treatments-xml/data/EF/98/AB/EF98AB20DF14505B922050381FC8848E.xml

<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.183.71167" ID-GBIF-Dataset="1b90f3b6-b67e-4c54-b926-40a08baf97b7" ID-PMC="PMC8520029" ID-Pensoft-Pub="1314-2003-183-1" ID-Pensoft-UUID="A19943DE378D5AA8ADB009DC2E94C86C" ID-PubMed="34720626" ModsDocID="1314-2003-183-1" checkinTime="1633693789008" checkinUser="pensoft" docAuthor="Inuthai, Janejaree, Chantanaorrapint, Sahut, Poopath, Manop, Tetsana, Naiyana, Kiewbang, Wittawat &amp; Suddee, Somran" docDate="2021" docId="EF98AB20DF14505B922050381FC8848E" docLanguage="en" docName="PhytoKeys 183: 1-7" docOrigin="PhytoKeys 183" docPubDate="2021-10-08" docSource="http://dx.doi.org/10.3897/phytokeys.183.71167" docTitle="Corybas papillatus Inuthai, Chantanaorr. &amp; Suddee 2021, sp. nov." docType="treatment" docVersion="4" id="A19943DE378D5AA8ADB009DC2E94C86C" lastPageNumber="1" masterDocId="A19943DE378D5AA8ADB009DC2E94C86C" masterDocTitle="Corybas papillatus (Orchidaceae), a new orchid species from peninsular Thailand" masterLastPageNumber="7" masterPageNumber="1" pageNumber="1" updateTime="1668140461286" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Corybas papillatus (Orchidaceae), a new orchid species from peninsular Thailand</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Inuthai, Janejaree</mods:namePart>
<mods:affiliation>Department of Biotechnology, Faculty of Science and Technology, Thammasat University, Lampang Campus, Hang Chat, Lampang, 52190, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Chantanaorrapint, Sahut</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-9739-0994</mods:nameIdentifier>
<mods:affiliation>PSU Herbarium, Division of Biological Science, Faculty of Science, Prince of Songkla University, Hat Yai, Songkhla, 90110, Thailand</mods:affiliation>
<mods:nameIdentifier type="email">sahut.c@psu.ac.th</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Poopath, Manop</mods:namePart>
<mods:affiliation>Forest Herbarium, Department of National Parks, Wildlife and Plant Conservation, 61 Phahonyothin Road, Chatuchak, Bangkok 10900, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tetsana, Naiyana</mods:namePart>
<mods:affiliation>Forest Herbarium, Department of National Parks, Wildlife and Plant Conservation, 61 Phahonyothin Road, Chatuchak, Bangkok 10900, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kiewbang, Wittawat</mods:namePart>
<mods:affiliation>Forest Economics Bureau, Royal Forest Department, Bangkok 10900, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Suddee, Somran</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4031-9721</mods:nameIdentifier>
<mods:affiliation>Forest Herbarium, Department of National Parks, Wildlife and Plant Conservation, 61 Phahonyothin Road, Chatuchak, Bangkok 10900, Thailand</mods:affiliation>
</mods:name>
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<mods:titleInfo>
<mods:title>PhytoKeys</mods:title>
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<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-10-08</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>183</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>7</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/phytokeys.183.71167</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.183.71167</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-183-1</mods:identifier>
<mods:identifier type="Pensoft-UUID">A19943DE378D5AA8ADB009DC2E94C86C</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="189469141" LSID="urn:lsid:plazi:treatment:EF98AB20DF14505B922050381FC8848E" httpUri="http://treatment.plazi.org/id/EF98AB20DF14505B922050381FC8848E" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="EF98AB20-DF14-505B-9220-50381FC8848E" authority="Inuthai, Chantanaorr. &amp; Suddee" authorityName="Inuthai, Chantanaorr. &amp; Suddee" authorityYear="2021" class="Liliopsida" family="Orchidaceae" genus="Corybas" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Corybas papillatus" order="Asparagales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="papillatus" status="sp. nov.">Corybas papillatus Inuthai, Chantanaorr. &amp; Suddee</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Corybas papillatus Inuthai, Chantanaorr. &amp; Suddee A, B plants in natural habitat, on humus associated with Bazzania sp. and Acroporium sp. C plant with immature fruit D, E flowers D front view E lateral view F immature flower G underground parts with tubers. Photographs by W. Kiewbang." figureDoi="10.3897/phytokeys.183.71167.figure1" httpUri="https://binary.pensoft.net/fig/595910" pageId="0" pageNumber="1">Figs 1</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Corybas papillatus Inuthai, Chantanaorr. &amp; Suddee A whole plant with flower and tubers B plant with immature flower C mature flower, lateral view D plant with immature fruit E floral bract, F, G dorsal sepal F ventral view G lateral view H lateral sepals and petals I labellum J, K column, J lateral view, K front view. Drawn by J. Inuthai." figureDoi="10.3897/phytokeys.183.71167.figure2" httpUri="https://binary.pensoft.net/fig/595911" pageId="0" pageNumber="1">, 2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
Similar to 
<taxonomicName class="Liliopsida" family="Orchidaceae" genus="Corybas" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Corybas villosus" order="Asparagales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="villosus">
<emphasis italics="true" pageId="0" pageNumber="1">Corybas villosus</emphasis>
</taxonomicName>
J. Dransf. &amp; Gord. Sm., but differs in the absence of dorsal sepal keel and the lateral sepals adnate laterally at the base to the connate petals.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">
<typeStatus>Type</typeStatus>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation collectingDate="2018-07-04" collectorName="M. Poopath, N. Tetsana, W. Kiewbang, C. Hemrat, S. Jirakorn" country="Thailand" elevation="1700" latitude="8.493556" location="Khao Luang Mt." longLatPrecision="1" longitude="99.72747" municipality="Khao Luang National Park" specimenCount="1" stateProvince="Nakhon Si Thammarat province" typeStatus="holotype">
<collectingCountry name="Thailand">Thailand</collectingCountry>
. 
<collectingRegion country="Thailand" name="Nakhon Si Thammarat">Nakhon Si Thammarat province</collectingRegion>
, 
<collectingMunicipality>Khao Luang National Park</collectingMunicipality>
, near summit of 
<location LSID="urn:lsid:plazi:treatment:EF98AB20DF14505B922050381FC8848E:5157FF49B53C11D58B082CAA34CD766E" country="Thailand" latitude="8.493556" longLatPrecision="1" longitude="99.72747" municipality="Khao Luang National Park" name="Khao Luang Mt." stateProvince="Nakhon Si Thammarat province">Khao Luang Mt.</location>
, 
<geoCoordinate degrees="08" direction="north" minutes="29" orientation="latitude" precision="1" seconds="36.8" value="8.493556">08°29'36.8&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="099" direction="east" minutes="43" orientation="longitude" precision="1" seconds="38.9" value="99.72747">099°43'38.9&quot;E</geoCoordinate>
, ca. 
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">1,700 m</elevation>
</quantity>
alt., 
<collectingDate value="2018-07-04">4 July 2018</collectingDate>
, 
<emphasis italics="true" pageId="0" pageNumber="1">
<collectorName>M. Poopath</collectorName>
, 
<collectorName>N. Tetsana</collectorName>
, 
<collectorName>W. Kiewbang</collectorName>
, 
<collectorName>C. Hemrat</collectorName>
&amp; 
<collectorName>S. Jirakorn</collectorName>
2201
</emphasis>
(
<typeStatus>holotype</typeStatus>
BKF!, spirit material)
</materialsCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Small terrestrial herb with underground tubers. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Tubers</emphasis>
globose or ovoid, fleshy, 
<quantity metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3-4 mm</quantity>
diam. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Stem</emphasis>
erect, whitish-green, 
<quantity metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="12.0" metricValueMin="5.0" unit="mm" value="8.5" valueMax="12.0" valueMin="5.0">5-12 mm</quantity>
long, 
<quantity metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">1-1.5 mm</quantity>
diam., with a basal sheath; stolon whitish, hairy, up to 
<quantity metricMagnitude="-2" metricUnit="m" metricValue="2.0" unit="cm" value="2.0">2 cm</quantity>
long, 
<quantity metricMagnitude="-3" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" unit="mm" value="1.1" valueMax="1.2" valueMin="1.0">1-1.2 mm</quantity>
diam. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Foliage leaf</emphasis>
sessile, cordate, long acuminate at apex, glabrous, 
<quantity metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="10.0" metricValueMin="6.0" unit="mm" value="8.0" valueMax="10.0" valueMin="6.0">6-10 mm</quantity>
long, 
<quantity metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" unit="mm" value="7.0" valueMax="8.0" valueMin="6.0">6-8 mm</quantity>
wide at the widest point, flat, only slightly undulate along margin, pale green with paler veins, the veins scarcely conspicuous. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Inflorescence</emphasis>
one-flowered, terminal; bract pale green, lanceolate-triangular, long acuminate, 
<quantity metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">5-6 mm</quantity>
long, recurved. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Flower</emphasis>
dark purple. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Dorsal sepal</emphasis>
purplish, erect below, then strongly curved above, hooded and clasping labellum throughout its length, spathulate, ca. 
<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.5" unit="mm" value="15.0">15 mm</quantity>
long, 
<quantity metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">8-10 mm</quantity>
wide, rounded at apex, apical margins denticulate, abaxial (dorsal) surface bearing irregular papillae in the upper half. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Lateral sepals</emphasis>
greenish-white, linear-triangular to antenna-like, ca. 
<quantity metricMagnitude="-2" metricUnit="m" metricValue="2.5" unit="mm" value="25.0">25 mm</quantity>
long, laterally adnate to the petals in the basal ca. 
<quantity metricMagnitude="-3" metricUnit="m" metricValue="1.0" unit="mm" value="1.0">1 mm</quantity>
. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Petals</emphasis>
greenish-white, connate at the base for ca. 
<quantity metricMagnitude="-3" metricUnit="m" metricValue="1.0" unit="mm" value="1.0">1 mm</quantity>
, free above, similar to lateral sepals but shorter, ca. 
<quantity metricMagnitude="-2" metricUnit="m" metricValue="2.2" unit="mm" value="22.0">22 mm</quantity>
long. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Labellum</emphasis>
differentiated into 2 parts, erect in basal half and strongly reflexed above, ca. 
<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.7" unit="mm" value="17.0">17 mm</quantity>
long in total length; the basal half with margins strongly inrolled throughout, and forming a tube with the dorsal sepal, ca. 
<quantity metricMagnitude="-3" metricUnit="m" metricValue="7.0" unit="mm" value="7.0">7 mm</quantity>
long; the upper half reflexed and expanded, orbicular, ca. 
<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.0" unit="mm" value="10.0">10 mm</quantity>
diam., margin erose, reddish-purple excepted for a white zone just below the margins; mouth of throat very deeply V-shaped; adaxial surface bearing short hairs; spurs 2, divergent, conical, ca. 
<quantity metricMagnitude="-3" metricUnit="m" metricValue="3.0" unit="mm" value="3.0">3 mm</quantity>
long. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Column</emphasis>
very short, ca. 
<quantity metricMagnitude="-3" metricUnit="m" metricValue="1.5" unit="mm" value="1.5">1.5 mm</quantity>
long; stigma rounded, ca. 
<quantity metricMagnitude="-4" metricUnit="m" metricValue="5.0" unit="mm" value="0.5">0.5 mm</quantity>
diam.; anther erect, smooth; pollinia not seen. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Ovary</emphasis>
green, glabrous, ca. 
<quantity metricMagnitude="-3" metricUnit="m" metricValue="2.0" unit="mm" value="2.0">2 mm</quantity>
long. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Capsule</emphasis>
(immature) erect, ellipsoid or fusiform, 
<quantity metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">5-7 mm</quantity>
long, 
<quantity metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" unit="mm" value="2.75" valueMax="3.0" valueMin="2.5">2.5-3 mm</quantity>
diam. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Seed</emphasis>
not seen.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="phenology">
<paragraph pageId="0" pageNumber="1">Phenology.</paragraph>
<paragraph pageId="0" pageNumber="1">Flowering and fruiting observed in July.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="habitat">
<paragraph pageId="0" pageNumber="1">Habitat and ecology.</paragraph>
<paragraph pageId="0" pageNumber="1">
The new species was found growing amongst bryophytes (Fig. 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Corybas papillatus Inuthai, Chantanaorr. &amp; Suddee A, B plants in natural habitat, on humus associated with Bazzania sp. and Acroporium sp. C plant with immature fruit D, E flowers D front view E lateral view F immature flower G underground parts with tubers. Photographs by W. Kiewbang." figureDoi="10.3897/phytokeys.183.71167.figure1" httpUri="https://binary.pensoft.net/fig/595910" pageId="0" pageNumber="1">1A-C</figureCitation>
), such as 
<taxonomicName authorityName="Mitten" authorityYear="1868" class="Bryopsida" family="Sematophyllaceae" genus="Acroporium" kingdom="Plantae" lsidName="Acroporium" order="Hypnales" pageId="0" pageNumber="1" phylum="Bryophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Acroporium</emphasis>
</taxonomicName>
sp., 
<taxonomicName authorityName="S.F.Gray" authorityYear="1821" class="Jungermanniopsida" family="Lepidoziaceae" genus="Bazzania" kingdom="Plantae" lsidName="Bazzania" order="Jungermanniales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Bazzania</emphasis>
</taxonomicName>
sp., and 
<taxonomicName class="Jungermanniopsida" family="Schistochilaceae" genus="Schistochila" kingdom="Plantae" lsidName="Schistochila aligera" order="Jungermanniales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="species" species="aligera">
<emphasis italics="true" pageId="0" pageNumber="1">Schistochila aligera</emphasis>
</taxonomicName>
(Nees &amp; Blume) J.B. Jack &amp; Steph., on humus in shade in montane forest, ca. 1,700 m above sea level.
</paragraph>
<caption doi="10.3897/phytokeys.183.71167.figure1" httpUri="https://binary.pensoft.net/fig/595910" pageId="0" pageNumber="1" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 1.</emphasis>
<taxonomicName authorityName="Inuthai, Chantanaorr. &amp; Suddee" authorityYear="2021" class="Liliopsida" family="Orchidaceae" genus="Corybas" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Corybas papillatus" order="Asparagales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="papillatus">
<emphasis italics="true" pageId="0" pageNumber="1">Corybas papillatus</emphasis>
</taxonomicName>
Inuthai, Chantanaorr. &amp; Suddee 
<emphasis bold="true" pageId="0" pageNumber="1">A, B</emphasis>
plants in natural habitat, on humus associated with 
<taxonomicName authorityName="S.F.Gray" authorityYear="1821" class="Jungermanniopsida" family="Lepidoziaceae" genus="Bazzania" kingdom="Plantae" lsidName="Bazzania" order="Jungermanniales" pageId="0" pageNumber="1" phylum="Marchantiophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Bazzania</emphasis>
</taxonomicName>
sp. and 
<taxonomicName authorityName="Mitten" authorityYear="1868" class="Bryopsida" family="Sematophyllaceae" genus="Acroporium" kingdom="Plantae" lsidName="Acroporium" order="Hypnales" pageId="0" pageNumber="1" phylum="Bryophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Acroporium</emphasis>
</taxonomicName>
sp. 
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
plant with immature fruit 
<emphasis bold="true" pageId="0" pageNumber="1">D, E</emphasis>
flowers 
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
front view 
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
lateral view 
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
immature flower 
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
underground parts with tubers. Photographs by W. Kiewbang.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Inuthai, Chantanaorr. &amp; Suddee" authorityYear="2021" class="Liliopsida" family="Orchidaceae" genus="Corybas" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Corybas papillatus" order="Asparagales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="papillatus">
<emphasis italics="true" pageId="0" pageNumber="1">Corybas papillatus</emphasis>
</taxonomicName>
is only known from the type locality (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Type locality of Corybas papillatus Inuthai, Chantanaorr. &amp; Suddee (•)." figureDoi="10.3897/phytokeys.183.71167.figure3" httpUri="https://binary.pensoft.net/fig/595912" pageId="0" pageNumber="1">3</figureCitation>
), however, it may also occur in other areas in peninsular Thailand with similar vegetation type.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
The specific epithet ' 
<taxonomicName authorityName="Inuthai, Chantanaorr. &amp; Suddee" authorityYear="2021" class="Liliopsida" family="Orchidaceae" genus="Corybas" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Corybas papillatus" order="Asparagales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="papillatus">
<emphasis italics="true" pageId="0" pageNumber="1">Corybas papillatus</emphasis>
</taxonomicName>
' alludes to occurrence of irregular papillae in upper portion of abaxial (dorsal) surface of dorsal sepal.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="conservation status">
<paragraph pageId="0" pageNumber="1">Conservation status.</paragraph>
<paragraph pageId="0" pageNumber="1">
We consider it likely that if a formal assessment were performed, this species would be categorized as Critically Endangered (CR (D)) based on a preliminary risk of extinction assessment using the IUCN red list categories and criteria (
<bibRefCitation author="IUCN" journalOrPublisher="Phytotaxa" pageId="0" pageNumber="1" publicationUrl="http://www.iucnredlist.org/documents/RedListGuidelines.pdf" refId="B6" refString="IUCN, 2019. Guidelines for using the IUCN Red List Categories and Criteria. Version 14. Prepared by Standards and Petitions Committee. http://www.iucnredlist.org/documents/RedListGuidelines.pdf" title="Guidelines for using the IUCN Red List Categories and Criteria. Version 14. Prepared by Standards and Petitions Committee." url="http://www.iucnredlist.org/documents/RedListGuidelines.pdf" year="2019">IUCN 2019</bibRefCitation>
). This species is known from only four individuals from the type locality which attracts high numbers of camping tourists. Although we returned to the same locality and tried to find more specimens in 2020 it could not be found again. The species is, however, easily be overlooked in the field because of its small size.
</paragraph>
<caption doi="10.3897/phytokeys.183.71167.figure2" httpUri="https://binary.pensoft.net/fig/595911" pageId="0" pageNumber="1" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 2.</emphasis>
<taxonomicName authorityName="Inuthai, Chantanaorr. &amp; Suddee" authorityYear="2021" class="Liliopsida" family="Orchidaceae" genus="Corybas" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Corybas papillatus" order="Asparagales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="papillatus">
<emphasis italics="true" pageId="0" pageNumber="1">Corybas papillatus</emphasis>
</taxonomicName>
Inuthai, Chantanaorr. &amp; Suddee 
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
whole plant with flower and tubers 
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
plant with immature flower 
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
mature flower, lateral view 
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
plant with immature fruit 
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
floral bract, 
<emphasis bold="true" pageId="0" pageNumber="1">F, G</emphasis>
dorsal sepal 
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
ventral view 
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
lateral view 
<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
lateral sepals and petals 
<emphasis bold="true" pageId="0" pageNumber="1">I</emphasis>
labellum 
<emphasis bold="true" pageId="0" pageNumber="1">J, K</emphasis>
column, 
<emphasis bold="true" pageId="0" pageNumber="1">J</emphasis>
lateral view, 
<emphasis bold="true" pageId="0" pageNumber="1">K</emphasis>
front view. Drawn by J. Inuthai.
</paragraph>
</caption>
<caption doi="10.3897/phytokeys.183.71167.figure3" httpUri="https://binary.pensoft.net/fig/595912" pageId="0" pageNumber="1" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 3.</emphasis>
Type locality of 
<taxonomicName authorityName="Inuthai, Chantanaorr. &amp; Suddee" authorityYear="2021" class="Liliopsida" family="Orchidaceae" genus="Corybas" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Corybas papillatus" order="Asparagales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="papillatus">
<emphasis italics="true" pageId="0" pageNumber="1">Corybas papillatus</emphasis>
</taxonomicName>
Inuthai, Chantanaorr. &amp; Suddee (•).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>