366 lines
47 KiB
XML
366 lines
47 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.938.51234" ID-GBIF-Dataset="0223fbe1-35f8-4987-b029-29b62cdce2e6" ID-PMC="PMC7289900" ID-Pensoft-Pub="1313-2970-938-137" ID-Pensoft-UUID="258F1856E4025778B55F9D603A15DD8C" ID-PubMed="32550789" ID-ZooBank="C57570A14A484EA3889E0646632DCA9B" ModsDocID="1313-2970-938-137" checkinTime="1591376751164" checkinUser="pensoft" docAuthor="Likhitrakarn, Natdanai, Golovatch, Sergei I., Thach, Phanara, Chhuoy, Samol, Ngor, Peng Bun, Srisonchai, Ruttapon, Sutcharit, Chirasak & Panha, Somsak" docDate="2020" docId="5C16212414185C3B96ED0B45CC259ED9" docLanguage="en" docName="ZooKeys 938: 137-151" docOrigin="ZooKeys 938" docSource="http://dx.doi.org/10.3897/zookeys.938.51234" docTitle="Plusioglyphiulus khmer Likhitrakarn & Golovatch & Thach & Chhuoy & Ngor & Srisonchai & Sutcharit & Panha 2020, sp. nov." docType="treatment" docUuid="7721A428-FA76-4226-9E3E-42CF2100475B" docUuidSource="ZooBank" docVersion="4" id="258F1856E4025778B55F9D603A15DD8C" lastPageNumber="137" masterDocId="258F1856E4025778B55F9D603A15DD8C" masterDocTitle="Two new species of the millipede genus Plusioglyphiulus Silvestri, 1923 from Cambodia (Diplopoda, Spirostreptida)" masterLastPageNumber="151" masterPageNumber="137" pageNumber="137" updateTime="1668168908888" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Two new species of the millipede genus Plusioglyphiulus Silvestri, 1923 from Cambodia (Diplopoda, Spirostreptida)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Likhitrakarn, Natdanai</mods:namePart>
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<mods:affiliation>Division of Plant Protection, Faculty of Agricultural Production, Maejo University, Chiang Mai 50290, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Golovatch, Sergei I.</mods:namePart>
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<mods:affiliation>Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071, Russia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Thach, Phanara</mods:namePart>
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<mods:affiliation>Inland Fisheries Research and Development Institute (IFReDI) No. 86, Norodom Blvd., PO Box 582, Phnom Penh, Cambodia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Chhuoy, Samol</mods:namePart>
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<mods:affiliation>Inland Fisheries Research and Development Institute (IFReDI) No. 86, Norodom Blvd., PO Box 582, Phnom Penh, Cambodia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ngor, Peng Bun</mods:namePart>
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<mods:affiliation>Inland Fisheries Research and Development Institute (IFReDI) No. 86, Norodom Blvd., PO Box 582, Phnom Penh, Cambodia</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-3659-6577</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Srisonchai, Ruttapon</mods:namePart>
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<mods:affiliation>Department of Biology, Faculty of Science, Khon Kaen University, Khon Kaen, 40002, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Panha, Somsak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>938</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>137</mods:start>
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<mods:end>151</mods:end>
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</mods:part>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.938.51234</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.938.51234</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-938-137</mods:identifier>
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<treatment ID-GBIF-Taxon="164479804" LSID="urn:lsid:zoobank.org:act:7721A428-FA76-4226-9E3E-42CF2100475B" httpUri="http://treatment.plazi.org/id/5C16212414185C3B96ED0B45CC259ED9" lastPageNumber="137" pageId="0" pageNumber="137">
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<subSubSection pageId="0" pageNumber="137" type="nomenclature">
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<paragraph pageId="0" pageNumber="137">
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<taxonomicName LSID="5C162124-1418-5C3B-96ED-0B45CC259ED9" authority="Likhitrakarn & Golovatch & Thach & Chhuoy & Ngor & Srisonchai & Sutcharit & Panha, 2020" authorityName="Likhitrakarn & Golovatch & Thach & Chhuoy & Ngor & Srisonchai & Sutcharit & Panha" authorityYear="2020" class="Diplopoda" family="Cambalopsidae" genus="Plusioglyphiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plusioglyphiulus khmer" order="Spirostreptida" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="khmer" status="sp. nov.">Plusioglyphiulus khmer</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="137">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus, live coloration A Plusioglyphiulus boutini Mauries, 1970, ♂ from Prasat Phnom Totong Temple B Plusioglyphiulus biserratus sp. nov., ♀ paratype C Plusioglyphiulus khmer sp. nov., ♂ paratype. All pictures by R. Srisonchai, not taken to scale." figureDoi="10.3897/zookeys.938.51234.figure1" httpUri="https://binary.pensoft.net/fig/418283" pageId="0" pageNumber="137">Figures 1C</figureCitation>
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Plusioglyphiulus khmer sp. nov., ♂ paratype A-C anterior part of body, lateral, dorsal and ventral views, respectively D, E midbody segments, dorsal and lateral views, respectively F cross-section of a midbody segment G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.938.51234.figure4" httpUri="https://binary.pensoft.net/fig/418286" pageId="0" pageNumber="137">, 4</figureCitation>
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Plusioglyphiulus khmer sp. nov. A, B ♂ paratype C-L ♂ holotype A gnathochilarium, ventral view B collum, dorsal view C antenna, lateral view D, E ♂ legs 1, anterior and posterior views, respectively F ♂ legs 2, posterior view G ♂ legs 3, posterior view H, I anterior gonopods, anterior and posterior views, respectively J, K posterior gonopods, posterior and anterior views, respectively L midbody leg, anterior view. Abbreviations: cxp 1 1 st coxosternal process cxp 2 2 nd coxosternal process te telopodites d terminal medial spike k a small bifid process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.938.51234.figure5" httpUri="https://binary.pensoft.net/fig/418287" pageId="0" pageNumber="137">, 5</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="137" type="type material">
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<paragraph pageId="0" pageNumber="137">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="137">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="137">Holotype</emphasis>
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♂ (CUMZ-CAM186), Cambodia, Kep Province, Damnak
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<normalizedToken originalValue="Chang’aeur">Chang'aeur</normalizedToken>
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District, Ou Krasar, Phnom Sorsia Temple (locality code C047),
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<geoCoordinate degrees="10" direction="north" minutes="33" orientation="latitude" precision="15" seconds="54" value="10.565001">10°33'54"N</geoCoordinate>
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,
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<geoCoordinate degrees="104" direction="east" minutes="17" orientation="longitude" precision="15" seconds="2" value="104.28389">104°17'2"E</geoCoordinate>
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, 16.09.2019, leg. E. Jeratthitikul and R. Srisonchai.
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</paragraph>
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<paragraph pageId="0" pageNumber="137">
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<emphasis bold="true" pageId="0" pageNumber="137">
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<emphasis italics="true" pageId="0" pageNumber="137">Paratypes</emphasis>
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.
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</emphasis>
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16 ♂, 17 ♀, 6 juveniles (CUMZ-CAM186), 2 ♂, 2 ♀ (ZMUM), 2 ♂, 2 ♀ (NHMD), 2 ♂, 2 ♀ (ZRC), same locality, together with holotype.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="137" type="name">
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<paragraph pageId="0" pageNumber="137">Name.</paragraph>
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<paragraph pageId="0" pageNumber="137">
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To emphasize "
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<taxonomicName authorityName="Likhitrakarn & Golovatch & Thach & Chhuoy & Ngor & Srisonchai & Sutcharit & Panha" authorityYear="2020" class="Diplopoda" family="Cambalopsidae" genus="Plusioglyphiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="khmer" order="Spirostreptida" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="khmer">khmer</taxonomicName>
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", referring to the main people of Cambodia; a noun in apposition.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="137" type="diagnosis">
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<paragraph pageId="0" pageNumber="137">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="137">
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This new species differs from all congeners by all crests on the collum being undivided, mostly complete (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Plusioglyphiulus khmer sp. nov. A, B ♂ paratype C-L ♂ holotype A gnathochilarium, ventral view B collum, dorsal view C antenna, lateral view D, E ♂ legs 1, anterior and posterior views, respectively F ♂ legs 2, posterior view G ♂ legs 3, posterior view H, I anterior gonopods, anterior and posterior views, respectively J, K posterior gonopods, posterior and anterior views, respectively L midbody leg, anterior view. Abbreviations: cxp 1 1 st coxosternal process cxp 2 2 nd coxosternal process te telopodites d terminal medial spike k a small bifid process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.938.51234.figure5" httpUri="https://binary.pensoft.net/fig/418287" pageId="0" pageNumber="137">5B</figureCitation>
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) and male legs 1 with very long and one-segmented telopodites (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Plusioglyphiulus khmer sp. nov. A, B ♂ paratype C-L ♂ holotype A gnathochilarium, ventral view B collum, dorsal view C antenna, lateral view D, E ♂ legs 1, anterior and posterior views, respectively F ♂ legs 2, posterior view G ♂ legs 3, posterior view H, I anterior gonopods, anterior and posterior views, respectively J, K posterior gonopods, posterior and anterior views, respectively L midbody leg, anterior view. Abbreviations: cxp 1 1 st coxosternal process cxp 2 2 nd coxosternal process te telopodites d terminal medial spike k a small bifid process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.938.51234.figure5" httpUri="https://binary.pensoft.net/fig/418287" pageId="0" pageNumber="137">5D, E</figureCitation>
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), as well as by the presence of 2+2 paramedian, characteristically long, slender, coxosternal processes of the anterior gonopods, a shorter and only apically curved posterior (
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<emphasis bold="true" pageId="0" pageNumber="137">cxp1</emphasis>
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) pair, and a longer, mesally micropapillate and regularly curved anterior (
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<emphasis bold="true" pageId="0" pageNumber="137">cxp2</emphasis>
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) one (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Plusioglyphiulus khmer sp. nov. A, B ♂ paratype C-L ♂ holotype A gnathochilarium, ventral view B collum, dorsal view C antenna, lateral view D, E ♂ legs 1, anterior and posterior views, respectively F ♂ legs 2, posterior view G ♂ legs 3, posterior view H, I anterior gonopods, anterior and posterior views, respectively J, K posterior gonopods, posterior and anterior views, respectively L midbody leg, anterior view. Abbreviations: cxp 1 1 st coxosternal process cxp 2 2 nd coxosternal process te telopodites d terminal medial spike k a small bifid process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.938.51234.figure5" httpUri="https://binary.pensoft.net/fig/418287" pageId="0" pageNumber="137">5H, I</figureCitation>
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); the posterior gonopods are oblong-subtrapeziform, membranous, each with a small apical spike (
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<emphasis bold="true" pageId="0" pageNumber="137">d</emphasis>
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) and a bifid process (
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<emphasis bold="true" pageId="0" pageNumber="137">k</emphasis>
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) at about 1/3 telopodite length on anterior face (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Plusioglyphiulus khmer sp. nov. A, B ♂ paratype C-L ♂ holotype A gnathochilarium, ventral view B collum, dorsal view C antenna, lateral view D, E ♂ legs 1, anterior and posterior views, respectively F ♂ legs 2, posterior view G ♂ legs 3, posterior view H, I anterior gonopods, anterior and posterior views, respectively J, K posterior gonopods, posterior and anterior views, respectively L midbody leg, anterior view. Abbreviations: cxp 1 1 st coxosternal process cxp 2 2 nd coxosternal process te telopodites d terminal medial spike k a small bifid process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.938.51234.figure5" httpUri="https://binary.pensoft.net/fig/418287" pageId="0" pageNumber="137">5J, K</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="137" type="description">
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<paragraph pageId="0" pageNumber="137">Description.</paragraph>
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<paragraph pageId="0" pageNumber="137">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="137">Length</emphasis>
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of holotype ca 26 mm; adult paratypes 19.3-25.4 (♂) or 14.1-27.0 mm (♀), juveniles 8.5-11.5 mm long; midbody segments round in cross-section (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Plusioglyphiulus khmer sp. nov., ♂ paratype A-C anterior part of body, lateral, dorsal and ventral views, respectively D, E midbody segments, dorsal and lateral views, respectively F cross-section of a midbody segment G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.938.51234.figure4" httpUri="https://binary.pensoft.net/fig/418286" pageId="0" pageNumber="137">4F</figureCitation>
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), their width (horizontal diameter) shorter than height (vertical diameter), width in holotype 1.2 mm; paratypes 1.1-1.5 (♂), 1.1-1.6 mm (♀) or 0.8-1.1 mm (juveniles).
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</paragraph>
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<caption doi="10.3897/zookeys.938.51234.figure4" httpUri="https://binary.pensoft.net/fig/418286" pageId="0" pageNumber="137" start="Figure 4" startId="F4">
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<paragraph pageId="0" pageNumber="137">
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<emphasis bold="true" pageId="0" pageNumber="137">Figure 4.</emphasis>
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<taxonomicName authorityName="Likhitrakarn & Golovatch & Thach & Chhuoy & Ngor & Srisonchai & Sutcharit & Panha" authorityYear="2020" class="Diplopoda" family="Cambalopsidae" genus="Plusioglyphiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plusioglyphiulus khmer" order="Spirostreptida" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="khmer">
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<emphasis italics="true" pageId="0" pageNumber="137">Plusioglyphiulus khmer</emphasis>
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</taxonomicName>
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sp. nov., ♂ paratype
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<emphasis bold="true" pageId="0" pageNumber="137">A-C</emphasis>
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anterior part of body, lateral, dorsal and ventral views, respectively
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<emphasis bold="true" pageId="0" pageNumber="137">D, E</emphasis>
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midbody segments, dorsal and lateral views, respectively
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<emphasis bold="true" pageId="0" pageNumber="137">F</emphasis>
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cross-section of a midbody segment
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<emphasis bold="true" pageId="0" pageNumber="137">G-I</emphasis>
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posterior part of body, lateral, dorsal and ventral views, respectively.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="137">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="137">Coloration</emphasis>
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of live animals light brown to chocolate brown (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus, live coloration A Plusioglyphiulus boutini Mauries, 1970, ♂ from Prasat Phnom Totong Temple B Plusioglyphiulus biserratus sp. nov., ♀ paratype C Plusioglyphiulus khmer sp. nov., ♂ paratype. All pictures by R. Srisonchai, not taken to scale." figureDoi="10.3897/zookeys.938.51234.figure1" httpUri="https://binary.pensoft.net/fig/418283" pageId="0" pageNumber="137">1C</figureCitation>
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) with blackish or dark brown anterior and posterior parts of body; antennae light brown to brown; venter and legs light yellowish to yellowish white; rather contrasting dark brownish, lateral, longitudinal stripes above ozopores on each side, both interrupted mid-dorsally by a light wide axial stripe; ommatidia brown to blackish (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Plusioglyphiulus khmer sp. nov., ♂ paratype A-C anterior part of body, lateral, dorsal and ventral views, respectively D, E midbody segments, dorsal and lateral views, respectively F cross-section of a midbody segment G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.938.51234.figure4" httpUri="https://binary.pensoft.net/fig/418286" pageId="0" pageNumber="137">4A, C</figureCitation>
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); coloration in alcohol, after six months of preservation (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Plusioglyphiulus khmer sp. nov., ♂ paratype A-C anterior part of body, lateral, dorsal and ventral views, respectively D, E midbody segments, dorsal and lateral views, respectively F cross-section of a midbody segment G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.938.51234.figure4" httpUri="https://binary.pensoft.net/fig/418286" pageId="0" pageNumber="137">4</figureCitation>
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), similar to live one, but body red-brownish or dark castaneous brown to grey-brown; vertex dark brown to yellow-brown; antennae dark brown to blackish; venter and legs light yellowish to yellowish red (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Plusioglyphiulus khmer sp. nov., ♂ paratype A-C anterior part of body, lateral, dorsal and ventral views, respectively D, E midbody segments, dorsal and lateral views, respectively F cross-section of a midbody segment G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.938.51234.figure4" httpUri="https://binary.pensoft.net/fig/418286" pageId="0" pageNumber="137">4A-C, E-G, I</figureCitation>
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).
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</paragraph>
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<paragraph pageId="0" pageNumber="137">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="137">Adult body</emphasis>
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with 53p+2a+T (holotype); paratypes with 46-57p+1-3a+T (♂), 44-55p+2-4a+T (♀) or 35-42+3-4a +T (juveniles). Eye patches transversely ovoid, with 11-18 ommatidia arranged in three or four longitudinal rows (adult) (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Plusioglyphiulus khmer sp. nov., ♂ paratype A-C anterior part of body, lateral, dorsal and ventral views, respectively D, E midbody segments, dorsal and lateral views, respectively F cross-section of a midbody segment G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.938.51234.figure4" httpUri="https://binary.pensoft.net/fig/418286" pageId="0" pageNumber="137">4A, C</figureCitation>
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) or with 6-10 ommatidia in three longitudinal rows (juveniles). Clypeus with three teeth anteromedially (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Plusioglyphiulus khmer sp. nov., ♂ paratype A-C anterior part of body, lateral, dorsal and ventral views, respectively D, E midbody segments, dorsal and lateral views, respectively F cross-section of a midbody segment G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.938.51234.figure4" httpUri="https://binary.pensoft.net/fig/418286" pageId="0" pageNumber="137">4C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="137">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="137">Antennae</emphasis>
|
||
short and clavate (Figs
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus, live coloration A Plusioglyphiulus boutini Mauries, 1970, ♂ from Prasat Phnom Totong Temple B Plusioglyphiulus biserratus sp. nov., ♀ paratype C Plusioglyphiulus khmer sp. nov., ♂ paratype. All pictures by R. Srisonchai, not taken to scale." figureDoi="10.3897/zookeys.938.51234.figure1" httpUri="https://binary.pensoft.net/fig/418283" pageId="0" pageNumber="137">1C</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Plusioglyphiulus khmer sp. nov., ♂ paratype A-C anterior part of body, lateral, dorsal and ventral views, respectively D, E midbody segments, dorsal and lateral views, respectively F cross-section of a midbody segment G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.938.51234.figure4" httpUri="https://binary.pensoft.net/fig/418286" pageId="0" pageNumber="137">4A, C</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Plusioglyphiulus khmer sp. nov. A, B ♂ paratype C-L ♂ holotype A gnathochilarium, ventral view B collum, dorsal view C antenna, lateral view D, E ♂ legs 1, anterior and posterior views, respectively F ♂ legs 2, posterior view G ♂ legs 3, posterior view H, I anterior gonopods, anterior and posterior views, respectively J, K posterior gonopods, posterior and anterior views, respectively L midbody leg, anterior view. Abbreviations: cxp 1 1 st coxosternal process cxp 2 2 nd coxosternal process te telopodites d terminal medial spike k a small bifid process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.938.51234.figure5" httpUri="https://binary.pensoft.net/fig/418287" pageId="0" pageNumber="137">5C</figureCitation>
|
||
), extending behind segment 3 laterally, antennomeres 5 and 6 each with a small apicodorsal field or corolla of bacilliform sensilla (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Plusioglyphiulus khmer sp. nov. A, B ♂ paratype C-L ♂ holotype A gnathochilarium, ventral view B collum, dorsal view C antenna, lateral view D, E ♂ legs 1, anterior and posterior views, respectively F ♂ legs 2, posterior view G ♂ legs 3, posterior view H, I anterior gonopods, anterior and posterior views, respectively J, K posterior gonopods, posterior and anterior views, respectively L midbody leg, anterior view. Abbreviations: cxp 1 1 st coxosternal process cxp 2 2 nd coxosternal process te telopodites d terminal medial spike k a small bifid process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.938.51234.figure5" httpUri="https://binary.pensoft.net/fig/418287" pageId="0" pageNumber="137">5C</figureCitation>
|
||
). Gnathochilarium oligotrichous, each lamella lingualis with two or three setae; promentum bare, separated from eumentum by a distinct transverse suture (
|
||
<emphasis italics="true" pageId="0" pageNumber="137">n</emphasis>
|
||
= 2) (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Plusioglyphiulus khmer sp. nov. A, B ♂ paratype C-L ♂ holotype A gnathochilarium, ventral view B collum, dorsal view C antenna, lateral view D, E ♂ legs 1, anterior and posterior views, respectively F ♂ legs 2, posterior view G ♂ legs 3, posterior view H, I anterior gonopods, anterior and posterior views, respectively J, K posterior gonopods, posterior and anterior views, respectively L midbody leg, anterior view. Abbreviations: cxp 1 1 st coxosternal process cxp 2 2 nd coxosternal process te telopodites d terminal medial spike k a small bifid process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.938.51234.figure5" httpUri="https://binary.pensoft.net/fig/418287" pageId="0" pageNumber="137">5A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.938.51234.figure5" httpUri="https://binary.pensoft.net/fig/418287" pageId="0" pageNumber="137" start="Figure 5" startId="F5">
|
||
<paragraph pageId="0" pageNumber="137">
|
||
<emphasis bold="true" pageId="0" pageNumber="137">Figure 5.</emphasis>
|
||
<taxonomicName authorityName="Likhitrakarn & Golovatch & Thach & Chhuoy & Ngor & Srisonchai & Sutcharit & Panha" authorityYear="2020" class="Diplopoda" family="Cambalopsidae" genus="Plusioglyphiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plusioglyphiulus khmer" order="Spirostreptida" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="khmer">
|
||
<emphasis italics="true" pageId="0" pageNumber="137">Plusioglyphiulus khmer</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="137">A, B</emphasis>
|
||
♂ paratype
|
||
<emphasis bold="true" pageId="0" pageNumber="137">C-L</emphasis>
|
||
♂ holotype
|
||
<emphasis bold="true" pageId="0" pageNumber="137">A</emphasis>
|
||
gnathochilarium, ventral view
|
||
<emphasis bold="true" pageId="0" pageNumber="137">B</emphasis>
|
||
collum, dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="137">C</emphasis>
|
||
antenna, lateral view
|
||
<emphasis bold="true" pageId="0" pageNumber="137">D, E</emphasis>
|
||
♂ legs 1, anterior and posterior views, respectively
|
||
<emphasis bold="true" pageId="0" pageNumber="137">F</emphasis>
|
||
♂ legs 2, posterior view
|
||
<emphasis bold="true" pageId="0" pageNumber="137">G</emphasis>
|
||
♂ legs 3, posterior view
|
||
<emphasis bold="true" pageId="0" pageNumber="137">H, I</emphasis>
|
||
anterior gonopods, anterior and posterior views, respectively
|
||
<emphasis bold="true" pageId="0" pageNumber="137">J, K</emphasis>
|
||
posterior gonopods, posterior and anterior views, respectively
|
||
<emphasis bold="true" pageId="0" pageNumber="137">L</emphasis>
|
||
midbody leg, anterior view. Abbreviations:
|
||
<emphasis bold="true" pageId="0" pageNumber="137">cxp1</emphasis>
|
||
1st coxosternal process
|
||
<emphasis bold="true" pageId="0" pageNumber="137">cxp2</emphasis>
|
||
2nd coxosternal process
|
||
<emphasis bold="true" pageId="0" pageNumber="137">te</emphasis>
|
||
telopodites
|
||
<emphasis bold="true" pageId="0" pageNumber="137">d</emphasis>
|
||
terminal medial spike
|
||
<emphasis bold="true" pageId="0" pageNumber="137">k</emphasis>
|
||
a small bifid process. Scale bars: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="137">
|
||
<taxonomicName genus="Postcollum" lsidName="Postcollum" pageId="0" pageNumber="137" rank="genus">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="137">Postcollum</emphasis>
|
||
</taxonomicName>
|
||
constriction evident, but collum only moderately enlarged (Figs
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus, live coloration A Plusioglyphiulus boutini Mauries, 1970, ♂ from Prasat Phnom Totong Temple B Plusioglyphiulus biserratus sp. nov., ♀ paratype C Plusioglyphiulus khmer sp. nov., ♂ paratype. All pictures by R. Srisonchai, not taken to scale." figureDoi="10.3897/zookeys.938.51234.figure1" httpUri="https://binary.pensoft.net/fig/418283" pageId="0" pageNumber="137">1C</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Plusioglyphiulus khmer sp. nov., ♂ paratype A-C anterior part of body, lateral, dorsal and ventral views, respectively D, E midbody segments, dorsal and lateral views, respectively F cross-section of a midbody segment G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.938.51234.figure4" httpUri="https://binary.pensoft.net/fig/418286" pageId="0" pageNumber="137">4A-C</figureCitation>
|
||
). Carinotaxic formula of collum: 1+ta+2+3+4+ta+5(5p/t)+ta+P+ma (Figs
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Plusioglyphiulus khmer sp. nov., ♂ paratype A-C anterior part of body, lateral, dorsal and ventral views, respectively D, E midbody segments, dorsal and lateral views, respectively F cross-section of a midbody segment G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.938.51234.figure4" httpUri="https://binary.pensoft.net/fig/418286" pageId="0" pageNumber="137">4A-C</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Plusioglyphiulus khmer sp. nov. A, B ♂ paratype C-L ♂ holotype A gnathochilarium, ventral view B collum, dorsal view C antenna, lateral view D, E ♂ legs 1, anterior and posterior views, respectively F ♂ legs 2, posterior view G ♂ legs 3, posterior view H, I anterior gonopods, anterior and posterior views, respectively J, K posterior gonopods, posterior and anterior views, respectively L midbody leg, anterior view. Abbreviations: cxp 1 1 st coxosternal process cxp 2 2 nd coxosternal process te telopodites d terminal medial spike k a small bifid process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.938.51234.figure5" httpUri="https://binary.pensoft.net/fig/418287" pageId="0" pageNumber="137">5B</figureCitation>
|
||
). Carinotaxy of metatergum 2, 8/8+m/m+8/8; of metaterga 3 and 4, 7/7+m/m+7/7 (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Plusioglyphiulus khmer sp. nov., ♂ paratype A-C anterior part of body, lateral, dorsal and ventral views, respectively D, E midbody segments, dorsal and lateral views, respectively F cross-section of a midbody segment G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.938.51234.figure4" httpUri="https://binary.pensoft.net/fig/418286" pageId="0" pageNumber="137">4A, B</figureCitation>
|
||
); formula of metaterga 5 and following metaterga, except last few, usually 3/3+I/i+3/3/3+m/m+3/3/3+I/i+3/3 (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Plusioglyphiulus khmer sp. nov., ♂ paratype A-C anterior part of body, lateral, dorsal and ventral views, respectively D, E midbody segments, dorsal and lateral views, respectively F cross-section of a midbody segment G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.938.51234.figure4" httpUri="https://binary.pensoft.net/fig/418286" pageId="0" pageNumber="137">4A, B, D-H</figureCitation>
|
||
); of legless segments, usually 7+m+7 (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Plusioglyphiulus khmer sp. nov., ♂ paratype A-C anterior part of body, lateral, dorsal and ventral views, respectively D, E midbody segments, dorsal and lateral views, respectively F cross-section of a midbody segment G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.938.51234.figure4" httpUri="https://binary.pensoft.net/fig/418286" pageId="0" pageNumber="137">4G, H</figureCitation>
|
||
); all crests and tubercles, including poriferous cones, rather low. Dorsal crests on several posteriormost segments lower than others (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Plusioglyphiulus khmer sp. nov., ♂ paratype A-C anterior part of body, lateral, dorsal and ventral views, respectively D, E midbody segments, dorsal and lateral views, respectively F cross-section of a midbody segment G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.938.51234.figure4" httpUri="https://binary.pensoft.net/fig/418286" pageId="0" pageNumber="137">4A, B, D-H</figureCitation>
|
||
). Porosteles large, low, conical, round, directed caudolaterad, wider than high (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Plusioglyphiulus khmer sp. nov., ♂ paratype A-C anterior part of body, lateral, dorsal and ventral views, respectively D, E midbody segments, dorsal and lateral views, respectively F cross-section of a midbody segment G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.938.51234.figure4" httpUri="https://binary.pensoft.net/fig/418286" pageId="0" pageNumber="137">4D-H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="137">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="137">Tegument</emphasis>
|
||
finely alveolate-areolate (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Plusioglyphiulus khmer sp. nov., ♂ paratype A-C anterior part of body, lateral, dorsal and ventral views, respectively D, E midbody segments, dorsal and lateral views, respectively F cross-section of a midbody segment G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.938.51234.figure4" httpUri="https://binary.pensoft.net/fig/418286" pageId="0" pageNumber="137">4A, B, D, E, G, H</figureCitation>
|
||
), dull throughout. Metatergal setae absent. Pleural regions of segments 2-4 conspicuously elongated, flap-shaped, especially well so on segment 3 (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Plusioglyphiulus khmer sp. nov., ♂ paratype A-C anterior part of body, lateral, dorsal and ventral views, respectively D, E midbody segments, dorsal and lateral views, respectively F cross-section of a midbody segment G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.938.51234.figure4" httpUri="https://binary.pensoft.net/fig/418286" pageId="0" pageNumber="137">4A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="137">
|
||
<taxonomicName genus="Epiproct" lsidName="Epiproct" pageId="0" pageNumber="137" rank="genus">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="137">Epiproct</emphasis>
|
||
</taxonomicName>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Plusioglyphiulus khmer sp. nov., ♂ paratype A-C anterior part of body, lateral, dorsal and ventral views, respectively D, E midbody segments, dorsal and lateral views, respectively F cross-section of a midbody segment G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.938.51234.figure4" httpUri="https://binary.pensoft.net/fig/418286" pageId="0" pageNumber="137">4G-I</figureCitation>
|
||
) broadly rounded apically, with 2+2 small, paramedian tubercles at midway. Paraprocts rather clearly flattened, each with a faint premarginal sulcus medially (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Plusioglyphiulus khmer sp. nov., ♂ paratype A-C anterior part of body, lateral, dorsal and ventral views, respectively D, E midbody segments, dorsal and lateral views, respectively F cross-section of a midbody segment G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.938.51234.figure4" httpUri="https://binary.pensoft.net/fig/418286" pageId="0" pageNumber="137">4G, I</figureCitation>
|
||
). Hypoproct clearly emarginate at caudal margin (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Plusioglyphiulus khmer sp. nov., ♂ paratype A-C anterior part of body, lateral, dorsal and ventral views, respectively D, E midbody segments, dorsal and lateral views, respectively F cross-section of a midbody segment G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.938.51234.figure4" httpUri="https://binary.pensoft.net/fig/418286" pageId="0" pageNumber="137">4I</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="137">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="137">Ventral</emphasis>
|
||
flaps behind gonopod aperture on male segment 7 evident, distinguishable as rather high swellings with rounded flaps bent abruptly caudad (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Plusioglyphiulus khmer sp. nov., ♂ paratype A-C anterior part of body, lateral, dorsal and ventral views, respectively D, E midbody segments, dorsal and lateral views, respectively F cross-section of a midbody segment G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.938.51234.figure4" httpUri="https://binary.pensoft.net/fig/418286" pageId="0" pageNumber="137">4C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="137">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="137">Legs</emphasis>
|
||
short, on midbody segments about 1/3 as long as body diameter (Figs
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Plusioglyphiulus khmer sp. nov., ♂ paratype A-C anterior part of body, lateral, dorsal and ventral views, respectively D, E midbody segments, dorsal and lateral views, respectively F cross-section of a midbody segment G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.938.51234.figure4" httpUri="https://binary.pensoft.net/fig/418286" pageId="0" pageNumber="137">4E-G</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Plusioglyphiulus khmer sp. nov. A, B ♂ paratype C-L ♂ holotype A gnathochilarium, ventral view B collum, dorsal view C antenna, lateral view D, E ♂ legs 1, anterior and posterior views, respectively F ♂ legs 2, posterior view G ♂ legs 3, posterior view H, I anterior gonopods, anterior and posterior views, respectively J, K posterior gonopods, posterior and anterior views, respectively L midbody leg, anterior view. Abbreviations: cxp 1 1 st coxosternal process cxp 2 2 nd coxosternal process te telopodites d terminal medial spike k a small bifid process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.938.51234.figure5" httpUri="https://binary.pensoft.net/fig/418287" pageId="0" pageNumber="137">5L</figureCitation>
|
||
), claw at base with a strong, spiniform, accessory claw about 1/3 as long as claw itself (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Plusioglyphiulus khmer sp. nov. A, B ♂ paratype C-L ♂ holotype A gnathochilarium, ventral view B collum, dorsal view C antenna, lateral view D, E ♂ legs 1, anterior and posterior views, respectively F ♂ legs 2, posterior view G ♂ legs 3, posterior view H, I anterior gonopods, anterior and posterior views, respectively J, K posterior gonopods, posterior and anterior views, respectively L midbody leg, anterior view. Abbreviations: cxp 1 1 st coxosternal process cxp 2 2 nd coxosternal process te telopodites d terminal medial spike k a small bifid process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.938.51234.figure5" httpUri="https://binary.pensoft.net/fig/418287" pageId="0" pageNumber="137">5L</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="137">
|
||
Male legs 1 with a usual, strong, central hook (actually a double structure of tightly appressed hooks), regularly curved forward; a pair of 1-segmented, sac-shaped and very long telopodites, the latter almost as long as central hook (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Plusioglyphiulus khmer sp. nov. A, B ♂ paratype C-L ♂ holotype A gnathochilarium, ventral view B collum, dorsal view C antenna, lateral view D, E ♂ legs 1, anterior and posterior views, respectively F ♂ legs 2, posterior view G ♂ legs 3, posterior view H, I anterior gonopods, anterior and posterior views, respectively J, K posterior gonopods, posterior and anterior views, respectively L midbody leg, anterior view. Abbreviations: cxp 1 1 st coxosternal process cxp 2 2 nd coxosternal process te telopodites d terminal medial spike k a small bifid process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.938.51234.figure5" httpUri="https://binary.pensoft.net/fig/418287" pageId="0" pageNumber="137">5D, E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="137">
|
||
Male legs 2 strongly enlarged, with very high and large coxae; telopodites hirsute on anterior face; penes subconical, truncate apically, fused at base (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Plusioglyphiulus khmer sp. nov. A, B ♂ paratype C-L ♂ holotype A gnathochilarium, ventral view B collum, dorsal view C antenna, lateral view D, E ♂ legs 1, anterior and posterior views, respectively F ♂ legs 2, posterior view G ♂ legs 3, posterior view H, I anterior gonopods, anterior and posterior views, respectively J, K posterior gonopods, posterior and anterior views, respectively L midbody leg, anterior view. Abbreviations: cxp 1 1 st coxosternal process cxp 2 2 nd coxosternal process te telopodites d terminal medial spike k a small bifid process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.938.51234.figure5" httpUri="https://binary.pensoft.net/fig/418287" pageId="0" pageNumber="137">5F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="137">
|
||
Male legs 3 modified as usual, with particularly elongate and slender coxae and shortened telopodites (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Plusioglyphiulus khmer sp. nov. A, B ♂ paratype C-L ♂ holotype A gnathochilarium, ventral view B collum, dorsal view C antenna, lateral view D, E ♂ legs 1, anterior and posterior views, respectively F ♂ legs 2, posterior view G ♂ legs 3, posterior view H, I anterior gonopods, anterior and posterior views, respectively J, K posterior gonopods, posterior and anterior views, respectively L midbody leg, anterior view. Abbreviations: cxp 1 1 st coxosternal process cxp 2 2 nd coxosternal process te telopodites d terminal medial spike k a small bifid process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.938.51234.figure5" httpUri="https://binary.pensoft.net/fig/418287" pageId="0" pageNumber="137">5G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="137">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="137">Anterior gonopods</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Plusioglyphiulus khmer sp. nov. A, B ♂ paratype C-L ♂ holotype A gnathochilarium, ventral view B collum, dorsal view C antenna, lateral view D, E ♂ legs 1, anterior and posterior views, respectively F ♂ legs 2, posterior view G ♂ legs 3, posterior view H, I anterior gonopods, anterior and posterior views, respectively J, K posterior gonopods, posterior and anterior views, respectively L midbody leg, anterior view. Abbreviations: cxp 1 1 st coxosternal process cxp 2 2 nd coxosternal process te telopodites d terminal medial spike k a small bifid process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.938.51234.figure5" httpUri="https://binary.pensoft.net/fig/418287" pageId="0" pageNumber="137">5H, I</figureCitation>
|
||
) rather complex, with 2+2 paramedian, long, slender, coxosternal processes: a shorter and only apically curved posterior (
|
||
<emphasis bold="true" pageId="0" pageNumber="137">cxp1</emphasis>
|
||
) pair, and a longer, mesally micropapillate and regularly curved anterior (
|
||
<emphasis bold="true" pageId="0" pageNumber="137">cxp2</emphasis>
|
||
) one (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Plusioglyphiulus khmer sp. nov. A, B ♂ paratype C-L ♂ holotype A gnathochilarium, ventral view B collum, dorsal view C antenna, lateral view D, E ♂ legs 1, anterior and posterior views, respectively F ♂ legs 2, posterior view G ♂ legs 3, posterior view H, I anterior gonopods, anterior and posterior views, respectively J, K posterior gonopods, posterior and anterior views, respectively L midbody leg, anterior view. Abbreviations: cxp 1 1 st coxosternal process cxp 2 2 nd coxosternal process te telopodites d terminal medial spike k a small bifid process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.938.51234.figure5" httpUri="https://binary.pensoft.net/fig/418287" pageId="0" pageNumber="137">5H, I</figureCitation>
|
||
); telopodites (
|
||
<emphasis bold="true" pageId="0" pageNumber="137">te</emphasis>
|
||
) finger-shaped, subcylindrical, long, apically setose, about as high as
|
||
<emphasis bold="true" pageId="0" pageNumber="137">cxp2</emphasis>
|
||
, attached to coxal region caudolaterally, probably capable of movement.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="137">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="137">Posterior gonopods</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Plusioglyphiulus khmer sp. nov. A, B ♂ paratype C-L ♂ holotype A gnathochilarium, ventral view B collum, dorsal view C antenna, lateral view D, E ♂ legs 1, anterior and posterior views, respectively F ♂ legs 2, posterior view G ♂ legs 3, posterior view H, I anterior gonopods, anterior and posterior views, respectively J, K posterior gonopods, posterior and anterior views, respectively L midbody leg, anterior view. Abbreviations: cxp 1 1 st coxosternal process cxp 2 2 nd coxosternal process te telopodites d terminal medial spike k a small bifid process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.938.51234.figure5" httpUri="https://binary.pensoft.net/fig/418287" pageId="0" pageNumber="137">5J, K</figureCitation>
|
||
) rather long (high), simple; coxites well-separated from sternum, fused only basally, oblong-subtrapeziform, membranous, with traces of telopodites in the form of a small, latero-parabasal field of microsetae on posterior face of each gonopod; each coxite with a terminal medial spike (
|
||
<emphasis bold="true" pageId="0" pageNumber="137">d</emphasis>
|
||
), a subterminal lobule at
|
||
<emphasis bold="true" pageId="0" pageNumber="137">d</emphasis>
|
||
base, and a small bifid process (
|
||
<emphasis bold="true" pageId="0" pageNumber="137">k</emphasis>
|
||
) at about 1/3 coxite length on frontal face; distal half lamellose, fringed and with a deep fovea subapically.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="137" type="remarks">
|
||
<paragraph pageId="0" pageNumber="137">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="137">
|
||
This species was found together with a single male specimen of
|
||
<taxonomicName baseAuthorityName="Saussure" baseAuthorityYear="1860" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha coarctata" order="Polydesmida" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="coarctata">
|
||
<emphasis italics="true" pageId="0" pageNumber="137">Orthomorpha coarctata</emphasis>
|
||
</taxonomicName>
|
||
(De Saussure, 1860).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |