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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.68.729" ID-GBIF-Dataset="84c8a2ac-d7b8-46f3-9888-ce6c1485ac65" ID-PMC="PMC3088436" ID-Pensoft-Pub="1313-2970-68-1" ID-PubMed="21594036" ModsDocAuthor="" ModsDocDate="2010" ModsDocID="1313-2970-68-1" ModsDocOrigin="ZooKeys 68" ModsDocTitle="Skeletons in confusion: a review of astrophorid sponges with (dicho)calthrops as structural megascleres (Porifera, Demospongiae, Astrophorida)" checkinTime="1451250894870" checkinUser="pensoft" docAuthor="Van Soest, Rob W. M., Beglinger, Elly J. &amp; De Voogd, Nicole J." docDate="2010" docId="CE798D4C1566762D9D7ADA59C7E8983D" docLanguage="en" docName="ZooKeys 68: 1-88" docOrigin="ZooKeys 68" docSource="http://dx.doi.org/10.3897/zookeys.68.729" docTitle="Calthropella (Pachataxa) enigmatica Levi &amp; Levi 1983" docType="treatment" docVersion="5" lastPageNumber="60" masterDocId="5D0AFFB7C87613675E79FFF7DA38AE2E" masterDocTitle="Skeletons in confusion: a review of astrophorid sponges with (dicho-) calthrops as structural megascleres (Porifera, Demospongiae, Astrophorida)" masterLastPageNumber="88" masterPageNumber="1" pageNumber="58" updateTime="1668164561298" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Skeletons in confusion: a review of astrophorid sponges with (dicho-) calthrops as structural megascleres (Porifera, Demospongiae, Astrophorida)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Van Soest, Rob W. M.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Beglinger, Elly J.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>De Voogd, Nicole J.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2010</mods:date>
<mods:detail type="volume">
<mods:number>68</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>88</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.68.729</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.68.729</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-68-1</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="159360730" LSID="urn:lsid:plazi:treatment:CE798D4C1566762D9D7ADA59C7E8983D" httpUri="http://treatment.plazi.org/id/CE798D4C1566762D9D7ADA59C7E8983D" lastPageId="59" lastPageNumber="60" pageId="57" pageNumber="58">
<subSubSection pageId="57" pageNumber="58" type="nomenclature">
<paragraph pageId="57" pageNumber="58">
<taxonomicName authority="Levi &amp; Levi, 1983" authorityName="Levi &amp; Levi" authorityYear="1983" class="Demospongiae" family="Calthropellidae" genus="Calthropella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calthropella (Pachataxa) enigmatica" order="Tetractinellida" pageId="57" pageNumber="58" phylum="Porifera" rank="species" species="enigmatica" subGenus="Pachataxa">
Calthropella Pachataxa enigmatica
<normalizedToken originalValue="Lévi">Levi</normalizedToken>
&amp;
<normalizedToken originalValue="Lévi">Levi</normalizedToken>
, 1983
</taxonomicName>
Figs 30
<normalizedToken originalValue="AF">A-F</normalizedToken>
</paragraph>
</subSubSection>
<subSubSection pageId="57" pageNumber="58" type="reference_group">
<paragraph pageId="57" pageNumber="58">
<taxonomicName class="Demospongiae" family="Ancorinidae" genus="Pachataxa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachataxa enigmatica" order="Tetractinellida" pageId="57" pageNumber="58" phylum="Porifera" rank="species" species="enigmatica">Pachataxa enigmatica</taxonomicName>
<bibRefCitation author="Levi, C" journalOrPublisher="Bulletin du Museum national d'Histoire naturelle (4)" pageId="69" pageNumber="70" pagination="101 - 168" title="Eponges Tetractinellides et Lithistides bathyales de Nouvelle- Caledonie." volume="5" year="1983">
<normalizedToken originalValue="Lévi">Levi</normalizedToken>
and
<normalizedToken originalValue="Lévi">Levi</normalizedToken>
1983
</bibRefCitation>
: 151, fig. 29.
</paragraph>
</subSubSection>
<subSubSection pageId="57" pageNumber="58" type="material examined">
<paragraph pageId="57" pageNumber="58">Material examined.</paragraph>
<paragraph pageId="57" pageNumber="58">Holotype MNHN DCL 2805; paratypes DCL 2895 (4 small sponges), both from New Caledonia, 22.8°S; 167.15°E, depth 355-360 m.</paragraph>
</subSubSection>
<subSubSection lastPageId="59" lastPageNumber="60" pageId="58" pageNumber="59" type="description">
<paragraph pageId="58" pageNumber="59">
<pageBreakToken pageId="58" pageNumber="59" start="start">Description</pageBreakToken>
.
</paragraph>
<paragraph pageId="58" pageNumber="59">
Massive with rounded flattened lobate outgrowths (Fig. 30A). Size of holotype 3.5
<normalizedToken originalValue="×3.5×">x3.5x</normalizedToken>
1.2 cm, paratypes slightly smaller to about 1 cm3. Brownish beige in alcohol. Small slightly raised oscules of less than 1 mm diameter are scattered in vague groups on the upper surface. Consistency firm to hard, crumbly.
</paragraph>
<paragraph pageId="58" pageNumber="59">Skeleton: confused, with a crust of microscleres carried by the triaene megascleres. A few broken oxea-like spicules were present in the slides made from the largest paratype.</paragraph>
<paragraph pageId="58" pageNumber="59">Spicules (Fig. 30B): Calthrops, spheroxyasters, ataxasters.</paragraph>
<paragraph pageId="58" pageNumber="59">
Calthrops (Fig. 30
<normalizedToken originalValue="BC">B-C</normalizedToken>
), occasionally with five-claded mesotriaene-like forms (called
<normalizedToken originalValue="centrotriaenes">'centrotriaenes'</normalizedToken>
by
<normalizedToken originalValue="Lévi">Levi</normalizedToken>
and
<normalizedToken originalValue="Lévi">Levi</normalizedToken>
), cladi 70
<normalizedToken originalValue="215.4">-215.4-</normalizedToken>
450
<normalizedToken originalValue="×">x</normalizedToken>
13
<normalizedToken originalValue="30.6">-30.6-</normalizedToken>
54
<normalizedToken originalValue="µm">µm</normalizedToken>
, cladomes 120
<normalizedToken originalValue="311">-311-</normalizedToken>
510
<normalizedToken originalValue="µm">µm</normalizedToken>
.
</paragraph>
<paragraph pageId="58" pageNumber="59">
Spheroxyasters (Figs 30
<normalizedToken originalValue="DE">D-E</normalizedToken>
), called
<normalizedToken originalValue="chiasters">'chiasters'</normalizedToken>
by
<normalizedToken originalValue="Lévi">Levi</normalizedToken>
and
<normalizedToken originalValue="Lévi">Levi</normalizedToken>
, but they do not have blunt rays), small, 4
<normalizedToken originalValue="8.7">-8.7-</normalizedToken>
12
<normalizedToken originalValue="µm">µm</normalizedToken>
diameter (but not as small as 4-5
<normalizedToken originalValue="µm">µm</normalizedToken>
as
<normalizedToken originalValue="Lévi">Levi</normalizedToken>
and
<normalizedToken originalValue="Lévi">Levi</normalizedToken>
stated).
</paragraph>
<paragraph pageId="59" pageNumber="60">
<pageBreakToken pageId="59" pageNumber="60" start="start">Ataxasters</pageBreakToken>
(Fig. 30F) are microspined, basically ovoid microrhabds, with many forms branched or irregular (shapes may be termed centrotylote rhabds, rods with side branch(es), round balls, cross-shaped, or rarely polyangular/aster-like), size ranging from 9
<normalizedToken originalValue="×">x</normalizedToken>
4 to 42
<normalizedToken originalValue="×">x</normalizedToken>
16
<normalizedToken originalValue="µm">µm</normalizedToken>
.
</paragraph>
<caption pageId="59" pageNumber="60">
<paragraph pageId="59" pageNumber="60">
Figure 30.
<taxonomicName class="Demospongiae" family="Calthropellidae" genus="Calthropella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calthropella (Pachataxa) enigmatica" order="Tetractinellida" pageId="59" pageNumber="60" phylum="Porifera" rank="species" species="enigmatica" subGenus="Pachataxa">Calthropella (Pachataxa) enigmatica</taxonomicName>
, paratype MNHN DCL 2895 from New Caledonia, A habit (photo P.
<normalizedToken originalValue="Cárdenas">Cardenas</normalizedToken>
, scale bar 1 cm) B overview of spicules C calthrops D larger oxyaster E smaller oxyaster F various ataxasters.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="59" pageNumber="60" type="habitat">
<paragraph pageId="59" pageNumber="60">Habitat.</paragraph>
<paragraph pageId="59" pageNumber="60">Deep water, 355 m.</paragraph>
</subSubSection>
<subSubSection pageId="59" pageNumber="60" type="distribution">
<paragraph pageId="59" pageNumber="60">Distribution.</paragraph>
<paragraph pageId="59" pageNumber="60">Known only from off New Caledonia.</paragraph>
</subSubSection>
<subSubSection pageId="59" pageNumber="60" type="remarks">
<paragraph pageId="59" pageNumber="60">Remarks.</paragraph>
<paragraph pageId="59" pageNumber="60">
The New Caledonian species differs from the Caribbean
<taxonomicName class="Demospongiae" family="Calthropellidae" genus="Calthropella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calthropella (Pachataxa) lithistina" order="Tetractinellida" pageId="59" pageNumber="60" phylum="Porifera" rank="species" species="lithistina" subGenus="Pachataxa">Calthropella (Pachataxa) lithistina</taxonomicName>
in overall spicule size and particularly in the small size of the oxyasters; that species is much larger and has a plate-shape.
<taxonomicName class="Demospongiae" family="Calthropellidae" genus="Calthropella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calthropella (Pachataxa) pyrifera" order="Tetractinellida" pageId="59" pageNumber="60" phylum="Porifera" rank="species" species="pyrifera" subGenus="Pachataxa">Calthropella (Pachataxa) pyrifera</taxonomicName>
sp. n. (see below) differs likewise in having larger oxyasters up to 24
<normalizedToken originalValue="µm">µm</normalizedToken>
diameter and asymmetrical pear-shaped ataxasters.
</paragraph>
</subSubSection>
</treatment>
</document>