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<document id="B59192C5922AAAE43388410E7A27AD4E" ID-DOI="10.11646/zootaxa.1887.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5133869" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1627159130724" checkinUser="felipe" docAuthor="Mendes, Humberto Fonseca &amp; Andersen, Trond" docDate="2008" docId="5C31878AFFD4FFFA6AADFC665375A630" docLanguage="en" docName="zt01887p075.pdf" docOrigin="Zootaxa 1887 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.1887.1.1" docStyle="DocumentStyle:C4791ACFF01C5C140D2C1A203DB3C9B0.4:Zootaxa.2007-2008.monograph" docStyleId="C4791ACFF01C5C140D2C1A203DB3C9B0" docStyleName="Zootaxa.2007-2008.monograph" docStyleVersion="4" docTitle="Gravatamberus guatemaltecus Mendes &amp; Andersen 2008, sp. n." docType="treatment" docVersion="5" lastPageNumber="49" masterDocId="A008FFF2FFFAFFCA6A3AFFBC5729A47F" masterDocTitle="A review of Antillocladius Saether and Litocladius Mendes, Andersen et Saether, with the description of two new Neotropical genera (Diptera, Chironomidae, Orthocladiinae)" masterLastPageNumber="75" masterPageNumber="1" pageNumber="47" updateTime="1699021915548" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="A2A1882A29BFEE6D9528E8585C732FD2" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="57F08B3B8267ACD2922DAA7EEDA035B0">
<mods:title id="4BE14607EB59E3C49FD4DC2BB3A27812">A review of Antillocladius Saether and Litocladius Mendes, Andersen et Saether, with the description of two new Neotropical genera (Diptera, Chironomidae, Orthocladiinae)</mods:title>
</mods:titleInfo>
<mods:name id="85EFCAB8D2DE5EDE926C3390DB72BE2B" type="personal">
<mods:role id="850089A7824243248BD04510D49CB6FF">
<mods:roleTerm id="205505559A94E7BE033EA251AB1F9015">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AF563BD6808994596D66495599A0D483">Mendes, Humberto Fonseca</mods:namePart>
</mods:name>
<mods:name id="43D66B799724B5B23EE36388D31D5C10" type="personal">
<mods:role id="E9830E9FC1F7D9235E92AED670A29652">
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</mods:role>
<mods:namePart id="A1344739A2426E048C5B21571397F513">Andersen, Trond</mods:namePart>
</mods:name>
<mods:typeOfResource id="437E5E547348467789C75113D41955C5">text</mods:typeOfResource>
<mods:relatedItem id="510D12115C8AFBCF7F9201402B544A88" type="host">
<mods:titleInfo id="A9E7C36DF9A179E520E7DFBE492B20F0">
<mods:title id="DFD831FB9133330866179BC1AA03A1C6">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="D97E67725D48888F2652FE0D06DCA3AC">
<mods:date id="55DA1B842F1149F5617DED55EA2AB25D">2008</mods:date>
<mods:detail id="2C753C3D1230859B8CD04D18754164A5" type="pubDate">
<mods:number id="DD60CD6C3CE3768020374FA8DF464F99">2008-09-29</mods:number>
</mods:detail>
<mods:detail id="63955FD82BA7B27DB58F86329BDA21B0" type="volume">
<mods:number id="4389F743402BDCDC462BB3ECE4FF54BB">1887</mods:number>
</mods:detail>
<mods:detail id="7EF854839CB5095B76592BCF98F011E4" type="issue">
<mods:number id="6E333F253E4FABE969BD4A80F9E55734">1</mods:number>
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<mods:start id="7C2AB7A39518605CAA51B90F72CEA7EF">1</mods:start>
<mods:end id="5ED9CE06854D805071169361013B0DC0">75</mods:end>
</mods:extent>
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<mods:location id="8488D6C8278167B5253A400273BF6C96">
<mods:url id="E899E5FADE5DC36B84B6381CA827922A">https://biotaxa.org/Zootaxa/article/view/zootaxa.1887.1.1</mods:url>
</mods:location>
<mods:classification id="2A427F469BD49BEBCE257060BB6AD9DB">journal article</mods:classification>
<mods:identifier id="2584FBD14D991C2F45CE72DF67380215" type="DOI">10.11646/zootaxa.1887.1.1</mods:identifier>
<mods:identifier id="78F5ECC4A82A50E26FF3530E8E48D7CC" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="91EFD62EAE12166B686D91EE549C85B2" type="Zenodo-Dep">5133869</mods:identifier>
</mods:mods>
<treatment id="5C31878AFFD4FFFA6AADFC665375A630" ID-DOI="http://doi.org/10.5281/zenodo.5134531" ID-Zenodo-Dep="5134531" LSID="urn:lsid:plazi:treatment:5C31878AFFD4FFFA6AADFC665375A630" httpUri="http://treatment.plazi.org/id/5C31878AFFD4FFFA6AADFC665375A630" lastPageId="48" lastPageNumber="49" pageId="46" pageNumber="47">
<subSubSection id="9C826517FFD4FFE46AADFC665579A78B" box="[151,592,986,1012]" pageId="46" pageNumber="47" type="nomenclature">
<paragraph id="D427369CFFD4FFE46AADFC665579A78B" blockId="46.[151,592,986,1047]" box="[151,592,986,1012]" pageId="46" pageNumber="47">
<heading id="8F6F81F0FFD4FFE46AADFC665579A78B" bold="true" box="[151,592,986,1012]" fontSize="11" level="1" pageId="46" pageNumber="47" reason="1">
<emphasis id="E6ECEA8EFFD4FFE46AADFC665579A78B" bold="true" box="[151,592,986,1012]" pageId="46" pageNumber="47">
<taxonomicName id="13984D1FFFD4FFE46AADFC66552EA78B" ID-CoL="3HBNB" authority="Mendes &amp; Andersen, 2008" authorityName="Mendes &amp; Andersen" authorityYear="2008" box="[151,519,986,1012]" class="Insecta" family="Chironomidae" genus="Gravatamberus" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="guatemaltecus" status="sp. nov.">
<emphasis id="E6ECEA8EFFD4FFE46AADFC66552EA78B" bold="true" box="[151,519,986,1012]" italics="true" pageId="46" pageNumber="47">Gravatamberus guatemaltecus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FDDF57F5FFD4FFE46834FC665579A78B" box="[526,592,986,1012]" pageId="46" pageNumber="47" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="9C826517FFD4FFE46AADFC415604A068" box="[151,301,1021,1047]" pageId="46" pageNumber="47" type="description">
<paragraph id="D427369CFFD4FFE46AADFC415604A068" blockId="46.[151,592,986,1047]" box="[151,301,1021,1047]" pageId="46" pageNumber="47">
(
<figureCitation id="4CA32A19FFD4FFE46AA5FC41560CA068" box="[159,293,1021,1047]" captionStart="FIGURES 53–57" captionStartId="47.[151,269,1750,1774]" captionTargetBox="[239,1374,196,1732]" captionTargetId="figure-132@47.[195,1405,177,1765]" captionTargetPageId="47" captionText="FIGURES 53–57. Gravatamberus guatemaltecus sp. n., male. 53—tentorium, stipes, and cibarial pump; 54—thorax; 55—wing; 56–anal point and tergite IX and dorsal aspect of left gonocoxite and gonostylus; 57—hypopygium with anal point and tergite IX removed, dorsal aspect to the left and ventral aspect to the right." figureDoi="http://doi.org/10.5281/zenodo.5133921" httpUri="https://zenodo.org/record/5133921/files/figure.png" pageId="46" pageNumber="47">Figs 53–57</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="9C826517FFD4FFE46AADFBF157FBA178" pageId="46" pageNumber="47" type="materials_examined">
<paragraph id="D427369CFFD4FFE46AADFBF153D9A0F0" blockId="46.[151,1436,1101,1937]" pageId="46" pageNumber="47">
<emphasis id="E6ECEA8EFFD4FFE46AADFBF1567CA018" bold="true" box="[151,341,1101,1127]" pageId="46" pageNumber="47">Type material:</emphasis>
<materialsCitation id="64F03CC1FFD4FFE46B59FBF153D9A0F0" collectingDate="1999-11-09" collectionCode="ZMBN" collectorName="A. C. Baily &amp; J. Monzon" country="Guatemala" county="Pueblo Nueve Vinas" elevation="1800" location="Bosque Pino-Eucino" municipality="Finca Maria Mundo" pageId="46" pageNumber="47" specimenCount="1" stateProvince="Santa Rosa" typeStatus="holotype">
<typeStatus id="0B23883EFFD4FFE46B59FBF156F9A018" box="[355,464,1101,1127]" pageId="46" pageNumber="47" type="holotype">Holotype</typeStatus>
male,
<emphasis id="E6ECEA8EFFD4FFE46816FBF15456A018" bold="true" box="[556,895,1101,1127]" pageId="46" pageNumber="47">
<collectingCountry id="AC8F760CFFD4FFE46816FBF155E2A019" box="[556,715,1101,1127]" name="Guatemala" pageId="46" pageNumber="47">GUATEMALA</collectingCountry>
:
<collectingRegion id="165CF87EFFD4FFE468D8FBF15453A018" box="[738,890,1101,1127]" country="Guatemala" name="Santa Rosa" pageId="46" pageNumber="47">Santa Rosa</collectingRegion>
:
</emphasis>
<collectingCounty id="3D464E10FFD4FFE469B7FBF153A2A018" box="[909,1163,1101,1127]" pageId="46" pageNumber="47">Pueblo Nueve Viñas</collectingCounty>
,
<collectingMunicipality id="3443ACE6FFD4FFE46EA7FBF152B1A018" box="[1181,1432,1101,1127]" pageId="46" pageNumber="47">Finca Maria Mundo</collectingMunicipality>
,
<location id="D1476047FFD4FFE46AADFBC956A2A0F0" LSID="urn:lsid:plazi:treatment:5C31878AFFD4FFFA6AADFC665375A630:D1476047FFD4FFE46AADFBC956A2A0F0" box="[151,395,1141,1167]" country="Guatemala" county="Pueblo Nueve Vinas" municipality="Finca Maria Mundo" name="Bosque Pino-Eucino" pageId="46" pageNumber="47" stateProvince="Santa Rosa">Bosque Pino-Eucino</location>
,
<elevation id="5FB5D1AFFFD4FFE46BADFBC95506A0F0" box="[407,559,1141,1167]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="46" pageNumber="47" unit="m" value="1800.0">
<quantity id="13609B79FFD4FFE46BADFBC956D9A0F0" box="[407,496,1141,1167]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="46" pageNumber="47" unit="m" value="1800.0">1800 m</quantity>
a.s.l.
</elevation>
,
<date id="A026105CFFD4FFE46806FBC95584A0F0" box="[572,685,1141,1167]" pageId="46" pageNumber="47" value="1999-11-09">
<collectingDate id="B062E9B4FFD4FFE46806FBC95584A0F0" box="[572,685,1141,1167]" pageId="46" pageNumber="47" value="1999-11-09">9.xi.1999</collectingDate>
</date>
,
<collectingMethod id="0DD94E8BFFD4FFE46883FBC95464A0F0" box="[697,845,1141,1167]" pageId="46" pageNumber="47">Malaise trap</collectingMethod>
,
<collectorName id="796D534AFFD4FFE46963FBC954FEA0F0" box="[857,983,1141,1167]" pageId="46" pageNumber="47">A.C. Baily</collectorName>
&amp;
<collectorName id="796D534AFFD4FFE469C7FBC95350A0F0" box="[1021,1145,1141,1167]" pageId="46" pageNumber="47">J. Monzon</collectorName>
(
<collectionCode id="B289AE59FFD4FFE46EB3FBC953CDA0F0" box="[1161,1252,1141,1167]" country="Norway" httpUri="http://grbio.org/cool/0tqf-efed" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="46" pageNumber="47">ZMBN</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="D427369CFFD4FFE46AFFFB2157FBA178" blockId="46.[151,1436,1101,1937]" pageId="46" pageNumber="47">
<emphasis id="E6ECEA8EFFD4FFE46AFFFB2156C8A0C8" bold="true" box="[197,481,1181,1207]" pageId="46" pageNumber="47">Diagnostic characters:</emphasis>
The species can be separated from all other members of the genus by having more than 10 setae in cell m proximal to RM, costal extension ending close to wing tip, and AR lower than 0.50.
</paragraph>
</subSubSection>
<subSubSection id="9C826517FFD4FFFA6AFFFAA95691A658" lastPageId="48" lastPageNumber="49" pageId="46" pageNumber="47" type="etymology">
<paragraph id="D427369CFFD4FFE46AFFFAA95630A128" blockId="46.[151,1436,1101,1937]" pageId="46" pageNumber="47">
<emphasis id="E6ECEA8EFFD4FFE46AFFFAA9567FA150" bold="true" box="[197,342,1301,1327]" pageId="46" pageNumber="47">Etymology:</emphasis>
From Spanish
<emphasis id="E6ECEA8EFFD4FFE46830FAA95581A151" box="[522,680,1301,1326]" italics="true" pageId="46" pageNumber="47">guatemalteco</emphasis>
, a native of
<collectingCountry id="AC8F760CFFD4FFE46907FAA954E9A150" box="[829,960,1301,1327]" name="Guatemala" pageId="46" pageNumber="47">Guatemala</collectingCountry>
; the name is to be regarded as a noun in apposition.
</paragraph>
<paragraph id="D427369CFFD4FFE46AFFFAD95467A1B0" blockId="46.[151,1436,1101,1937]" pageId="46" pageNumber="47">
<emphasis id="E6ECEA8EFFD4FFE46AFFFAD9562DA100" bold="true" box="[197,260,1381,1407]" pageId="46" pageNumber="47">Male</emphasis>
(n = 1). Total length 1.45 mm. Wing length 1.23 mm. Total length / wing length 1.17. Wing length / length of profemur 2.53. Coloration brown; thorax dark brown with markings on preepisternum, median anepisternum and notum; legs and tarsi uniformly light brown.
</paragraph>
<paragraph id="D427369CFFD4FFE46AFFFA6156ABA2E8" blockId="46.[151,1436,1101,1937]" pageId="46" pageNumber="47">
<emphasis id="E6ECEA8EFFD4FFE46AFFFA615625A189" box="[197,268,1501,1526]" italics="true" pageId="46" pageNumber="47">Head.</emphasis>
AR 0.26. Ultimate flagellomere
<quantity id="13609B79FFD4FFE468ACFA6155DBA188" box="[662,754,1501,1527]" metricMagnitude="-1" metricUnit="m" metricValue="1.1199999999999999" pageId="46" pageNumber="47" unit="mm" value="112.0">112 µm</quantity>
long, stout subapical seta
<quantity id="13609B79FFD4FFE46E15FA615354A188" box="[1071,1149,1501,1527]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="46" pageNumber="47" unit="mm" value="40.0">40 µm</quantity>
long. Temporal setae 9, including 4 inner verticals, 3 outer verticals, and 2 postorbitals. Clypeus with 13 setae. Tentorium, stipes, and cibarial pump as in
<figureCitation id="4CA32A19FFD4FFE46BBBF99156DCA238" box="[385,501,1581,1607]" captionStart="FIGURES 53–57" captionStartId="47.[151,269,1750,1774]" captionTargetBox="[239,1374,196,1732]" captionTargetId="figure-132@47.[195,1405,177,1765]" captionTargetPageId="47" captionText="FIGURES 53–57. Gravatamberus guatemaltecus sp. n., male. 53—tentorium, stipes, and cibarial pump; 54—thorax; 55—wing; 56–anal point and tergite IX and dorsal aspect of left gonocoxite and gonostylus; 57—hypopygium with anal point and tergite IX removed, dorsal aspect to the left and ventral aspect to the right." figureDoi="http://doi.org/10.5281/zenodo.5133921" httpUri="https://zenodo.org/record/5133921/files/figure.png" pageId="46" pageNumber="47">Figure 53</figureCitation>
. Tentorium
<quantity id="13609B79FFD4FFE468BEF99155F8A238" box="[644,721,1581,1607]" metricMagnitude="-2" metricUnit="m" metricValue="9.1" pageId="46" pageNumber="47" unit="mm" value="91.0">91 µm</quantity>
long,
<quantity id="13609B79FFD4FFE46926F9915443A238" box="[796,874,1581,1607]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="46" pageNumber="47" unit="mm" value="18.0">18 µm</quantity>
wide. Stipes
<quantity id="13609B79FFD4FFE46E33F991537FA238" box="[1033,1110,1581,1607]" metricMagnitude="-2" metricUnit="m" metricValue="9.5" pageId="46" pageNumber="47" unit="mm" value="95.0">95 µm</quantity>
long. Palp segment lengths (in µm): 18, 27, 82, 95, fifth palpomere not measurable. Third palpomere with 2 sensilla clavata subapically, longest
<quantity id="13609B79FFD4FFE46AC9F9C15616A2E8" box="[243,319,1661,1687]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="46" pageNumber="47" unit="mm" value="14.0">14 µm</quantity>
long.
</paragraph>
<paragraph id="D427369CFFD4FFE46AFFF9195411A298" blockId="46.[151,1436,1101,1937]" pageId="46" pageNumber="47">
<emphasis id="E6ECEA8EFFD4FFE46AFFF9195633A2C1" box="[197,282,1701,1726]" italics="true" pageId="46" pageNumber="47">Thorax</emphasis>
(
<figureCitation id="4CA32A19FFD4FFE46B10F91956ABA2C0" box="[298,386,1701,1727]" captionStart="FIGURES 53–57" captionStartId="47.[151,269,1750,1774]" captionTargetBox="[239,1374,196,1732]" captionTargetId="figure-132@47.[195,1405,177,1765]" captionTargetPageId="47" captionText="FIGURES 53–57. Gravatamberus guatemaltecus sp. n., male. 53—tentorium, stipes, and cibarial pump; 54—thorax; 55—wing; 56–anal point and tergite IX and dorsal aspect of left gonocoxite and gonostylus; 57—hypopygium with anal point and tergite IX removed, dorsal aspect to the left and ventral aspect to the right." figureDoi="http://doi.org/10.5281/zenodo.5133921" httpUri="https://zenodo.org/record/5133921/files/figure.png" pageId="46" pageNumber="47">Fig. 54</figureCitation>
). Antepronotum with 5 setae. Dorsocentrals 18; acrostichals 6, all scalpellate; prealars 6, extended anteriorly; no supraalar. Scutellum with 2 setae.
</paragraph>
<paragraph id="D427369CFFD4FFE46AFFF9495304A3F2" blockId="46.[151,1436,1101,1937]" pageId="46" pageNumber="47">
<emphasis id="E6ECEA8EFFD4FFE46AFFF9495628A371" box="[197,257,1781,1806]" italics="true" pageId="46" pageNumber="47">Wing</emphasis>
(
<figureCitation id="4CA32A19FFD4FFE46B28F9495643A370" box="[274,362,1781,1807]" captionStart="FIGURES 53–57" captionStartId="47.[151,269,1750,1774]" captionTargetBox="[239,1374,196,1732]" captionTargetId="figure-132@47.[195,1405,177,1765]" captionTargetPageId="47" captionText="FIGURES 53–57. Gravatamberus guatemaltecus sp. n., male. 53—tentorium, stipes, and cibarial pump; 54—thorax; 55—wing; 56–anal point and tergite IX and dorsal aspect of left gonocoxite and gonostylus; 57—hypopygium with anal point and tergite IX removed, dorsal aspect to the left and ventral aspect to the right." figureDoi="http://doi.org/10.5281/zenodo.5133921" httpUri="https://zenodo.org/record/5133921/files/figure.png" pageId="46" pageNumber="47">Fig. 55</figureCitation>
). VR 1.33. C extension
<quantity id="13609B79FFD4FFE468BFF94955C8A370" box="[645,737,1781,1807]" metricMagnitude="-1" metricUnit="m" metricValue="1.61" pageId="46" pageNumber="47" unit="mm" value="161.0">161 µm</quantity>
long. Brachiolum with 1 seta, Sc with 16 seta, C extension with 23 non-marginal setae, R with 15 setae, R
<subScript id="481C34D9FFD4FFE46883F89155E8A344" attach="left" box="[697,705,1837,1851]" fontSize="6" pageId="46" pageNumber="47">1</subScript>
with 11 setae, R with 20 setae, M
<subScript id="481C34D9FFD4FFE46E5FF8915357A344" attach="left" box="[1125,1150,1837,1851]" fontSize="6" pageId="46" pageNumber="47">1+2</subScript>
with 45 setae, M
<subScript id="481C34D9FFD4FFE46F72F8915249A344" attach="left" box="[1352,1376,1837,1851]" fontSize="6" pageId="46" pageNumber="47">3+4</subScript>
with 24
<taxonomicName id="13984D1FFFD4FFE46A86F8F45601A31D" authority=", Cu" authorityName="Cu" box="[188,296,1864,1890]" class="Insecta" family="Chironomidae" genus="Lopescladius" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="setae">setae, Cu</taxonomicName>
with 17
<taxonomicName id="13984D1FFFD4FFE46BA8F8F456D7A31D" authority=", Cu" authorityName="Cu" box="[402,510,1864,1890]" class="Insecta" family="Chironomidae" genus="Lopescladius" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="setae">setae, Cu</taxonomicName>
<subScript id="481C34D9FFD4FFE46BC4F8E4552FA319" attach="left" box="[510,518,1880,1894]" fontSize="6" pageId="46" pageNumber="47">1</subScript>
with 14 setae, PCu with 36
<taxonomicName id="13984D1FFFD4FFE4696FF8F454EBA31D" authority=", An" authorityName="An" box="[853,962,1864,1890]" class="Insecta" family="Chironomidae" genus="Lopescladius" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="setae">setae, An</taxonomicName>
with 20 setae. Cell m with 19 setae, r with 224 setae, m
<subScript id="481C34D9FFD4FFE46B5FF83F5656A3EE" attach="left" box="[357,383,1923,1937]" fontSize="6" pageId="46" pageNumber="47">1+2</subScript>
with 223 setae, m
<subScript id="481C34D9FFD4FFE4686FF83F5544A3EE" attach="left" box="[597,621,1923,1937]" fontSize="6" pageId="46" pageNumber="47">3+4</subScript>
with 96
<taxonomicName id="13984D1FFFD4FFE468EFF8CF5413A3F2" authority=", cu" authorityName="Cu" box="[725,826,1907,1933]" class="Insecta" family="Chironomidae" genus="Lopescladius" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="setae">setae, cu</taxonomicName>
+ an with 227 setae.
</paragraph>
<paragraph id="D427369CFFD4FFE469B9F89154B4A344" blockId="46.[151,1436,1101,1937]" box="[899,925,1837,1851]" pageId="46" pageNumber="47">
<subScript id="481C34D9FFD4FFE469B9F89154B4A344" attach="left" box="[899,925,1837,1851]" fontSize="6" pageId="46" pageNumber="47">4+5</subScript>
</paragraph>
<paragraph id="D427369CFFD4FFE46FB9F8E452B5A319" blockId="46.[151,1436,1101,1937]" box="[1411,1436,1880,1894]" pageId="46" pageNumber="47">
<subScript id="481C34D9FFD4FFE46FB9F8E452B5A319" attach="left" box="[1411,1436,1880,1894]" fontSize="6" pageId="46" pageNumber="47">4+5</subScript>
</paragraph>
<caption id="80E76614FFD5FFE56AADF96A5333A34C" ID-DOI="http://doi.org/10.5281/zenodo.5133921" ID-Zenodo-Dep="5133921" httpUri="https://zenodo.org/record/5133921/files/figure.png" pageId="47" pageNumber="48" startId="47.[151,269,1750,1774]" targetBox="[239,1374,196,1732]" targetPageId="47">
<paragraph id="D427369CFFD5FFE56AADF96A5333A34C" blockId="47.[151,1436,1750,1843]" pageId="47" pageNumber="48">
<emphasis id="E6ECEA8EFFD5FFE56AADF96A5676A291" bold="true" box="[151,351,1750,1774]" pageId="47" pageNumber="48">FIGURES 53–57.</emphasis>
<taxonomicName id="13984D1FFFD5FFE56B52F96B559BA291" authority="Mendes &amp; Andersen, 2008" authorityName="Mendes &amp; Andersen" authorityYear="2008" box="[360,690,1751,1774]" class="Insecta" family="Chironomidae" genus="Gravatamberus" kingdom="Animalia" order="Diptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="guatemaltecus" status="sp. nov.">
<emphasis id="E6ECEA8EFFD5FFE56B52F96B559BA291" box="[360,690,1751,1774]" italics="true" pageId="47" pageNumber="48">Gravatamberus guatemaltecus</emphasis>
</taxonomicName>
<emphasis id="E6ECEA8EFFD5FFE56881F96A55D0A291" bold="true" box="[699,761,1750,1774]" pageId="47" pageNumber="48">
<taxonomicNameLabel id="FDDF57F5FFD5FFE56881F96A55D0A291" box="[699,761,1750,1774]" pageId="47" pageNumber="48" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, male.
<emphasis id="E6ECEA8EFFD5FFE56976F96A54A8A291" bold="true" box="[844,897,1750,1774]" pageId="47" pageNumber="48">53—</emphasis>
tentorium, stipes, and cibarial pump;
<emphasis id="E6ECEA8EFFD5FFE56F21F96A5279A291" bold="true" box="[1307,1360,1750,1774]" pageId="47" pageNumber="48">54—</emphasis>
thorax;
<emphasis id="E6ECEA8EFFD5FFE56AADF94557E5A36E" bold="true" box="[151,204,1785,1809]" pageId="47" pageNumber="48">55—</emphasis>
wing;
<emphasis id="E6ECEA8EFFD5FFE56B2AF9455611A36E" bold="true" box="[272,312,1785,1809]" pageId="47" pageNumber="48">56–</emphasis>
anal point and tergite IX and dorsal aspect of left gonocoxite and gonostylus;
<emphasis id="E6ECEA8EFFD5FFE56E4CF9455382A36E" bold="true" box="[1142,1195,1785,1809]" pageId="47" pageNumber="48">57—</emphasis>
hypopygium with anal point and tergite IX removed, dorsal aspect to the left and ventral aspect to the right.
</paragraph>
</caption>
<paragraph id="D427369CFFD5FFE56AFFF8DB539FA3AE" blockId="47.[151,1436,1895,2001]" pageId="47" pageNumber="48">
<emphasis id="E6ECEA8EFFD5FFE56AFFF8DB562DA3FF" box="[197,260,1895,1920]" italics="true" pageId="47" pageNumber="48">Legs.</emphasis>
Spur of foretibia
<quantity id="13609B79FFD5FFE56BE0F8DB5501A3FE" box="[474,552,1895,1921]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" pageId="47" pageNumber="48" unit="mm" value="29.0">29 µm</quantity>
long, spurs of midtibia
<quantity id="13609B79FFD5FFE5697DF8DB54BFA3FE" box="[839,918,1895,1921]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="47" pageNumber="48" unit="mm" value="32.0">32 µm</quantity>
and
<quantity id="13609B79FFD5FFE569EBF8DB5336A3FE" box="[977,1055,1895,1921]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="47" pageNumber="48" unit="mm" value="16.0">16 µm</quantity>
long, spurs of hind tibia
<quantity id="13609B79FFD5FFE56F74F8DB52B5A3FE" box="[1358,1436,1895,1921]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" pageId="47" pageNumber="48" unit="mm" value="39.0">39 µm</quantity>
and
<quantity id="13609B79FFD5FFE56AF1F8335630A3D6" box="[203,281,1935,1961]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="47" pageNumber="48" unit="mm" value="18.0">18 µm</quantity>
long. Width at apex of foretibia
<quantity id="13609B79FFD5FFE5689DF83355D1A3D6" box="[679,760,1935,1961]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" pageId="47" pageNumber="48" unit="mm" value="23.0">23 µm</quantity>
, of midtibia
<quantity id="13609B79FFD5FFE569AEF83354CCA3D6" box="[916,997,1935,1961]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="47" pageNumber="48" unit="mm" value="25.0">25 µm</quantity>
, of hind tibia
<quantity id="13609B79FFD5FFE56EAAF83353CBA3D6" box="[1168,1250,1935,1961]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="47" pageNumber="48" unit="mm" value="32.0">32 µm</quantity>
. Comb with 10 setae, longest
<quantity id="13609B79FFD5FFE56B06F80B56A3A3AE" box="[316,394,1975,2001]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="47" pageNumber="48" unit="mm" value="36.0">36 µm</quantity>
, shortest
<quantity id="13609B79FFD5FFE56BC3F80B556FA3AE" box="[505,582,1975,2001]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="47" pageNumber="48" unit="mm" value="16.0">16 µm</quantity>
long. Lengths and proportions of legs as in
<tableCitation id="991A0327FFD5FFE56E73F80B539BA3AE" box="[1097,1202,1975,2001]" captionStart="TABLE 10" captionStartId="48.[151,239,151,175]" captionText="TABLE 10. Lengths (in µm) and proportions of legs of Gravatamberus guatemaltecus sp. n., male (n = 1)." pageId="47" pageNumber="48">Table 10</tableCitation>
.
</paragraph>
<caption id="80E76614FFCAFFFA6AADFF2B5227A4D0" box="[151,1294,151,175]" pageId="48" pageNumber="49" startId="48.[151,239,151,175]">
<paragraph id="D427369CFFCAFFFA6AADFF2B5227A4D0" blockId="48.[151,1294,151,175]" box="[151,1294,151,175]" pageId="48" pageNumber="49">
<emphasis id="E6ECEA8EFFCAFFFA6AADFF2B563FA4D0" bold="true" box="[151,278,151,175]" pageId="48" pageNumber="49">TABLE 10.</emphasis>
Lengths (in µm) and proportions of legs of
<taxonomicName id="13984D1FFFCAFFFA68D6FF24531DA4D0" authority="Mendes &amp; Andersen, 2008" authorityName="Mendes &amp; Andersen" authorityYear="2008" box="[748,1076,152,175]" class="Insecta" family="Chironomidae" genus="Gravatamberus" kingdom="Animalia" order="Diptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="guatemaltecus" status="sp. nov.">
<emphasis id="E6ECEA8EFFCAFFFA68D6FF24531DA4D0" box="[748,1076,152,175]" italics="true" pageId="48" pageNumber="49">Gravatamberus guatemaltecus</emphasis>
</taxonomicName>
<emphasis id="E6ECEA8EFFCAFFFA6E00FF2B535FA4D0" bold="true" box="[1082,1142,151,175]" pageId="48" pageNumber="49">
<taxonomicNameLabel id="FDDF57F5FFCAFFFA6E00FF2B535FA4D0" box="[1082,1142,151,175]" pageId="48" pageNumber="49" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, male (n = 1).
</paragraph>
</caption>
<paragraph id="D427369CFFCAFFFA6AD5FF6C524CA510" pageId="48" pageNumber="49">
<table id="A698C43CFFCA00356AA5FF6C5241A50C" box="[159,1384,208,371]" colsContinueFrom="25.[159,1382,208,371]" colsContinueIn="58.[159,1386,208,371]" gridcols="12" gridrows="4" pageId="48" pageNumber="49">
<tr id="6AA834DEFFCA00356AA5FF6C5241A494" box="[159,1384,208,235]" gridrow="0" pageId="48" pageNumber="49" rowspan-0="1">
<th id="29795DA2FFCA00356AD5FF6C563EA494" box="[239,279,208,235]" gridcol="1" gridrow="0" pageId="48" pageNumber="49">fe</th>
<th id="29795DA2FFCA00356B64FF6C56AFA494" box="[350,390,208,235]" gridcol="2" gridrow="0" pageId="48" pageNumber="49">ti</th>
<th id="29795DA2FFCA00356BF7FF6C56DCA494" box="[461,501,208,235]" gridcol="3" gridrow="0" pageId="48" pageNumber="49">ta1</th>
<th id="29795DA2FFCA00356801FF6C554AA494" box="[571,611,208,235]" gridcol="4" gridrow="0" pageId="48" pageNumber="49">ta2</th>
<th id="29795DA2FFCA00356890FF6C55FBA494" box="[682,722,208,235]" gridcol="5" gridrow="0" pageId="48" pageNumber="49">ta3</th>
<th id="29795DA2FFCA00356923FF6C541CA494" box="[793,821,208,235]" gridcol="6" gridrow="0" pageId="48" pageNumber="49">ta4</th>
<th id="29795DA2FFCA003569B2FF6C548AA494" box="[904,931,208,235]" gridcol="7" gridrow="0" pageId="48" pageNumber="49">ta5</th>
<th id="29795DA2FFCA003569CDFF6C530FA494" box="[1015,1062,208,235]" gridcol="8" gridrow="0" pageId="48" pageNumber="49">LR</th>
<th id="29795DA2FFCA00356E5CFF6C53BCA494" box="[1126,1173,208,235]" gridcol="9" gridrow="0" pageId="48" pageNumber="49">BV</th>
<th id="29795DA2FFCA00356EEFFF6C522DA494" box="[1237,1284,208,235]" gridcol="10" gridrow="0" pageId="48" pageNumber="49">SV</th>
<th id="29795DA2FFCA00356F7EFF6C5241A494" box="[1348,1384,208,235]" gridcol="11" gridrow="0" pageId="48" pageNumber="49">BR</th>
</tr>
<tr id="6AA834DEFFCA00356AA5FF415241A566" box="[159,1384,253,281]" gridrow="1" pageId="48" pageNumber="49">
<th id="29795DA2FFCA00356AA5FF41579AA566" box="[159,179,253,281]" gridcol="0" gridrow="1" pageId="48" pageNumber="49">p1</th>
<td id="29795DA2FFCA00356AD5FF41563EA566" box="[239,279,253,281]" gridcol="1" gridrow="1" pageId="48" pageNumber="49">475</td>
<td id="29795DA2FFCA00356B64FF4156AFA566" box="[350,390,253,281]" gridcol="2" gridrow="1" pageId="48" pageNumber="49">522</td>
<td id="29795DA2FFCA00356BF7FF4156DCA566" box="[461,501,253,281]" gridcol="3" gridrow="1" pageId="48" pageNumber="49">353</td>
<td id="29795DA2FFCA00356801FF41554AA566" box="[571,611,253,281]" gridcol="4" gridrow="1" pageId="48" pageNumber="49">184</td>
<td id="29795DA2FFCA00356890FF4155FBA566" box="[682,722,253,281]" gridcol="5" gridrow="1" pageId="48" pageNumber="49">140</td>
<td id="29795DA2FFCA00356923FF41541CA566" box="[793,821,253,281]" gridcol="6" gridrow="1" pageId="48" pageNumber="49">90</td>
<td id="29795DA2FFCA003569B2FF41548AA566" box="[904,931,253,281]" gridcol="7" gridrow="1" pageId="48" pageNumber="49">43</td>
<td id="29795DA2FFCA003569CDFF41530FA566" box="[1015,1062,253,281]" gridcol="8" gridrow="1" pageId="48" pageNumber="49">0.68</td>
<td id="29795DA2FFCA00356E5CFF4153BCA566" box="[1126,1173,253,281]" gridcol="9" gridrow="1" pageId="48" pageNumber="49">2.95</td>
<td id="29795DA2FFCA00356EEFFF41522DA566" box="[1237,1284,253,281]" gridcol="10" gridrow="1" pageId="48" pageNumber="49">2.83</td>
<td id="29795DA2FFCA00356F7EFF415241A566" box="[1348,1384,253,281]" gridcol="11" gridrow="1" pageId="48" pageNumber="49">3.7</td>
</tr>
<tr id="6AA834DEFFCA00356AA5FE965241A539" box="[159,1384,298,326]" gridrow="2" pageId="48" pageNumber="49">
<th id="29795DA2FFCA00356AA5FE96579AA539" box="[159,179,298,326]" gridcol="0" gridrow="2" pageId="48" pageNumber="49">p2</th>
<td id="29795DA2FFCA00356AD5FE96563EA539" box="[239,279,298,326]" gridcol="1" gridrow="2" pageId="48" pageNumber="49">443</td>
<td id="29795DA2FFCA00356B64FE9656AFA539" box="[350,390,298,326]" gridcol="2" gridrow="2" pageId="48" pageNumber="49">443</td>
<td id="29795DA2FFCA00356BF7FE9656DCA539" box="[461,501,298,326]" gridcol="3" gridrow="2" pageId="48" pageNumber="49">216</td>
<td id="29795DA2FFCA00356801FE96554AA539" box="[571,611,298,326]" gridcol="4" gridrow="2" pageId="48" pageNumber="49">101</td>
<td id="29795DA2FFCA00356890FE9655FBA539" box="[682,722,298,326]" gridcol="5" gridrow="2" pageId="48" pageNumber="49">72</td>
<td id="29795DA2FFCA00356923FE96541CA539" box="[793,821,298,326]" gridcol="6" gridrow="2" pageId="48" pageNumber="49">47</td>
<td id="29795DA2FFCA003569B2FE96548AA539" box="[904,931,298,326]" gridcol="7" gridrow="2" pageId="48" pageNumber="49">32</td>
<td id="29795DA2FFCA003569CDFE96530FA539" box="[1015,1062,298,326]" gridcol="8" gridrow="2" pageId="48" pageNumber="49">0.49</td>
<td id="29795DA2FFCA00356E5CFE9653BCA539" box="[1126,1173,298,326]" gridcol="9" gridrow="2" pageId="48" pageNumber="49">4.37</td>
<td id="29795DA2FFCA00356EEFFE96522DA539" box="[1237,1284,298,326]" gridcol="10" gridrow="2" pageId="48" pageNumber="49">4.10</td>
<td id="29795DA2FFCA00356F7EFE965241A539" box="[1348,1384,298,326]" gridcol="11" gridrow="2" pageId="48" pageNumber="49">4.3</td>
</tr>
<tr id="6AA834DEFFCA00356AA5FEE45241A50C" box="[159,1384,344,371]" gridrow="3" pageId="48" pageNumber="49">
<th id="29795DA2FFCA00356AA5FEE4579AA50C" box="[159,179,344,371]" gridcol="0" gridrow="3" pageId="48" pageNumber="49">p3</th>
<td id="29795DA2FFCA00356AD5FEE4563EA50C" box="[239,279,344,371]" gridcol="1" gridrow="3" pageId="48" pageNumber="49">468</td>
<td id="29795DA2FFCA00356B64FEE456AFA50C" box="[350,390,344,371]" gridcol="2" gridrow="3" pageId="48" pageNumber="49">511</td>
<td id="29795DA2FFCA00356BF7FEE456DCA50C" box="[461,501,344,371]" gridcol="3" gridrow="3" pageId="48" pageNumber="49">277</td>
<td id="29795DA2FFCA00356801FEE4554AA50C" box="[571,611,344,371]" gridcol="4" gridrow="3" pageId="48" pageNumber="49">137</td>
<td id="29795DA2FFCA00356890FEE455FBA50C" box="[682,722,344,371]" gridcol="5" gridrow="3" pageId="48" pageNumber="49">126</td>
<td id="29795DA2FFCA00356923FEE4541CA50C" box="[793,821,344,371]" gridcol="6" gridrow="3" pageId="48" pageNumber="49">54</td>
<td id="29795DA2FFCA003569B2FEE4548AA50C" box="[904,931,344,371]" gridcol="7" gridrow="3" pageId="48" pageNumber="49">32</td>
<td id="29795DA2FFCA003569CDFEE4530FA50C" box="[1015,1062,344,371]" gridcol="8" gridrow="3" pageId="48" pageNumber="49">0.54</td>
<td id="29795DA2FFCA00356E5CFEE453BCA50C" box="[1126,1173,344,371]" gridcol="9" gridrow="3" pageId="48" pageNumber="49">3.60</td>
<td id="29795DA2FFCA00356EEFFEE4522DA50C" box="[1237,1284,344,371]" gridcol="10" gridrow="3" pageId="48" pageNumber="49">3.53</td>
<td id="29795DA2FFCA00356F7EFEE45241A50C" box="[1348,1384,344,371]" gridcol="11" gridrow="3" pageId="48" pageNumber="49">6.6</td>
</tr>
</table>
</paragraph>
<paragraph id="D427369CFFCAFFFA6AFFFE015691A658" blockId="48.[151,1436,445,591]" pageId="48" pageNumber="49">
<emphasis id="E6ECEA8EFFCAFFFA6AFFFE015672A5A9" box="[197,347,445,470]" italics="true" pageId="48" pageNumber="49">Hypopygium</emphasis>
(
<figureCitation id="4CA32A19FFCAFFFA6B56FE0156DEA5A8" box="[364,503,445,471]" captionStart="FIGURES 53–57" captionStartId="47.[151,269,1750,1774]" captionTargetBox="[239,1374,196,1732]" captionTargetId="figure-132@47.[195,1405,177,1765]" captionTargetPageId="47" captionText="FIGURES 53–57. Gravatamberus guatemaltecus sp. n., male. 53—tentorium, stipes, and cibarial pump; 54—thorax; 55—wing; 56–anal point and tergite IX and dorsal aspect of left gonocoxite and gonostylus; 57—hypopygium with anal point and tergite IX removed, dorsal aspect to the left and ventral aspect to the right." figureDoi="http://doi.org/10.5281/zenodo.5133921" httpUri="https://zenodo.org/record/5133921/files/figure.png" pageId="48" pageNumber="49">Figs 56–57</figureCitation>
). Tergite IX with 2 setae, laterosternite IX with 5 setae. Phallapodeme
<quantity id="13609B79FFCAFFFA6F77FE0152B5A5A8" box="[1357,1436,445,471]" metricMagnitude="-2" metricUnit="m" metricValue="5.7" pageId="48" pageNumber="49" unit="mm" value="57.0">57 µm</quantity>
long, transverse sternapodeme
<quantity id="13609B79FFCAFFFA683BFE595564A580" box="[513,589,485,511]" metricMagnitude="-2" metricUnit="m" metricValue="5.7" pageId="48" pageNumber="49" unit="mm" value="57.0">57 µm</quantity>
long. Gonocoxite
<quantity id="13609B79FFCAFFFA691CFE595456A580" box="[806,895,485,511]" metricMagnitude="-1" metricUnit="m" metricValue="1.2" pageId="48" pageNumber="49" unit="mm" value="120.0">120 µm</quantity>
long. Gonostylus
<quantity id="13609B79FFCAFFFA6E6CFE59538BA580" box="[1110,1186,485,511]" metricMagnitude="-2" metricUnit="m" metricValue="5.7" pageId="48" pageNumber="49" unit="mm" value="57.0">57 µm</quantity>
long, megaseta
<quantity id="13609B79FFCAFFFA6F65FE5952B5A580" box="[1375,1436,485,511]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="48" pageNumber="49" unit="mm" value="7.0">7 µm</quantity>
long. HR 2.12, HV 2.55.
</paragraph>
</subSubSection>
<subSubSection id="9C826517FFCAFFFA6AFFFD895375A630" box="[197,1116,565,591]" pageId="48" pageNumber="49" type="distribution">
<paragraph id="D427369CFFCAFFFA6AFFFD895375A630" blockId="48.[151,1436,445,591]" box="[197,1116,565,591]" pageId="48" pageNumber="49">
<emphasis id="E6ECEA8EFFCAFFFA6AFFFD89564EA630" bold="true" box="[197,359,565,591]" pageId="48" pageNumber="49">Distribution:</emphasis>
This species is known only from the
<typeStatus id="0B23883EFFCAFFFA6925FD895478A630" box="[799,849,565,591]" pageId="48" pageNumber="49">type</typeStatus>
locality in
<collectingCountry id="AC8F760CFFCAFFFA69EFFD895371A630" box="[981,1112,565,591]" name="Guatemala" pageId="48" pageNumber="49">Guatemala</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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