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<document id="BDE6AF2BCF7091A1825C4CD4F07D462E" ID-CLB-Dataset="7385" ID-DOI="10.11646/zootaxa.4951.3.5" ID-GBIF-Dataset="caee27e5-0786-4e95-84fb-ee5394fa79e3" ID-ISSN="1175-5326" ID-Zenodo-Dep="4668365" ID-ZooBank="9BEB98F0-DEA7-4131-8E37-DFC4D5DFCB7B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617780432061" checkinUser="plazi" docAuthor="Bogorodsky, Sergey V., Zajonz, Uwe, Saeed, Fouad N. &amp; Weigmann, Simon" docDate="2021" docId="03B851366641FFADFF6C10AAE310ECD5" docLanguage="en" docName="zootaxa.4951.3.5.pdf" docOrigin="Zootaxa 4951 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhynchobatus australiae Whitley 1939" docType="treatment" docVersion="7" lastPageNumber="519" masterDocId="FF81294E6645FFA5FFFB1531E100ED7B" masterDocTitle="Notes on batoid fishes of the Socotra Archipelago (north-western Indian Ocean) with four new records" masterLastPageNumber="528" masterPageNumber="511" pageNumber="515" updateTime="1698936112385" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="61A5A2574885F961087F25811BD6B102">Notes on batoid fishes of the Socotra Archipelago (north-western Indian Ocean) with four new records</mods:title>
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<mods:namePart id="853CE425BEBB2D7E840DDD5400A498BA">Bogorodsky, Sergey V.</mods:namePart>
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<mods:namePart id="0FAADFBF78EECD6D809678B0DC8B9186">Zajonz, Uwe</mods:namePart>
<mods:affiliation id="E12E415FEB882C5825F88EDABB870B9A">Senckenberg oesearch fns ṫẚṫuṫe and Museum of ka ṫure (SMc) I Sec ṫẚon fch ṫhyology &amp; Senckenberg B ẚodẚversẚṫy and Cl ẚmaṫe oesearch Cen ṫre (SB ẚh-c) I B ẚogeography-Senckenberganlage 2 RI SMP 2 R crankfur ṫ am Ma ẚnI GermanyK</mods:affiliation>
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<mods:namePart id="02AB2FB1BEC00076FF944438BB2B710B">Saeed, Fouad N.</mods:namePart>
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<mods:namePart id="4290CC32416983B6641B4E37A694AE02">Weigmann, Simon</mods:namePart>
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<mods:date id="CE621EE67D9DAE85B1BDCD59DBE0E003">2021</mods:date>
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<mods:number id="894BC0A076B9C73982D20D756DDCAFA1">4951</mods:number>
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<treatment id="03B851366641FFADFF6C10AAE310ECD5" ID-DOI="http://doi.org/10.5281/zenodo.4685067" ID-GBIF-Taxon="181003439" ID-Zenodo-Dep="4685067" LSID="urn:lsid:plazi:treatment:03B851366641FFADFF6C10AAE310ECD5" httpUri="http://treatment.plazi.org/id/03B851366641FFADFF6C10AAE310ECD5" lastPageId="8" lastPageNumber="519" pageId="4" pageNumber="515">
<subSubSection id="C30BB3AB6641FFA1FF6C10AAE089E8A1" pageId="4" pageNumber="515" type="nomenclature">
<paragraph id="8BAEE0206641FFA1FF6C10AAE378E8CD" blockId="4.[151,632,1435,1535]" box="[151,632,1435,1462]" pageId="4" pageNumber="515">
<heading id="D0E6574C6641FFA1FF6C10AAE378E8CD" bold="true" box="[151,632,1435,1462]" fontSize="11" level="1" pageId="4" pageNumber="515" reason="1">
<taxonomicName id="4C119BA36641FFA1FF6C10AAE378E8CD" authority="Whitley, 1939" authorityName="Whitley" authorityYear="1939" box="[151,632,1435,1462]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhynchobatus" kingdom="Animalia" order="Rajiformes" pageId="4" pageNumber="515" phylum="Chordata" rank="species" species="australiae">
<emphasis id="B9653C326641FFA1FF6C10AAE378E8CD" bold="true" box="[151,632,1435,1462]" pageId="4" pageNumber="515">
<emphasis id="B9653C326641FFA1FF6C10AAE0C6E8CD" bold="true" box="[151,454,1435,1462]" italics="true" pageId="4" pageNumber="515">Rhynchobatus australiae</emphasis>
<bibRefCitation id="EF809DD16641FFA1FE3710ADE378E8CD" author="Whitley, G. P." box="[460,632,1436,1462]" pageId="4" pageNumber="515" pagination="227 - 262" refId="ref13266" refString="Whitley, G. P. (1939) Taxonomic notes on sharks and rays. Aus tralẚan Zoolog ẚst, 9 (3), 227 - 262." type="journal article" year="1939">Whitley, 1939</bibRefCitation>
</emphasis>
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</heading>
</paragraph>
<paragraph id="8BAEE0206641FFA1FF6C10F1E089E8A1" blockId="4.[151,632,1435,1535]" box="[151,393,1472,1498]" pageId="4" pageNumber="515">Bottlenose wedgefish</paragraph>
</subSubSection>
<subSubSection id="C30BB3AB6641FFA1FF6C10D5E489EB3B" pageId="4" pageNumber="515" type="description">
<paragraph id="8BAEE0206641FFA1FF6C10D5E05DE885" blockId="4.[151,632,1435,1535]" box="[151,349,1508,1535]" pageId="4" pageNumber="515">
<figureCitation id="132AFCA56641FFA1FF6C10D5E02FE884" box="[151,303,1508,1535]" captionStart="FIGURE 5" captionStartId="6.[151,250,1897,1921]" captionTargetBox="[189,1399,181,1873]" captionTargetId="figure-23@6.[189,1399,181,1873]" captionTargetPageId="6" captionText="FIGURE 5. ohynchobaṫus spp., dorsal view, fresh colouration. A: oK ausṫralẚae, SMF uncatalogued, female, 73.5 cm TL, fish market in Hadibo, Socotra Island; B: oK ausṫralẚae, SMF uncatalogued, male, 106.5 cm TL, Ras Hawlaf, Socotra Island; C: oK djẚddensẚs, female, 130 cm TL, Socotra Island; D: oK laevẚs, SMF uncatalogued, female, 115 cm TL, Qatar, the Gulf. Photos by S.V. Bogorodsky &amp; U. Zajonz (A, B and D), R. Bonfil (C)." figureDoi="http://doi.org/10.5281/zenodo.4668381" httpUri="https://zenodo.org/record/4668381/files/figure.png" pageId="4" pageNumber="515">Figs. 5A &amp; B</figureCitation>
,
<figureCitation id="132AFCA56641FFA1FEC010D5E05DE885" box="[315,349,1508,1534]" captionStart="FIGURE 6" captionStartId="7.[151,250,1964,1988]" captionTargetBox="[151,1437,1497,1941]" captionTargetId="figure-686@7.[151,1437,1497,1941]" captionTargetPageId="7" captionText="FIGURE 6. ohynchobaṫus spp., underside of snout. A: oK ausṫralẚae, SMF uncatalogued, male, 106.5 cm TL, Ras Hawlaf, Socotra Island; B: oK laevẚs, SMF uncatalogued, female, 115 cm TL, Qatar, the Gulf. Photos by S.V. Bogorodsky &amp; U. Zajonz." figureDoi="http://doi.org/10.5281/zenodo.4668385" httpUri="https://zenodo.org/record/4668385/files/figure.png" pageId="4" pageNumber="515">6A</figureCitation>
</paragraph>
<paragraph id="8BAEE0206641FFA1FF6C1319E489EB3B" blockId="4.[151,1417,1576,1601]" box="[151,1417,1576,1601]" pageId="4" pageNumber="515">
<emphasis id="B9653C326641FFA1FF6C1319E301EB3B" box="[151,513,1576,1600]" italics="true" pageId="4" pageNumber="515">ohynchobaṫus djẚddensẚs ausṫralẚae</emphasis>
<materialsCitation id="3B79EA7D6641FFA1FDFC1319E485EB3B" ID-GBIF-Occurrence="3071284305" box="[519,1413,1576,1601]" country="Australia" location="New South Wales" pageId="4" pageNumber="515" specimenCount="1" stateProvince="New South Wales" typeStatus="holotype">
<bibRefCitation id="EF809DD16641FFA1FDFC1319E3CCEB3B" author="Whitley, G. P." box="[519,716,1576,1601]" pageId="4" pageNumber="515" pagination="227 - 262" refId="ref13266" refString="Whitley, G. P. (1939) Taxonomic notes on sharks and rays. Aus tralẚan Zoolog ẚst, 9 (3), 227 - 262." type="journal article" year="1939">Whitley, 1939: 245</bibRefCitation>
;
<typeStatus id="54AA5E826641FFA1FD2C1319E234EB3B" box="[727,820,1576,1600]" pageId="4" pageNumber="515" type="holotype">holotype</typeStatus>
: AMS IA.4959; type locality:
<collectingRegion id="49D52EC26641FFA1FB9E1318E41CEB3B" box="[1125,1308,1576,1600]" country="Australia" name="New South Wales" pageId="4" pageNumber="515">New South Wales</collectingRegion>
,
<collectingCountry id="F306A0B06641FFA1FADE1319E485EB3B" box="[1317,1413,1576,1600]" name="Australia" pageId="4" pageNumber="515">Australia</collectingCountry>
</materialsCitation>
.
</paragraph>
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<subSubSection id="C30BB3AB6641FFA2FF6C135FE48EED81" lastPageId="7" lastPageNumber="518" pageId="4" pageNumber="515" type="materials_examined">
<paragraph id="8BAEE0206641FFA1FF6C135FE0CBEBAA" blockId="4.[151,1436,1646,1997]" pageId="4" pageNumber="515">
<emphasis id="B9653C326641FFA1FF6C135FE082EBF3" bold="true" box="[151,386,1646,1672]" pageId="4" pageNumber="515">Material examined:</emphasis>
<materialsCitation id="3B79EA7D6641FFA1FE73135EE0DFEBD6" ID-GBIF-Occurrence="3071284301" collectingDate="2019-04-10" collectionCode="SMF" country="Socotra Island" location="Hadibo" pageId="4" pageNumber="515" specimenCount="1">
<collectionCode id="ED0078E56641FFA1FE73135EE0C0EBF2" box="[392,448,1647,1673]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="4" pageNumber="515">SMF</collectionCode>
uncatalogued [tissue sample SOC19-447], female,
<quantity id="4CE94DC56641FFA1FC07135FE555EBF2" box="[1020,1109,1646,1673]" metricMagnitude="-1" metricUnit="m" metricValue="7.35" pageId="4" pageNumber="515" unit="cm" value="73.5">73.5 cm</quantity>
TL,
<collectingCountry id="F306A0B06641FFA1FB71135EE42DEBF2" box="[1162,1325,1647,1673]" name="Socotra Island" pageId="4" pageNumber="515">Socotra Island</collectingCountry>
, fish market in
<location id="8ECEB6FB6641FFA1FF2513A2E033EBD6" LSID="urn:lsid:plazi:treatment:03B851366641FFADFF6C10AAE310ECD5:8ECEB6FB6641FFA1FF2513A2E033EBD6" box="[222,307,1683,1709]" country="Socotra Island" name="Hadibo" pageId="4" pageNumber="515">Hadibo</location>
,
<date id="FFAFC6E06641FFA1FEC413A2E0DFEBD6" box="[319,479,1683,1709]" pageId="4" pageNumber="515" value="2019-04-10">
<collectingDate id="EFEB3F086641FFA1FEC413A2E0DFEBD6" box="[319,479,1683,1709]" pageId="4" pageNumber="515" value="2019-04-10">10 April 2019</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B79EA7D6641FFA1FE1713A2E0C7EBAA" ID-GBIF-Occurrence="3071284307" collectingDate="2019-04-11" collectionCode="SMF" country="Socotra Island" location="Ras Hawlaf" pageId="4" pageNumber="515" specimenCount="1">
<collectionCode id="ED0078E56641FFA1FE1713A2E324EBD6" box="[492,548,1683,1709]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="4" pageNumber="515">SMF</collectionCode>
uncatalogued [tissue sample SOC19-448], male,
<quantity id="4CE94DC56641FFA1FBA813A3E5BBEBD6" box="[1107,1211,1682,1709]" metricMagnitude="0" metricUnit="m" metricValue="1.065" pageId="4" pageNumber="515" unit="cm" value="106.5">106.5 cm</quantity>
TL,
<collectingCountry id="F306A0B06641FFA1FB0F13A2E498EBD6" box="[1268,1432,1683,1709]" name="Socotra Island" pageId="4" pageNumber="515">Socotra Island</collectingCountry>
,
<location id="8ECEB6FB6641FFA1FF6C1386E01DEBAA" LSID="urn:lsid:plazi:treatment:03B851366641FFADFF6C10AAE310ECD5:8ECEB6FB6641FFA1FF6C1386E01DEBAA" box="[151,285,1719,1745]" country="Socotra Island" name="Ras Hawlaf" pageId="4" pageNumber="515">Ras Hawlaf</location>
,
<date id="FFAFC6E06641FFA1FED21386E0C7EBAA" box="[297,455,1719,1745]" pageId="4" pageNumber="515" value="2019-04-11">
<collectingDate id="EFEB3F086641FFA1FED21386E0C7EBAA" box="[297,455,1719,1745]" pageId="4" pageNumber="515" value="2019-04-11">11 April 2019</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="8BAEE0206641FFA1FF3C13EBE0C9EA1A" blockId="4.[151,1436,1646,1997]" pageId="4" pageNumber="515">
<emphasis id="B9653C326641FFA1FF3C13EBE0D5EB8E" bold="true" box="[199,469,1754,1781]" pageId="4" pageNumber="515">Distinctive characters:</emphasis>
<emphasis id="B9653C326641FFA1FE2713EBE3F4EB8C" box="[476,756,1754,1783]" italics="true" pageId="4" pageNumber="515">ohynchobaṫus ausṫralẚae</emphasis>
is characterised by having a long and pointed, bottle-shaped snout that is slightly constricted near tip; posterior tip of pectoral fins ending before pelvic fins; no prominent ridges on head and body; spiracle with two folds; upper caudal-fin lobe longer than the lower one (
<bibRefCitation id="EF809DD16641FFA1FB5F1212E460EA46" author="Weigmann, S." box="[1188,1376,1827,1853]" pageId="4" pageNumber="515" pagination="249 - 312" refId="ref13016" refString="Weigmann, S. (2011) Contribution to the taxonomy and distribution of eight ray species (Chondrichthyes, Batoidea) from coastal waters of Thailand. mroceed ẚngs of the Soc ẚety of ka tural Sc ẚences ẚn eamburg, 46, 249 - 312." type="journal article" year="2011">Weigmann 2011</bibRefCitation>
; Last
<emphasis id="B9653C326641FFA1FF6C1277E1C8EA1A" box="[151,200,1862,1889]" italics="true" pageId="4" pageNumber="515">eṫ al</emphasis>
. 2016b;
<bibRefCitation id="EF809DD16641FFA1FED01276E0BDEA1A" author="Jabado, R. W." box="[299,445,1863,1889]" pageId="4" pageNumber="515" refId="ref10382" refString="Jabado, R. W. (2019) tedgef ẚshes and G ẚant Gu ẚtarfẚshes: a Gu ẚde to spec ẚes ẚdentẚfẚcatẚon. Wildlife Conservation Society, New York, 30 pp." type="book" year="2019">Jabado 2019</bibRefCitation>
).
</paragraph>
<paragraph id="8BAEE0206641FFA2FF3C125DE48EED81" blockId="4.[151,1436,1646,1997]" lastBlockId="7.[151,1437,151,1474]" lastPageId="7" lastPageNumber="518" pageId="4" pageNumber="515">
<emphasis id="B9653C326641FFA1FF3C125DE050EAFC" box="[199,336,1900,1927]" italics="true" pageId="4" pageNumber="515">Colouraṫẚon</emphasis>
: Olivaceous to grey-brown dorsally; head without transverse black bands between eye and spiracles; side of body, posterior half of pectoral fins and sometimes distal third of pelvic fins with sparse coverage of small white spots, which do not extent posteriorly behind rear tip of first dorsal fin; black pectoral marking surrounded by a distinctive pattern of white spots (three or four spots around the marking plus one or two in front of it). Large individuals usually entirely dark, with white spots and pectoral marking hardly discernible (Last
<emphasis id="B9653C326642FFA2FAF2158BE439EDAE" box="[1289,1337,186,213]" italics="true" pageId="7" pageNumber="518">eṫ al</emphasis>
. 2016b). Ventral surface almost uniformly white; underside of snout whitish in the middle, with violet hue laterally (
<figureCitation id="132AFCA56642FFA2FAD215EEE482ED81" box="[1321,1410,223,250]" captionStart="FIGURE 6" captionStartId="7.[151,250,1964,1988]" captionTargetBox="[151,1437,1497,1941]" captionTargetId="figure-686@7.[151,1437,1497,1941]" captionTargetPageId="7" captionText="FIGURE 6. ohynchobaṫus spp., underside of snout. A: oK ausṫralẚae, SMF uncatalogued, male, 106.5 cm TL, Ras Hawlaf, Socotra Island; B: oK laevẚs, SMF uncatalogued, female, 115 cm TL, Qatar, the Gulf. Photos by S.V. Bogorodsky &amp; U. Zajonz." figureDoi="http://doi.org/10.5281/zenodo.4668385" httpUri="https://zenodo.org/record/4668385/files/figure.png" pageId="7" pageNumber="518">Fig. 6A</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF6EB0A86640FFA0FF6C12A6E232EAAF" ID-DOI="http://doi.org/10.5281/zenodo.4668379" ID-Zenodo-Dep="4668379" httpUri="https://zenodo.org/record/4668379/files/figure.png" pageId="5" pageNumber="516" startId="5.[151,250,1943,1967]" targetBox="[153,1434,181,1920]" targetPageId="5">
<paragraph id="8BAEE0206640FFA0FF6C12A6E232EAAF" blockId="5.[151,1436,1943,2004]" pageId="5" pageNumber="516">
<emphasis id="B9653C326640FFA0FF6C12A6E015EAD4" bold="true" box="[151,277,1943,1967]" pageId="5" pageNumber="516">FIGURE 4.</emphasis>
<emphasis id="B9653C326640FFA0FEE512A6E0E3EAD4" box="[286,483,1943,1967]" italics="true" pageId="5" pageNumber="516">ohẚna ancylosṫoma</emphasis>
, fresh colouration, female, 225 cm TL, Abd al-Kuri Island, Socotra Archipelago.
<emphasis id="B9653C326640FFA0FAC012A6E455EAD4" bold="true" box="[1339,1365,1943,1967]" pageId="5" pageNumber="516">A:</emphasis>
dorsal view;
<emphasis id="B9653C326640FFA0FF2D128DE1F0EAAF" bold="true" box="[214,240,1980,2004]" pageId="5" pageNumber="516">B:</emphasis>
head close-up. Photos by S.V. Bogorodsky &amp; U. Zajonz.
</paragraph>
</caption>
<caption id="DF6EB0A86643FFA3FF6C1258E3EDEA95" ID-DOI="http://doi.org/10.5281/zenodo.4668381" ID-Zenodo-Dep="4668381" httpUri="https://zenodo.org/record/4668381/files/figure.png" pageId="6" pageNumber="517" startId="6.[151,250,1897,1921]" targetBox="[189,1399,181,1873]" targetPageId="6">
<paragraph id="8BAEE0206643FFA3FF6C1258E3EDEA95" blockId="6.[151,1436,1897,2030]" pageId="6" pageNumber="517">
<emphasis id="B9653C326643FFA3FF6C1258E013EAF9" bold="true" box="[151,275,1897,1922]" pageId="6" pageNumber="517">FIGURE 5.</emphasis>
<emphasis id="B9653C326643FFA3FEE11258E0A9EAFA" box="[282,425,1897,1921]" italics="true" pageId="6" pageNumber="517">ohynchobaṫus</emphasis>
spp., dorsal view, fresh colouration.
<emphasis id="B9653C326643FFA3FCDA1258E23CEAFA" bold="true" box="[801,828,1897,1921]" pageId="6" pageNumber="517">A:</emphasis>
<emphasis id="B9653C326643FFA3FCB8125BE2C6EAFA" box="[835,966,1897,1921]" italics="true" pageId="6" pageNumber="517">oK ausṫralẚae</emphasis>
, SMF uncatalogued, female, 73.5 cm TL, fish market in Hadibo, Socotra Island;
<emphasis id="B9653C326643FFA3FE0E12BFE30FEADD" bold="true" box="[501,527,1934,1958]" pageId="6" pageNumber="517">B:</emphasis>
<emphasis id="B9653C326643FFA3FDEE12BFE399EADE" box="[533,665,1933,1957]" italics="true" pageId="6" pageNumber="517">oK ausṫralẚae</emphasis>
, SMF uncatalogued, male, 106.5 cm TL, Ras Hawlaf, Socotra Island;
<emphasis id="B9653C326643FFA3FA9E12BCE480EADE" bold="true" box="[1381,1408,1933,1957]" pageId="6" pageNumber="517">C:</emphasis>
<emphasis id="B9653C326643FFA3FA7D12BFE1FEEAB2" italics="true" pageId="6" pageNumber="517">oK djẚddensẚs</emphasis>
, female, 130 cm TL, Socotra Island;
<emphasis id="B9653C326643FFA3FD8C1283E391EAB1" bold="true" box="[631,657,1970,1994]" pageId="6" pageNumber="517">D:</emphasis>
<emphasis id="B9653C326643FFA3FD6C1283E3EEEAB2" box="[663,750,1969,1993]" italics="true" pageId="6" pageNumber="517">oK laevẚs</emphasis>
, SMF uncatalogued, female, 115 cm TL, Qatar, the Gulf. Photos by S.V. Bogorodsky &amp; U. Zajonz (A, B and D), R. Bonfil (C).
</paragraph>
</caption>
<subSubSection id="C30BB3AB6642FFA2FF3C1433E028ECD5" pageId="7" pageNumber="518" type="distribution">
<paragraph id="8BAEE0206642FFA2FF3C1433E028ECD5" blockId="7.[151,1437,151,1474]" pageId="7" pageNumber="518">
<emphasis id="B9653C326642FFA2FF3C1433E063EC67" bold="true" box="[199,355,258,284]" pageId="7" pageNumber="518">Distribution:</emphasis>
Known from the
<collectingCountry id="F306A0B06642FFA2FDD71432E382EC66" box="[556,642,259,285]" name="Yemen" pageId="7" pageNumber="518">Socotra</collectingCountry>
Archipelago southward to
<collectingCountry id="F306A0B06642FFA2FC481432E51BEC66" box="[947,1051,259,285]" name="Tanzania" pageId="7" pageNumber="518">Tanzania</collectingCountry>
, eastward to eastern
<collectingCountry id="F306A0B06642FFA2FAFE1432E46FEC66" box="[1285,1391,259,285]" name="Australia" pageId="7" pageNumber="518">Australia</collectingCountry>
, the
<collectingCountry id="F306A0B06642FFA2FF6C1416E015EC3A" box="[151,277,295,321]" name="Philippines" pageId="7" pageNumber="518">Philippines</collectingCountry>
and
<collectingCountry id="F306A0B06642FFA2FEB51416E32CEC3A" box="[334,556,295,321]" name="Papua New Guinea" pageId="7" pageNumber="518">Papua New Guinea</collectingCountry>
. The presence off
<collectingCountry id="F306A0B06642FFA2FD071416E261EC3A" box="[764,865,295,321]" name="Tanzania" pageId="7" pageNumber="518">Tanzania</collectingCountry>
is confirmed by a specimen that nested within the same genetic lineage as
<emphasis id="B9653C326642FFA2FE5D147AE337EC1C" box="[422,567,331,359]" italics="true" pageId="7" pageNumber="518">oK ausṫralẚae</emphasis>
(unpublished data). Last
<emphasis id="B9653C326642FFA2FCAE147BE287EC1E" box="[853,903,330,357]" italics="true" pageId="7" pageNumber="518">eṫ al</emphasis>
. (2016b) included
<collectingCountry id="F306A0B06642FFA2FBA7147AE5F0EC1E" box="[1116,1264,331,357]" name="Mozambique" pageId="7" pageNumber="518">Mozambique</collectingCountry>
in the distribution range of the species but the respective specimen from off
<collectingCountry id="F306A0B06642FFA2FCAF145EE2E8ECF2" box="[852,1000,367,393]" name="Mozambique" pageId="7" pageNumber="518">Mozambique</collectingCountry>
is referred to as
<emphasis id="B9653C326642FFA2FB5E145EE436ECF0" box="[1189,1334,367,395]" italics="true" pageId="7" pageNumber="518">oK djẚddensẚs</emphasis>
by Giles
<emphasis id="B9653C326642FFA2FF6C14A3E1C8ECD6" box="[151,200,402,429]" italics="true" pageId="7" pageNumber="518">eṫ al</emphasis>
. (2016).
</paragraph>
</subSubSection>
<subSubSection id="C30BB3AB6642FFADFF3C1486E310ECD5" lastPageId="8" lastPageNumber="519" pageId="7" pageNumber="518" type="discussion">
<paragraph id="8BAEE0206642FFA2FF3C1486E437EE6E" blockId="7.[151,1437,151,1474]" pageId="7" pageNumber="518">
<emphasis id="B9653C326642FFA2FF3C1486E03DECAA" bold="true" box="[199,317,439,465]" pageId="7" pageNumber="518">Remarks:</emphasis>
Giles
<emphasis id="B9653C326642FFA2FE721487E0BBECAA" box="[393,443,438,465]" italics="true" pageId="7" pageNumber="518">eṫ al</emphasis>
. (2016) provided comprehensive data on the genetic structure and phenotypic variation within
<emphasis id="B9653C326642FFA2FF1C14EAE078EC8C" box="[231,376,475,503]" italics="true" pageId="7" pageNumber="518">oK ausṫralẚae</emphasis>
but no specimens from the Arabian region were examined in their study. Jabado
<emphasis id="B9653C326642FFA2FAEA14EBE443EC8E" box="[1297,1347,474,501]" italics="true" pageId="7" pageNumber="518">eṫ al</emphasis>
. (2017) summarized the conservation status of sharks, rays and chimaeras in the Arabian region and included three species of
<emphasis id="B9653C326642FFA2FF431713E056EF46" box="[184,342,546,573]" italics="true" pageId="7" pageNumber="518">ohynchobaṫus</emphasis>
:
<emphasis id="B9653C326642FFA2FE931712E0FBEF44" box="[360,507,547,575]" italics="true" pageId="7" pageNumber="518">oK ausṫralẚae</emphasis>
,
<emphasis id="B9653C326642FFA2FDF71712E39FEF44" box="[524,671,547,575]" italics="true" pageId="7" pageNumber="518">oK djẚddensẚs</emphasis>
(
<bibRefCitation id="EF809DD16642FFA2FD4B1713E25AEF46" author="Forsskal, P. S." box="[688,858,546,573]" pageId="7" pageNumber="518" refId="ref9644" refString="Forsskal, P. S. (1775) oaja dj ẚddensẚs. fn: Niebuhr, C. (Ed.), aescr ẚptẚones an ẚmalẚum av ẚumI amph ẚbẚorumI p ẚscẚumI ẚnsectorumI verm ẚum; quae ẚn ẚtẚnere or ẚentalẚ observav ẚtKKK mos t mor tem auc torẚs ed ẚdẚt Cars ten k ẚebuhrK ex officina Molleri, Hauniae, pp. 18 + viii. https: // doi. org / 10.5962 / bhl. title. 2154" type="book" year="1775">Forsskål, 1775</bibRefCitation>
) and
<emphasis id="B9653C326642FFA2FC651712E501EF44" box="[926,1025,547,575]" italics="true" pageId="7" pageNumber="518">oK laevẚs</emphasis>
(
<bibRefCitation id="EF809DD16642FFA2FBE91712E43AEF46" author="Bloch, M. E. &amp; Schneider, J. G." box="[1042,1338,547,573]" pageId="7" pageNumber="518" refId="ref8596" refString="Bloch, M. E. &amp; Schneider, J. G. (1801) MKbK Bloch ẚẚ xKKKz Sys tema ẚchthyologẚae ẚconẚbus Cu ẚllustratumK mos t ob ẚtum auc torẚs opus ẚnchoatum absolv ẚtI correx ẚtI ẚnterpolavẚt goK Go ttlob Schne ẚderI Saxo. Sumtibus auctoris impressum et bibliopolio Sanderiano commissum, Berolini, LX + 584 pp. https: // doi. org / 10.5962 / bhl. title. 5750" type="book" year="1801">Bloch &amp; Schneider, 1801</bibRefCitation>
). Subsequently, Jabado (2018) listed those three species as present in the Gulf and
<collectingCountry id="F306A0B06642FFA2FC0D1776E53AEF1A" box="[1014,1082,583,609]" name="Oman" pageId="7" pageNumber="518">Oman</collectingCountry>
waters; whereby both publications lack evidence for the presence of
<emphasis id="B9653C326642FFA2FDB5175AE3DEEFFC" box="[590,734,619,647]" italics="true" pageId="7" pageNumber="518">oK ausṫralẚae</emphasis>
in the region. So far, only one confirmed record of the species from the south-eastern part of the Arabian Sea existed based on two specimens landed at
<collectingCountry id="F306A0B06642FFA2FB8D17BEE5C7EFD2" box="[1142,1223,655,681]" name="India" pageId="7" pageNumber="518">Cochin</collectingCountry>
Fisheries Harbour,
<collectingRegion id="49D52EC26642FFA2FF6C1782E1E2EFB6" box="[151,226,691,717]" country="India" name="Kerala" pageId="7" pageNumber="518">Kerala</collectingRegion>
(Bineesh
<emphasis id="B9653C326642FFA2FEA81783E083EFB6" box="[339,387,690,717]" italics="true" pageId="7" pageNumber="518">eṫ al</emphasis>
. 2014).
<bibRefCitation id="EF809DD16642FFA2FE1B1782E37FEFB6" author="Jabado, R. W." box="[480,639,691,717]" pageId="7" pageNumber="518" refId="ref10382" refString="Jabado, R. W. (2019) tedgef ẚshes and G ẚant Gu ẚtarfẚshes: a Gu ẚde to spec ẚes ẚdentẚfẚcatẚon. Wildlife Conservation Society, New York, 30 pp." type="book" year="2019">Jabado (2019)</bibRefCitation>
provided an illustrated guide for the identification of all known species of
<taxonomicName id="4C119BA36642FFA2FF4E17E6E01AEF8A" authorityName="Muller &amp; Henle" authorityYear="1841" box="[181,282,727,753]" class="Elasmobranchii" family="Rhinidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rajiformes" pageId="7" pageNumber="518" phylum="Chordata" rank="family">Rhinidae</taxonomicName>
based on fresh specimens and fins. She noted the misidentification of species of the genus as a common problem for fisheries management and accurate identification as being critical for conservation programs.
</paragraph>
<paragraph id="8BAEE0206642FFA2FF3C162FE41CE906" blockId="7.[151,1437,151,1474]" pageId="7" pageNumber="518">
<emphasis id="B9653C326642FFA2FF3C162FE0D9EE40" box="[199,473,798,827]" italics="true" pageId="7" pageNumber="518">ohynchobaṫus djẚddensẚs</emphasis>
, described from the Red Sea, ranges southward to
<collectingCountry id="F306A0B06642FFA2FBEB162EE59FEE42" box="[1040,1183,799,825]" name="South Africa" pageId="7" pageNumber="518">South Africa</collectingCountry>
, northward to the Gulf and into the north-eastern part of the Arabian Sea (Last
<emphasis id="B9653C326642FFA2FCFE1673E236EE26" box="[773,822,834,861]" italics="true" pageId="7" pageNumber="518">eṫ al</emphasis>
. 2016b; Kizhakudan
<emphasis id="B9653C326642FFA2FBDD1673E556EE26" box="[1062,1110,834,861]" italics="true" pageId="7" pageNumber="518">eṫ al</emphasis>
. 2018), with positive records in the region from the Red Sea, south coast of
<collectingCountry id="F306A0B06642FFA2FD611656E3E7EEFA" box="[666,743,871,897]" name="Yemen" pageId="7" pageNumber="518">Yemen</collectingCountry>
(
<bibRefCitation id="EF809DD16642FFA2FD0D1656E508EEFA" author="Bonfil, R. &amp; Abdallah, M." box="[758,1032,871,897]" pageId="7" pageNumber="518" refId="ref8738" refString="Bonfil, R. &amp; Abdallah, M. (2004) cAO Spec ẚes fden tẚfẚcatẚon Gu ẚde for c ẚshery murposesK c ẚeld fden tẚfẚcatẚon Gu ẚde to the Sharks and oays of the oed Sea and Gulf of AdenK Food and Agriculture Organization of the United Nations, Rome, IX + 71 pp." type="book" year="2004">Bonfil &amp; Abdallah 2004</bibRefCitation>
),
<collectingCountry id="F306A0B06642FFA2FBE01656E55EEEFA" box="[1051,1118,871,897]" name="Oman" pageId="7" pageNumber="518">Oman</collectingCountry>
(Henderson
<emphasis id="B9653C326642FFA2FB151657E425EEFA" box="[1262,1317,870,897]" italics="true" pageId="7" pageNumber="518">eṫ alK</emphasis>
2015) and the Gulf (Almojil
<emphasis id="B9653C326642FFA2FEA716BBE08BEEDE" box="[348,395,906,933]" italics="true" pageId="7" pageNumber="518">eṫ al</emphasis>
. 2015; Henderson
<emphasis id="B9653C326642FFA2FDA016BBE38AEEDE" box="[603,650,906,933]" italics="true" pageId="7" pageNumber="518">eṫ al</emphasis>
. 2015).
<bibRefCitation id="EF809DD16642FFA2FD1F16BAE277EEDE" author="Kemp, J. M." box="[740,887,907,933]" pageId="7" pageNumber="518" pagination="919 - 933" refId="ref10654" refString="Kemp, J. M. (1998) Zoogeography of the coral reef fishes of the Socotra Archipelago. gournal of B ẚogeography, 25 (5), 919 - 933. https: // doi. org / 10.1046 / j. 1365 - 2699.1998.00249. x" type="journal article" year="1998">Kemp (1998)</bibRefCitation>
listed
<emphasis id="B9653C326642FFA2FC4416BAE54CEEDC" box="[959,1100,907,935]" italics="true" pageId="7" pageNumber="518">oK djẚddensẚs</emphasis>
from the
<collectingCountry id="F306A0B06642FFA2FB4E16BAE409EEDE" box="[1205,1289,907,933]" name="Yemen" pageId="7" pageNumber="518">Socotra</collectingCountry>
Archipelago, yet without sample or photographic evidence.
<bibRefCitation id="EF809DD16642FFA2FD65169EE2BFEEB2" author="Bonfil, R. &amp; Abdallah, M." box="[670,959,943,969]" pageId="7" pageNumber="518" refId="ref8738" refString="Bonfil, R. &amp; Abdallah, M. (2004) cAO Spec ẚes fden tẚfẚcatẚon Gu ẚde for c ẚshery murposesK c ẚeld fden tẚfẚcatẚon Gu ẚde to the Sharks and oays of the oed Sea and Gulf of AdenK Food and Agriculture Organization of the United Nations, Rome, IX + 71 pp." type="book" year="2004">Bonfil &amp; Abdallah (2004)</bibRefCitation>
subsequently included a photograph taken from off
<collectingCountry id="F306A0B06642FFA2FF0C16E2E094EE96" box="[247,404,979,1005]" name="Socotra Island" pageId="7" pageNumber="518">Socotra Island</collectingCountry>
in their FAO Guide (Ramon Bonfil, personal communication), confirming its occurrence there.
<emphasis id="B9653C326642FFA2FF6C16C7E079E968" box="[151,377,1014,1043]" italics="true" pageId="7" pageNumber="518">ohynchobaṫus laevẚs</emphasis>
is reported in the region from off
<collectingCountry id="F306A0B06642FFA2FD0A16C6E234E96A" box="[753,820,1015,1041]" name="Oman" pageId="7" pageNumber="518">Oman</collectingCountry>
(Randall 1995 as
<emphasis id="B9653C326642FFA2FC0616C6E58BE968" box="[1021,1163,1015,1043]" italics="true" pageId="7" pageNumber="518">oK djẚddensẚs</emphasis>
; Henderson
<emphasis id="B9653C326642FFA2FAEC16C7E447E96A" box="[1303,1351,1014,1041]" italics="true" pageId="7" pageNumber="518">eṫ al</emphasis>
. 2015), the Gulf (Almojil
<emphasis id="B9653C326642FFA2FEA0112BE08AE94E" box="[347,394,1050,1077]" italics="true" pageId="7" pageNumber="518">eṫ al</emphasis>
. 2015; Henderson
<emphasis id="B9653C326642FFA2FDA3112BE386E94E" box="[600,646,1050,1077]" italics="true" pageId="7" pageNumber="518">eṫ al</emphasis>
. 2015), and
<collectingCountry id="F306A0B06642FFA2FCF6112AE26AE94E" box="[781,874,1051,1077]" name="Pakistan" pageId="7" pageNumber="518">Pakistan</collectingCountry>
(Psomadakis
<emphasis id="B9653C326642FFA2FBF9112BE531E94E" box="[1026,1073,1050,1077]" italics="true" pageId="7" pageNumber="518">eṫ al</emphasis>
. 2015).
<bibRefCitation id="EF809DD16642FFA2FB71112AE49CE94E" author="Spaet, J. Y. L. &amp; Berumen, M. L." box="[1162,1436,1050,1077]" pageId="7" pageNumber="518" pagination="356 - 364" refId="ref12912" refString="Spaet, J. Y. L. &amp; Berumen, M. L. (2015) Fish market surveys indicate unsustainable elasmobranch fisheries in the Saudi Arabian Red Sea. c ẚsherẚes oesearch, 161, 356 - 364. https: // doi. org / 10.1016 / j. fishres. 2014.08.022" type="journal article" year="2015">Spaet &amp; Berumen (2015)</bibRefCitation>
reported the presence of two species of
<emphasis id="B9653C326642FFA2FDB5110FE3EAE922" box="[590,746,1086,1113]" italics="true" pageId="7" pageNumber="518">ohynchobaṫus</emphasis>
in the Red Sea, yet without providing photographs or descriptions. Until now, no evidence for the occurrence of
<emphasis id="B9653C326642FFA2FD331152E255E904" box="[712,853,1123,1151]" italics="true" pageId="7" pageNumber="518">oK ausṫralẚae</emphasis>
in the Arabian region had been provided.
</paragraph>
<paragraph id="8BAEE0206642FFA2FF3C11B7E09EE8B9" blockId="7.[151,1437,151,1474]" pageId="7" pageNumber="518">
<emphasis id="B9653C326642FFA2FF3C11B7E0E0E9D8" box="[199,480,1158,1187]" italics="true" pageId="7" pageNumber="518">ohynchobaṫus djẚddensẚs</emphasis>
differs from other species by having a large black, circular pectoral marking about as large as eye (may be indistinct or absent in adults), a dense pattern of white spots (sometimes also rings) on the dorsal surface, which do not or slightly extend forward of pectoral markings on mid-disc, absence of white spots on the pelvic fins, and prominent mask-like markings between the eyes (Last
<emphasis id="B9653C326642FFA2FC0611C3E52EE876" box="[1021,1070,1266,1293]" italics="true" pageId="7" pageNumber="518">eṫ al</emphasis>
. 2016b;
<bibRefCitation id="EF809DD16642FFA2FB6A11C2E420E876" author="Jabado, R. W." box="[1169,1312,1267,1293]" pageId="7" pageNumber="518" refId="ref10382" refString="Jabado, R. W. (2019) tedgef ẚshes and G ẚant Gu ẚtarfẚshes: a Gu ẚde to spec ẚes ẚdentẚfẚcatẚon. Wildlife Conservation Society, New York, 30 pp." type="book" year="2019">Jabado 2019</bibRefCitation>
) (
<figureCitation id="132AFCA56642FFA2FACC11C2E490E876" box="[1335,1424,1266,1293]" captionStart="FIGURE 5" captionStartId="6.[151,250,1897,1921]" captionTargetBox="[189,1399,181,1873]" captionTargetId="figure-23@6.[189,1399,181,1873]" captionTargetPageId="6" captionText="FIGURE 5. ohynchobaṫus spp., dorsal view, fresh colouration. A: oK ausṫralẚae, SMF uncatalogued, female, 73.5 cm TL, fish market in Hadibo, Socotra Island; B: oK ausṫralẚae, SMF uncatalogued, male, 106.5 cm TL, Ras Hawlaf, Socotra Island; C: oK djẚddensẚs, female, 130 cm TL, Socotra Island; D: oK laevẚs, SMF uncatalogued, female, 115 cm TL, Qatar, the Gulf. Photos by S.V. Bogorodsky &amp; U. Zajonz (A, B and D), R. Bonfil (C)." figureDoi="http://doi.org/10.5281/zenodo.4668381" httpUri="https://zenodo.org/record/4668381/files/figure.png" pageId="7" pageNumber="518">Fig. 5C</figureCitation>
).
<emphasis id="B9653C326642FFA2FF6C1027E081E848" box="[151,385,1302,1331]" italics="true" pageId="7" pageNumber="518">ohynchobaṫus laevẚs</emphasis>
differs from its congeners by having a relatively broad snout, rows of small white spots on the dorsal surface of body and tail, spots beginning distinctly behind black pectoral markings (usually the dorsal surface anterior to first dorsal-fin origin remains unspotted), a pectoral marking that usually is a black ocellus with pale brown centre surrounded by 57 small white spots, and a blackish underside of snout (Last
<emphasis id="B9653C326642FFA2FB4510B3E5EEE8E6" box="[1214,1262,1410,1437]" italics="true" pageId="7" pageNumber="518">eṫ al</emphasis>
. 2016b;
<bibRefCitation id="EF809DD16642FFA2FAB510B2E1D1E8BA" author="Jabado, R. W." pageId="7" pageNumber="518" refId="ref10382" refString="Jabado, R. W. (2019) tedgef ẚshes and G ẚant Gu ẚtarfẚshes: a Gu ẚde to spec ẚes ẚdentẚfẚcatẚon. Wildlife Conservation Society, New York, 30 pp." type="book" year="2019">Jabado 2019</bibRefCitation>
) (
<figureCitation id="132AFCA56642FFA2FF131096E049E8BA" box="[232,329,1446,1473]" captionStart="FIGURE 5" captionStartId="6.[151,250,1897,1921]" captionTargetBox="[189,1399,181,1873]" captionTargetId="figure-23@6.[189,1399,181,1873]" captionTargetPageId="6" captionText="FIGURE 5. ohynchobaṫus spp., dorsal view, fresh colouration. A: oK ausṫralẚae, SMF uncatalogued, female, 73.5 cm TL, fish market in Hadibo, Socotra Island; B: oK ausṫralẚae, SMF uncatalogued, male, 106.5 cm TL, Ras Hawlaf, Socotra Island; C: oK djẚddensẚs, female, 130 cm TL, Socotra Island; D: oK laevẚs, SMF uncatalogued, female, 115 cm TL, Qatar, the Gulf. Photos by S.V. Bogorodsky &amp; U. Zajonz (A, B and D), R. Bonfil (C)." figureDoi="http://doi.org/10.5281/zenodo.4668381" httpUri="https://zenodo.org/record/4668381/files/figure.png" pageId="7" pageNumber="518">Figs. 5D</figureCitation>
&amp;
<figureCitation id="132AFCA56642FFA2FE961096E092E8B9" box="[365,402,1447,1474]" captionStart="FIGURE 6" captionStartId="7.[151,250,1964,1988]" captionTargetBox="[151,1437,1497,1941]" captionTargetId="figure-686@7.[151,1437,1497,1941]" captionTargetPageId="7" captionText="FIGURE 6. ohynchobaṫus spp., underside of snout. A: oK ausṫralẚae, SMF uncatalogued, male, 106.5 cm TL, Ras Hawlaf, Socotra Island; B: oK laevẚs, SMF uncatalogued, female, 115 cm TL, Qatar, the Gulf. Photos by S.V. Bogorodsky &amp; U. Zajonz." figureDoi="http://doi.org/10.5281/zenodo.4668385" httpUri="https://zenodo.org/record/4668385/files/figure.png" pageId="7" pageNumber="518">6B</figureCitation>
).
</paragraph>
<caption id="DF6EB0A86642FFA2FF6C129DE481EA92" ID-DOI="http://doi.org/10.5281/zenodo.4668385" ID-Zenodo-Dep="4668385" httpUri="https://zenodo.org/record/4668385/files/figure.png" pageId="7" pageNumber="518" startId="7.[151,250,1964,1988]" targetBox="[151,1437,1497,1941]" targetPageId="7">
<paragraph id="8BAEE0206642FFA2FF6C129DE481EA92" blockId="7.[151,1436,1964,2025]" pageId="7" pageNumber="518">
<emphasis id="B9653C326642FFA2FF6C129DE013EABE" bold="true" box="[151,275,1964,1989]" pageId="7" pageNumber="518">FIGURE 6.</emphasis>
<emphasis id="B9653C326642FFA2FEE2129DE0A8EABF" box="[281,424,1964,1988]" italics="true" pageId="7" pageNumber="518">ohynchobaṫus</emphasis>
spp., underside of snout.
<emphasis id="B9653C326642FFA2FD57129DE3C7EABF" bold="true" box="[684,711,1964,1988]" pageId="7" pageNumber="518">A:</emphasis>
<emphasis id="B9653C326642FFA2FD36129CE250EABF" box="[717,848,1964,1988]" italics="true" pageId="7" pageNumber="518">oK ausṫralẚae</emphasis>
, SMF uncatalogued, male, 106.5 cm TL, Ras Hawlaf, Socotra Island;
<emphasis id="B9653C326642FFA2FEE012E0E035EA92" bold="true" box="[283,309,2001,2025]" pageId="7" pageNumber="518">B:</emphasis>
<emphasis id="B9653C326642FFA2FEC012E0E092EA93" box="[315,402,2000,2024]" italics="true" pageId="7" pageNumber="518">oK laevẚs</emphasis>
, SMF uncatalogued, female, 115 cm TL, Qatar, the Gulf. Photos by S.V. Bogorodsky &amp; U. Zajonz.
</paragraph>
</caption>
<paragraph id="8BAEE020664DFFADFF3C15A6E310ECD5" blockId="8.[151,1437,150,430]" pageId="8" pageNumber="519">
Both examined specimens match well the descriptions of
<emphasis id="B9653C32664DFFADFC9115A7E585EDC8" box="[874,1157,150,179]" italics="true" pageId="8" pageNumber="519">ohynchobaṫus ausṫralẚae</emphasis>
provided by
<bibRefCitation id="EF809DD1664DFFADFADF15A6E1E1EDAE" author="Weigmann, S." pageId="8" pageNumber="519" pagination="249 - 312" refId="ref13016" refString="Weigmann, S. (2011) Contribution to the taxonomy and distribution of eight ray species (Chondrichthyes, Batoidea) from coastal waters of Thailand. mroceed ẚngs of the Soc ẚety of ka tural Sc ẚences ẚn eamburg, 46, 249 - 312." type="journal article" year="2011">Weigmann (2011)</bibRefCitation>
and Last
<emphasis id="B9653C32664DFFADFEB0158BE07BEDAE" box="[331,379,186,213]" italics="true" pageId="8" pageNumber="519">eṫ al</emphasis>
. (2016b), representing the first confirmed record for the Arabian region. Elsewhere, the species typically lives on sedimentary substrata of shallow lagoons and coastal reef areas at depths of
<quantity id="4CE94DC5664DFFADFB4615EEE413ED82" box="[1213,1299,223,249]" metricMagnitude="1" metricUnit="m" metricValue="3.0" metricValueMax="6.0" metricValueMin="0.0" pageId="8" pageNumber="519" unit="m" value="30.0" valueMax="60.0" valueMin="0.0">060 m</quantity>
(
<bibRefCitation id="EF809DD1664DFFADFADA15EEE1D4EC65" author="Weigmann, S." pageId="8" pageNumber="519" pagination="837 - 1037" refId="ref13064" refString="Weigmann, S. (2016) Annotated checklist of the living sharks, batoids and chimaeras (Chondrichthyes) of the world, with a focus on biogeographical diversity. gournal of c ẚsh B ẚology, 88 (3), 837 - 1037. https: // doi. org / 10.1111 / jfb. 12874" type="journal article" year="2016">Weigmann 2016</bibRefCitation>
). The female was taken from the fish market in Hadibo without precise collection data on locality and habitat, while the male was caught by two of the authors (SVB and FNS) in a large lagoon with silty sand bottom in an area mixed with small stones and sea weeds at a depth of
<quantity id="4CE94DC5664DFFADFCE3147BE25FEC1E" box="[792,863,330,357]" metricMagnitude="0" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="8" pageNumber="519" unit="m" value="4.5" valueMax="5.0" valueMin="4.0">45 m</quantity>
in Ras Hawlaf bay, northern part of
<collectingCountry id="F306A0B0664DFFADFB0E147AE495EC1E" box="[1269,1429,331,357]" name="Socotra Island" pageId="8" pageNumber="519">Socotra Island</collectingCountry>
. The species is included as critically endangered in IUCN Red List Assessments and urgently requires conservation management (Kyne
<emphasis id="B9653C32664DFFADFE8114A3E0ABECD6" box="[378,427,402,429]" italics="true" pageId="8" pageNumber="519">eṫ al</emphasis>
. 2019b).
</paragraph>
</subSubSection>
</treatment>
</document>