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<mods:title id="6C872CE009A41B61199351C13E284F50">Crabs (Crustacea, Decapoda) from the Sea off East and Southeast Asia Collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 2. Timor Sea</mods:title>
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<treatment id="039387ECAF5FFF85FC892CF38876FA0B" ID-DOI="http://doi.org/10.5281/zenodo.12571397" ID-GBIF-Taxon="233808313" ID-Zenodo-Dep="12571397" LSID="urn:lsid:plazi:treatment:039387ECAF5FFF85FC892CF38876FA0B" httpUri="http://treatment.plazi.org/id/039387ECAF5FFF85FC892CF38876FA0B" lastPageId="8" lastPageNumber="13" pageId="6" pageNumber="11">
<subSubSection id="C3206571AF5FFF8BFC892CF38EEEF957" box="[783,1159,1685,1710]" pageId="6" pageNumber="11" type="nomenclature">
<paragraph id="8B8536FAAF5FFF8BFC892CF38EEEF957" blockId="6.[712,1231,1616,1739]" box="[783,1159,1685,1710]" pageId="6" pageNumber="11">
<heading id="D0CD8196AF5FFF8BFC892CF38EEEF957" box="[783,1159,1685,1710]" centered="true" fontSize="9" level="2" pageId="6" pageNumber="11" reason="4">
<taxonomicName id="4C3A4D79AF5FFF8BFC892CF38EEEF957" ID-CoL="6H583" authority="Ng and Ho, 2003" authorityName="Ng and Ho. A" authorityYear="2003" box="[783,1159,1685,1710]" class="Malacostraca" family="Ethusidae" genus="Ethusina" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="11" phylum="Arthropoda" rank="species" species="chenae">
<emphasis id="B94EEAE8AF5FFF8BFC892CF389A3F954" bold="true" box="[783,970,1685,1709]" italics="true" pageId="6" pageNumber="11">Ethusina chenae</emphasis>
<bibRefCitation id="EFAB4B0BAF5FFF8BFC562CF08EEEF957" author="Ng, P. K. L. &amp; P. - H. Ho" box="[976,1159,1686,1710]" pageId="6" pageNumber="11" pagination="71 - 85" refId="ref12453" refString="Ng, P. K. L. and P. - H. Ho 2003. On the deep-water dorip- pid crabs of the genus Ethusina Smith, 1884 (Crustacea: Decapoda: Brachyura) from Taiwan. The Raffles Bulletin of Zoology 51: 71 - 85." type="journal article" year="2003">Ng and Ho, 2003</bibRefCitation>
</taxonomicName>
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<paragraph id="8B8536FAAF5FFF8BFC2F2CD38987F932" blockId="6.[712,1231,1616,1739]" box="[937,1006,1717,1739]" pageId="6" pageNumber="11">
(
<figureCitation id="13012A7FAF5FFF8BFC362CD3898FF932" box="[944,998,1717,1739]" captionStart="Fig" captionStartId="7.[163,196,1073,1095]" captionTargetBox="[234,1142,214,1039]" captionTargetId="figure-311@7.[233,1143,213,1040]" captionTargetPageId="7" captionText="Fig. 4. Ethusina chenae Ng and Ho. AB: Female (NSMT-Cr 29255: CB 9.3mm×CL 10.9mm) from sta. 26 (Timor Sea, 610690m depth). C: Juvenile (NSMT-Cr 29256: CB 4.0mm×CL 4.2mm) from sta. 26 (Timor Sea; 610690m depth)." figureDoi="http://doi.org/10.5281/zenodo.12571362" httpUri="https://zenodo.org/record/12571362/files/figure.png" pageId="6" pageNumber="11">Fig. 4</figureCitation>
)
</paragraph>
<paragraph id="8B8536FAAF5FFF8AFD5B2C898B76FA73" blockId="6.[733,1236,1775,1799]" lastBlockId="7.[140,670,1185,1800]" lastPageId="7" lastPageNumber="12" pageId="6" pageNumber="11">
<emphasis id="B94EEAE8AF5FFF8BFD5B2C8989D6F8FE" box="[733,959,1775,1799]" italics="true" pageId="6" pageNumber="11">Material examined</emphasis>
. RV
<emphasis id="B94EEAE8AF5FFF8BFBA02C898EBDF8FE" box="[1062,1236,1775,1799]" italics="true" pageId="6" pageNumber="11">Hakuhō Maru</emphasis>
KH-72-1 cruise, sta. 26 (Timor Sea;
<geoCoordinate id="EE0E503DAF5EFF8AFDA92EC488FEFB43" box="[559,663,1185,1210]" degrees="09" direction="south" minutes="27.0" orientation="latitude" pageId="7" pageNumber="12" precision="92" value="-9.45">09°27.0S</geoCoordinate>
,
<geoCoordinate id="EE0E503DAF5EFF8AFF0A2EA38B6BFB24" box="[140,258,1219,1245]" degrees="127" direction="east" minutes="58.6" orientation="longitude" pageId="7" pageNumber="12" precision="92" value="127.97667">127°58.6E</geoCoordinate>
09°285.0S,
<geoCoordinate id="EE0E503DAF5EFF8AFE1D2EA3887DFB25" box="[411,532,1219,1245]" degrees="127" direction="east" minutes="56.1" orientation="longitude" pageId="7" pageNumber="12" precision="92" value="127.935">127°56.1E</geoCoordinate>
,
<quantity id="4CC29B1FAF5EFF8AFDAE2EA388F4FB25" box="[552,669,1221,1245]" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="6.9" metricValueMin="6.1" pageId="7" pageNumber="12" unit="m" value="650.0" valueMax="690.0" valueMin="610.0">610690 m</quantity>
deep),
<quantity id="4CC29B1FAF5EFF8AFF5A2E818B68FB06" box="[220,257,1255,1279]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="12" unit="m" value="3.0">3 m</quantity>
beam trawl; 19 June, 1972;
<specimenCount id="9D3CFD73AF5EFF8AFDDD2E8188F7FB06" box="[603,670,1255,1279]" count="1" pageId="7" pageNumber="12" type="generic">1 juv.</specimenCount>
(NSMT-Cr 29256: CB 4.0 mm, CL
<quantity id="4CC29B1FAF5EFF8AFDB92F6C88F8FADB" box="[575,657,1290,1314]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="7" pageNumber="12" unit="mm" value="4.2">4.2 mm</quantity>
),
<specimenCount id="9D3CFD73AF5EFF8AFF0A2F4B8AA1FABC" box="[140,200,1325,1349]" count="1" pageId="7" pageNumber="12" type="generic">1 juv.</specimenCount>
(NSMT-Cr 29257, dried up: CB 5.0 mm, CL
<quantity id="4CC29B1FAF5EFF8AFF332F298B6EFA9E" box="[181,263,1359,1383]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="7" pageNumber="12" unit="mm" value="5.5">5.5 mm</quantity>
), 1 ♀ (NSMT-Cr 29255: CB
<quantity id="4CC29B1FAF5EFF8AFDCE2F2988F0FA9E" box="[584,665,1359,1383]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="7" pageNumber="12" unit="mm" value="9.3">9.3 mm</quantity>
, CL
<quantity id="4CC29B1FAF5EFF8AFF322F148B7DFA73" box="[180,276,1394,1418]" metricMagnitude="-2" metricUnit="m" metricValue="1.09" pageId="7" pageNumber="12" unit="mm" value="10.9">10.9 mm</quantity>
).
</paragraph>
<caption id="DF456672AF5EFF8AFF252E578BCEFB7B" ID-DOI="http://doi.org/10.5281/zenodo.12571362" ID-Zenodo-Dep="12571362" httpUri="https://zenodo.org/record/12571362/files/figure.png" pageId="7" pageNumber="12" startId="7.[163,196,1073,1095]" targetBox="[234,1142,214,1039]" targetPageId="7" targetType="figure">
<paragraph id="8B8536FAAF5EFF8AFF252E578BCEFB7B" blockId="7.[163,1214,1072,1154]" pageId="7" pageNumber="12">
Fig. 4.
<taxonomicName id="4C3A4D79AF5EFF8AFF7F2E57884AFBBE" authority="Ng and Ho. A" authorityName="Ng and Ho. A" authorityYear="2003" box="[249,547,1073,1095]" class="Malacostraca" family="Ethusidae" genus="Ethusina" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="12" phylum="Arthropoda" rank="species" species="chenae">
<emphasis id="B94EEAE8AF5EFF8AFF7F2E578BFBFBBF" box="[249,402,1073,1094]" italics="true" pageId="7" pageNumber="12">Ethusina chenae</emphasis>
Ng and Ho. A
</taxonomicName>
B: Female (NSMT-Cr 29255: CB 9.3 mm×CL 10.9mm) from sta. 26 (Timor Sea, 610690 m depth). C: Juvenile (NSMT-Cr 29256: CB 4.0 mm×CL 4.2 mm) from sta. 26 (Timor Sea; 610690 m depth).
</paragraph>
</caption>
<paragraph id="8B8536FAAF5EFF8AFF212FF38E04F9EC" blockId="7.[140,670,1185,1800]" lastBlockId="7.[707,1237,1186,1799]" pageId="7" pageNumber="12">
<emphasis id="B94EEAE8AF5EFF8AFF212FF38B6BFA54" box="[167,258,1429,1453]" italics="true" pageId="7" pageNumber="12">Remarks</emphasis>
.
<taxonomicName id="4C3A4D79AF5EFF8AFE902FF3881CFA54" authority="(Miers, 1886)" baseAuthorityName="Miers" baseAuthorityYear="1886" box="[278,629,1429,1453]" class="Malacostraca" family="Ethusidae" genus="Ethusina" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="12" phylum="Arthropoda" rank="species" species="robusta">
<emphasis id="B94EEAE8AF5EFF8AFE902FF38BA6FA54" box="[278,463,1429,1453]" italics="true" pageId="7" pageNumber="12">Ethusina robusta</emphasis>
(
<bibRefCitation id="EFAB4B0BAF5EFF8AFE652FF38807FA54" author="Miers, E. J." box="[483,622,1429,1453]" pageId="7" pageNumber="12" pagination="1 - 29" refId="ref11996" refString="Miers, E. J. 1886. Report on the Brachyura collected by H. M. S. Challenger during the years 1873 - 76. In: Report on the Scientific Results of the Voyage of H. M. S. Challenger during the years 1873 - 1876 under the command of Captain George S. Nares, R. N., F. R. S. and the late Captain Frank Tourle Thomson, R. N. prepared under the Superintendence of the late Sir C. Wyville Thomson, Knt., F. R. S. &amp; c. Regius Professor of Natural History in the University of Edinburgh of the civilian scientific staff on board and now of John Mur- ray, one of the naturalists of the Expedition. Zoology, 17: i - L, 1 - 362, pls. 1 - 29." type="journal article" year="1886">Miers, 1886</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C3A4D79AF5EFF8AFD002FF38B8DFA36" authority="(Alcock, 1896)" baseAuthorityName="Alcock" baseAuthorityYear="1896" class="Malacostraca" family="Ethusidae" genus="Ethusina" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="12" phylum="Arthropoda" rank="species" species="investigatoris">
<emphasis id="B94EEAE8AF5EFF8AFD002FF38B73FA36" italics="true" pageId="7" pageNumber="12">E. investigatoris</emphasis>
(
<bibRefCitation id="EFAB4B0BAF5EFF8AFEBF2FD18BB5FA36" author="Alcock, A." box="[313,476,1463,1487]" pageId="7" pageNumber="12" pagination="134 - 296" refId="ref9898" refString="Alcock, A. 1896. Materials for a carcinological fauna of India. No. 2. The Brachyura Oxystoma. Journal of the Asiatic Society of Bengal, (II) 65: 134 - 296, pls. 6 - 8." type="journal article" year="1896">Alcock, 1896</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C3A4D79AF5EFF8AFD862FDE8B46FA0B" authority="(Alcock, 1896)" baseAuthorityName="Alcock" baseAuthorityYear="1896" class="Malacostraca" family="Ethusidae" genus="Ethusina" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="12" phylum="Arthropoda" rank="species" species="desciscens">
<emphasis id="B94EEAE8AF5EFF8AFD862FDE88F4FA36" box="[512,669,1463,1487]" italics="true" pageId="7" pageNumber="12">E. desciscens</emphasis>
(
<bibRefCitation id="EFAB4B0BAF5EFF8AFF152FBC8B4EFA0B" author="Alcock, A." box="[147,295,1498,1522]" pageId="7" pageNumber="12" pagination="134 - 296" refId="ref9898" refString="Alcock, A. 1896. Materials for a carcinological fauna of India. No. 2. The Brachyura Oxystoma. Journal of the Asiatic Society of Bengal, (II) 65: 134 - 296, pls. 6 - 8." type="journal article" year="1896">Alcock, 1896</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C3A4D79AF5EFF8AFEBA2FBD8803FA0B" authority="Chen, 2000" authorityName="Chen" authorityYear="2000" box="[316,618,1498,1522]" class="Malacostraca" family="Ethusidae" genus="Ethusina" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="12" phylum="Arthropoda" rank="species" species="vanuatuensis">
<emphasis id="B94EEAE8AF5EFF8AFEBA2FBD8B8AFA0B" box="[316,483,1499,1522]" italics="true" pageId="7" pageNumber="12">E. vanuatuensis</emphasis>
<bibRefCitation id="EFAB4B0BAF5EFF8AFE6B2FBC8803FA0B" author="Chen, H." box="[493,618,1498,1522]" pageId="7" pageNumber="12" pagination="425 - 435" refId="ref10587" refString="Chen, H. 2000. Crustacea Decapoda: new species and new records of Ethusinae (Dorippidae) from Vanuatu. In Crosnier, A. (ed.): Resultats des campagnes MUSORSTOM. Volume 21. Memoires du Museum national d`Histoire naturelle, Paris, (A) 184: 425 - 435." type="journal article" year="2000">Chen, 2000</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C3A4D79AF5EFF8AFF0A2F9B8BAFF9EC" authority="Ng and Ho, 2003" authorityName="Ng and Ho" authorityYear="2003" box="[140,454,1533,1557]" class="Malacostraca" family="Trichopeltariidae" genus="Trichopeltarion" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="12" phylum="Arthropoda" rank="species" species="alcocki">
<emphasis id="B94EEAE8AF5EFF8AFF0A2F9B8A91F9EC" box="[140,248,1533,1557]" italics="true" pageId="7" pageNumber="12">E. alcocki</emphasis>
<bibRefCitation id="EFAB4B0BAF5EFF8AFE852F9B8BAFF9EC" author="Ng, P. K. L. &amp; P. - H. Ho" box="[259,454,1533,1557]" pageId="7" pageNumber="12" pagination="71 - 85" refId="ref12453" refString="Ng, P. K. L. and P. - H. Ho 2003. On the deep-water dorip- pid crabs of the genus Ethusina Smith, 1884 (Crustacea: Decapoda: Brachyura) from Taiwan. The Raffles Bulletin of Zoology 51: 71 - 85." type="journal article" year="2003">Ng and Ho, 2003</bibRefCitation>
</taxonomicName>
are closely related, differing from each other mainly in details of the front-orbital tuberculation. Their complex synonymies were thoroughly clarified by
<bibRefCitation id="EFAB4B0BAF5EFF8AFD842C0388F3F985" author="Castro, P." box="[514,666,1636,1661]" pageId="7" pageNumber="12" pagination="499 - 600" refId="ref10299" refString="Castro, P. 2005. Crabs of the subfamily Ethusinae Guinot, 1977 (Crustacea, Decapoda, Brachyura, Dorippidae) of the Indo-West Pacific region. Zoosystema 27: 499 - 600." type="journal article" year="2005">Castro (2005)</bibRefCitation>
: 1)
<taxonomicName id="4C3A4D79AF5EFF8AFF362CE18BC0F966" baseAuthorityName="Alcock" baseAuthorityYear="1896" box="[176,425,1671,1695]" class="Malacostraca" family="Ethusidae" genus="Ethusina" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="12" phylum="Arthropoda" rank="species" species="investigatoris">
<emphasis id="B94EEAE8AF5EFF8AFF362CE18BC0F966" box="[176,425,1671,1695]" italics="true" pageId="7" pageNumber="12">Ethusina investigatoris</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3A4D79AF5EFF8AFE6D2CEE8832F966" authorityName="Ng and Ho" authorityYear="2003" box="[491,603,1671,1695]" class="Malacostraca" family="Trichopeltariidae" genus="Trichopeltarion" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="12" phylum="Arthropoda" rank="species" species="alcocki">
<emphasis id="B94EEAE8AF5EFF8AFE6D2CEE8832F966" box="[491,603,1671,1695]" italics="true" pageId="7" pageNumber="12">E. alcocki</emphasis>
</taxonomicName>
were synonymized with
<taxonomicName id="4C3A4D79AF5EFF8AFED52CCD8BA9F93B" baseAuthorityName="Miers" baseAuthorityYear="1886" box="[339,448,1706,1730]" class="Malacostraca" family="Ethusidae" genus="Ethusina" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="12" phylum="Arthropoda" rank="species" species="robusta">
<emphasis id="B94EEAE8AF5EFF8AFED52CCD8BA9F93B" box="[339,448,1706,1730]" italics="true" pageId="7" pageNumber="12">E. robusta</emphasis>
</taxonomicName>
; 2)
<taxonomicName id="4C3A4D79AF5EFF8AFE6C2CCC88F4F93B" baseAuthorityName="Miers" baseAuthorityYear="1886" box="[490,669,1706,1730]" class="Malacostraca" family="Ethusidae" genus="Ethusina" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="12" phylum="Arthropoda" rank="species" species="robusta">
<emphasis id="B94EEAE8AF5EFF8AFE6C2CCC88F4F93B" box="[490,669,1706,1730]" italics="true" pageId="7" pageNumber="12">Ethusina robusta</emphasis>
</taxonomicName>
identified by
<bibRefCitation id="EFAB4B0BAF5EFF8AFE902CAB8BF3F91C" author="Chen, H." box="[278,410,1741,1765]" pageId="7" pageNumber="12" pagination="118 - 139" refId="ref10437" refString="Chen, H. 1986. Studies on the Dorippidae (Crustacea, Brachyura) of Chinese waters. Transactions of the Chi- nese Crustacean Society 1: 118 - 139 (In Chinese with English summary)." type="journal article" year="1986">Chen (1986)</bibRefCitation>
was corrected to
<emphasis id="B94EEAE8AF5EFF8AFDD52CAB8ADBF8FE" italics="true" pageId="7" pageNumber="12">E. chenae</emphasis>
<bibRefCitation id="EFAB4B0BAF5EFF8AFF3C2C968B10F8FE" author="Ng, P. K. L. &amp; P. - H. Ho" box="[186,377,1775,1800]" pageId="7" pageNumber="12" pagination="71 - 85" refId="ref12453" refString="Ng, P. K. L. and P. - H. Ho 2003. On the deep-water dorip- pid crabs of the genus Ethusina Smith, 1884 (Crustacea: Decapoda: Brachyura) from Taiwan. The Raffles Bulletin of Zoology 51: 71 - 85." type="journal article" year="2003">Ng and Ho, 2003</bibRefCitation>
; 3) the identifications of
<taxonomicName id="4C3A4D79AF5EFF8AFD002C968958FB43" baseAuthorityName="Alcock" baseAuthorityYear="1896" box="[646,817,1186,1799]" class="Malacostraca" family="Ethusidae" genus="Ethusina" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="12" phylum="Arthropoda" rank="species" species="desciscens">
<emphasis id="B94EEAE8AF5EFF8AFD002C968958FB43" box="[646,817,1186,1799]" italics="true" pageId="7" pageNumber="12">E. desciscens</emphasis>
</taxonomicName>
identified by
<bibRefCitation id="EFAB4B0BAF5EFF8AFC492EC48E38FB43" author="Chen, H." box="[975,1105,1186,1210]" pageId="7" pageNumber="12" pagination="118 - 139" refId="ref10437" refString="Chen, H. 1986. Studies on the Dorippidae (Crustacea, Brachyura) of Chinese waters. Transactions of the Chi- nese Crustacean Society 1: 118 - 139 (In Chinese with English summary)." type="journal article" year="1986">Chen (1986</bibRefCitation>
,
<bibRefCitation id="EFAB4B0BAF5EFF8AFBE62EC48EF1FB43" author="Chen, H." box="[1120,1176,1186,1210]" pageId="7" pageNumber="12" pagination="228 - 264" refId="ref10531" refString="Chen, H. 1998. Further study on the crabs (Crustacea: Brachyura) from Nansha Islands and adjacent waters. Studies on Marine Flora and Fauna and Biogeography of the Nansha Islands and Neighboring Waters 3: 228 - 264. Ocean Press, Beijing (In Chinese with English summary)." type="journal article" year="1998">1998</bibRefCitation>
), and
<bibRefCitation id="EFAB4B0BAF5EFF8AFD452EA389CAFB24" author="Chen, H. &amp; H. Sun" box="[707,931,1221,1245]" pageId="7" pageNumber="12" refId="ref10645" refString="Chen, H. and H. Sun 2002. Marine Primitive Crabs, Brachyura, Arthropoda Crustacea. Fauna Sinica, Inver- tebrata 30: xiii + 597, 16 pls. Science Press, Beijing (In Chinese with English summary)." type="book" year="2002">Chen and Sun (2002)</bibRefCitation>
were corrected to
<emphasis id="B94EEAE8AF5EFF8AFBE12EA3889EFB06" italics="true" pageId="7" pageNumber="12">E. taiwanensis</emphasis>
<bibRefCitation id="EFAB4B0BAF5EFF8AFD792E8E89D3FB06" author="Ng, P. K. L. &amp; P. - H. Ho" box="[767,954,1255,1280]" pageId="7" pageNumber="12" pagination="71 - 85" refId="ref12453" refString="Ng, P. K. L. and P. - H. Ho 2003. On the deep-water dorip- pid crabs of the genus Ethusina Smith, 1884 (Crustacea: Decapoda: Brachyura) from Taiwan. The Raffles Bulletin of Zoology 51: 71 - 85." type="journal article" year="2003">Ng and Ho, 2003</bibRefCitation>
; 4)
<taxonomicName id="4C3A4D79AF5EFF8AFC652E818EFEFB06" baseAuthorityName="Miers" baseAuthorityYear="1886" box="[995,1175,1255,1279]" class="Malacostraca" family="Ethusidae" genus="Ethusina" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="12" phylum="Arthropoda" rank="species" species="robusta">
<emphasis id="B94EEAE8AF5EFF8AFC652E818EFEFB06" box="[995,1175,1255,1279]" italics="true" pageId="7" pageNumber="12">Ethusina robusta</emphasis>
</taxonomicName>
identified by
<bibRefCitation id="EFAB4B0BAF5EFF8AFCA52F6C89C0FADB" author="Chen, H." box="[803,937,1290,1314]" pageId="7" pageNumber="12" pagination="315 - 345" refId="ref10476" refString="Chen, H. 1993. Crustacea Decapoda: Dorippidae of New Caledonia, Indonesia and the Philippines. In Crosnier, A. (ed.): Resultats des campagnes MUSORSTOM. Volume 10. Memoires du Museum national d`Histoire naturelle, Paris, (A) 156: 315 - 345." type="journal article" year="1993">Chen (1993)</bibRefCitation>
was referred to
<emphasis id="B94EEAE8AF5EFF8AFBDE2F6D889EFABD" italics="true" pageId="7" pageNumber="12">E. vanuatuensis</emphasis>
<bibRefCitation id="EFAB4B0BAF5EFF8AFC802F4B89E0FABC" author="Chen, H." box="[774,905,1325,1349]" pageId="7" pageNumber="12" pagination="315 - 345" refId="ref10476" refString="Chen, H. 1993. Crustacea Decapoda: Dorippidae of New Caledonia, Indonesia and the Philippines. In Crosnier, A. (ed.): Resultats des campagnes MUSORSTOM. Volume 10. Memoires du Museum national d`Histoire naturelle, Paris, (A) 156: 315 - 345." type="journal article" year="1993">Chen, 1993</bibRefCitation>
,
<taxonomicName id="4C3A4D79AF5EFF8AFC1D2F4B8893FA9E" authority="Ng and Ho, 2003" authorityName="Ng and Ho" authorityYear="2003" class="Malacostraca" family="Ethusidae" genus="Ethusina" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="12" phylum="Arthropoda" rank="species" species="macrospina">
<emphasis id="B94EEAE8AF5EFF8AFC1D2F4B8E55FABD" box="[923,1084,1325,1348]" italics="true" pageId="7" pageNumber="12">E. macrospina</emphasis>
<bibRefCitation id="EFAB4B0BAF5EFF8AFBCC2F4B8893FA9E" author="Ng, P. K. L. &amp; P. - H. Ho" pageId="7" pageNumber="12" pagination="71 - 85" refId="ref12453" refString="Ng, P. K. L. and P. - H. Ho 2003. On the deep-water dorip- pid crabs of the genus Ethusina Smith, 1884 (Crustacea: Decapoda: Brachyura) from Taiwan. The Raffles Bulletin of Zoology 51: 71 - 85." type="journal article" year="2003">Ng and Ho, 2003</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C3A4D79AF5EFF8AFCB12F368E05FA9E" authority="Castro, 2005" authorityName="Castro" authorityYear="2005" box="[823,1132,1359,1383]" class="Malacostraca" family="Ethusidae" genus="Ethusina" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="12" phylum="Arthropoda" rank="species" species="stenommata">
<emphasis id="B94EEAE8AF5EFF8AFCB12F3689BFFA9E" box="[823,982,1360,1383]" italics="true" pageId="7" pageNumber="12">E. stenommata</emphasis>
<bibRefCitation id="EFAB4B0BAF5EFF8AFC662F298E05FA9E" author="Castro, P." box="[992,1132,1359,1383]" pageId="7" pageNumber="12" pagination="499 - 600" refId="ref10299" refString="Castro, P. 2005. Crabs of the subfamily Ethusinae Guinot, 1977 (Crustacea, Decapoda, Brachyura, Dorippidae) of the Indo-West Pacific region. Zoosystema 27: 499 - 600." type="journal article" year="2005">Castro, 2005</bibRefCitation>
</taxonomicName>
; 5)
<taxonomicName id="4C3A4D79AF5EFF8AFB1C2F298938FA73" baseAuthorityName="Miers" baseAuthorityYear="1886" class="Malacostraca" family="Ethusidae" genus="Ethusina" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="12" phylum="Arthropoda" rank="species" species="robusta">
<emphasis id="B94EEAE8AF5EFF8AFB1C2F298938FA73" italics="true" pageId="7" pageNumber="12">Ethusina robusta</emphasis>
</taxonomicName>
identified by
<bibRefCitation id="EFAB4B0BAF5EFF8AFB812F148EFEFA73" author="Chen, H." box="[1031,1175,1394,1418]" pageId="7" pageNumber="12" pagination="425 - 435" refId="ref10587" refString="Chen, H. 2000. Crustacea Decapoda: new species and new records of Ethusinae (Dorippidae) from Vanuatu. In Crosnier, A. (ed.): Resultats des campagnes MUSORSTOM. Volume 21. Memoires du Museum national d`Histoire naturelle, Paris, (A) 184: 425 - 435." type="journal article" year="2000">Chen (2000)</bibRefCitation>
was reidentified as
<taxonomicName id="4C3A4D79AF5EFF8AFCD92FF38EEDFA55" authority="Castro, 2005" authorityName="Castro" authorityYear="2005" box="[863,1156,1428,1453]" class="Malacostraca" family="Ethusidae" genus="Ethusina" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="12" phylum="Arthropoda" rank="species" species="ciliacirrata">
<emphasis id="B94EEAE8AF5EFF8AFCD92FF3899DFA54" box="[863,1012,1429,1453]" italics="true" pageId="7" pageNumber="12">E. ciliacirrata</emphasis>
<bibRefCitation id="EFAB4B0BAF5EFF8AFC7D2FF38EEDFA55" author="Castro, P." box="[1019,1156,1428,1453]" pageId="7" pageNumber="12" pagination="499 - 600" refId="ref10299" refString="Castro, P. 2005. Crabs of the subfamily Ethusinae Guinot, 1977 (Crustacea, Decapoda, Brachyura, Dorippidae) of the Indo-West Pacific region. Zoosystema 27: 499 - 600." type="journal article" year="2005">Castro, 2005</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C3A4D79AF5EFF8AFB3B2FF389D8FA36" authority="Castro, 2005" authorityName="Castro" authorityYear="2005" class="Malacostraca" family="Ethusidae" genus="Ethusina" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="12" phylum="Arthropoda" rank="species" species="coronata">
<emphasis id="B94EEAE8AF5EFF8AFB3B2FF38949FA36" italics="true" pageId="7" pageNumber="12">E. coronata</emphasis>
<bibRefCitation id="EFAB4B0BAF5EFF8AFCA12FD189D8FA36" author="Castro, P." box="[807,945,1463,1487]" pageId="7" pageNumber="12" pagination="499 - 600" refId="ref10299" refString="Castro, P. 2005. Crabs of the subfamily Ethusinae Guinot, 1977 (Crustacea, Decapoda, Brachyura, Dorippidae) of the Indo-West Pacific region. Zoosystema 27: 499 - 600." type="journal article" year="2005">Castro, 2005</bibRefCitation>
</taxonomicName>
; and 6) the identification of
<taxonomicName id="4C3A4D79AF5EFF8AFD452FBD8959FA0B" baseAuthorityName="Miers" baseAuthorityYear="1886" box="[707,816,1498,1522]" class="Malacostraca" family="Ethusidae" genus="Ethusina" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="12" phylum="Arthropoda" rank="species" species="robusta">
<emphasis id="B94EEAE8AF5EFF8AFD452FBD8959FA0B" box="[707,816,1498,1522]" italics="true" pageId="7" pageNumber="12">E. robusta</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFAB4B0BAF5EFF8AFCDD2FBC8E57FA0B" author="Chen, H. &amp; H. Sun" box="[859,1086,1498,1522]" pageId="7" pageNumber="12" refId="ref10645" refString="Chen, H. and H. Sun 2002. Marine Primitive Crabs, Brachyura, Arthropoda Crustacea. Fauna Sinica, Inver- tebrata 30: xiii + 597, 16 pls. Science Press, Beijing (In Chinese with English summary)." type="book" year="2002">Chen and Sun (2002)</bibRefCitation>
was similarly corrected to
<taxonomicName id="4C3A4D79AF5EFF8AFCC32F9B8E00F9EC" authority="Ng and Ho, 2003" authorityName="Ng and Ho. A" authorityYear="2003" box="[837,1129,1533,1557]" class="Malacostraca" family="Ethusidae" genus="Ethusina" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="12" phylum="Arthropoda" rank="species" species="chenae">
<emphasis id="B94EEAE8AF5EFF8AFCC32F9B89C2F9EC" box="[837,939,1533,1557]" italics="true" pageId="7" pageNumber="12">E. chenae</emphasis>
<bibRefCitation id="EFAB4B0BAF5EFF8AFC342F9B8E00F9EC" author="Ng, P. K. L. &amp; P. - H. Ho" box="[946,1129,1533,1557]" pageId="7" pageNumber="12" pagination="71 - 85" refId="ref12453" refString="Ng, P. K. L. and P. - H. Ho 2003. On the deep-water dorip- pid crabs of the genus Ethusina Smith, 1884 (Crustacea: Decapoda: Brachyura) from Taiwan. The Raffles Bulletin of Zoology 51: 71 - 85." type="journal article" year="2003">Ng and Ho, 2003</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8B8536FAAF5EFF85FD5B2C798BA8FC7B" blockId="7.[707,1237,1186,1799]" lastBlockId="8.[140,670,215,1522]" lastPageId="8" lastPageNumber="13" pageId="7" pageNumber="12">
After detailed comparison with the literature, the female at hand (
<figureCitation id="13012A7FAF5EFF8AFC2E2C248E48F9A3" box="[936,1057,1602,1626]" captionStart="Fig" captionStartId="7.[163,196,1073,1095]" captionTargetBox="[234,1142,214,1039]" captionTargetId="figure-311@7.[233,1143,213,1040]" captionTargetPageId="7" captionText="Fig. 4. Ethusina chenae Ng and Ho. AB: Female (NSMT-Cr 29255: CB 9.3mm×CL 10.9mm) from sta. 26 (Timor Sea, 610690m depth). C: Juvenile (NSMT-Cr 29256: CB 4.0mm×CL 4.2mm) from sta. 26 (Timor Sea; 610690m depth)." figureDoi="http://doi.org/10.5281/zenodo.12571362" httpUri="https://zenodo.org/record/12571362/files/figure.png" pageId="7" pageNumber="12">Fig. 4AB</figureCitation>
), which is fully mature, was identified with
<taxonomicName id="4C3A4D79AF5EFF8AFB852C038E1BF984" authorityName="Ng and Ho. A" authorityYear="2003" box="[1027,1138,1637,1661]" class="Malacostraca" family="Ethusidae" genus="Ethusina" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="12" phylum="Arthropoda" rank="species" species="chenae">
<emphasis id="B94EEAE8AF5EFF8AFB852C038E1BF984" box="[1027,1138,1637,1661]" italics="true" pageId="7" pageNumber="12">E. chenae</emphasis>
</taxonomicName>
. Among the species mentioned above,
<taxonomicName id="4C3A4D79AF5EFF8AFB8B2CEE8E10F966" authorityName="Ng and Ho. A" authorityYear="2003" box="[1037,1145,1671,1695]" class="Malacostraca" family="Ethusidae" genus="Ethusina" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="12" phylum="Arthropoda" rank="species" species="chenae">
<emphasis id="B94EEAE8AF5EFF8AFB8B2CEE8E10F966" box="[1037,1145,1671,1695]" italics="true" pageId="7" pageNumber="12">E. chenae</emphasis>
</taxonomicName>
is characteristic in that 1) the carapace shape is not constricted behind the external orbital spine that is directed nearly forward or only slightly outward, with the tip reaching nearly to the tip of the median frontal tooth, 2) each median frontal tooth is triangular in dorsal view, with a sharp tip, and the lateral frontal tooth is directed forward, similar in shape to the outer orbital spine, extending beyond the tip of the median frontal tooth, 3) the median frontal sinus is comparatively wide and shallow, and the orbital sinus is distinctly shallow, with the orbital margin gently retreating toward the external orbital spine. The characters observed as above in the present female seem to be minor but reliable, agreeing well with the original description and figures by
<bibRefCitation id="EFAB4B0BAF51FF85FF0A28FC8B3BFD4B" author="Ng, P. K. L. &amp; P. - H. Ho" box="[140,338,666,690]" pageId="8" pageNumber="13" pagination="71 - 85" refId="ref12453" refString="Ng, P. K. L. and P. - H. Ho 2003. On the deep-water dorip- pid crabs of the genus Ethusina Smith, 1884 (Crustacea: Decapoda: Brachyura) from Taiwan. The Raffles Bulletin of Zoology 51: 71 - 85." type="journal article" year="2003">Ng and Ho (2003)</bibRefCitation>
, and with comments by
<bibRefCitation id="EFAB4B0BAF51FF85FDD128FC8ABFFD2D" author="Castro, P." pageId="8" pageNumber="13" pagination="499 - 600" refId="ref10299" refString="Castro, P. 2005. Crabs of the subfamily Ethusinae Guinot, 1977 (Crustacea, Decapoda, Brachyura, Dorippidae) of the Indo-West Pacific region. Zoosystema 27: 499 - 600." type="journal article" year="2005">Castro (2005)</bibRefCitation>
. In the
<typeStatus id="54818858AF51FF85FEC328DB8BCBFD2C" box="[325,418,701,725]" pageId="8" pageNumber="13" type="holotype">holotype</typeStatus>
female, both of the median frontal teeth seem to be blunt, probably due to damage during lifetime. The figures represented by
<bibRefCitation id="EFAB4B0BAF51FF85FF7C29438B89FCC4" author="Chen, H. &amp; H. Sun" box="[250,480,805,829]" pageId="8" pageNumber="13" refId="ref10645" refString="Chen, H. and H. Sun 2002. Marine Primitive Crabs, Brachyura, Arthropoda Crustacea. Fauna Sinica, Inver- tebrata 30: xiii + 597, 16 pls. Science Press, Beijing (In Chinese with English summary)." type="book" year="2002">Chen and Sun (2002)</bibRefCitation>
as
<taxonomicName id="4C3A4D79AF51FF85FD802943881CFCC4" baseAuthorityName="Miers" baseAuthorityYear="1886" box="[518,629,805,829]" class="Malacostraca" family="Ethusidae" genus="Ethusina" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="13" phylum="Arthropoda" rank="species" species="robusta">
<emphasis id="B94EEAE8AF51FF85FD802943881CFCC4" box="[518,629,805,829]" italics="true" pageId="8" pageNumber="13">E. robusta</emphasis>
</taxonomicName>
are somewhat schematic, but indicative of the specific characters of
<emphasis id="B94EEAE8AF51FF85FED6290D8BA8FC7B" box="[336,449,874,898]" italics="true" pageId="8" pageNumber="13">
<taxonomicName id="4C3A4D79AF51FF85FED6290D8BD4FC7B" authorityName="Ng and Ho. A" authorityYear="2003" box="[336,445,874,898]" class="Malacostraca" family="Ethusidae" genus="Ethusina" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="13" phylum="Arthropoda" rank="species" species="chenae">E. chenae</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8B8536FAAF51FF85FF2129EB8809FADB" blockId="8.[140,670,215,1522]" pageId="8" pageNumber="13">
Two small juveniles at hand belong without doubt to
<taxonomicName id="4C3A4D79AF51FF85FF7029C98B3AFC3E" authorityName="Smith" authorityYear="1884" box="[246,339,943,967]" class="Malacostraca" family="Ethusidae" genus="Ethusina" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAE8AF51FF85FF7029C98B3AFC3E" box="[246,339,943,967]" italics="true" pageId="8" pageNumber="13">Ethusina</emphasis>
</taxonomicName>
by having the swollen basal antennular article pushing the antenna and the long ambulatory legs, but it is remarkable in having the prominent eyestalks visible from above, different from most of the congeners and somewhat similar to the
<taxonomicName id="4C3A4D79AF51FF85FEED2E3B887BFB8C" box="[363,530,1117,1141]" class="Malacostraca" family="Ethusidae" genus="Ethusa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="13" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B94EEAE8AF51FF85FEED2E3B8BDDFB8C" box="[363,436,1117,1141]" italics="true" pageId="8" pageNumber="13">Ethusa</emphasis>
species.
</taxonomicName>
They keyed out close to
<taxonomicName id="4C3A4D79AF51FF85FE942EE68BF6FB6E" baseAuthorityName="Alcock" baseAuthorityYear="1896" box="[274,415,1151,1175]" class="Malacostraca" family="Ethusidae" genus="Ethusina" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="13" phylum="Arthropoda" rank="species" species="desciscens">
<emphasis id="B94EEAE8AF51FF85FE942EE68BF6FB6E" box="[274,415,1151,1175]" italics="true" pageId="8" pageNumber="13">E. desciscens</emphasis>
</taxonomicName>
in the key provided by
<bibRefCitation id="EFAB4B0BAF51FF85FF0A2EC48B4BFB43" author="Castro, P." box="[140,290,1186,1210]" pageId="8" pageNumber="13" pagination="499 - 600" refId="ref10299" refString="Castro, P. 2005. Crabs of the subfamily Ethusinae Guinot, 1977 (Crustacea, Decapoda, Brachyura, Dorippidae) of the Indo-West Pacific region. Zoosystema 27: 499 - 600." type="journal article" year="2005">Castro (2005)</bibRefCitation>
. The semitransparent soft carapace, with sharp but not spiniform frontal and external orbital teeth, and the distinct eyestalks may be referable to the immaturity of the specimens.
</paragraph>
<paragraph id="8B8536FAAF51FF85FF212F4B8876FA0B" blockId="8.[140,670,215,1522]" pageId="8" pageNumber="13">
<emphasis id="B94EEAE8AF51FF85FF212F4B8B40FABC" box="[167,297,1325,1349]" italics="true" pageId="8" pageNumber="13">Distribution</emphasis>
. East
<collectingCountry id="F32D766AAF51FF85FEF52F4B8BDAFABC" box="[371,435,1325,1349]" name="China" pageId="8" pageNumber="13">China</collectingCountry>
Sea (
<bibRefCitation id="EFAB4B0BAF51FF85FE732F4B8811FABC" author="Chen, H." box="[501,632,1325,1349]" pageId="8" pageNumber="13" pagination="118 - 139" refId="ref10437" refString="Chen, H. 1986. Studies on the Dorippidae (Crustacea, Brachyura) of Chinese waters. Transactions of the Chi- nese Crustacean Society 1: 118 - 139 (In Chinese with English summary)." type="journal article" year="1986">Chen, 1986</bibRefCitation>
, as
<taxonomicName id="4C3A4D79AF51FF85FF0A2F368A96FA9E" baseAuthorityName="Miers" baseAuthorityYear="1886" box="[140,255,1359,1383]" class="Malacostraca" family="Ethusidae" genus="Ethusina" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="13" phylum="Arthropoda" rank="species" species="robusta">
<emphasis id="B94EEAE8AF51FF85FF0A2F368A96FA9E" box="[140,255,1359,1383]" italics="true" pageId="8" pageNumber="13">E. robusta</emphasis>
</taxonomicName>
),
<collectingCountry id="F32D766AAF51FF85FE9E2F298B0FFA9E" box="[280,358,1359,1383]" name="Taiwan" pageId="8" pageNumber="13">Taiwan</collectingCountry>
(Ng and Ho, 2002;
<bibRefCitation id="EFAB4B0BAF51FF85FDD72F298AACFA73" author="Castro, P." pageId="8" pageNumber="13" pagination="499 - 600" refId="ref10299" refString="Castro, P. 2005. Crabs of the subfamily Ethusinae Guinot, 1977 (Crustacea, Decapoda, Brachyura, Dorippidae) of the Indo-West Pacific region. Zoosystema 27: 499 - 600." type="journal article" year="2005">Castro, 2005</bibRefCitation>
), the
<collectingCountry id="F32D766AAF51FF85FE872F148B10FA73" box="[257,377,1394,1418]" name="Philippines" pageId="8" pageNumber="13">Philippines</collectingCountry>
(
<bibRefCitation id="EFAB4B0BAF51FF85FE0F2F148860FA73" author="Chen, H." box="[393,521,1394,1418]" pageId="8" pageNumber="13" pagination="179 - 203" refId="ref10384" refString="Chen, H. 1985. Decapod Crustacea: Dorippidae. In For- est, J. (ed.): Resultats des campagnes MUSORSTOM I et II. Philippines. Volume 2. Memoires du Museum national d`Histoire naturelle, Paris, (A) 133: 179 - 203." type="journal article" year="1985">Chen, 1985</bibRefCitation>
, as
<emphasis id="B94EEAE8AF51FF85FDB22F158AD2FA55" italics="true" pageId="8" pageNumber="13">E. desciscens</emphasis>
;
<bibRefCitation id="EFAB4B0BAF51FF85FF502FF38B00FA55" author="Castro, P." box="[214,361,1428,1453]" pageId="8" pageNumber="13" pagination="499 - 600" refId="ref10299" refString="Castro, P. 2005. Crabs of the subfamily Ethusinae Guinot, 1977 (Crustacea, Decapoda, Brachyura, Dorippidae) of the Indo-West Pacific region. Zoosystema 27: 499 - 600." type="journal article" year="2005">Castro 2005</bibRefCitation>
),
<quantity id="4CC29B1FAF51FF85FE0E2FF28862FA55" box="[392,523,1428,1452]" metricMagnitude="3" metricUnit="m" metricValue="1.079" metricValueMax="1.649" metricValueMin="0.509" pageId="8" pageNumber="13" unit="m" value="1079.0" valueMax="1649.0" valueMin="509.0">5091649 m</quantity>
depth. The record of
<taxonomicName id="4C3A4D79AF51FF85FF732FDE8BEDFA36" baseAuthorityName="Alcock" baseAuthorityYear="1896" box="[245,388,1463,1487]" class="Malacostraca" family="Ethusidae" genus="Ethusina" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="13" phylum="Arthropoda" rank="species" species="desciscens">
<emphasis id="B94EEAE8AF51FF85FF732FDE8BEDFA36" box="[245,388,1463,1487]" italics="true" pageId="8" pageNumber="13">E. desciscens</emphasis>
</taxonomicName>
from
<collectingCountry id="F32D766AAF51FF85FE402FDE886AFA36" box="[454,515,1464,1487]" name="Japan" pageId="8" pageNumber="13">Japan</collectingCountry>
by
<bibRefCitation id="EFAB4B0BAF51FF85FDAA2FDE8B3EFA0B" author="Marumura, M. &amp; A. Kosaka" pageId="8" pageNumber="13" pagination="1 - 8" refId="ref11868" refString="Marumura, M. and A. Kosaka 2003. Catalogue of the Brachyuran and Anomuran Crabs donated by the late Mr. Seiji Nagai to the Wakayama Prefectural Museum of Natural History. Wakayama Prefectural Museum of Natural History, pp. 12 - 73, pls. 1 - 8 (In Japanese)." type="book chapter" year="2003">Marumura and Kosaka (2003)</bibRefCitation>
is not yet verified.
</paragraph>
</subSubSection>
</treatment>
</document>