plazi/treatments-xml
2
0
Fork 0
Code Issues Pull requests Releases Activity
treatments-xml/data/03/98/87/039887CBFFA3FF96F284F2B7FEA1A815.xml
maintenance bc2564723d added 03
2024-06-21 12:22:17 +02:00

472 lines
56 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="6A7623EF77D8173D674927BAD5311FD9" ID-CLB-Dataset="46369" ID-DOI="10.11646/zootaxa.3768.2.3" ID-GBIF-Dataset="c232760e-33e0-499b-adc1-ede336e64d29" ID-ISSN="1175-5326" ID-Zenodo-Dep="252057" ID-ZooBank="40B1B7AF-84E8-4EC0-97DA-68E4BD0C482A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460388281548" checkinUser="plazi" docAuthor="Maryan, Brad, Oliver, Paul M., Fitch, Alison J. &amp; O’Connell, Morgan" docDate="2014" docId="039887CBFFA3FF96F284F2B7FEA1A815" docLanguage="en" docName="zt03768p158.pdf" docOrigin="Zootaxa 3768 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Varanus hamersleyensis Maryan, Oliver, Fitch &amp; O’Connell, 2014, sp. nov." docType="treatment" docVersion="9" lastPageNumber="155" masterDocId="FFA1FFB3FFA8FF86F213F602FFAFAB00" masterDocTitle="Molecular and morphological assessment of Varanus pilbarensis (Squamata: Varanidae), with a description of a new species from the southern Pilbara, Western Australia" masterLastPageNumber="158" masterPageNumber="139" pageNumber="150" updateTime="1698297920950" updateUser="ExternalLinkService">
<mods:mods id="6888DEF7A60DFBEB5DD4F90FE8B22D9C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B0C5BD8D30CFAE9BCEBA5B7B0C2C1D62">
<mods:title id="15412692A0BFBE59D182972FA711D711">Molecular and morphological assessment of Varanus pilbarensis (Squamata: Varanidae), with a description of a new species from the southern Pilbara, Western Australia</mods:title>
</mods:titleInfo>
<mods:name id="8B54FF677F0F397504FD3F873742C308" type="personal">
<mods:role id="E1C8D3B29673584A5EF36F3449EDE35A">
<mods:roleTerm id="258E793EFAA3EA17FDA5DE05E07A208D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2D0907065014CB8DFB3F71D90424127E">Maryan, Brad</mods:namePart>
</mods:name>
<mods:name id="31F34E7E8AA6698852F118FB3F92CBED" type="personal">
<mods:role id="8B4F167EC52C7C8364B1905594F6BD79">
<mods:roleTerm id="A9E52F4766F15BBA89D9D4260E75DB09">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="733D178A87A7810C7A1B5E3635EACCE5">Oliver, Paul M.</mods:namePart>
</mods:name>
<mods:name id="F82252766574190CF58C00B798501C07" type="personal">
<mods:role id="4D0EBF23CBF5C4D945B72140F854F753">
<mods:roleTerm id="FC1739EE7E7405ABACF9539AFAE065D7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2135909412AC2423C89DD850E7A47838">Fitch, Alison J.</mods:namePart>
</mods:name>
<mods:name id="C3045A222CCBC7DB3F714F348EED51DA" type="personal">
<mods:role id="634B9408B89FDDBFD3D202458554B08E">
<mods:roleTerm id="13F5FDE68F10DE20156089B3ACFFEFAF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="964D0DDFF25FFD9E7BE62DEA739D6210">O’Connell, Morgan</mods:namePart>
</mods:name>
<mods:typeOfResource id="2B90FF886EBC564DAA1658D943424421">text</mods:typeOfResource>
<mods:relatedItem id="C73BEFC4F83DA0B1F42B9BE863D2761C" type="host">
<mods:titleInfo id="44B13021923DA67B0BA04C4EF1A1A43C">
<mods:title id="1AFDFC8367B33328DE767BB79D758138">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="CAE68636FF00E8DA4DC21F74A1153D88">
<mods:date id="ECA1D7B01F4C0A1F5C838C97B303743D">2014</mods:date>
<mods:detail id="53B36C9B106FFB71DCF60BC2CB275C60" type="volume">
<mods:number id="38F27A0119F8F9AA6FDF2B89396B2C27">3768</mods:number>
</mods:detail>
<mods:detail id="BBAECA9288E044BADE89A4A3E6A54177" type="issue">
<mods:number id="545FC37A0EDCC849756E17559742D182">2</mods:number>
</mods:detail>
<mods:extent id="04769E37017F1E280AC8A5D4AE8F9776" unit="page">
<mods:start id="282F2457DA8BA0B5C982738440A53886">139</mods:start>
<mods:end id="D06F422A5E22258AAE425D4D3834D9CD">158</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="741895B7F6BEE324E189D372F4D4B2CE">journal article</mods:classification>
<mods:identifier id="9962B7862A1AF65BBF1E99A80D833D95" type="CLB-Dataset">46369</mods:identifier>
<mods:identifier id="D11D5B148579F6DC68495650D27C03B4" type="DOI">10.11646/zootaxa.3768.2.3</mods:identifier>
<mods:identifier id="FEE88D0B02F20BED33844610C093EDA7" type="GBIF-Dataset">c232760e-33e0-499b-adc1-ede336e64d29</mods:identifier>
<mods:identifier id="72C74575B4A8FD3B29B71D3A57B86858" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="C9A73CA07625CD7D061C1CB3AD5BAACE" type="Zenodo-Dep">252057</mods:identifier>
<mods:identifier id="E3AE71697F2D8719DB5EA449A2C5E442" type="ZooBank">40B1B7AF-84E8-4EC0-97DA-68E4BD0C482A</mods:identifier>
</mods:mods>
<treatment id="039887CBFFA3FF96F284F2B7FEA1A815" ID-DOI="http://doi.org/10.5281/zenodo.5613439" ID-GBIF-Taxon="119416045" ID-Zenodo-Dep="5613439" LSID="urn:lsid:plazi:treatment:039887CBFFA3FF96F284F2B7FEA1A815" httpUri="http://treatment.plazi.org/id/039887CBFFA3FF96F284F2B7FEA1A815" lastPageId="16" lastPageNumber="155" pageId="11" pageNumber="150">
<subSubSection id="C32B6556FFA3FF8DF284F2B7FE94AE15" pageId="11" pageNumber="150" type="nomenclature">
<paragraph id="8B8E36DDFFA3FF8DF284F2B7FD8FAFD0" blockId="11.[151,544,1205,1301]" box="[151,544,1205,1232]" pageId="11" pageNumber="150">
<heading id="D0C681B1FFA3FF8DF284F2B7FD8FAFD0" bold="true" box="[151,544,1205,1232]" fontSize="11" level="1" pageId="11" pageNumber="150" reason="1">
<emphasis id="B945EACFFFA3FF8DF284F2B7FD8FAFD0" bold="true" box="[151,544,1205,1232]" pageId="11" pageNumber="150">
<taxonomicName id="4C314D5EFFA3FF8DF284F2B7FE14AFCF" box="[151,443,1205,1231]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="150" phylum="Chordata" rank="species" species="hamersleyensis" status="sp. nov.">
<emphasis id="B945EACFFFA3FF8DF284F2B7FE14AFCF" bold="true" box="[151,443,1205,1231]" italics="true" pageId="11" pageNumber="150">Varanus hamersleyensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27657B4FFA3FF8DF3D1F2B4FD8FAFD0" box="[450,544,1206,1232]" pageId="11" pageNumber="150" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8E36DDFFA3FF8DF284F2DAFE94AE15" blockId="11.[151,544,1205,1301]" pageId="11" pageNumber="150">
Southern Pilbara Rock Goanna (
<figureCitation id="130A2A58FFA3FF8DF28CF2FEFF42AE15" box="[159,237,1276,1301]" captionStart="FIGURE 8" captionStartId="12.[151,250,1621,1643]" captionTargetBox="[151,1436,176,1588]" captionTargetId="figure@12.[151,1436,157,1588]" captionTargetPageId="12" captionText="FIGURE 8. Preserved holotype WAM R 145733 of V. hamersleyensis sp. nov. in dorsal, ventral and lateral views." httpUri="https://zenodo.org/record/252064/files/figure.png" pageId="11" pageNumber="150">Figs. 8</figureCitation>
,
<figureCitation id="130A2A58FFA3FF8DF2E9F2FFFEA5AE15" box="[250,266,1277,1301]" captionStart="FIGURE 9" captionStartId="14.[151,250,1847,1869]" captionTargetBox="[161,1425,218,1791]" captionTargetId="figure@14.[151,1436,213,1803]" captionTargetPageId="14" captionText="FIGURE 9. Adult V. hamersleyensis sp. nov. photographed in life from (A) Marillana and (B) Ophthalmia Range (images—B. Maryan)." httpUri="https://zenodo.org/record/252065/files/figure.png" pageId="11" pageNumber="150">9</figureCitation>
,
<figureCitation id="130A2A58FFA3FF8DF305F2FFFE9CAE15" box="[278,307,1277,1301]" captionStart="FIGURE 10" captionStartId="15.[151,250,1045,1067]" captionTargetBox="[151,1436,176,1023]" captionTargetId="figure@15.[151,1436,167,1023]" captionTargetPageId="15" captionText="FIGURE 10. Variation in V. hamersleyensis sp. nov. from left to right, WAM R 125456, WAM R 125766, WAM R 140734, WAM R 164579." httpUri="https://zenodo.org/record/252066/files/figure.png" pageId="11" pageNumber="150">10</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32B6556FFA3FF8DF284F346FE12AD35" pageId="11" pageNumber="150" type="materials_examined">
<paragraph id="8B8E36DDFFA3FF8DF284F346FE12AD35" blockId="11.[151,1437,1348,2022]" pageId="11" pageNumber="150">
<emphasis id="B945EACFFFA3FF8DF284F346FDA0AE5D" bold="true" box="[151,527,1348,1373]" pageId="11" pageNumber="150">
Material examined.
<typeStatus id="548A887FFFA3FF8DF38AF346FDA5AE5D" box="[409,522,1348,1373]" pageId="11" pageNumber="150" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectionCode id="ED20AE18FFA3FF8DF00CF347FDCCAE5D" LSID="urn:lsid:biocol.org:col:34981" box="[543,611,1349,1373]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="11" pageNumber="150">WAM</collectionCode>
R145733, male, Hamersley Range, Weeli Wolli Creek (
<geoCoordinate id="EE05501AFFA3FF8DF70CF346FA33AE5D" box="[1311,1436,1348,1373]" direction="south" orientation="latitude" pageId="11" pageNumber="150" precision="15" value="-22.877777">22º52'40&quot;S</geoCoordinate>
<geoCoordinate id="EE05501AFFA3FF8DF284F36BFE8BAE81" box="[151,292,1385,1409]" direction="east" orientation="longitude" pageId="11" pageNumber="150" precision="15" value="119.24">119º14'24&quot;E</geoCoordinate>
), Western
<collectingCountry id="F326764DFFA3FF8DF388F36AFDAAAE81" box="[411,517,1384,1409]" name="Australia" pageId="11" pageNumber="150">Australia</collectingCountry>
,
<collectingCountry id="F326764DFFA3FF8DF003F36AFDD6AE81" box="[528,633,1384,1409]" name="Australia" pageId="11" pageNumber="150">Australia</collectingCountry>
, R. Teale &amp; G. Harold,
<date id="FF8F101DFFA3FF8DF192F36BFB8CAE81" box="[897,1059,1384,1409]" pageId="11" pageNumber="150" value="2003-04-28">28 April 2003</date>
. Fixed in 10% formalin, stored in 70% ethanol, liver sample stored in –80ºC ultrafreezer at
<collectionCode id="ED20AE18FFA3FF8DF123F38FFCD7AEA5" LSID="urn:lsid:biocol.org:col:34981" box="[816,888,1421,1445]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="11" pageNumber="150">WAM</collectionCode>
.
<emphasis id="B945EACFFFA3FF8DF197F38EFBAAAEA5" bold="true" box="[900,1029,1420,1445]" pageId="11" pageNumber="150">
<typeStatus id="548A887FFFA3FF8DF197F38EFBAFAEA5" box="[900,1024,1420,1445]" pageId="11" pageNumber="150" type="paratype">Paratypes</typeStatus>
:
</emphasis>
All from Western
<collectingCountry id="F326764DFFA3FF8DF6F3F38EFAE5AEA5" box="[1248,1354,1420,1445]" name="Australia" pageId="11" pageNumber="150">Australia</collectingCountry>
.
<collectionCode id="ED20AE18FFA3FF8DF744F38FFA33AEA5" LSID="urn:lsid:biocol.org:col:34981" box="[1367,1436,1421,1445]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="11" pageNumber="150">WAM</collectionCode>
R125456 (male)—
<quantity id="4CC99B38FFA3FF8DF37FF3B2FE1AAEC9" box="[364,437,1456,1481]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="11" pageNumber="150" unit="km" value="30.0">30 km</quantity>
E Newman (
<geoCoordinate id="EE05501AFFA3FF8DF05DF3B3FD0AAEC9" box="[590,677,1457,1481]" direction="south" orientation="latitude" pageId="11" pageNumber="150" precision="925" value="-23.316668">23º19'S</geoCoordinate>
<geoCoordinate id="EE05501AFFA3FF8DF0BDF3B3FCB9AEC9" box="[686,790,1457,1481]" direction="east" orientation="longitude" pageId="11" pageNumber="150" precision="925" value="120.03333">120º02'E</geoCoordinate>
);
<collectionCode id="ED20AE18FFA3FF8DF13EF3B3FCDEAEC9" LSID="urn:lsid:biocol.org:col:34981" box="[813,881,1457,1481]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="11" pageNumber="150">WAM</collectionCode>
R125766 (female)—Circular Pool, Dales Gorge (
<geoCoordinate id="EE05501AFFA3FF8DF28CF3D7FEB4AEED" box="[159,283,1492,1517]" direction="south" orientation="latitude" pageId="11" pageNumber="150" precision="15" value="-22.643333">22º38'36&quot;S</geoCoordinate>
<geoCoordinate id="EE05501AFFA3FF8DF335F3D7FE1CAEED" box="[294,435,1493,1517]" direction="east" orientation="longitude" pageId="11" pageNumber="150" precision="15" value="118.56306">118º33'47&quot;E</geoCoordinate>
);
<collectionCode id="ED20AE18FFA3FF8DF3DFF3D7FDBFAEED" LSID="urn:lsid:biocol.org:col:34981" box="[460,528,1493,1517]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="11" pageNumber="150">WAM</collectionCode>
R129628 (male)—
<quantity id="4CC99B38FFA3FF8DF0E1F3D6FCE5AEED" box="[754,842,1492,1517]" metricMagnitude="5" metricUnit="m" metricValue="1.2" pageId="11" pageNumber="150" unit="km" value="120.0">120 km</quantity>
NW Newman (
<geoCoordinate id="EE05501AFFA3FF8DF616F3D6FBF3AEED" box="[1029,1116,1492,1517]" direction="south" orientation="latitude" pageId="11" pageNumber="150" precision="925" value="-22.916666">22º55'S</geoCoordinate>
<geoCoordinate id="EE05501AFFA3FF8DF675F3D7FB62AEED" box="[1126,1229,1493,1517]" direction="east" orientation="longitude" pageId="11" pageNumber="150" precision="925" value="119.01667">119º01'E</geoCoordinate>
);
<collectionCode id="ED20AE18FFA3FF8DF6F5F3D7FA85AEED" LSID="urn:lsid:biocol.org:col:34981" box="[1254,1322,1493,1517]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="11" pageNumber="150">WAM</collectionCode>
R
<date id="FF8F101DFFA3FF8DF75BF3D7FA33AEED" box="[1352,1436,1493,1517]" pageId="11" pageNumber="150" value="34-07-14">140734</date>
(female)—Rocklea Station (
<geoCoordinate id="EE05501AFFA3FF8DF3FFF3FAFDC6AD11" box="[492,617,1528,1553]" direction="south" orientation="latitude" pageId="11" pageNumber="150" precision="15" value="-22.94861">22º56'55&quot;S</geoCoordinate>
117º16'0 7&quot;E);
<collectionCode id="ED20AE18FFA3FF8DF138F3FBFCC0AD11" LSID="urn:lsid:biocol.org:col:34981" box="[811,879,1529,1553]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="11" pageNumber="150">WAM</collectionCode>
R164579 (male)—
<quantity id="4CC99B38FFA3FF8DF677F3FAFB11AD11" box="[1124,1214,1528,1553]" metricMagnitude="3" metricUnit="m" metricValue="8.5" pageId="11" pageNumber="150" unit="km" value="8.5">8.5 km</quantity>
NE Mount Rica (21º57'0 8&quot;S
<geoCoordinate id="EE05501AFFA3FF8DF330F01EFE1EAD35" box="[291,433,1564,1589]" direction="east" orientation="longitude" pageId="11" pageNumber="150" precision="15" value="116.48833">116º29'18&quot;E</geoCoordinate>
).
</paragraph>
</subSubSection>
<subSubSection id="C32B6556FFA3FF8DF2D4F042FE35ACE5" pageId="11" pageNumber="150" type="diagnosis">
<paragraph id="8B8E36DDFFA3FF8DF2D4F042FEB6AC55" blockId="11.[151,1437,1348,2022]" pageId="11" pageNumber="150">
<emphasis id="B945EACFFFA3FF8DF2D4F042FDA8AD59" bold="true" box="[199,519,1600,1625]" pageId="11" pageNumber="150">Diagnosis and comparison.</emphasis>
Differs from all Australian congeners by the following combination of characters: size moderately small (SVL up to
<quantity id="4CC99B38FFA3FF8DF005F066FDD8AD7D" box="[534,631,1636,1661]" metricMagnitude="-1" metricUnit="m" metricValue="1.6" pageId="11" pageNumber="150" unit="mm" value="160.0">160 mm</quantity>
); slender build; tail long (between 168–207% of SVL) and thin, circular in cross section at midpoint, midbody scales 120–122; dorsal and lateral scales weakly keeled, enlarged keeled ventrolateral scales on each side of and posterior to vent in 4–6 rows, scales on top of head smooth; supraoculars gradually merging with larger interoculars; dorsal scales elongate, ovate; nostrils high and oriented dorsolaterally, dorsal and lateral surfaces of body with subdued irregular dark brown to black spots or vermiculations on a dark reddish-brown background, small ocelli on the dorsal surface of the hindlimbs only, and tail without prominent dark bands.
</paragraph>
<paragraph id="8B8E36DDFFA3FF8DF2D4F160FE35ACE5" blockId="11.[151,1437,1348,2022]" pageId="11" pageNumber="150">
<taxonomicName id="4C314D5EFFA3FF8DF2D4F160FE79AC79" box="[199,470,1888,1913]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="150" phylum="Chordata" rank="species" species="hamersleyensis" status="sp. nov.">
<emphasis id="B945EACFFFA3FF8DF2D4F160FE79AC79" box="[199,470,1888,1913]" italics="true" pageId="11" pageNumber="150">Varanus hamersleyensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27657B4FFA3FF8DF3CDF162FD97AC79" box="[478,568,1888,1913]" pageId="11" pageNumber="150" rank="species">
<emphasis id="B945EACFFFA3FF8DF3CDF162FD97AC79" bold="true" box="[478,568,1888,1913]" pageId="11" pageNumber="150">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from its sister species
<taxonomicName id="4C314D5EFFA3FF8DF19CF160FB88AC79" box="[911,1063,1888,1913]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="150" phylum="Chordata" rank="species" species="pilbarensis">
<emphasis id="B945EACFFFA3FF8DF19CF160FB88AC79" box="[911,1063,1888,1913]" italics="true" pageId="11" pageNumber="150">V. pilbarensis</emphasis>
</taxonomicName>
in its overall darker colouration, presence of small whitish ocelli usually on the dorsal surface of the limbs only (
<taxonomicName id="4C314D5EFFA3FF8DF607F187FBC8AC9D" box="[1044,1127,1924,1949]" pageId="11" pageNumber="150" rank="variety" variety="large">v. large</taxonomicName>
greyish ocelli on the dorsal and lateral surfaces of body and limbs), largely unbanded tail (
<taxonomicName id="4C314D5EFFA3FF8DF15CF1ABFC6AACC1" box="[847,965,1960,1985]" pageId="11" pageNumber="150" rank="variety" variety="strongly">v. strongly</taxonomicName>
banded tail) and smaller and less elongate dorsal scales (
<figureCitation id="130A2A58FFA3FF8DF327F1CFFED8ACE5" box="[308,375,1997,2022]" captionStart="FIGURE 3" captionStartId="5.[151,250,741,763]" captionTargetBox="[313,1273,193,701]" captionTargetId="figure@5.[313,1273,193,702]" captionTargetPageId="5" captionText="FIGURE 3. Details of primary dorsal scales and granules in (A) WAM R 102103, V. pilbarensis and (B) WAM R 129628, V. hamersleyensis sp. nov .. Scale bar 1 cm." httpUri="https://zenodo.org/record/252059/files/figure.png" pageId="11" pageNumber="150">Fig. 3</figureCitation>
B).
</paragraph>
</subSubSection>
<caption id="DF4E6655FFA4FF8AF284F057FAB6AD6B" box="[151,1305,1621,1644]" httpUri="https://zenodo.org/record/252064/files/figure.png" pageId="12" pageNumber="151" targetBox="[151,1436,176,1588]" targetPageId="12">
<paragraph id="8B8E36DDFFA4FF8AF284F057FAB6AD6B" blockId="12.[151,1305,1621,1644]" box="[151,1305,1621,1644]" pageId="12" pageNumber="151">
<emphasis id="B945EACFFFA4FF8AF284F057FEBDAD6B" bold="true" box="[151,274,1621,1643]" pageId="12" pageNumber="151">FIGURE 8.</emphasis>
Preserved holotype WAM R145733 of
<taxonomicName id="4C314D5EFFA4FF8AF0B7F055FCF7AD6B" box="[676,856,1621,1644]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="151" phylum="Chordata" rank="species" species="hamersleyensis" status="sp. nov.">
<emphasis id="B945EACFFFA4FF8AF0B7F055FCF7AD6B" box="[676,856,1621,1644]" italics="true" pageId="12" pageNumber="151">V. hamersleyensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27657B4FFA4FF8AF14DF054FC00AD6B" box="[862,943,1622,1643]" pageId="12" pageNumber="151" rank="species">
<emphasis id="B945EACFFFA4FF8AF14DF054FC00AD6B" bold="true" box="[862,943,1622,1643]" pageId="12" pageNumber="151">sp. nov.</emphasis>
</taxonomicNameLabel>
in dorsal, ventral and lateral views.
</paragraph>
</caption>
<subSubSection id="C32B6556FFA4FF96F2D4F09AFE45AAAD" lastPageId="16" lastPageNumber="155" pageId="12" pageNumber="151" type="description">
<paragraph id="8B8E36DDFFA4FF8AF2D4F09AFE33ACF5" blockId="12.[151,1437,1688,2037]" pageId="12" pageNumber="151">
<emphasis id="B945EACFFFA4FF8AF2D4F09AFE72ADB1" bold="true" box="[199,477,1688,1713]" pageId="12" pageNumber="151">
Description of
<typeStatus id="548A887FFFA4FF8AF365F09AFE72ADB1" box="[374,477,1688,1713]" pageId="12" pageNumber="151" type="holotype">holotype</typeStatus>
</emphasis>
(
<figureCitation id="130A2A58FFA4FF8AF3FEF09AFD80ADB1" box="[493,559,1688,1713]" captionStart="FIGURE 8" captionStartId="12.[151,250,1621,1643]" captionTargetBox="[151,1436,176,1588]" captionTargetId="figure@12.[151,1436,157,1588]" captionTargetPageId="12" captionText="FIGURE 8. Preserved holotype WAM R 145733 of V. hamersleyensis sp. nov. in dorsal, ventral and lateral views." httpUri="https://zenodo.org/record/252064/files/figure.png" pageId="12" pageNumber="151">Fig. 8</figureCitation>
). Small-bodied
<taxonomicName id="4C314D5EFFA4FF8AF0F7F09BFCEFADB0" box="[740,832,1689,1712]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="151" phylum="Chordata" rank="genus">
<emphasis id="B945EACFFFA4FF8AF0F7F09BFCEFADB0" box="[740,832,1689,1712]" italics="true" pageId="12" pageNumber="151">Varanus</emphasis>
</taxonomicName>
with depressed body. Head elongate and depressed in lateral view; upper rostrum slightly protruding over lower jaw, genial groove distinct. Tail long and gradually tapering, moderately dorsoventrally depressed proximally, circular centrally, becoming more triangular distally. Eyes large with scaly upper and lower eyelids; closer to nostril than to ear opening; supraocular brow present above eyes. Snout moderately long, rounded in dorsal and lateral views; nostrils large and visible from above, wider than high, oriented dorsolaterally, approximately equidistant between tip of snout and anterior edge of eye. Ear openings much higher than wide, oriented distally at 45º to the corner of jaw. Head scales small, smooth and unornamented; supraocular scales smaller than those on the rostrum, frontal and parietal regions; canthus rostralis weak; rostral scale twice as large as adjacent scales, almost rounded, slightly narrower anteriorly; mental scale higher than wide and almost rectangular.
</paragraph>
<paragraph id="8B8E36DDFFA5FF8BF2D4F695FCB8ABF8" blockId="13.[151,1437,151,1040]" pageId="13" pageNumber="152">Limbs well-developed and moderately slender; ventral surface of digits with enlarged paired circular lamellae, scales on manus and pes otherwise consisting of conical tubercles; claws darkly pigmented, relatively short, very thick, laterally compressed and with sharp, recurved tips.</paragraph>
<paragraph id="8B8E36DDFFA5FF8BF2D4F706FEF4AAF5" blockId="13.[151,1437,151,1040]" pageId="13" pageNumber="152">Dorsal primary scales small, non-overlapping, relatively uniform and ovate, approximately twice as long as wide and with a small central keel; bordered by row of small granules; ventral scales non-overlapping and more rectangular, approximately 50% longer and 100% wider than dorsal scales, grading smaller on the throat and vent; caudal scales undifferentiated on dorsal and ventral surfaces, at base of tail similar in size to dorsal scales on torso, grading to larger with moderate keels in central section of tail and narrow with sharp keels towards tip; cloacal region with paired set of 4-6 curved lateroventral rows of long, narrow pointed scales, increasing in height, size and spinosity distally.</paragraph>
<paragraph id="8B8E36DDFFA5FF8BF2D4F402FC95A9A8" blockId="13.[151,1437,151,1040]" pageId="13" pageNumber="152">In preservative, ground colour of dorsum dark brown, slightly lighter on head and neck. Patterning on head includes black spots on some individual scales and dark brown labial bars, dorsal surfaces of torso with mixture of irregular black spots on individual scales or spots coalesce to form narrow, wavy transverse lines. Light brown tail with irregular dark brown to black spots. Forelimbs light brown, hindlimbs darker brown with small whitish ocelli becoming more prominent distally. Ventral surfaces greyish.</paragraph>
<paragraph id="8B8E36DDFFA5FF8BF2D4F4B6FAD2A8A5" blockId="13.[151,1437,151,1040]" pageId="13" pageNumber="152">
<emphasis id="B945EACFFFA5FF8BF2D4F4B6FE90A9CD" bold="true" box="[199,319,692,717]" pageId="13" pageNumber="152">Variation.</emphasis>
Summary mensural and meristic data for the
<typeStatus id="548A887FFFA5FF8BF156F4B7FCDAA9CD" box="[837,885,693,717]" pageId="13" pageNumber="152">type</typeStatus>
series of
<taxonomicName id="4C314D5EFFA5FF8BF1F7F4B7FB03A9CD" box="[996,1196,692,717]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="152" phylum="Chordata" rank="species" species="hamersleyensis" status="sp. nov.">
<emphasis id="B945EACFFFA5FF8BF1F7F4B7FB03A9CD" box="[996,1196,692,717]" italics="true" pageId="13" pageNumber="152">V. hamersleyensis</emphasis>
</taxonomicName>
<emphasis id="B945EACFFFA5FF8BF6A7F4B6FAA1A9CD" bold="true" box="[1204,1294,692,717]" pageId="13" pageNumber="152">
<taxonomicNameLabel id="A27657B4FFA5FF8BF6A7F4B6FAA1A9CD" box="[1204,1294,692,717]" pageId="13" pageNumber="152" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is presented in
<tableCitation id="C6B30366FFA5FF8BF2A7F4D5FEBCA9F0" box="[180,275,727,752]" captionStart="TABLE 1" captionStartId="9.[151,239,152,175]" captionTargetBox="[159,1375,276,762]" captionTargetPageId="9" captionText="TABLE 1. The mean, range, ± one standard deviation (SD) and sample size (N) of the principal morphological measurements (mm) and counts for V. pilbarensis and V. hamersleyensis sp. nov." httpUri="http://table.plazi.org/id/DF4E6655FFA1FF8FF284F69AFC5BABCF" pageId="13" pageNumber="152" tableUuid="DF4E6655FFA1FF8FF284F69AFC5BABCF">Tables 1</tableCitation>
and
<tableCitation id="C6B30366FFA5FF8BF358F4DAFEF4A9F0" box="[331,347,728,752]" captionStart="TABLE 3" captionStartId="13.[151,239,1081,1104]" captionTargetBox="[156,1412,1164,1434]" captionTargetPageId="13" captionText="TABLE 3. Individual measurements (mm) and meristic counts for the type series of V. hamersleyensis sp. nov.. NA = not available." httpUri="http://table.plazi.org/id/DF4E6655FFA5FF8BF284F23BFE88AF70" pageId="13" pageNumber="152" tableUuid="DF4E6655FFA5FF8BF284F23BFE88AF70">3</tableCitation>
. In preservative, the ground colouration is always brown (
<figureCitation id="130A2A58FFA5FF8BF1FDF4D5FBEDA9F0" box="[1006,1090,727,752]" captionStart="FIGURE 10" captionStartId="15.[151,250,1045,1067]" captionTargetBox="[151,1436,176,1023]" captionTargetId="figure@15.[151,1436,167,1023]" captionTargetPageId="15" captionText="FIGURE 10. Variation in V. hamersleyensis sp. nov. from left to right, WAM R 125456, WAM R 125766, WAM R 140734, WAM R 164579." httpUri="https://zenodo.org/record/252066/files/figure.png" pageId="13" pageNumber="152">Fig. 10</figureCitation>
). All specimens have irregular dark spots on some individual scales or vermiculations across the dorsum, although these vary in degree and intensity; on one individual (WAM R125766) the entire dorsum has very fine spots with barely any indication of alignment, while at the other extreme (WAM R
<date id="FF8F101DFFA5FF8BF0AFF546FCBEA85C" box="[700,785,836,860]" pageId="13" pageNumber="152" value="34-07-14">140734</date>
) the dense spots align to form regular wavy lines. Most individuals have some dark spotting on the tail, although one individual (WAM R164579) has almost none. The venter sometimes features relatively bold greyish bands that become more narrow and wavy on neck and throat.
</paragraph>
<paragraph id="8B8E36DDFFA5FF8BF2D4F5B2FE3EAF10" blockId="13.[151,1437,151,1040]" pageId="13" pageNumber="152">
Two juvenile specimens (WAM R69695: SVL
<quantity id="4CC99B38FFA5FF8BF0C5F5ADFC92A8C7" box="[726,829,943,968]" metricMagnitude="-2" metricUnit="m" metricValue="6.16" pageId="13" pageNumber="152" unit="mm" value="61.6">61.6 mm</quantity>
; WAM R125100:
<quantity id="4CC99B38FFA5FF8BF619F5B2FBF4A8C8" box="[1034,1115,944,968]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="13" pageNumber="152" unit="mm" value="70.0">70 mm</quantity>
) are more brightly patterned than adults with much larger spots on dorsum, more contrasting and strongly defined ventral surface bands and obviously ringed tails.
</paragraph>
<caption id="DF4E6655FFA5FF8BF284F23BFE88AF70" ID-Table-UUID="DF4E6655FFA5FF8BF284F23BFE88AF70" httpUri="http://table.plazi.org/id/DF4E6655FFA5FF8BF284F23BFE88AF70" pageId="13" pageNumber="152" targetBox="[156,1412,1164,1434]" targetIsTable="true" targetPageId="13">
<paragraph id="8B8E36DDFFA5FF8BF284F23BFE88AF70" blockId="13.[151,1435,1080,1136]" pageId="13" pageNumber="152">
<emphasis id="B945EACFFFA5FF8BF284F23BFEA4AF50" bold="true" box="[151,267,1081,1104]" pageId="13" pageNumber="152">TABLE 3.</emphasis>
Individual measurements (mm) and meristic counts for the type series of
<taxonomicName id="4C314D5EFFA5FF8BF621F23BFB5FAF50" box="[1074,1264,1080,1104]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="152" phylum="Chordata" rank="species" species="hamersleyensis" status="sp. nov.">
<emphasis id="B945EACFFFA5FF8BF621F23BFB5FAF50" box="[1074,1264,1080,1104]" italics="true" pageId="13" pageNumber="152">V. hamersleyensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27657B4FFA5FF8BF6EAF23BFAE0AF50" box="[1273,1359,1081,1104]" pageId="13" pageNumber="152" rank="species">
<emphasis id="B945EACFFFA5FF8BF6EAF23BFAE0AF50" bold="true" box="[1273,1359,1081,1104]" pageId="13" pageNumber="152">sp. nov.</emphasis>
</taxonomicNameLabel>
. NA = not available.
</paragraph>
</caption>
<paragraph id="8B8E36DDFFA5FF8BF28FF28EFADCAE9A" pageId="13" pageNumber="152">
<table id="F931C47DFFA50079F28FF28EFA2BAE9A" box="[156,1412,1164,1434]" gridcols="12" gridrows="7" pageId="13" pageNumber="152">
<tr id="3501349FFFA50079F28FF28EFA2BAFA2" box="[156,1412,1164,1186]" gridrow="0" pageId="13" pageNumber="152">
<th id="76D05DE3FFA50079F28FF28EFF55AFA2" box="[156,250,1164,1186]" gridcol="0" gridrow="0" pageId="13" pageNumber="152">WAM #</th>
<th id="76D05DE3FFA50079F333F28EFECAAFA2" box="[288,357,1164,1186]" gridcol="1" gridrow="0" pageId="13" pageNumber="152">Sex</th>
<th id="76D05DE3FFA50079F380F28EFE63AFA2" box="[403,460,1164,1186]" gridcol="2" gridrow="0" pageId="13" pageNumber="152">SVL</th>
<th id="76D05DE3FFA50079F014F28EFD82AFA2" box="[519,557,1164,1186]" gridcol="3" gridrow="0" pageId="13" pageNumber="152">TL</th>
<th id="76D05DE3FFA50079F047F28EFD27AFA2" box="[596,648,1164,1186]" gridcol="4" gridrow="0" pageId="13" pageNumber="152">HNL</th>
<th id="76D05DE3FFA50079F0A6F28EFD4EAFA2" box="[693,737,1164,1186]" gridcol="5" gridrow="0" pageId="13" pageNumber="152">FLL</th>
<th id="76D05DE3FFA50079F13BF28EFCF5AFA2" box="[808,858,1164,1186]" gridcol="6" gridrow="0" pageId="13" pageNumber="152">HLL</th>
<th id="76D05DE3FFA50079F18FF28EFC6EAFA2" box="[924,961,1164,1186]" gridcol="7" gridrow="0" pageId="13" pageNumber="152">HD</th>
<th id="76D05DE3FFA50079F1EEF28EFB86AFA2" box="[1021,1065,1164,1186]" gridcol="8" gridrow="0" pageId="13" pageNumber="152">HW</th>
<th id="76D05DE3FFA50079F64EF28EFB4FAFA2" box="[1117,1248,1164,1186]" gridcol="9" gridrow="0" pageId="13" pageNumber="152">BL MSR</th>
<th id="76D05DE3FFA50079F718F28EFA83AFA2" box="[1291,1324,1164,1186]" gridcol="10" gridrow="0" pageId="13" pageNumber="152">VS</th>
<th id="76D05DE3FFA50079F74AF28EFA2BAFA2" box="[1369,1412,1164,1186]" gridcol="11" gridrow="0" pageId="13" pageNumber="152">PSL</th>
</tr>
<tr id="3501349FFFA50079F28FF2B7FA2BAFCB" box="[156,1412,1205,1227]" gridrow="1" pageId="13" pageNumber="152">
<th id="76D05DE3FFA50079F28FF2B7FF55AFCB" box="[156,250,1205,1227]" gridcol="0" gridrow="1" pageId="13" pageNumber="152">R125456</th>
<td id="76D05DE3FFA50079F333F2B7FECAAFCB" box="[288,357,1205,1227]" gridcol="1" gridrow="1" pageId="13" pageNumber="152">male</td>
<td id="76D05DE3FFA50079F380F2B7FE63AFCB" box="[403,460,1205,1227]" gridcol="2" gridrow="1" pageId="13" pageNumber="152">122.3</td>
<td id="76D05DE3FFA50079F014F2B7FD82AFCB" box="[519,557,1205,1227]" gridcol="3" gridrow="1" pageId="13" pageNumber="152">NA</td>
<td id="76D05DE3FFA50079F047F2B7FD27AFCB" box="[596,648,1205,1227]" gridcol="4" gridrow="1" pageId="13" pageNumber="152">46.7</td>
<td id="76D05DE3FFA50079F0A6F2B7FD4EAFCB" box="[693,737,1205,1227]" gridcol="5" gridrow="1" pageId="13" pageNumber="152">36.4</td>
<td id="76D05DE3FFA50079F13BF2B7FCF5AFCB" box="[808,858,1205,1227]" gridcol="6" gridrow="1" pageId="13" pageNumber="152">45.5</td>
<td id="76D05DE3FFA50079F18FF2B7FC6EAFCB" box="[924,961,1205,1227]" gridcol="7" gridrow="1" pageId="13" pageNumber="152">7.5</td>
<td id="76D05DE3FFA50079F1EEF2B7FB86AFCB" box="[1021,1065,1205,1227]" gridcol="8" gridrow="1" pageId="13" pageNumber="152">15</td>
<td id="76D05DE3FFA50079F64EF2B7FB4FAFCB" box="[1117,1248,1205,1227]" gridcol="9" gridrow="1" pageId="13" pageNumber="152">75.6 120</td>
<td id="76D05DE3FFA50079F718F2B7FA83AFCB" box="[1291,1324,1205,1227]" gridcol="10" gridrow="1" pageId="13" pageNumber="152">70</td>
<td id="76D05DE3FFA50079F74AF2B7FA2BAFCB" box="[1369,1412,1205,1227]" gridcol="11" gridrow="1" pageId="13" pageNumber="152">22</td>
</tr>
<tr id="3501349FFFA50079F28FF2DCFA2BAFF4" box="[156,1412,1246,1268]" gridrow="2" pageId="13" pageNumber="152">
<th id="76D05DE3FFA50079F28FF2DCFF55AFF4" box="[156,250,1246,1268]" gridcol="0" gridrow="2" pageId="13" pageNumber="152">R125766</th>
<td id="76D05DE3FFA50079F333F2DCFECAAFF4" box="[288,357,1246,1268]" gridcol="1" gridrow="2" pageId="13" pageNumber="152">female</td>
<td id="76D05DE3FFA50079F380F2DCFE63AFF4" box="[403,460,1246,1268]" gridcol="2" gridrow="2" pageId="13" pageNumber="152">121.2</td>
<td id="76D05DE3FFA50079F014F2DCFD82AFF4" box="[519,557,1246,1268]" gridcol="3" gridrow="2" pageId="13" pageNumber="152">217</td>
<td id="76D05DE3FFA50079F047F2DCFD27AFF4" box="[596,648,1246,1268]" gridcol="4" gridrow="2" pageId="13" pageNumber="152">48.7</td>
<td id="76D05DE3FFA50079F0A6F2DCFD4EAFF4" box="[693,737,1246,1268]" gridcol="5" gridrow="2" pageId="13" pageNumber="152">39.4</td>
<td id="76D05DE3FFA50079F13BF2DCFCF5AFF4" box="[808,858,1246,1268]" gridcol="6" gridrow="2" pageId="13" pageNumber="152">49</td>
<td id="76D05DE3FFA50079F18FF2DCFC6EAFF4" box="[924,961,1246,1268]" gridcol="7" gridrow="2" pageId="13" pageNumber="152">6</td>
<td id="76D05DE3FFA50079F1EEF2DCFB86AFF4" box="[1021,1065,1246,1268]" gridcol="8" gridrow="2" pageId="13" pageNumber="152">13.2</td>
<td id="76D05DE3FFA50079F64EF2DCFB4FAFF4" box="[1117,1248,1246,1268]" gridcol="9" gridrow="2" pageId="13" pageNumber="152">72.5 120</td>
<td id="76D05DE3FFA50079F718F2DCFA83AFF4" box="[1291,1324,1246,1268]" gridcol="10" gridrow="2" pageId="13" pageNumber="152">66</td>
<td id="76D05DE3FFA50079F74AF2DCFA2BAFF4" box="[1369,1412,1246,1268]" gridcol="11" gridrow="2" pageId="13" pageNumber="152">22</td>
</tr>
<tr id="3501349FFFA50079F28FF30AFA2BAE1E" box="[156,1412,1288,1310]" gridrow="3" pageId="13" pageNumber="152">
<th id="76D05DE3FFA50079F28FF30AFF55AE1E" box="[156,250,1288,1310]" gridcol="0" gridrow="3" pageId="13" pageNumber="152">R129628</th>
<td id="76D05DE3FFA50079F333F30AFECAAE1E" box="[288,357,1288,1310]" gridcol="1" gridrow="3" pageId="13" pageNumber="152">male</td>
<td id="76D05DE3FFA50079F380F30AFE63AE1E" box="[403,460,1288,1310]" gridcol="2" gridrow="3" pageId="13" pageNumber="152">160</td>
<td id="76D05DE3FFA50079F014F30AFD82AE1E" box="[519,557,1288,1310]" gridcol="3" gridrow="3" pageId="13" pageNumber="152">270</td>
<td id="76D05DE3FFA50079F047F30AFD27AE1E" box="[596,648,1288,1310]" gridcol="4" gridrow="3" pageId="13" pageNumber="152">64</td>
<td id="76D05DE3FFA50079F0A6F30AFD4EAE1E" box="[693,737,1288,1310]" gridcol="5" gridrow="3" pageId="13" pageNumber="152">44.4</td>
<td id="76D05DE3FFA50079F13BF30AFCF5AE1E" box="[808,858,1288,1310]" gridcol="6" gridrow="3" pageId="13" pageNumber="152">64</td>
<td id="76D05DE3FFA50079F18FF30AFC6EAE1E" box="[924,961,1288,1310]" gridcol="7" gridrow="3" pageId="13" pageNumber="152">9.4</td>
<td id="76D05DE3FFA50079F1EEF30AFB86AE1E" box="[1021,1065,1288,1310]" gridcol="8" gridrow="3" pageId="13" pageNumber="152">19.8</td>
<td id="76D05DE3FFA50079F64EF30AFB4FAE1E" box="[1117,1248,1288,1310]" gridcol="9" gridrow="3" pageId="13" pageNumber="152">95.9 122</td>
<td id="76D05DE3FFA50079F718F30AFA83AE1E" box="[1291,1324,1288,1310]" gridcol="10" gridrow="3" pageId="13" pageNumber="152">70</td>
<td id="76D05DE3FFA50079F74AF30AFA2BAE1E" box="[1369,1412,1288,1310]" gridcol="11" gridrow="3" pageId="13" pageNumber="152">21</td>
</tr>
<tr id="3501349FFFA50079F28FF333FA2BAE47" box="[156,1412,1329,1351]" gridrow="4" pageId="13" pageNumber="152">
<th id="76D05DE3FFA50079F28FF333FF55AE47" box="[156,250,1329,1351]" gridcol="0" gridrow="4" pageId="13" pageNumber="152">R140734</th>
<td id="76D05DE3FFA50079F333F333FECAAE47" box="[288,357,1329,1351]" gridcol="1" gridrow="4" pageId="13" pageNumber="152">female</td>
<td id="76D05DE3FFA50079F380F333FE63AE47" box="[403,460,1329,1351]" gridcol="2" gridrow="4" pageId="13" pageNumber="152">120</td>
<td id="76D05DE3FFA50079F014F333FD82AE47" box="[519,557,1329,1351]" gridcol="3" gridrow="4" pageId="13" pageNumber="152">NA</td>
<td id="76D05DE3FFA50079F047F333FD27AE47" box="[596,648,1329,1351]" gridcol="4" gridrow="4" pageId="13" pageNumber="152">47.3</td>
<td id="76D05DE3FFA50079F0A6F333FD4EAE47" box="[693,737,1329,1351]" gridcol="5" gridrow="4" pageId="13" pageNumber="152">33.7</td>
<td id="76D05DE3FFA50079F13BF333FCF5AE47" box="[808,858,1329,1351]" gridcol="6" gridrow="4" pageId="13" pageNumber="152">39.4</td>
<td id="76D05DE3FFA50079F18FF333FC6EAE47" box="[924,961,1329,1351]" gridcol="7" gridrow="4" pageId="13" pageNumber="152">6.5</td>
<td id="76D05DE3FFA50079F1EEF333FB86AE47" box="[1021,1065,1329,1351]" gridcol="8" gridrow="4" pageId="13" pageNumber="152">13.4</td>
<td id="76D05DE3FFA50079F64EF333FB4FAE47" box="[1117,1248,1329,1351]" gridcol="9" gridrow="4" pageId="13" pageNumber="152">72.9 122</td>
<td id="76D05DE3FFA50079F718F333FA83AE47" box="[1291,1324,1329,1351]" gridcol="10" gridrow="4" pageId="13" pageNumber="152">72</td>
<td id="76D05DE3FFA50079F74AF333FA2BAE47" box="[1369,1412,1329,1351]" gridcol="11" gridrow="4" pageId="13" pageNumber="152">21</td>
</tr>
<tr id="3501349FFFA50079F28FF358FA2BAE70" box="[156,1412,1370,1392]" gridrow="5" pageId="13" pageNumber="152">
<th id="76D05DE3FFA50079F28FF358FF55AE70" box="[156,250,1370,1392]" gridcol="0" gridrow="5" pageId="13" pageNumber="152">R145733</th>
<td id="76D05DE3FFA50079F333F358FECAAE70" box="[288,357,1370,1392]" gridcol="1" gridrow="5" pageId="13" pageNumber="152">male</td>
<td id="76D05DE3FFA50079F380F358FE63AE70" box="[403,460,1370,1392]" gridcol="2" gridrow="5" pageId="13" pageNumber="152">155.8</td>
<td id="76D05DE3FFA50079F014F358FD82AE70" box="[519,557,1370,1392]" gridcol="3" gridrow="5" pageId="13" pageNumber="152">295</td>
<td id="76D05DE3FFA50079F047F358FD27AE70" box="[596,648,1370,1392]" gridcol="4" gridrow="5" pageId="13" pageNumber="152">64.5</td>
<td id="76D05DE3FFA50079F0A6F358FD4EAE70" box="[693,737,1370,1392]" gridcol="5" gridrow="5" pageId="13" pageNumber="152">43.7</td>
<td id="76D05DE3FFA50079F13BF358FCF5AE70" box="[808,858,1370,1392]" gridcol="6" gridrow="5" pageId="13" pageNumber="152">58.4</td>
<td id="76D05DE3FFA50079F18FF358FC6EAE70" box="[924,961,1370,1392]" gridcol="7" gridrow="5" pageId="13" pageNumber="152">9.7</td>
<td id="76D05DE3FFA50079F1EEF358FB86AE70" box="[1021,1065,1370,1392]" gridcol="8" gridrow="5" pageId="13" pageNumber="152">17.7</td>
<td id="76D05DE3FFA50079F64EF358FB4FAE70" box="[1117,1248,1370,1392]" gridcol="9" gridrow="5" pageId="13" pageNumber="152">91.3 122</td>
<td id="76D05DE3FFA50079F718F358FA83AE70" box="[1291,1324,1370,1392]" gridcol="10" gridrow="5" pageId="13" pageNumber="152">74</td>
<td id="76D05DE3FFA50079F74AF358FA2BAE70" box="[1369,1412,1370,1392]" gridcol="11" gridrow="5" pageId="13" pageNumber="152">23</td>
</tr>
<tr id="3501349FFFA50079F28FF386FA2BAE9A" box="[156,1412,1412,1434]" gridrow="6" pageId="13" pageNumber="152">
<th id="76D05DE3FFA50079F28FF386FF55AE9A" box="[156,250,1412,1434]" gridcol="0" gridrow="6" pageId="13" pageNumber="152">R164579</th>
<td id="76D05DE3FFA50079F333F386FECAAE9A" box="[288,357,1412,1434]" gridcol="1" gridrow="6" pageId="13" pageNumber="152">male</td>
<td id="76D05DE3FFA50079F380F386FE63AE9A" box="[403,460,1412,1434]" gridcol="2" gridrow="6" pageId="13" pageNumber="152">124.8</td>
<td id="76D05DE3FFA50079F014F386FD82AE9A" box="[519,557,1412,1434]" gridcol="3" gridrow="6" pageId="13" pageNumber="152">226</td>
<td id="76D05DE3FFA50079F047F386FD27AE9A" box="[596,648,1412,1434]" gridcol="4" gridrow="6" pageId="13" pageNumber="152">48.9</td>
<td id="76D05DE3FFA50079F0A6F386FD4EAE9A" box="[693,737,1412,1434]" gridcol="5" gridrow="6" pageId="13" pageNumber="152">39.5</td>
<td id="76D05DE3FFA50079F13BF386FCF5AE9A" box="[808,858,1412,1434]" gridcol="6" gridrow="6" pageId="13" pageNumber="152">50.5</td>
<td id="76D05DE3FFA50079F18FF386FC6EAE9A" box="[924,961,1412,1434]" gridcol="7" gridrow="6" pageId="13" pageNumber="152">7.9</td>
<td id="76D05DE3FFA50079F1EEF386FB86AE9A" box="[1021,1065,1412,1434]" gridcol="8" gridrow="6" pageId="13" pageNumber="152">13.4</td>
<td id="76D05DE3FFA50079F64EF386FB4FAE9A" box="[1117,1248,1412,1434]" gridcol="9" gridrow="6" pageId="13" pageNumber="152">75.8 120</td>
<td id="76D05DE3FFA50079F718F386FA83AE9A" box="[1291,1324,1412,1434]" gridcol="10" gridrow="6" pageId="13" pageNumber="152">65</td>
<td id="76D05DE3FFA50079F74AF386FA2BAE9A" box="[1369,1412,1412,1434]" gridcol="11" gridrow="6" pageId="13" pageNumber="152">22</td>
</tr>
</table>
</paragraph>
<paragraph id="8B8E36DDFFA5FF8BF2D4F3D5FB60AC10" blockId="13.[151,1436,1495,2025]" pageId="13" pageNumber="152">
<emphasis id="B945EACFFFA5FF8BF2D4F3D5FE05AEF0" bold="true" box="[199,426,1495,1520]" pageId="13" pageNumber="152">Colouration in life.</emphasis>
The following description of colouration in life is based on
<figureCitation id="130A2A58FFA5FF8BF643F3D5FB33AEF0" box="[1104,1180,1495,1520]" captionStart="FIGURE 9" captionStartId="14.[151,250,1847,1869]" captionTargetBox="[161,1425,218,1791]" captionTargetId="figure@14.[151,1436,213,1803]" captionTargetPageId="14" captionText="FIGURE 9. Adult V. hamersleyensis sp. nov. photographed in life from (A) Marillana and (B) Ophthalmia Range (images—B. Maryan)." httpUri="https://zenodo.org/record/252065/files/figure.png" pageId="13" pageNumber="152">Figs. 9</figureCitation>
A, B and a photograph from Mount Channar near Paraburdoo (
<bibRefCitation id="EFA04B2CFFA5FF8BF05CF3FEFD5FAD14" author="Ehmann" box="[591,752,1532,1557]" pageId="13" pageNumber="152" refString="Ehmann, H. (1992) Encyclopedia of Australian Animals - Reptiles. Angus &amp; Robertson, New South Wales, Australia, 495 pp." type="book" year="1992">Ehmann 1992</bibRefCitation>
). Dorsal ground colour of head, torso and limbs dark reddishbrown, often tending slightly paler on head. Head and neck with irregular subdued small blackish-brown to black spots, snout forward of nostrils often dark brown, sides of head with yellowish tinge. Labial scales with blackishbrown vertical bars. Dorsal surface of torso with numerous small blackish-brown to black spots, often coalescing into narrow (one scale wide) transverse bars or wavy lines and forming a fine black reticulum. Forelimbs reddish brown with dark flecks, hindlimbs noticeably darker brown with small whitish ocelli. Digits on both limbs with indistinct thin pale yellowish bands. Ground colour of the tail light brown, extensively overlain with darker-brown to black spots along its length. Ventral surface of the throat, neck, limbs and body greyish-white.
</paragraph>
<paragraph id="8B8E36DDFFA5FF8BF2D4F11EFC36ACE9" blockId="13.[151,1436,1495,2025]" pageId="13" pageNumber="152">
<emphasis id="B945EACFFFA5FF8BF2D4F11EFE13AC35" bold="true" box="[199,444,1820,1845]" pageId="13" pageNumber="152">Habitat and ecology.</emphasis>
<taxonomicName id="4C314D5EFFA5FF8BF3D0F11FFD7FAC35" box="[451,720,1820,1845]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="152" phylum="Chordata" rank="species" species="hamersleyensis" status="sp. nov.">
<emphasis id="B945EACFFFA5FF8BF3D0F11FFD7FAC35" box="[451,720,1820,1845]" italics="true" pageId="13" pageNumber="152">Varanus hamersleyensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27657B4FFA5FF8BF0C4F11EFC9EAC35" box="[727,817,1820,1845]" pageId="13" pageNumber="152" rank="species">
<emphasis id="B945EACFFFA5FF8BF0C4F11EFC9EAC35" bold="true" box="[727,817,1820,1845]" pageId="13" pageNumber="152">sp. nov.</emphasis>
</taxonomicNameLabel>
is strictly saxicolous and closely associated with rocky habitats such as banded ironstone rock faces and gorges (
<figureCitation id="130A2A58FFA5FF8BF135F13DFCD6AC58" box="[806,889,1855,1880]" captionStart="FIGURE 11" captionStartId="15.[151,250,1979,2001]" captionTargetBox="[151,1436,1147,1956]" captionTargetId="figure@15.[151,1436,1147,1958]" captionTargetPageId="15" captionText="FIGURE 11. Typical habitat of V. hamersleyensis sp. nov. in the Hamersley Range: a banded ironstone rock face with horizontal crevices, Mount Robinson. Note the individual in the centre, indicated by arrow (image—B. Maryan)." httpUri="https://zenodo.org/record/252067/files/figure.png" pageId="13" pageNumber="152">Fig. 11</figureCitation>
). It shelters in overhangs, crevices, cavities and occasionally under exfoliated rock slabs. Typical vegetation on these faces varies, but often includes
<taxonomicName id="4C314D5EFFA5FF8BF70CF166FA34AC7D" box="[1311,1435,1892,1917]" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" kingdom="Plantae" order="Myrtales" pageId="13" pageNumber="152" phylum="Tracheophyta" rank="genus">
<emphasis id="B945EACFFFA5FF8BF70CF166FA34AC7D" box="[1311,1435,1892,1917]" italics="true" pageId="13" pageNumber="152">Eucalyptus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C314D5EFFA5FF8BF2DAF185FE35ACA0" authority="Spinifex" authorityName="Spinifex" box="[201,410,1927,1952]" class="Liliopsida" family="Poaceae" genus="Triodia" kingdom="Animalia" order="Poales" pageId="13" pageNumber="152" phylum="Tracheophyta" rank="genus">
<emphasis id="B945EACFFFA5FF8BF2DAF185FEB3ACA0" box="[201,284,1927,1952]" italics="true" pageId="13" pageNumber="152">Triodia</emphasis>
(Spinifex)
</taxonomicName>
. Field observations indicate a preference for flat, horizontal sheet ironstone with an eastwest aspect to allow optimum basking in the morning. It is extremely swift and agile; ascending vertical rock faces with ease, and quickly takes shelter in deep crevices when disturbed.
</paragraph>
<caption id="DF4E6655FFA6FF88F284F135FF5AAC6B" httpUri="https://zenodo.org/record/252065/files/figure.png" pageId="14" pageNumber="153" targetBox="[161,1425,218,1791]" targetPageId="14">
<paragraph id="8B8E36DDFFA6FF88F284F135FF5AAC6B" blockId="14.[151,1437,1847,1899]" pageId="14" pageNumber="153">
<emphasis id="B945EACFFFA6FF88F284F135FEBDAC4C" bold="true" box="[151,274,1847,1869]" pageId="14" pageNumber="153">FIGURE 9.</emphasis>
Adult
<taxonomicName id="4C314D5EFFA6FF88F34BF13AFDA3AC4D" box="[344,524,1847,1869]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="153" phylum="Chordata" rank="species" species="hamersleyensis" status="sp. nov.">
<emphasis id="B945EACFFFA6FF88F34BF13AFDA3AC4D" box="[344,524,1847,1869]" italics="true" pageId="14" pageNumber="153">V. hamersleyensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27657B4FFA6FF88F001F135FDCDAC4C" box="[530,610,1847,1868]" pageId="14" pageNumber="153" rank="species">
<emphasis id="B945EACFFFA6FF88F001F135FDCDAC4C" bold="true" box="[530,610,1847,1868]" pageId="14" pageNumber="153">sp. nov.</emphasis>
</taxonomicNameLabel>
photographed in life from (A) Marillana and (B) Ophthalmia Range (images—B. Maryan).
</paragraph>
</caption>
<caption id="DF4E6655FFA7FF89F284F217FE90AF49" httpUri="https://zenodo.org/record/252066/files/figure.png" pageId="15" pageNumber="154" targetBox="[151,1436,176,1023]" targetPageId="15">
<paragraph id="8B8E36DDFFA7FF89F284F217FE90AF49" blockId="15.[151,1436,1044,1097]" pageId="15" pageNumber="154">
<emphasis id="B945EACFFFA7FF89F284F217FE8DAF2A" bold="true" box="[151,290,1045,1067]" pageId="15" pageNumber="154">FIGURE 10.</emphasis>
Variation in
<taxonomicName id="4C314D5EFFA7FF89F3BDF214FDCAAF2A" box="[430,613,1044,1067]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="154" phylum="Chordata" rank="species" species="hamersleyensis" status="sp. nov.">
<emphasis id="B945EACFFFA7FF89F3BDF214FDCAAF2A" box="[430,613,1044,1067]" italics="true" pageId="15" pageNumber="154">V. hamersleyensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27657B4FFA7FF89F07EF217FD6FAF2A" box="[621,704,1045,1066]" pageId="15" pageNumber="154" rank="species">
<emphasis id="B945EACFFFA7FF89F07EF217FD6FAF2A" bold="true" box="[621,704,1045,1066]" pageId="15" pageNumber="154">sp. nov.</emphasis>
</taxonomicNameLabel>
from left to right, WAM R125456, WAM R125766, WAM R140734, WAM R164579.
</paragraph>
</caption>
<caption id="DF4E6655FFA7FF89F284F1B9FAA1ACF0" httpUri="https://zenodo.org/record/252067/files/figure.png" pageId="15" pageNumber="154" targetBox="[151,1436,1147,1956]" targetPageId="15">
<paragraph id="8B8E36DDFFA7FF89F284F1B9FAA1ACF0" blockId="15.[151,1436,1979,2032]" pageId="15" pageNumber="154">
<emphasis id="B945EACFFFA7FF89F284F1B9FE8CACD0" bold="true" box="[151,291,1979,2001]" pageId="15" pageNumber="154">FIGURE 11.</emphasis>
Typical habitat of
<taxonomicName id="4C314D5EFFA7FF89F3E6F1BEFD02ACD1" box="[501,685,1979,2001]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="154" phylum="Chordata" rank="species" species="hamersleyensis" status="sp. nov.">
<emphasis id="B945EACFFFA7FF89F3E6F1BEFD02ACD1" box="[501,685,1979,2001]" italics="true" pageId="15" pageNumber="154">V. hamersleyensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27657B4FFA7FF89F0ABF1B9FCA1ACD0" box="[696,782,1979,2000]" pageId="15" pageNumber="154" rank="species">
<emphasis id="B945EACFFFA7FF89F0ABF1B9FCA1ACD0" bold="true" box="[696,782,1979,2000]" pageId="15" pageNumber="154">sp. nov.</emphasis>
</taxonomicNameLabel>
in the Hamersley Range: a banded ironstone rock face with horizontal crevices, Mount Robinson. Note the individual in the centre, indicated by arrow (image—B. Maryan).
</paragraph>
</caption>
<paragraph id="8B8E36DDFFB8FF96F2D4F69AFE45AAAD" blockId="16.[151,1436,151,789]" pageId="16" pageNumber="155">
As with
<taxonomicName id="4C314D5EFFB8FF96F336F69BFE12ABB0" box="[293,445,151,176]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="155" phylum="Chordata" rank="species" species="pilbarensis">
<emphasis id="B945EACFFFB8FF96F336F69BFE12ABB0" box="[293,445,151,176]" italics="true" pageId="16" pageNumber="155">V. pilbarensis</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFA04B2CFFB8FF96F3D9F695FCCCABB0" author="Ehmann" box="[458,867,151,176]" pageId="16" pageNumber="155" refString="Ehmann, H. (1992) Encyclopedia of Australian Animals - Reptiles. Angus &amp; Robertson, New South Wales, Australia, 495 pp." type="book" year="1992">Ehmann 1992; B. Maryan pers. obs.</bibRefCitation>
), this species has been observed active after sunset on warm, humid nights. Field observations indicated that activity periods vary seasonally, peaking during the hotter and humid months from October–April and declining when conditions are dry from May–September. The sex ratio among the
<typeStatus id="548A887FFFB8FF96F306F707FEEAAA1D" box="[277,325,261,285]" pageId="16" pageNumber="155">type</typeStatus>
series of
<taxonomicName id="4C314D5EFFB8FF96F3A6F707FDD2AA1D" box="[437,637,260,285]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="155" phylum="Chordata" rank="species" species="hamersleyensis" status="sp. nov.">
<emphasis id="B945EACFFFB8FF96F3A6F707FDD2AA1D" box="[437,637,260,285]" italics="true" pageId="16" pageNumber="155">V. hamersleyensis</emphasis>
</taxonomicName>
<emphasis id="B945EACFFFB8FF96F097F706FD48AA1D" bold="true" box="[644,743,260,285]" pageId="16" pageNumber="155">
<taxonomicNameLabel id="A27657B4FFB8FF96F097F706FD4EAA1D" box="[644,737,260,285]" pageId="16" pageNumber="155" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
including the misidentified
<typeStatus id="548A887FFFB8FF96F635F707FB28AA1D" box="[1062,1159,261,285]" pageId="16" pageNumber="155" type="paratype">paratype</typeStatus>
of
<taxonomicName id="4C314D5EFFB8FF96F6BDF707FAE8AA1D" box="[1198,1351,260,285]" class="Reptilia" family="Varanidae" genus="Varanus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="155" phylum="Chordata" rank="species" species="pilbarensis">
<emphasis id="B945EACFFFB8FF96F6BDF707FAE8AA1D" box="[1198,1351,260,285]" italics="true" pageId="16" pageNumber="155">V. pilbarensis</emphasis>
</taxonomicName>
(WAM R14901) is skewed (
<specimenCount id="9D37FD54FFB8FF96F36DF725FE64AA40" box="[382,459,295,320]" pageId="16" pageNumber="155" type="adult">5 adult</specimenCount>
males and
<specimenCount id="9D37FD54FFB8FF96F05AF72AFD39AA40" box="[585,662,295,320]" pageId="16" pageNumber="155" type="adult">2 adult</specimenCount>
females), a bias which seems to be consistent with museum series of other opportunistically collected varanids (
<bibRefCitation id="EFA04B2CFFB8FF96F095F74EFCA3AA64" author="Greer" box="[646,780,332,356]" pageId="16" pageNumber="155" refString="Greer, A. E. (1989) The Biology and Evolution of Australian Lizards. Surrey Beatty, Sydney, New South Wales, Australia, 264 pp." type="book" year="1989">Greer 1989</bibRefCitation>
) and may reflect different activity periods of the sexes. During surveys in the eastern Hamersley Range, most individuals captured were also males (B. Maryan and M. O’Connell, unpublished data).
</paragraph>
</subSubSection>
<subSubSection id="C32B6556FFB8FF96F2D4F7B5FD3CA93C" pageId="16" pageNumber="155" type="distribution">
<paragraph id="8B8E36DDFFB8FF96F2D4F7B5FD3CA93C" blockId="16.[151,1436,151,789]" pageId="16" pageNumber="155">
<emphasis id="B945EACFFFB8FF96F2D4F7B5FECFAAD0" bold="true" box="[199,352,439,464]" pageId="16" pageNumber="155">Distribution.</emphasis>
Known only from the Hamersley Range, a heavily dissected region of banded ironstone occupying the southern part of the Pilbara craton of Western
<collectingCountry id="F326764DFFB8FF96F12FF7DEFC09AAF5" box="[828,934,476,501]" name="Australia" pageId="16" pageNumber="155">Australia</collectingCountry>
. Records span the Hamersley Range for over
<quantity id="4CC99B38FFB8FF96F284F402FF43A918" box="[151,236,511,536]" metricMagnitude="5" metricUnit="m" metricValue="3.0" pageId="16" pageNumber="155" unit="km" value="300.0">300 km</quantity>
from east of Newman (WAM R125100, WAM R125456) west to Mount Rica (WAM R164579) and south to Rocklea Station (WAM R
<date id="FF8F101DFFB8FF96F3C7F426FD86A93C" box="[468,553,548,572]" pageId="16" pageNumber="155" value="34-07-14">140734</date>
) (Fig. 2).
</paragraph>
</subSubSection>
<subSubSection id="C32B6556FFB8FF96F2D4F445FBE7A9A8" pageId="16" pageNumber="155" type="etymology">
<paragraph id="8B8E36DDFFB8FF96F2D4F445FBE7A9A8" blockId="16.[151,1436,151,789]" pageId="16" pageNumber="155">
<emphasis id="B945EACFFFB8FF96F2D4F445FEE1A960" bold="true" box="[199,334,583,608]" pageId="16" pageNumber="155">Etymology.</emphasis>
Named for the Hamersley Range of Western
<collectingCountry id="F326764DFFB8FF96F142F445FC15A960" box="[849,954,583,608]" name="Australia" pageId="16" pageNumber="155">Australia</collectingCountry>
, the region to which it is restricted. This is the most prominent mountainous area in Western
<collectingCountry id="F326764DFFB8FF96F0C6F46EFC94A985" box="[725,827,620,645]" name="Australia" pageId="16" pageNumber="155">Australia</collectingCountry>
(including Mount Meharry,
<quantity id="4CC99B38FFB8FF96F695F46EFB70A984" box="[1158,1247,620,645]" metricMagnitude="3" metricUnit="m" metricValue="1.2449999999999999" pageId="16" pageNumber="155" unit="m" value="1245.0">1245 m</quantity>
, and the highest peak in the state) and home to a suite of endemic taxa. Used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C32B6556FFB8FF96F2D4F4B6FEA1A815" pageId="16" pageNumber="155" type="discussion">
<paragraph id="8B8E36DDFFB8FF96F2D4F4B6FEA1A815" blockId="16.[151,1436,151,789]" pageId="16" pageNumber="155">
<emphasis id="B945EACFFFB8FF96F2D4F4B6FE94A9CD" bold="true" box="[199,315,692,717]" pageId="16" pageNumber="155">Remarks.</emphasis>
This species appears to be secure and widespread throughout the Hamersley Range, and occurs in popular tourist destinations such as at Karijini National Park (one of the largest National Parks in Western
<collectingCountry id="F326764DFFB8FF96F284F4FEFEACA815" box="[151,259,764,789]" name="Australia" pageId="16" pageNumber="155">Australia</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>
Reference in a new issue View git blame Copy permalink