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<document id="00749502023994109DA95DDF1378B73A" ID-CLB-Dataset="53922" ID-DOI="10.11646/zootaxa.5149.1.1" ID-GBIF-Dataset="caecca9d-72a3-4d9a-836d-9ac3e0ded512" ID-ISSN="1175-5326" ID-Zenodo-Dep="6605961" ID-ZooBank="BF98000F-04F9-4D22-9D93-4C3A2418D7EF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1654152510384" checkinUser="plazi" docAuthor="Probst, Rodolfo Da Silva &amp; Brandão, Carlos Roberto Ferreira" docDate="2022" docId="F62B8F24FFA6E37FFF0FFAAB85D1DA1F" docLanguage="en" docName="zootaxa.5149.1.1.pdf" docOrigin="Zootaxa 5149 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.16:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="16" docTitle="Basiceros manni Brown &amp; Kempf 1960" docType="treatment" docVersion="8" lastPageNumber="42" masterDocId="0A12F75CFF84E355FF98FF9B811BDB5E" masterDocTitle="A taxonomic revision of the dirt ants, Basiceros Schulz, 1906 (Hymenoptera, Formicidae)" masterLastPageNumber="75" masterPageNumber="1" pageNumber="35" updateTime="1702002095603" updateUser="plazi">
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<mods:title id="55FD10863A0E7B3A75C360F9B7D1CBD9">A taxonomic revision of the dirt ants, Basiceros Schulz, 1906 (Hymenoptera, Formicidae)</mods:title>
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<mods:namePart id="3419DCD9089E88EC27AC71C26980EAD1">Probst, Rodolfo Da Silva</mods:namePart>
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<mods:namePart id="7455E99B86D76B312FF08419C8AD41D1">Brandão, Carlos Roberto Ferreira</mods:namePart>
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<treatment id="F62B8F24FFA6E37FFF0FFAAB85D1DA1F" ID-DOI="http://doi.org/10.5281/zenodo.7636265" ID-GBIF-Taxon="209289996" ID-Zenodo-Dep="7636265" LSID="urn:lsid:plazi:treatment:F62B8F24FFA6E37FFF0FFAAB85D1DA1F" httpUri="http://treatment.plazi.org/id/F62B8F24FFA6E37FFF0FFAAB85D1DA1F" lastPageId="42" lastPageNumber="42" pageId="34" pageNumber="35">
<subSubSection id="36986DB9FFA6E377FF0FFAAB8366DE14" box="[151,637,1328,1354]" pageId="34" pageNumber="35" type="nomenclature">
<paragraph id="7E3D3E32FFA6E377FF0FFAAB8366DE14" blockId="34.[151,637,1328,1391]" box="[151,637,1328,1354]" pageId="34" pageNumber="35">
<heading id="2575895EFFA6E377FF0FFAAB8366DE14" bold="true" box="[151,637,1328,1354]" fontSize="11" level="2" pageId="34" pageNumber="35" reason="4">
<taxonomicName id="B98245B1FFA6E377FF0FFAAB8366DE14" ID-CoL="5WCVT" ID-ENA="411698" authority="Brown &amp; Kempf 1960" authorityName="Brown &amp; Kempf" authorityYear="1960" box="[151,637,1328,1354]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="manni">
<emphasis id="4CF6E220FFA6E377FF0FFAAB8366DE14" bold="true" box="[151,637,1328,1354]" pageId="34" pageNumber="35">
<emphasis id="4CF6E220FFA6E377FF0FFAAB8079DE14" bold="true" box="[151,354,1328,1354]" italics="true" pageId="34" pageNumber="35">Basiceros manni</emphasis>
<bibRefCitation id="1A1343C3FFA6E377FEF1FAAB8366DE14" author="Brown, W. L. Jr. &amp; Kempf, W. W." box="[361,637,1328,1354]" pageId="34" pageNumber="35" pagination="161 - 250" refId="ref61211" refString="Brown, W. L. Jr. &amp; Kempf, W. W. (1960) A world revision of the ant tribe Basicerotini (Hym. Formicidae). Studia Entomologica, New Series, 3, 161 - 250." type="journal article" year="1960">Brown &amp; Kempf 1960</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="36986DB9FFA6E377FF0FFACF804ADE30" box="[151,337,1364,1391]" pageId="34" pageNumber="35" type="description">
<paragraph id="7E3D3E32FFA6E377FF0FFACF804ADE30" blockId="34.[151,637,1328,1391]" box="[151,337,1364,1391]" pageId="34" pageNumber="35">
(
<figureCitation id="E6B922B7FFA6E377FF07FACF803BDE31" box="[159,288,1364,1391]" captionStart-0="FIGURE 13" captionStart-1="FIGURE 14" captionStart-2="FIGURE 15" captionStartId-0="36.[151,250,1616,1640]" captionStartId-1="37.[151,250,1692,1716]" captionStartId-2="39.[151,250,1854,1878]" captionTargetBox-0="[190,1397,921,1591]" captionTargetBox-1="[174,1414,849,1669]" captionTargetBox-2="[253,1332,489,1831]" captionTargetId-0="figure-328@36.[190,1397,921,1592]" captionTargetId-1="figure-301@37.[174,1414,849,1669]" captionTargetId-2="figure-167@39.[253,1332,489,1831]" captionTargetPageId-0="36" captionTargetPageId-1="37" captionTargetPageId-2="39" captionText-0="FIGURE 13. Basiceros manni, worker (CASENT0914888, Costa Rica: Heredia); A. full-face view, B. lateral view, C. dorsal view. Images by Michele Esposito (CAS)." captionText-1="FIGURE 14. Basiceros manni, gyne (MCZ 546486, Costa Rica: Heredia) and male (MCZ, Costa Rica: Heredia); A and D. fullface view, B and E. dorsal view, C and F. lateral view. Scale bars: A, D = 0.5 mm, B, C, E, F =1 mm." captionText-2="FIGURE 15. Basiceros manni, larva (MZSP, Nicaragua: Boaco); A. head in anteroventral view, B. dorsal pilosity, C. lateral view. Scale bars: A = 0.1 mm, C = 1 mm, B = 0.05 mm." figureDoi-0="http://doi.org/10.5281/zenodo.6605993" figureDoi-1="http://doi.org/10.5281/zenodo.6605995" figureDoi-2="http://doi.org/10.5281/zenodo.6605997" httpUri-0="https://zenodo.org/record/6605993/files/figure.png" httpUri-1="https://zenodo.org/record/6605995/files/figure.png" httpUri-2="https://zenodo.org/record/6605997/files/figure.png" pageId="34" pageNumber="35">Figs 1315</figureCitation>
,
<figureCitation id="E6B922B7FFA6E377FEB4FACF8052DE30" box="[300,329,1364,1390]" captionStart="FIGURE 30" captionStartId="70.[151,250,1588,1612]" captionTargetBox="[151,1437,453,1564]" captionTargetId="figure-102@70.[151,1437,453,1564]" captionTargetPageId="70" captionText="FIGURE 30. Distribution map of Basiceros species from the singularis clade." figureDoi="http://doi.org/10.5281/zenodo.6606035" httpUri="https://zenodo.org/record/6606035/files/figure.png" pageId="34" pageNumber="35">30</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="36986DB9FFA6E377FF0FFA0F8590DEB7" pageId="34" pageNumber="35" type="reference_group">
<paragraph id="7E3D3E32FFA6E377FF0FFA0F82F5DEF5" blockId="34.[151,1437,1427,1543]" box="[151,1006,1427,1451]" pageId="34" pageNumber="35">
<treatmentCitationGroup id="5E92191CFFA6E377FF0FFA0F82F5DEF5" box="[151,1006,1427,1451]" pageId="34" pageNumber="35">
<treatmentCitation id="FF231823FFA6E377FF0FFA0F837BDEF5" author="Brown, W. L. Jr. &amp; Kempf, W. W." box="[151,608,1427,1451]" page="177" pageId="34" pageNumber="35" year="1960">
<taxonomicName id="B98245B1FFA6E377FF0FFA0F837BDEF5" ID-CoL="5WCVT" authority="Brown &amp; Kempf 1960: 177" authorityName="Brown &amp; Kempf" authorityPageNumber="177" authorityYear="1960" box="[151,608,1427,1451]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="manni">
<emphasis id="4CF6E220FFA6E377FF0FFA0F8024DEF5" box="[151,319,1428,1451]" italics="true" pageId="34" pageNumber="35">Basiceros manni</emphasis>
<bibRefCitation id="1A1343C3FFA6E377FEDEFA0F837BDEF5" author="Brown, W. L. Jr. &amp; Kempf, W. W." box="[326,608,1427,1451]" pageId="34" pageNumber="35" pagination="161 - 250" refId="ref61211" refString="Brown, W. L. Jr. &amp; Kempf, W. W. (1960) A world revision of the ant tribe Basicerotini (Hym. Formicidae). Studia Entomologica, New Series, 3, 161 - 250." type="journal article" year="1960">Brown &amp; Kempf 1960: 177</bibRefCitation>
</taxonomicName>
</treatmentCitation>
,
<figureCitation id="E6B922B7FFA6E377FDF3FA0883B0DEF5" box="[619,683,1427,1451]" captionStart="FIGURE 3" captionStartId="14.[151,250,1673,1697]" captionTargetBox="[156,1432,921,1649]" captionTargetId="figure-348@14.[156,1432,921,1649]" captionTargetPageId="14" captionText="FIGURE 3. Basiceros browni sp. nov., holotype worker (MCZ, Ecuador: Los Ríos); A. full-face view, B. lateral view, and C. dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6605969" httpUri="https://zenodo.org/record/6605969/files/figure.png" pageId="34" pageNumber="35">Figs 3</figureCitation>
,
<figureCitation id="E6B922B7FFA6E377FD2EFA0883D5DEF5" box="[694,718,1427,1451]" captionStart="FIGURE 11" captionStartId="29.[151,250,1643,1667]" captionTargetBox="[154,1433,885,1620]" captionTargetId="figure-311@29.[154,1433,885,1620]" captionTargetPageId="29" captionText="FIGURE 11. Basiceros disciger, worker (MZSP, Brazil: São Paulo); A. full-face view, B. lateral view. C. dorsal view. Scale bars: A = 0.5 mm, BC =1 mm." figureDoi="http://doi.org/10.5281/zenodo.6605989" httpUri="https://zenodo.org/record/6605989/files/figure.png" pageId="34" pageNumber="35">11</figureCitation>
(worker, gyne):
<collectingCountry id="06957EA2FFA6E377FCE0FA0882F1DEF5" box="[888,1002,1427,1451]" name="Costa Rica" pageId="34" pageNumber="35">Costa Rica</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="7E3D3E32FFA6E377FF0FFA298590DEB7" blockId="34.[151,1437,1427,1543]" pageId="34" pageNumber="35">
<bibRefCitation id="1A1343C3FFA6E377FF0FFA2980D5DE94" author="Wheeler, G. C. &amp; Wheeler, J." box="[151,462,1458,1482]" pageId="34" pageNumber="35" pagination="111 - 145" refId="ref63133" refString="Wheeler, G. C. &amp; Wheeler, J. (1955) [&quot; 1954 &quot;]) The ant larvae of the myrmicine tribes Basicerotini and Dacetini. Psyche, Cambridge, 61, 111 - 145. https: // doi. org / 10.1155 / 1954 / 49606" type="journal article" year="1955">Wheeler &amp; Wheeler 1955: 112</bibRefCitation>
, Plate 10,
<figureCitation id="E6B922B7FFA6E377FDADFA2983BDDE94" box="[565,678,1458,1482]" captionStart-0="FIGURE 16" captionStart-1="FIGURE 17" captionStartId-0="41.[151,250,1112,1136]" captionStartId-1="43.[151,250,1542,1566]" captionTargetBox-0="[158,1430,525,1089]" captionTargetBox-1="[148,1439,741,1519]" captionTargetId-0="figure-182@41.[158,1430,525,1089]" captionTargetId-1="figure-261@43.[148,1439,741,1519]" captionTargetPageId-0="41" captionTargetPageId-1="43" captionText-0="FIGURE 16. Basiceros manni colony in abandoned Meliponini nest at La Selva Biological Station, Costa Rica: Heredia; A. Exterior view of nest entrance, B. Detail of chamber with immatures below nest entrance, C. Brood chamber. Images: courtesy of Ronald Garcia." captionText-1="FIGURE 17. Basiceros militaris, worker (CASENT0280778, Ecuador: Tiputini); A. full-face view, B. lateral view, C. dorsal view. Image by Estella Ortega (CAS), edited." figureDoi-0="http://doi.org/10.5281/zenodo.6605999" figureDoi-1="http://doi.org/10.5281/zenodo.6606003" httpUri-0="https://zenodo.org/record/6605999/files/figure.png" httpUri-1="https://zenodo.org/record/6606003/files/figure.png" pageId="34" pageNumber="35">Figs 1617</figureCitation>
(immature of
<taxonomicName id="B98245B1FFA6E377FCAEFA2982C5DE97" ID-CoL="5WCVT" authorityName="Brown &amp; Kempf" authorityYear="1960" box="[822,990,1458,1481]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="manni">
<emphasis id="4CF6E220FFA6E377FCAEFA2982C5DE97" box="[822,990,1458,1481]" italics="true" pageId="34" pageNumber="35">Basiceros manni</emphasis>
</taxonomicName>
identified as prepupa of
<taxonomicName id="B98245B1FFA6E377FB4FFA298446DE97" box="[1239,1373,1458,1481]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="4CF6E220FFA6E377FB4FFA298421DE97" box="[1239,1338,1458,1481]" italics="true" pageId="34" pageNumber="35">Basiceros</emphasis>
sp.
</taxonomicName>
Hamburg Farm,
<collectingCountry id="06957EA2FFA6E377FEA6FA4A80ABDEB6" box="[318,432,1488,1513]" name="Costa Rica" pageId="34" pageNumber="35">Costa Rica</collectingCountry>
; collected with the type-series of what would be described as
<taxonomicName id="B98245B1FFA6E377FBBAFA4A8566DEB6" ID-CoL="5WCVT" authorityName="Brown &amp; Kempf" authorityYear="1960" box="[1058,1149,1489,1512]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="manni">
<emphasis id="4CF6E220FFA6E377FBBAFA4A8566DEB6" box="[1058,1149,1489,1512]" italics="true" pageId="34" pageNumber="35">B. manni</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="36986DB9FFA6E377FF0FFA74833BDD59" box="[151,544,1519,1543]" pageId="34" pageNumber="35" type="description">
<paragraph id="7E3D3E32FFA6E377FF0FFA74833BDD59" blockId="34.[151,1437,1427,1543]" box="[151,544,1519,1543]" pageId="34" pageNumber="35">
<bibRefCitation id="1A1343C3FFA6E377FF0FFA7480D4DD59" author="Wheeler, G. C. &amp; Wheeler, J." box="[151,463,1519,1543]" pageId="34" pageNumber="35" pagination="601 - 610" refId="ref63295" refString="Wheeler, G. C. &amp; Wheeler, J. (1983) Supplementary studies on ant larvae: Myrmicinae. Transactions of the American Entomological Society, 108, 601 - 610." type="journal article" year="1983">Wheeler &amp; Wheeler 1983: 607</bibRefCitation>
(larva).
</paragraph>
</subSubSection>
<subSubSection id="36986DB9FFA6E377FF0FF9AB858BDC7D" pageId="34" pageNumber="35" type="materials_examined">
<paragraph id="7E3D3E32FFA6E377FF0FF9AB858BDC7D" blockId="34.[151,1437,1584,2043]" pageId="34" pageNumber="35">
<emphasis id="4CF6E220FFA6E377FF0FF9AB805CDD14" bold="true" box="[151,327,1584,1610]" pageId="34" pageNumber="35">Type material.</emphasis>
<materialsCitation id="CEEA346FFFA6E377FEC8F9AB8379DD31" ID-GBIF-Occurrence="4029332318" collectionCode="USNM" collectorName="F. Nevermann" country="Costa Rica" location="Limon" pageId="34" pageNumber="35" specimenCount="1" specimenCount-worker="1" stateProvince="Limon" typeStatus="holotype">
<collectingCountry id="06957EA2FFA6E377FEC8F9AB80ECDD14" box="[336,503,1584,1610]" name="Costa Rica" pageId="34" pageNumber="35">COSTA RICA</collectingCountry>
:
<collectingRegion id="BC46F0D0FFA6E377FD9CF9AB8354DD14" box="[516,591,1584,1610]" country="Costa Rica" name="Limon" pageId="34" pageNumber="35">Limón</collectingRegion>
(“
<collectorName id="D3775BE4FFA6E377FDF1F9AB83E3DD15" box="[617,760,1584,1611]" pageId="34" pageNumber="35">Santa Clara</collectorName>
”):
<collectorName id="D3775BE4FFA6E377FC8FF9AB82DCDD15" box="[791,967,1584,1611]" pageId="34" pageNumber="35">Hamburg Farm</collectorName>
, 1924,
<collectorName id="D3775BE4FFA6E377FB84F9AA85A7DD15" box="[1052,1212,1585,1611]" pageId="34" pageNumber="35">F. Nevermann</collectorName>
col., n. 65065 (
<specimenCount id="6884F5BBFFA6E377FAEAF9AB81F3DD31" pageId="34" pageNumber="35" type="worker">one worker</specimenCount>
<typeStatus id="A1398090FFA6E377FE9DF9CF807CDD31" box="[261,359,1620,1647]" pageId="34" pageNumber="35" type="holotype">holotype</typeStatus>
) [
<collectionCode id="1893A6F7FFA6E377FEE3F9C880C9DD30" box="[379,466,1619,1646]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="34" pageNumber="35" type="Museum">USNM</collectionCode>
] (examined)
</materialsCitation>
;
<materialsCitation id="CEEA346FFFA6E377FDF5F9CE8443DD31" ID-GBIF-Occurrence="4029332426" box="[621,1368,1619,1647]" collectionCode="USNM" country="Costa Rica" location="Limon" pageId="34" pageNumber="35" specimenCount="1" specimenCount-worker="1" stateProvince="Limon" typeStatus="holotype">
same data, n. 65065 (
<specimenCount id="6884F5BBFFA6E377FCC1F9CF82F1DD30" box="[857,1002,1620,1646]" pageId="34" pageNumber="35" type="worker">five workers</specimenCount>
, one gyne) [
<collectionCode id="1893A6F7FFA6E377FBE9F9C885D3DD30" box="[1137,1224,1619,1646]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="34" pageNumber="35" type="Museum">USNM</collectionCode>
] (examined)
</materialsCitation>
;
<materialsCitation id="CEEA346FFFA6E377FAFBF9CE8279DDCD" ID-GBIF-Occurrence="4029332429" collectionCode="MCZ" country="Costa Rica" location="Limon" pageId="34" pageNumber="35" specimenCount="1" specimenCount-worker="1" stateProvince="Limon" typeStatus="paratype">
same data, n. 30878 (
<specimenCount id="6884F5BBFFA6E377FED5F9E380F0DDCD" box="[333,491,1656,1683]" pageId="34" pageNumber="35" type="worker">three workers</specimenCount>
<typeStatus id="A1398090FFA6E377FD90F9E3836FDDCD" box="[520,628,1656,1683]" pageId="34" pageNumber="35" type="paratype">paratypes</typeStatus>
) [
<collectionCode id="1893A6F7FFA6E377FD14F9EC83D5DDCC" box="[652,718,1655,1682]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="34" pageNumber="35" type="Museum">MCZ</collectionCode>
] (examined)
</materialsCitation>
;
<materialsCitation id="CEEA346FFFA6E377FCF7F9E283E3DD85" ID-GBIF-Occurrence="4029332413" collectingDate="1920-03" collectionCode="MZSP" collectorName="W. M. Mann" country="Costa Rica" location="Colombiana Farm" pageId="34" pageNumber="35" specimenCount="2" specimenCount-worker="2" stateProvince="Limon" typeStatus="paratype">
same data (fours workers, one gyne—
<typeStatus id="A1398090FFA6E377FAB0F9E2848EDDCD" box="[1320,1429,1657,1683]" pageId="34" pageNumber="35" type="paratype">paratypes</typeStatus>
) [
<collectionCode id="1893A6F7FFA6E377FF06F90081F6DDE8" box="[158,237,1691,1718]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="34" pageNumber="35" type="Museum">MZSP</collectionCode>
] (examined); (
<specimenCount id="6884F5BBFFA6E377FE0CF907832ADDE9" box="[404,561,1692,1719]" pageId="34" pageNumber="35" type="worker">eight workers</specimenCount>
<typeStatus id="A1398090FFA6E377FDD6F90783A1DDE9" box="[590,698,1692,1719]" pageId="34" pageNumber="35" type="paratype">paratypes</typeStatus>
) (not examined, location unknown);
<location id="7B5D68E9FFA6E377FBFAF9078429DDE9" LSID="urn:lsid:plazi:treatment:F62B8F24FFA6E37FFF0FFAAB85D1DA1F:7B5D68E9FFA6E377FBFAF9078429DDE9" box="[1122,1330,1692,1719]" country="Costa Rica" name="Colombiana Farm" pageId="34" pageNumber="35">Colombiana Farm</location>
,
<date id="0A3C18F2FFA6E377FAA7F9078483DDE8" box="[1343,1432,1692,1718]" pageId="34" pageNumber="35" value="1920-03">
<collectingDate id="1A78E11AFFA6E377FAA7F9078483DDE8" box="[1343,1432,1692,1718]" pageId="34" pageNumber="35" value="1920-03">iii.1920</collectingDate>
</date>
,
<collectorName id="D3775BE4FFA6E377FF0FF95A803DDD85" box="[151,294,1729,1755]" pageId="34" pageNumber="35">W. M. Mann</collectorName>
col. (
<specimenCount id="6884F5BBFFA6E377FEFDF95B8319DD85" box="[357,514,1728,1755]" pageId="34" pageNumber="35" type="worker">three workers</specimenCount>
) [
<collectionCode id="1893A6F7FFA6E377FD80F924837CDD84" box="[536,615,1727,1754]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="34" pageNumber="35" type="Museum">MZSP</collectionCode>
] (examined)
</materialsCitation>
;
<materialsCitation id="CEEA346FFFA6E377FC9BF95A8058DDA1" ID-GBIF-Occurrence="4029332330" collectingDate="1920-03" collectionCode="MZSP" collectorName="W. M. Mann" country="Costa Rica" location="Colombiana Farm" pageId="34" pageNumber="35" specimenCount="1" specimenCount-worker="1" stateProvince="Limon" typeStatus="paratype">
same data (
<specimenCount id="6884F5BBFFA6E377FC19F95B8505DD85" box="[897,1054,1728,1755]" pageId="34" pageNumber="35" type="worker">three workers</specimenCount>
<typeStatus id="A1398090FFA6E377FBA2F95B85BDDD85" box="[1082,1190,1728,1755]" pageId="34" pageNumber="35" type="paratype">paratypes</typeStatus>
) (not examined, location unknown)
</materialsCitation>
.
<materialsCitation id="CEEA346FFFA6E377FEC8F97F806ADC7D" ID-GBIF-Occurrence="4029332302" collectingDate="1924-05" collectionCode="MCZ" collectorName="W. M. Mann" country="Honduras" location="HONDURAS" pageId="34" pageNumber="35" specimenCount="1" specimenCount-worker="1" typeStatus="paratype">
<collectingCountry id="06957EA2FFA6E377FEC8F97F80EBDDA0" box="[336,496,1764,1790]" name="Honduras" pageId="34" pageNumber="35">HONDURAS</collectingCountry>
:
<collectorName id="D3775BE4FFA6E377FE66F97F835CDDA0" box="[510,583,1764,1790]" pageId="34" pageNumber="35">Colón</collectorName>
:
<collectorName id="D3775BE4FFA6E377FDCCF97F83C8DDA0" box="[596,723,1764,1790]" pageId="34" pageNumber="35">Sangrelaya</collectorName>
,
<date id="0A3C18F2FFA6E377FD7AF97F822BDDA0" box="[738,816,1764,1790]" pageId="34" pageNumber="35" value="1924-05">
<collectingDate id="1A78E11AFFA6E377FD7AF97F822BDDA0" box="[738,816,1764,1790]" pageId="34" pageNumber="35" value="1924-05">v.1924</collectingDate>
</date>
,
<collectorName id="D3775BE4FFA6E377FCA6F97E82D3DDA1" box="[830,968,1765,1791]" pageId="34" pageNumber="35">W.M. Mann</collectorName>
col., n. 546483 (
<specimenCount id="6884F5BBFFA6E377FB0BF97F840CDDA1" box="[1171,1303,1764,1791]" pageId="34" pageNumber="35" type="worker">one worker</specimenCount>
<typeStatus id="A1398090FFA6E377FAACF97F848EDDA1" box="[1332,1429,1764,1791]" pageId="34" pageNumber="35" type="paratype">paratype</typeStatus>
) [
<collectionCode id="1893A6F7FFA6E377FF06F89C81FBDC7C" box="[158,224,1799,1826]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="34" pageNumber="35" type="Museum">MCZ</collectionCode>
] (examined)
</materialsCitation>
;
<materialsCitation id="CEEA346FFFA6E377FEE4F8928597DC7D" ID-GBIF-Occurrence="4029332402" box="[380,1164,1800,1827]" collectingDate="1924-05" collectionCode="MCZ" collectorName="W. M. Mann" country="Honduras" location="HONDURAS" pageId="34" pageNumber="35" specimenCount="1" specimenCount-worker="1" typeStatus="paratype">
same data (
<specimenCount id="6884F5BBFFA6E377FE62F8938360DC7D" box="[506,635,1800,1827]" pageId="34" pageNumber="35" type="worker">one worker</specimenCount>
<typeStatus id="A1398090FFA6E377FD00F89383E2DC7D" box="[664,761,1800,1827]" pageId="34" pageNumber="35" type="paratype">paratype</typeStatus>
) (not examined, location unknown)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="36986DB9FFA6E377FF5FF8B78527DCA4" pageId="34" pageNumber="35" type="diagnosis">
<paragraph id="7E3D3E32FFA6E377FF5FF8B78527DCA4" blockId="34.[151,1437,1584,2043]" pageId="34" pageNumber="35">
<emphasis id="4CF6E220FFA6E377FF5FF8B78059DC18" bold="true" box="[199,322,1836,1862]" pageId="34" pageNumber="35">Diagnosis.</emphasis>
Comparatively large size; color light to dark brown coloration. Specialized pilosity distributed in a double layer: a basal subdecumbent plumose, and a fine erect and clavate layers. Clypeomandibular space broad; dorsum of mandibles and clypeus with piligerous punctuations. Petiole claviform, elongated and with a depressed node. Anterior margin of first gastral sternite continuous, without longitudinal projection in the form of carina or prominence. Additionally, adult workers and gynes are usually densely covered with soil and litter particles, which adhere to the double layer of specialized hairs (see
<figureCitation id="E6B922B7FFA6E377FD4CF87B8201DCA4" box="[724,794,2016,2042]" captionStart="FIGURE 1" captionStartId="7.[151,250,1879,1903]" captionTargetBox="[149,1439,181,1854]" captionTargetId="figure-17@7.[149,1439,181,1854]" captionTargetPageId="7" captionText="FIGURE 1. Wing venation of Basiceros males. A. forewing and B. hindwing of B. conjugans (RSPPC, Peru). C. forewing of B. manni male (MCZ, Costa Rica: Heredia). Fused veins are indicated with a plus sign and crossveins by a dash. Cell names (in white) are indicated in italics. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6605965" httpUri="https://zenodo.org/record/6605965/files/figure.png" pageId="34" pageNumber="35">Fig1A</figureCitation>
from
<bibRefCitation id="1A1343C3FFA6E377FCC7F87B852BDCA4" author="Probst, R. S. &amp; Wray, B. D. &amp; Moreau, C. S. &amp; Brandao, C. R. F." box="[863,1072,2016,2043]" pageId="34" pageNumber="35" pagination="1 - 12" refId="ref62641" refString="Probst, R. S., Wray, B. D., Moreau, C. S. &amp; Brandao, C. R. F. (2019) A phylogenetic analysis of the dirt ants, Basiceros (Formicidae: Myrmicinae): inferring life histories through morphological convergence. Insect Systematics and Diversity, 3 (4), 1 - 12. https: // doi. org / 10.1093 / isd / ixz 013" type="journal article" year="2019">
Probst
<emphasis id="4CF6E220FFA6E377FC35F87A82C5DCA4" box="[941,990,2016,2042]" italics="true" pageId="34" pageNumber="35">et al</emphasis>
., 2019
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="36986DB9FFA7E372FF5FFF0D845FDA8F" lastPageId="39" lastPageNumber="40" pageId="35" pageNumber="36" type="description">
<paragraph id="7E3D3E32FFA7E376FF5FFF0D8082DA3B" blockId="35.[151,1437,150,2013]" pageId="35" pageNumber="36">
<emphasis id="4CF6E220FFA7E376FF5FFF0D8042DBEE" bold="true" box="[199,345,150,176]" pageId="35" pageNumber="36">Description.</emphasis>
<emphasis id="4CF6E220FFA7E376FEF9FF0D80A9DBEE" box="[353,434,150,176]" italics="true" pageId="35" pageNumber="36">Worker</emphasis>
(n=4). HL 1.501.53, HL2 1.561.62, HW1 1.311.39, MdL 0.971.00, SL1 1.181.22, SL2 1.191.20, PDL 0.140.15, A3L 0.040.06, AFL 0.470.50, FuL 1.301.34, EL 0.220.25, EW 0.190.22, ML 2.282.40, MfL 1.872.03, MtL 1.511.62,
<collectingCountry id="06957EA2FFA7E376FD2BFF4483CCDBA7" box="[691,727,223,249]" name="Philippines" pageId="35" pageNumber="36">PH</collectingCountry>
0.370.40, PL 1.031.12, PW 0.360.39, PPL 0.590.65, PPW 0.560.65, GL 2.00
<emphasis id="4CF6E220FFA7E376FE1FFE9980A3DA42" bold="true" box="[391,440,258,284]" pageId="35" pageNumber="36">2.09</emphasis>
, GW 1.311.40, TL 8.438.68, CI 8892, CS 1.421.44, MCI 6466, SI 8590, ESI 1820, SAI2 237253, EI1 0.280.33, MFI 6570, PTI 264283. (In bold, measurement for worker presenting slightly dilated gaster).
</paragraph>
<paragraph id="7E3D3E32FFA7E376FF5FFEF5832CDE9F" blockId="35.[151,1437,150,2013]" pageId="35" pageNumber="36">
Size large compared to other
<taxonomicName id="B98245B1FFA7E376FD92FEF483CEDAD6" box="[522,725,366,392]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="4CF6E220FFA7E376FD92FEF4836CDAD6" box="[522,631,367,392]" italics="true" pageId="35" pageNumber="36">Basiceros</emphasis>
species.
</taxonomicName>
Color (when integument is visible and not densely covered with particles) brown to reddish-brown; appendices slightly lighter. Mandibles and appendages slightly reddish-brown; mandibular dorsa covered with minute piligerous punctuations; apex with short yellowish setae; interdental setae present, yellowish and brush-shaped, slightly longer than teeth. Basimandibular setae present, narrow and erect, located at the anteroventral limit of mandibular peduncle. Sparse squamiform hairs on the anterodorsal margin of the mandibular peduncle, close to the basimandibular setae. Stipes covered by fine, medium, and subdecumbent pilosity; each dorsum with a suberect hair close to its anterior margin. Labrum densely covered with piligerous punctuations bearing short and decumbent setae. Median frontoclypeal portion covered by coarse piligerous punctuations; lateral limits of clypeus with decumbent, short and squamiform yellowish hairs. Head dorsum predominantly covered by piligerous fovea bearing either squamiform or plumose-like hairs, subdecumbent; sparse long erect to suberect hairs on the anterior portion of head, more abundant near the posterolateral region of head and vertexal margin. Ventral surface of head covered by piligerous punctuations; long, erect hairs near the sides of head and squamiform hairs marginating the ventral limit of antennal scrobe. Pilosity on meso- and metasoma present as a double layer of specialized hairs. The “basal” layer is composed of yellowish and subdecumbent plumose-like hairs, present on meso- and metasoma in the following conformation: dense, surrounding the anterior margin of pronotum; slightly sparse on the dorsum of pronotum and mesonotum, dorsum and lateral of propodeum, mesopleuron near the epicnemial fossa, upper portion of mesopleuron, dorsal surface of petiolar node and postpetiole; sides of petiolar peduncle with plumose hairs, following the petiolar node; dorsum of gaster with plumose and squamiform hairs, relatively sparse, on the anterior face and anterolateral portion of procoxae and sparser on the dorsum of metacoxae. The second layer is composed of erect to suberect hairs, long and slightly clavate, present in the meso- and metasoma in the following conformation: dense on the dorsal surface of pronotum, petiolar node, postpetiole, and gaster (relatively sparse on the anterior region of first gastral sternite); one pair present on metanotal suture; 45 pairs on the back and sides of propodeum; slightly thinner and sparse on petiolar peduncle; two pairs on each side of anterior margin of postpetiolar sternite; present on the anterior face and anterolateral portion of procoxae; two to three pairs on the dorsum of meso- and metacoxae. Femora and tibiae covered by short, subdecumbent hairs (on tibiae, with a flatter aspect); minute appressed and clavate hairs also present on legs, more easily recognized on ventral surface of femora, tibiae (dorsum of meso- and metatibia as well) and basitarsi. Antennal pilosity unique: subdecumbent squamiform hairs on the basal lobe of scape, dense hairs, erect to suberect on scape dorsum; external margin with long erect hairs, slightly swollen on its median portion and distributed along the crenulation; short clavate and apically curved hairs also following the external margin; ventral face with short, subdecumbent setae; funicles densely covered with short, decumbent setae.
</paragraph>
<paragraph id="7E3D3E32FFA7E376FF5FFA5182FBDDBE" blockId="35.[151,1437,150,2013]" pageId="35" pageNumber="36">Body mostly smooth and shiny on glabrous regions. Dorsum of mandibles and clypeus covered with punctuations. Anteroventral portion of mandibles slightly alveolate. Anterior margin of labrum finely reticulate. Head dorsally rugose-foveate, irregular rugae forming fovea of different sizes; antennal fossa reticulate; antennal scrobe surface punctuate-foveate on its posterior half; head ventral face coarse and sparse punctuations. Pronotum foveate on anterior margin and sides. Mesonotum with lateral costulae, extending to the anterior face of propodeum. Gaster densely punctuate-reticulate; tergite of abdominal segments V, VI, and VII finely and densely punctate, slightly opaque, tergal margins smooth and shiny, slightly yellowish; more sparse sculpture on the first gastral sternite, especially in the anteromedial portion. Scapes finely reticulate. Legs smooth.</paragraph>
<paragraph id="7E3D3E32FFA7E371FF5FF9718364DA3B" blockId="35.[151,1437,150,2013]" lastBlockId="36.[151,1437,151,897]" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
Head oblong, sides emarginate from the eye insertion to posterolateral region and behind the head; posterior portion of head slightly depressed, projected posteriorly; vertexal margin with rounded corners, median portion broadly concave. Cervical margin strongly carinate. Palp formula 2,2; palps strongly fused; maxillary palp slightly larger and wider than labial, with one sensillum; labial palp apically clavate, with two sensilla. Stipes subrectangular. Labrum lunate, laterally curved (apex
<quantity id="B97A93D7FFA7E376FDE8F8E183C3DCCB" box="[624,728,1914,1941]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="35" pageNumber="36" unit="mm" value="0.07">0.07 mm</quantity>
); distal margin rounded, medial knob present. Mandibles triangular; in full-face view, lateral margins slightly concave; basal margin conspicuously concave, clypeomandibular space wide; basal angle rounded, followed by about 20 slightly rounded denticles and gradually decreasing in size along the masticatory margin; in lateral view mandibular apex slightly curved ventrally. Clypeus lamellated laterally and on the anterolateral portion; anterior margin concave medially. Short frontal sulcus present separating the dorsal convexity into two parts. Compound eyes large. Carina-like rugae below eyes, projecting from the middle of the antennal fossa margin and extending to the side of the head, like an arc. Scape with lamellar basal angle slightly rounded, external margin crenulate and lamellar. Antennal fossa deep. Antennal scrobe deep on the anterior half, posterior and ventral limits weakly distinct.
</paragraph>
<paragraph id="7E3D3E32FFA0E371FF5FFEF48314D8DF" blockId="36.[151,1437,151,897]" pageId="36" pageNumber="37">Mesosoma slender; lateral profile of promesonotal complex continuously convex, strongly sloping caudad; metanotal suture wide. Mesopleuron bordered anteriorly, interrupted at the meeting of a narrow epicnemial fossa. In lateral view, anterior portion of the propodeum short and slightly oblique posteriorly, dorsal surface of propodeum slightly sloping posteriorly. Propodeal slope laterally carinate and with strong transverse lamellar carina connecting to sharp and tapered, slightly curved upwards propodeal projection. Opening of propodeal spiracle round. Bulb of metapleural gland prominent, projected and slightly narrowed; opening transversal and covered by cuticular lamella. Dorsal surface of petiolar peduncle careened longitudinally. In dorsal view, propodeal spiracle slightly projected dorsally. In lateral view, petiole long and claviform; postpetiole subcircular, subequal to the length of petiolar node. Subpetiolar process variable: from anterior process slightly curved anteriorly, followed by 48 spines of various sizes to slightly bifid process followed by simple or double spines and vestigial processes on the posterior portion of peduncle. In dorsal view, petiolar node ogive-shape; posterior margin of postpetiole convex and widely inserted to anterior concavity of gaster. Postpetiolar sternite conspicuously carinate anteromedially, prora present, lamellar and slightly darkened. Anterodorsal lamellar margin of gaster yellowish; median longitudinal sulcus present on first tergite, shallow and glabrous. Calcar of strigil pectinate. Meso- and metabasitarsi conspicuously long, about ¾ of tibial length. Tarsal claws simple.</paragraph>
<caption id="2AFD6EBAFFA0E371FF0FF9CB8358DDD2" ID-DOI="http://doi.org/10.5281/zenodo.6605993" ID-Zenodo-Dep="6605993" httpUri="https://zenodo.org/record/6605993/files/figure.png" pageId="36" pageNumber="37" startId="36.[151,250,1616,1640]" targetBox="[190,1397,921,1591]" targetPageId="36">
<paragraph id="7E3D3E32FFA0E371FF0FF9CB8358DDD2" blockId="36.[151,1436,1616,1676]" pageId="36" pageNumber="37">
<emphasis id="4CF6E220FFA0E371FF0FF9CB803ADD36" bold="true" box="[151,289,1616,1640]" pageId="36" pageNumber="37">FIGURE 13.</emphasis>
<taxonomicName id="B98245B1FFA0E371FEB0F9CA80CADD36" authorityName="Brown &amp; Kempf" authorityYear="1960" box="[296,465,1617,1640]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="manni">
<emphasis id="4CF6E220FFA0E371FEB0F9CA80CADD36" box="[296,465,1617,1640]" italics="true" pageId="36" pageNumber="37">Basiceros manni</emphasis>
</taxonomicName>
, worker (CASENT0914888, Costa Rica: Heredia); A. full-face view, B. lateral view, C. dorsal view. Images by Michele Esposito (CAS).
</paragraph>
</caption>
<paragraph id="7E3D3E32FFA0E371FF5FF920850EDCD6" blockId="36.[151,1437,1722,2037]" pageId="36" pageNumber="37">
<emphasis id="4CF6E220FFA0E371FF5FF9208019DD8B" box="[199,258,1723,1749]" italics="true" pageId="36" pageNumber="37">Gyne</emphasis>
(n=3). HL 1.48155, HL2 1.531.62, HW1 1.331.40, MdL 1.00, SL1 1.191.22, SL2 1.181.20, PDL 0.150.16, A3L 0.050.06, AFL 0.480.50, FuL 1.311.36, EL 0.310.33, EW 0.250.28, LOD 0.060.09, MOD 0.080.09, OOD 0.300.37, ML 2.342.47, MSL 1.061.15, MSW 0.931.00, MLL 0.280.34, MLW 0.440.45, MfL 1.952.00, MtL 1.501.60,
<collectingCountry id="06957EA2FFA0E371FD8BF8BC832CDC1F" box="[531,567,1831,1857]" name="Philippines" pageId="36" pageNumber="37">PH</collectingCountry>
0.350.40, PL 1.031.15, PW 0.400.48, PPL 0.650.69, PPW 0.670.70, GL 2.112.28, GW 1.501.53, TL 8.759.08, CI 8991, CS 1.401.47, MCI 6467, SI 8592, ESI 2127, SAI2 240245, EI1 0.370.41, MTI 8094, MLI 127163, MFI 6770, PTI 284292.
</paragraph>
<paragraph id="7E3D3E32FFA0E370FF5FF809830CD867" blockId="36.[151,1437,1722,2037]" lastBlockId="37.[151,1437,151,825]" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37">
Coloration and sculpture similar to conspecific workers; size slightly larger. Cephalic dorsum with three ocelli: median ocelli inserted at the top of the frontal sulcus and lateral ocelli inserted slightly above. Head pilosity as in conspecific workers. Double layer of specialized pilosity on pronotum denser than in workers. Mesoscutum almost lacking basal pilosity, only two subdecumbent hairs present, with a plumose aspect, anteriorly to each parapsis, and four hairs close to the margin of the scutoscutellar suture; erect pilosity relatively sparse, present primarily on lateral margins of mesoscutum; central portion of mesoscutum nearly glabrous. Mesoscutellum with double-layer of pilosity. Median region of metanotum with a pair of long, clavate and suberect hairs and four hairs from the basal layer with a plumose appearance. Posterior corner of mesoanepisternum, just below forewing insertion, with basal pilosity. Ventral and posterior limits of mesokatepisternum with basal pilosity, as well as the ventral portion of metakatepisternum. Dorsum of meso- and metacoxa with specialized pilosity. Remaining regions with pilosity shape and configuration similar to conspecific workers. Mesoscutum with irregular longitudinal rugae, interspersed with rugulae forming fovea of different sizes. Mesoscutellum irregularly rugo-foveate. Mesopleuron smooth, subopaque (weakly granular aspect) on most of its extension, except for foveate posterolateral portion of mesoanepisternum. Anapleural sulcus smooth. Anterior portion of petiolar peduncle finely reticulate. Gastral sculpture deeper than on workers. Other regions of the body ornamented as conspecific workers. In dorsal view, humeral angles strongly projected, dividing pronotum into two portions. Mesoscutum anteriorly rounded and slightly wedge-shaped; anteromedial carina present, short and longitudinal; posterior margin broadly convex at the meeting with the scutoscutellar suture; notauli indistinct; parapsidal lines obsolete, involved by the sculpture; parapsis subreniform, slightly deep, tegulae narrow, elongated, apical margin rounded. Pre-scutellum narrow; axillae projected posteriorly, rounded and slightly depressed. Scutoscutellar suture well-marked, transversally costulate. Mesoscutellum trapezoidal, posterior limit concave. First gastral sternite with a slightly projecting median region on basal half. Forewing
<typeStatus id="A1398090FFA1E370FA93FD608420D84B" box="[1291,1339,763,789]" pageId="37" pageNumber="38">type</typeStatus>
1; hindwing with five submedian hamuli.
</paragraph>
<caption id="2AFD6EBAFFA1E370FF0FF907858EDD86" ID-DOI="http://doi.org/10.5281/zenodo.6605995" ID-Zenodo-Dep="6605995" httpUri="https://zenodo.org/record/6605995/files/figure.png" pageId="37" pageNumber="38" startId="37.[151,250,1692,1716]" targetBox="[174,1414,849,1669]" targetPageId="37">
<paragraph id="7E3D3E32FFA1E370FF0FF907858EDD86" blockId="37.[151,1437,1692,1753]" pageId="37" pageNumber="38">
<emphasis id="4CF6E220FFA1E370FF0FF9078004DDEA" bold="true" box="[151,287,1692,1716]" pageId="37" pageNumber="38">FIGURE 14.</emphasis>
<taxonomicName id="B98245B1FFA1E370FEBDF90680D6DDEA" authorityName="Brown &amp; Kempf" authorityYear="1960" box="[293,461,1693,1716]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="manni">
<emphasis id="4CF6E220FFA1E370FEBDF90680D6DDEA" box="[293,461,1693,1716]" italics="true" pageId="37" pageNumber="38">Basiceros manni</emphasis>
</taxonomicName>
, gyne (MCZ 546486, Costa Rica: Heredia) and male (MCZ, Costa Rica: Heredia); A and D. fullface view, B and E. dorsal view, C and F. lateral view. Scale bars: A, D = 0.5 mm, B, C, E, F =1 mm.
</paragraph>
</caption>
<paragraph id="7E3D3E32FFA1E370FF5FF89C81EFDCA4" blockId="37.[151,1437,1799,2042]" pageId="37" pageNumber="38">
<emphasis id="4CF6E220FFA1E370FF5FF89C801ADC7F" box="[199,257,1799,1825]" italics="true" pageId="37" pageNumber="38">Male</emphasis>
(n=2).
<materialsCitation id="CEEA346FFFA1E370FECCF89C81EADCA4" ID-GBIF-Occurrence="4029332370" collectionCode="MOD, PL, PW, CS" country="Philippines" location="PH" pageId="37" pageNumber="38" specimenCode="ML 1.88" specimenCount="1" specimenCount-male="1">
HL 0.930.96, HW1 0.95, HW2 1.10, MdL 0.550.56, SL2 0.200.23, PDL 0.130.15, A3L 0.48, AFL 0.610.63, EL 0.350.38, EW 0.280.31, LOD 0.090.10,
<collectionCode id="1893A6F7FFA1E370FC1FF8B082D3DC1B" box="[903,968,1835,1861]" country="Italy" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14382" lsid="urn:lsid:biocol.org:col:14382" name="University of Modena and Reggio Emilia, Department of Biology" pageId="37" pageNumber="38" type="University or college">MOD</collectionCode>
0.09, OOD 0.320.37,
<specimenCode id="2E249649FFA1E370FB74F8B08448DC1B" box="[1260,1363,1835,1861]" collectionCode="ML" country="France" name="Musee de Lectoure" pageId="37" pageNumber="38">ML 1.88</specimenCode>
, MSL 0.931.05, MSW 0.780.80, MLL 0.330.38, MLW 0.450.46, MfL 1.801.85, MtL 1.291.33,
<collectingCountry id="06957EA2FFA1E370FB4BF8D485ECDC37" box="[1235,1271,1871,1897]" name="Philippines" pageId="37" pageNumber="38">PH</collectingCountry>
0.250.30,
<collectionCode id="1893A6F7FFA1E370FAE5F8D48486DC37" box="[1405,1437,1871,1897]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="37" pageNumber="38" type="Museum">PL</collectionCode>
0.880.90,
<collectionCode id="1893A6F7FFA1E370FE8DF8E88025DCD3" box="[277,318,1907,1933]" country="Thailand" name="Paleontological Collections" pageId="37" pageNumber="38">PW</collectionCode>
0.350.38, PPL 0.43, PPW 0.43
<emphasis id="4CF6E220FFA1E370FD2EF8E883FCDCD3" bold="true" box="[694,743,1907,1933]" pageId="37" pageNumber="38">0.50</emphasis>
, GL 1.631.80, GW 1.151.25, TL 6.356.45, CI 98102,
<collectionCode id="1893A6F7FFA1E370FAE2F8E88487DCD3" box="[1402,1436,1907,1933]" country="France" name="Musee des Dinosaures d'Esperaza (Aude)" pageId="37" pageNumber="38">CS</collectionCode>
0.940.96, MCI 5859, SI 2123, ESI 166175, SAI 4247, SAI2 3236, EI1 4247, EI2 120123, MTI 7881, MLI 123138, MFI 5152, PTI 300350. (In bold: for
<specimenCount id="6884F5BBFFA1E370FD64F820827FDC8B" box="[764,868,1979,2005]" pageId="37" pageNumber="38" type="male">one male</specimenCount>
, wings partially covered the lateral margin of postpetiole)
</materialsCitation>
.
</paragraph>
<paragraph id="7E3D3E32FFA2E373FF5FFF0C832DDAAB" blockId="38.[151,1437,151,2013]" pageId="38" pageNumber="39">
Slightly smaller than conspecific gyne. Coloration dark brown to black, with sides of pronotum and mesoscutum, posterodorsal region of scutellum and portion of dorsum of petiole brown; postpetiole and anterior to anterolateral region of gaster testaceous; appendages brown to light brown; antennas with opaque appearance. Wings whitish to yellowish. Mandibles with long yellow hairs close to their apexes; dorsum with fine yellowish hairs and subdecumbent, slightly shorter. Head with two main
<typeStatus id="A1398090FFA2E373FD6AFEBC8236DA1F" box="[754,813,295,321]" pageId="38" pageNumber="39">types</typeStatus>
of hair: medium to short, yellow and fine, erect to suberect, sparse; and long and yellowish, generally with a curved apex and primarily on frons, vertexal, and ventral regions. The second hair
<typeStatus id="A1398090FFA2E373FE2DFEF480FEDAD7" box="[437,485,367,393]" pageId="38" pageNumber="39">type</typeStatus>
widely present throughout the body: on the dorsum of mesosoma, waist and gaster, coxae and mesokatepisternum suture. Filiform subdecumbent and short hairs present on the first gastral tergite and sternite. Antennomeres with short appressed whitish setae. Legs with suberect setae sparsely distributed among short decumbent to decumbent setae.
</paragraph>
<paragraph id="7E3D3E32FFA2E373FF5FFE648229D9DB" blockId="38.[151,1437,151,2013]" pageId="38" pageNumber="39">Body uniformly punctuate-reticulate (sculpture changing diameter and degree of impression). Distal half of mandibles smooth and shiny. Irregular longitudinal rugae on the central disc of clypeus, on head dorsum, and neck. Irregular transverse rugae near the occipital carina; on the dorsum of mesosoma; on pronotum sides, and propodeum. Mesoanepisternum shiny on its anterodorsal portion.</paragraph>
<paragraph id="7E3D3E32FFA2E373FF5FFD1485C3DF4F" blockId="38.[151,1437,151,2013]" pageId="38" pageNumber="39">Head piriform; occipital margin wide and lamellar. Palp formula 1,1; palps apically subclavate and subequal; maxillary palp slightly wider than labial; apexes of palpi with long, filiform sensillum. In ventral view, mentum arrow-shaped. Labrum trapezoidal; basal region transversely striate; distal margin weakly concave; corners angled; 1012 long and erect setae present. Mandibles triangular, elongated and gently curved apically; masticatory margin with 1214 conical teeth, the basal and the subbasal sometimes extremely short and apical tooth slightly falcate. Clypeus with convex central disc and lateral regions depressed; anterior margin lamellar and moderately flat; frontoclypeal region excavated. Carina genal present on head side, extending posteriorly to occipital carina. Frons with arc-shaped carina above the antennal insertions and connected to frontal lobes. Antennal scrobes deep and marginally lamellar. Antennal arch expanded to posterolateral lobe, hiding the antennal bulb in full-face view. Pedicel longer than wide, third antennomere about five times longer than pedicel. Eyes large, globular, protruding from cephalic capsule; posterior margin emarginated. Ocelli pearl-shaped, projected on carinate crests.</paragraph>
<paragraph id="7E3D3E32FFA2E373FF5FFB8085CCDDC4" blockId="38.[151,1437,151,2013]" pageId="38" pageNumber="39">
In dorsal view, mesoscutum keel-shaped; pronotum transversally projected anterodorsad by a carinate conical or rounded process. This carina extends laterally through the pronotum, dividing it into a lower and an upper portion and producing triangular lateral flaps. Broad and partially shiny carina present on the anteromedial region of mesoscutum, extending to the dorsal region. Anterior portion of notauli separating the mesoscutum into two distinct regions: an anterior, narrow and slightly higher and obliquely convex in profile, delimited by the notauli lateroposterad and forming a projection at this region; and one posterior region, subquadrate. Notauli broad and somewhat deep, converging posteriorly on a sinuous longitudinal carina almost continuously covering mesoscutal surface. Parapsidal lines shiny; curved on anterior portion, subparallel and directed anterolaterally to the parascutal flange; posteriorly reaching the notauli. Parapsides broad, deep and reniform. Transscutal articulation convex. Axillae strongly curved posteriorly, projected and hook-shaped. Anapleural sulcus broad, deep—mesoanepisternum considerably elevated above the mesokatepisternum; scrobiculate and slit-shaped; posterior limit oblique. Scutoscutellar sulcus wide and longitudinally scrobiculate. Mesoscutellum subquadrate; posterior margin concave, crenulate and moderately depressed. In dorsal view, metanotum distinct, posterior margin projected, lateral margins lamellar. Propodeal spines short and triangular; carinate as in conspecific females. Propodeal lobes auricular. Calcar of strigil pectinate. Tarsal claws simple; short arolia present. Shape of petiole similar to conspecific females, claviform; in dorsal view, petiolar spiracle projected; anteroventral process followed by minute triangular spines. Postpetiole short; in lateral view, dorsally convex; spiracle projected; sternite with transverse lamellar prora near the anterior margin. Forewing
<typeStatus id="A1398090FFA2E373FF0FF9E481DCDDC7" box="[151,199,1663,1689]" pageId="38" pageNumber="39">type</typeStatus>
1; hindwing with five to six submedian hamuli (four in
<specimenCount id="6884F5BBFFA2E373FCA5F9E482C2DDC7" box="[829,985,1663,1689]" pageId="38" pageNumber="39" type="generic">one specimen</specimenCount>
, probably due to loss).
</paragraph>
<paragraph id="7E3D3E32FFA2E372FF5FF938845FDA8F" blockId="38.[151,1437,151,2013]" lastBlockId="39.[151,1437,151,465]" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39">
<emphasis id="4CF6E220FFA2E373FF5FF9388011DDE2" box="[199,266,1699,1724]" italics="true" pageId="38" pageNumber="39">Larva</emphasis>
. Approximate length through spiracles:
<quantity id="B97A93D7FFA2E373FD78F9398227DDE3" box="[736,828,1698,1725]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="38" pageNumber="39" unit="mm" value="5.1">5.1 mm</quantity>
; profile pogonomyrmecoid: larger diameter near the middle of abdominal region, slender thorax; curved ventrally. Anus ventral, anal opening weakly convex, anal flap present. Spiracles minute. Integument of ventral region densely covered by spinules arranged in transverse rows. Pilosity moderate, denser on abdominal dorsum and ventral margin, sparse on rest of the body; long hairs—especially on part of head and ventral margin of thorax—(
<quantity id="B97A93D7FFA2E373FC9EF8A982B6DC13" box="[774,941,1842,1869]" metricMagnitude="-4" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" pageId="38" pageNumber="39" unit="mm" value="0.22499999999999998" valueMax="0.3" valueMin="0.15">0.150.30 mm</quantity>
) flexuous and denticulate; alveolus and articular membrane present or not. Cranium slightly suboctagonal, occipital margin with median impression; antenna extremely minute. Clypeus protruding. Labrum bilobed, approximately three times wider than long; ventral surface densely spinous; anteroventral margin of each lobe with about six rounded sensilla; anterodorsal border with four isolated short setae resembling trichoid sensilla. Mandibles pogonomyrmecoid, long and narrow, curved; subapical tooth slightly forward to the plane of apical tooth; basal tooth strongly projected into the inner margin; anteroventral margin densely covered by subtransverse lines of short and arcuate spinules. Maxillae parabolic, dorsum with five setae, apex spinose; maxillary palps and galeae digitiform, subequal in diameter; maxillary palp with five sensilla: two apical, encapsulated, two lateral with one spinule, and one subapical spinulate; galeae with two apical spinulate sensilla. Labium covered by short transverse rows of spinules; spinules developed, capillary (about
<quantity id="B97A93D7FFA3E372FB66FEBC847FDA1F" box="[1278,1380,295,321]" metricMagnitude="-5" metricUnit="m" metricValue="2.0" pageId="39" pageNumber="40" unit="mm" value="0.02">0.02 mm</quantity>
) and densely covering the area surrounding the opening of the sericeous gland, shorter spinules on the rest of labium surface; labial palp papillary, with five apical sensilla (two encapsulated and three with spinules); three trichoid sensilla isolated under the labial palp, in triangular configuration. Opening of sericeous gland transversal, each side with a short and isolated sensillum. Hypopharynx densely spinous; spinules arranged in short subtransverse rows.
</paragraph>
</subSubSection>
<caption id="2AFD6EBAFFA3E372FF0FF8A583D6DC24" ID-DOI="http://doi.org/10.5281/zenodo.6605997" ID-Zenodo-Dep="6605997" httpUri="https://zenodo.org/record/6605997/files/figure.png" pageId="39" pageNumber="40" startId="39.[151,250,1854,1878]" targetBox="[253,1332,489,1831]" targetPageId="39">
<paragraph id="7E3D3E32FFA3E372FF0FF8A583D6DC24" blockId="39.[151,1436,1854,1914]" pageId="39" pageNumber="40">
<emphasis id="4CF6E220FFA3E372FF0FF8A5803ADC08" bold="true" box="[151,289,1854,1878]" pageId="39" pageNumber="40">FIGURE 15.</emphasis>
<taxonomicName id="B98245B1FFA3E372FEB1F8A480C8DC08" authorityName="Brown &amp; Kempf" authorityYear="1960" box="[297,467,1855,1878]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="manni">
<emphasis id="4CF6E220FFA3E372FEB1F8A480C8DC08" box="[297,467,1855,1878]" italics="true" pageId="39" pageNumber="40">Basiceros manni</emphasis>
</taxonomicName>
, larva (MZSP, Nicaragua: Boaco); A. head in anteroventral view, B. dorsal pilosity, C. lateral view. Scale bars: A = 0.1 mm, C = 1 mm, B = 0.05 mm.
</paragraph>
</caption>
<subSubSection id="36986DB9FFA3E37DFF5FF83280C8DA43" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40" type="etymology">
<paragraph id="7E3D3E32FFA3E37DFF5FF83280C8DA43" blockId="39.[151,1436,1960,2023]" lastBlockId="40.[151,1437,151,2013]" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40">
<emphasis id="4CF6E220FFA3E372FF5FF8328055DC9D" bold="true" box="[199,334,1961,1987]" pageId="39" pageNumber="40">Etymology.</emphasis>
This species was named after Dr. William M. Mann (18861960), a prominent American entomologist and myrmecologist. Dr. Mann was a doctoral student of William Morton Wheeler at Harvard University and worked at the
<collectingCountry id="06957EA2FFACE37DFEABFF0C80D3DBEF" box="[307,456,151,177]" name="United States of America" pageId="40" pageNumber="41">United States</collectingCountry>
Department of Agriculture (USDA) as an entomologist, conducting several expeditions, mainly in Central and South America.According to
<bibRefCitation id="1A1343C3FFACE37DFD4EFF208507DB8B" author="Brown, W. L. Jr. &amp; Kempf, W. W." box="[726,1052,187,213]" pageId="40" pageNumber="41" pagination="161 - 250" refId="ref61211" refString="Brown, W. L. Jr. &amp; Kempf, W. W. (1960) A world revision of the ant tribe Basicerotini (Hym. Formicidae). Studia Entomologica, New Series, 3, 161 - 250." type="journal article" year="1960">Brown &amp; Kempf (1960: 178)</bibRefCitation>
, Dr. Mann was the first to discover this taxon (he collected much of the type-series on expeditions in
<collectingCountry id="06957EA2FFACE37DFCE5FF4482ECDBA7" box="[893,1015,223,249]" name="Costa Rica" pageId="40" pageNumber="41">Costa Rica</collectingCountry>
and
<collectingCountry id="06957EA2FFACE37DFBB7FF448585DBA4" box="[1071,1182,223,250]" name="Honduras" pageId="40" pageNumber="41">Honduras</collectingCountry>
) and to recognize it as a new species (
<bibRefCitation id="1A1343C3FFACE37DFED8FE9880DDDA43" author="Mann, W. M." box="[320,454,259,285]" pageId="40" pageNumber="41" pagination="1 - 54" refId="ref62206" refString="Mann, W. M. (1922) Ants from Honduras and Guatemala. Proceedings of the United States National Museum, 61, 1 - 54. https: // doi. org / 10.5479 / si. 00963801.61 - 2434.1" type="journal article" year="1922">Mann 1922</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="36986DB9FFACE37DFF5FFEBD83FAD993" pageId="40" pageNumber="41" type="discussion">
<paragraph id="7E3D3E32FFACE37DFF5FFEBD8427D947" blockId="40.[151,1437,151,2013]" pageId="40" pageNumber="41">
<emphasis id="4CF6E220FFACE37DFF5FFEBD8054DA1E" bold="true" box="[199,335,294,320]" pageId="40" pageNumber="41">Comments.</emphasis>
<taxonomicName id="B98245B1FFACE37DFEC1FEBC830DDA1E" authorityName="Brown &amp; Kempf" authorityYear="1960" box="[345,534,295,320]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="manni">
<emphasis id="4CF6E220FFACE37DFEC1FEBC830DDA1E" box="[345,534,295,320]" italics="true" pageId="40" pageNumber="41">Basiceros manni</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="B98245B1FFACE37DFD75FEBC8265DA1F" baseAuthorityName="Smith" baseAuthorityYear="1858" box="[749,894,295,321]" class="Insecta" family="Formicidae" genus="Meranoplus" kingdom="Animalia" order="Hymenoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="singularis">
<emphasis id="4CF6E220FFACE37DFD75FEBC8265DA1F" box="[749,894,295,321]" italics="true" pageId="40" pageNumber="41">B. singularis</emphasis>
</taxonomicName>
in several characteristics, such as size, general morphology, and pilosity. Specimens of both species are easily mistaken when covered with litter and soil particles. In terms of variation, there are differences in the shape of the subpetiolar process, sculpture, and in a lesser degree in pilosity for the female castes. For males, the variation observed is mainly in the sculpture (especially in the mesosoma) and shape of the transverse keel of the anterior margin of the pronotum, whose apex ranges from conical to slightly rounded.
<bibRefCitation id="1A1343C3FFACE37DFEC4FE4080F1DAAB" author="Dietz, B. H." box="[348,490,475,501]" pageId="40" pageNumber="41" refId="ref61457" refString="Dietz, B. H. (2004) Uma revisao de Basicerotini Brown, 1948 (Formicidae: Myrmicinae), suas relacoes filogeneticas internas e com outras tribos dacetineas (Dacetini e Phalacromyrmecini). Instituto de Biociencias da Universidade de Sao Paulo (IB / USP), Sao Paulo, 245 pp." type="book" year="2004">Dietz (2004)</bibRefCitation>
mentions that in dorsal view it is impossible to visualize the metanotal flange on the male of
<taxonomicName id="B98245B1FFACE37DFF6BFE648043D946" authorityName="Brown &amp; Kempf" authorityYear="1960" box="[243,344,511,536]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="manni">
<emphasis id="4CF6E220FFACE37DFF6BFE648043D946" box="[243,344,511,536]" italics="true" pageId="40" pageNumber="41">B. manni</emphasis>
</taxonomicName>
, however part of the metanotum is present on specimens examined for the present study.
</paragraph>
<paragraph id="7E3D3E32FFACE37DFF5FFDB8820AD9DB" blockId="40.[151,1437,151,2013]" pageId="40" pageNumber="41">
<materialsCitation id="CEEA346FFFACE37DFF5FFDB88217D9DB" ID-GBIF-Occurrence="4029332333" collectorName="E. O. Wilson &amp; B. Holldobler" country="Costa Rica" location="La Selva Biological Station" pageId="40" pageNumber="41" specimenCount="1">
Most males examined here are part of nest series collected by
<collectorName id="D3775BE4FFACE37DFC1AFDB88517D963" box="[898,1036,547,573]" pageId="40" pageNumber="41">E.O. Wilson</collectorName>
and
<collectorName id="D3775BE4FFACE37DFBDBFDB885C6D963" box="[1091,1245,547,573]" pageId="40" pageNumber="41">B. Hölldobler</collectorName>
at
<location id="7B5D68E9FFACE37DFA98FDB88024D93F" LSID="urn:lsid:plazi:treatment:F62B8F24FFA6E37FFF0FFAAB85D1DA1F:7B5D68E9FFACE37DFA98FDB88024D93F" country="Costa Rica" name="La Selva Biological Station" pageId="40" pageNumber="41">La Selva Biological Station</location>
, in
<collectingCountry id="06957EA2FFACE37DFEF4FDDC80F0D93F" box="[364,491,583,609]" name="Costa Rica" pageId="40" pageNumber="41">Costa Rica</collectingCountry>
, with
<specimenCount id="6884F5BBFFACE37DFDADFDDC83F0D93F" box="[565,747,583,609]" pageId="40" pageNumber="41" type="generic">three specimens</specimenCount>
obtained by
<bibRefCitation id="1A1343C3FFACE37DFC1BFDDC85D1D93C" author="Holldobler, B. &amp; Wilson, E. O." box="[899,1226,583,610]" pageId="40" pageNumber="41" pagination="12 - 20" refId="ref61910" refString="Holldobler, B. &amp; Wilson, E. O. (1986). Soil-binding pilosity and camouflage in ants of the tribes Basicerotini and Stegomyrmecini. Zoomorphology, 106, 12 - 20. https: // doi. org / 10.1007 / BF 00311942" type="journal article" year="1986">Wilson &amp; Hölldobler (1986)</bibRefCitation>
from lab colonies during an ethological project involving those nest series
</materialsCitation>
.
</paragraph>
<paragraph id="7E3D3E32FFACE37DFF5FFD1483FAD993" blockId="40.[151,1437,151,2013]" pageId="40" pageNumber="41">
A brachypterous intercaste have been collected for this species in
<collectingCountry id="06957EA2FFACE37DFC5CFD14852CD9F7" box="[964,1079,655,681]" name="Nicaragua" pageId="40" pageNumber="41">Nicaragua</collectingCountry>
(Dr. Michael Branstetter, pers. comm.). However, this specimen was not examined.
</paragraph>
</subSubSection>
<subSubSection id="36986DB9FFACE37CFF5FFD4D845EDE17" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41" type="distribution">
<paragraph id="7E3D3E32FFACE37DFF5FFD4D830EDF07" blockId="40.[151,1437,151,2013]" pageId="40" pageNumber="41">
<emphasis id="4CF6E220FFACE37DFF5FFD4D807AD9AE" bold="true" box="[199,353,726,752]" pageId="40" pageNumber="41">Distribution.</emphasis>
This species is recorded from low elevations, in primary or low anthropogenically disturbed forests, in
<collectingCountry id="06957EA2FFACE37DFF72FD60807DD84B" box="[234,358,763,789]" name="Costa Rica" pageId="40" pageNumber="41">Costa Rica</collectingCountry>
,
<collectingCountry id="06957EA2FFACE37DFEE9FD6080FAD84B" box="[369,481,763,789]" name="Honduras" pageId="40" pageNumber="41">Honduras</collectingCountry>
, and
<collectingCountry id="06957EA2FFACE37DFD82FD608396D84B" box="[538,653,763,789]" name="Nicaragua" pageId="40" pageNumber="41">Nicaragua</collectingCountry>
and is likely restricted to this geographic distribution. Records for this species for
<collectingCountry id="06957EA2FFACE37DFE80FC84806CD867" box="[280,375,799,825]" name="Ecuador" pageId="40" pageNumber="41">Ecuador</collectingCountry>
,
<collectingCountry id="06957EA2FFACE37DFE1BFC8480C4D867" box="[387,479,799,825]" name="Panama" pageId="40" pageNumber="41">Panama</collectingCountry>
,
<collectingCountry id="06957EA2FFACE37DFE72FC8483CCD867" box="[490,727,799,825]" name="Trinidad and Tobago" pageId="40" pageNumber="41">Trinidad and Tobago</collectingCountry>
, and islands in the Western Caribbean Sea (case of
<collectingCountry id="06957EA2FFACE37DFAB8FC84806ED800" name="Saint Vincent and the Grenadines" pageId="40" pageNumber="41">St. Vincent and the Grenadines</collectingCountry>
) are not supported in the present study. Based on the specimens examined during this study,
<taxonomicName id="B98245B1FFACE37DFF0FFCFC81E6D8DE" authorityName="Brown &amp; Kempf" authorityYear="1960" box="[151,253,871,896]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="manni">
<emphasis id="4CF6E220FFACE37DFF0FFCFC81E6D8DE" box="[151,253,871,896]" italics="true" pageId="40" pageNumber="41">B. manni</emphasis>
</taxonomicName>
does not occur in
<collectingCountry id="06957EA2FFACE37DFE57FCFC8337D8DF" box="[463,556,871,897]" name="Ecuador" pageId="40" pageNumber="41">Ecuador</collectingCountry>
or
<collectingCountry id="06957EA2FFACE37DFDCAFCFC8224D8DF" box="[594,831,871,897]" name="Trinidad and Tobago" pageId="40" pageNumber="41">Trinidad and Tobago</collectingCountry>
, and some specimens of
<taxonomicName id="B98245B1FFACE37DFBC2FCFC85F2D8DF" baseAuthorityName="Smith" baseAuthorityYear="1858" box="[1114,1257,871,897]" class="Insecta" family="Formicidae" genus="Meranoplus" kingdom="Animalia" order="Hymenoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="singularis">
<emphasis id="4CF6E220FFACE37DFBC2FCFC85F2D8DF" box="[1114,1257,871,897]" italics="true" pageId="40" pageNumber="41">B. singularis</emphasis>
</taxonomicName>
examined from these two points were wrongly identified as belonging to
<taxonomicName id="B98245B1FFACE37DFC87FC10829FD8FA" authorityName="Brown &amp; Kempf" authorityYear="1960" box="[799,900,907,932]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="manni">
<emphasis id="4CF6E220FFACE37DFC87FC10829FD8FA" box="[799,900,907,932]" italics="true" pageId="40" pageNumber="41">B. manni</emphasis>
</taxonomicName>
. It was not possible to access material from the other locations mentioned and there is no way to ensure that this species occurs there. The MCZ online database highlights that those
<collectingCountry id="06957EA2FFACE37DFE1BFC488340D8B3" box="[387,603,979,1005]" name="Trinidad and Tobago" pageId="40" pageNumber="41">Trinidad &amp; Tobago</collectingCountry>
specimens might have unmatched specimen label information in terms of locality. Additionally, a worker collected at
<collectingCountry id="06957EA2FFACE37DFDE7FC6C823EDF4F" box="[639,805,1015,1041]" name="French Guiana" pageId="40" pageNumber="41">French Guiana</collectingCountry>
by The Joint Research Unit Ecology of Guianan Forests (EcoFoG) (AntWeb specimen code ECOFOG-IT14-0106-61) and identified as
<taxonomicName id="B98245B1FFACE37DFB8AFB808563DF6A" authorityName="Brown &amp; Kempf" authorityYear="1960" box="[1042,1144,1051,1076]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="manni">
<emphasis id="4CF6E220FFACE37DFB8AFB808563DF6A" box="[1042,1144,1051,1076]" italics="true" pageId="40" pageNumber="41">B. manni</emphasis>
</taxonomicName>
by Franco
<emphasis id="4CF6E220FFACE37DFB60FB808432DF6B" box="[1272,1321,1051,1077]" italics="true" pageId="40" pageNumber="41">et al</emphasis>
. (2019) is actually a worker of
<taxonomicName id="B98245B1FFACE37DFEE7FBA48315DF07" baseAuthorityName="Smith" baseAuthorityYear="1858" box="[383,526,1087,1113]" class="Insecta" family="Formicidae" genus="Meranoplus" kingdom="Animalia" order="Hymenoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="singularis">
<emphasis id="4CF6E220FFACE37DFEE7FBA48315DF07" box="[383,526,1087,1113]" italics="true" pageId="40" pageNumber="41">B. singularis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="7E3D3E32FFACE37DFF5FFBF885D9DEC3" blockId="40.[151,1437,151,2013]" pageId="40" pageNumber="41">
<emphasis id="4CF6E220FFACE37DFF5FFBF8808BDF22" bold="true" box="[199,400,1122,1149]" pageId="40" pageNumber="41">Natural History:</emphasis>
<taxonomicName id="B98245B1FFACE37DFE0FFBF88348DF22" authorityName="Brown &amp; Kempf" authorityYear="1960" box="[407,595,1123,1148]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="manni">
<emphasis id="4CF6E220FFACE37DFE0FFBF88348DF22" box="[407,595,1123,1148]" italics="true" pageId="40" pageNumber="41">Basiceros manni</emphasis>
</taxonomicName>
is the species of the genus with the greatest amount of behavioral ecology data, thanks largely to the work of
<bibRefCitation id="1A1343C3FFACE37DFD8EFB1C8249DFFC" author="Holldobler, B. &amp; Wilson, E. O." box="[534,850,1159,1186]" pageId="40" pageNumber="41" pagination="12 - 20" refId="ref61910" refString="Holldobler, B. &amp; Wilson, E. O. (1986). Soil-binding pilosity and camouflage in ants of the tribes Basicerotini and Stegomyrmecini. Zoomorphology, 106, 12 - 20. https: // doi. org / 10.1007 / BF 00311942" type="journal article" year="1986">Wilson &amp; Hölldobler (1986)</bibRefCitation>
and
<bibRefCitation id="1A1343C3FFACE37DFC1EFB1C85D9DFFC" author="Holldobler, B. &amp; Wilson, E. O." box="[902,1218,1159,1186]" pageId="40" pageNumber="41" pagination="12 - 20" refId="ref61910" refString="Holldobler, B. &amp; Wilson, E. O. (1986). Soil-binding pilosity and camouflage in ants of the tribes Basicerotini and Stegomyrmecini. Zoomorphology, 106, 12 - 20. https: // doi. org / 10.1007 / BF 00311942" type="journal article" year="1986">Hölldobler &amp; Wilson (1986)</bibRefCitation>
and to the information recorded on the page http://ants.biology.utah.edu/genera/basiceros/species/manni/manni.html, maintained by Dr. John Longino. Foraging workers of this species are usually covered by a thick layer of particles, as mentioned in the diagnosis.
<bibRefCitation id="1A1343C3FFACE37DFECCFB68838BDE50" author="Holldobler, B. &amp; Wilson, E. O." box="[340,656,1267,1294]" pageId="40" pageNumber="41" pagination="12 - 20" refId="ref61910" refString="Holldobler, B. &amp; Wilson, E. O. (1986). Soil-binding pilosity and camouflage in ants of the tribes Basicerotini and Stegomyrmecini. Zoomorphology, 106, 12 - 20. https: // doi. org / 10.1007 / BF 00311942" type="journal article" year="1986">Hölldobler &amp; Wilson (1986)</bibRefCitation>
speculate that the specialized double-layer pilosity acts with the layer of erect and clavate hairs capturing soil and litter particles from the environment and the layer of subdecumbent plumose hairs adhering them to the ants body. Individuals of
<taxonomicName id="B98245B1FFACE37DFCD7FAA082AFDE0A" authorityName="Brown &amp; Kempf" authorityYear="1960" box="[847,948,1339,1364]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="manni">
<emphasis id="4CF6E220FFACE37DFCD7FAA082AFDE0A" box="[847,948,1339,1364]" italics="true" pageId="40" pageNumber="41">B. manni</emphasis>
</taxonomicName>
present thanatosis and this behavior, added to the cryptic coloration presented by this species—a result of the already mentioned dense cover of soil particles and litter adhered to its integument—makes field observation of this dirt ant an ungrateful task.
</paragraph>
<paragraph id="7E3D3E32FFACE37DFF5FFA3C836BDDE3" blockId="40.[151,1437,151,2013]" pageId="40" pageNumber="41">
Colonies of
<taxonomicName id="B98245B1FFACE37DFED7FA3C80AFDE9E" authorityName="Brown &amp; Kempf" authorityYear="1960" box="[335,436,1447,1472]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="manni">
<emphasis id="4CF6E220FFACE37DFED7FA3C80AFDE9E" box="[335,436,1447,1472]" italics="true" pageId="40" pageNumber="41">B. manni</emphasis>
</taxonomicName>
are usually found nesting in decayed trunks at different levels of composition, in the leaflitter located at the base of trees.
<bibRefCitation id="1A1343C3FFACE37DFD90FA50825DDEB8" author="Holldobler, B. &amp; Wilson, E. O." box="[520,838,1483,1510]" pageId="40" pageNumber="41" pagination="12 - 20" refId="ref61910" refString="Holldobler, B. &amp; Wilson, E. O. (1986). Soil-binding pilosity and camouflage in ants of the tribes Basicerotini and Stegomyrmecini. Zoomorphology, 106, 12 - 20. https: // doi. org / 10.1007 / BF 00311942" type="journal article" year="1986">Wilson &amp; Hölldobler (1986)</bibRefCitation>
mention the collection of a colony nesting in a rotten fava bean of
<taxonomicName id="B98245B1FFACE37DFEB6FA74835BDD57" box="[302,576,1519,1545]" class="Magnoliopsida" family="Fabaceae" genus="Pentaclethra" kingdom="Plantae" order="Fabales" pageId="40" pageNumber="41" phylum="Tracheophyta" rank="species" species="macroloba">
<emphasis id="4CF6E220FFACE37DFEB6FA74835BDD57" box="[302,576,1519,1545]" italics="true" pageId="40" pageNumber="41">Pentaclethra macroloba</emphasis>
</taxonomicName>
partially buried in the leaf-litter. Ronald Garcia, field biologist at La Selva Biological Station in
<collectingCountry id="06957EA2FFACE37DFE1CF988831BDD73" box="[388,512,1555,1581]" name="Costa Rica" pageId="40" pageNumber="41">Costa Rica</collectingCountry>
, share the following data on a colony nesting in an abandoned nest of what appears to be a Meliponini bee (
<figureCitation id="E6B922B7FFACE37DFE3DF9AC80EADD0C" box="[421,497,1591,1618]" captionStart="FIGURE 16" captionStartId="41.[151,250,1112,1136]" captionTargetBox="[158,1430,525,1089]" captionTargetId="figure-182@41.[158,1430,525,1089]" captionTargetPageId="41" captionText="FIGURE 16. Basiceros manni colony in abandoned Meliponini nest at La Selva Biological Station, Costa Rica: Heredia; A. Exterior view of nest entrance, B. Detail of chamber with immatures below nest entrance, C. Brood chamber. Images: courtesy of Ronald Garcia." figureDoi="http://doi.org/10.5281/zenodo.6605999" httpUri="https://zenodo.org/record/6605999/files/figure.png" pageId="40" pageNumber="41">Fig 16</figureCitation>
), with the following report: “The nest was in a tree fork of a living tree, to more or less,
<quantity id="B97A93D7FFACE37DFF49F9C08003DD2B" box="[209,280,1627,1653]" metricMagnitude="-1" metricUnit="m" metricValue="4.0" pageId="40" pageNumber="41" unit="cm" value="40.0">40 cm</quantity>
from the ground. The nest got my attention because the entrance looks like some stingless bees, this entrance is unbelievable! It is in the Aerial Tram, in the foothill of the Braulio Carrillo Mountain,
<quantity id="B97A93D7FFACE37DFB75F9E58436DDC7" box="[1261,1325,1662,1689]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="40" pageNumber="41" unit="m" value="500.0">500m</quantity>
a.s.l. The size of the nest is a little more than a fist”.
</paragraph>
<paragraph id="7E3D3E32FFACE37CFF5FF95C8076DAAB" blockId="40.[151,1437,151,2013]" lastBlockId="41.[151,1437,151,501]" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41">
<bibRefCitation id="1A1343C3FFACE37DFF5FF95C831DDDBC" author="Holldobler, B. &amp; Wilson, E. O." box="[199,518,1735,1762]" pageId="40" pageNumber="41" pagination="12 - 20" refId="ref61910" refString="Holldobler, B. &amp; Wilson, E. O. (1986). Soil-binding pilosity and camouflage in ants of the tribes Basicerotini and Stegomyrmecini. Zoomorphology, 106, 12 - 20. https: // doi. org / 10.1007 / BF 00311942" type="journal article" year="1986">Wilson &amp; Hölldobler (1986)</bibRefCitation>
, studying three colonies in the laboratory for about a month, report that ovaries are maximally developed in younger workers, associated with the laying of trophic eggs. In contrast, the venom gland reservoir reaches its maximum size in older workers, associated with an increase in predatory activity. These ants exhibit an exceptionally simple behavioral repertoire (Wilson &amp; Hölldobler
<emphasis id="4CF6E220FFACE37DFC7DF8A8852ADC12" box="[997,1073,1843,1868]" italics="true" pageId="40" pageNumber="41">op. cit.</emphasis>
). There is a temporal division of labor, with young workers acting as nurses and older workers leaving the nest in search of food. Dr. Jack Longino reports on his website (see address above) that he found this species in lowland forests of
<collectingCountry id="06957EA2FFACE37DFB12F8E0841CDCCB" box="[1162,1287,1915,1941]" name="Costa Rica" pageId="40" pageNumber="41">Costa Rica</collectingCountry>
, usually with workers foraging alone at night or collected from litter samples submitted to a Winkler extractor. In addition, he observed a nest in a primary forest at Los Patos area in the Corcovado National Park, also in
<collectingCountry id="06957EA2FFACE37DFB00F858840EDC83" box="[1176,1301,1987,2013]" name="Costa Rica" pageId="40" pageNumber="41">Costa Rica</collectingCountry>
. The colony was on a trunk in an advanced state of decomposition at the base of a small tree, and the workers were scattered in several chambers. One chamber had almost all the immatures, in another some larvae and four empty shells of gastropods of the same species, with a long and pointed spiral (also see the natural history topic under
<taxonomicName id="B98245B1FFADE37CFAB7FF44801CDA43" baseAuthorityName="Smith" baseAuthorityYear="1858" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="singularis">
<emphasis id="4CF6E220FFADE37CFAB7FF44801CDA43" italics="true" pageId="41" pageNumber="42">Basiceros singularis</emphasis>
</taxonomicName>
). Dr. Longino comments that Dr. Ted Schultz (USNM) collected a colony in a primary forest area of humid tropical forest at La Selva Station. The contents of the nest included (in addition to workers, gynes, males and immatures) a variety of empty gastropod shells, various termite abdomens and other unidentified objects reminiscent of
<taxonomicName id="B98245B1FFADE37CFE93FEF48082DAD7" authorityName="Mayr" authorityYear="1861" box="[267,409,367,393]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="genus">
<emphasis id="4CF6E220FFADE37CFE93FEF48082DAD7" box="[267,409,367,393]" italics="true" pageId="41" pageNumber="42">Camponotus</emphasis>
</taxonomicName>
(
<taxonomicName id="B98245B1FFADE37CFE3FFEF48336DAD7" box="[423,557,367,393]" class="Insecta" family="Formicidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="family">Formicidae</taxonomicName>
: Formicinae) gasters. Unfortunately,
<bibRefCitation id="1A1343C3FFADE37CFC53FEF48407DAD4" author="Wilson, E. O. &amp; Holldobler, B." box="[971,1308,367,394]" pageId="41" pageNumber="42" pagination="70 - 84" refId="ref63455" refString="Wilson, E. O. &amp; Holldobler, B. (1986) Ecology and behavior of the Neotropical cryptobiotic ant Basiceros manni (Hymenoptera: Formicidae: Basicerotini). Insectes Sociaux, 33, 70 - 84. https: // doi. org / 10.1007 / BF 02224036" type="journal article" year="1986">Wilson &amp; Hölldoblers (1986)</bibRefCitation>
“cafeteria” experiments did not include land snails, and their captive colonies of
<taxonomicName id="B98245B1FFADE37CFC31FE088514DAF2" authorityName="Brown &amp; Kempf" authorityYear="1960" box="[937,1039,403,428]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="manni">
<emphasis id="4CF6E220FFADE37CFC31FE088514DAF2" box="[937,1039,403,428]" italics="true" pageId="41" pageNumber="42">B. manni</emphasis>
</taxonomicName>
accepted beetle larvae, centipedes, termites, and freshly killed
<taxonomicName id="B98245B1FFADE37CFE48FE2C8354DA8F" box="[464,591,439,465]" class="Agaricomycetes" family="Psathyrellaceae" genus="Drosophila" kingdom="Fungi" order="Agaricales" pageId="41" pageNumber="42" phylum="Basidiomycota" rank="genus">
<emphasis id="4CF6E220FFADE37CFE48FE2C8354DA8F" box="[464,591,439,465]" italics="true" pageId="41" pageNumber="42">Drosophila</emphasis>
</taxonomicName>
adults. Prey was offered directly to the larvae. The workers foraged alone and did not recruit.
</paragraph>
<caption id="2AFD6EBAFFADE37CFF0FFBC38056DFE6" ID-DOI="http://doi.org/10.5281/zenodo.6605999" ID-Zenodo-Dep="6605999" httpUri="https://zenodo.org/record/6605999/files/figure.png" pageId="41" pageNumber="42" startId="41.[151,250,1112,1136]" targetBox="[158,1430,525,1089]" targetPageId="41">
<paragraph id="7E3D3E32FFADE37CFF0FFBC38056DFE6" blockId="41.[151,1437,1112,1208]" pageId="41" pageNumber="42">
<emphasis id="4CF6E220FFADE37CFF0FFBC3803ADF2E" bold="true" box="[151,289,1112,1136]" pageId="41" pageNumber="42">FIGURE 16.</emphasis>
<taxonomicName id="B98245B1FFADE37CFEB1FBC280C8DF2E" authorityName="Brown &amp; Kempf" authorityYear="1960" box="[297,467,1113,1136]" class="Insecta" family="Formicidae" genus="Basiceros" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="manni">
<emphasis id="4CF6E220FFADE37CFEB1FBC280C8DF2E" box="[297,467,1113,1136]" italics="true" pageId="41" pageNumber="42">Basiceros manni</emphasis>
</taxonomicName>
colony in abandoned Meliponini nest at La Selva Biological Station, Costa Rica: Heredia; A. Exterior view of nest entrance, B. Detail of chamber with immatures below nest entrance, C. Brood chamber. Images: courtesy of Ronald Garcia.
</paragraph>
</caption>
<paragraph id="7E3D3E32FFADE37CFF5FFB7C845EDE17" blockId="41.[151,1437,1255,2038]" pageId="41" pageNumber="42">
This scenario of likely specialized predation on land snails was reinforced by Dr. Michael Branstetter. According to him, all colonies collected in
<collectingCountry id="06957EA2FFADE37CFDBBFA90838DDE7B" box="[547,662,1291,1317]" name="Nicaragua" pageId="41" pageNumber="42">Nicaragua</collectingCountry>
had empty gastropod shells. The shells appear to belong to a “pulmonated” zonitid mollusk (
<taxonomicName id="B98245B1FFADE37CFE3DFAB4830DDE14" authorityName="Morch" authorityYear="1864" box="[421,534,1327,1354]" class="Gastropoda" family="Zonitidae" kingdom="Animalia" order="Stylommatophora" pageId="41" pageNumber="42" phylum="Mollusca" rank="family">Zonitidae</taxonomicName>
), probably from the genus
<taxonomicName id="B98245B1FFADE37CFCDBFAB482CEDE17" authorityName="E. von Martens" authorityYear="1892" box="[835,981,1327,1353]" class="Gastropoda" family="Gastrodontidae" genus="Glyphyalinia" kingdom="Animalia" order="Stylommatophora" pageId="41" pageNumber="42" phylum="Mollusca" rank="genus">
<emphasis id="4CF6E220FFADE37CFCDBFAB482CEDE17" box="[835,981,1327,1353]" italics="true" pageId="41" pageNumber="42">Glyphyalinia</emphasis>
</taxonomicName>
(Dr. Jaime Jardim pers. comm.).
</paragraph>
</subSubSection>
<subSubSection id="36986DB9FFADE37FFF5FFAC985D1DA1F" lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42" type="materials_examined">
<paragraph id="7E3D3E32FFADE37FFF5FFAC985D1DA1F" blockId="41.[151,1437,1255,2038]" lastBlockId="42.[151,1437,150,322]" lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42">
<emphasis id="4CF6E220FFADE37CFF5FFAC980A8DE32" bold="true" box="[199,435,1362,1388]" pageId="41" pageNumber="42">Material examined.</emphasis>
<materialsCitation id="CEEA346FFFADE37CFE23FAC88096DECF" ID-GBIF-Occurrence="4029332440" collectingDate="1974-02-04" collectingDateMax="1991-10-14" collectingDateMin="1974-02-04" collectionCode="LACM, MCZ, T" collectorName="I. Perfecto" country="Costa Rica" latitude="10.433333" location="La Selva Biological Station" longLatPrecision="1298" longitude="-84.0" pageId="41" pageNumber="42" specimenCount="24" specimenCount-male="3" specimenCount-worker="21" stateProvince="Cartago">
<collectingCountry id="06957EA2FFADE37CFE23FAC88378DE33" box="[443,611,1363,1389]" name="Costa Rica" pageId="41" pageNumber="42">COSTA RICA</collectingCountry>
:
<collectingRegion id="BC46F0D0FFADE37CFDE8FAC883D6DE33" box="[624,717,1363,1389]" country="Costa Rica" name="Cartago" pageId="41" pageNumber="42">Cartago</collectingRegion>
:
<collectorName id="D3775BE4FFADE37CFD41FAC88258DE33" box="[729,835,1363,1389]" pageId="41" pageNumber="42">Turrialba</collectorName>
, summer of 1988,
<collectorName id="D3775BE4FFADE37CFB84FAC88589DE33" box="[1052,1170,1363,1389]" pageId="41" pageNumber="42">I. Perfecto</collectorName>
col., n. 17—n. 326567 (
<specimenCount id="6884F5BBFFADE37CFF38FAEC8009DECC" box="[160,274,1399,1426]" pageId="41" pageNumber="42" type="worker">2 workers</specimenCount>
) [
<collectionCode id="1893A6F7FFADE37CFEB1FAED8098DECF" box="[297,387,1398,1425]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="41" pageNumber="42" type="Museum">LACM</collectionCode>
];
</materialsCitation>
<materialsCitation id="CEEA346FFFADE37CFE0DFAEC8077DEEB" ID-GBIF-Occurrence="4029332411" collectingDate="1974-02-04" collectingDateMax="1991-10-14" collectingDateMin="1974-02-04" collectionCode="LACM, MCZ, T" collectorName="Talbot &amp; VanDeVender" country="Costa Rica" latitude="10.433333" location="La Selva Biological Station" longLatPrecision="1298" longitude="-84.0" pageId="41" pageNumber="42" specimenCount="24" specimenCount-male="3" specimenCount-worker="21" stateProvince="Cartago">
<collectingRegion id="BC46F0D0FFADE37CFE0DFAEC80E9DECF" box="[405,498,1399,1425]" country="Costa Rica" name="Heredia" pageId="41" pageNumber="42">Heredia</collectingRegion>
:
<location id="7B5D68E9FFADE37CFE66FAEC8223DECF" LSID="urn:lsid:plazi:treatment:F62B8F24FFA6E37FFF0FFAAB85D1DA1F:7B5D68E9FFADE37CFE66FAEC8223DECF" box="[510,824,1399,1425]" country="Costa Rica" latitude="10.433333" longLatPrecision="1298" longitude="-84.0" name="La Selva Biological Station" pageId="41" pageNumber="42" stateProvince="Cartago">La Selva Biological Station</location>
,
<date id="0A3C18F2FFADE37CFCDCFAEC82B6DECF" box="[836,941,1399,1425]" pageId="41" pageNumber="42" value="1974-02-04">
<collectingDate id="1A78E11AFFADE37CFCDCFAEC82B6DECF" box="[836,941,1399,1425]" pageId="41" pageNumber="42" value="1974-02-04">4.II.1974</collectingDate>
</date>
,
<collectorName id="D3775BE4FFADE37CFC21FAEC851BDECF" box="[953,1024,1399,1425]" pageId="41" pageNumber="42">Talbot</collectorName>
&amp; VanDeVender cols., n. 326565 (
<specimenCount id="6884F5BBFFADE37CFA16FAEC81E8DEE8" pageId="41" pageNumber="42" type="worker">3 workers</specimenCount>
) [
<collectionCode id="1893A6F7FFADE37CFE90FA018079DEEB" box="[264,354,1434,1461]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="41" pageNumber="42" type="Museum">LACM</collectionCode>
];
</materialsCitation>
<materialsCitation id="CEEA346FFFADE37CFEEBFA008507DEEB" ID-GBIF-Occurrence="4029332305" box="[371,1052,1434,1462]" collectingDate="1974-02-04" collectingDateMax="1991-10-14" collectingDateMin="1974-02-04" collectionCode="LACM, MCZ, T" collectorName="E. O. Wilson" country="Costa Rica" latitude="10.433333" location="La Selva Biological Station" longLatPrecision="1298" longitude="-84.0" pageId="41" pageNumber="42" specimenCount="24" specimenCount-male="3" specimenCount-worker="21" stateProvince="Cartago">
<date id="0A3C18F2FFADE37CFEEBFA0080F4DEE8" box="[371,495,1435,1462]" pageId="41" pageNumber="42" value="1985-03-24">
<collectingDate id="1A78E11AFFADE37CFEEBFA0080F4DEE8" box="[371,495,1435,1462]" pageId="41" pageNumber="42" value="1985-03-24">24.iii.1985</collectingDate>
</date>
,
<collectorName id="D3775BE4FFADE37CFE62FA008391DEEB" box="[506,650,1435,1461]" pageId="41" pageNumber="42">E. O. Wilson</collectorName>
col., n. 546485 (
<specimenCount id="6884F5BBFFADE37CFCDDFA0082ADDEE8" box="[837,950,1435,1462]" pageId="41" pageNumber="42" type="worker">2 workers</specimenCount>
) [
<collectionCode id="1893A6F7FFADE37CFC53FA01850BDEEB" box="[971,1040,1434,1461]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="41" pageNumber="42" type="Museum">MCZ</collectionCode>
];
</materialsCitation>
<materialsCitation id="CEEA346FFFADE37CFBBAFA0083EEDE87" ID-GBIF-Occurrence="4029332415" collectingDate="1974-02-04" collectingDateMax="1991-10-14" collectingDateMin="1974-02-04" collectionCode="LACM, MCZ, T" collectorName="B. Hölldobler &amp; E. O. Wilson" country="Costa Rica" latitude="10.433333" location="La Selva Biological Station" longLatPrecision="1298" longitude="-84.0" pageId="41" pageNumber="42" specimenCount="24" specimenCount-male="3" specimenCount-worker="21" stateProvince="Cartago">
same data,
<date id="0A3C18F2FFADE37CFB06FA008401DEE8" box="[1182,1306,1435,1462]" pageId="41" pageNumber="42" value="1985-03-26">
<collectingDate id="1A78E11AFFADE37CFB06FA008401DEE8" box="[1182,1306,1435,1462]" pageId="41" pageNumber="42" value="1985-03-26">26.iii.1985</collectingDate>
</date>
,
<collectorName id="D3775BE4FFADE37CFABDFA0081D8DE87" pageId="41" pageNumber="42">B. Hölldobler</collectorName>
&amp;
<collectorName id="D3775BE4FFADE37CFF7FFA248063DE87" box="[231,376,1471,1497]" pageId="41" pageNumber="42">E. O. Wilson</collectorName>
cols. (
<specimenCount id="6884F5BBFFADE37CFE5BFA24832DDE87" box="[451,566,1471,1497]" pageId="41" pageNumber="42" type="worker">2 workers</specimenCount>
,
<specimenCount id="6884F5BBFFADE37CFDD9FA248395DE84" box="[577,654,1471,1498]" pageId="41" pageNumber="42" type="male">1 male</specimenCount>
) [
<collectionCode id="1893A6F7FFADE37CFD3CFA2583F2DE87" box="[676,745,1470,1497]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="41" pageNumber="42" type="Museum">MCZ</collectionCode>
];
</materialsCitation>
<materialsCitation id="CEEA346FFFADE37CFD63FA248330DEA3" ID-GBIF-Occurrence="4029332401" collectingDate="1974-02-04" collectingDateMax="1991-10-14" collectingDateMin="1974-02-04" collectionCode="LACM, MCZ, T" collectorName="E. O. Wilson &amp; B. Holldobler" country="Costa Rica" latitude="10.433333" location="La Selva Biological Station" longLatPrecision="1298" longitude="-84.0" pageId="41" pageNumber="42" specimenCount="24" specimenCount-male="3" specimenCount-worker="21" stateProvince="Cartago">
same data,
<date id="0A3C18F2FFADE37CFCE1FA2482EEDE84" box="[889,1013,1471,1498]" pageId="41" pageNumber="42" value="1985-03-29">
<collectingDate id="1A78E11AFFADE37CFCE1FA2482EEDE84" box="[889,1013,1471,1498]" pageId="41" pageNumber="42" value="1985-03-29">29.iii.1985</collectingDate>
</date>
,
<collectorName id="D3775BE4FFADE37CFB99FA248581DE87" box="[1025,1178,1471,1497]" pageId="41" pageNumber="42">B. Hölldobler</collectorName>
&amp;
<collectorName id="D3775BE4FFADE37CFB27FA248452DE87" box="[1215,1353,1471,1497]" pageId="41" pageNumber="42">E.O. Wilson</collectorName>
col., n. 546486 (
<specimenCount id="6884F5BBFFADE37CFF63FA788076DEA3" box="[251,365,1507,1533]" pageId="41" pageNumber="42" type="worker">3 workers</specimenCount>
, 1 gyne) [
<collectionCode id="1893A6F7FFADE37CFE42FA798304DEA3" box="[474,543,1506,1533]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="41" pageNumber="42" type="Museum">MCZ</collectionCode>
];
</materialsCitation>
<materialsCitation id="CEEA346FFFADE37CFDA9FA788506DD37" ID-GBIF-Occurrence="4029332301" collectingDate="1974-02-04" collectingDateMax="1991-10-14" collectingDateMin="1974-02-04" collectionCode="LACM, MCZ, T" collectorName="E. O. Wilson &amp; B. Holldobler" country="Costa Rica" latitude="10.433333" location="La Selva Biological Station" longLatPrecision="1298" longitude="-84.0" pageId="41" pageNumber="42" specimenCount="24" specimenCount-male="3" specimenCount-worker="21" stateProvince="Cartago">
same data,
<date id="0A3C18F2FFADE37CFD36FA788231DEA0" box="[686,810,1507,1534]" pageId="41" pageNumber="42" value="1985-03-29">
<collectingDate id="1A78E11AFFADE37CFD36FA788231DEA0" box="[686,810,1507,1534]" pageId="41" pageNumber="42" value="1985-03-29">29.iii.1985</collectingDate>
</date>
,
<collectorName id="D3775BE4FFADE37CFCADFA7882D5DEA3" box="[821,974,1507,1533]" pageId="41" pageNumber="42">B. Hölldobler</collectorName>
&amp;
<collectorName id="D3775BE4FFADE37CFC6AFA788560DEA3" box="[1010,1147,1507,1533]" pageId="41" pageNumber="42">E.O. Wilson</collectorName>
cols. (
<specimenCount id="6884F5BBFFADE37CFB5DFA78842CDEA0" box="[1221,1335,1507,1534]" pageId="41" pageNumber="42" type="worker">2 workers</specimenCount>
) [
<collectionCode id="1893A6F7FFADE37CFAD4FA79848ADEA3" box="[1356,1425,1506,1533]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="41" pageNumber="42" type="Museum">MCZ</collectionCode>
]; same data,
<date id="0A3C18F2FFADE37CFE8CF99C8095DD7C" box="[276,398,1543,1570]" pageId="41" pageNumber="42" value="1985-11-19">
<collectingDate id="1A78E11AFFADE37CFE8CF99C8095DD7C" box="[276,398,1543,1570]" pageId="41" pageNumber="42" value="1985-11-19">19.xi.1985</collectingDate>
</date>
,
<collectorName id="D3775BE4FFADE37CFE01F99C8338DD7F" box="[409,547,1543,1569]" pageId="41" pageNumber="42">E.O. Wilson</collectorName>
col., (
<specimenCount id="6884F5BBFFADE37CFDF4F99C83CBDD7F" box="[620,720,1543,1569]" pageId="41" pageNumber="42" type="worker">1 worker</specimenCount>
,
<specimenCount id="6884F5BBFFADE37CFD43F99C823CDD7C" box="[731,807,1543,1570]" pageId="41" pageNumber="42" type="male">1 male</specimenCount>
) [
<collectionCode id="1893A6F7FFADE37CFCA5F99D8299DD7F" box="[829,898,1542,1569]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="41" pageNumber="42" type="Museum">MCZ</collectionCode>
]; same data,
<date id="0A3C18F2FFADE37CFB88F99C8571DD7F" box="[1040,1130,1543,1569]" pageId="41" pageNumber="42" value="1987-03">
<collectingDate id="1A78E11AFFADE37CFB88F99C8571DD7F" box="[1040,1130,1543,1569]" pageId="41" pageNumber="42" value="1987-03">iii.1987</collectingDate>
</date>
,
<collectorName id="D3775BE4FFADE37CFBECF99C841EDD7F" box="[1140,1285,1543,1569]" pageId="41" pageNumber="42">E. O. Wilson</collectorName>
col. (
<specimenCount id="6884F5BBFFADE37CFADCF99C81A2DD1B" pageId="41" pageNumber="42" type="worker">3 workers</specimenCount>
) [
<collectionCode id="1893A6F7FFADE37CFF57F9B1800FDD1B" box="[207,276,1578,1605]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="41" pageNumber="42" type="Museum">MCZ</collectionCode>
]; same data,
<date id="0A3C18F2FFADE37CFE3AF9B08305DD1B" box="[418,542,1579,1605]" pageId="41" pageNumber="42" value="1987-03-14">
<collectingDate id="1A78E11AFFADE37CFE3AF9B08305DD1B" box="[418,542,1579,1605]" pageId="41" pageNumber="42" value="1987-03-14">14.iii.1987</collectingDate>
</date>
,
<collectorName id="D3775BE4FFADE37CFDB1F9B083A8DD1B" box="[553,691,1579,1605]" pageId="41" pageNumber="42">E.O. Wilson</collectorName>
col., (
<specimenCount id="6884F5BBFFADE37CFD63F9B08244DD1B" box="[763,863,1579,1605]" pageId="41" pageNumber="42" type="worker">1 worker</specimenCount>
,
<specimenCount id="6884F5BBFFADE37CFCF2F9B082ACDD18" box="[874,951,1579,1606]" pageId="41" pageNumber="42" type="male">1 male</specimenCount>
) [
<collectionCode id="1893A6F7FFADE37CFC54F9B1850ADD1B" box="[972,1041,1578,1605]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="41" pageNumber="42" type="Museum">MCZ</collectionCode>
]; same data,
<date id="0A3C18F2FFADE37CFB38F9B08400DD1B" box="[1184,1307,1579,1605]" pageId="41" pageNumber="42" value="1988-03-23">
<collectingDate id="1A78E11AFFADE37CFB38F9B08400DD1B" box="[1184,1307,1579,1605]" pageId="41" pageNumber="42" value="1988-03-23">23.iii.1988</collectingDate>
</date>
,
<collectorName id="D3775BE4FFADE37CFABFF9B08487DD1B" box="[1319,1436,1579,1605]" pageId="41" pageNumber="42">S.P. Cover</collectorName>
col., n. 546463 (
<specimenCount id="6884F5BBFFADE37CFED2F9D480A0DD34" box="[330,443,1615,1642]" pageId="41" pageNumber="42" type="worker">2 workers</specimenCount>
) [
<collectionCode id="1893A6F7FFADE37CFE57F9D5830FDD37" box="[463,532,1614,1641]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="41" pageNumber="42" type="Museum">MCZ</collectionCode>
]; same data,
<geoCoordinate id="1BB658F5FFADE37CFD07F9D4821BDD37" box="[671,768,1615,1641]" degrees="10" direction="north" minutes="26" orientation="latitude" pageId="41" pageNumber="42" precision="925" value="10.433333">10°26N</geoCoordinate>
<geoCoordinate id="1BB658F5FFADE37CFC9EF9D48276DD37" box="[774,877,1615,1641]" degrees="84" direction="west" minutes="00" orientation="longitude" pageId="41" pageNumber="42" precision="925" value="-84.0">84°00W</geoCoordinate>
,
<date id="0A3C18F2FFADE37CFCEFF9D482F2DD37" box="[887,1001,1615,1641]" pageId="41" pageNumber="42" value="1991-10-14">
<collectingDate id="1A78E11AFFADE37CFCEFF9D482F2DD37" box="[887,1001,1615,1641]" pageId="41" pageNumber="42" value="1991-10-14">14.x.1991</collectingDate>
</date>
, J.
<collectionCode id="1893A6F7FFADE37CFB93F9D48506DD37" box="[1035,1053,1615,1641]" name="Tavera, Department of Geology and Geophysics" pageId="41" pageNumber="42">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="CEEA346FFFADE37CFBBFF9D48321DDD0" ID-GBIF-Occurrence="4029332351" collectionCode="INBIO-CRI" collectorName="Longino" country="Costa Rica" pageId="41" pageNumber="42" specimenCode="INBIO-CRI001237650" specimenCount="1" specimenCount-worker="1" stateProvince="Cartago">
<collectorName id="D3775BE4FFADE37CFBBFF9D4859DDD37" box="[1063,1158,1615,1641]" pageId="41" pageNumber="42">Longino</collectorName>
col., n. J.
<collectorName id="D3775BE4FFADE37CFB72F9D48456DD37" box="[1258,1357,1615,1641]" pageId="41" pageNumber="42">Longino</collectorName>
#3078- 5,
<specimenCode id="2E249649FFADE37CFF2CF9E880ACDDD0" box="[180,439,1651,1678]" collectionCode="INBIO-CRI" pageId="41" pageNumber="42">INBIO-CRI001237650</specimenCode>
(
<specimenCount id="6884F5BBFFADE37CFE52F9E88329DDD0" box="[458,562,1651,1678]" pageId="41" pageNumber="42" type="worker">1 worker</specimenCount>
)
</materialsCitation>
,
<materialsCitation id="CEEA346FFFADE37CFDDDF9E982ECDDEF" ID-GBIF-Occurrence="4029332311" collectingDate="1992-07-02" collectionCode="JTL, T, R" collectorName="Longino" country="Costa Rica" location="Puerto" longitude="8.53864" pageId="41" pageNumber="42" specimenCount="1" specimenCount-male="1" specimenCount-worker="1" stateProvince="Cartago">
[
<collectionCode id="1893A6F7FFADE37CFDD4F9E9839ADDD3" box="[588,641,1650,1677]" pageId="41" pageNumber="42">JTL</collectionCode>
]; same data,
<date id="0A3C18F2FFADE37CFC8AF9E8828FDDD3" box="[786,916,1651,1677]" pageId="41" pageNumber="42" value="1992-07-02">
<collectingDate id="1A78E11AFFADE37CFC8AF9E8828FDDD3" box="[786,916,1651,1677]" pageId="41" pageNumber="42" value="1992-07-02">02.vii.1992</collectingDate>
</date>
,
<collectionCode id="1893A6F7FFADE37CFC07F9E882AADDD3" box="[927,945,1651,1677]" name="Tavera, Department of Geology and Geophysics" pageId="41" pageNumber="42">T</collectionCode>
.
<collectionCode id="1893A6F7FFADE37CFC25F9E882C8DDD3" box="[957,979,1651,1677]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="41" pageNumber="42">R</collectionCode>
.
<collectorName id="D3775BE4FFADE37CFC46F9E88529DDD3" box="[990,1074,1651,1677]" pageId="41" pageNumber="42">Schultz</collectorName>
col., n. TRS920702-08, 538640 e 538643 (
<specimenCount id="6884F5BBFFADE37CFE8CF90C8060DDEF" box="[276,379,1687,1713]" pageId="41" pageNumber="42" type="worker">1 worker</specimenCount>
, 1 gyne,
<specimenCount id="6884F5BBFFADE37CFE7DF90C832FDDEC" box="[485,564,1687,1714]" pageId="41" pageNumber="42" type="male">1 male</specimenCount>
) [
<collectionCode id="1893A6F7FFADE37CFDD3F90D83BEDDEF" box="[587,677,1686,1713]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="41" pageNumber="42" type="Museum">USNM</collectionCode>
];
<location id="7B5D68E9FFADE37CFD21F90C8219DDEF" LSID="urn:lsid:plazi:treatment:F62B8F24FFA6E37FFF0FFAAB85D1DA1F:7B5D68E9FFADE37CFD21F90C8219DDEF" box="[697,770,1687,1713]" country="Costa Rica" longitude="8.53864" name="Puerto" pageId="41" pageNumber="42" stateProvince="Cartago">Puerto</location>
Viejo,
<date id="0A3C18F2FFADE37CFCCFF90C82C9DDEC" box="[855,978,1687,1714]" pageId="41" pageNumber="42" value="1965-08-03">
<collectingDate id="1A78E11AFFADE37CFCCFF90C82C9DDEC" box="[855,978,1687,1714]" pageId="41" pageNumber="42" value="1965-08-03">3.viii.1965</collectingDate>
</date>
,
<collectionCode id="1893A6F7FFADE37CFC78F90C82ECDDEF" box="[992,1015,1687,1713]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="41" pageNumber="42">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="CEEA346FFFADE37CFC63F90C8353DDA4" ID-GBIF-Occurrence="4029332344" collectingDate="1987-03-02" collectionCode="ANIC, LACM" collectorName="J. Hamton &amp; D. Olson" country="Costa Rica" elevation="50" location="Long." longitude="8.53864" municipality="Lat" pageId="41" pageNumber="42" specimenCode="DM029, CASENT0914888" specimenCount="6" specimenCount-worker="6" stateProvince="Cartago">
<collectorName id="D3775BE4FFADE37CFC63F90C8575DDEF" box="[1019,1134,1687,1713]" pageId="41" pageNumber="42">J. Hamton</collectorName>
col., (
<specimenCount id="6884F5BBFFADE37CFB22F90C8436DDEC" box="[1210,1325,1687,1714]" pageId="41" pageNumber="42" type="worker">3 workers</specimenCount>
) [
<collectionCode id="1893A6F7FFADE37CFADCF90D848ADDEF" box="[1348,1425,1686,1713]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="41" pageNumber="42">ANIC</collectionCode>
]; same data, n. 326566 (
<specimenCount id="6884F5BBFFADE37CFE03F9208314DD8B" box="[411,527,1723,1749]" pageId="41" pageNumber="42" type="worker">2 workers</specimenCount>
, 1 gyne) [
<collectionCode id="1893A6F7FFADE37CFD1AF92183C7DD8B" box="[642,732,1722,1749]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="41" pageNumber="42" type="Museum">LACM</collectionCode>
];
<collectingMunicipality id="9E59A448FFADE37CFD77F920820FDD8B" box="[751,788,1723,1749]" pageId="41" pageNumber="42">Lat</collectingMunicipality>
10.4333
<location id="7B5D68E9FFADE37CFC19F92082DFDD8B" LSID="urn:lsid:plazi:treatment:F62B8F24FFA6E37FFF0FFAAB85D1DA1F:7B5D68E9FFADE37CFC19F92082DFDD8B" box="[897,964,1723,1749]" municipality="Lat" name="Long." pageId="41" pageNumber="42">Long.</location>
-83.9833,
<quantity id="B97A93D7FFADE37CFBD8F920856FDD88" box="[1088,1140,1723,1750]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="41" pageNumber="42" unit="m" value="50.0">
<elevation id="F5AFD901FFADE37CFBD8F920856FDD88" box="[1088,1140,1723,1750]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="41" pageNumber="42" unit="m" value="50.0">50m</elevation>
</quantity>
,
<date id="0A3C18F2FFADE37CFB19F92085F4DD8B" box="[1153,1263,1723,1749]" pageId="41" pageNumber="42" value="1987-03-02">
<collectingDate id="1A78E11AFFADE37CFB19F92085F4DD8B" box="[1153,1263,1723,1749]" pageId="41" pageNumber="42" value="1987-03-02">2.iii.1987</collectingDate>
</date>
,
<collectorName id="D3775BE4FFADE37CFB64F9208478DD8B" box="[1276,1379,1723,1749]" pageId="41" pageNumber="42">D. Olson</collectorName>
col.,
<specimenCode id="2E249649FFADE37CFF0FF94481F6DDA7" box="[151,237,1759,1785]" collectionCode="DM" country="New Zealand" name="Dominion Museum" pageId="41" pageNumber="42">DM029</specimenCode>
/
<specimenCode id="2E249649FFADE37CFF6EF94480DEDDA7" box="[246,453,1759,1785]" collectionCode="CASENT" pageId="41" pageNumber="42">CASENT0914888</specimenCode>
(
<specimenCount id="6884F5BBFFADE37CFE42F944835BDDA4" box="[474,576,1759,1786]" pageId="41" pageNumber="42" type="worker">1 worker</specimenCount>
)
</materialsCitation>
<materialsCitation id="CEEA346FFFADE37CFDCAF94582E1DC40" ID-GBIF-Occurrence="4029332324" accessionNumber="BLF10423" collectingDate="2004-08-04" collectingDateMax="2004-08-14" collectingDateMin="2004-08-04" collectionCode="UCDC" collectorName="B. L. Fisher" country="Costa Rica" elevation="50" latitude="10.433333" location="La Selva Biological" longLatPrecision="1298" longitude="-84.0" pageId="41" pageNumber="42" specimenCode="CASENT0052769" specimenCount="1" specimenCount-worker="1" stateProvince="Cartago">
[
<collectionCode id="1893A6F7FFADE37CFDC0F94583B5DDA7" box="[600,686,1758,1785]" country="USA" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="41" pageNumber="42" type="Museum">UCDC</collectionCode>
];
<location id="7B5D68E9FFADE37CFD5BF94482BCDDA7" LSID="urn:lsid:plazi:treatment:F62B8F24FFA6E37FFF0FFAAB85D1DA1F:7B5D68E9FFADE37CFD5BF94482BCDDA7" box="[707,935,1759,1785]" country="Costa Rica" latitude="10.433333" longLatPrecision="1298" longitude="-84.0" name="La Selva Biological" pageId="41" pageNumber="42" stateProvince="Cartago">La Selva Biological</location>
Station-AntCourse 2004,
<geoCoordinate id="1BB658F5FFADE37CFB4CF944842EDDA7" box="[1236,1333,1759,1785]" degrees="10" direction="north" minutes="26" orientation="latitude" pageId="41" pageNumber="42" precision="925" value="10.433333">10°26N</geoCoordinate>
<geoCoordinate id="1BB658F5FFADE37CFAA7F944848CDDA7" box="[1343,1431,1759,1785]" degrees="84" direction="west" minutes="0" orientation="longitude" pageId="41" pageNumber="42" precision="925" value="-84.0">84°0W</geoCoordinate>
,
<quantity id="B97A93D7FFADE37CFF0FF89881D0DC40" box="[151,203,1795,1822]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="41" pageNumber="42" unit="m" value="50.0">
<elevation id="F5AFD901FFADE37CFF0FF89881D0DC40" box="[151,203,1795,1822]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="41" pageNumber="42" unit="m" value="50.0">50m</elevation>
</quantity>
,
<date id="0A3C18F2FFADE37CFF41F8988064DC43" box="[217,383,1795,1821]" pageId="41" pageNumber="42" value="2004-08-04" valueMax="2004-08-14" valueMin="2004-08-04">
<collectingDate id="1A78E11AFFADE37CFF41F8988064DC43" box="[217,383,1795,1821]" pageId="41" pageNumber="42" value="2004-08-04" valueMax="2004-08-14" valueMin="2004-08-04">414.viii.2004</collectingDate>
</date>
,
<collectorName id="D3775BE4FFADE37CFE15F8988301DC43" box="[397,538,1795,1821]" pageId="41" pageNumber="42">B. L. Fisher</collectorName>
, BLF10423/
<specimenCode id="2E249649FFADE37CFD3EF898826CDC40" box="[678,887,1795,1822]" collectionCode="CASENT" pageId="41" pageNumber="42">CASENT0052769</specimenCode>
(
<specimenCount id="6884F5BBFFADE37CFC14F89882E9DC40" box="[908,1010,1795,1822]" pageId="41" pageNumber="42" type="worker">1 worker</specimenCount>
)
</materialsCitation>
<materialsCitation id="CEEA346FFFADE37CFB9BF89982D1DC1F" ID-GBIF-Occurrence="4029332422" accessionNumber="BLF10423" collectingDate="1924-03" collectingDateMax="2010-03-07" collectingDateMin="1924-03" collectionCode="CAS, MCZ, USNM, MZSP, LACM, T" collectorName="W. M. Mann" country="Costa Rica" county="Around Guapiles" elevation="50" latitude="10.433333" location="Peninsula Osa" longLatPrecision="1298" longitude="-84.0" municipality="Rio Toro Amarillo" pageId="41" pageNumber="42" specimenCount="24" specimenCount-worker="24" stateProvince="Limon">
[
<collectionCode id="1893A6F7FFADE37CFB93F8998552DC43" box="[1035,1097,1794,1821]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="41" pageNumber="42" type="Museum">CAS</collectionCode>
].
<collectingRegion id="BC46F0D0FFADE37CFBC5F89885B3DC43" box="[1117,1192,1795,1821]" country="Costa Rica" name="Limon" pageId="41" pageNumber="42">Limón</collectingRegion>
(“Santa Clara”):
<collectorName id="D3775BE4FFADE37CFAEAF898805DDC1F" pageId="41" pageNumber="42">Colombiana Farm</collectorName>
:
<date id="0A3C18F2FFADE37CFED7F8BC80B3DC1F" box="[335,424,1831,1857]" pageId="41" pageNumber="42" value="1924-03">
<collectingDate id="1A78E11AFFADE37CFED7F8BC80B3DC1F" box="[335,424,1831,1857]" pageId="41" pageNumber="42" value="1924-03">iii.1924</collectingDate>
</date>
,
<collectorName id="D3775BE4FFADE37CFE2AF8BC8323DC1F" box="[434,568,1831,1857]" pageId="41" pageNumber="42">W.M. Mann</collectorName>
col., n. 546484 (
<specimenCount id="6884F5BBFFADE37CFD6BF8BC827FDC1C" box="[755,868,1831,1858]" pageId="41" pageNumber="42" type="worker">2 workers</specimenCount>
) [
<collectionCode id="1893A6F7FFADE37CFCE1F8BD82A5DC1F" box="[889,958,1830,1857]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="41" pageNumber="42" type="Museum">MCZ</collectionCode>
];
</materialsCitation>
<materialsCitation id="CEEA346FFFADE37CFC48F8BC8301DCAB" ID-GBIF-Occurrence="4029332373" accessionNumber="BLF10423" collectingDate="1924-03" collectingDateMax="2010-03-07" collectingDateMin="1924-03" collectionCode="CAS, MCZ, USNM, MZSP, LACM, T" collectorName="F. Nevermann" country="Costa Rica" county="Around Guapiles" elevation="50" latitude="10.433333" location="Peninsula Osa" longLatPrecision="1298" longitude="-84.0" municipality="Rio Toro Amarillo" pageId="41" pageNumber="42" specimenCount="24" specimenCount-worker="24" stateProvince="Limon">
same data (
<specimenCount id="6884F5BBFFADE37CFBD5F8BC85D7DC1F" box="[1101,1228,1831,1857]" pageId="41" pageNumber="42" type="worker">11 workers</specimenCount>
, 1 gyne) [
<collectionCode id="1893A6F7FFADE37CFAA0F8BD8489DC1F" box="[1336,1426,1830,1857]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="41" pageNumber="42" type="Museum">USNM</collectionCode>
]; Hamburg Farm: 1924,
<collectorName id="D3775BE4FFADE37CFE01F8D08323DC3B" box="[409,568,1867,1893]" pageId="41" pageNumber="42">F. Nevermann</collectorName>
col., n. 546487 (
<specimenCount id="6884F5BBFFADE37CFD63F8D0827BDC38" box="[763,864,1867,1894]" pageId="41" pageNumber="42" type="worker">1 worker</specimenCount>
) [
<collectionCode id="1893A6F7FFADE37CFCEDF8D182ACDC3B" box="[885,951,1866,1893]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="41" pageNumber="42" type="Museum">MCZ</collectionCode>
] (
<typeStatus id="A1398090FFADE37CFC56F8D0851FDC3B" box="[974,1028,1867,1893]" pageId="41" pageNumber="42">type</typeStatus>
?);
<collectingCounty id="975C46BEFFADE37CFBBBF8D085FDDC3B" box="[1059,1254,1867,1893]" pageId="41" pageNumber="42">Around Guapiles</collectingCounty>
,
<collectingMunicipality id="9E59A448FFADE37CFB6AF8D081D3DCD7" pageId="41" pageNumber="42">Rio Toro Amarillo</collectingMunicipality>
,
<date id="0A3C18F2FFADE37CFF4DF8F4809FDCD4" box="[213,388,1903,1930]" pageId="41" pageNumber="42" value="1966-02-25" valueMax="1966-03-09" valueMin="1966-02-25">
<collectingDate id="1A78E11AFFADE37CFF4DF8F4809FDCD4" box="[213,388,1903,1930]" pageId="41" pageNumber="42" value="1966-02-25" valueMax="1966-03-09" valueMin="1966-02-25">25.ii9.iii.1966</collectingDate>
</date>
,
<collectorName id="D3775BE4FFADE37CFE08F8F48300DCD7" box="[400,539,1903,1929]" pageId="41" pageNumber="42">W.L. Brown</collectorName>
col., n. 546481 (
<specimenCount id="6884F5BBFFADE37CFD47F8F4824ADCD4" box="[735,849,1903,1930]" pageId="41" pageNumber="42" type="worker">3 workers</specimenCount>
) [
<collectionCode id="1893A6F7FFADE37CFCF0F8F582B6DCD7" box="[872,941,1902,1929]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="41" pageNumber="42" type="Museum">MCZ</collectionCode>
]; same data, n. 546480 (
<specimenCount id="6884F5BBFFADE37CFB5BF8F4842EDCD4" box="[1219,1333,1903,1930]" pageId="41" pageNumber="42" type="worker">3 workers</specimenCount>
) [
<collectionCode id="1893A6F7FFADE37CFAD4F8F5848ADCD7" box="[1356,1425,1902,1929]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="41" pageNumber="42" type="Museum">MCZ</collectionCode>
]; same data (
<specimenCount id="6884F5BBFFADE37CFE8FF8088091DCF0" box="[279,394,1939,1966]" pageId="41" pageNumber="42" type="worker">2 workers</specimenCount>
) [
<collectionCode id="1893A6F7FFADE37CFE38F80980E9DCF3" box="[416,498,1938,1965]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="41" pageNumber="42" type="Museum">MZSP</collectionCode>
]; same data, n. 326564 (
<specimenCount id="6884F5BBFFADE37CFC9EF8088270DCF0" box="[774,875,1939,1966]" pageId="41" pageNumber="42" type="worker">1 worker</specimenCount>
) [
<collectionCode id="1893A6F7FFADE37CFC19F80982C0DCF3" box="[897,987,1938,1965]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="41" pageNumber="42" type="Museum">LACM</collectionCode>
];
<location id="7B5D68E9FFADE37CFC75F8088589DCF3" LSID="urn:lsid:plazi:treatment:F62B8F24FFA6E37FFF0FFAAB85D1DA1F:7B5D68E9FFADE37CFC75F8088589DCF3" box="[1005,1170,1939,1965]" county="Around Guapiles" municipality="Rio Toro Amarillo" name="Peninsula Osa" pageId="41" pageNumber="42" stateProvince="Limon">Península Osa</location>
:
<location id="7B5D68E9FFADE37CFB05F8088407DCF3" LSID="urn:lsid:plazi:treatment:F62B8F24FFA6E37FFF0FFAAB85D1DA1F:7B5D68E9FFADE37CFB05F8088407DCF3" box="[1181,1308,1939,1965]" county="Around Guapiles" municipality="Rio Toro Amarillo" name="Corcovado" pageId="41" pageNumber="42" stateProvince="Limon">Corcovado</location>
:
<location id="7B5D68E9FFADE37CFABFF808848CDCF3" LSID="urn:lsid:plazi:treatment:F62B8F24FFA6E37FFF0FFAAB85D1DA1F:7B5D68E9FFADE37CFABFF808848CDCF3" box="[1319,1431,1939,1965]" county="Around Guapiles" municipality="Rio Toro Amarillo" name="Los Patos" pageId="41" pageNumber="42" stateProvince="Limon">Los Patos</location>
,
<date id="0A3C18F2FFADE37CFF0FF82C8008DC8F" box="[151,275,1975,2001]" pageId="41" pageNumber="42" value="1981-03-01">
<collectingDate id="1A78E11AFFADE37CFF0FF82C8008DC8F" box="[151,275,1975,2001]" pageId="41" pageNumber="42" value="1981-03-01">01.iii.1981</collectingDate>
</date>
,
<collectorName id="D3775BE4FFADE37CFE87F82C808CDC8F" box="[287,407,1975,2001]" pageId="41" pageNumber="42">J. Longino</collectorName>
col., n. 546482 (
<specimenCount id="6884F5BBFFADE37CFDC1F82C83A5DC8C" box="[601,702,1975,2002]" pageId="41" pageNumber="42" type="worker">1 worker</specimenCount>
) [
<collectionCode id="1893A6F7FFADE37CFD4CF82D8202DC8F" box="[724,793,1974,2001]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="41" pageNumber="42" type="Museum">MCZ</collectionCode>
];
<collectingRegion id="BC46F0D0FFADE37CFCB4F82C82B0DC8F" box="[812,939,1975,2001]" country="Costa Rica" name="Puntarenas" pageId="41" pageNumber="42">Puntarenas</collectingRegion>
:
<locationDeviation id="6F8F2710FFADE37CFC2EF82C85EFDC8F" box="[950,1268,1975,2002]" location="Puerto Jimenez" pageId="41" pageNumber="42">
<quantity id="B97A93D7FFADE37CFC2EF82C82E6DC8F" box="[950,1021,1975,2002]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="41" pageNumber="42" unit="km" value="15.0">15 km</quantity>
SSW Puerto Jimenez
</locationDeviation>
, Lat. 8.407665
<location id="7B5D68E9FFADE37CFF0FF84081C1DCAB" LSID="urn:lsid:plazi:treatment:F62B8F24FFA6E37FFF0FFAAB85D1DA1F:7B5D68E9FFADE37CFF0FF84081C1DCAB" box="[151,218,2011,2037]" county="Around Guapiles" municipality="Rio Toro Amarillo" name="Long." pageId="41" pageNumber="42" stateProvince="Limon">Long.</location>
-83.32788,
<date id="0A3C18F2FFADE37CFEFCF84080FBDCAB" box="[356,480,2011,2037]" pageId="41" pageNumber="42" value="2010-03-07">
<collectingDate id="1A78E11AFFADE37CFEFCF84080FBDCAB" box="[356,480,2011,2037]" pageId="41" pageNumber="42" value="2010-03-07">07.iii.2010</collectingDate>
</date>
, J.
<collectionCode id="1893A6F7FFADE37CFD90F8408301DCAB" box="[520,538,2011,2037]" name="Tavera, Department of Geology and Geophysics" pageId="41" pageNumber="42">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="CEEA346FFFADE37CFDBFF840843BDCA8" ID-GBIF-Occurrence="4029332353" box="[551,1312,2011,2038]" collectionCode="CASENT" collectorName="Longino" country="Costa Rica" pageId="41" pageNumber="42" specimenCode="CASENT0634078" specimenCount="1" specimenCount-worker="1" stateProvince="Limon">
<collectorName id="D3775BE4FFADE37CFDBFF840839DDCAB" box="[551,646,2011,2037]" pageId="41" pageNumber="42">Longino</collectorName>
col., n. J.
<collectorName id="D3775BE4FFADE37CFC98F8408245DCA8" box="[768,862,2011,2038]" pageId="41" pageNumber="42">Longino</collectorName>
#6901.45/
<specimenCode id="2E249649FFADE37CFC57F8408584DCA8" box="[975,1183,2011,2038]" collectionCode="CASENT" pageId="41" pageNumber="42">CASENT0634078</specimenCode>
(
<specimenCount id="6884F5BBFFADE37CFB2BF8408402DCA8" box="[1203,1305,2011,2038]" pageId="41" pageNumber="42" type="worker">1 worker</specimenCount>
)
</materialsCitation>
<materialsCitation id="CEEA346FFFADE37FFAB2F8418302DBEF" ID-GBIF-Occurrence="4029332408" collectionCode="JTL" country="Costa Rica" lastPageId="42" lastPageNumber="43" location="Gary Umphrey" municipality="No" pageId="41" pageNumber="42" specimenCount="1" stateProvince="Limon">
[
<collectionCode id="1893A6F7FFADE37CFAA9F841847DDCAB" box="[1329,1382,2010,2037]" pageId="41" pageNumber="42">JTL</collectionCode>
];
<collectingMunicipality id="9E59A448FFADE37CFAE2F8408487DCAB" box="[1402,1436,2011,2037]" pageId="41" pageNumber="42">No</collectingMunicipality>
label (according to
<location id="7B5D68E9FFAEE37FFEF5FF0C8302DBEF" LSID="urn:lsid:plazi:treatment:F62B8F24FFA6E37FFF0FFAAB85D1DA1F:7B5D68E9FFAEE37FFEF5FF0C8302DBEF" box="[365,537,151,177]" country="Costa Rica" municipality="No" name="Gary Umphrey" pageId="42" pageNumber="43" stateProvince="Limon">Gary Umphrey</location>
</materialsCitation>
:
<materialsCitation id="CEEA346FFFAEE37FFDBBFF0C8482DBEF" ID-GBIF-Occurrence="4029332369" box="[547,1433,150,178]" collectionCode="MCZ" collectorName="La Selva &amp; E. O. Wilson" country="Costa Rica" location="COSTA RICA" pageId="42" pageNumber="43" specimenCount="2" specimenCount-male="1" specimenCount-worker="1">
<collectingCountry id="06957EA2FFAEE37FFDBBFF0C83D3DBEF" box="[547,712,151,177]" name="Costa Rica" pageId="42" pageNumber="43">COSTA RICA</collectingCountry>
:
<collectorName id="D3775BE4FFAEE37FFD4AFF0C822DDBEF" box="[722,822,151,177]" pageId="42" pageNumber="43">La Selva</collectorName>
/III-14-87,
<collectorName id="D3775BE4FFAEE37FFC37FF0C8525DBEF" box="[943,1086,151,177]" pageId="42" pageNumber="43">E. O. Wilson</collectorName>
col. (
<specimenCount id="6884F5BBFFAEE37FFBE6FF0C85F9DBEF" box="[1150,1250,151,177]" pageId="42" pageNumber="43" type="worker">1 worker</specimenCount>
,
<specimenCount id="6884F5BBFFAEE37FFB74FF0C8423DBEC" box="[1260,1336,151,178]" pageId="42" pageNumber="43" type="male">1 male</specimenCount>
) [
<collectionCode id="1893A6F7FFAEE37FFAD5FF0D8489DBEF" box="[1357,1426,150,177]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="42" pageNumber="43" type="Museum">MCZ</collectionCode>
]
</materialsCitation>
.
<materialsCitation id="CEEA346FFFAEE37FFF0FFF208086DBA7" ID-GBIF-Occurrence="4029332419" collectingDate="2011-04-17" collectingDateMax="2011-04-20" collectingDateMin="2011-04-17" collectionCode="MGB" collectorName="El Porton &amp; Lat. &amp; Long." country="Nicaragua" elevation="750" location="Boaco" pageId="42" pageNumber="43" specimenCode="MGB1723" specimenCount="5" specimenCount-larva="1" specimenCount-pupa="1" specimenCount-worker="3" stateProvince="Boaco">
<collectingCountry id="06957EA2FFAEE37FFF0FFF20805FDB8B" box="[151,324,187,213]" name="Nicaragua" pageId="42" pageNumber="43">NICARAGUA</collectingCountry>
:
<collectingRegion id="BC46F0D0FFAEE37FFED7FF208080DB8B" box="[335,411,187,213]" country="Nicaragua" name="Boaco" pageId="42" pageNumber="43">Boaco</collectingRegion>
:
<quantity id="B97A93D7FFAEE37FFE3EFF2080C3DB8B" box="[422,472,187,213]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="42" pageNumber="43" unit="km" value="8.0">8km</quantity>
LSE de
<collectorName id="D3775BE4FFAEE37FFDA4FF2083B0DB8B" box="[572,683,187,213]" pageId="42" pageNumber="43">El Porton</collectorName>
,
<collectorName id="D3775BE4FFAEE37FFD2FFF2083FFDB8B" box="[695,740,187,213]" pageId="42" pageNumber="43">Lat.</collectorName>
12.58465
<collectorName id="D3775BE4FFAEE37FFCC5FF208284DB8B" box="[861,927,187,213]" pageId="42" pageNumber="43">Long.</collectorName>
-85.4656,
<quantity id="B97A93D7FFAEE37FFB82FF218546DB8B" box="[1050,1117,186,213]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="42" pageNumber="43" unit="m" value="750.0">
<elevation id="F5AFD901FFAEE37FFB82FF218546DB8B" box="[1050,1117,186,213]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="42" pageNumber="43" unit="m" value="750.0">750m</elevation>
</quantity>
, n.
<specimenCode id="2E249649FFAEE37FFB1EFF2085E5DB8B" box="[1158,1278,187,213]" collectionCode="MGB" country="Spain" httpUri="http://grbio.org/cool/d3z2-v7v8" name="Museo de Geologia (del Seminario Diocesano) de Barcelona" pageId="42" pageNumber="43">MGB1723</specimenCode>
(
<specimenCount id="6884F5BBFFAEE37FFA97FF208499DB8B" box="[1295,1410,187,213]" pageId="42" pageNumber="43" type="worker">3 workers</specimenCount>
,
<specimenCount id="6884F5BBFFAEE37FFA16FF2081CBDBA7" pageId="42" pageNumber="43" type="pupa">1 pupa</specimenCount>
,
<specimenCount id="6884F5BBFFAEE37FFF42FF448033DBA4" box="[218,296,223,250]" pageId="42" pageNumber="43" type="larva">1 larva</specimenCount>
) [
<collectionCode id="1893A6F7FFAEE37FFEA4FF458089DBA7" box="[316,402,222,249]" pageId="42" pageNumber="43">MBPC</collectionCode>
];
</materialsCitation>
<materialsCitation id="CEEA346FFFAEE37FFE3BFF44823BDA43" ID-GBIF-Occurrence="4029332436" collectingDate="2011-04-17" collectingDateMax="2011-04-20" collectingDateMin="2011-04-17" collectionCode="MBPC" collectorName="Lat. &amp; Long." country="Nicaragua" elevation="740" location="4 km NE de Cuapa" pageId="42" pageNumber="43" specimenCode="MGB1735" specimenCount="5" specimenCount-worker="5" stateProvince="Chontales">
<collectingRegion id="BC46F0D0FFAEE37FFE3BFF44830CDBA7" box="[419,535,223,249]" country="Nicaragua" name="Chontales" pageId="42" pageNumber="43">Chontales</collectingRegion>
:
<location id="7B5D68E9FFAEE37FFDB9FF4483F5DBA7" LSID="urn:lsid:plazi:treatment:F62B8F24FFA6E37FFF0FFAAB85D1DA1F:7B5D68E9FFAEE37FFDB9FF4483F5DBA7" box="[545,750,223,249]" country="Nicaragua" name="4 km NE de Cuapa" pageId="42" pageNumber="43" stateProvince="Chontales">
<quantity id="B97A93D7FFAEE37FFDB9FF448348DBA7" box="[545,595,223,249]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="42" pageNumber="43" unit="km" value="4.0">4km</quantity>
NE de Cuapa
</location>
,
<date id="0A3C18F2FFAEE37FFD60FF448274DBA7" box="[760,879,223,249]" pageId="42" pageNumber="43" value="2011-04-20">
<collectingDate id="1A78E11AFFAEE37FFD60FF448274DBA7" box="[760,879,223,249]" pageId="42" pageNumber="43" value="2011-04-20">20.iv.2011</collectingDate>
</date>
,
<collectorName id="D3775BE4FFAEE37FFCE2FF4482BCDBA7" box="[890,935,223,249]" pageId="42" pageNumber="43">Lat.</collectorName>
12.2903
<collectorName id="D3775BE4FFAEE37FFB95FF448554DBA7" box="[1037,1103,223,249]" pageId="42" pageNumber="43">Long.</collectorName>
-85.35436,
<quantity id="B97A93D7FFAEE37FFB4CFF44840CDBA7" box="[1236,1303,223,249]" metricMagnitude="2" metricUnit="m" metricValue="7.4" pageId="42" pageNumber="43" unit="m" value="740.0">
<elevation id="F5AFD901FFAEE37FFB4CFF44840CDBA7" box="[1236,1303,223,249]" metricMagnitude="2" metricUnit="m" metricValue="7.4" pageId="42" pageNumber="43" unit="m" value="740.0">740m</elevation>
</quantity>
,
<date id="0A3C18F2FFAEE37FFAB8FF448483DBA7" box="[1312,1432,223,249]" pageId="42" pageNumber="43" value="2011-04-17">
<collectingDate id="1A78E11AFFAEE37FFAB8FF448483DBA7" box="[1312,1432,223,249]" pageId="42" pageNumber="43" value="2011-04-17">17.iv.2011</collectingDate>
</date>
, n.
<specimenCode id="2E249649FFAEE37FFF2DFE998036DA43" box="[181,301,258,285]" collectionCode="MGB" country="Spain" httpUri="http://grbio.org/cool/d3z2-v7v8" name="Museo de Geologia (del Seminario Diocesano) de Barcelona" pageId="42" pageNumber="43">MGB1735</specimenCode>
(
<specimenCount id="6884F5BBFFAEE37FFEA7FE9980AFDA43" box="[319,436,258,285]" pageId="42" pageNumber="43" type="worker">5 workers</specimenCount>
, a pupa, some larvae) [
<collectionCode id="1893A6F7FFAEE37FFD27FE99820EDA43" box="[703,789,258,285]" pageId="42" pageNumber="43">MBPC</collectionCode>
];
</materialsCitation>
<collectionCode id="1893A6F7FFAEE37FFCB2FE988265DA43" box="[810,894,259,285]" pageId="42" pageNumber="43">RAAN</collectionCode>
:
<materialsCitation id="CEEA346FFFAEE37FFC12FE988392DA1F" ID-GBIF-Occurrence="4029332431" collectingDate="2011-05-07" collectionCode="LLAMA" country="Nicaragua" county="Parque Nacional Cerro Saslaya" elevation="310" location="Long." municipality="Lat." pageId="42" pageNumber="43" specimenCount="1" stateProvince="Chontales">
<collectingCounty id="975C46BEFFAEE37FFC12FE9885EBDA43" box="[906,1264,259,285]" pageId="42" pageNumber="43">Parque Nacional Cerro Saslaya</collectingCounty>
,
<collectingMunicipality id="9E59A448FFAEE37FFB66FE988430DA43" box="[1278,1323,259,285]" pageId="42" pageNumber="43">Lat.</collectingMunicipality>
13.77065
<location id="7B5D68E9FFAEE37FFF0FFEBC81C1DA1F" LSID="urn:lsid:plazi:treatment:F62B8F24FFA6E37FFF0FFAAB85D1DA1F:7B5D68E9FFAEE37FFF0FFEBC81C1DA1F" box="[151,218,295,321]" country="Nicaragua" county="Parque Nacional Cerro Saslaya" municipality="Lat." name="Long." pageId="42" pageNumber="43" stateProvince="Chontales">Long.</location>
-08498018,
<quantity id="B97A93D7FFAEE37FFEF0FEBC80B1DA1F" box="[360,426,295,321]" metricMagnitude="2" metricUnit="m" metricValue="3.1" pageId="42" pageNumber="43" unit="m" value="310.0">
<elevation id="F5AFD901FFAEE37FFEF0FEBC80B1DA1F" box="[360,426,295,321]" metricMagnitude="2" metricUnit="m" metricValue="3.1" pageId="42" pageNumber="43" unit="m" value="310.0">310m</elevation>
</quantity>
,
<date id="0A3C18F2FFAEE37FFE2DFEBC830CDA1F" box="[437,535,295,321]" pageId="42" pageNumber="43" value="2011-05-07">
<collectingDate id="1A78E11AFFAEE37FFE2DFEBC830CDA1F" box="[437,535,295,321]" pageId="42" pageNumber="43" value="2011-05-07">7.v.2011</collectingDate>
</date>
,
<collectionCode id="1893A6F7FFAEE37FFDBBFEBC8392DA1F" box="[547,649,295,321]" pageId="42" pageNumber="43">LLAMA</collectionCode>
</materialsCitation>
#
<materialsCitation id="CEEA346FFFAEE37FFD0FFEBC8560DA1C" ID-GBIF-Occurrence="4029332336" box="[663,1147,295,322]" collectionCode="CASENT" country="Nicaragua" pageId="42" pageNumber="43" specimenCode="CASENT0628761" specimenCount="1" specimenCount-worker="1" stateProvince="Chontales">
Wa-D-02-44,
<specimenCode id="2E249649FFAEE37FFCB7FEBC82E5DA1F" box="[815,1022,295,321]" collectionCode="CASENT" pageId="42" pageNumber="43">CASENT0628761</specimenCode>
(
<specimenCount id="6884F5BBFFAEE37FFB88FEBC856FDA1C" box="[1040,1140,295,322]" pageId="42" pageNumber="43" type="worker">1 worker</specimenCount>
)
</materialsCitation>
[
<collectionCode id="1893A6F7FFAEE37FFB12FEBD85A4DA1F" box="[1162,1215,294,321]" pageId="42" pageNumber="43">JTL</collectionCode>
].
</paragraph>
</subSubSection>
</treatment>
</document>