" rank="species" species="landanaensis">
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<emphasis id="15C98433FFADE455FF05FB96FE6717EA" bold="true" box="[151,421,1126,1152]" italics="true" pageId="20" pageNumber="501">Onuphis landanaensis</emphasis>
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<bibRefCitation id="432C25D0FFADE455FE3FFB96FDA217EA" author="Augener" box="[429,608,1126,1152]" pageId="20" pageNumber="501" refString="Augener, H. (1918) Polychaeta. In: Michaelsen, W. (Ed.), Beitrage zur Kenntnis der Meeresfauna Westafrikas. 2 (2). Friedrichsen & Co, Hamburg, pp. 67 - 625." type="journal article" year="1918">Augener, 1918</bibRefCitation>
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</taxonomicName>
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emended
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="27025821FFADE455FF05FB79FDC117C8" blockId="20.[151,726,1126,1186]" box="[151,515,1161,1186]" pageId="20" pageNumber="501">
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||
<figureCitation id="BF8644A4FFADE455FF05FB79FEC317C8" box="[151,257,1161,1186]" captionStart="FIGURE 2" captionStartId="7.[151,250,1937,1959]" captionTargetBox="[151,1436,193,1915]" captionTargetId="figure@7.[151,1436,193,1916]" captionTargetPageId="7" captionText="FIGURE 2. Photomicrographs. Onuphis augeneri sp. nov. Holotype A, B (ZMH V. 868), Onuphis landanaensis lectotype C, D (ZMH V. 8674). A, anterior end, dorsal view; B, detailed view of dorsum under methyl blue staining; C, anterior end, dorsal view; D, detailed view of anterior end." httpUri="http://zenodo.org/record/155273/files/figure.png" pageId="20" pageNumber="501">Figures 2</figureCitation>
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C, D, 3, 13;
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<tableCitation id="6A3F6D9AFFADE455FE1BFB79FE2517C8" box="[393,487,1161,1186]" captionStart="TABLE 1" captionStartId="5.[125,202,650,670]" captionTargetBox="[125,1883,695,1111]" captionText="TABLE 1. Comparison of key features and habitats of the eastern Atlantic species of the Onuphis eremita complex." httpUri="http://table.plazi.org/id/73C208A9FFBCE444FFEFFD7AFB6011F5" pageId="20" pageNumber="501" tableUuid="73C208A9FFBCE444FFEFFD7AFB6011F5">Tables 1</tableCitation>
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,
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<tableCitation id="6A3F6D9AFFADE455FE67FB7AFDC117C8" box="[501,515,1162,1186]" captionStart="TABLE 2" captionStartId="6.[151,242,1166,1189]" captionTargetBox="[159,1374,1216,2029]" captionText="TABLE 2. Distinguishing features among O. augeneri sp. nov., O. landanaensis and O. texana." httpUri="http://table.plazi.org/id/73C208A9FFBFE447FF05FB7EFB5817CF" pageId="20" pageNumber="501" tableUuid="73C208A9FFBFE447FF05FB7EFB5817CF">2</tableCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="6FA70BAAFFADE455FF05FB21FB20178D" box="[151,1250,1233,1255]" pageId="20" pageNumber="501" type="reference_group">
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||
<paragraph id="27025821FFADE455FF05FB21FB20178D" blockId="20.[151,1250,1233,1255]" box="[151,1250,1233,1255]" pageId="20" pageNumber="501">
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<treatmentCitationGroup id="07AD7F0FFFADE455FF05FB21FB20178D" box="[151,1250,1233,1255]" pageId="20" pageNumber="501">
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||
<taxonomicName id="E0BD23A2FFADE455FF05FB21FD8B178D" ID-CoL="49QK6" authority="Augener, 1918: 339" authorityName="Augener" authorityPageNumber="339" authorityYear="1918" box="[151,585,1233,1255]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="20" pageNumber="501" phylum="Annelida" rank="species" species="landanaensis">
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<emphasis id="15C98433FFADE455FF05FB21FEBB178D" box="[151,377,1233,1255]" italics="true" pageId="20" pageNumber="501">Onuphis landanaensis</emphasis>
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<treatmentCitation id="A61C7E30FFADE455FEEDFB22FD8B178D" author="Augener" box="[383,585,1233,1255]" page="339" pageId="20" pageNumber="501" year="1918">
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||
<bibRefCitation id="432C25D0FFADE455FEEDFB22FDDA178D" author="Augener" box="[383,536,1233,1255]" pageId="20" pageNumber="501" refString="Augener, H. (1918) Polychaeta. In: Michaelsen, W. (Ed.), Beitrage zur Kenntnis der Meeresfauna Westafrikas. 2 (2). Friedrichsen & Co, Hamburg, pp. 67 - 625." type="journal article" year="1918">Augener, 1918</bibRefCitation>
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: 339
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</treatmentCitation>
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||
</taxonomicName>
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–343, Tafel V Figs. 135–138, Tafel VI Fig. 197, Textfig XXXVL.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="6FA70BAAFFADE455FF05FAE4FB75161F" pageId="20" pageNumber="501" type="materials_examined">
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<paragraph id="27025821FFADE455FF05FAE4FCC5163B" blockId="20.[151,1437,1300,2009]" pageId="20" pageNumber="501">
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<materialsCitation id="97D5527CFFADE455FF05FAE4FA551646" ID-GBIF-Occurrence="1313442840" box="[151,1431,1300,1325]" collectionCode="ZMH" collectorName="A. Hupfer" country="Benin" location="Whydah" pageId="20" pageNumber="501" specimenCount="2" stateProvince="West" typeStatus="lectotype">
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<emphasis id="15C98433FFADE455FF05FAE4FE131647" bold="true" box="[151,465,1300,1325]" pageId="20" pageNumber="501">
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<typeStatus id="F806E683FFADE455FF05FAE4FF101647" box="[151,210,1300,1325]" pageId="20" pageNumber="501">Type</typeStatus>
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material.
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<typeStatus id="F806E683FFADE455FEC9FAE4FE131647" box="[347,465,1300,1325]" pageId="20" pageNumber="501" type="lectotype">Lectotype</typeStatus>
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</emphasis>
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(
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<collectionCode id="41ACC0E4FFADE455FE7AFAE4FDEA1646" box="[488,552,1300,1324]" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="20" pageNumber="501">ZMH</collectionCode>
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V.8674),
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<location id="22620EFAFFADE455FD34FAE4FCCA1647" LSID="urn:lsid:plazi:treatment:AF14E937FFADE454FF05FB96FD42141A:22620EFAFFADE455FD34FAE4FCCA1647" box="[678,776,1300,1325]" country="Benin" name="Whydah" pageId="20" pageNumber="501" stateProvince="West">Whydah</location>
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,
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<collectingCountry id="5FAA18B1FFADE455FC89FAE4FCA31647" box="[795,865,1300,1325]" name="Benin" pageId="20" pageNumber="501">Benin</collectingCountry>
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||
,
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<collectingRegion id="E57996C3FFADE455FCE6FAE5FC691646" box="[884,939,1301,1324]" country="Cameroon" name="West" pageId="20" pageNumber="501">West</collectingRegion>
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Africa, coll.
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<collectorName id="8A483DF7FFADE455FBC5FAE4FB121647" box="[1111,1232,1300,1325]" pageId="20" pageNumber="501">A. Hupfer</collectorName>
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(prior to 1917)
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</materialsCitation>
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;
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<materialsCitation id="97D5527CFFADE455FF05FAC8FCC1163B" ID-GBIF-Occurrence="1313442841" box="[151,771,1336,1361]" collectionCode="ZMH" collectorName="A. Hupfer" country="Benin" location="Whydah" pageId="20" pageNumber="501" specimenCount="1" stateProvince="West" typeStatus="paralectotype">
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<emphasis id="15C98433FFADE455FF05FAC8FEF8163B" bold="true" box="[151,314,1336,1361]" pageId="20" pageNumber="501">
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||
<typeStatus id="F806E683FFADE455FF05FAC8FEF8163B" box="[151,314,1336,1361]" pageId="20" pageNumber="501" type="paralectotype">paralectotype</typeStatus>
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||
</emphasis>
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||
(
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<collectionCode id="41ACC0E4FFADE455FEDAFAC8FE4A163A" box="[328,392,1336,1360]" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="20" pageNumber="501">ZMH</collectionCode>
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P 27822), same data as lectotype
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="27025821FFADE455FF55FAACFB75161F" blockId="20.[151,1437,1300,2009]" box="[199,1207,1372,1397]" pageId="20" pageNumber="501">
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<materialsCitation id="97D5527CFFADE455FF55FAACFB71161F" ID-GBIF-Occurrence="1313442845" box="[199,1203,1372,1397]" country="Benin" county="Off Whydah" location="eastern Atlantic" municipality="Gulf of Guinea" pageId="20" pageNumber="501" specimenCount="1" stateProvince="West" typeStatus="holotype">
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<emphasis id="15C98433FFADE455FF55FAACFEA7161F" bold="true" box="[199,357,1372,1397]" pageId="20" pageNumber="501">
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||
<typeStatus id="F806E683FFADE455FF55FAACFEC0161F" box="[199,258,1372,1397]" pageId="20" pageNumber="501">Type</typeStatus>
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locality.
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</emphasis>
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<collectingCounty id="CE6320ADFFADE455FEFEFAACFE39161F" box="[364,507,1372,1397]" pageId="20" pageNumber="501">Off Whydah</collectingCounty>
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,
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<collectingCountry id="5FAA18B1FFADE455FD94FAACFD8E161F" box="[518,588,1372,1397]" name="Benin" pageId="20" pageNumber="501">Benin</collectingCountry>
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,
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<collectingMunicipality id="C766C25BFFADE455FDC5FAACFCC6161F" box="[599,772,1372,1397]" pageId="20" pageNumber="501">Gulf of Guinea</collectingMunicipality>
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,
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<collectingRegion id="E57996C3FFADE455FC9CFAADFC87161E" box="[782,837,1373,1396]" country="Cameroon" name="West" pageId="20" pageNumber="501">West</collectingRegion>
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Africa, tropical
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<location id="22620EFAFFADE455FC6DFAADFB71161F" LSID="urn:lsid:plazi:treatment:AF14E937FFADE454FF05FB96FD42141A:22620EFAFFADE455FC6DFAADFB71161F" box="[1023,1203,1372,1397]" country="Benin" county="Off Whydah" municipality="Gulf of Guinea" name="eastern Atlantic" pageId="20" pageNumber="501" stateProvince="West">eastern Atlantic</location>
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</materialsCitation>
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||
.
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</paragraph>
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</subSubSection>
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<subSubSection id="6FA70BAAFFADE455FF55FA70FC1E15D3" pageId="20" pageNumber="501" type="diagnosis">
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<paragraph id="27025821FFADE455FF55FA70FC1E15D3" blockId="20.[151,1437,1300,2009]" pageId="20" pageNumber="501">
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<emphasis id="15C98433FFADE455FF55FA70FE8316F3" bold="true" box="[199,321,1408,1433]" pageId="20" pageNumber="501">Diagnosis.</emphasis>
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Prostomium anteriorly extended, one pair of small eyespots present at anterolateral end of prostomium. Palps reaching chaetiger 1, lateral antennae reaching chaetiger 8–9 and median antenna reaching chaetiger 2–3. Ceratophores long and strongly ringed, palpophores with 14–15 rings, lateral antennophores with 17–19 rings and median antennophore with 14–15 rings. Subulate ventral cirri in first six chaetigers; distinct subulate postchaetal lobes in first 10 chaetigers. Tridentate pseudocompound hooks in first two chaetigers, slender long-appendaged hooks absent. Subacicular hooks from chaetiger 9. Flat, distally oblique pectinate chaetae with 10 teeth from chaetiger 9. Single branchial filaments from chaetiger 1 to 19–20, thereafter number increasing rapidly to maximum of three to four. Peristomium and first five chaetigers brown coloured and following chaetigers with two brown transverse bands per segment, anterior band wider than second.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="6FA70BAAFFADE454FF55F934FD2A10CE" lastPageId="21" lastPageNumber="502" pageId="20" pageNumber="501" type="description">
|
||
<paragraph id="27025821FFADE455FF55F934FD84144F" blockId="20.[151,1437,1300,2009]" pageId="20" pageNumber="501">
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||
<emphasis id="15C98433FFADE455FF55F934FE9B15B7" bold="true" box="[199,345,1732,1757]" pageId="20" pageNumber="501">Description.</emphasis>
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Based on
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<typeStatus id="F806E683FFADE455FE44F934FDFD15B7" box="[470,575,1732,1757]" pageId="20" pageNumber="501" type="lectotype">lectotype</typeStatus>
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||
, with variation of
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<typeStatus id="F806E683FFADE455FC8AF934FC6D15B7" box="[792,943,1732,1757]" pageId="20" pageNumber="501" type="paralectotype">paralectotype</typeStatus>
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included. Small and slender species. Both
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<typeStatus id="F806E683FFADE455FF05F919FF05146B" box="[151,199,1769,1793]" pageId="20" pageNumber="501">type</typeStatus>
|
||
specimens incomplete. Length of
|
||
<typeStatus id="F806E683FFADE455FDC1F918FD7B146B" box="[595,697,1768,1793]" pageId="20" pageNumber="501" type="lectotype">lectotype</typeStatus>
|
||
<quantity id="E045F5C4FFADE455FD51F918FCD9146A" box="[707,795,1768,1793]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="20" pageNumber="501" unit="mm" value="5.2">5.2 mm</quantity>
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||
for 37 chaetigers, width
|
||
<quantity id="E045F5C4FFADE455FBAEF918FB55146A" box="[1084,1175,1768,1792]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="20" pageNumber="501" unit="mm" value="0.4">0.4 mm</quantity>
|
||
;
|
||
<typeStatus id="F806E683FFADE455FB37F918FAFE146B" box="[1189,1340,1768,1793]" pageId="20" pageNumber="501" type="paralectotype">paralectotype</typeStatus>
|
||
<quantity id="E045F5C4FFADE455FAD7F918FA5F146A" box="[1349,1437,1768,1793]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="20" pageNumber="501" unit="mm" value="7.5">7.5 mm</quantity>
|
||
long,
|
||
<quantity id="E045F5C4FFADE455FF45F8FCFEEF144E" box="[215,301,1804,1829]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="20" pageNumber="501" unit="mm" value="0.5">0.5 mm</quantity>
|
||
wide with 43 chaetigers.
|
||
</paragraph>
|
||
<paragraph id="27025821FFADE455FF55F8C0FCF914FA" blockId="20.[151,1437,1300,2009]" pageId="20" pageNumber="501">
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||
Colour pattern consisting of peristomium and first five chaetigers completely brown coloured and following chaetigers with two brown transverse bands per segment, anterior band wider than the second, posterior band sometimes broken into two lateral patches (
|
||
<figureCitation id="BF8644A4FFADE455FDECF888FD0414FA" box="[638,710,1912,1937]" captionStart="FIGURE 2" captionStartId="7.[151,250,1937,1959]" captionTargetBox="[151,1436,193,1915]" captionTargetId="figure@7.[151,1436,193,1916]" captionTargetPageId="7" captionText="FIGURE 2. Photomicrographs. Onuphis augeneri sp. nov. Holotype A, B (ZMH V. 868), Onuphis landanaensis lectotype C, D (ZMH V. 8674). A, anterior end, dorsal view; B, detailed view of dorsum under methyl blue staining; C, anterior end, dorsal view; D, detailed view of anterior end." httpUri="http://zenodo.org/record/155273/files/figure.png" pageId="20" pageNumber="501">Figs 2</figureCitation>
|
||
C, D; 3
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||
<collectingRegion id="E57996C3FFADE455FC84F888FCEC14FA" box="[790,814,1912,1936]" country="Nigeria" name="Rivers" pageId="20" pageNumber="501">A</collectingRegion>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="27025821FFADE454FF55F86CFBB9122A" blockId="20.[151,1437,1300,2009]" lastBlockId="21.[151,1437,151,1904]" lastPageId="21" lastPageNumber="502" pageId="20" pageNumber="501">
|
||
Prostomium subtriangular with pair of conical frontal lips (
|
||
<figureCitation id="BF8644A4FFADE455FCF1F86CFC7714DE" box="[867,949,1948,1973]" captionStart="FIGURE 13" captionStartId="22.[151,250,1953,1975]" captionTargetBox="[309,1268,248,1878]" captionTargetId="figure@22.[162,1425,193,1932]" captionTargetPageId="22" captionText="FIGURE 13. Onuphis landanaensis. Lectotype (ZMH V. 8674). Line drawings. A, detailed view of prostomium, antennae omitted; B – D, tridentate pseudocompound hooks from chaetiger 1; E, pectinate chaeta from chaetiger 13." httpUri="http://zenodo.org/record/155284/files/figure.png" pageId="20" pageNumber="501">Fig. 13</figureCitation>
|
||
<collectingRegion id="E57996C3FFADE455FC27F86CFC0F14DE" box="[949,973,1948,1972]" country="Nigeria" name="Rivers" pageId="20" pageNumber="501">A</collectingRegion>
|
||
). Small pair of eyespots present, situated close to bases of lateral antennae (
|
||
<figureCitation id="BF8644A4FFADE455FDB2F830FDB014B2" box="[544,626,1984,2009]" captionStart="FIGURE 13" captionStartId="22.[151,250,1953,1975]" captionTargetBox="[309,1268,248,1878]" captionTargetId="figure@22.[162,1425,193,1932]" captionTargetPageId="22" captionText="FIGURE 13. Onuphis landanaensis. Lectotype (ZMH V. 8674). Line drawings. A, detailed view of prostomium, antennae omitted; B – D, tridentate pseudocompound hooks from chaetiger 1; E, pectinate chaeta from chaetiger 13." httpUri="http://zenodo.org/record/155284/files/figure.png" pageId="20" pageNumber="501">Fig. 13</figureCitation>
|
||
<collectingRegion id="E57996C3FFADE455FDE0F830FD4814B2" box="[626,650,1984,2008]" country="Nigeria" name="Rivers" pageId="20" pageNumber="501">A</collectingRegion>
|
||
). Palps reaching chaetiger 1 with 13–14 basal rings and longer distal ring in palpophores. Lateral antennae reaching chaetiger 8–9 with 16–18 basal rings in antennophores, median antenna reaching chaetiger 2–3 with 13–14 basal rings in antennophore, all three antennophores with longer distal ring (
|
||
<figureCitation id="BF8644A4FFACE454FF44FF2FFEDF1392" box="[214,285,223,248]" captionStart="FIGURE 2" captionStartId="7.[151,250,1937,1959]" captionTargetBox="[151,1436,193,1915]" captionTargetId="figure@7.[151,1436,193,1916]" captionTargetPageId="7" captionText="FIGURE 2. Photomicrographs. Onuphis augeneri sp. nov. Holotype A, B (ZMH V. 868), Onuphis landanaensis lectotype C, D (ZMH V. 8674). A, anterior end, dorsal view; B, detailed view of dorsum under methyl blue staining; C, anterior end, dorsal view; D, detailed view of anterior end." httpUri="http://zenodo.org/record/155273/files/figure.png" pageId="21" pageNumber="502">Fig. 2</figureCitation>
|
||
D). Palpostyles and antennostyles with irregular longitudinal rows of sensory buds. Nuchal grooves straight with narrow mid-dorsal separation. Peristomial cirri inserted distally on peristomium slightly lateral to lateral antennae, longer than peristomium, not exceeding anterior margin of prostomium.
|
||
</paragraph>
|
||
<paragraph id="27025821FFACE454FF55FEBCFDFC1172" blockId="21.[151,1437,151,1904]" pageId="21" pageNumber="502">
|
||
Anterior chaetigers (1–3) slighty longer than those following (
|
||
<figureCitation id="BF8644A4FFACE454FC18FEBCFC11120E" box="[906,979,332,357]" captionStart="FIGURE 2" captionStartId="7.[151,250,1937,1959]" captionTargetBox="[151,1436,193,1915]" captionTargetId="figure@7.[151,1436,193,1916]" captionTargetPageId="7" captionText="FIGURE 2. Photomicrographs. Onuphis augeneri sp. nov. Holotype A, B (ZMH V. 868), Onuphis landanaensis lectotype C, D (ZMH V. 8674). A, anterior end, dorsal view; B, detailed view of dorsum under methyl blue staining; C, anterior end, dorsal view; D, detailed view of anterior end." httpUri="http://zenodo.org/record/155273/files/figure.png" pageId="21" pageNumber="502">Figs 2</figureCitation>
|
||
C, D; 3
|
||
<collectingRegion id="E57996C3FFACE454FBB4FEBCFBFC120E" box="[1062,1086,332,356]" country="Nigeria" name="Rivers" pageId="21" pageNumber="502">A</collectingRegion>
|
||
). First three pairs of parapodia modified, not enlarged, directed slightly anterolaterally, with low prechaetal fold, triangular prechaetal lobe and spindle-shaped postchaetal lobe, longer than base of parapodium; digitate dorsal cirrus and ventral cirrus about as long as postchaetal lobe (
|
||
<figureCitation id="BF8644A4FFACE454FE24FE47FE3812BA" box="[438,506,439,464]" captionStart="FIGURE 3" captionStartId="8.[151,250,1953,1975]" captionTargetBox="[256,1322,247,1874]" captionTargetId="figure@8.[190,1397,193,1932]" captionTargetPageId="8" captionText="FIGURE 3. Onuphis landanaensis original line drawings of Augener (1918). A, anterior end, dorsal view; B, first parapodium, anterior view; C, median parapodium, anterior view; D, mandibles; E, maxillae; F, subacicular hook of median chaetiger; G, tridentate pseudocompound hook of modified chaetiger." httpUri="http://zenodo.org/record/155274/files/figure.png" pageId="21" pageNumber="502">Fig. 3</figureCitation>
|
||
B). Subulate ventral cirri in first six chaetigers and pad-like thereafter. Postchaetal lobe as distinct subulate lobe in first 10 chaetigers, becoming smaller and conical in shape thereafter. Interramal papilla at base of dorsal cirrus absent.
|
||
</paragraph>
|
||
<paragraph id="27025821FFACE454FF55FDD4FCCC110A" blockId="21.[151,1437,151,1904]" pageId="21" pageNumber="502">
|
||
Branchiae as single filament from first chaetiger to chaetiger 19–20 (
|
||
<figureCitation id="BF8644A4FFACE454FC6FFDD4FB861156" box="[1021,1092,548,573]" captionStart="FIGURE 3" captionStartId="8.[151,250,1953,1975]" captionTargetBox="[256,1322,247,1874]" captionTargetId="figure@8.[190,1397,193,1932]" captionTargetPageId="8" captionText="FIGURE 3. Onuphis landanaensis original line drawings of Augener (1918). A, anterior end, dorsal view; B, first parapodium, anterior view; C, median parapodium, anterior view; D, mandibles; E, maxillae; F, subacicular hook of median chaetiger; G, tridentate pseudocompound hook of modified chaetiger." httpUri="http://zenodo.org/record/155274/files/figure.png" pageId="21" pageNumber="502">Fig. 3</figureCitation>
|
||
B), then number of filaments increasing rapidly to maximum of three by chaetiger 40.
|
||
</paragraph>
|
||
<paragraph id="27025821FFACE454FF55FD9CFD2A10CE" blockId="21.[151,1437,151,1904]" pageId="21" pageNumber="502">
|
||
Aciculae yellowish with pointed tips, usually three per parapodium. Hooded pseudocompound hooks in first two chaetigers. First two parapodia (
|
||
<figureCitation id="BF8644A4FFACE454FDA4FD7FFDBA11C2" box="[566,632,655,680]" captionStart="FIGURE 3" captionStartId="8.[151,250,1953,1975]" captionTargetBox="[256,1322,247,1874]" captionTargetId="figure@8.[190,1397,193,1932]" captionTargetPageId="8" captionText="FIGURE 3. Onuphis landanaensis original line drawings of Augener (1918). A, anterior end, dorsal view; B, first parapodium, anterior view; C, median parapodium, anterior view; D, mandibles; E, maxillae; F, subacicular hook of median chaetiger; G, tridentate pseudocompound hook of modified chaetiger." httpUri="http://zenodo.org/record/155274/files/figure.png" pageId="21" pageNumber="502">Fig. 3</figureCitation>
|
||
B) with following chaetal complement from superior to inferior part of chaetal fan: one to two simple chaetae, three protruding distal tips of aciculae, two to three slightly longappendaged tridentate pseudocompound hooks (
|
||
<figureCitation id="BF8644A4FFACE454FD42FD27FCDE119A" box="[720,796,727,752]" captionStart="FIGURE 3" captionStartId="8.[151,250,1953,1975]" captionTargetBox="[256,1322,247,1874]" captionTargetId="figure@8.[190,1397,193,1932]" captionTargetPageId="8" captionText="FIGURE 3. Onuphis landanaensis original line drawings of Augener (1918). A, anterior end, dorsal view; B, first parapodium, anterior view; C, median parapodium, anterior view; D, mandibles; E, maxillae; F, subacicular hook of median chaetiger; G, tridentate pseudocompound hook of modified chaetiger." httpUri="http://zenodo.org/record/155274/files/figure.png" pageId="21" pageNumber="502">Figs 3</figureCitation>
|
||
G, 13B, C), two to three short-appendaged tridentate pseudocompound hooks (
|
||
<figureCitation id="BF8644A4FFACE454FE2FFD0CFDD1107E" box="[445,531,764,789]" captionStart="FIGURE 13" captionStartId="22.[151,250,1953,1975]" captionTargetBox="[309,1268,248,1878]" captionTargetId="figure@22.[162,1425,193,1932]" captionTargetPageId="22" captionText="FIGURE 13. Onuphis landanaensis. Lectotype (ZMH V. 8674). Line drawings. A, detailed view of prostomium, antennae omitted; B – D, tridentate pseudocompound hooks from chaetiger 1; E, pectinate chaeta from chaetiger 13." httpUri="http://zenodo.org/record/155284/files/figure.png" pageId="21" pageNumber="502">Fig. 13</figureCitation>
|
||
D). All hooks of almost equal thickness. Pseudocompound hooks replaced by limbate chaetae from chaetiger 3. Flat, distally oblique pectinate chaetae with about 10 teeth from chaetiger 9, usually single (
|
||
<figureCitation id="BF8644A4FFACE454FEADFCB4FE4D1036" box="[319,399,836,861]" captionStart="FIGURE 13" captionStartId="22.[151,250,1953,1975]" captionTargetBox="[309,1268,248,1878]" captionTargetId="figure@22.[162,1425,193,1932]" captionTargetPageId="22" captionText="FIGURE 13. Onuphis landanaensis. Lectotype (ZMH V. 8674). Line drawings. A, detailed view of prostomium, antennae omitted; B – D, tridentate pseudocompound hooks from chaetiger 1; E, pectinate chaeta from chaetiger 13." httpUri="http://zenodo.org/record/155284/files/figure.png" pageId="21" pageNumber="502">Fig. 13</figureCitation>
|
||
E). Hooded bidentate subacicular hooks from chaetiger 9 (
|
||
<figureCitation id="BF8644A4FFACE454FB8AFCB4FB981036" box="[1048,1114,836,861]" captionStart="FIGURE 3" captionStartId="8.[151,250,1953,1975]" captionTargetBox="[256,1322,247,1874]" captionTargetId="figure@8.[190,1397,193,1932]" captionTargetPageId="8" captionText="FIGURE 3. Onuphis landanaensis original line drawings of Augener (1918). A, anterior end, dorsal view; B, first parapodium, anterior view; C, median parapodium, anterior view; D, mandibles; E, maxillae; F, subacicular hook of median chaetiger; G, tridentate pseudocompound hook of modified chaetiger." httpUri="http://zenodo.org/record/155274/files/figure.png" pageId="21" pageNumber="502">Fig. 3</figureCitation>
|
||
F). Mandibles (
|
||
<figureCitation id="BF8644A4FFACE454FA97FCB4FA841036" box="[1285,1350,836,861]" captionStart="FIGURE 3" captionStartId="8.[151,250,1953,1975]" captionTargetBox="[256,1322,247,1874]" captionTargetId="figure@8.[190,1397,193,1932]" captionTargetPageId="8" captionText="FIGURE 3. Onuphis landanaensis original line drawings of Augener (1918). A, anterior end, dorsal view; B, first parapodium, anterior view; C, median parapodium, anterior view; D, mandibles; E, maxillae; F, subacicular hook of median chaetiger; G, tridentate pseudocompound hook of modified chaetiger." httpUri="http://zenodo.org/record/155274/files/figure.png" pageId="21" pageNumber="502">Fig. 3</figureCitation>
|
||
D) with white calcified cutting plates and slender shafts. Maxillary formula:
|
||
<collectingCountry id="5FAA18B1FFACE454FC1CFC98FC771015" box="[910,949,872,895]" name="Mexico" pageId="21" pageNumber="502">Mx</collectingCountry>
|
||
I = 1 + 1,
|
||
<collectingCountry id="5FAA18B1FFACE454FBB5FC98FB8C1015" box="[1063,1102,872,895]" name="Mexico" pageId="21" pageNumber="502">Mx</collectingCountry>
|
||
II = 8 + 7,
|
||
<collectingCountry id="5FAA18B1FFACE454FB5BFC98FB321015" box="[1225,1264,872,895]" name="Mexico" pageId="21" pageNumber="502">Mx</collectingCountry>
|
||
III = 8 + 0,
|
||
<collectingCountry id="5FAA18B1FFACE454FAE7FC98FA5E1015" box="[1397,1436,872,895]" name="Mexico" pageId="21" pageNumber="502">Mx</collectingCountry>
|
||
IV =?7 +?7,
|
||
<collectingCountry id="5FAA18B1FFACE454FEA1FC7DFE9810CE" box="[307,346,909,932]" name="Mexico" pageId="21" pageNumber="502">Mx</collectingCountry>
|
||
V = 1 + 1,
|
||
<collectingCountry id="5FAA18B1FFACE454FE49FC7DFDC010CE" box="[475,514,909,932]" name="Mexico" pageId="21" pageNumber="502">Mx</collectingCountry>
|
||
VI absent (
|
||
<figureCitation id="BF8644A4FFACE454FD16FC7CFD0410CE" box="[644,710,908,933]" captionStart="FIGURE 3" captionStartId="8.[151,250,1953,1975]" captionTargetBox="[256,1322,247,1874]" captionTargetId="figure@8.[190,1397,193,1932]" captionTargetPageId="8" captionText="FIGURE 3. Onuphis landanaensis original line drawings of Augener (1918). A, anterior end, dorsal view; B, first parapodium, anterior view; C, median parapodium, anterior view; D, mandibles; E, maxillae; F, subacicular hook of median chaetiger; G, tridentate pseudocompound hook of modified chaetiger." httpUri="http://zenodo.org/record/155274/files/figure.png" pageId="21" pageNumber="502">Fig. 3</figureCitation>
|
||
E).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6FA70BAAFFACE454FF55FC5FFD5C175F" pageId="21" pageNumber="502" type="distribution">
|
||
<paragraph id="27025821FFACE454FF55FC5FFD5C175F" blockId="21.[151,1437,151,1904]" pageId="21" pageNumber="502">
|
||
<emphasis id="15C98433FFACE454FF55FC5FFE3910A2" bold="true" box="[199,507,943,968]" pageId="21" pageNumber="502">Distribution and ecology.</emphasis>
|
||
In the original description by
|
||
<bibRefCitation id="432C25D0FFACE454FCF4FC40FBDC10A2" author="Augener" box="[870,1054,944,968]" pageId="21" pageNumber="502" refString="Augener, H. (1918) Polychaeta. In: Michaelsen, W. (Ed.), Beitrage zur Kenntnis der Meeresfauna Westafrikas. 2 (2). Friedrichsen & Co, Hamburg, pp. 67 - 625." type="journal article" year="1918">Augener (1918)</bibRefCitation>
|
||
,
|
||
<taxonomicName id="E0BD23A2FFACE454FBBCFC40FB1910A2" box="[1070,1243,944,968]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="21" pageNumber="502" phylum="Annelida" rank="species" species="landanensis">
|
||
<emphasis id="15C98433FFACE454FBBCFC40FB1910A2" box="[1070,1243,944,968]" italics="true" pageId="21" pageNumber="502">O. landanensis</emphasis>
|
||
</taxonomicName>
|
||
was reported as having a wide distribution in
|
||
<collectingRegion id="E57996C3FFACE454FE76FC25FDD91086" box="[484,539,981,1004]" country="Cameroon" name="West" pageId="21" pageNumber="502">West</collectingRegion>
|
||
Africa, from
|
||
<collectingCountry id="5FAA18B1FFACE454FD25FC24FCCA1087" box="[695,776,980,1005]" name="Liberia" pageId="21" pageNumber="502">Liberia</collectingCountry>
|
||
to
|
||
<collectingCountry id="5FAA18B1FFACE454FCBCFC24FC461087" box="[814,900,980,1005]" name="Angola" pageId="21" pageNumber="502">Angola</collectingCountry>
|
||
. However, herein its distribution is restricted to the shallow subtidal sandy bottoms of Whydah (the
|
||
<typeStatus id="F806E683FFACE454FD4BFC08FCCB177A" box="[729,777,1016,1040]" pageId="21" pageNumber="502">type</typeStatus>
|
||
locality of the
|
||
<typeStatus id="F806E683FFACE454FC22FC07FBDB177A" box="[944,1049,1015,1040]" pageId="21" pageNumber="502" type="lectotype">lectotype</typeStatus>
|
||
) in
|
||
<collectingCountry id="5FAA18B1FFACE454FBD1FC07FB4B177A" box="[1091,1161,1015,1040]" name="Benin" pageId="21" pageNumber="502">Benin</collectingCountry>
|
||
, inner part of the Gulf of
|
||
<collectingCountry id="5FAA18B1FFACE454FF05FBECFF29175F" box="[151,235,1052,1077]" name="Guinea-Bissau" pageId="21" pageNumber="502">Guinea</collectingCountry>
|
||
,
|
||
<collectingRegion id="E57996C3FFACE454FF64FBEDFEEF175E" box="[246,301,1053,1076]" country="Cameroon" name="West" pageId="21" pageNumber="502">West</collectingRegion>
|
||
Africa, tropical eastern Atlantic.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6FA70BAAFFACE454FF55FBCFFD42141A" pageId="21" pageNumber="502" type="discussion">
|
||
<paragraph id="27025821FFACE454FF55FBCFFA5F153A" blockId="21.[151,1437,151,1904]" pageId="21" pageNumber="502">
|
||
<emphasis id="15C98433FFACE454FF55FBCFFEF91732" bold="true" box="[199,315,1087,1112]" pageId="21" pageNumber="502">Remarks.</emphasis>
|
||
<taxonomicName id="E0BD23A2FFACE454FED3FBB0FDF81732" box="[321,570,1088,1112]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="21" pageNumber="502" phylum="Annelida" rank="species" species="landanaensis">
|
||
<emphasis id="15C98433FFACE454FED3FBB0FDF81732" box="[321,570,1088,1112]" italics="true" pageId="21" pageNumber="502">Onuphis landanaensis</emphasis>
|
||
</taxonomicName>
|
||
was synonymised with
|
||
<taxonomicName id="E0BD23A2FFACE454FCD6FBB0FC79173D" box="[836,955,1088,1111]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="21" pageNumber="502" phylum="Annelida" rank="species" species="eremita">
|
||
<emphasis id="15C98433FFACE454FCD6FBB0FC79173D" box="[836,955,1088,1111]" italics="true" pageId="21" pageNumber="502">O. eremita</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="432C25D0FFACE454FC71FBCFFB421732" author="Fauvel" box="[995,1152,1087,1112]" pageId="21" pageNumber="502" refString="Fauvel, P. (1923) Faune de France: Polychetes errantes. Paul Lechevalier, Paris, 488 pp." type="book" year="1923">Fauvel (1923)</bibRefCitation>
|
||
and has been regarded as such for nearly a century. But re-examination of the syntypes of
|
||
<taxonomicName id="E0BD23A2FFACE454FCFCFB94FBE71716" box="[878,1061,1124,1148]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="21" pageNumber="502" phylum="Annelida" rank="species" species="landanaensis">
|
||
<emphasis id="15C98433FFACE454FCFCFB94FBE71716" box="[878,1061,1124,1148]" italics="true" pageId="21" pageNumber="502">O. landanaensis</emphasis>
|
||
</taxonomicName>
|
||
in the ZMH (Hamburg) revealed the presence of not just one, but two species morphologically different from
|
||
<emphasis id="15C98433FFACE454FC64FB78FBB717F5" box="[1014,1141,1160,1183]" italics="true" pageId="21" pageNumber="502">
|
||
<taxonomicName id="E0BD23A2FFACE454FC64FB78FBB317F5" box="[1014,1137,1160,1183]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="21" pageNumber="502" phylum="Annelida" rank="species" species="eremita">O. eremita</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Of these two species, one differs considerably from the original description and thus it is described as a new species,
|
||
<taxonomicName id="E0BD23A2FFACE454FB22FB5CFAFA17AE" box="[1200,1336,1196,1220]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="21" pageNumber="502" phylum="Annelida" rank="species" species="augeneri" status="sp. nov.">
|
||
<emphasis id="15C98433FFACE454FB22FB5CFAFA17AE" box="[1200,1336,1196,1220]" italics="true" pageId="21" pageNumber="502">O. augeneri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="15C98433FFACE454FAD2FB5CFA5E17AF" bold="true" box="[1344,1436,1196,1221]" pageId="21" pageNumber="502">
|
||
<taxonomicNameLabel id="0EFA3948FFACE454FAD2FB5CFA5E17AF" box="[1344,1436,1196,1221]" pageId="21" pageNumber="502" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(see above) and the other, represented by two specimens from Whydah, Benin, displays a number of features that aligns it with the original description by
|
||
<bibRefCitation id="432C25D0FFACE454FDF1FB04FCDB1666" author="Augener" box="[611,793,1268,1292]" pageId="21" pageNumber="502" refString="Augener, H. (1918) Polychaeta. In: Michaelsen, W. (Ed.), Beitrage zur Kenntnis der Meeresfauna Westafrikas. 2 (2). Friedrichsen & Co, Hamburg, pp. 67 - 625." type="journal article" year="1918">Augener (1918)</bibRefCitation>
|
||
. Since
|
||
<bibRefCitation id="432C25D0FFACE454FCF8FB04FBDF1666" author="Augener" box="[874,1053,1268,1292]" pageId="21" pageNumber="502" refString="Augener, H. (1918) Polychaeta. In: Michaelsen, W. (Ed.), Beitrage zur Kenntnis der Meeresfauna Westafrikas. 2 (2). Friedrichsen & Co, Hamburg, pp. 67 - 625." type="journal article" year="1918">Augener (1918)</bibRefCitation>
|
||
did not choose a holotype, I have designated a lectotype and one paralectotype and thus formally reinstated
|
||
<taxonomicName id="E0BD23A2FFACE454FC7EFAE8FB66165A" box="[1004,1188,1304,1328]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="21" pageNumber="502" phylum="Annelida" rank="species" species="landanaensis">
|
||
<emphasis id="15C98433FFACE454FC7EFAE8FB66165A" box="[1004,1188,1304,1328]" italics="true" pageId="21" pageNumber="502">O. landanaensis</emphasis>
|
||
</taxonomicName>
|
||
. The main difference between the lectotype and paralectotype herein designated and the data from the original description is the number of anterior chaetigers with pseudocompound hooks, being two in the lectotype and paralectotype and reported as three in Augener’s description. This mistake can be easily explained, since at that time the possession of hooks on the first three chaetigers was considered a generic character of
|
||
<taxonomicName id="E0BD23A2FFACE454FCFEFA58FC0916AA" box="[876,971,1448,1472]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="21" pageNumber="502" phylum="Annelida" rank="genus">
|
||
<emphasis id="15C98433FFACE454FCFEFA58FC0916AA" box="[876,971,1448,1472]" italics="true" pageId="21" pageNumber="502">Onuphis</emphasis>
|
||
</taxonomicName>
|
||
, and often it was taken for granted. The remaining features fit quite well with the description and illustrations in
|
||
<bibRefCitation id="432C25D0FFACE454FC4AFA3CFB4E168E" author="Augener" box="[984,1164,1484,1508]" pageId="21" pageNumber="502" refString="Augener, H. (1918) Polychaeta. In: Michaelsen, W. (Ed.), Beitrage zur Kenntnis der Meeresfauna Westafrikas. 2 (2). Friedrichsen & Co, Hamburg, pp. 67 - 625." type="journal article" year="1918">Augener (1918)</bibRefCitation>
|
||
(
|
||
<figureCitation id="BF8644A4FFACE454FB0FFA3CFB21168E" box="[1181,1251,1484,1509]" captionStart="FIGURE 3" captionStartId="8.[151,250,1953,1975]" captionTargetBox="[256,1322,247,1874]" captionTargetId="figure@8.[190,1397,193,1932]" captionTargetPageId="8" captionText="FIGURE 3. Onuphis landanaensis original line drawings of Augener (1918). A, anterior end, dorsal view; B, first parapodium, anterior view; C, median parapodium, anterior view; D, mandibles; E, maxillae; F, subacicular hook of median chaetiger; G, tridentate pseudocompound hook of modified chaetiger." httpUri="http://zenodo.org/record/155274/files/figure.png" pageId="21" pageNumber="502">Fig. 3</figureCitation>
|
||
). Another small difference is related to the number of branchial filaments, reported as five or six by Augener (
|
||
<figureCitation id="BF8644A4FFACE454FA96FA1FFA881562" box="[1284,1354,1519,1544]" captionStart="FIGURE 3" captionStartId="8.[151,250,1953,1975]" captionTargetBox="[256,1322,247,1874]" captionTargetId="figure@8.[190,1397,193,1932]" captionTargetPageId="8" captionText="FIGURE 3. Onuphis landanaensis original line drawings of Augener (1918). A, anterior end, dorsal view; B, first parapodium, anterior view; C, median parapodium, anterior view; D, mandibles; E, maxillae; F, subacicular hook of median chaetiger; G, tridentate pseudocompound hook of modified chaetiger." httpUri="http://zenodo.org/record/155274/files/figure.png" pageId="21" pageNumber="502">Fig. 3</figureCitation>
|
||
C) and appearing as maximum of three in the lectotype and paralectotype. However, this is not a major issue since the number of branchial filaments is a growth-related feature and the statement could have related to a larger specimen.
|
||
</paragraph>
|
||
<paragraph id="27025821FFACE454FF55F9ACFD42141A" blockId="21.[151,1437,151,1904]" pageId="21" pageNumber="502">
|
||
<taxonomicName id="E0BD23A2FFACE454FF55F9ACFE03151E" box="[199,449,1628,1652]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="21" pageNumber="502" phylum="Annelida" rank="species" species="landanaensis">
|
||
<emphasis id="15C98433FFACE454FF55F9ACFE03151E" box="[199,449,1628,1652]" italics="true" pageId="21" pageNumber="502">Onuphis landanaensis</emphasis>
|
||
</taxonomicName>
|
||
can be readily distingished from the
|
||
<typeStatus id="F806E683FFACE454FCF3F9ADFC53151F" box="[865,913,1629,1653]" pageId="21" pageNumber="502">type</typeStatus>
|
||
species (
|
||
<taxonomicName id="E0BD23A2FFACE454FC6BF9ACFBB2151E" box="[1017,1136,1628,1652]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="21" pageNumber="502" phylum="Annelida" rank="species" species="eremita">
|
||
<emphasis id="15C98433FFACE454FC6BF9ACFBB2151E" box="[1017,1136,1628,1652]" italics="true" pageId="21" pageNumber="502">O. eremita</emphasis>
|
||
</taxonomicName>
|
||
) by the possession of only two anterior chaetigers with pseudocompound hooks (three in
|
||
<taxonomicName id="E0BD23A2FFACE454FCC2F970FC0415FD" box="[848,966,1664,1687]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="21" pageNumber="502" phylum="Annelida" rank="species" species="eremita">
|
||
<emphasis id="15C98433FFACE454FCC2F970FC0415FD" box="[848,966,1664,1687]" italics="true" pageId="21" pageNumber="502">O. eremita</emphasis>
|
||
</taxonomicName>
|
||
), the absence of slender long-appendaged ones and the interramal papillae at base of dorsal cirri (both characters present in
|
||
<taxonomicName id="E0BD23A2FFACE454FBB0F954FB5A15D6" box="[1058,1176,1700,1724]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="21" pageNumber="502" phylum="Annelida" rank="species" species="eremita">
|
||
<emphasis id="15C98433FFACE454FBB0F954FB5A15D6" box="[1058,1176,1700,1724]" italics="true" pageId="21" pageNumber="502">O. eremita</emphasis>
|
||
</taxonomicName>
|
||
) and the appearance of the SAHs from chaetiger 9 (from chaetiger
|
||
<quantity id="E045F5C4FFACE454FD04F938FD10158A" box="[662,722,1735,1760]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="21" pageNumber="502" unit="in" value="10.0">10 in</quantity>
|
||
<taxonomicName id="E0BD23A2FFACE454FD4EF938FC9715B5" box="[732,853,1736,1759]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="21" pageNumber="502" phylum="Annelida" rank="species" species="eremita">
|
||
<emphasis id="15C98433FFACE454FD4EF938FC9715B5" box="[732,853,1736,1759]" italics="true" pageId="21" pageNumber="502">O. eremita</emphasis>
|
||
</taxonomicName>
|
||
). Other distinguishing features between these two species and the other eastern Atlantic members of the
|
||
<taxonomicName id="E0BD23A2FFACE454FC9FF91CFC45146E" box="[781,903,1772,1796]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="21" pageNumber="502" phylum="Annelida" rank="species" species="eremita">
|
||
<emphasis id="15C98433FFACE454FC9FF91CFC45146E" box="[781,903,1772,1796]" italics="true" pageId="21" pageNumber="502">O. eremita</emphasis>
|
||
</taxonomicName>
|
||
complex are summarised in
|
||
<tableCitation id="6A3F6D9AFFACE454FB45F91CFAEC146E" box="[1239,1326,1772,1797]" captionStart="TABLE 1" captionStartId="5.[125,202,650,670]" captionTargetBox="[125,1883,695,1111]" captionText="TABLE 1. Comparison of key features and habitats of the eastern Atlantic species of the Onuphis eremita complex." httpUri="http://table.plazi.org/id/73C208A9FFBCE444FFEFFD7AFB6011F5" pageId="21" pageNumber="502" tableUuid="73C208A9FFBCE444FFEFFD7AFB6011F5">Table 1</tableCitation>
|
||
.
|
||
<taxonomicName id="E0BD23A2FFACE454FAAFF91CFEE91442" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="21" pageNumber="502" phylum="Annelida" rank="species" species="landanaensis">
|
||
<emphasis id="15C98433FFACE454FAAFF91CFEE91442" italics="true" pageId="21" pageNumber="502">Onuphis landanaensis</emphasis>
|
||
</taxonomicName>
|
||
closely resembles
|
||
<taxonomicName id="E0BD23A2FFACE454FD94F8E0FD4C144D" box="[518,654,1808,1831]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="21" pageNumber="502" phylum="Annelida" rank="species" species="augeneri" status="sp. nov.">
|
||
<emphasis id="15C98433FFACE454FD94F8E0FD4C144D" box="[518,654,1808,1831]" italics="true" pageId="21" pageNumber="502">O. augeneri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="15C98433FFACE454FD05F8FFFD311442" bold="true" box="[663,755,1807,1832]" pageId="21" pageNumber="502">
|
||
<taxonomicNameLabel id="0EFA3948FFACE454FD05F8FFFD311442" box="[663,755,1807,1832]" pageId="21" pageNumber="502" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="E0BD23A2FFACE454FCBCF8E0FC5F1442" box="[814,925,1808,1832]" class="Polychaeta" family="Onuphidae" genus="Onuphis" kingdom="Animalia" order="Eunicida" pageId="21" pageNumber="502" phylum="Annelida" rank="species" species="texana">
|
||
<emphasis id="15C98433FFACE454FCBCF8E0FC5F1442" box="[814,925,1808,1832]" italics="true" pageId="21" pageNumber="502">O. texana</emphasis>
|
||
</taxonomicName>
|
||
but differ from these two species by having only tridentate pseudocompound hooks on the first two chaetigers. Additional distinctive characteristics among these three species are presented in
|
||
<tableCitation id="6A3F6D9AFFACE454FDB5F8A7FDBE141A" box="[551,636,1879,1904]" captionStart="TABLE 2" captionStartId="6.[151,242,1166,1189]" captionTargetBox="[159,1374,1216,2029]" captionText="TABLE 2. Distinguishing features among O. augeneri sp. nov., O. landanaensis and O. texana." httpUri="http://table.plazi.org/id/73C208A9FFBFE447FF05FB7EFB5817CF" pageId="21" pageNumber="502" tableUuid="73C208A9FFBFE447FF05FB7EFB5817CF">Table 2</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |