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<document id="060118CCA039C96F27953343E25E6AC9" ID-DOI="10.5281/zenodo.183391" ID-GBIF-Dataset="46dd9791-f59e-4019-b1cc-30f1ddf8629f" ID-ISSN="1175-5326" ID-Zenodo-Dep="183391" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459905098680" checkinUser="plazi" docAuthor="Guinot, Danièle" docDate="2008" docId="20528B6EFFD0FFA7D8E5F93EFE6DF9B6" docLanguage="en" docName="zt01850p026.pdf" docOrigin="Zootaxa 1850" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Acanthodromiinae" docType="treatment" docVersion="9" lastPageNumber="8" masterDocId="DC6BF316FFD5FFA0D872FFE9FFCFFFE4" masterDocTitle="A re-evaluation of the Dynomenidae Ortmann, 1892 (Crustacea, Decapoda, Brachyura, Podotremata), with the recognition of four subfamilies" masterLastPageNumber="26" masterPageNumber="1" pageNumber="6" updateTime="1698228993237" updateUser="plazi">
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<mods:title id="B1F5BA3EC32105F4DB9D9A82D4A66637">A re-evaluation of the Dynomenidae Ortmann, 1892 (Crustacea, Decapoda, Brachyura, Podotremata), with the recognition of four subfamilies</mods:title>
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<treatment id="20528B6EFFD0FFA7D8E5F93EFE6DF9B6" ID-DOI="http://doi.org/10.5281/zenodo.6234451" ID-GBIF-Taxon="119373364" ID-Zenodo-Dep="6234451" LSID="urn:lsid:plazi:treatment:20528B6EFFD0FFA7D8E5F93EFE6DF9B6" httpUri="http://treatment.plazi.org/id/20528B6EFFD0FFA7D8E5F93EFE6DF9B6" lastPageId="7" lastPageNumber="8" pageId="5" pageNumber="6">
<subSubSection id="E0E169F3FFD0FFA5D8E5F93EFE53F8F0" pageId="5" pageNumber="6" type="nomenclature">
<paragraph id="A8443A78FFD0FFA5D8E5F93EFDC4F915" blockId="5.[151,523,1751,1812]" box="[151,523,1751,1777]" pageId="5" pageNumber="6">
<heading id="F30C8D14FFD0FFA5D8E5F93EFDC4F915" bold="true" box="[151,523,1751,1777]" fontSize="11" level="1" pageId="5" pageNumber="6" reason="1">
<emphasis id="9A8FE66AFFD0FFA5D8E5F93EFDC4F915" bold="true" box="[151,523,1751,1777]" pageId="5" pageNumber="6">
<taxonomicName id="6FFB41FBFFD0FFA5D8E5F93EFE4EF915" ID-CoL="8479G" box="[151,385,1751,1777]" class="Malacostraca" family="Dynomenidae" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="subFamily" subFamily="Acanthodromiinae">Acanthodromiinae</taxonomicName>
n. subfam.
</emphasis>
</heading>
</paragraph>
<paragraph id="A8443A78FFD0FFA5D8E5F913FE53F8F0" blockId="5.[151,523,1751,1812]" box="[151,412,1786,1812]" pageId="5" pageNumber="6">
(
<figureCitation id="30C026FDFFD0FFA5D8EDF913FEC5F8F0" box="[159,266,1786,1812]" captionStart="FIGURE 1" captionStartId="4.[151,255,1602,1626]" captionTargetBox="[283,1303,425,1571]" captionTargetId="figure@4.[283,1304,425,1574]" captionTargetPageId="4" captionText="FIGURE 1. Representative species of the four subfamilies of Dynomenidae, overall view. A, Acanthodromiinae n. subfam.: Acanthodromia margarita (Alcock, 1899), Balicasag Island, Panglao, Bohol, Visayas, Philippines (ZRC) (see McLay &amp; Ng 2005: 18); B, Paradynomeninae n. subfam.: Paradynomene tuberculata Sakai, 1963, Balicasag Island, Panglao, Bohol, Visayas, Philippines (ZRC) (see McLay &amp; Ng 2004: 4); C, Metadynomeninae n. subfam.: Metadynomene tanensis (Yokoya, 1933) with its undulating tomentum, Balicasag Island, Panglao, Bohol, Visayas, Philippines (ZRC) (see McLay &amp; Ng 2005: 25); D, Dynomeninae Ortmann, 1892: Hirsutodynomene vespertilio McLay &amp; Ng, 2005, Balicasag Island, Panglao, Bohol, Visayas, Philippines (ZRC) (see McLay &amp; Ng 2005: 21). E, Paradynomene tuberculata Sakai, 1963, New Caledonia, frontal view showing the characteristic “ face ” (ZRC). (All photographs courtesy of P. K. L. Ng)." httpUri="https://zenodo.org/record/183392/files/figure.png" pageId="5" pageNumber="6">Figures 1</figureCitation>
A, 2, 5OP)
</paragraph>
</subSubSection>
<subSubSection id="E0E169F3FFD0FFA5D8E5F8A3FCCCF865" pageId="5" pageNumber="6" type="reference_group">
<paragraph id="A8443A78FFD0FFA5D8E5F8A3FCCCF865" blockId="5.[151,1436,1866,1921]" pageId="5" pageNumber="6">
<taxonomicName id="6FFB41FBFFD0FFA5D8E5F8A3FEE3F885" ID-CoL="847F5" box="[151,300,1866,1889]" class="Malacostraca" family="Dynomenidae" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="subFamily" subFamily="Dynomeninae">Dynomeninae</taxonomicName>
<bibRefCitation id="CC6A4789FFD0FFA5D947F8A3FD7BF885" author="Milne-Edwards" box="[309,692,1866,1889]" pageId="5" pageNumber="25" refString="Milne-Edwards, A. &amp; Bouvier, E. - L. (1899) Crustaces Decapodes provenant des Campagnes de l'Hirondelle (Supplement) et de la Princesse Alice (1891 - 1897). Brachyures et Anomoures. Resultats des Campagnes scientifiques du Prince Albert 1 er de Monaco, 13, 1 - 106." type="journal article" year="1899">A. Milne-Edwards &amp; Bouvier 1899</bibRefCitation>
<emphasis id="9A8FE66AFFD0FFA5DACFF8A3FCEEF885" box="[701,801,1866,1889]" italics="true" pageId="5" pageNumber="6">pro parte</emphasis>
: 9;
<bibRefCitation id="CC6A4789FFD0FFA5DB3CF8A3FC11F885" author="Alcock" box="[846,990,1866,1889]" pageId="5" pageNumber="23" refString="Alcock, A. (1900) Materials for a Carcinological Fauna of India. N ° 5. The Brachyura Primigenia or Dromiacea. Journal of the Asiatic Society of Bengal, 68 (2) 3 [1899], 123 - 169." type="journal article" year="1900">Alcock, 1900</bibRefCitation>
<emphasis id="9A8FE66AFFD0FFA5DB95F8A3FB84F885" box="[999,1099,1866,1889]" italics="true" pageId="5" pageNumber="6">pro parte</emphasis>
: 127, 133; 1901
<emphasis id="9A8FE66AFFD0FFA5DD74F8A3FAA4F885" box="[1286,1387,1866,1889]" italics="true" pageId="5" pageNumber="6">pro parte</emphasis>
: 34, 74;
<bibRefCitation id="CC6A4789FFD0FFA5D89DF883FDA5F865" author="Milne-Edwards" box="[239,618,1898,1921]" pageId="5" pageNumber="25" refString="Milne-Edwards, A. &amp; Bouvier, E. - L. (1902) Reports on the results of dredging, under the supervision of Alexander Agassiz, in the Gulf of Mexico (1877 - 78), in the Caribbean Sea (1878 - 79), and along the Atlantic Coast of the United States (1880), by the U. S. Coast Survey Steamer &quot; Blake &quot;. XXXIX. Les Dromiaces et Oxystomes. Memoirs of the Museum of Comparative Zoology at Harvard College, 27 (1), 1 - 127." type="journal article" year="1902">A. Milne-Edwards &amp; Bouvier 1902</bibRefCitation>
<emphasis id="9A8FE66AFFD0FFA5DA02F883FD1CF865" box="[624,723,1898,1921]" italics="true" pageId="5" pageNumber="6">pro parte</emphasis>
: 22.
</paragraph>
</subSubSection>
<subSubSection id="E0E169F3FFD0FFA5D8E5F85BFE97F810" pageId="5" pageNumber="6" type="materials_examined">
<paragraph id="A8443A78FFD0FFA5D8E5F85BFE97F810" blockId="5.[151,1436,1970,2036]" pageId="5" pageNumber="6">
<emphasis id="9A8FE66AFFD0FFA5D8E5F85BFEE7F828" bold="true" box="[151,296,1970,1996]" pageId="5" pageNumber="6">
<typeStatus id="774084DAFFD0FFA5D8E5F85BFF1BF828" box="[151,212,1970,1996]" pageId="5" pageNumber="6">Type</typeStatus>
genus
</emphasis>
.
<taxonomicName id="6FFB41FBFFD0FFA5D948F85AFCE8F828" authority="A. Milne-Edwards, 1880" authorityName="A. Milne-Edwards" authorityYear="1880" box="[314,807,1970,1996]" class="Malacostraca" family="Dynomenidae" genus="Acanthodromia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="9A8FE66AFFD0FFA5D948F85AFE20F828" box="[314,495,1971,1996]" italics="true" pageId="5" pageNumber="6">Acanthodromia</emphasis>
<bibRefCitation id="CC6A4789FFD0FFA5D989F85BFCE8F828" author="Milne-Edwards" box="[507,807,1970,1996]" pageId="5" pageNumber="25" refString="Milne-Edwards, A. (1880) Etudes preliminaires sur les Crustaces, 1 ere Partie. Reports on the Results of Dredging under the Supervision of Alexander Agassiz, in the Gulf of Mexico, and in the Caribbean Sea, 1877, ' 78, ' 79, by the U. S. Coast Survey Steamer &quot; Blake &quot;, Lieut. - Commander C. D. Sigsbee, U. S. N., and Commander J. R. Bartlett, U. S. N., Commanding, VIII. Bulletin of the Museum of Comparative Zoology at Harvard College, 8 (1), 1 - 68." type="journal article" year="1880">A. Milne-Edwards, 1880</bibRefCitation>
</taxonomicName>
(
<typeStatus id="774084DAFFD0FFA5DB48F85BFCA1F828" box="[826,878,1970,1996]" pageId="5" pageNumber="6">type</typeStatus>
species by monotypy:
<taxonomicName id="6FFB41FBFFD0FFA5DCFFF85AFAD6F828" box="[1165,1305,1971,1996]" class="Malacostraca" family="Dynomenidae" genus="Acanthodromia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="erinacea">
<emphasis id="9A8FE66AFFD0FFA5DCFFF85AFAD6F828" box="[1165,1305,1971,1996]" italics="true" pageId="5" pageNumber="6">A. erinacea</emphasis>
</taxonomicName>
A. Milne-
<bibRefCitation id="CC6A4789FFD0FFA5D8E5F833FE84F810" author="Edwards" box="[151,331,2010,2036]" pageId="5" pageNumber="25" refString="Milne-Edwards, A. (1880) Etudes preliminaires sur les Crustaces, 1 ere Partie. Reports on the Results of Dredging under the Supervision of Alexander Agassiz, in the Gulf of Mexico, and in the Caribbean Sea, 1877, ' 78, ' 79, by the U. S. Coast Survey Steamer &quot; Blake &quot;, Lieut. - Commander C. D. Sigsbee, U. S. N., and Commander J. R. Bartlett, U. S. N., Commanding, VIII. Bulletin of the Museum of Comparative Zoology at Harvard College, 8 (1), 1 - 68." type="journal article" year="1880">Edwards, 1880</bibRefCitation>
).
</paragraph>
</subSubSection>
<caption id="FC846AF0FFD3FFA6D8E5FA75FC5BF970" httpUri="https://zenodo.org/record/183393/files/figure.png" pageId="6" pageNumber="7" targetBox="[281,1304,184,1402]" targetPageId="6">
<paragraph id="A8443A78FFD3FFA6D8E5FA75FC5BF970" blockId="6.[151,1436,1436,1684]" pageId="6" pageNumber="7">
<emphasis id="9A8FE66AFFD3FFA6D8E5FA75FED4FA50" bold="true" box="[151,283,1436,1460]" pageId="6" pageNumber="7">FIGURE 2.</emphasis>
<taxonomicName id="6FFB41FBFFD3FFA6D951FA74FD12FA50" authority="A. Milne-Edwards, 1880" authorityName="A. Milne-Edwards" authorityYear="1880" box="[291,733,1437,1460]" class="Malacostraca" family="Dynomenidae" genus="Acanthodromia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="9A8FE66AFFD3FFA6D951FA74FE07FA50" box="[291,456,1437,1460]" italics="true" pageId="6" pageNumber="7">Acanthodromia</emphasis>
A. Milne-Edwards, 1880
</taxonomicName>
(
<taxonomicName id="6FFB41FBFFD3FFA6DA9FFA74FC7AFA50" box="[749,949,1437,1460]" class="Malacostraca" family="Dynomenidae" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="subFamily" subFamily="Acanthodromiinae">Acanthodromiinae</taxonomicName>
<emphasis id="9A8FE66AFFD3FFA6DBCFFA75FBF8FA50" bold="true" box="[957,1079,1436,1460]" pageId="6" pageNumber="7">n. subfam.</emphasis>
).
<emphasis id="9A8FE66AFFD3FFA6DC3CFA75FB4DFA50" bold="true" box="[1102,1154,1436,1460]" pageId="6" pageNumber="7">AC</emphasis>
,
<taxonomicName id="6FFB41FBFFD3FFA6DCE3FA74FE6AFA30" authority="A. Milne-Edwards, 1880" authorityName="A. Milne-Edwards" authorityYear="1880" class="Malacostraca" family="Dynomenidae" genus="Acanthodromia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="erinacea">
<emphasis id="9A8FE66AFFD3FFA6DCE3FA74FA53FA50" box="[1169,1436,1437,1460]" italics="true" pageId="6" pageNumber="7">Acanthodromia erinacea</emphasis>
A. Milne-Edwards, 1880
</taxonomicName>
, ovigerous female, 14.7 x 12 mm, west of Puerto Rico (USNM 124263).
<emphasis id="9A8FE66AFFD3FFA6DCCDFA55FB1DFA30" bold="true" box="[1215,1234,1468,1492]" pageId="6" pageNumber="7">A</emphasis>
, thoracic sternum;
<emphasis id="9A8FE66AFFD3FFA6D8E5FA35FF66FA10" bold="true" box="[151,169,1500,1524]" pageId="6" pageNumber="7">B</emphasis>
, ventral surface without abdomen;
<emphasis id="9A8FE66AFFD3FFA6DA55FA35FDF5FA10" bold="true" box="[551,570,1500,1524]" pageId="6" pageNumber="7">C</emphasis>
, abdomen.
<emphasis id="9A8FE66AFFD3FFA6DACAFA35FD04FA10" bold="true" box="[696,715,1500,1524]" pageId="6" pageNumber="7">D</emphasis>
,
<taxonomicName id="6FFB41FBFFD3FFA6DAA8FA34FB52FA10" authority="Alcock, 1899" authorityName="Alcock" authorityYear="1899" box="[730,1181,1501,1524]" class="Malacostraca" family="Dynomenidae" genus="Acanthodromia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="margarita">
<emphasis id="9A8FE66AFFD3FFA6DAA8FA34FC3EFA10" box="[730,1009,1501,1524]" italics="true" pageId="6" pageNumber="7">Acanthodromia margarita</emphasis>
(Alcock, 1899)
</taxonomicName>
, male, 11.6 x 12.8 mm, Balicasag Island, Panglao, Bohol, Visayas, Philippines, McLay &amp; Ng det. (MNHN-B28812) [The abdominal sutures are erroneously too much pronouced in these figures since there is a fusion of abdominal somites 36]. cx1cx5, coxae of P1P5; d5d6, sterno-coxal depressions 5 and 6; g, female gonopore; 26, sternites 26 (1 and 2 fused into shield; 3, individualised thoracic sternite 3; 3/4, limit between thoracic sternites 3 and 4; 4/55/6, thoracic sternal sutures 4/55/6 (indistinct); 7/8, distinct thoracic sternal suture 7/8. Scale bars: 2.5 mm.
</paragraph>
</caption>
<subSubSection id="E0E169F3FFD3FFA7D8B4F92BFD8FFB0E" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="diagnosis">
<paragraph id="A8443A78FFD3FFA7D8B4F92BFD8FFB0E" blockId="6.[151,1437,1730,2036]" lastBlockId="7.[151,1437,152,1618]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<emphasis id="9A8FE66AFFD3FFA6D8B4F92BFEF0F938" bold="true" box="[198,319,1730,1756]" pageId="6" pageNumber="7">Diagnosis</emphasis>
. Carapace longer than wide, oblong; dorsal surface convex, ornamented with close-set spines. Cervical, branchial grooves not well visible, branchiocardiac groove crescent- shaped. Anterolateral margins poorly defined, joining corners of buccal cavity, obscured by numerous spines. Posterior margin markedly concave. Frontal margin forming narrow projection; supraorbital margin oblique, continuous above orbits, eave-like, rimmed, not notched, spinous, prolonged in straight groove delineating subhepatic region; infraorbital margin concave, ornamented with spines. Orbits obliquely located, elongated, clearly visible from dorsal view; eyestalks short. Antenna with urinal article extended transversely, not medially beaked; second article with firmly fixed exopod. Proepistome short. Epistome narrow. Anteroventrally, front, inflated subhepatic pterygostomial portion forming together with merus of mxp3 a weak “face”. Anterior border of endostome forming raised wall, laterally notched by exhalant orifices. Mxp3 operculiform, angled; basis separated from ischium by nearly complete suture; ischium narrow at base, merus not extended laterally, narrow. Pleural line as wide, decalcified zone; branchiostegite of normal texture. Thoracic sternum extremely narrow, entirely (apart from sternites 1, 2) covered by male abdomen, abdominal margins close to P2, P3 coxae. Sternites 1, 2 fused into small, narrow shield, inserted between bases of mxp3; sternite 3 distinct but short, expanded laterally, delimited posteriorly by convex crest, corresponding to suture 3/4; sternites 38 not exposed laterally when abdomen closed; sternite 8 tilted. Sterno-coxal depressions very deep. Female sutures 7/8 ending well apart, along internal border of P3 sterno-coxal depression; spermathecal aperture small, slightly behind level of female gonopore. Sterno-abdominal depression narrow, deep, with steep sides, medially a flat floor. Abdomen broad, curved, extending over entire sternum (sternites 38) and reaching mxp3; first somite dorsal, with same concave curvature as posterior margin of carapace in which it is inserted; subsequent somites narrow, subequal in width; telson long, broadly triangular, slightly wider at base; somite 4 with a prominent, pearlshaped double tubercule. Abdominal somites 36 fused in males, probably also in females; sutures obscured by spines, only partially visible; suture between somites 5, 6 absent in males, more clearly visible in females (at least in
<taxonomicName id="6FFB41FBFFD2FFA7D968FD19FD82FCED" box="[282,589,752,777]" class="Malacostraca" family="Dynomenidae" genus="Acanthodromia" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="margarita">
<emphasis id="9A8FE66AFFD2FFA7D968FD19FD82FCED" box="[282,589,752,777]" italics="true" pageId="7" pageNumber="8">Acanthodromia margarita</emphasis>
</taxonomicName>
); in females somites 35 each with lateral portions produced as flanges overlapping following somites; flange of somite 6 more pronounced, raised. Male uropod small, showing as narrow transverse plate, slightly mobile. Abdomen tightly locked in both sexes by mechanism involving coxae of 4 thoracopods: coxa of mxp3 with spinules overhanging posterior part of telson; coxae of P1P3 with several spinules or granules overhanging telson at P1, P2 levels, somite 6 with uropods, and somite 5 at P3 level (in
<taxonomicName id="6FFB41FBFFD2FFA7D940FC51FDA6FC35" box="[306,617,952,977]" class="Malacostraca" family="Dynomenidae" genus="Acanthodromia" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="margarita">
<emphasis id="9A8FE66AFFD2FFA7D940FC51FDA6FC35" box="[306,617,952,977]" italics="true" pageId="7" pageNumber="8">Acanthodromia margarita</emphasis>
</taxonomicName>
all spinules better developed in females than in males). Chelipeds equal, more robust than P2; dactylus strongly curved downwards; fixed finger almost straight; both fingers spoon-tipped. P2P4 ornamented with spines; dactyli curved, bearing numerous spines; 4 or 5 spines on inferior margin. P5 conspicuously reduced, sexually dimorphic; basis-ischium fused to merus, basis-ischiummerus proportionally thicker than distal articles; dactylus rudimentary ending in subchelate mechanism, obsolete in males, more noticeable in females. P5 coxa of males modified, extended to enclose penis. Pl1 vestigial in females. Vestigial pleopods on somites
<quantity id="6F03979DFFD2FFA7DAF2FB41FD06FB26" box="[640,713,1192,1218]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" metricValueMax="1.27" metricValueMin="0.762" pageId="7" pageNumber="8" unit="in" value="4.0" valueMax="5.0" valueMin="3.0">35 in</quantity>
males. Gonopod 1 stout, forming half-rolled tube. G2 needlelike, with row of small distal spines.
</paragraph>
</subSubSection>
<subSubSection id="E0E169F3FFD2FFA7D8B4FB1EFE6DF9B6" pageId="7" pageNumber="8" type="discussion">
<paragraph id="A8443A78FFD2FFA7D8B4FB1EFE6DF9B6" blockId="7.[151,1437,152,1618]" pageId="7" pageNumber="8">
<emphasis id="9A8FE66AFFD2FFA7D8B4FB1EFEF7FAF5" bold="true" box="[198,312,1271,1297]" pageId="7" pageNumber="8">Remarks</emphasis>
. The subfamily is monotypic, and
<taxonomicName id="6FFB41FBFFD2FFA7DAADFB11FC5BFAF5" box="[735,916,1272,1297]" class="Malacostraca" family="Dynomenidae" genus="Acanthodromia" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="9A8FE66AFFD2FFA7DAADFB11FC5BFAF5" box="[735,916,1272,1297]" italics="true" pageId="7" pageNumber="8">Acanthodromia</emphasis>
</taxonomicName>
comprises only two species (
<tableCitation id="E5790FC3FFD2FFA7DC8AFB11FA9CFAF6" box="[1272,1363,1272,1298]" captionStart="TABLE 1" captionStartId="1.[151,239,391,415]" captionText="TABLE 1. List of extant genera and species currently placed in the family Dynomenidae Ortmann, 1892 (after Ng et al. 2008: 37). Generic names in square parentheses are the genera in which the species were originally described" pageId="7" pageNumber="8">Table 1</tableCitation>
). The morphology is masked by the spines which cover the whole body, so the precise outline and the grooves on the dorsal surface of carapace are not evident, which implies that potentially related fossil taxa are not easily assigned to this subfamily. The initial assignment of
<taxonomicName id="6FFB41FBFFD2FFA7DB6EFA99FC1DFA6D" box="[796,978,1392,1417]" class="Malacostraca" family="Dynomenidae" genus="Acanthodromia" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="9A8FE66AFFD2FFA7DB6EFA99FC1DFA6D" box="[796,978,1392,1417]" italics="true" pageId="7" pageNumber="8">Acanthodromia</emphasis>
</taxonomicName>
to the
<taxonomicName id="6FFB41FBFFD2FFA7DC5EFA99FB66FA6E" box="[1068,1193,1392,1418]" class="Malacostraca" family="Dromiidae" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="family">Dromiidae</taxonomicName>
(
<bibRefCitation id="CC6A4789FFD2FFA7DCCCFA99FF1AFA56" author="Milne-Edwards" pageId="7" pageNumber="25" refString="Milne-Edwards, A. (1880) Etudes preliminaires sur les Crustaces, 1 ere Partie. Reports on the Results of Dredging under the Supervision of Alexander Agassiz, in the Gulf of Mexico, and in the Caribbean Sea, 1877, ' 78, ' 79, by the U. S. Coast Survey Steamer &quot; Blake &quot;, Lieut. - Commander C. D. Sigsbee, U. S. N., and Commander J. R. Bartlett, U. S. N., Commanding, VIII. Bulletin of the Museum of Comparative Zoology at Harvard College, 8 (1), 1 - 68." type="journal article" year="1880">A. Milne-Edwards 1880</bibRefCitation>
;
<bibRefCitation id="CC6A4789FFD2FFA7D895FA71FE4AFA56" author="Alcock" box="[231,389,1432,1458]" pageId="7" pageNumber="23" refString="Alcock, A. (1899) An Account of the Deep-Sea Brachyura collected by the Royal Indian Marine Survey Ship &quot; Investigator &quot;. Indian Museum, Calcutta, 85 pp." type="book" year="1899">Alcock 1899</bibRefCitation>
) and subsequent transfer to the
<taxonomicName id="6FFB41FBFFD2FFA7DB6CFA71FC6CFA56" box="[798,931,1432,1458]" class="Malacostraca" family="Prosopidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="family">Prosopidae</taxonomicName>
, subfamily Pithonotinae Glaessner, 1933 (
<bibRefCitation id="CC6A4789FFD2FFA7D8ECFA29FE71FA3E" author="Wright" box="[158,446,1472,1498]" pageId="7" pageNumber="26" refString="Wright, C. W. &amp; Collins, J. S. H. (1972) British Cretaceous crabs. Monograph of the Palaeontographical Society, London, 126 (533), 1 - 114." type="journal article" year="1972">Wright &amp; Collins 1972</bibRefCitation>
), illustrates well the “curious mixture” (
<bibRefCitation id="CC6A4789FFD2FFA7DBDBFA29FB4AFA3E" author="McLay" box="[937,1157,1472,1498]" pageId="7" pageNumber="25" refString="McLay, C. L. (1999) Crustacea Decapoda: Revision of the Family Dynomenidae. In: Crosnier, A. (Ed.), Resultats des Campagnes MUSORSTOM, Volume 20. Memoires du Museum national d'Histoire naturelle, 180, 427 - 569." type="journal article" year="1999">McLay 1999: 534</bibRefCitation>
) of characters, already pointed out by
<bibRefCitation id="CC6A4789FFD2FFA7D935FA01FD30F9E6" author="Milne-Edwards" box="[327,767,1512,1538]" pageId="7" pageNumber="25" refString="Milne-Edwards, A. &amp; Bouvier, E. - L. (1902) Reports on the results of dredging, under the supervision of Alexander Agassiz, in the Gulf of Mexico (1877 - 78), in the Caribbean Sea (1878 - 79), and along the Atlantic Coast of the United States (1880), by the U. S. Coast Survey Steamer &quot; Blake &quot;. XXXIX. Les Dromiaces et Oxystomes. Memoirs of the Museum of Comparative Zoology at Harvard College, 27 (1), 1 - 127." type="journal article" year="1902">A. Milne-Edwards &amp; Bouvier (1902)</bibRefCitation>
. The present study has shown that there is strong support for the interpretation that the Recent
<taxonomicName id="6FFB41FBFFD2FFA7DA3DF9F9FCCBF9CD" box="[591,772,1552,1577]" class="Malacostraca" family="Dynomenidae" genus="Acanthodromia" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="9A8FE66AFFD2FFA7DA3DF9F9FCCBF9CD" box="[591,772,1552,1577]" italics="true" pageId="7" pageNumber="8">Acanthodromia</emphasis>
</taxonomicName>
retains ancestral characters and is the most primitive of all extant dynomenids.
</paragraph>
</subSubSection>
</treatment>
</document>