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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1154.86237" ID-Pensoft-Pub="1313-2970-1154-49" ID-Pensoft-UUID="E4F41DD1A11059DAB4A979F00E85D50F" ID-ZooBank="4E410C2ACC03438F8AC7C5370819DE6C" ModsDocID="1313-2970-1154-49" checkinTime="1679436496911" checkinUser="pensoft" docAuthor="Wu, Min, Shen, Wang &amp; Chen, Zhong-Guang" docDate="2023" docId="4436028B101256E5AD42AC719F02AC3C" docLanguage="en" docName="ZooKeys 1154: 49-147" docOrigin="ZooKeys 1154" docPubDate="2023-03-20" docSource="http://dx.doi.org/10.3897/zookeys.1154.86237" docTitle="Laeocathaica nordsiecki Wu 2023, sp. nov." docType="treatment" docUuid="BBBD09C3-661E-4E70-AA52-59A2B8E818A3" docUuidSource="ZooBank" docVersion="2" id="E4F41DD1A11059DAB4A979F00E85D50F" lastPageNumber="49" masterDocId="E4F41DD1A11059DAB4A979F00E85D50F" masterDocTitle="Land snail diversity in central China: revision of Laeocathaica Moellendorff, 1899 (Gastropoda, Camaenidae), with descriptions of seven new species" masterLastPageNumber="147" masterPageNumber="49" pageNumber="49" updateTime="1679436803143" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Land snail diversity in central China: revision of Laeocathaica Moellendorff, 1899 (Gastropoda, Camaenidae), with descriptions of seven new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wu, Min</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5434-5544</mods:nameIdentifier>
<mods:affiliation>School of Life Sciences, Nanjing University, Nanjing 210023, China</mods:affiliation>
<mods:nameIdentifier type="email">minwu1969@aliyun.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Shen, Wang</mods:namePart>
<mods:affiliation>School of Life Sciences, Nanjing University, Nanjing 210023, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Chen, Zhong-Guang</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-2689-3321</mods:nameIdentifier>
<mods:affiliation>Jiangxi Province Key Laboratory of Watershed Ecosystem Change and Biodiversity, Center for Watershed Ecology, Institute of Life Science and School of Life Sciences, Nanchang University, Nanchang 330031, China</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-03-20</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1154</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>49</mods:start>
<mods:end>147</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1154.86237</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1154.86237</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1154-49</mods:identifier>
<mods:identifier type="ZooBank">4E410C2ACC03438F8AC7C5370819DE6C</mods:identifier>
<mods:identifier type="Pensoft-UUID">E4F41DD1A11059DAB4A979F00E85D50F</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:zoobank.org:act:BBBD09C3-661E-4E70-AA52-59A2B8E818A3" httpUri="http://treatment.plazi.org/id/4436028B101256E5AD42AC719F02AC3C" lastPageNumber="49" pageId="0" pageNumber="49">
<subSubSection pageId="0" pageNumber="49" type="nomenclature">
<paragraph pageId="0" pageNumber="49">
<taxonomicName LSID="https://zoobank.org/BBBD09C3-661E-4E70-AA52-59A2B8E818A3" authority="Wu" authorityName="Wu" authorityYear="2023" class="Gastropoda" family="Camaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica nordsiecki" order="Stylommatophora" pageId="0" pageNumber="49" phylum="Mollusca" rank="species" species="nordsiecki" status="sp. nov.">Laeocathaica nordsiecki Wu</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="49">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="49" type="description">
<paragraph pageId="0" pageNumber="49">
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. A distribution map of all known species of Laeocathaica Moellendorff, 1899 except L. anceyi (Moellendorff in Ancey, 1889), L. hisanoi Pall-Gergely, 2022, and L. leucorhaphe Moellendorff, 1899 whose precise localities are not known; white dots: the localities where no Laeocathaica species was ever found B distribution of the new species described in this paper, yellow dots: L. qingchuanensis Wu, sp. nov., green dot: L. zhengpingliui Wu, sp. nov., brown dots: L. parapolytyla Wu, sp. nov., white dot: L. qiminglii Wu, sp. nov., pink dot: L. cheni Wu, sp. nov., blue dots: L. qishilii Wu, sp. nov., orange dots: L. nordsiecki Wu, sp. nov. C distribution of L. carinifera (H. Adams, 1870) (green dots), L. qingchuanensis Wu, sp. nov. (yellow dots), and L. stenochone Moellendorff, 1899 (blue dots) D distribution of L. amdoana Moellendorff, 1899 (green dots), L. distinguenda Moellendorff, 1899 (pink dots), and L. tropidorhaphe Moellendorff, 1899 (blue dots) C equivalent to the larger box in (A), B, D equivalent to the smaller box in (A)." figureDoi="10.3897/zookeys.1154.86237.figure2" httpUri="https://binary.pensoft.net/fig/827345" pageId="0" pageNumber="49">Figs 2A, B</figureCitation>
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. A Laeocathaica zhengpingliui Wu, sp. nov., HBUMM 05553 - spec. 1, holotype B L. cheni Wu, sp. nov., HBUMM 05553 b-spec. 1, holotype C L. qingchuanensis Wu, sp. nov., HBUMM 01179 a-spec. 1, holotype D L. nordsiecki Wu, sp. nov., HBUMM 08446 - spec. 1, holotype." figureDoi="10.3897/zookeys.1154.86237.figure37" httpUri="https://binary.pensoft.net/fig/827380" pageId="0" pageNumber="49">, 37D</figureCitation>
<figureCitation captionStart="Figure 47" captionStartId="F47" captionText="Figure 47. SEM micrographs A, B Laeocathaica odophora Moellendorff, 1899, HBUMM 08430 A protoconch B teleoconch C, D L. dolani (Pilsbry, 1934), HBUMM 08439 C protoconch D teleoconch E, F L. nordsiecki Wu, sp. nov., HBUMM 08446, paratype, subadult E protoconch F teleoconch G, H L. pewzowi Moellendorff, 1899, HBUMM 08452 G protoconch H teleoconch I, J L. carinalis Chen &amp; Zhang, 2004, HBUMM 08453 I protoconch J teleoconch K, L L. zhengpingliui Wu, sp. nov., HBUMM 05553 - spec. 2, paratype, juvenile K protoconch L teleoconch." figureDoi="10.3897/zookeys.1154.86237.figure47" httpUri="https://binary.pensoft.net/fig/827390" pageId="0" pageNumber="49">, 47E, F</figureCitation>
<figureCitation captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Habitats (excluding D) A, B Laeocathaica qishilii Wu, sp. nov., border of Jiuzhaigou County and Wen County C L. nordsiecki Wu, sp. nov., Guoyuanxiang, Jiuzhaigou D L. qiminglii Wu, sp. nov., an indoor photograph E L. qingchuanensis Wu, sp. nov., Dagou Nature Reserve, Qingchuan County F L. cheni Wu, sp. nov., Hengdan, Wenxian." figureDoi="10.3897/zookeys.1154.86237.figure49" httpUri="https://binary.pensoft.net/fig/827392" pageId="0" pageNumber="49">, 49C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="49" type="type material">
<paragraph pageId="0" pageNumber="49">Type material.</paragraph>
<paragraph pageId="0" pageNumber="49">
<emphasis bold="true" italics="true" pageId="0" pageNumber="49">Holotype</emphasis>
HBUMM08446-spec.1, fully mature shell with immature genital system, Guoyuanxiang [郭元乡], Jiuzhaigouxian, Sichuan Province; near point (
<geoCoordinate degrees="33.125506" direction="north" orientation="latitude" precision="1" value="33.125507">33.125506°N</geoCoordinate>
,
<geoCoordinate degrees="104.329876" direction="east" orientation="longitude" precision="1" value="104.32988">104.329876°E</geoCoordinate>
); 2021-IX-24, coll. Chen, Z.-G.
<emphasis bold="true" italics="true" pageId="0" pageNumber="49">Paratypes</emphasis>
HBUMM08446-spec.2-3, 2 fms; HBUMM08446-spec.4-5, 2 subadults; same data of holotype. HBUMM08447-spec.1-2, 2 fms and 3 living juvs (in rearing), Qinglongcun [青龙村], Guoyuanxiang, Jiuzhaigouxian, Sichuan Province; (
<geoCoordinate degrees="33.085615" direction="north" orientation="latitude" precision="1" value="33.085613">33.085615°N</geoCoordinate>
,
<geoCoordinate degrees="104.348250" direction="east" orientation="longitude" precision="1" value="104.34825">104.348250°E</geoCoordinate>
); 2020-VII, coll. Chen, Z.-G.
</paragraph>
<caption doi="10.3897/zookeys.1154.86237.figure44" httpUri="https://binary.pensoft.net/fig/827387" pageId="0" pageNumber="49" start="Figure 44" startId="F44">
<paragraph pageId="0" pageNumber="49">
<emphasis bold="true" pageId="0" pageNumber="49">Figure 44.</emphasis>
Internal view of penis
<emphasis bold="true" pageId="0" pageNumber="49">A, B</emphasis>
<taxonomicName authorityName="Moellendorff" authorityYear="1899" class="Gastropoda" family="Camaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica tropidorhaphe" order="Stylommatophora" pageId="0" pageNumber="49" phylum="Mollusca" rank="species" species="tropidorhaphe">
<emphasis italics="true" pageId="0" pageNumber="49">Laeocathaica tropidorhaphe</emphasis>
</taxonomicName>
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
, 1899
<emphasis bold="true" pageId="0" pageNumber="49">A</emphasis>
general view, HBUMM05619-spec.1
<emphasis bold="true" pageId="0" pageNumber="49">B</emphasis>
proximal penis, HBUMM05664-spec.1
<emphasis bold="true" pageId="0" pageNumber="49">C</emphasis>
<taxonomicName lsidName="L. qishilii" pageId="0" pageNumber="49" rank="species" species="qishilii">
<emphasis italics="true" pageId="0" pageNumber="49">L. qishilii</emphasis>
</taxonomicName>
Wu, sp. nov., general view, HBUMM08298-spec.1, holotype
<emphasis bold="true" pageId="0" pageNumber="49">D</emphasis>
<taxonomicName lsidName="L. zhengpingliui" pageId="0" pageNumber="49" rank="species" species="zhengpingliui">
<emphasis italics="true" pageId="0" pageNumber="49">L. zhengpingliui</emphasis>
</taxonomicName>
Wu, sp. nov., general view, HBUMM05553-spec.1, holotype
<emphasis bold="true" pageId="0" pageNumber="49">E</emphasis>
<taxonomicName lsidName="L. qingchuanensis" pageId="0" pageNumber="49" rank="species" species="qingchuanensis">
<emphasis italics="true" pageId="0" pageNumber="49">L. qingchuanensis</emphasis>
</taxonomicName>
Wu, sp. nov., general view, HBUMM01179-spec.1, holotype
<emphasis bold="true" pageId="0" pageNumber="49">F-H</emphasis>
<taxonomicName lsidName="L. cheni" pageId="0" pageNumber="49" rank="species" species="cheni">
<emphasis italics="true" pageId="0" pageNumber="49">L. cheni</emphasis>
</taxonomicName>
Wu, sp. nov.
<emphasis bold="true" pageId="0" pageNumber="49">F, G</emphasis>
HBUMM05553b-spec.1, holotype
<emphasis bold="true" pageId="0" pageNumber="49">F</emphasis>
general view
<emphasis bold="true" pageId="0" pageNumber="49">G</emphasis>
proximal penis
<emphasis bold="true" pageId="0" pageNumber="49">H</emphasis>
HBUMM08428-spec.1, paratype, partial internal view of distal penis. Arrows indicate the position where two penial internal pilasters fuse into one distal pilaster.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="49" type="description">
<paragraph pageId="0" pageNumber="49">Measurement of holotype.</paragraph>
<paragraph pageId="0" pageNumber="49">Shell height 3.7 mm, maximum diameter 10.8 mm, aperture height 3.0 mm, aperture breadth 3.3 mm, umbilicus diameter 5.0 mm, protoconch whorls 11/2, whorls 51/8.</paragraph>
<caption doi="10.3897/zookeys.1154.86237.figure45" httpUri="https://binary.pensoft.net/fig/827388" pageId="0" pageNumber="49" start="Figure 45" startId="F45">
<paragraph pageId="0" pageNumber="49">
<emphasis bold="true" pageId="0" pageNumber="49">Figure 45.</emphasis>
SEM micrographs
<emphasis bold="true" pageId="0" pageNumber="49">A, B</emphasis>
<taxonomicName authorityName="Moellendorff" authorityYear="1899" class="Gastropoda" family="Camaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica potanini" order="Stylommatophora" pageId="0" pageNumber="49" phylum="Mollusca" rank="species" species="potanini">
<emphasis italics="true" pageId="0" pageNumber="49">Laeocathaica potanini</emphasis>
</taxonomicName>
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
, 1899, HBUMM00633
<emphasis bold="true" pageId="0" pageNumber="49">A</emphasis>
protoconch
<emphasis bold="true" pageId="0" pageNumber="49">B</emphasis>
teleoconch
<emphasis bold="true" pageId="0" pageNumber="49">C, D</emphasis>
<taxonomicName lsidName="L. dityla" pageId="0" pageNumber="49" rank="species" species="dityla">
<emphasis italics="true" pageId="0" pageNumber="49">L. dityla</emphasis>
</taxonomicName>
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
, 1899, HBUMM00698
<emphasis bold="true" pageId="0" pageNumber="49">C</emphasis>
protoconch
<emphasis bold="true" pageId="0" pageNumber="49">D</emphasis>
teleoconch
<emphasis bold="true" pageId="0" pageNumber="49">E, F</emphasis>
<taxonomicName lsidName="L. qingchuanensis" pageId="0" pageNumber="49" rank="species" species="qingchuanensis">
<emphasis italics="true" pageId="0" pageNumber="49">L. qingchuanensis</emphasis>
</taxonomicName>
Wu, sp. nov., HBUMM01179a-spec.6, paratype
<emphasis bold="true" pageId="0" pageNumber="49">E</emphasis>
protoconch
<emphasis bold="true" pageId="0" pageNumber="49">F</emphasis>
teleoconch
<emphasis bold="true" pageId="0" pageNumber="49">G, H</emphasis>
<taxonomicName lsidName="L. christinae" pageId="0" pageNumber="49" rank="species" species="christinae">
<emphasis italics="true" pageId="0" pageNumber="49">L. christinae</emphasis>
</taxonomicName>
(H. Adams, 1870), HBUMM01251a
<emphasis bold="true" pageId="0" pageNumber="49">G</emphasis>
protoconch
<emphasis bold="true" pageId="0" pageNumber="49">H</emphasis>
teleoconch
<emphasis bold="true" pageId="0" pageNumber="49">I, J</emphasis>
<taxonomicName lsidName="L. carinifera" pageId="0" pageNumber="49" rank="species" species="carinifera">
<emphasis italics="true" pageId="0" pageNumber="49">L. carinifera</emphasis>
</taxonomicName>
(H. Adams, 1870), HBUMM04162
<emphasis bold="true" pageId="0" pageNumber="49">I</emphasis>
protoconch
<emphasis bold="true" pageId="0" pageNumber="49">J</emphasis>
teleoconch
<emphasis bold="true" pageId="0" pageNumber="49">K, L</emphasis>
<taxonomicName lsidName="L. polytyla" pageId="0" pageNumber="49" rank="species" species="polytyla">
<emphasis italics="true" pageId="0" pageNumber="49">L. polytyla</emphasis>
</taxonomicName>
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
, 1899, HBUMM05437
<emphasis bold="true" pageId="0" pageNumber="49">K</emphasis>
protoconch
<emphasis bold="true" pageId="0" pageNumber="49">L</emphasis>
teleoconch.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="49" type="diagnosis">
<paragraph pageId="0" pageNumber="49">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="49">
Protoconch with dense fine granules. Umbilicus very broad, ~ 1/2 of maximum diameter. Body whorl shouldered above periphery. On shoulder with a white band. Peristome continuous. Shell strongly glossy as in
<taxonomicName authorityName="Mollendorff" authorityYear="1899" class="Gastropoda" family="Camaenidae" genus="Stilpnodiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stilpnodiscus" order="Stylommatophora" pageId="0" pageNumber="49" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="49">Stilpnodiscus</emphasis>
</taxonomicName>
, almost transparent.
</paragraph>
<caption doi="10.3897/zookeys.1154.86237.figure46" httpUri="https://binary.pensoft.net/fig/827389" pageId="0" pageNumber="49" start="Figure 46" startId="F46">
<paragraph pageId="0" pageNumber="49">
<emphasis bold="true" pageId="0" pageNumber="49">Figure 46.</emphasis>
SEM micrographs
<emphasis bold="true" pageId="0" pageNumber="49">A, B</emphasis>
<taxonomicName authorityName="Moellendorff" authorityYear="1899" class="Gastropoda" family="Camaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica distinguenda" order="Stylommatophora" pageId="0" pageNumber="49" phylum="Mollusca" rank="species" species="distinguenda">
<emphasis italics="true" pageId="0" pageNumber="49">Laeocathaica distinguenda</emphasis>
</taxonomicName>
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
, 1899, HBUMM05479
<emphasis bold="true" pageId="0" pageNumber="49">A</emphasis>
protoconch
<emphasis bold="true" pageId="0" pageNumber="49">B</emphasis>
teleoconch
<emphasis bold="true" pageId="0" pageNumber="49">C, D</emphasis>
<taxonomicName lsidName="L. stenochone" pageId="0" pageNumber="49" rank="species" species="stenochone">
<emphasis italics="true" pageId="0" pageNumber="49">L. stenochone</emphasis>
</taxonomicName>
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
, 1899, HBUMM05495
<emphasis bold="true" pageId="0" pageNumber="49">C</emphasis>
protoconch
<emphasis bold="true" pageId="0" pageNumber="49">D</emphasis>
teleoconch
<emphasis bold="true" pageId="0" pageNumber="49">E, F</emphasis>
<taxonomicName lsidName="L. prionotropis" pageId="0" pageNumber="49" rank="species" species="prionotropis">
<emphasis italics="true" pageId="0" pageNumber="49">L. prionotropis</emphasis>
</taxonomicName>
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
, 1899, HBUMM05549
<emphasis bold="true" pageId="0" pageNumber="49">E</emphasis>
protoconch
<emphasis bold="true" pageId="0" pageNumber="49">F</emphasis>
teleoconch
<emphasis bold="true" pageId="0" pageNumber="49">G, H</emphasis>
<taxonomicName lsidName="L. amdoana" pageId="0" pageNumber="49" rank="species" species="amdoana">
<emphasis italics="true" pageId="0" pageNumber="49">L. amdoana</emphasis>
</taxonomicName>
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
, 1899, HBUMM05640
<emphasis bold="true" pageId="0" pageNumber="49">G</emphasis>
protoconch
<emphasis bold="true" pageId="0" pageNumber="49">H</emphasis>
teleoconch
<emphasis bold="true" pageId="0" pageNumber="49">I, J</emphasis>
<taxonomicName lsidName="L. tropidorhaphe" pageId="0" pageNumber="49" rank="species" species="tropidorhaphe">
<emphasis italics="true" pageId="0" pageNumber="49">L. tropidorhaphe</emphasis>
</taxonomicName>
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
, 1899, HBUMM05664
<emphasis bold="true" pageId="0" pageNumber="49">I</emphasis>
protoconch
<emphasis bold="true" pageId="0" pageNumber="49">J</emphasis>
teleoconch
<emphasis bold="true" pageId="0" pageNumber="49">K, L</emphasis>
<taxonomicName lsidName="L. cheni" pageId="0" pageNumber="49" rank="species" species="cheni">
<emphasis italics="true" pageId="0" pageNumber="49">L. cheni</emphasis>
</taxonomicName>
Wu, sp. nov., HBUMM08368, paratype, juvenile
<emphasis bold="true" pageId="0" pageNumber="49">K</emphasis>
protoconch
<emphasis bold="true" pageId="0" pageNumber="49">L</emphasis>
teleoconch.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="49" type="description">
<paragraph pageId="0" pageNumber="49">Description of shell.</paragraph>
<paragraph pageId="0" pageNumber="49">
Sinistral, depressed, thin but somewhat solid. Shell with 5-51/8 convex whorls. Suture impressed. Protoconch 11/2-15/8 whorls, densely with tiny granules (each ~ 10
<normalizedToken originalValue="µm">µm</normalizedToken>
long) which are obscured by erosion or weathering. Growth lines indistinct. Shell in pale brown, strongly glossy, almost transparent. Body whorl shouldered above periphery, with a white band on shoulder. Aperture oblique, round, abruptly descending in front. A white thickening within aperture and thickened callus forming a continuous peristome. Peristome expanded and slightly reflexed at lower part. Columella oblique. Umbilicus very broad, ~ 1/2 of maximum diameter. Protoconch visible through umbilicus.
</paragraph>
<caption doi="10.3897/zookeys.1154.86237.figure47" httpUri="https://binary.pensoft.net/fig/827390" pageId="0" pageNumber="49" start="Figure 47" startId="F47">
<paragraph pageId="0" pageNumber="49">
<emphasis bold="true" pageId="0" pageNumber="49">Figure 47.</emphasis>
SEM micrographs
<emphasis bold="true" pageId="0" pageNumber="49">A, B</emphasis>
<taxonomicName authorityName="Moellendorff" authorityYear="1899" class="Gastropoda" family="Camaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica odophora" order="Stylommatophora" pageId="0" pageNumber="49" phylum="Mollusca" rank="species" species="odophora">
<emphasis italics="true" pageId="0" pageNumber="49">Laeocathaica odophora</emphasis>
</taxonomicName>
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
, 1899, HBUMM08430
<emphasis bold="true" pageId="0" pageNumber="49">A</emphasis>
protoconch
<emphasis bold="true" pageId="0" pageNumber="49">B</emphasis>
teleoconch
<emphasis bold="true" pageId="0" pageNumber="49">C, D</emphasis>
<taxonomicName lsidName="L. dolani" pageId="0" pageNumber="49" rank="species" species="dolani">
<emphasis italics="true" pageId="0" pageNumber="49">L. dolani</emphasis>
</taxonomicName>
(Pilsbry, 1934), HBUMM08439
<emphasis bold="true" pageId="0" pageNumber="49">C</emphasis>
protoconch
<emphasis bold="true" pageId="0" pageNumber="49">D</emphasis>
teleoconch
<emphasis bold="true" pageId="0" pageNumber="49">E, F</emphasis>
<taxonomicName lsidName="L. nordsiecki" pageId="0" pageNumber="49" rank="species" species="nordsiecki">
<emphasis italics="true" pageId="0" pageNumber="49">L. nordsiecki</emphasis>
</taxonomicName>
Wu, sp. nov., HBUMM08446, paratype, subadult
<emphasis bold="true" pageId="0" pageNumber="49">E</emphasis>
protoconch
<emphasis bold="true" pageId="0" pageNumber="49">F</emphasis>
teleoconch
<emphasis bold="true" pageId="0" pageNumber="49">G, H</emphasis>
<taxonomicName lsidName="L. pewzowi" pageId="0" pageNumber="49" rank="species" species="pewzowi">
<emphasis italics="true" pageId="0" pageNumber="49">L. pewzowi</emphasis>
</taxonomicName>
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
, 1899, HBUMM08452
<emphasis bold="true" pageId="0" pageNumber="49">G</emphasis>
protoconch
<emphasis bold="true" pageId="0" pageNumber="49">H</emphasis>
teleoconch
<emphasis bold="true" pageId="0" pageNumber="49">I, J</emphasis>
<taxonomicName lsidName="L. carinalis" pageId="0" pageNumber="49" rank="species" species="carinalis">
<emphasis italics="true" pageId="0" pageNumber="49">L. carinalis</emphasis>
</taxonomicName>
Chen &amp; Zhang, 2004, HBUMM08453
<emphasis bold="true" pageId="0" pageNumber="49">I</emphasis>
protoconch
<emphasis bold="true" pageId="0" pageNumber="49">J</emphasis>
teleoconch
<emphasis bold="true" pageId="0" pageNumber="49">K, L</emphasis>
<taxonomicName lsidName="L. zhengpingliui" pageId="0" pageNumber="49" rank="species" species="zhengpingliui">
<emphasis italics="true" pageId="0" pageNumber="49">L. zhengpingliui</emphasis>
</taxonomicName>
Wu, sp. nov., HBUMM05553-spec.2, paratype, juvenile
<emphasis bold="true" pageId="0" pageNumber="49">K</emphasis>
protoconch
<emphasis bold="true" pageId="0" pageNumber="49">L</emphasis>
teleoconch.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="49" type="etymology">
<paragraph pageId="0" pageNumber="49">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="49">This cute new species is named in memory of Hartmut Nordsieck, a German malacologist who showed strong interest in the snails of the South Gansu Plateau and was a good friend of the first author.</paragraph>
<caption doi="10.3897/zookeys.1154.86237.figure48" httpUri="https://binary.pensoft.net/fig/827391" pageId="0" pageNumber="49" start="Figure 48" startId="F48">
<paragraph pageId="0" pageNumber="49">
<emphasis bold="true" pageId="0" pageNumber="49">Figure 48.</emphasis>
SEM micrographs
<emphasis bold="true" pageId="0" pageNumber="49">A, B</emphasis>
<taxonomicName authorityName="Wu" authorityYear="2023" class="Gastropoda" family="Camaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica qishilii" order="Stylommatophora" pageId="0" pageNumber="49" phylum="Mollusca" rank="species" species="qishilii">
<emphasis italics="true" pageId="0" pageNumber="49">Laeocathaica qishilii</emphasis>
</taxonomicName>
Wu, sp. nov., HBUMM08298, paratype
<emphasis bold="true" pageId="0" pageNumber="49">A</emphasis>
protoconch
<emphasis bold="true" pageId="0" pageNumber="49">B</emphasis>
teleoconch
<emphasis bold="true" pageId="0" pageNumber="49">C, D</emphasis>
<taxonomicName lsidName="L. qiminglii" pageId="0" pageNumber="49" rank="species" species="qiminglii">
<emphasis italics="true" pageId="0" pageNumber="49">L. qiminglii</emphasis>
</taxonomicName>
Wu, sp. nov., HBUMM08448, paratype
<emphasis bold="true" pageId="0" pageNumber="49">C</emphasis>
protoconch
<emphasis bold="true" pageId="0" pageNumber="49">D</emphasis>
teleoconch
<emphasis bold="true" pageId="0" pageNumber="49">E, F</emphasis>
<taxonomicName lsidName="L. phaeomphala" pageId="0" pageNumber="49" rank="species" species="phaeomphala">
<emphasis italics="true" pageId="0" pageNumber="49">L. phaeomphala</emphasis>
</taxonomicName>
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
, 1899, CZG202008-w3, subadult
<emphasis bold="true" pageId="0" pageNumber="49">E</emphasis>
protoconch
<emphasis bold="true" pageId="0" pageNumber="49">F</emphasis>
teleoconch
<emphasis bold="true" pageId="0" pageNumber="49">G, H</emphasis>
<taxonomicName lsidName="L. filippina" pageId="0" pageNumber="49" rank="species" species="filippina">
<emphasis italics="true" pageId="0" pageNumber="49">L. filippina</emphasis>
</taxonomicName>
(Heude, 1882), HBUMM04166
<emphasis bold="true" pageId="0" pageNumber="49">G</emphasis>
protoconch
<emphasis bold="true" pageId="0" pageNumber="49">H</emphasis>
teleoconch
<emphasis bold="true" pageId="0" pageNumber="49">I, J</emphasis>
<taxonomicName lsidName="L. parapolytyla" pageId="0" pageNumber="49" rank="species" species="parapolytyla">
<emphasis italics="true" pageId="0" pageNumber="49">L. parapolytyla</emphasis>
</taxonomicName>
Wu, sp. nov., HBUMM06640-spec.9, paratype
<emphasis bold="true" pageId="0" pageNumber="49">I</emphasis>
protoconch
<emphasis bold="true" pageId="0" pageNumber="49">J</emphasis>
teleoconch.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="49" type="ecology">
<paragraph pageId="0" pageNumber="49">Ecology.</paragraph>
<paragraph pageId="0" pageNumber="49">This species was found on bare earth with a few broken rocks.</paragraph>
<caption doi="10.3897/zookeys.1154.86237.figure49" httpUri="https://binary.pensoft.net/fig/827392" pageId="0" pageNumber="49" start="Figure 49" startId="F49">
<paragraph pageId="0" pageNumber="49">
<emphasis bold="true" pageId="0" pageNumber="49">Figure 49.</emphasis>
Habitats (excluding
<emphasis bold="true" pageId="0" pageNumber="49">D</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="49">A, B</emphasis>
<taxonomicName authorityName="Wu" authorityYear="2023" class="Gastropoda" family="Camaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica qishilii" order="Stylommatophora" pageId="0" pageNumber="49" phylum="Mollusca" rank="species" species="qishilii">
<emphasis italics="true" pageId="0" pageNumber="49">Laeocathaica qishilii</emphasis>
</taxonomicName>
Wu, sp. nov., border of Jiuzhaigou County and Wen County
<emphasis bold="true" pageId="0" pageNumber="49">C</emphasis>
<taxonomicName lsidName="L. nordsiecki" pageId="0" pageNumber="49" rank="species" species="nordsiecki">
<emphasis italics="true" pageId="0" pageNumber="49">L. nordsiecki</emphasis>
</taxonomicName>
Wu, sp. nov., Guoyuanxiang, Jiuzhaigou
<emphasis bold="true" pageId="0" pageNumber="49">D</emphasis>
<taxonomicName lsidName="L. qiminglii" pageId="0" pageNumber="49" rank="species" species="qiminglii">
<emphasis italics="true" pageId="0" pageNumber="49">L. qiminglii</emphasis>
</taxonomicName>
Wu, sp. nov., an indoor photograph
<emphasis bold="true" pageId="0" pageNumber="49">E</emphasis>
<taxonomicName lsidName="L. qingchuanensis" pageId="0" pageNumber="49" rank="species" species="qingchuanensis">
<emphasis italics="true" pageId="0" pageNumber="49">L. qingchuanensis</emphasis>
</taxonomicName>
Wu, sp. nov., Dagou Nature Reserve, Qingchuan County
<emphasis bold="true" pageId="0" pageNumber="49">F</emphasis>
<taxonomicName lsidName="L. cheni" pageId="0" pageNumber="49" rank="species" species="cheni">
<emphasis italics="true" pageId="0" pageNumber="49">L. cheni</emphasis>
</taxonomicName>
Wu, sp. nov., Hengdan, Wenxian.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="49" type="distribution">
<paragraph pageId="0" pageNumber="49">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="49">Only known from two localities where the types were found.</paragraph>
<caption doi="10.3897/zookeys.1154.86237.figure50" httpUri="https://binary.pensoft.net/fig/827393" pageId="0" pageNumber="49" start="Figure 50" startId="F50">
<paragraph pageId="0" pageNumber="49">
<emphasis bold="true" pageId="0" pageNumber="49">Figure 50.</emphasis>
Scatter plots of canonical variate 1 against canonical variate 2 (Canonical Variate Analysis), showing the shell morphological relationships among
<emphasis bold="true" pageId="0" pageNumber="49">A</emphasis>
<taxonomicName authorityName="Moellendorff" authorityYear="1899" class="Gastropoda" family="Camaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica distinguenda" order="Stylommatophora" pageId="0" pageNumber="49" phylum="Mollusca" rank="species" species="distinguenda">
<emphasis italics="true" pageId="0" pageNumber="49">Laeocathaica distinguenda</emphasis>
</taxonomicName>
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
, 1899 (green dots; a, average shape),
<taxonomicName lsidName="L. amdoana" pageId="0" pageNumber="49" rank="species" species="amdoana">
<emphasis italics="true" pageId="0" pageNumber="49">L. amdoana</emphasis>
</taxonomicName>
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
, 1899 (blue dots; b, average shape) and
<taxonomicName lsidName="L. tropidorhaphe" pageId="0" pageNumber="49" rank="species" species="tropidorhaphe">
<emphasis italics="true" pageId="0" pageNumber="49">L. tropidorhaphe</emphasis>
</taxonomicName>
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
, 1899 (orange dots; c, average shape)
<emphasis bold="true" pageId="0" pageNumber="49">B</emphasis>
<taxonomicName lsidName="L. qingchuanensis" pageId="0" pageNumber="49" rank="species" species="qingchuanensis">
<emphasis italics="true" pageId="0" pageNumber="49">L. qingchuanensis</emphasis>
</taxonomicName>
Wu, sp. nov. (green dots; a, average shape),
<taxonomicName lsidName="L. stenochone" pageId="0" pageNumber="49" rank="species" species="stenochone">
<emphasis italics="true" pageId="0" pageNumber="49">L. stenochone</emphasis>
</taxonomicName>
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
, 1899 (orange dots; b, average shape) and
<taxonomicName lsidName="L. carinifera" pageId="0" pageNumber="49" rank="species" species="carinifera">
<emphasis italics="true" pageId="0" pageNumber="49">L. carinifera</emphasis>
</taxonomicName>
(H. Adams, 1870) (blue dots; c, average shape). Encircled dots: 1, SMF 8959, lectotype; 2, HBUMM05436-spec.1; 3, SMF 8952, lectotype; 4, SMF 9074, lectotype; 5, paratype of
<taxonomicName lsidName="L. dangchangensis" pageId="0" pageNumber="49" rank="species" species="dangchangensis">
<emphasis italics="true" pageId="0" pageNumber="49">L. dangchangensis</emphasis>
</taxonomicName>
Chen &amp; Zhang, 2004; 6, SMF 9071, lectotype; 7, syntype, NHMUK 1870.7.16.7. Data for 5 from fig. 7D, 6 and 7 from fig. 12 in
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.1086.77408" author="Pall-Gergely, B" journalOrPublisher="ZooKeys" pageId="0" pageNumber="49" pagination="33 - 76" refId="B46" refString="Pall-Gergely, B, Hunyadi, A, Meng, K, Fekete, J, 2022. A review of the genus Laeocathaica Moellendorff, 1899 (Gastropoda, Pulmonata, Camaenidae). ZooKeys 1086: 33 - 76, DOI: https://doi.org/10.3897/zookeys.1086.77408" title="A review of the genus Laeocathaica Moellendorff, 1899 (Gastropoda, Pulmonata, Camaenidae)." url="https://doi.org/10.3897/zookeys.1086.77408" volume="1086" year="2022">
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
et al. 2022
</bibRefCitation>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="49" type="remarks">
<paragraph pageId="0" pageNumber="49">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="49">
The new species is the smallest species in
<taxonomicName authorityName="Moellendorff" authorityYear="1899" class="Gastropoda" family="Camaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica" order="Stylommatophora" pageId="0" pageNumber="49" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="49">Laeocathaica</emphasis>
</taxonomicName>
, where it is provisionally placed due to its chirality, granules on the protoconch and the similarity of the shell to that of
<taxonomicName authorityName="Moellendorff" authorityYear="1899" class="Gastropoda" family="Camaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica dityla" order="Stylommatophora" pageId="0" pageNumber="49" phylum="Mollusca" rank="species" species="dityla">
<emphasis italics="true" pageId="0" pageNumber="49">Laeocathaica dityla</emphasis>
</taxonomicName>
. The very glossy shell also makes this species distinctive, reminiscent of the genus
<taxonomicName authorityName="Mollendorff" authorityYear="1899" class="Gastropoda" family="Camaenidae" genus="Stilpnodiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stilpnodiscus" order="Stylommatophora" pageId="0" pageNumber="49" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="49">Stilpnodiscus</emphasis>
</taxonomicName>
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
, 1899, which appears in the phylogram (Fig.
<figureCitation captionStart="Figure 51" captionStartId="F51" captionText="Figure 51. Bayesian phylogram of camaenid species (for detail see in Table 1) based on the concatenated partial mitochondrial 16 S and partial ITS 2 sequences. The tree is rooted with Helix pomatia. Numbers near nodes indicating Bayesian posterior probabilities and Maximum-likelihood bootstrap values are given as BPP BP. Black or orange part shows the topology where the result yielded by using Bayesian-Inference method agrees with that by using Maximum-likelihood method or not, respectively. An asterisk indicates the branch is exactly one third shortened in length. Scale bar is for substitutions per site." figureDoi="10.3897/zookeys.1154.86237.figure51" httpUri="https://binary.pensoft.net/fig/827394" pageId="0" pageNumber="49">51</figureCitation>
) as the sister group of all the
<taxonomicName authorityName="Moellendorff" authorityYear="1899" class="Gastropoda" family="Camaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica" order="Stylommatophora" pageId="0" pageNumber="49" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="49">Laeocathaica</emphasis>
</taxonomicName>
species. Undoubtedly, final generic assignment of this species will depend on further anatomical information and molecular studies.
</paragraph>
<caption doi="10.3897/zookeys.1154.86237.figure51" httpUri="https://binary.pensoft.net/fig/827394" pageId="0" pageNumber="49" start="Figure 51" startId="F51">
<paragraph pageId="0" pageNumber="49">
<emphasis bold="true" pageId="0" pageNumber="49">Figure 51.</emphasis>
Bayesian phylogram of camaenid species (for detail see in Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Some character states of the terminal genitalia in Laeocathaica Moellendorff, 1899." httpUri="http://table.plazi.org/id/9AAE1A30B00EA431759A83DD4FF4F907" pageId="0" pageNumber="49" tableUuid="9AAE1A30B00EA431759A83DD4FF4F907">1</tableCitation>
) based on the concatenated partial mitochondrial 16S and partial ITS2 sequences. The tree is rooted with
<taxonomicName authorityName="Linnaeus" authorityYear="1758" class="Gastropoda" family="Camaenidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix pomatia" order="Stylommatophora" pageId="0" pageNumber="49" phylum="Mollusca" rank="species" species="pomatia">
<emphasis italics="true" pageId="0" pageNumber="49">Helix pomatia</emphasis>
</taxonomicName>
. Numbers near nodes indicating Bayesian posterior probabilities and Maximum-likelihood bootstrap values are given as BPP BP. Black or orange part shows the topology where the result yielded by using Bayesian-Inference method agrees with that by using Maximum-likelihood method or not, respectively. An asterisk indicates the branch is exactly one third shortened in length. Scale bar is for substitutions per site.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>