383 lines
47 KiB
XML
383 lines
47 KiB
XML
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<mods:title id="0ABF04F33F63C0D18E4E9FAB2F54D868">Contributions to the knowledge of Neotropical Lycaenidae: taxonomy of Johnsonita Salazar & Constantino, 1995 with descriptions of seven new species (Theclinae Eumaeini)</mods:title>
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<mods:namePart id="B41015E87EFB0CC0712180E0F6FB16E1">Bálint, Zsolt</mods:namePart>
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<mods:affiliation id="85739E0F4C62E47DBEE0F649F545EB27">Hungarian Natural History Museum, Department of Zoology, H- 1088 Budapest, Baross utca 13, Hungary. balint. zsolt @ nhmus. hu; https: // orcid. org / 0000 - 0001 - 8174 - 878 X</mods:affiliation>
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<mods:affiliation id="E36BB89E94C16D1E75B122492CE68DFE">Universidad Nacional de San Agustín de Arequipa, Museo de Historia Natural, Av. Alcides Carrión s / n, Arequipa, Peru. cerdenajoseal @ yahoo. es; https: // orcid. org / 0000 - 0003 - 3930 - 6480 jjackie 4 u @ gmail. com; https: // orcid. org / 0000 - 0001 - 5913 - 0756</mods:affiliation>
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<mods:affiliation id="932A25BF00B67D05F49A6020CBA6E06A">Universidad Nacional de San Agustín de Arequipa, Museo de Historia Natural, Av. Alcides Carrión s / n, Arequipa, Peru. cerdenajoseal @ yahoo. es; https: // orcid. org / 0000 - 0003 - 3930 - 6480 jjackie 4 u @ gmail. com; https: // orcid. org / 0000 - 0001 - 5913 - 0756</mods:affiliation>
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<mods:affiliation id="A3CF2CF7803E97A2F88A44B4188F1EDC">Department of Invertebrate Evolution, Institute of Zoology and Biomedical Research, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland.</mods:affiliation>
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<treatment id="D5008F50FFE3B307FF76FE55971CFB94" ID-DOI="http://doi.org/10.5281/zenodo.4696414" ID-GBIF-Taxon="181531946" ID-Zenodo-Dep="4696414" LSID="urn:lsid:plazi:treatment:D5008F50FFE3B307FF76FE55971CFB94" httpUri="http://treatment.plazi.org/id/D5008F50FFE3B307FF76FE55971CFB94" lastPageId="11" lastPageNumber="22" pageId="10" pageNumber="21">
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<emphasis id="6FDDE254FFE3B306FF76FE5597BCFDC3" bold="true" box="[151,405,499,526]" italics="true" pageId="10" pageNumber="21">Johnsonita catadupa</emphasis>
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(
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<bibRefCitation id="393843B7FFE3B306FE45FE52944FFDC3" author="Hewitson" box="[420,614,500,526]" firstAuthor="Hewitson" journalOrPublisher="John Van Voorst, London, (4), 115 - 136, 14 a - 14 h, supplementary" pageId="10" pageNumber="21" pagination="1 - 5" refId="ref22793" refString="Hewitson, W. C. (1869) In: Illustrations of diurnal Lepidoptera. Part I. (4). Lycaenidae. John Van Voorst, London, (4), 115 - 136, 14 a - 14 h, supplementary pp. 1 - 16, pls. 47 - 54, 3 a - 3 c, supplementary pls. 1 - 5." type="book chapter" volumeTitle="Illustrations of diurnal Lepidoptera. Part I. (4). Lycaenidae" year="1869">Hewitson, 1869</bibRefCitation>
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<bibRefCitation id="393843B7FFE3B306FD97FE559525FDC3" author="Balint" box="[630,780,499,526]" firstAuthor="Balint" journalOrPublisher="Folia entomologica hungarica" pageId="10" pageNumber="21" pagination="243 - 250" part="64" refId="ref21464" refString="Balint, Zs. (2003) Description of Johnsonita johnbanksi sp. n. from Peru, with notes on the genus (Lepidoptera: Lycaenidae). Folia entomologica hungarica, 64, 243 - 250." title="Description of Johnsonita johnbanksi sp. n. from Peru, with notes on the genus (Lepidoptera: Lycaenidae)" type="journal article" year="2003">Bálint, 2003</bibRefCitation>
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(
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Figs
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,
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<figureCitation id="C59222C3FFE3B306FF15FDBE9716FDFF" box="[244,319,536,562]" captionStart="FIGURE 17" captionStartId="27.[151,250,1943,1967]" captionTargetBox="[272,1345,182,1893]" captionTargetId="figure-18@27.[175,1413,181,1920]" captionTargetPageId="27" captionText="FIGURE 17. Johnsonita genitalia armatures, entire male organs in lateral (left column) and ventral (right column) views.A–B: J. auda, Ecuador, Morona-Santiago, 9 de Octubre-Macas; C–D: J. carpia, Ecuador, Napo, Res. Yanayacu; E–F: J. catadupa, Ecuador, Loja, Yangana; G–H: J. chlamydem, Peru, Pasco, Oxapampa, La Antena (CEP–MZUJ specimens) (scale as indicated)." figureDoi="http://doi.org/10.5281/zenodo.4711291" httpUri="https://zenodo.org/record/4711291/files/figure.png" pageId="10" pageNumber="21">17E–F</figureCitation>
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,
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<figureCitation id="C59222C3FFE3B306FEABFDBE97BDFDFF" box="[330,404,536,562]" captionStart="FIGURE 20" captionStartId="30.[151,250,1909,1933]" captionTargetBox="[183,1405,181,1885]" captionTargetId="figure-23@30.[183,1405,181,1885]" captionTargetPageId="30" captionText="FIGURE 20. Entire Johnsonita female organs in lateral (left column), ductus in lateral (central column) and ventral (right column) views. A–C: J. auda, Venezuela, Edo. Barinas, Minas de San Isidro; D–F: J. carpia, Ecuador, Prov. Carchi, Res. Forest Golondrinas; G–I: J. catadupa, Peru, Dept. Amazonas, Molinopampa, Tingo Ocol (CEP–MZUJ specimens) (scale as indicated)." figureDoi="http://doi.org/10.5281/zenodo.4711297" httpUri="https://zenodo.org/record/4711297/files/figure.png" pageId="10" pageNumber="21">20G–I</figureCitation>
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,
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<figureCitation id="C59222C3FFE3B306FE41FDBE97F8FDFF" box="[416,465,536,562]" captionStart="FIGURE 23" captionStartId="33.[151,250,1570,1594]" captionTargetBox="[390,1203,198,1539]" captionTargetId="figure-18@33.[378,1210,181,1545]" captionTargetPageId="33" captionText="FIGURE 23. Johnsonita mail trait ventral forewing subcubital scent patch indicated by arrow. A: J. carpia, B: J. catadupa, C: J. chlamydem, D: J. ianusca, E: J. johnsoni, F: J. pardoa, G: J. turquisca, H: J. zubkova." figureDoi="http://doi.org/10.5281/zenodo.4711305" httpUri="https://zenodo.org/record/4711305/files/figure.png" pageId="10" pageNumber="21">23B</figureCitation>
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,
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<figureCitation id="C59222C3FFE3B306FE3DFDBE97D3FDFF" box="[476,506,536,562]" captionStart="FIGURE 27" captionStartId="36.[151,250,1590,1614]" captionTargetBox="[163,1421,184,1559]" captionTargetId="figure-23@36.[151,1436,181,1566]" captionTargetPageId="36" captionText="FIGURE 27. Thecla catadupa Hewitson, 1869. NHMUK lectotype specimen and labels (scale bar: 10 mm)." figureDoi="http://doi.org/10.5281/zenodo.4711317" httpUri="https://zenodo.org/record/4711317/files/figure.png" pageId="10" pageNumber="21">27</figureCitation>
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,
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<figureCitation id="C59222C3FFE3B306FDE7FDBE940AFDFF" box="[518,547,536,562]" captionStart="FIGURE 30" captionStartId="39.[151,250,1742,1766]" captionTargetBox="[251,1396,204,1718]" captionTargetId="figure-18@39.[192,1396,181,1718]" captionTargetPageId="39" captionText="FIGURE 30. Johnsonita distributions based on museum material examinded. J. assula = red circle, J. carpia = yellow trinagle, J. catadupa = blue diamond, J. chalamydem = brown triangle, J. ianusca = green square, J. oxalida = purple diamond." figureDoi="http://doi.org/10.5281/zenodo.4711325" httpUri="https://zenodo.org/record/4711325/files/figure.png" pageId="10" pageNumber="21">30</figureCitation>
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)
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<emphasis id="6FDDE254FFE3B306FF76FDFA9769FDB9" box="[151,320,604,628]" italics="true" pageId="10" pageNumber="21">Thecla catadupa</emphasis>
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–
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|
||
<bibRefCitation id="393843B7FFE3B306FDD5FDFA949EFDB8" author="Draudt" box="[564,695,604,629]" editor="Seitz, A." firstAuthor="Draudt" journalOrPublisher="Stuttgart, Alfred Kernen" pageId="10" pageNumber="21" pagination="144 - 159" refId="ref22147" refString="Draudt, M. W. K. (1919 - 21). Familie: Lycaenidae. In: Seitz, A. (Ed.), Die Gross- Schmetterlinge der Erde. Vol. 5. Stuttgart, Alfred Kernen, pp. 744 - 752 (24 November 1919) + 753 - 768 (6 December 1919) + 769 - 776 (12 January 1920) + 777 - 808 (3 February 1920) + 809 - 816 (10 December 1920) + 817 - 824 (11 January 1921) + 825 - 831 (24 January 1921), pls. 144 - 159." title="Familie: Lycaenidae" type="book chapter" volumeTitle="Die Gross- Schmetterlinge der Erde. Vol. 5" year="1919">Draudt 1919</bibRefCitation>
|
||
-1921: 757;
|
||
<treatmentCitation id="DC081857FFE3B306FCCEFDFA922FFDB8" author="d'Abrera" box="[815,1030,604,629]" page="1138" pageId="10" pageNumber="21" year="1995">
|
||
<bibRefCitation id="393843B7FFE3B306FCCEFDFA922FFDB8" author="d'Abrera" box="[815,1030,604,629]" firstAuthor="d'Abrera" journalOrPublisher="Hill House, Black Rock, Victoria" page="1138" pageId="10" pageNumber="21" pagination="1098 - 1270" refId="ref22067" refString="D'Abrera, B. (1995) In: Butterflies of the Neotropical Region. Part VII. Lycaenidae. Hill House, Black Rock, Victoria, pp. i - xi + 1098 - 1270." type="book chapter" volumeTitle="Butterflies of the Neotropical Region. Part VII. Lycaenidae" year="1995">d’Abrera 1995: 1138</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="5D163E46FFE3B306FF76FDDD9584FD5E" blockId="10.[151,1103,604,690]" box="[151,941,635,659]" pageId="10" pageNumber="21">
|
||
<treatmentCitationGroup id="7DB91968FFE3B306FF76FDDD9584FD5E" box="[151,941,635,659]" pageId="10" pageNumber="21">
|
||
<taxonomicName id="9AA945C5FFE3B306FF76FDDD97F7FD5E" authority="(Hewitson)" baseAuthorityName="Hewitson" baseAuthorityYear="1869" box="[151,478,635,659]" class="Insecta" family="Lycaenidae" genus="Radissima" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="21" phylum="Arthropoda" rank="species" species="catadupa">
|
||
<emphasis id="6FDDE254FFE3B306FF76FDDD974CFD5E" box="[151,357,635,659]" italics="true" pageId="10" pageNumber="21">Radissima catadupa</emphasis>
|
||
(Hewitson)
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="DC081857FFE3B306FE16FDDD949FFD5E" author="Johnson" box="[503,694,635,659]" page="174" pageId="10" pageNumber="21" year="1992">
|
||
<bibRefCitation id="393843B7FFE3B306FE16FDDD949FFD5E" author="Johnson" bookContentInfo="205 figs." box="[503,694,635,659]" firstAuthor="Johnson" journalOrPublisher="Reports of the Museum of Natural History, University of Wisconsin, Stevens Point" page="174" pageId="10" pageNumber="21" pagination="1 - 135" part="2" refId="ref22864" refString="Johnson, K. (1992) Genera and species of the Neotropical " elfin " - like hairstreak butterflies (Lepidoptera, Lycaenidae, Theclinae). Reports of the Museum of Natural History, University of Wisconsin, Stevens Point, 22 (1 & 2), 1 - 135, 205 figs. & 136 - 279 + 241, 2 figs." title="Genera and species of the Neotropical " elfin " - like hairstreak butterflies (Lepidoptera, Lycaenidae, Theclinae)" type="book chapter" year="1992">Johnson 1992: 174</bibRefCitation>
|
||
</treatmentCitation>
|
||
(
|
||
<typeStatus id="821280E4FFE3B306FD25FDDD950AFD5E" box="[708,803,635,659]" pageId="10" pageNumber="21" type="lectotype">lectotype</typeStatus>
|
||
designation).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="5D163E46FFE3B307FF76FD3C9715FED3" blockId="10.[151,1103,604,690]" lastBlockId="11.[151,1437,150,1113]" lastPageId="11" lastPageNumber="22" pageId="10" pageNumber="21">
|
||
<taxonomicName id="9AA945C5FFE3B306FF76FD3C97CAFD7F" authority="(Hewitson)" authorityName="Balint" authorityYear="2003" baseAuthorityName="Hewitson" baseAuthorityYear="1869" box="[151,483,666,690]" class="Insecta" family="Lycaenidae" genus="Johnsonita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="21" phylum="Arthropoda" rank="species" species="catadupa">
|
||
<emphasis id="6FDDE254FFE3B306FF76FD3C9742FD7F" box="[151,363,666,690]" italics="true" pageId="10" pageNumber="21">Johnsonita catadupa</emphasis>
|
||
(Hewitson)
|
||
</taxonomicName>
|
||
–
|
||
<bibRefCitation id="393843B7FFE3B306FE1CFD3C948EFD7F" author="Balint" box="[509,679,666,690]" firstAuthor="Balint" journalOrPublisher="Folia entomologica hungarica" page="249" pageId="10" pageNumber="21" pagination="243 - 250" part="64" refId="ref21464" refString="Balint, Zs. (2003) Description of Johnsonita johnbanksi sp. n. from Peru, with notes on the genus (Lepidoptera: Lycaenidae). Folia entomologica hungarica, 64, 243 - 250." title="Description of Johnsonita johnbanksi sp. n. from Peru, with notes on the genus (Lepidoptera: Lycaenidae)" type="journal article" year="2003">Bálint 2003: 249</bibRefCitation>
|
||
(new combination);
|
||
<bibRefCitation id="393843B7FFE3B306FC9AFD3C9262FD7F" author="Robbins" box="[891,1099,666,690]" editor="Lamas, G." firstAuthor="Robbins" journalOrPublisher="Scientific Publishers, Gainesville, Florida" page="120" pageId="10" pageNumber="21" pagination="118 - 137" refId="ref23342" refString="Robbins, R. K. (2004 b) Lycaenidae. Theclinae. Tribe Eumaeini,. In: Lamas, G. (Ed.), Checklist: Part 4 A. Hesperioidea - Papilionoidea. In: Heppner, J. B. (Ed.), Atlas of Neotropical Lepidoptera. Vol. 5 A. Association for Tropical Lepidoptera, Scientific Publishers, Gainesville, Florida, pp. 118 - 137." title="Lycaenidae. Theclinae. Tribe Eumaeini" type="book chapter" volumeTitle="Checklist: Part 4 A. Hesperioidea - Papilionoidea. In: Heppner, J. B" year="2004" yearSuffix="b">Robbins 2004b: 120</bibRefCitation>
|
||
.
|
||
<emphasis id="6FDDE254FFE3B306FF76F8E8971DF8AA" box="[151,308,1869,1895]" italics="true" pageId="10" pageNumber="21">Type material</emphasis>
|
||
(
|
||
<figureCitation id="C59222C3FFE3B306FEA7F8EB97B5F8AA" box="[326,412,1869,1895]" captionStart="FIGURE 27" captionStartId="36.[151,250,1590,1614]" captionTargetBox="[163,1421,184,1559]" captionTargetId="figure-23@36.[151,1436,181,1566]" captionTargetPageId="36" captionText="FIGURE 27. Thecla catadupa Hewitson, 1869. NHMUK lectotype specimen and labels (scale bar: 10 mm)." figureDoi="http://doi.org/10.5281/zenodo.4711317" httpUri="https://zenodo.org/record/4711317/files/figure.png" pageId="10" pageNumber="21">Fig. 27</figureCitation>
|
||
). The species was described on the basis of an unstated number of male specimens from
|
||
<collectingCountry id="25BE7ED6FFE3B306FF76F8D796DEF846" box="[151,247,1905,1931]" name="Ecuador" pageId="10" pageNumber="21">Ecuador</collectingCountry>
|
||
.
|
||
<bibRefCitation id="393843B7FFE3B306FEE3F8D79798F846" author="Johnson" bookContentInfo="205 figs." box="[258,433,1905,1931]" firstAuthor="Johnson" journalOrPublisher="Reports of the Museum of Natural History, University of Wisconsin, Stevens Point" pageId="10" pageNumber="21" pagination="1 - 135" part="2" refId="ref22864" refString="Johnson, K. (1992) Genera and species of the Neotropical " elfin " - like hairstreak butterflies (Lepidoptera, Lycaenidae, Theclinae). Reports of the Museum of Natural History, University of Wisconsin, Stevens Point, 22 (1 & 2), 1 - 135, 205 figs. & 136 - 279 + 241, 2 figs." title="Genera and species of the Neotropical " elfin " - like hairstreak butterflies (Lepidoptera, Lycaenidae, Theclinae)" type="book chapter" year="1992">Johnson (1992)</bibRefCitation>
|
||
designated the
|
||
<typeStatus id="821280E4FFE3B306FD82F8D794E3F846" box="[611,714,1905,1931]" pageId="10" pageNumber="21" type="lectotype">lectotype</typeStatus>
|
||
from the Hewitson collection. This specimen has the following labels: (1) “
|
||
<emphasis id="6FDDE254FFE3B306FEFEF8339756F862" box="[287,383,1941,1967]" italics="true" pageId="10" pageNumber="21">
|
||
<collectingCountry id="25BE7ED6FFE3B306FEFEF8339756F862" box="[287,383,1941,1967]" name="Ecuador" pageId="10" pageNumber="21">Ecuador</collectingCountry>
|
||
</emphasis>
|
||
[//] Hewitson Coll. [//] 79—69. [//]
|
||
<taxonomicName id="9AA945C5FFE3B306FCC3F8339547F862" ID-CoL="9CM8J" authorityName="Fabricius" authorityYear="1807" box="[802,878,1941,1967]" class="Insecta" family="Lycaenidae" genus="Thecla" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="21" phylum="Arthropoda" rank="genus">Thecla</taxonomicName>
|
||
[//]
|
||
<emphasis id="6FDDE254FFE3B306FC42F833927AF862" box="[931,1107,1941,1967]" italics="true" pageId="10" pageNumber="21">Catadupa Hew.</emphasis>
|
||
[//]
|
||
<emphasis id="6FDDE254FFE3B306FB66F8339286F862" box="[1159,1199,1941,1967]" italics="true" pageId="10" pageNumber="21">(3.)</emphasis>
|
||
” (white label,
|
||
<emphasis id="6FDDE254FFE3B306FABAF83396CFF81E" italics="true" pageId="10" pageNumber="21">handwritten</emphasis>
|
||
and printed in black), (2) “Type” (white confetti–like label with red frame, printed); (3) “B.M. TYPE [//] No. Rh.
|
||
<emphasis id="6FDDE254FFE3B306FF18F87B970AF83A" box="[249,291,2013,2039]" italics="true" pageId="10" pageNumber="21">595</emphasis>
|
||
” (white label, printed and
|
||
<emphasis id="6FDDE254FFE3B306FDB9F87B94F6F83A" box="[600,735,2013,2039]" italics="true" pageId="10" pageNumber="21">handwritten</emphasis>
|
||
in black); (4) “B.M.(N.H) [//] Rhopalocera [//] Slide No. [//]
|
||
<emphasis id="6FDDE254FFE2B307FF76FF3196C2FF7C" box="[151,235,151,177]" italics="true" pageId="11" pageNumber="22">
|
||
<date id="29171886FFE2B307FF76FF3196C2FF7C" box="[151,235,151,177]" pageId="11" pageNumber="22" value="1540-05">V. 1540</date>
|
||
</emphasis>
|
||
” (white label, printed in red and
|
||
<emphasis id="6FDDE254FFE2B307FD81FF3194CEFF7C" box="[608,743,151,177]" italics="true" pageId="11" pageNumber="22">handwritten</emphasis>
|
||
in black); (5) “
|
||
<emphasis id="6FDDE254FFE2B307FC74FF319209FF7C" box="[917,1056,151,177]" italics="true" pageId="11" pageNumber="22">Genitalia by</emphasis>
|
||
[//]
|
||
<emphasis id="6FDDE254FFE2B307FBB0FF3192D5FF7C" box="[1105,1276,151,177]" italics="true" pageId="11" pageNumber="22">K. Johnson ‘83</emphasis>
|
||
” (white label,
|
||
<emphasis id="6FDDE254FFE2B307FF76FF1D9737FF18" box="[151,286,187,213]" italics="true" pageId="11" pageNumber="22">handwritten</emphasis>
|
||
); (6) “
|
||
<emphasis id="6FDDE254FFE2B307FE8AFF1D942AFF18" box="[363,515,187,213]" italics="true" pageId="11" pageNumber="22">
|
||
<typeStatus id="821280E4FFE2B307FE8AFF1D942AFF18" box="[363,515,187,213]" pageId="11" pageNumber="22" type="lectotype">LECTOTYPE</typeStatus>
|
||
</emphasis>
|
||
[//]
|
||
<typeStatus id="821280E4FFE2B307FDD2FF1D9498FF18" box="[563,689,187,213]" pageId="11" pageNumber="22" type="syntype">SYNTYPE</typeStatus>
|
||
<emphasis id="6FDDE254FFE2B307FD59FF1D94C5FF18" box="[696,748,187,213]" italics="true" pageId="11" pageNumber="22">(m#)</emphasis>
|
||
[//]
|
||
<taxonomicName id="9AA945C5FFE2B307FCFCFF1D9540FF18" ID-CoL="9CM8J" authorityName="Fabricius" authorityYear="1807" box="[797,873,187,213]" class="Insecta" family="Lycaenidae" genus="Thecla" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="22" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6FDDE254FFE2B307FCFCFF1D9540FF18" box="[797,873,187,213]" italics="true" pageId="11" pageNumber="22">Thecla</emphasis>
|
||
</taxonomicName>
|
||
[//]
|
||
<taxonomicName id="9AA945C5FFE2B307FC78FF1D9228FF18" authorityName="Balint" authorityYear="2003" baseAuthorityName="Hewitson" baseAuthorityYear="1869" box="[921,1025,187,213]" class="Insecta" family="Lycaenidae" genus="Johnsonita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="22" phylum="Arthropoda" rank="species" species="catadupa">
|
||
<emphasis id="6FDDE254FFE2B307FC78FF1D9228FF18" box="[921,1025,187,213]" italics="true" pageId="11" pageNumber="22">catadupa</emphasis>
|
||
</taxonomicName>
|
||
[//]
|
||
<emphasis id="6FDDE254FFE2B307FBD0FF1D92B2FF19" box="[1073,1179,187,212]" italics="true" pageId="11" pageNumber="22">Hewitson</emphasis>
|
||
[//]
|
||
<emphasis id="6FDDE254FFE2B307FB2AFF1D935AFF19" box="[1227,1395,186,213]" italics="true" pageId="11" pageNumber="22">London, VI. 26</emphasis>
|
||
[//] Zs. Bálint, 2000” (red label with black frame,
|
||
<emphasis id="6FDDE254FFE2B307FD43FF799500FF34" box="[674,809,223,249]" italics="true" pageId="11" pageNumber="22">handwritten</emphasis>
|
||
and printed in black); (7) “BMNH(E) # 266559”(white label, printed).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="09D66ECEFFE3B306FF76F94595D9F8D2" ID-DOI="http://doi.org/10.5281/zenodo.4711259" ID-Zenodo-Dep="4711259" httpUri="https://zenodo.org/record/4711259/files/figure.png" pageId="10" pageNumber="21" startId="10.[151,250,1763,1787]" targetBox="[155,1427,721,1734]" targetPageId="10">
|
||
<paragraph id="5D163E46FFE3B306FF76F94595D9F8D2" blockId="10.[151,1437,1763,1823]" pageId="10" pageNumber="21">
|
||
<emphasis id="6FDDE254FFE3B306FF76F945973DF936" bold="true" box="[151,276,1763,1787]" pageId="10" pageNumber="21">FIGURE 5.</emphasis>
|
||
<taxonomicName id="9AA945C5FFE3B306FEFAF9459480F936" authority="(Hewitson, 1869)" authorityName="Balint" authorityYear="2003" baseAuthorityName="Hewitson" baseAuthorityYear="1869" box="[283,681,1763,1787]" class="Insecta" family="Lycaenidae" genus="Johnsonita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="21" phylum="Arthropoda" rank="species" species="catadupa">
|
||
<emphasis id="6FDDE254FFE3B306FEFAF94597C6F936" box="[283,495,1763,1787]" italics="true" pageId="10" pageNumber="21">Johnsonita catadupa</emphasis>
|
||
(
|
||
<bibRefCitation id="393843B7FFE3B306FE1FF945948BF936" author="Hewitson" box="[510,674,1763,1787]" firstAuthor="Hewitson" journalOrPublisher="John Van Voorst, London, (4), 115 - 136, 14 a - 14 h, supplementary" pageId="10" pageNumber="21" pagination="1 - 5" refId="ref22793" refString="Hewitson, W. C. (1869) In: Illustrations of diurnal Lepidoptera. Part I. (4). Lycaenidae. John Van Voorst, London, (4), 115 - 136, 14 a - 14 h, supplementary pp. 1 - 16, pls. 47 - 54, 3 a - 3 c, supplementary pls. 1 - 5." type="book chapter" volumeTitle="Illustrations of diurnal Lepidoptera. Part I. (4). Lycaenidae" year="1869">Hewitson, 1869</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
imagines (left column: recto, right column: verso). A– B: male (Ecuador: Loja), C– D: female (Peru: Amazonas) (CEP–MZUJ specimens) (scale bar: 10 mm).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="15B36DCDFFE2B307FF26FE81971CFB94" pageId="11" pageNumber="22" type="materials_examined">
|
||
<paragraph id="5D163E46FFE2B307FF26FE8194D4FE60" blockId="11.[151,1437,150,1113]" pageId="11" pageNumber="22">
|
||
<emphasis id="6FDDE254FFE2B307FF26FE8197B0FE8C" box="[199,409,295,321]" italics="true" pageId="11" pageNumber="22">Material examined</emphasis>
|
||
(n=4, two dissections).
|
||
<materialsCitation id="EDC1341BFFE2B307FD41FE8197DFFE44" ID-GBIF-Occurrence="3083794315" collectingDate="1996-02-25" collectingDateMax="1997-07" collectingDateMin="1996-02-25" collectionCode="CEP, MZUJ, CB" collectorName=" J. Lorenc" country="Ecuador" elevation="2150" location="Zambi" municipality="South region" pageId="11" pageNumber="22" specimenCount="1" stateProvince="Loja">
|
||
<collectingCountry id="25BE7ED6FFE2B307FD41FE819505FE8C" box="[672,812,295,321]" name="Ecuador" pageId="11" pageNumber="22">ECUADOR</collectingCountry>
|
||
:
|
||
<collectingMunicipality id="BD72A43CFFE2B307FCD5FE8195ECFE8C" box="[820,965,295,321]" pageId="11" pageNumber="22">South region</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="9F6DF0A4FFE2B307FC2FFE81922DFE8C" box="[974,1028,295,321]" country="Ecuador" name="Loja" pageId="11" pageNumber="22">Loja</collectingRegion>
|
||
, before 1996, coll.
|
||
<collectorName id="F05C5B90FFE2B307FB32FE819364FE8C" box="[1235,1357,295,321]" pageId="11" pageNumber="22">A. Jasiński</collectorName>
|
||
(
|
||
<collectionCode id="3BB8A683FFE2B307FAB8FE8193A4FE8C" box="[1369,1421,295,321]" pageId="11" pageNumber="22">CEP</collectionCode>
|
||
–
|
||
<collectionCode id="3BB8A683FFE2B307FF76FEED96CCFEA8" box="[151,229,331,357]" pageId="11" pageNumber="22">MZUJ</collectionCode>
|
||
: male);
|
||
<collectingRegion id="9F6DF0A4FFE2B307FEDAFEED9758FEA8" box="[315,369,331,357]" country="Ecuador" name="Loja" pageId="11" pageNumber="22">Loja</collectingRegion>
|
||
,
|
||
<location id="5876689DFFE2B307FE9BFEED97EFFEA8" LSID="urn:lsid:plazi:treatment:D5008F50FFE3B307FF76FE55971CFB94:5876689DFFE2B307FE9BFEED97EFFEA8" box="[378,454,331,357]" country="Ecuador" municipality="South region" name="Zambi" pageId="11" pageNumber="22" stateProvince="Loja">Zambi</location>
|
||
,
|
||
<quantity id="9A5193A3FFE2B307FE31FEED9445FEA8" box="[464,620,331,357]" metricMagnitude="3" metricUnit="m" metricValue="2.15" metricValueMax="2.3" metricValueMin="2.0" pageId="11" pageNumber="22" unit="m" value="2150.0" valueMax="2300.0" valueMin="2000.0">
|
||
<elevation id="D684D975FFE2B307FE31FEED9445FEA8" box="[464,620,331,357]" metricMagnitude="3" metricUnit="m" metricValue="2.15" metricValueMax="2.3" metricValueMin="2.0" pageId="11" pageNumber="22" unit="m" value="2150.0" valueMax="2300.0" valueMin="2000.0">2000–2300 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="29171886FFE2B307FD97FEEC94C4FEA8" box="[630,749,330,357]" pageId="11" pageNumber="22" value="1996-02-25">
|
||
<collectingDate id="3953E16EFFE2B307FD97FEEC94C4FEA8" box="[630,749,330,357]" pageId="11" pageNumber="22" value="1996-02-25">25.II.1996</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="F05C5B90FFE2B307FD16FEED957DFEA8" box="[759,852,331,357]" pageId="11" pageNumber="22">P. Boyer</collectorName>
|
||
(
|
||
<collectionCode id="3BB8A683FFE2B307FC82FEED95A4FEA8" box="[867,909,331,357]" country="Australia" name="The CB Rhizobium Collection" pageId="11" pageNumber="22">CB</collectionCode>
|
||
: male);
|
||
<collectingRegion id="9F6DF0A4FFE2B307FC02FEED9230FEA8" box="[995,1049,331,357]" country="Ecuador" name="Loja" pageId="11" pageNumber="22">Loja</collectingRegion>
|
||
,
|
||
<collectorName id="F05C5B90FFE2B307FBC3FEED92AFFEA8" box="[1058,1158,331,357]" pageId="11" pageNumber="22">Yangana</collectorName>
|
||
,
|
||
<date id="29171886FFE2B307FB6FFEED92D1FEA8" box="[1166,1272,331,357]" pageId="11" pageNumber="22" value="1997-07">
|
||
<collectingDate id="3953E16EFFE2B307FB6FFEED92D1FEA8" box="[1166,1272,331,357]" pageId="11" pageNumber="22" value="1997-07">VII.1997</collectingDate>
|
||
</date>
|
||
. (
|
||
<collectionCode id="3BB8A683FFE2B307FAE9FEED9314FEA8" box="[1288,1341,331,357]" pageId="11" pageNumber="22">CEP</collectionCode>
|
||
–
|
||
<collectionCode id="3BB8A683FFE2B307FAADFEED93B0FEA8" box="[1356,1433,331,357]" pageId="11" pageNumber="22">MZUJ</collectionCode>
|
||
: male, gen. prep. 383/
|
||
<collectorName id="F05C5B90FFE2B307FE65FEC997C4FE44" box="[388,493,367,393]" pageId="11" pageNumber="22">J. Lorenc</collectorName>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="EDC1341BFFE2B307FE1EFEC994D0FE60" ID-GBIF-Occurrence="3083794329" collectingDate="1998-07-05" collectionCode="T, CEP, MZUJ" collectorName="Pyrcz & J. Lorenc" country="Peru" elevation="2450" location="Tingo Ocol" municipality="Molinopampa" pageId="11" pageNumber="22" specimenCount="1">
|
||
<collectingCountry id="25BE7ED6FFE2B307FE1EFEC99462FE44" box="[511,587,367,393]" name="Peru" pageId="11" pageNumber="22">PERU</collectingCountry>
|
||
: Dept. Amazonas,
|
||
<collectingMunicipality id="BD72A43CFFE2B307FCFAFEC99592FE44" box="[795,955,367,393]" pageId="11" pageNumber="22">Molinopampa</collectingMunicipality>
|
||
,
|
||
<location id="5876689DFFE2B307FC25FEC9926FFE44" LSID="urn:lsid:plazi:treatment:D5008F50FFE3B307FF76FE55971CFB94:5876689DFFE2B307FC25FEC9926FFE44" box="[964,1094,367,393]" country="Peru" municipality="Molinopampa" name="Tingo Ocol" pageId="11" pageNumber="22">Tingo Ocol</location>
|
||
,
|
||
<quantity id="9A5193A3FFE2B307FBB1FEC892C4FE44" box="[1104,1261,366,393]" metricMagnitude="3" metricUnit="m" metricValue="2.45" metricValueMax="2.5" metricValueMin="2.4" pageId="11" pageNumber="22" unit="m" value="2450.0" valueMax="2500.0" valueMin="2400.0">
|
||
<elevation id="D684D975FFE2B307FBB1FEC892C4FE44" box="[1104,1261,366,393]" metricMagnitude="3" metricUnit="m" metricValue="2.45" metricValueMax="2.5" metricValueMin="2.4" pageId="11" pageNumber="22" unit="m" value="2450.0" valueMax="2500.0" valueMin="2400.0">2400–2500 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="29171886FFE2B307FB19FEC89351FE44" box="[1272,1400,366,393]" pageId="11" pageNumber="22" value="1998-07-05">
|
||
<collectingDate id="3953E16EFFE2B307FB19FEC89351FE44" box="[1272,1400,366,393]" pageId="11" pageNumber="22" value="1998-07-05">5.VII.1998</collectingDate>
|
||
</date>
|
||
.,
|
||
<collectionCode id="3BB8A683FFE2B307FA67FEC993B1FE44" box="[1414,1432,367,393]" name="Tavera, Department of Geology and Geophysics" pageId="11" pageNumber="22">T</collectionCode>
|
||
.
|
||
<collectorName id="F05C5B90FFE2B307FF76FE3596FEFE60" box="[151,215,403,429]" pageId="11" pageNumber="22">Pyrcz</collectorName>
|
||
(
|
||
<collectionCode id="3BB8A683FFE2B307FF07FE359732FE60" box="[230,283,403,429]" pageId="11" pageNumber="22">CEP</collectionCode>
|
||
–
|
||
<collectionCode id="3BB8A683FFE2B307FECBFE35975EFE60" box="[298,375,403,429]" pageId="11" pageNumber="22">MZUJ</collectionCode>
|
||
: female; gen. prep. 384/
|
||
<collectorName id="F05C5B90FFE2B307FD66FE3594D9FE60" box="[647,752,403,429]" pageId="11" pageNumber="22">J. Lorenc</collectorName>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="5D163E46FFE2B307FF26FE1193A5FD00" blockId="11.[151,1437,150,1113]" pageId="11" pageNumber="22">
|
||
<emphasis id="6FDDE254FFE2B307FF26FE11977AFE1D" box="[199,339,439,464]" italics="true" pageId="11" pageNumber="22">Description.</emphasis>
|
||
Wings. FW costa length:
|
||
<quantity id="9A5193A3FFE2B307FD61FE1094D2FE1C" box="[640,763,438,465]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="11" pageNumber="22" unit="mm" value="13.5" valueMax="15.0" valueMin="12.0">12–15 mm</quantity>
|
||
(n=3,
|
||
<collectionCode id="3BB8A683FFE2B307FCA8FE119555FE1C" box="[841,892,439,465]" pageId="11" pageNumber="22">CEP</collectionCode>
|
||
–
|
||
<collectionCode id="3BB8A683FFE2B307FC6BFE1195FAFE1C" box="[906,979,439,465]" pageId="11" pageNumber="22">MZUJ</collectionCode>
|
||
specimens), costa apical area and outer margin not straight resulting somewhat rounded apex; male dorsal wingsurface blue colouration bright azure; dorsal FW surface black border inner margin reaching vein M3 erection; HW dorsal surface discal area structurally coloured; ventral FW submedial and medial pattern running parallel, HW medial pattern composed of light ruptive dashes supplemented distally with dark spots, submarginal line indistinct and inner margin medial area with brown scaling. Genitalia. Male valva with rounded costal margin flip and pointed valval terminus (slightly longer than total valval length). Female ductus with parallel edges, without lower plate, upper plate symmetric with slightly sclerotized frame pointed in both sides. Androconia present in FW along Cubital 2 vein as narrow patch in medial area.
|
||
</paragraph>
|
||
<paragraph id="5D163E46FFE2B307FF26FD719457FCF4" blockId="11.[151,1437,150,1113]" pageId="11" pageNumber="22">
|
||
<emphasis id="6FDDE254FFE2B307FF26FD719769FD3D" box="[199,320,727,752]" italics="true" pageId="11" pageNumber="22">Diagnosis.</emphasis>
|
||
This species can be easily distinguished from all the congeners by the dorsal wing colouration, because it is the most vivid blue in the genus (all the other species are darker or duller), and also by the complete lack of gleaming scales on ventral wing surface.
|
||
</paragraph>
|
||
<paragraph id="5D163E46FFE2B307FF26FCE59476FC4C" blockId="11.[151,1437,150,1113]" pageId="11" pageNumber="22">
|
||
<emphasis id="6FDDE254FFE2B307FF26FCE59771FC90" box="[199,344,835,861]" italics="true" pageId="11" pageNumber="22">Distribution.</emphasis>
|
||
<collectingCountry id="25BE7ED6FFE2B307FE81FCE597CEFC90" box="[352,487,835,861]" name="Ecuador" pageId="11" pageNumber="22">ECUADOR</collectingCountry>
|
||
(
|
||
<collectingRegion id="9F6DF0A4FFE2B307FE16FCE59419FC93" box="[503,560,835,862]" country="Ecuador" name="Loja" pageId="11" pageNumber="22">Loja</collectingRegion>
|
||
),
|
||
<collectingCountry id="25BE7ED6FFE2B307FDA4FCE594A4FC90" box="[581,653,835,861]" name="Peru" pageId="11" pageNumber="22">PERU</collectingCountry>
|
||
(Amazonas) (
|
||
<figureCitation id="C59222C3FFE2B307FCCFFCE595ADFC90" box="[814,900,835,861]" captionStart="FIGURE 30" captionStartId="39.[151,250,1742,1766]" captionTargetBox="[251,1396,204,1718]" captionTargetId="figure-18@39.[192,1396,181,1718]" captionTargetPageId="39" captionText="FIGURE 30. Johnsonita distributions based on museum material examinded. J. assula = red circle, J. carpia = yellow trinagle, J. catadupa = blue diamond, J. chalamydem = brown triangle, J. ianusca = green square, J. oxalida = purple diamond." figureDoi="http://doi.org/10.5281/zenodo.4711325" httpUri="https://zenodo.org/record/4711325/files/figure.png" pageId="11" pageNumber="22">Fig. 30</figureCitation>
|
||
); spatial: recorded between
|
||
<quantity id="9A5193A3FFE2B307FB22FCE49376FC90" box="[1219,1375,834,861]" metricMagnitude="3" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="11" pageNumber="22" unit="m" value="2250.0" valueMax="2500.0" valueMin="2000.0">2000–2500 m</quantity>
|
||
a.s.l; temporal: recorded in February and July.
|
||
</paragraph>
|
||
<paragraph id="5D163E46FFE2B307FF26FC2D971CFB94" blockId="11.[151,1437,150,1113]" pageId="11" pageNumber="22">
|
||
<emphasis id="6FDDE254FFE2B307FF26FC2D972DFC69" box="[199,260,907,932]" italics="true" pageId="11" pageNumber="22">Note.</emphasis>
|
||
The placement of the nominal taxon
|
||
<taxonomicName id="9AA945C5FFE2B307FD4CFC2D951BFC68" authorityName="Balint" authorityYear="2003" baseAuthorityName="Hewitson" baseAuthorityYear="1869" box="[685,818,907,933]" class="Insecta" family="Lycaenidae" genus="Johnsonita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="22" phylum="Arthropoda" rank="species" species="catadupa">
|
||
<emphasis id="6FDDE254FFE2B307FD4CFC2D951BFC68" box="[685,818,907,933]" italics="true" pageId="11" pageNumber="22">T. catadupa</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<taxonomicName id="9AA945C5FFE2B307FCB9FC2D925EFC68" authority="Johnson, 1992" authorityName="Johnson" authorityYear="1992" box="[856,1143,907,933]" class="Insecta" family="Lycaenidae" genus="Radissima" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="22" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6FDDE254FFE2B307FCB9FC2D95E4FC68" box="[856,973,907,933]" italics="true" pageId="11" pageNumber="22">Radissima</emphasis>
|
||
<bibRefCitation id="393843B7FFE2B307FC35FC2D925EFC68" author="Johnson" bookContentInfo="205 figs." box="[980,1143,907,933]" firstAuthor="Johnson" journalOrPublisher="Reports of the Museum of Natural History, University of Wisconsin, Stevens Point" pageId="11" pageNumber="22" pagination="1 - 135" part="2" refId="ref22864" refString="Johnson, K. (1992) Genera and species of the Neotropical " elfin " - like hairstreak butterflies (Lepidoptera, Lycaenidae, Theclinae). Reports of the Museum of Natural History, University of Wisconsin, Stevens Point, 22 (1 & 2), 1 - 135, 205 figs. & 136 - 279 + 241, 2 figs." title="Genera and species of the Neotropical " elfin " - like hairstreak butterflies (Lepidoptera, Lycaenidae, Theclinae)" type="book chapter" year="1992">Johnson, 1992</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<typeStatus id="821280E4FFE2B307FB66FC2D9290FC68" box="[1159,1209,907,933]" pageId="11" pageNumber="22">type</typeStatus>
|
||
species:
|
||
<taxonomicName id="9AA945C5FFE2B307FAC0FC2D975BFC04" authority="Geyer, 1837" authorityName="Geyer" authorityYear="1837" class="Insecta" family="Lycaenidae" genus="Sithon" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="22" phylum="Arthropoda" rank="species" species="umbratus">
|
||
<emphasis id="6FDDE254FFE2B307FAC0FC2D96F6FC04" italics="true" pageId="11" pageNumber="22">Sithon umbratus</emphasis>
|
||
Geyer, 1837
|
||
</taxonomicName>
|
||
, by original designation) is erroneous and based on the superficial similarity of wing shape and ventral wing surface pattern. The genus
|
||
<taxonomicName id="9AA945C5FFE2B307FDB3FC7594EEFC20" authorityName="Johnson" authorityYear="1992" box="[594,711,979,1005]" class="Insecta" family="Lycaenidae" genus="Radissima" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="22" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6FDDE254FFE2B307FDB3FC7594EEFC20" box="[594,711,979,1005]" italics="true" pageId="11" pageNumber="22">Radissima</emphasis>
|
||
</taxonomicName>
|
||
lacks the characteristically incurved inner margin behind the HW tornus and the postmedian pattern composed of dashed lines (cf. specimen documentation in
|
||
<bibRefCitation id="393843B7FFE2B307FB42FC5193B9FBDC" author="d'Abrera" box="[1187,1424,1014,1041]" firstAuthor="d'Abrera" journalOrPublisher="Hill House, Black Rock, Victoria" page="1137" pageId="11" pageNumber="22" pagination="1098 - 1270" refId="ref22067" refString="D'Abrera, B. (1995) In: Butterflies of the Neotropical Region. Part VII. Lycaenidae. Hill House, Black Rock, Victoria, pp. i - xi + 1098 - 1270." type="book chapter" volumeTitle="Butterflies of the Neotropical Region. Part VII. Lycaenidae" year="1995">d’Abrera 1995: 1137</bibRefCitation>
|
||
). It was placed in synonymy of
|
||
<taxonomicName id="9AA945C5FFE2B307FE0EFBBD94C5FBF8" authority="Kaye, 1904" authorityName="Kaye" authorityYear="1904" box="[495,748,1051,1077]" class="Insecta" family="Lycaenidae" genus="Paiwarria" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="22" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6FDDE254FFE2B307FE0EFBBD944BFBF9" box="[495,610,1051,1076]" italics="true" pageId="11" pageNumber="22">Paiwarria</emphasis>
|
||
Kaye, 1904
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="393843B7FFE2B307FCF9FBBD95F0FBFB" author="Robbins" box="[792,985,1051,1078]" editor="Lamas, G." firstAuthor="Robbins" journalOrPublisher="Scientific Publishers, Gainesville, Florida" pageId="11" pageNumber="22" pagination="118 - 137" refId="ref23342" refString="Robbins, R. K. (2004 b) Lycaenidae. Theclinae. Tribe Eumaeini,. In: Lamas, G. (Ed.), Checklist: Part 4 A. Hesperioidea - Papilionoidea. In: Heppner, J. B. (Ed.), Atlas of Neotropical Lepidoptera. Vol. 5 A. Association for Tropical Lepidoptera, Scientific Publishers, Gainesville, Florida, pp. 118 - 137." title="Lycaenidae. Theclinae. Tribe Eumaeini" type="book chapter" volumeTitle="Checklist: Part 4 A. Hesperioidea - Papilionoidea. In: Heppner, J. B" year="2004" yearSuffix="b">Robbins (2004b)</bibRefCitation>
|
||
, but the justification of such action was not presented.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |