302 lines
36 KiB
XML
302 lines
36 KiB
XML
<document id="C4592B50F8300A80184B90F9710AB7CB" ID-DOI="10.5281/zenodo.210542" ID-GBIF-Dataset="c9e5cc54-2a89-4b75-91aa-a4fce79f17fc" ID-ISSN="1175-5326" ID-Zenodo-Dep="210542" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460500319311" checkinUser="plazi" docAuthor="Ramírez-Ponce, Andrés & Morón, Miguel Angel" docDate="2012" docId="A567C22F585AFF84FF64A2CC50C585C0" docLanguage="en" docName="zt03394p024.pdf" docOrigin="Zootaxa 3394" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Pachystethus nutans Bates 1888, new combination" docType="treatment" docVersion="9" lastPageNumber="13" masterDocId="595EBA575850FF88FFF3A25B52478434" masterDocTitle="Revision of the genus Pachystethus Blanchard with description of three new species from Mexico (Coleoptera: Scarabaeidae: Rutelinae: Anomalini)" masterLastPageNumber="24" masterPageNumber="1" pageNumber="11" updateTime="1698311598655" updateUser="plazi">
|
||
<mods:mods id="BAF4B0FB9B9409F76613A77B986254F5" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="309CB526C500935DEC7A9CF869EAEEDA">
|
||
<mods:title id="6CEC40642117DDB3B762F56C84C92E80">Revision of the genus Pachystethus Blanchard with description of three new species from Mexico (Coleoptera: Scarabaeidae: Rutelinae: Anomalini)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="C1CDBF26CC29A60F892B1E98A664C237" type="personal">
|
||
<mods:role id="12E7DF85BFFCA153C118177638BD3756">
|
||
<mods:roleTerm id="45398CCBEE05C17644D5348FD4609625">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="58303815D7FAEB20AA2093A87C4544B6">Ramírez-Ponce, Andrés</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="FAD475960BE4EA5EC2D3ED77E188CBA9" type="personal">
|
||
<mods:role id="68ED965BA41CAFA4FACBD8552BFEB620">
|
||
<mods:roleTerm id="3D344E91CEE1207BAC583B129BE0EC6B">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="AEE7A8B36C691174AD3EA375D8319722">Morón, Miguel Angel</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="62657E1B05A4514EDB7E7002E48FCEBA">text</mods:typeOfResource>
|
||
<mods:relatedItem id="8DA06DDCB907076A5E5F4107889119D1" type="host">
|
||
<mods:titleInfo id="B28AD2F03E1F09EEA56AF11A69E002B5">
|
||
<mods:title id="9AC98B02CBC83A3FEC68697C39B7DAE5">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="915C777C803768AF3080031C1D721996">
|
||
<mods:date id="0FFE8293E53ABA831005C91BE94EA073">2012</mods:date>
|
||
<mods:detail id="376E9A724E680FB203AADE10DB9E9823" type="volume">
|
||
<mods:number id="72E184D41A1A7002BD07797945F28451">3394</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="9855A59E515C195C3BBD6F10F912A53F" unit="page">
|
||
<mods:start id="B082D340BE49638957ABF5E957C3BB1C">1</mods:start>
|
||
<mods:end id="529A1C08536F4FA813E6719FABA3C5A8">24</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="2920EF9FAA2C440E0B7A53F1B6B268E2">journal article</mods:classification>
|
||
<mods:identifier id="C683785271302765AE2DC985834735CC" type="DOI">10.5281/zenodo.210542</mods:identifier>
|
||
<mods:identifier id="693D0FF0DF86C83BD0B3C8CB237A3472" type="GBIF-Dataset">c9e5cc54-2a89-4b75-91aa-a4fce79f17fc</mods:identifier>
|
||
<mods:identifier id="B0B0A45BA3D44053491746286854B2D0" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="8CFF900C52C0CD64ED53C4E0984F8A48" type="Zenodo-Dep">210542</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="A567C22F585AFF84FF64A2CC50C585C0" ID-DOI="http://doi.org/10.5281/zenodo.6179198" ID-GBIF-Taxon="119521052" ID-Zenodo-Dep="6179198" LSID="urn:lsid:plazi:treatment:A567C22F585AFF84FF64A2CC50C585C0" httpUri="http://treatment.plazi.org/id/A567C22F585AFF84FF64A2CC50C585C0" lastPageId="12" lastPageNumber="13" pageId="10" pageNumber="11">
|
||
<subSubSection id="65D420B2585AFF82FF64A2CC53EE84E6" pageId="10" pageNumber="11" type="nomenclature">
|
||
<paragraph id="2D717339585AFF82FF64A2CC51518485" blockId="10.[151,790,151,210]" box="[151,790,151,177]" pageId="10" pageNumber="11">
|
||
<heading id="7639C455585AFF82FF64A2CC51518485" bold="true" box="[151,790,151,177]" fontSize="11" level="1" pageId="10" pageNumber="11" reason="1">
|
||
<emphasis id="1FBAAF2B585AFF82FF64A2CC51518485" bold="true" box="[151,790,151,177]" pageId="10" pageNumber="11">
|
||
<taxonomicName id="EACE08BA585AFF82FF64A2CC507F8485" authority="Bates, 1888" authorityName="Bates" authorityYear="1888" box="[151,568,151,177]" class="Insecta" family="Rutelidae" genus="Pachystethus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="nutans" status="comb. nov.">
|
||
<emphasis id="1FBAAF2B585AFF82FF64A2CC53D68485" bold="true" box="[151,401,151,177]" italics="true" pageId="10" pageNumber="11">Pachystethus nutans</emphasis>
|
||
(
|
||
<bibRefCitation id="495F0EC8585AFF82FE53A2CC50778485" author="Bates" box="[416,560,151,177]" pageId="10" pageNumber="11" refString="Bates, H. W. (1888) Pectinicornia and Lamellicornia. In: O. Salvin & F. D. Godwin (eds.), Biologia Centrali-Americana. Insecta Coleoptera. Volume 2, part 2: 1 - 432." type="book chapter" year="1888">Bates, 1888</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="04891250585AFF82FDB3A2CC51518485" box="[576,790,151,177]" pageId="10" pageNumber="11">new combination</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="2D717339585AFF82FF64A2E153EE84E6" blockId="10.[151,790,151,210]" box="[151,425,186,210]" pageId="10" pageNumber="11">
|
||
(
|
||
<figureCitation id="B5F56FBC585AFF82FF6CA2E1536084E6" box="[159,295,186,210]" captionStart="FIGURES 14 – 19" captionStartId="11.[151,264,1976,1999]" captionTargetBox="[167,1348,573,1945]" captionTargetId="figure@11.[151,1436,553,1965]" captionTargetPageId="11" captionText="FIGURES 14 – 19. Diagnostic structures in P. nu t an s. 14) right proleg, 15) right antenna, 16) male genitalia in dorsal view, 17) male genitalia in lateral view, 18) spiculum gastrale, 19) female genital plates. Scale lines 1 mm." httpUri="https://zenodo.org/record/210546/files/figure.png" pageId="10" pageNumber="11">Figs. 14–19</figureCitation>
|
||
,
|
||
<figureCitation id="B5F56FBC585AFF82FEC1A2E1530884E6" box="[306,335,186,210]" captionStart="FIGURES 42 – 47" captionStartId="19.[151,264,1909,1932]" captionTargetBox="[214,1377,207,1832]" captionTargetId="figure@19.[198,1392,193,1847]" captionTargetPageId="19" captionText="FIGURES 42 – 47. Dorsal view of Pachystethus species: 42 a, b) P. viduus males; 42 c) P. viduus female; 42 d) P. viduus female, lectotype; 43 a) P. nectoctenus male; 43 b) P. nectoctenus female; 44 a) P. crassesculptus male; 44 b) P. crassesculptus female; 45 a, b) P. nutans males; 45 c) P. nu t an s female; 46 a, b) P. matzapatlecus males; 46 c) P. matzapatlecus female; 47 a) P. ixtacomitanus male; 47 b) P. ixtacomitanus female. Scale lines 1 mm." httpUri="https://zenodo.org/record/210550/files/figure.png" pageId="10" pageNumber="11">45</figureCitation>
|
||
a–c, 51)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="65D420B2585AFF82FF64A35953AA852C" box="[151,493,258,280]" pageId="10" pageNumber="11" type="reference_group">
|
||
<paragraph id="2D717339585AFF82FF64A35953AA852C" blockId="10.[151,493,258,280]" box="[151,493,258,280]" pageId="10" pageNumber="11">
|
||
<treatmentCitationGroup id="0DDE5417585AFF82FF64A35953AA852C" box="[151,493,258,280]" pageId="10" pageNumber="11">
|
||
<taxonomicName id="EACE08BA585AFF82FF64A35953AA852C" authority="Bates, 1888: 246" authorityName="Bates" authorityPageNumber="246" authorityYear="1888" box="[151,493,258,280]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="nutans">
|
||
<emphasis id="1FBAAF2B585AFF82FF64A359537A852C" box="[151,317,258,280]" italics="true" pageId="10" pageNumber="11">Anomala nutans</emphasis>
|
||
<treatmentCitation id="AC6F5528585AFF82FEB1A35853AA852C" author="Bates" box="[322,493,258,280]" page="246" pageId="10" pageNumber="11" year="1888">
|
||
<bibRefCitation id="495F0EC8585AFF82FEB1A35853FA852C" author="Bates" box="[322,445,259,280]" pageId="10" pageNumber="11" refString="Bates, H. W. (1888) Pectinicornia and Lamellicornia. In: O. Salvin & F. D. Godwin (eds.), Biologia Centrali-Americana. Insecta Coleoptera. Volume 2, part 2: 1 - 432." type="book chapter" year="1888">Bates, 1888</bibRefCitation>
|
||
: 246
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="65D420B2585AFF82FF64A31E571186F1" pageId="10" pageNumber="11" type="materials_examined">
|
||
<paragraph id="2D717339585AFF82FF64A31E57148569" blockId="10.[151,1437,325,2005]" box="[151,1363,325,350]" pageId="10" pageNumber="11">
|
||
<emphasis id="1FBAAF2B585AFF82FF64A31E531E856A" bold="true" box="[151,345,325,350]" pageId="10" pageNumber="11">
|
||
<typeStatus id="F275CD9B585AFF82FF64A31E52958569" box="[151,210,325,349]" pageId="10" pageNumber="11">Type</typeStatus>
|
||
specimens.
|
||
</emphasis>
|
||
Male and female
|
||
<typeStatus id="F275CD9B585AFF82FDD0A31D50C2856A" box="[547,645,326,350]" pageId="10" pageNumber="11" type="syntype">syntypes</typeStatus>
|
||
examined, deposited in
|
||
<collectionCode id="4BDFEBFC585AFF82FC65A31D519C8569" box="[918,987,326,349]" pageId="10" pageNumber="11">NHM</collectionCode>
|
||
.
|
||
<typeStatus id="F275CD9B585AFF82FC16A31D565B856A" box="[997,1052,326,350]" pageId="10" pageNumber="11">Type</typeStatus>
|
||
locality: “Juquila,
|
||
<collectingCountry id="55D933A9585AFF82FB01A31E570D8569" box="[1266,1354,325,349]" name="Mexico" pageId="10" pageNumber="11">Mexico</collectingCountry>
|
||
”
|
||
</paragraph>
|
||
<paragraph id="2D717339585AFF82FF3AA332571186F1" blockId="10.[151,1437,325,2005]" pageId="10" pageNumber="11">
|
||
<emphasis id="1FBAAF2B585AFF82FF3AA332538A85B5" bold="true" box="[201,461,360,385]" pageId="10" pageNumber="11">
|
||
<typeStatus id="F275CD9B585AFF82FF3AA332534385B5" box="[201,260,361,385]" pageId="10" pageNumber="11">Type</typeStatus>
|
||
material revised
|
||
</emphasis>
|
||
(
|
||
<specimenCount id="3BC8B8B0585AFF82FE13A332506D85B5" box="[480,554,361,385]" pageId="10" pageNumber="11" type="male">1 male</specimenCount>
|
||
<typeStatus id="F275CD9B585AFF82FDC0A33250DA85B5" box="[563,669,361,385]" pageId="10" pageNumber="11" type="lectotype">lectotype</typeStatus>
|
||
,
|
||
<specimenCount id="3BC8B8B0585AFF82FD5BA332514D85B5" box="[680,778,361,385]" pageId="10" pageNumber="11" type="female">1 female</specimenCount>
|
||
<typeStatus id="F275CD9B585AFF82FCE1A33251EA85B5" box="[786,941,361,385]" pageId="10" pageNumber="11" type="paralectotype">paralectotype</typeStatus>
|
||
:
|
||
<collectionCode id="4BDFEBFC585AFF82FC4AA33251BB85B5" box="[953,1020,361,385]" pageId="10" pageNumber="11">NHM</collectionCode>
|
||
) labeled:
|
||
<emphasis id="1FBAAF2B585AFF82FB9FA332576585B5" bold="true" box="[1132,1314,361,385]" pageId="10" pageNumber="11">
|
||
<typeStatus id="F275CD9B585AFF82FB9FA332569085B5" box="[1132,1239,361,385]" pageId="10" pageNumber="11" type="lectotype">lectotype</typeStatus>
|
||
male:
|
||
</emphasis>
|
||
a) “MEX- ICO: [Oaxaca] Juquila. Salle Coll. B. C. A., Coll., II (2)” (typeface), b) “
|
||
<taxonomicName id="EACE08BA585AFF82FC21A3D657118591" authority="Bates, 1888" authorityName="Bates" authorityYear="1888" box="[978,1366,397,421]" class="Insecta" family="Scarabaeidae" genus="Pachyststhus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="nutans">
|
||
<emphasis id="1FBAAF2B585AFF82FC21A3D656F08591" box="[978,1207,397,421]" italics="true" pageId="10" pageNumber="11">Pachyststhus nutans</emphasis>
|
||
(
|
||
<bibRefCitation id="495F0EC8585AFF82FB35A3D657098591" author="Bates" box="[1222,1358,397,421]" pageId="10" pageNumber="11" refString="Bates, H. W. (1888) Pectinicornia and Lamellicornia. In: O. Salvin & F. D. Godwin (eds.), Biologia Centrali-Americana. Insecta Coleoptera. Volume 2, part 2: 1 - 432." type="book chapter" year="1888">Bates, 1888</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
LEC- TOTYPE male Ramírez-Ponce y Morón det. 2011” (typeface;
|
||
<collectionCode id="4BDFEBFC585AFF82FC60A3EA519D85FD" box="[915,986,433,457]" pageId="10" pageNumber="11">NHM</collectionCode>
|
||
).
|
||
<emphasis id="1FBAAF2B585AFF82FC00A3EA577C85FD" bold="true" box="[1011,1339,432,457]" pageId="10" pageNumber="11">
|
||
<typeStatus id="F275CD9B585AFF82FC00A3EA562E85FD" box="[1011,1129,433,457]" pageId="10" pageNumber="11" type="lectotype">Lectotype</typeStatus>
|
||
here designated
|
||
</emphasis>
|
||
for the taxonomic purpose to preserve nomenclatural stability by selecting a single specimen as the name-bearing
|
||
<typeStatus id="F275CD9B585AFF82FABEA38D573A85DA" box="[1357,1405,470,494]" pageId="10" pageNumber="11">type</typeStatus>
|
||
of the name of this taxon.
|
||
<emphasis id="1FBAAF2B585AFF82FE59A3A250F68625" bold="true" box="[426,689,504,529]" pageId="10" pageNumber="11">
|
||
<typeStatus id="F275CD9B585AFF82FE59A3A250088625" box="[426,591,505,529]" pageId="10" pageNumber="11" type="paralectotype">Paralectotype</typeStatus>
|
||
female:
|
||
</emphasis>
|
||
“
|
||
<collectingCountry id="55D933A9585AFF82FD37A3A2517E8625" box="[708,825,505,529]" name="Mexico" pageId="10" pageNumber="11">MEXICO</collectingCountry>
|
||
: [Oaxaca] Juquila. Salle Coll. B. C. A., Coll., II (2)” (typeface), b) “
|
||
<taxonomicName id="EACE08BA585AFF82FEB3A04650858601" authority="Bates, 1888" authorityName="Bates" authorityYear="1888" box="[320,706,541,565]" class="Insecta" family="Rutelidae" genus="Pachystethus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="nutans">
|
||
<emphasis id="1FBAAF2B585AFF82FEB3A04650618601" box="[320,550,541,565]" italics="true" pageId="10" pageNumber="11">Pachystethus nutans</emphasis>
|
||
(
|
||
<bibRefCitation id="495F0EC8585AFF82FDC7A04650FD8601" author="Bates" box="[564,698,541,565]" pageId="10" pageNumber="11" refString="Bates, H. W. (1888) Pectinicornia and Lamellicornia. In: O. Salvin & F. D. Godwin (eds.), Biologia Centrali-Americana. Insecta Coleoptera. Volume 2, part 2: 1 - 432." type="book chapter" year="1888">Bates, 1888</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<typeStatus id="F275CD9B585AFF82FD3AA04651F68601" box="[713,945,541,565]" pageId="10" pageNumber="11" type="paralectotype">PARALECTOTYPE</typeStatus>
|
||
female Ramírez-Ponce y Morón des. 2011” (typeface;
|
||
<collectionCode id="4BDFEBFC585AFF82FEE3A01A5311866D" box="[272,342,577,601]" pageId="10" pageNumber="11">NHM</collectionCode>
|
||
).
|
||
<emphasis id="1FBAAF2B585AFF82FE99A01B50F1866D" bold="true" box="[362,694,576,601]" pageId="10" pageNumber="11">Additional material revised</emphasis>
|
||
(
|
||
<specimenCount id="3BC8B8B0585AFF82FD3BA01A5163866D" box="[712,804,577,601]" pageId="10" pageNumber="11" type="male">3 males</specimenCount>
|
||
;
|
||
<collectionCode id="4BDFEBFC585AFF82FCC2A01A513F866D" box="[817,888,577,601]" httpUri="http://grbio.org/cool/qbc0-xx4g" name="Coleccion Nacional de Insectos, Universidad Nacional Autonoma de Mexico" pageId="10" pageNumber="11">CNIN</collectionCode>
|
||
,
|
||
<collectionCode id="4BDFEBFC585AFF82FC75A01A5183866D" box="[902,964,577,601]" pageId="10" pageNumber="11">DDC</collectionCode>
|
||
,
|
||
<collectionCode id="4BDFEBFC585AFF82FC22A01A5662866D" box="[977,1061,577,601]" pageId="10" pageNumber="11">MXAL</collectionCode>
|
||
) labeled:
|
||
<collectingCountry id="55D933A9585AFF82FB64A01A56B6866D" box="[1175,1265,577,601]" name="Mexico" pageId="10" pageNumber="11">Mexico</collectingCountry>
|
||
: Oaxaca,
|
||
<quantity id="EA36DEDC585AFF82FA92A01A57DB866D" box="[1377,1436,577,601]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="10" pageNumber="11" unit="km" value="6.0">6 km</quantity>
|
||
SE Lachao, El Polvorín,
|
||
<quantity id="EA36DEDC585AFF82FE42A03E504E8649" box="[433,521,613,637]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="10" pageNumber="11" unit="m" value="1900.0">1900 m</quantity>
|
||
,
|
||
<date id="597055F9585AFF82FDE5A03E50D68649" box="[534,657,613,637]" pageId="10" pageNumber="11" value="2006-05-27">27-V-2006</date>
|
||
, D. Curoe (typeface) (
|
||
<specimenCount id="3BC8B8B0585AFF82FC61A03E51988649" box="[914,991,613,637]" pageId="10" pageNumber="11" type="male">1 male</specimenCount>
|
||
, DCC); a)
|
||
<taxonomicName id="EACE08BA585AFF82FBAFA03E5710864A" authority="Deyz" authorityName="Deyz" box="[1116,1367,613,638]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="nutans">
|
||
<emphasis id="1FBAAF2B585AFF82FBAFA03E57538649" box="[1116,1300,613,637]" italics="true" pageId="10" pageNumber="11">Anomala nutans</emphasis>
|
||
Deyz
|
||
</taxonomicName>
|
||
Veracruz (typeface), b) “
|
||
<taxonomicName id="EACE08BA585AFF82FE8AA0D2502D8695" authority="Bat." authorityName="Bat." box="[377,618,649,673]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="nutans">Anomala nutans Bat.</taxonomicName>
|
||
” (handwritten), c) “1670” (typeface), d) “Ex-colección Eugenio Dugés” (typeface) (
|
||
<specimenCount id="3BC8B8B0585AFF82FEE8A0F6532086F1" box="[283,359,685,709]" pageId="10" pageNumber="11" type="male">1 male</specimenCount>
|
||
,
|
||
<collectionCode id="4BDFEBFC585AFF82FE81A0F653FB86F1" box="[370,444,685,709]" httpUri="http://grbio.org/cool/qbc0-xx4g" name="Coleccion Nacional de Insectos, Universidad Nacional Autonoma de Mexico" pageId="10" pageNumber="11">CNIN</collectionCode>
|
||
); a) “
|
||
<collectingCountry id="55D933A9585AFF82FE05A0F6501786F1" box="[502,592,685,709]" name="Mexico" pageId="10" pageNumber="11">Mexico</collectingCountry>
|
||
, Jalapa, Hoge”, b) “
|
||
<taxonomicName id="EACE08BA585AFF82FCDCA0F6564286F1" authority="Bts." authorityName="Bts." box="[815,1029,685,709]" class="Insecta" family="Scarabaeidae" genus="Spilota" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="nutans">Spilota nutans Bts.</taxonomicName>
|
||
” (typeface) (
|
||
<specimenCount id="3BC8B8B0585AFF82FB69A0F656A086F1" box="[1178,1255,685,709]" pageId="10" pageNumber="11" type="male">1 male</specimenCount>
|
||
,
|
||
<collectionCode id="4BDFEBFC585AFF82FB01A0F6570D86F1" box="[1266,1354,685,709]" pageId="10" pageNumber="11">MXAL</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="65D420B2585AFF83FF3AA08A50ED850B" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="description">
|
||
<paragraph id="2D717339585AFF82FF3AA08A57318315" blockId="10.[151,1437,325,2005]" pageId="10" pageNumber="11">
|
||
<emphasis id="1FBAAF2B585AFF82FF3AA08A53FA86DE" bold="true" box="[201,445,721,746]" pageId="10" pageNumber="11">Redescription. Male</emphasis>
|
||
(
|
||
<emphasis id="1FBAAF2B585AFF82FE23A089539986DD" box="[464,478,722,745]" italics="true" pageId="10" pageNumber="11">n</emphasis>
|
||
= 4). Length
|
||
<quantity id="EA36DEDC585AFF82FD77A08A515786DD" box="[644,784,721,745]" metricMagnitude="-3" metricUnit="m" metricValue="9.35" metricValueMax="9.6" metricValueMin="9.1" pageId="10" pageNumber="11" unit="mm" value="9.35" valueMax="9.6" valueMin="9.1">9.1–9.6 mm</quantity>
|
||
, width
|
||
<quantity id="EA36DEDC585AFF82FC9BA08A518386DD" box="[872,964,721,745]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="10" pageNumber="11" unit="mm" value="4.2">4.2 mm</quantity>
|
||
. Color reddish yellow on scape, pedicel, funicle, lateral margins of pronotum and elytra; dark red on clypeal margins, apexes of femora, tibiae, distal tarsi, and claws; shiny black on antennal club, frons, disc of pronotum, scutelum, elytral suture, and legs (except their apices).
|
||
<emphasis id="1FBAAF2B585AFF82FF07A166537E8761" box="[244,313,829,853]" italics="true" pageId="10" pageNumber="11">Head.</emphasis>
|
||
Clypeus subrectangular, clearly wider than long; anterior border straight, strongly upturned and not directed forward, anterior angles rounded; clypeal surface with irregular, dense, deep, large punctures. Frontoclypeal suture straight, partially hidden by punctures. Frons flattened, glabrous, densely rugopunctate, punctures irregularly elongate and fused at center, more disperse on lateral and posterior margins. Punctures around eyes deep but not coalescent, with some long setae. Antennal club longer than the length of all preceding antennomeres combined; lamellar shape ovate, curved inward (
|
||
<figureCitation id="B5F56FBC585AFF82FD7CA1AA50A3803D" box="[655,740,1009,1033]" captionStart="FIGURES 14 – 19" captionStartId="11.[151,264,1976,1999]" captionTargetBox="[167,1348,573,1945]" captionTargetId="figure@11.[151,1436,553,1965]" captionTargetPageId="11" captionText="FIGURES 14 – 19. Diagnostic structures in P. nu t an s. 14) right proleg, 15) right antenna, 16) male genitalia in dorsal view, 17) male genitalia in lateral view, 18) spiculum gastrale, 19) female genital plates. Scale lines 1 mm." httpUri="https://zenodo.org/record/210546/files/figure.png" pageId="10" pageNumber="11">Fig. 15</figureCitation>
|
||
). Eyes medium size, ovate; ocular margin noticeable and setose. Eye canthus thick with glabrous and rounded apex, raised surface, irregular deep punctures. Maxillary basistipe convex, with irregular wide punctation; labrum wide and sinuate; labium sinuate, slightly longer than wide, irregularly punctate; prementum setose.
|
||
<emphasis id="1FBAAF2B585AFF82FD54A60551518041" box="[679,790,1118,1141]" italics="true" pageId="10" pageNumber="11">Pronotum</emphasis>
|
||
wider than long, moderately convex; surface with abundant deep, large, elongate punctures; lateral borders rounded, with bulky sides and weak lateral pits; posterior angles raised. Mesoepimera hidden under humeral calli. Space between mesocoxae wide; mesometasternal process long, projecting anteriorly past mesocoxa but do not reach procoxae with wide rounded, apex directed downward; metasternum moderately and sparsely punctate, sides uniformly setiferous, mesial region glabrous with weak longitudinal sulcus.
|
||
<emphasis id="1FBAAF2B585AFF82FEA9A74A53E7811D" box="[346,416,1297,1321]" italics="true" pageId="10" pageNumber="11">Elytra</emphasis>
|
||
convex, wide, ovate; striae deeply impressed with uniform, continuous, large punctures; elytral costae absent, suture raised towards posterior half; wide preapical slope; lateral margins with wide notch posterior to the end of epipleural fold (
|
||
<figureCitation id="B5F56FBC585AFF82FDA4A70250ED8145" box="[599,682,1369,1393]" captionStart="FIGURES 48 – 53" captionStartId="20.[151,264,1259,1282]" captionTargetBox="[200,1411,240,1184]" captionTargetId="figure@20.[151,1436,193,1238]" captionTargetPageId="20" captionText="FIGURES 48 – 53. Lateral view of Pachystethus species: 48) P. viduus a) male, b) female; 49) P. nectoctenus a) male, b) female; 50) P. crassesculptus male; 51) P. n u t a n s male; 52) P. matzapatlecus male; 53) P. ixtacomitanus male. Scale lines 1 mm." httpUri="https://zenodo.org/record/210551/files/figure.png" pageId="10" pageNumber="11">Fig. 51</figureCitation>
|
||
) and discontinuous row of sparse, short setae along entire length. Epipleural fold wide, uniformly swollen in lateral view, gradually narrowed after metacoxa.
|
||
<taxonomicName id="EACE08BA585AFF82FB34A726577281A1" box="[1223,1333,1405,1429]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Corynexochida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1FBAAF2B585AFF82FB34A726577281A1" box="[1223,1333,1405,1429]" italics="true" pageId="10" pageNumber="11">Scutellum</emphasis>
|
||
</taxonomicName>
|
||
: slightly longer than wide; lateral margins angled, apex acute; surface with sparse, deep, large punctures.
|
||
<emphasis id="1FBAAF2B585AFF82FB36A7FA5768818D" box="[1221,1327,1441,1465]" italics="true" pageId="10" pageNumber="11">Abdomen</emphasis>
|
||
: sternites nearly completely covered with setae and moderate punctation.
|
||
<emphasis id="1FBAAF2B585AFF82FC94A79D51DB81E9" box="[871,924,1478,1501]" italics="true" pageId="10" pageNumber="11">Legs</emphasis>
|
||
: protarsomeres 1–4 shortened, slightly wider than and as long as protarsomere 5. Protibia stout; lateral borders convergent towards apex; external border with proximal denticle short, acute; distal denticle long, curved (
|
||
<figureCitation id="B5F56FBC585AFF82FCC6A45651CE8211" box="[821,905,1549,1573]" captionStart="FIGURES 14 – 19" captionStartId="11.[151,264,1976,1999]" captionTargetBox="[167,1348,573,1945]" captionTargetId="figure@11.[151,1436,553,1965]" captionTargetPageId="11" captionText="FIGURES 14 – 19. Diagnostic structures in P. nu t an s. 14) right proleg, 15) right antenna, 16) male genitalia in dorsal view, 17) male genitalia in lateral view, 18) spiculum gastrale, 19) female genital plates. Scale lines 1 mm." httpUri="https://zenodo.org/record/210546/files/figure.png" pageId="10" pageNumber="11">Fig. 14</figureCitation>
|
||
). Mesofemora with a well-defined longitudinal keel, with long setae. Metafemora with incomplete setiferous punctures, transverse keel. Metatibia with scattered, setiferous, deep, large punctures; middle and apex of metatibia clearly swollen. Metatarsomeres 1–2 short, as long as metatarsomeres 3–5.
|
||
<emphasis id="1FBAAF2B585AFF82FE5AA422505482A5" box="[425,531,1657,1681]" italics="true" pageId="10" pageNumber="11">Genitalia</emphasis>
|
||
(
|
||
<figureCitation id="B5F56FBC585AFF82FDD7A42250F782A5" box="[548,688,1657,1681]" captionStart="FIGURES 14 – 19" captionStartId="11.[151,264,1976,1999]" captionTargetBox="[167,1348,573,1945]" captionTargetId="figure@11.[151,1436,553,1965]" captionTargetPageId="11" captionText="FIGURES 14 – 19. Diagnostic structures in P. nu t an s. 14) right proleg, 15) right antenna, 16) male genitalia in dorsal view, 17) male genitalia in lateral view, 18) spiculum gastrale, 19) female genital plates. Scale lines 1 mm." httpUri="https://zenodo.org/record/210546/files/figure.png" pageId="10" pageNumber="11">Figs. 16–18</figureCitation>
|
||
): parameres simple, shorter than the length of tectum, with upper part concave and smooth, strongly narrowed towards apex; lower part of parameres curved backwards. Basal half of tectum as wide as distal half, with outstanding posterolateral calli. Basal piece with long, curved, sharply pointed apodema.
|
||
<emphasis id="1FBAAF2B585AFF82FEF9A4BE539182C9" box="[266,470,1765,1789]" italics="true" pageId="10" pageNumber="11">Spiculum gastrale</emphasis>
|
||
(
|
||
<figureCitation id="B5F56FBC585AFF82FE16A4BE507182C9" box="[485,566,1765,1789]" captionStart="FIGURES 14 – 19" captionStartId="11.[151,264,1976,1999]" captionTargetBox="[167,1348,573,1945]" captionTargetId="figure@11.[151,1436,553,1965]" captionTargetPageId="11" captionText="FIGURES 14 – 19. Diagnostic structures in P. nu t an s. 14) right proleg, 15) right antenna, 16) male genitalia in dorsal view, 17) male genitalia in lateral view, 18) spiculum gastrale, 19) female genital plates. Scale lines 1 mm." httpUri="https://zenodo.org/record/210546/files/figure.png" pageId="10" pageNumber="11">Fig. 18</figureCitation>
|
||
) narrow, basal part slightly bend, briefly shorter than lateral sclerites; associated plates large, subquadrangular, with irregular apical margin clearly sinuated, and long setae near the border.
|
||
</paragraph>
|
||
<paragraph id="2D717339585AFF83FF3AA57551D884C3" blockId="10.[151,1437,325,2005]" lastBlockId="11.[151,1436,151,500]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
|
||
<emphasis id="1FBAAF2B585AFF82FF3AA57553678372" bold="true" box="[201,288,1838,1862]" pageId="10" pageNumber="11">Female</emphasis>
|
||
(
|
||
<emphasis id="1FBAAF2B585AFF82FEC3A57553798371" box="[304,318,1838,1861]" italics="true" pageId="10" pageNumber="11">n</emphasis>
|
||
= 1). Length
|
||
<quantity id="EA36DEDC585AFF82FE2BA57650768371" box="[472,561,1837,1861]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="10" pageNumber="11" unit="mm" value="9.5">9.5 mm</quantity>
|
||
, width
|
||
<quantity id="EA36DEDC585AFF82FD70A576509B8371" box="[643,732,1837,1861]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="10" pageNumber="11" unit="mm" value="4.2">4.2 mm</quantity>
|
||
. Color bright reddish yellow on clypeus, anterior half of frons, scape, pedicel, funicle, lateral margins of pronotum, elytra, coxae, trochanter, and femora; reddish brown on protibiae, protarsi, mesotarsi, and tarsal claws of all legs; shiny black on antennal club, posterior half of frons, pronotal disc,
|
||
<taxonomicName id="EACE08BA585AFF82FF21A5C253058385" box="[210,322,1945,1969]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Corynexochida" pageId="10" pageNumber="18" phylum="Arthropoda" rank="genus">scutellum</taxonomicName>
|
||
, abdomen, apices of femora, tibiae, and metatarsi. Antennal club as long as the length of the scape, pedicel, funicle combined. Body similar to male; elytral striae deep marked, with uniform large punctures; epipleural fold swollen; protibiae and protarsi narrower than in male, protarsomeres 1–4 slightly longer than in male; metatarsomeres wider than in male. Genital plates large, elongate, subtriangular, with acute apex; irregular inner and outer borders with long setae over nearly entire surface (
|
||
<figureCitation id="B5F56FBC585BFF83FCCCA28451D584C3" box="[831,914,223,247]" captionStart="FIGURES 14 – 19" captionStartId="11.[151,264,1976,1999]" captionTargetBox="[167,1348,573,1945]" captionTargetId="figure@11.[151,1436,553,1965]" captionTargetPageId="11" captionText="FIGURES 14 – 19. Diagnostic structures in P. nu t an s. 14) right proleg, 15) right antenna, 16) male genitalia in dorsal view, 17) male genitalia in lateral view, 18) spiculum gastrale, 19) female genital plates. Scale lines 1 mm." httpUri="https://zenodo.org/record/210546/files/figure.png" pageId="11" pageNumber="12">Fig. 19</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="2D717339585BFF83FF3AA35850ED850B" blockId="11.[151,1436,151,500]" pageId="11" pageNumber="12">
|
||
<emphasis id="1FBAAF2B585BFF83FF3AA35853708528" bold="true" box="[201,311,259,284]" pageId="11" pageNumber="12">Variation</emphasis>
|
||
. The punctation on pronotum and the black color on elytral suture can be more stressed and lighter respectively. Some males are completely black.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="65D420B2585BFF83FF3AA310517B85C0" pageId="11" pageNumber="12" type="diagnosis">
|
||
<paragraph id="2D717339585BFF83FF3AA310517B85C0" blockId="11.[151,1436,151,500]" pageId="11" pageNumber="12">
|
||
<emphasis id="1FBAAF2B585BFF83FF3AA31053048550" bold="true" box="[201,323,331,356]" pageId="11" pageNumber="12">Diagnosis.</emphasis>
|
||
Clypeus subrectangular, anterior margin strongly upturned. Pronotum with elongate, large sized (approximate size 0.035 x
|
||
<quantity id="EA36DEDC585BFF83FE36A334507B85B3" box="[453,572,367,391]" metricMagnitude="-5" metricUnit="m" metricValue="7.199999999999999" pageId="11" pageNumber="12" unit="mm" value="0.072">0.072 mm</quantity>
|
||
), deep punctures. Elytral striae deeply indicated with punctures; lateral margins with wide, central notch. Protarsomeres 1–4 shortened, slightly wider than protarsomere 5 but with equal length. Protibia stout, lateral borders convergent towards apex. Parameres short and simple, with upper part concave and smooth, strongly narrowed towards apex (Table 1).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="79B123B1585BFF83FF64A5E3562E83D9" httpUri="https://zenodo.org/record/210546/files/figure.png" pageId="11" pageNumber="12" targetBox="[167,1348,573,1945]" targetPageId="11">
|
||
<paragraph id="2D717339585BFF83FF64A5E3562E83D9" blockId="11.[151,1435,1976,2029]" pageId="11" pageNumber="12">
|
||
<emphasis id="1FBAAF2B585BFF83FF64A5E3530A83FA" bold="true" box="[151,333,1976,1999]" pageId="11" pageNumber="12">FIGURES 14–19</emphasis>
|
||
. Diagnostic structures in
|
||
<emphasis id="1FBAAF2B585BFF83FDA2A5E150EC83FB" box="[593,683,1978,1999]" italics="true" pageId="11" pageNumber="12">P. nu t an s</emphasis>
|
||
.
|
||
<emphasis id="1FBAAF2B585BFF83FD48A5E3509B83FB" bold="true" box="[699,732,1976,1999]" pageId="11" pageNumber="12">14)</emphasis>
|
||
right proleg,
|
||
<emphasis id="1FBAAF2B585BFF83FC94A5E351CE83FB" bold="true" box="[871,905,1976,1999]" pageId="11" pageNumber="12">15)</emphasis>
|
||
right antenna,
|
||
<emphasis id="1FBAAF2B585BFF83FBD2A5E3560483FB" bold="true" box="[1057,1091,1976,1999]" pageId="11" pageNumber="12">16)</emphasis>
|
||
male genitalia in dorsal view,
|
||
<emphasis id="1FBAAF2B585BFF83FA89A5E357DC83FB" bold="true" box="[1402,1435,1976,1999]" pageId="11" pageNumber="12">17)</emphasis>
|
||
male genitalia in lateral view,
|
||
<emphasis id="1FBAAF2B585BFF83FE3BA58D53AE83D9" bold="true" box="[456,489,2006,2029]" pageId="11" pageNumber="12">18)</emphasis>
|
||
spiculum gastrale,
|
||
<emphasis id="1FBAAF2B585BFF83FD5DA58D509783D9" bold="true" box="[686,720,2006,2029]" pageId="11" pageNumber="12">19)</emphasis>
|
||
female genital plates. Scale lines 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="65D420B2585CFF84FF3AA2C3513884C3" pageId="12" pageNumber="13" type="etymology">
|
||
<paragraph id="2D717339585CFF84FF3AA2C3513884C3" blockId="12.[151,1436,151,500]" pageId="12" pageNumber="13">
|
||
<emphasis id="1FBAAF2B585CFF84FF3AA2C353088484" bold="true" box="[201,335,152,176]" pageId="12" pageNumber="13">Etymology.</emphasis>
|
||
Although not cited in the original description, the name is derived from the Latin term
|
||
<taxonomicName id="EACE08BA585CFF84FAA2A2C257DB8484" box="[1361,1436,153,176]" class="Insecta" family="Scarabaeidae" genus="Spilota" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="nutans">
|
||
<emphasis id="1FBAAF2B585CFF84FAA2A2C257DB8484" box="[1361,1436,153,176]" italics="true" pageId="12" pageNumber="13">nutans</emphasis>
|
||
</taxonomicName>
|
||
(nodding, unsteady) (
|
||
<bibRefCitation id="495F0EC8585CFF84FE74A2E7505484E0" author="Jaeger" box="[391,531,188,212]" pageId="12" pageNumber="13" refString="Jaeger, E. C., (1978) A Source Book of Biological Names and Terms, 3 rd edition. Thomas Publisher, Springfield, Illinois." type="book" year="1978">Jaeger 1978</bibRefCitation>
|
||
), perhaps indicating that the author was uncertain of the species. The name was assigned to specimens in the collection of Sallé (
|
||
<bibRefCitation id="495F0EC8585CFF84FD44A284513484C3" author="Bates" box="[695,883,223,247]" pageId="12" pageNumber="13" refString="Bates, H. W. (1888) Pectinicornia and Lamellicornia. In: O. Salvin & F. D. Godwin (eds.), Biologia Centrali-Americana. Insecta Coleoptera. Volume 2, part 2: 1 - 432." type="book chapter" year="1888">Bates 1888: 247</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="65D420B2585CFF84FF3AA358507F85B3" pageId="12" pageNumber="13" type="distribution">
|
||
<paragraph id="2D717339585CFF84FF3AA358507F85B3" blockId="12.[151,1436,151,500]" pageId="12" pageNumber="13">
|
||
<emphasis id="1FBAAF2B585CFF84FF3AA35850908528" bold="true" box="[201,727,259,284]" pageId="12" pageNumber="13">Distribution, phenology, and natural history</emphasis>
|
||
. This species is distributed on the slopes of Sierra de Miahuatlan, Oaxaca, between
|
||
<quantity id="EA36DEDC585CFF84FE7FA37C5060850B" box="[396,551,295,319]" metricMagnitude="3" metricUnit="m" metricValue="1.6" metricValueMax="1.9" metricValueMin="1.3" pageId="12" pageNumber="13" unit="m" value="1600.0" valueMax="1900.0" valueMin="1300.0">1300–1900 m</quantity>
|
||
in elevation (
|
||
<figureCitation id="B5F56FBC585CFF84FD33A37C5154850B" box="[704,787,295,319]" captionStart="FIGURE 54" captionStartId="21.[151,250,1234,1257]" captionTargetBox="[157,1436,382,1193]" captionTargetId="figure@21.[157,1442,373,1193]" captionTargetPageId="21" captionText="FIGURE 54. Distribution of species of Pachystethus." httpUri="https://zenodo.org/record/210552/files/figure.png" pageId="12" pageNumber="13">Fig. 54</figureCitation>
|
||
), where pine forests and cloud forests are mixed with secondary vegetation. Adults are active during May. No plants have been recorded as hosts for adults, and the immature stages remain undescribed.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="65D420B2585CFF84FF3AA3CF50C585C0" pageId="12" pageNumber="13" type="discussion">
|
||
<paragraph id="2D717339585CFF84FF3AA3CF50C585C0" blockId="12.[151,1436,151,500]" pageId="12" pageNumber="13">
|
||
<emphasis id="1FBAAF2B585CFF84FF3AA3CF537A8598" bold="true" box="[201,317,404,428]" pageId="12" pageNumber="13">Remarks.</emphasis>
|
||
The record from “Jalapa, Hoge” is in doubt. It may correspond with the well-known confusion of some samples collected by Höge in the southern Pacific slopes and wrongly labeled as “Jalapa,” that many authors interpreted as Jalapa de Enriquez, Veracruz.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |