199 lines
20 KiB
XML
199 lines
20 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.883.36193" ID-GBIF-Dataset="4f6d7148-d281-4073-8bc1-24ce51c3717b" ID-PMC="PMC6828828" ID-Pensoft-Pub="1313-2970-883-1" ID-Pensoft-UUID="EAC9B5058CE55C5AA9A344F6FEFA25D5" ID-PubMed="31719773" ID-ZooBank="7ABDE7F0DD424B968A1380E1E59B1515" ModsDocID="1313-2970-883-1" checkinTime="1572422626326" checkinUser="pensoft" docAuthor="Wiklund, Helena, Neal, Lenka, Glover, Adrian G., Drennan, Regan, Muriel Rabone, & Dahlgren, Thomas G." docDate="2019" docId="CE792592061A5F2BA4A06F798C76CC2D" docLanguage="en" docName="ZooKeys 883: 1-82" docOrigin="ZooKeys 883" docSource="http://dx.doi.org/10.3897/zookeys.883.36193" docTitle="Capitellidae sp. (NHM _ 1486" docType="treatment" docVersion="5" id="EAC9B5058CE55C5AA9A344F6FEFA25D5" lastPageNumber="1" masterDocId="EAC9B5058CE55C5AA9A344F6FEFA25D5" masterDocTitle="Abyssal fauna of polymetallic nodule exploration areas, eastern Clarion-Clipperton Zone, central Pacific Ocean: Annelida: Capitellidae, Opheliidae, Scalibregmatidae, and Travisiidae" masterLastPageNumber="82" masterPageNumber="1" pageNumber="1" updateTime="1668167974120" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Abyssal fauna of polymetallic nodule exploration areas, eastern Clarion-Clipperton Zone, central Pacific Ocean: Annelida: Capitellidae, Opheliidae, Scalibregmatidae, and Travisiidae</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wiklund, Helena</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Neal, Lenka</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Glover, Adrian G.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Drennan, Regan</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Muriel Rabone,</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Dahlgren, Thomas G.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="volume">
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<mods:number>883</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>82</mods:end>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.883.36193</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.883.36193</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-883-1</mods:identifier>
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<mods:identifier type="ZooBank">7ABDE7F0DD424B968A1380E1E59B1515</mods:identifier>
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<mods:identifier type="Pensoft-UUID">EAC9B5058CE55C5AA9A344F6FEFA25D5</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="159909629" LSID="urn:lsid:plazi:treatment:CE792592061A5F2BA4A06F798C76CC2D" httpUri="http://treatment.plazi.org/id/CE792592061A5F2BA4A06F798C76CC2D" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="CE792592-061A-5F2B-A4A0-6F798C76CC2D" authority="sp. (NHM _ 1486)" family="Capitellidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Capitellida" pageId="0" pageNumber="1" rank="family">Capitellidae sp. (NHM_1486)</taxonomicName>
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Capitellidae sp. NHM _ 1486 (specimen NHM _ 1486). A Lab image, whole specimen B Live image, anterior C Lab image, anterior, highlighting first three chaetigers (faded stain, pr = prostomium, pb = proboscis) D Lab image, thoracic capillary chaetae E Lab image, transitionary chaetigers between thorax and abdomen (faded stain) (t = thorax, h = hooks, a = abdomen) F Lab image, hooks G Lab image, whole specimen (methyl green stain). Scale bars: 1 mm (A, C); 25 μm (D, F)." figureDoi="10.3897/zookeys.883.36193.figure2" httpUri="https://binary.pensoft.net/fig/350254" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.883.36193.figure2">
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Fig. 2
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<normalizedToken originalValue="A–G">A-G</normalizedToken>
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</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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NHM_613 NHMUK ANEA 2019.7105, coll. 17 Feb. 2015,
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<geoCoordinate degrees="12" direction="north" minutes="23.175" orientation="latitude" precision="1" value="12.38625">12°23.175N</geoCoordinate>
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,
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<geoCoordinate degrees="116" direction="west" minutes="32.92" orientation="longitude" precision="9" value="-116.54867">116°32.92W</geoCoordinate>
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, 4202 m http://data.nhm.ac.uk/object/be34eb86-fc0c-411c-8eb9-e86774c6515a; NHM_1486 NHMUK ANEA 2019.7106, coll. 04 Mar. 2015,
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<geoCoordinate degrees="12" direction="north" minutes="29.70" orientation="latitude" precision="9" value="12.495">12°29.70N</geoCoordinate>
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,
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<geoCoordinate degrees="116" direction="west" minutes="39.01" orientation="longitude" precision="9" value="-116.65017">116°39.01W</geoCoordinate>
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, 4260 m http://data.nhm.ac.uk/object/05d46c60-8b4d-4623-b7ff-2e8089453d7e.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Species represented by one anterior fragment and one body fragment only. Specimen NHM_1486 posteriorly incomplete, 8 mm long and 0.6 mm wide for about 22 chaetigers (posterior part of fragment is damaged). Preserved specimens creamy white in ethanol (
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Capitellidae sp. NHM _ 1486 (specimen NHM _ 1486). A Lab image, whole specimen B Live image, anterior C Lab image, anterior, highlighting first three chaetigers (faded stain, pr = prostomium, pb = proboscis) D Lab image, thoracic capillary chaetae E Lab image, transitionary chaetigers between thorax and abdomen (faded stain) (t = thorax, h = hooks, a = abdomen) F Lab image, hooks G Lab image, whole specimen (methyl green stain). Scale bars: 1 mm (A, C); 25 μm (D, F)." figureDoi="10.3897/zookeys.883.36193.figure2" httpUri="https://binary.pensoft.net/fig/350254" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.883.36193.figure2">Fig. 2A</figureCitation>
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); live specimens creamy yellow to white semi-translucent (
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Capitellidae sp. NHM _ 1486 (specimen NHM _ 1486). A Lab image, whole specimen B Live image, anterior C Lab image, anterior, highlighting first three chaetigers (faded stain, pr = prostomium, pb = proboscis) D Lab image, thoracic capillary chaetae E Lab image, transitionary chaetigers between thorax and abdomen (faded stain) (t = thorax, h = hooks, a = abdomen) F Lab image, hooks G Lab image, whole specimen (methyl green stain). Scale bars: 1 mm (A, C); 25 μm (D, F)." figureDoi="10.3897/zookeys.883.36193.figure2" httpUri="https://binary.pensoft.net/fig/350254" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.883.36193.figure2">Fig. 2B</figureCitation>
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). Epithelium smooth (
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Capitellidae sp. NHM _ 1486 (specimen NHM _ 1486). A Lab image, whole specimen B Live image, anterior C Lab image, anterior, highlighting first three chaetigers (faded stain, pr = prostomium, pb = proboscis) D Lab image, thoracic capillary chaetae E Lab image, transitionary chaetigers between thorax and abdomen (faded stain) (t = thorax, h = hooks, a = abdomen) F Lab image, hooks G Lab image, whole specimen (methyl green stain). Scale bars: 1 mm (A, C); 25 μm (D, F)." figureDoi="10.3897/zookeys.883.36193.figure2" httpUri="https://binary.pensoft.net/fig/350254" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.883.36193.figure2">Fig. 2</figureCitation>
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).
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</paragraph>
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<caption doi="10.3897/zookeys.883.36193.figure2" httpUri="https://binary.pensoft.net/fig/350254" pageId="0" pageNumber="1" start="Figure 2" startId="F2">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 2.</emphasis>
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<taxonomicName family="Capitellidae" lsidName="" pageId="0" pageNumber="1" rank="family">Capitellidae</taxonomicName>
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sp. NHM_1486 (specimen NHM_1486).
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<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
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Lab image, whole specimen
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<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
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Live image, anterior
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<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
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Lab image, anterior, highlighting first three chaetigers (faded stain, pr = prostomium, pb = proboscis)
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<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
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Lab image, thoracic capillary chaetae
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<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
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Lab image, transitionary chaetigers between thorax and abdomen (faded stain) (t = thorax, h = hooks, a = abdomen)
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<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
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Lab image, hooks
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<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
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Lab image, whole specimen (methyl green stain). Scale bars: 1 mm (
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<emphasis bold="true" pageId="0" pageNumber="1">A, C</emphasis>
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); 25
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<normalizedToken originalValue="μm">μm</normalizedToken>
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(
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<emphasis bold="true" pageId="0" pageNumber="1">D, F</emphasis>
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).
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="1">
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Prostomium conical, anteriorly broadly rounded, slightly longer than wide (
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Capitellidae sp. NHM _ 1486 (specimen NHM _ 1486). A Lab image, whole specimen B Live image, anterior C Lab image, anterior, highlighting first three chaetigers (faded stain, pr = prostomium, pb = proboscis) D Lab image, thoracic capillary chaetae E Lab image, transitionary chaetigers between thorax and abdomen (faded stain) (t = thorax, h = hooks, a = abdomen) F Lab image, hooks G Lab image, whole specimen (methyl green stain). Scale bars: 1 mm (A, C); 25 μm (D, F)." figureDoi="10.3897/zookeys.883.36193.figure2" httpUri="https://binary.pensoft.net/fig/350254" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.883.36193.figure2">Fig. 2C</figureCitation>
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). Eyespots not observed. Nuchal organs everted, lightly pigmented, located at posterior border of prostomium. Peristomium as a narrow, smooth, achateous ring.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Chaetigers 1-10 (= thorax) with capillaries only. First chaetiger with chaetae in notopodia only, subsequent nine chaetigers with chaetae in both noto- and neuropodia. All thoracic chaetae slender, bilimbate capillaries (
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Capitellidae sp. NHM _ 1486 (specimen NHM _ 1486). A Lab image, whole specimen B Live image, anterior C Lab image, anterior, highlighting first three chaetigers (faded stain, pr = prostomium, pb = proboscis) D Lab image, thoracic capillary chaetae E Lab image, transitionary chaetigers between thorax and abdomen (faded stain) (t = thorax, h = hooks, a = abdomen) F Lab image, hooks G Lab image, whole specimen (methyl green stain). Scale bars: 1 mm (A, C); 25 μm (D, F)." figureDoi="10.3897/zookeys.883.36193.figure2" httpUri="https://binary.pensoft.net/fig/350254" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.883.36193.figure2">Fig. 2D</figureCitation>
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), arranged in two rows, with alternating longer and shorter capillaries, about 10 chaetae per ramus. Genital pores or lateral organs not observed under light microscopy.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Chaetigers 11-12 are considered transitional between thorax and abdomen marked by appearance of hooded hooks only in neuropodia, but segments are of similar thickness to those in anterior part of body (
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Capitellidae sp. NHM _ 1486 (specimen NHM _ 1486). A Lab image, whole specimen B Live image, anterior C Lab image, anterior, highlighting first three chaetigers (faded stain, pr = prostomium, pb = proboscis) D Lab image, thoracic capillary chaetae E Lab image, transitionary chaetigers between thorax and abdomen (faded stain) (t = thorax, h = hooks, a = abdomen) F Lab image, hooks G Lab image, whole specimen (methyl green stain). Scale bars: 1 mm (A, C); 25 μm (D, F)." figureDoi="10.3897/zookeys.883.36193.figure2" httpUri="https://binary.pensoft.net/fig/350254" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.883.36193.figure2">Fig. 2E</figureCitation>
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).
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Abdominal segments enlarged (inflated), without lobe (
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Capitellidae sp. NHM _ 1486 (specimen NHM _ 1486). A Lab image, whole specimen B Live image, anterior C Lab image, anterior, highlighting first three chaetigers (faded stain, pr = prostomium, pb = proboscis) D Lab image, thoracic capillary chaetae E Lab image, transitionary chaetigers between thorax and abdomen (faded stain) (t = thorax, h = hooks, a = abdomen) F Lab image, hooks G Lab image, whole specimen (methyl green stain). Scale bars: 1 mm (A, C); 25 μm (D, F)." figureDoi="10.3897/zookeys.883.36193.figure2" httpUri="https://binary.pensoft.net/fig/350254" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.883.36193.figure2">Fig. 2A</figureCitation>
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). All abdominal chaetigers with capillary chaetae only in notopodia and hooded hooks only in neuropodia. Abdominal capillaries similar to those in thorax, about 15 per ramus. Hooded hooks with long and slender shaft, with swelling around mid-point of shaft; with a main fang and about three rows of small teeth (
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Capitellidae sp. NHM _ 1486 (specimen NHM _ 1486). A Lab image, whole specimen B Live image, anterior C Lab image, anterior, highlighting first three chaetigers (faded stain, pr = prostomium, pb = proboscis) D Lab image, thoracic capillary chaetae E Lab image, transitionary chaetigers between thorax and abdomen (faded stain) (t = thorax, h = hooks, a = abdomen) F Lab image, hooks G Lab image, whole specimen (methyl green stain). Scale bars: 1 mm (A, C); 25 μm (D, F)." figureDoi="10.3897/zookeys.883.36193.figure2" httpUri="https://binary.pensoft.net/fig/350254" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.883.36193.figure2">Fig. 2F</figureCitation>
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); about 15 hooks per ramus. Remainder of body unknown.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="methyl green stain">
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<paragraph pageId="0" pageNumber="1">Methyl green stain.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Prostomium, chaetigers 4-6 and abdominal chaetigers do not stain (or at best stain very lightly). Peristomium, chaetigers 1-3 and 7-12/13 stain more strongly (
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Capitellidae sp. NHM _ 1486 (specimen NHM _ 1486). A Lab image, whole specimen B Live image, anterior C Lab image, anterior, highlighting first three chaetigers (faded stain, pr = prostomium, pb = proboscis) D Lab image, thoracic capillary chaetae E Lab image, transitionary chaetigers between thorax and abdomen (faded stain) (t = thorax, h = hooks, a = abdomen) F Lab image, hooks G Lab image, whole specimen (methyl green stain). Scale bars: 1 mm (A, C); 25 μm (D, F)." figureDoi="10.3897/zookeys.883.36193.figure2" httpUri="https://binary.pensoft.net/fig/350254" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.883.36193.figure2">Fig. 2G</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="genetic data">
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<paragraph pageId="0" pageNumber="1">Genetic data.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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GenBank MN217415-MN217416 for 16S and MN217495-MN217496 for 18S. COI was unsuccessful for this species, no identical matches on GenBank for 16S or 18S. The species is distinct from all other specimens in this study and in our phylogenetic analyses forms an unresolved clade with
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<taxonomicName class="Polychaeta" family="Capitellidae" genus="Barantolla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Barantolla lepte" pageId="0" pageNumber="1" phylum="Annelida" rank="species" species="lepte">
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<emphasis italics="true" pageId="0" pageNumber="1">Barantolla lepte</emphasis>
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</taxonomicName>
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Hutchings, 1974, three
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<taxonomicName class="Polychaeta" family="Capitellidae" genus="Notomastus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Notomastus" pageId="0" pageNumber="1" phylum="Annelida" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Notomastus</emphasis>
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</taxonomicName>
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M. Sars, 1851 and one
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<taxonomicName class="Polychaeta" family="Capitellidae" genus="Heteromastus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heteromastus" pageId="0" pageNumber="1" phylum="Annelida" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Heteromastus</emphasis>
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</taxonomicName>
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Eisig, 1887 species (
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Phylogenetic analysis of Capitellidae. 50 % majority rule tree from the Bayesian analyses using 18 S and 16 S, with posterior probability values on nodes. Twenty-four taxa from GenBank were included, and Echiura was chosen as outgroup following the annelid phylogeny of Weigert and Bleidorn (2016)." figureDoi="10.3897/zookeys.883.36193.figure4" httpUri="https://binary.pensoft.net/fig/350256" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.883.36193.figure4">Fig. 4</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="remarks">
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<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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This species is unusual amongst
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<taxonomicName family="Capitellidae" lsidName="" pageId="0" pageNumber="1" rank="family">Capitellidae</taxonomicName>
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due to its large number of mixed segments (with capillaries only in notopodia and hooded hooks only in neuropodia). Usually, the abdominal chaetigers in
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<taxonomicName family="Capitellidae" lsidName="" pageId="0" pageNumber="1" rank="family">Capitellidae</taxonomicName>
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bear hooded hooks only, or there are a small number of posterior thoracic and/or anterior abdominal segments that bear both capillaries and hooks. Of the known genera, only
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<taxonomicName class="Polychaeta" family="Capitellidae" genus="Promastobranchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Promastobranchus" pageId="0" pageNumber="1" phylum="Annelida" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Promastobranchus</emphasis>
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</taxonomicName>
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Gallardo, 1968 shows such chaetal distribution. However, it can be distinguished from the other UKSR-collected species in having the first chaetiger with both noto- and neurochaetae and possessing 12 to 13 instead of 10 chaetigers with capillary chaetae in both rami. Therefore, this material represents not only a new species, but based on current morphological criteria supported also by genetic data, a new genus as well. Because the material is not complete (morphology for the posterior end is missing) the species is here not formally described.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="ecology">
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<paragraph pageId="0" pageNumber="1">Ecology.</paragraph>
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<paragraph pageId="0" pageNumber="1">Found in the eastern polymetallic nodule province of the CCZ.</paragraph>
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</subSubSection>
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</treatment>
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</document> |