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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.246.4112" ID-GBIF-Dataset="383fd8fc-74c6-4d8a-a6f8-df0c3bd37863" ID-PMC="PMC3520144" ID-Pensoft-Pub="1313-2970-246-27" ID-PubMed="23275749" ModsDocAuthor="" ModsDocDate="2012" ModsDocID="1313-2970-246-27" ModsDocOrigin="ZooKeys 246" ModsDocTitle="A new species of Lesticus Dejean, 1828 (Coleoptera, Carabidae) from the Finisterre Range, Papua New Guinea and a key to the genera of pterostichine-like Harpalinae of New Guinea" checkinTime="1451247896536" checkinUser="pensoft" docAuthor="Will, Kipling &amp; Kavanaugh, David H." docDate="2012" docId="25952439F93AB8226FA5A9E6D1315B9C" docLanguage="en" docName="ZooKeys 246: 27-37" docOrigin="ZooKeys 246" docSource="http://dx.doi.org/10.3897/zookeys.246.4112" docTitle="Lesticus finisterrae Will and Kavanaugh, sp. n." docType="treatment" docVersion="4" lastPageNumber="33" masterDocId="5A5B28367F0D7D66E01CFFDAFF9EC77B" masterDocTitle="A new species of Lesticus Dejean, 1828 (Coleoptera, Carabidae) from the Finisterre Range, Papua New Guinea and a key to the genera of pterostichine-like Harpalinae of New Guinea" masterLastPageNumber="37" masterPageNumber="27" pageNumber="29" updateTime="1668154969986" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A new species of Lesticus Dejean, 1828 (Coleoptera, Carabidae) from the Finisterre Range, Papua New Guinea and a key to the genera of pterostichine-like Harpalinae of New Guinea</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Will, Kipling</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kavanaugh, David H.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2012</mods:date>
<mods:detail type="volume">
<mods:number>246</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>27</mods:start>
<mods:end>37</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.246.4112</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.246.4112</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-246-27</mods:identifier>
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<treatment ID-GBIF-Taxon="152039057" LSID="urn:lsid:zoobank.org:act:52A9C3D7-A304-4D05-83DC-D1E385182B6F" httpUri="http://treatment.plazi.org/id/25952439F93AB8226FA5A9E6D1315B9C" lastPageId="6" lastPageNumber="33" pageId="2" pageNumber="29">
<subSubSection pageId="2" pageNumber="29" type="nomenclature">
<paragraph pageId="2" pageNumber="29">
<taxonomicName LSID="urn:lsid:zoobank.org:act:52A9C3D7-A304-4D05-83DC-D1E385182B6F" authority="Will and Kavanaugh" class="Insecta" family="Carabidae" genus="Lesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lesticus finisterrae" order="Coleoptera" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="finisterrae">Lesticus finisterrae Will and Kavanaugh</taxonomicName>
<taxonomicNameLabel pageId="2" pageNumber="29">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="29" type="holotype">
<paragraph pageId="2" pageNumber="29">Holotype.</paragraph>
<paragraph pageId="2" pageNumber="29">
Male. &quot;PAPUA NEW GUINEA, Madang/Morobe Province border, Finisterre Range, Teptep area, 3.5 air km WNW of Kewieng No.4 village, 3050 m, 28 March 1989 stop #89-50// D. H. Kavanaugh, G. E. Ball &amp; N. D. Penny collectors, Cal. Acad. Sci. Specimen//Papua New Guinea Expedition - 1989// California Academy of Sciences Type No. 18684//HOLOTYPE
<taxonomicName class="Insecta" family="Carabidae" genus="Lesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lesticus finisterrae" order="Coleoptera" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="finisterrae">Lesticus finisterrae</taxonomicName>
Will &amp; Kavanaugh [red label]&quot;. Deposited CASC.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="29" type="type locality">
<paragraph pageId="2" pageNumber="29">Type locality.</paragraph>
<paragraph pageId="2" pageNumber="29">
Finisterre Range, Papua New Guinea.
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-5.99778">5.99778°S</geoCoordinate>
, 52280°W.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="29" type="diagnosis">
<paragraph pageId="2" pageNumber="29">Diagnosis.</paragraph>
<paragraph pageId="2" pageNumber="29">
This species shares with other
<taxonomicName class="Insecta" family="Carabidae" genus="Lesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lesticus" order="Coleoptera" pageId="2" pageNumber="29" phylum="Arthropoda" rank="genus">Lesticus</taxonomicName>
species sharply hooked mandibular apices, relatively broad mentum, extremely wide gula (nearly the width of the mentum) and has the general form of a modified species of the
<taxonomicName class="Insecta" family="Carabidae" genus="Lesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lesticus chloronotus" order="Coleoptera" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="chloronotus">Lesticus chloronotus</taxonomicName>
group. It is distinguished from all other species of
<taxonomicName class="Insecta" family="Carabidae" genus="Lesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lesticus" order="Coleoptera" pageId="2" pageNumber="29" phylum="Arthropoda" rank="genus">Lesticus</taxonomicName>
, including the New Guinea species covered in
<normalizedToken originalValue="Darlingtons">Darlington's</normalizedToken>
keys and descriptions (1962, 1971) and
<bibRefCitation author="Dubault, G" journalOrPublisher="Bulletin de la Societe entomolologique de France" pageId="6" pageNumber="33" pagination="239 - 248" title="Les generes des &quot; trigonotomi &quot;: Pareuyaptus n. gen. et revision des Euryaptus Bates, 1892 (Coleoptera, Pterostichidae)." volume="113" year="2008">Dubault et al.'s key (2008)</bibRefCitation>
, by the combination of three or more setae in elytral intervals 3 and 5, the absence of transverse sulci of abdominal ventrites 4-5 and the presence of extremely prominent eyes.
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="30" pageId="2" pageNumber="29" type="description">
<paragraph pageId="2" pageNumber="29">Description.</paragraph>
<paragraph lastPageId="3" lastPageNumber="30" pageId="2" pageNumber="29">
(Fig. 3), Size. Overall length (sbl) 20.0mm, greatest width over elytra 8.5mm. Color. Dorsal and ventral surfaces black to brunneous. Legs, mouthparts, and antennae slightly paler, lateral margins of pronotum and lateral and sutural areas of elytra margins piceous to rufopiceous. Luster. Dorsally and ventrally shiny. Iridescence. Elytra and ventral surface of body without spectral iridescence. Head. Dorsal microsculpture absent, entire surface with micropunctulae, clypeal-ocular sulci impression absent, shallowly rugose above eyes, with broad, shallow paramedial depressions, ocular ratio (greatest width over eyes/width between eyes at level of anterior supraorbital setae) 1.29, eyes moderately large size, very prominently
<normalizedToken originalValue="“bulging”">&quot;bulging&quot;</normalizedToken>
. Labrum with anterior margin slightly emarginate, with six setae of which medial four setae equally distributed in medial half, distance from outermost medial seta to lateral seta about twice distance between medial setae. Antenna overall length moderately long, antennomere 11 just reaching beyond pronotal base. Thorax. Pronotum transverse, lateral margin sinuate near base, medial and basal setae touching lateral marginal bead; basal impressions obsolete; anterior angles scarcely produced; microsculpture not visible at 50
<normalizedToken originalValue="×">x</normalizedToken>
magnification; entire surface covered with minute punctulae. Elytral striae extremely shallow, scarcely impressed or absent; base of elytra not margined; humeri prominent but rounded; parascutellar punctures present at base of striae 1; interval 1 with two punctures at apex; interval 3 with three (right) or six (left) punctures; interval 5 with three (right) or four (left) punctures; interval 7 with one puncture near apex; interval 9
<pageBreakToken pageId="3" pageNumber="30" start="start">with</pageBreakToken>
20 evenly spaced punctures; elytral microsculpture visible at 50
<normalizedToken originalValue="×">x</normalizedToken>
as irregular isodiametric mesh of microlines; entire surface with micropunctulae. Male protaromeres 1-3 expanded with squamose setae ventrally. Tarsomere 5 on all legs ventral setose. Metatrochanter without setae. Metacoxa with single lateral seta. Abdomen. Abdominal ventrites smooth, glabrous except for very shallowly impressed transverse sulcus on last ventrite. Male with two setae on last ventrite. Aedeagus (Fig. 4) with small, dissimilar conchoid parameres; right paramere with large dorsal process (Fig. 4D); left paramere with a minute transverse apophysis (Fig. 4C). Median lobe with apex truncate; ostium dorsal; endophallus with small median sclerite.
</paragraph>
<caption pageId="3" pageNumber="30">
<paragraph pageId="3" pageNumber="30">
Figure 3. Dorsal habitus of Holotype of
<taxonomicName class="Insecta" family="Carabidae" genus="Lesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lesticus finisterrae" order="Coleoptera" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="finisterrae">Lesticus finisterrae</taxonomicName>
.
</paragraph>
</caption>
<caption pageId="3" pageNumber="30">
<paragraph pageId="3" pageNumber="30">
Figure 4. Male genitalia of holotype of
<taxonomicName class="Insecta" family="Carabidae" genus="Lesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lesticus finisterrae" order="Coleoptera" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="finisterrae">Lesticus finisterrae</taxonomicName>
. Median lobe. A left lateral B right lateral E ventral and F dorsal views. Parameres. C left D right. ta transverse apophysis.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="3" pageNumber="30" type="etymology">
<paragraph pageId="3" pageNumber="30">Etymology.</paragraph>
<paragraph pageId="3" pageNumber="30">The specific epithet finisterrae refers to the type locality, the Finisterre Range. Additionally, Finisterre is a contraction that derives from the Latin Finis Terrae, meaning &quot;End of the Earth&quot;, an appropriate metaphor for this remote and entomologically little-known region. Although terrae is the genitive form, the contraction is treated as a noun in apposition since the &quot;end of the Earth&quot; is used as the name for the location.</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="30" type="habitat">
<paragraph pageId="3" pageNumber="30">Habitat.</paragraph>
<paragraph pageId="3" pageNumber="30">
Only a single specimen of
<taxonomicName class="Insecta" family="Carabidae" genus="Lesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lesticus finisterrae" order="Coleoptera" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="finisterrae">Lesticus finisterrae</taxonomicName>
is known. It was collected in an unbaited pitfall trap placed in upper montane moss forest at an elevation of 3050 meters. The area was near the upper limit of forest growth and adjacent to open grassland (Fig. 2). Under the forest canopy, a fairly dense understory of low vegetation was present (Fig. 1).
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="33" pageId="3" pageNumber="30" type="identification and systematics">
<paragraph pageId="3" pageNumber="30">Identification and systematics.</paragraph>
<paragraph lastPageId="4" lastPageNumber="31" pageId="3" pageNumber="30">
<normalizedToken originalValue="“Trigonotomi”">&quot;Trigonotomi&quot;</normalizedToken>
sensu
<bibRefCitation author="Dubault, G" journalOrPublisher="Bulletin de la Societe entomolologique de France" pageId="6" pageNumber="33" pagination="239 - 248" title="Les generes des &quot; trigonotomi &quot;: Pareuyaptus n. gen. et revision des Euryaptus Bates, 1892 (Coleoptera, Pterostichidae)." volume="113" year="2008">Dubault et al. (2008)</bibRefCitation>
includes
<taxonomicName class="Insecta" family="Carabidae" genus="Lesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lesticus" order="Coleoptera" pageId="3" pageNumber="30" phylum="Arthropoda" rank="genus">Lesticus</taxonomicName>
,
<taxonomicName class="Insecta" family="Carabidae" genus="Trigonotoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trigonotoma" order="Coleoptera" pageId="3" pageNumber="30" phylum="Arthropoda" rank="genus">Trigonotoma</taxonomicName>
,
<taxonomicName class="Insecta" family="Carabidae" genus="Euryaptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Euryaptus" order="Coleoptera" pageId="3" pageNumber="30" phylum="Arthropoda" rank="genus">Euryaptus</taxonomicName>
and
<taxonomicName class="Insecta" family="Carabidae" genus="Pareuryaptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pareuryaptus" order="Coleoptera" pageId="3" pageNumber="30" phylum="Arthropoda" rank="genus">Pareuryaptus</taxonomicName>
. Although the group has not been rigorously tested for monophyly, it is reasonable to maintain its use as a working hypothesis based on similarity. Additionally, the genus
<taxonomicName class="Insecta" family="Carabidae" genus="Nesites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nesites" order="Coleoptera" pageId="3" pageNumber="30" phylum="Arthropoda" rank="genus">Nesites</taxonomicName>
Andrewes (cotype examined), which has the same combination of characters as
<taxonomicName class="Insecta" family="Carabidae" genus="Trigonotoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trigonotoma" order="Coleoptera" pageId="3" pageNumber="30" phylum="Arthropoda" rank="genus">Trigonotoma</taxonomicName>
, should be included. It is likely that
<taxonomicName class="Insecta" family="Carabidae" genus="Trigonotoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trigonotoma" order="Coleoptera" pageId="3" pageNumber="30" phylum="Arthropoda" rank="genus">Trigonotoma</taxonomicName>
and
<taxonomicName class="Insecta" family="Carabidae" genus="Nesites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nesites" order="Coleoptera" pageId="3" pageNumber="30" phylum="Arthropoda" rank="genus">Nesites</taxonomicName>
will prove to be synonymous.
<taxonomicName class="Insecta" family="Carabidae" genus="Lesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lesticus finisterrae" order="Coleoptera" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="finisterrae">Lesticus finisterrae</taxonomicName>
<pageBreakToken pageId="4" pageNumber="31" start="start">has</pageBreakToken>
the following states for characters given by
<bibRefCitation author="Dubault, G" journalOrPublisher="Bulletin de la Societe entomolologique de France" pageId="6" pageNumber="33" pagination="239 - 248" title="Les generes des &quot; trigonotomi &quot;: Pareuyaptus n. gen. et revision des Euryaptus Bates, 1892 (Coleoptera, Pterostichidae)." volume="113" year="2008">Dubault et al. (2008)</bibRefCitation>
: antennomere 1 less than combined length of antennomeres 2+3; setae at the anterior margin of the labrum more or less equally distributed; three or more setigerous punctures on interval 3 of the elytral disc. Females of
<taxonomicName class="Insecta" family="Carabidae" genus="Lesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lesticus" order="Coleoptera" pageId="4" pageNumber="31" phylum="Arthropoda" rank="genus">Lesticus</taxonomicName>
have four setae (two pairs) on the last ventrite, but the female for
<taxonomicName class="Insecta" family="Carabidae" genus="Lesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lesticus finisterrae" order="Coleoptera" pageId="4" pageNumber="31" phylum="Arthropoda" rank="species" species="finisterrae">Lesticus finisterrae</taxonomicName>
is unknown. The parascutellar striae are generally present and long in
<taxonomicName class="Insecta" family="Carabidae" genus="Lesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lesticus" order="Coleoptera" pageId="4" pageNumber="31" phylum="Arthropoda" rank="genus">Lesticus</taxonomicName>
species; however in
<taxonomicName class="Insecta" family="Carabidae" genus="Lesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lesticus finisterrae" order="Coleoptera" pageId="4" pageNumber="31" phylum="Arthropoda" rank="species" species="finisterrae">Lesticus finisterrae</taxonomicName>
all striae are very shallow or not impressed. There is no indication of the parascutellar striae.
<taxonomicName class="Insecta" family="Carabidae" genus="Lesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lesticus finisterrae" order="Coleoptera" pageId="4" pageNumber="31" phylum="Arthropoda" rank="species" species="finisterrae">Lesticus finisterrae</taxonomicName>
deviates from other species of
<taxonomicName class="Insecta" family="Carabidae" genus="Lesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lesticus" order="Coleoptera" pageId="4" pageNumber="31" phylum="Arthropoda" rank="genus">Lesticus</taxonomicName>
by the extremely prominent eyes, depressed form and lack of sulci on the abdominal ventrites. Based on general similarity and adjacencies of ranges, it seems probable that
<taxonomicName class="Insecta" family="Carabidae" genus="Lesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lesticus finisterrae" order="Coleoptera" pageId="4" pageNumber="31" phylum="Arthropoda" rank="species" species="finisterrae">Lesticus finisterrae</taxonomicName>
is an isolated and derived member of the
<taxonomicName class="Insecta" family="Carabidae" genus="Lesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lesticus chloronotus" order="Coleoptera" pageId="4" pageNumber="31" phylum="Arthropoda" rank="species" species="chloronotus">Lesticus chloronotus</taxonomicName>
group.
</paragraph>
<subSection lastPageId="5" lastPageNumber="32" pageId="4" pageNumber="31" type="key to pterostichine-like harpalinae">
<paragraph pageId="4" pageNumber="31">
Key to pterostichine-like
<taxonomicName lsidName="" pageId="4" pageNumber="31" rank="subFamily" subFamily="Harpalinae">Harpalinae</taxonomicName>
</paragraph>
<paragraph pageId="4" pageNumber="31">
Below is a revised key to the genera of
<normalizedToken originalValue="“pterostichites”">&quot;pterostichites&quot;</normalizedToken>
of New Guinea based on the key by
<bibRefCitation author="Darlington, PJ Jr" journalOrPublisher="Bulletin of the Museum of Comparative Zoology" pageId="6" pageNumber="33" pagination="319 - 564" title="The carabid beetles of New Guinea. Part I. Cicindelinae, Carabinae, Harpalinae through Pterostichini." volume="126" year="1962">Darlington (1962</bibRefCitation>
: 499-500). We have excluded
<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax" order="Coleoptera" pageId="4" pageNumber="31" phylum="Arthropoda" rank="genus">Mecyclothorax</taxonomicName>
Sharp (
<taxonomicName lsidName="" pageId="4" pageNumber="31" rank="tribe" tribe="Moriomorphini">Moriomorphini</taxonomicName>
,
<bibRefCitation author="Liebherr, JK" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="6" pageNumber="33" pagination="15 - 76" title="The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of West Maui, Hawaii: taxonomy, biogeography, and conservation." url="10.1002/mmnd.201100005" volume="58" year="2011">Liebherr 2011</bibRefCitation>
), which was included in
<normalizedToken originalValue="Darlingtons">Darlington's</normalizedToken>
key, as it is not a member of the
<taxonomicName lsidName="" pageId="4" pageNumber="31" rank="subFamily" subFamily="Harpalinae">Harpalinae</taxonomicName>
and only distantly related. It is easily distinguished from all the included taxa by the presence of a fixed seta in the mandibular scrobe.
</paragraph>
<paragraph lastPageId="5" lastPageNumber="32" pageId="4" pageNumber="31">
<table lastPageId="5" lastPageNumber="32" pageId="4" pageNumber="31">
<tr pageId="4" pageNumber="31">
<td colspan="1" pageId="4" pageNumber="31" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Morion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Morion" order="Coleoptera" pageId="4" pageNumber="31" phylum="Arthropoda" rank="genus">Morion</taxonomicName>
<taxonomicName lsidName="" pageId="4" pageNumber="31" rank="tribe" tribe="Morionini">Morionini</taxonomicName>
</td>
</tr>
<tr pageId="4" pageNumber="31">
<td colspan="1" pageId="4" pageNumber="31" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Brachidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Brachidius" order="Coleoptera" pageId="4" pageNumber="31" phylum="Arthropoda" rank="genus">Brachidius</taxonomicName>
<taxonomicName lsidName="" pageId="4" pageNumber="31" rank="tribe" tribe="Cratocerini">Cratocerini</taxonomicName>
</td>
</tr>
<tr pageId="4" pageNumber="31">
<td colspan="1" pageId="4" pageNumber="31" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Caelostomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caelostomus" order="Coleoptera" pageId="4" pageNumber="31" phylum="Arthropoda" rank="genus">Caelostomus</taxonomicName>
<taxonomicName lsidName="" pageId="4" pageNumber="31" rank="tribe" tribe="Drimostomatini">Drimostomatini</taxonomicName>
</td>
</tr>
<tr pageId="4" pageNumber="31">
<td colspan="1" pageId="4" pageNumber="31" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Stegazopteryx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stegazopteryx" order="Coleoptera" pageId="4" pageNumber="31" phylum="Arthropoda" rank="genus">Stegazopteryx</taxonomicName>
<taxonomicName lsidName="" pageId="4" pageNumber="31" rank="tribe" tribe="Drimostomatini">Drimostomatini</taxonomicName>
</td>
</tr>
<tr pageId="4" pageNumber="31">
<td colspan="1" pageId="4" pageNumber="31" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Abacetus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Abacetus" order="Coleoptera" pageId="4" pageNumber="31" phylum="Arthropoda" rank="genus">Abacetus</taxonomicName>
<taxonomicName lsidName="" pageId="4" pageNumber="31" rank="tribe" tribe="Abacetini">Abacetini</taxonomicName>
</td>
</tr>
<tr pageId="4" pageNumber="31">
<td colspan="1" pageId="4" pageNumber="31" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Catadromus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Catadromus" order="Coleoptera" pageId="4" pageNumber="31" phylum="Arthropoda" rank="genus">Catadromus</taxonomicName>
<taxonomicName lsidName="" pageId="4" pageNumber="31" rank="tribe" tribe="Pterostichini">Pterostichini</taxonomicName>
</td>
</tr>
<tr pageId="5" pageNumber="32">
<td colspan="1" pageId="5" pageNumber="32" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Prosopogmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prosopogmus" order="Coleoptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="genus">
<pageBreakToken pageId="5" pageNumber="32" start="start">Prosopogmus</pageBreakToken>
</taxonomicName>
<taxonomicName lsidName="" pageId="5" pageNumber="32" rank="tribe" tribe="Pterostichini">Pterostichini</taxonomicName>
</td>
</tr>
<tr pageId="5" pageNumber="32">
<td colspan="1" pageId="5" pageNumber="32" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Lesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lesticus" order="Coleoptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="genus">Lesticus</taxonomicName>
<taxonomicName lsidName="" pageId="5" pageNumber="32" rank="tribe" tribe="Pterostichini">Pterostichini</taxonomicName>
</td>
</tr>
<tr pageId="5" pageNumber="32">
<td colspan="1" pageId="5" pageNumber="32" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Rhytiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhytiferonia" order="Coleoptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="genus">Rhytiferonia</taxonomicName>
<taxonomicName lsidName="" pageId="5" pageNumber="32" rank="tribe" tribe="Pterostichini">Pterostichini</taxonomicName>
</td>
</tr>
<tr pageId="5" pageNumber="32">
<td colspan="1" pageId="5" pageNumber="32" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Rhytisternus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhytisternus laevis" order="Coleoptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="species" species="laevis">Rhytisternus laevis</taxonomicName>
<taxonomicName lsidName="" pageId="5" pageNumber="32" rank="tribe" tribe="Pterostichini">Pterostichini</taxonomicName>
</td>
</tr>
<tr pageId="5" pageNumber="32">
<td colspan="1" pageId="5" pageNumber="32" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Brachidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Brachidius" order="Coleoptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="genus">Brachidius</taxonomicName>
</td>
<td colspan="1" pageId="5" pageNumber="32" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Cosmodiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cosmodiscus" order="Coleoptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="genus">Cosmodiscus</taxonomicName>
<taxonomicName lsidName="" pageId="5" pageNumber="32" rank="tribe" tribe="Abacetini">Abacetini</taxonomicName>
</td>
</tr>
<tr pageId="5" pageNumber="32">
<td colspan="1" pageId="5" pageNumber="32" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Homalonesiota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Homalonesiota" order="Coleoptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="genus">Homalonesiota</taxonomicName>
<taxonomicName lsidName="" pageId="5" pageNumber="32" rank="tribe" tribe="Loxandrini">Loxandrini</taxonomicName>
</td>
</tr>
<tr pageId="5" pageNumber="32">
<td colspan="1" pageId="5" pageNumber="32" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Haploferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haploferonia" order="Coleoptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="genus">Haploferonia</taxonomicName>
<taxonomicName lsidName="" pageId="5" pageNumber="32" rank="tribe" tribe="Loxandrini">Loxandrini</taxonomicName>
</td>
</tr>
<tr pageId="5" pageNumber="32">
<td colspan="1" pageId="5" pageNumber="32" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Lesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lesticus" order="Coleoptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="genus">Lesticus</taxonomicName>
<taxonomicName lsidName="" pageId="5" pageNumber="32" rank="tribe" tribe="Pterostichini">Pterostichini</taxonomicName>
</td>
</tr>
<tr pageId="5" pageNumber="32">
<td colspan="1" pageId="5" pageNumber="32" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Loxandrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Loxandrus" order="Coleoptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="genus">Loxandrus</taxonomicName>
<taxonomicName lsidName="" pageId="5" pageNumber="32" rank="tribe" tribe="Loxandrini">Loxandrini</taxonomicName>
</td>
</tr>
<tr pageId="5" pageNumber="32">
<td colspan="1" pageId="5" pageNumber="32" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Nebrioferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nebrioferonia" order="Coleoptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="genus">Nebrioferonia</taxonomicName>
<taxonomicName lsidName="" pageId="5" pageNumber="32" rank="tribe" tribe="Loxandrini">Loxandrini</taxonomicName>
</td>
</tr>
<tr pageId="5" pageNumber="32">
<td colspan="1" pageId="5" pageNumber="32" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Tiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tiferonia" order="Coleoptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="genus">Tiferonia</taxonomicName>
<taxonomicName lsidName="" pageId="5" pageNumber="32" rank="tribe" tribe="Abacetini">Abacetini</taxonomicName>
</td>
</tr>
<tr pageId="5" pageNumber="32">
<td colspan="1" pageId="5" pageNumber="32" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Platycaelus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platycaelus" order="Coleoptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="genus">Platycaelus</taxonomicName>
<taxonomicName lsidName="" pageId="5" pageNumber="32" rank="tribe" tribe="Pterostichini">Pterostichini</taxonomicName>
</td>
</tr>
<tr pageId="5" pageNumber="32">
<td colspan="1" pageId="5" pageNumber="32" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Analoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Analoma" order="Coleoptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="genus">Analoma</taxonomicName>
<taxonomicName lsidName="" pageId="5" pageNumber="32" rank="tribe" tribe="Pterostichini">Pterostichini</taxonomicName>
</td>
</tr>
</table>
</paragraph>
<caption pageId="5" pageNumber="32">
<paragraph pageId="5" pageNumber="32">
Figure 5. Male genitalia of
<taxonomicName class="Insecta" family="Carabidae" genus="Stegazopteryx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stegazopteryx ivimkaensis" order="Coleoptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="species" species="ivimkaensis">Stegazopteryx ivimkaensis</taxonomicName>
. Median lobe. A left lateral B right lateral C ventral and D dorsal views.
</paragraph>
</caption>
</subSection>
<subSection pageId="5" pageNumber="32" type="additional records">
<paragraph pageId="5" pageNumber="32">Additional records</paragraph>
<paragraph pageId="5" pageNumber="32">
<taxonomicName class="Insecta" family="Carabidae" genus="Stegazopteryx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stegazopteryx ivimkaensis" order="Coleoptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="species" species="ivimkaensis">Stegazopteryx ivimkaensis</taxonomicName>
Will
</paragraph>
<paragraph pageId="5" pageNumber="32">
A male specimen labeled &quot;INDONESIA W-PAPUA 130km SE Kalmana, Omba (=Yamor) river 10-20km from coast,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-4.096945">4°05'49&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="134.9025">134°54'09&quot;E</geoCoordinate>
, 10-20m, 09.-11.II.2011 leg. A. Skale (008).&quot; Specimen deposited ZSM.
</paragraph>
<paragraph pageId="5" pageNumber="32">This specimen record extends the range of this species into eastern West-Papua. This specimen, the two female syntype specimens and the further records below, are all from elevations of 120m and below. This species is now known to range across nearly all of New Guinea. The examined male specimen is consistent with the description: of the genus based on females as given by Will (2004), with the following additions. sbl=5.4mm, greatest width of elytra 2.3mm. Secondary sexual characters- male protarsi not expanded but with spatulate setae ventrally on tarsomeres 1-3. Last ventrite with one pair of setae. Aedeagus right side up in repose. Right paramere larger and conchoid, left paramere small, peg-like. Median lobe of adeagus simple, no evident sclerotized structure on the endophallus (Fig. 5).</paragraph>
<paragraph pageId="5" pageNumber="32">Additional locality records (not examined, Martin Baehr, in litt.). PAPUA NEW GUINEA Canopy Mission, Madang Province, Baiteta, Light, Leg. Olivier Missa: 1 male- AR 53, 30-V-1996; 1 male, 1 female- T 2, 31-V-1993; 1 female, X G, 24-IV-1996; 1 female, M 2, 30-IV-1996. One male and two females deposited IRSNB, one male and one female ZSM.</paragraph>
</subSection>
<subSection pageId="6" pageNumber="33" type="rhytisternus laevis (macleay)">
<paragraph pageId="6" pageNumber="33">
<taxonomicName class="Insecta" family="Carabidae" genus="Rhytisternus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhytisternus laevis" order="Coleoptera" pageId="6" pageNumber="33" phylum="Arthropoda" rank="species" species="laevis">
<pageBreakToken pageId="6" pageNumber="33" start="start">Rhytisternus</pageBreakToken>
laevis
</taxonomicName>
(Macleay)
</paragraph>
<paragraph pageId="6" pageNumber="33">A female specimen labeled &quot;New Guinea, Weam, Aug 1962// H. Olmus collector&quot;. Deposited ANIC. This species is distributed across northern Australia and it has not been previously reported from New Guinea. It is likely a recent accidental introduction or dispersal. We have only seen this single individual from New Guinea and it is unknown if the species is established.</paragraph>
</subSection>
</subSubSection>
</treatment>
</document>