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<document id="59569F65304A71C9C09386EE85DB7616" ID-DOI="10.5281/zenodo.173979" ID-GBIF-Dataset="ecab4131-0772-49ca-9926-404c65ec981b" ID-ISSN="1175­5326" ID-Zenodo-Dep="173979" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459787568423" checkinUser="plazi" docAuthor="Smetana, Aleš" docDate="2006" docId="039387D3FFFA9C2FF707FB00883E798C" docLanguage="en" docName="zt01317p047.pdf" docOrigin="Zootaxa 1317" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Sphaerobulbus biplagiatus Smetana, 2006, spec. nov." docType="treatment" docVersion="9" lastPageNumber="45" masterDocId="FFAAFFABFFF89C2BF60FFFBE8B42785A" masterDocTitle="Contributions to the knowledge of the “ Staphylinus­complex ” (Coleoptera: Staphylinidae: Staphylinini) of China. Part 13. The genus Sphaerobulbus Smetana 2003. Section 3" masterLastPageNumber="47" masterPageNumber="41" pageNumber="43" updateTime="1698218796660" updateUser="plazi">
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<mods:title id="6C0052981FC4C59D2B9FE82E65F33387">Contributions to the knowledge of the “ Staphylinus­complex ” (Coleoptera: Staphylinidae: Staphylinini) of China. Part 13. The genus Sphaerobulbus Smetana 2003. Section 3</mods:title>
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<mods:namePart id="ADC332F5EABA93E05B469BF3621EFF05">Smetana, Aleš</mods:namePart>
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<mods:date id="2C7088179E3DE2C7DF59290B8F37729B">2006</mods:date>
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<treatment id="039387D3FFFA9C2FF707FB00883E798C" ID-DOI="http://doi.org/10.5281/zenodo.6258610" ID-GBIF-Taxon="119349523" ID-Zenodo-Dep="6258610" LSID="urn:lsid:plazi:treatment:039387D3FFFA9C2FF707FB00883E798C" httpUri="http://treatment.plazi.org/id/039387D3FFFA9C2FF707FB00883E798C" lastPageId="4" lastPageNumber="45" pageId="2" pageNumber="43">
<subSubSection id="C320654EFFFA9C29F707FB008ACA7CA1" pageId="2" pageNumber="43" type="nomenclature">
<paragraph id="8B8536C5FFFA9C29F707FB0089887C82" blockId="2.[264,714,1214,1275]" box="[264,714,1214,1240]" pageId="2" pageNumber="43">
<heading id="D0CD81A9FFFA9C29F707FB0089887C82" bold="true" box="[264,714,1214,1240]" fontSize="11" level="1" pageId="2" pageNumber="43" reason="1">
<emphasis id="B94EEAD7FFFA9C29F707FB0089887C82" bold="true" box="[264,714,1214,1240]" pageId="2" pageNumber="43">
<taxonomicName id="4C3A4D46FFFA9C29F707FB00890E7C82" ID-CoL="4YPP8" box="[264,588,1214,1240]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="43" phylum="Arthropoda" rank="species" species="biplagiatus" status="sp. nov.">
<emphasis id="B94EEAD7FFFA9C29F707FB00890E7C82" bold="true" box="[264,588,1214,1240]" italics="true" pageId="2" pageNumber="43">Sphaerobulbus biplagiatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27D57ACFFFA9C29F45CFB0089887C82" box="[595,714,1214,1240]" pageId="2" pageNumber="43" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8536C5FFFA9C29F707FB5F8ACA7CA1" blockId="2.[264,714,1214,1275]" box="[264,392,1249,1275]" pageId="2" pageNumber="43">
(
<figureCitation id="13012A40FFFA9C29F71EFB5F8A3D7CA1" box="[273,383,1249,1275]" captionStart="FIGURES 1 – 11" captionStartId="1.[264,382,1771,1795]" captionTargetBox="[282,1312,334,1749]" captionTargetId="figure@1.[274,1315,329,1751]" captionTargetPageId="1" captionText="FIGURES 1 – 11. Details of Sphaerobulbus biplagiatus (1 – 5) and S. ornatus (6 – 11). 1, 6 — sternite 9 of male genital segment; 2, 7 — tergite 10 of male genital segment; 3, 8 — aedoeagus, ventral view; 4, 9 — apical portion of median lobe, ventral view, paramere removed; 5, 10 — underside of apical portion of paramere; 11 — tergite 10 of female genital segment." httpUri="https://zenodo.org/record/173980/files/figure.png" pageId="2" pageNumber="43">Figs. 1–5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C320654EFFFA9C29F707FA8F898F7D29" pageId="2" pageNumber="43" type="materials_examined">
<paragraph id="8B8536C5FFFA9C29F707FA8F898F7D29" blockId="2.[264,1325,1329,1875]" pageId="2" pageNumber="43">
<emphasis id="B94EEAD7FFFA9C29F707FA8F8A847D11" bold="true" box="[264,454,1329,1355]" pageId="2" pageNumber="43">
<typeStatus id="54818867FFFA9C29F707FA8F8A077D11" box="[264,325,1329,1355]" pageId="2" pageNumber="43">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54818867FFFA9C29F7D8FA8F89067D11" box="[471,580,1329,1355]" pageId="2" pageNumber="43" type="holotype">Holotype</typeStatus>
(male):
<emphasis id="B94EEAD7FFFA9C29F4B6FA8F88E37D11" bold="true" box="[697,929,1329,1355]" pageId="2" pageNumber="43">
<collectingCountry id="F32D7655FFFA9C29F4B6FA8F885D7D11" box="[697,799,1329,1355]" name="China" pageId="2" pageNumber="43">CHINA</collectingCountry>
: Yunnan:
</emphasis>
“
<collectingCountry id="F32D7655FFFA9C29F5B4FA8F8F5C7D11" box="[955,1054,1329,1355]" name="China" pageId="2" pageNumber="43">CHINA</collectingCountry>
, W Yunnan Dequen, 15–
<date id="FF841005FFFA9C29F73AFAE78A297D29" box="[309,363,1369,1395]" pageId="2" pageNumber="43" value="2002-07-18">18.7</date>
. 2002 Lgt. E. Kučera” (ASC).
</paragraph>
</subSubSection>
<subSubSection id="C320654EFFFA9C2FF737FA3F8E4A7904" lastPageId="4" lastPageNumber="45" pageId="2" pageNumber="43" type="description">
<paragraph id="8B8536C5FFFA9C28F737FA3F8FB57BEC" blockId="2.[264,1325,1329,1875]" lastBlockId="3.[264,1325,284,1870]" lastPageId="3" lastPageNumber="44" pageId="2" pageNumber="43">
<emphasis id="B94EEAD7FFFA9C29F737FA3F8A8D7DC1" bold="true" box="[312,463,1409,1435]" pageId="2" pageNumber="43">Description.</emphasis>
Black, fore body rather dull; visible abdominal tergites 4 and 5 each with patch of golden tomentose pubescence in middle; maxillary and labial palpi piceous, antennae black, becoming vaguely paler toward apex, legs black with vaguely paler tarsi, front tarsi brownish. Head of rounded quadrangular shape, with entirely rounded posterior angles, wider than long (ratio 1.27), eyes small, slightly convex, tempora considerably longer than eyes from above (ratio 2.1), dorsal surface of head densely and finely punctate and pubescent, punctation gradually becoming less dense anteromediad; interspaces between punctures with dense, semimeshed microsculpture. Antenna moderately long, segment 3 slightly longer than segment 2 (ratio 1.18), segments 4 to 8 longer than wide, becoming gradually shorter, segments 9 and 10 about as long as wide, last segment shorter than two preceding segments combined. Pronotum slightly longer than wide (ratio 1.09), slightly narrowed anteriad, narrow marginal groove disappearing downwards at about anterior third of pronotal length; disc of pronotum with complete, indistinct, impunctate midline, dilated in front of posterior margin into slightly shiny (due to rudimentary microsculpture) mirror; punctation and pubescence similar to that on head, gradually becoming somewhat sparser mediad, interspaces between punctures with microsculpture similar to that on head. Scutellum sparsely, finely punctate and setose, surface rather shiny, with extremely fine, rudimentary microsculpture mediobasally. Elytra short, somewhat widened posteriad, base distinctly depressed, at suture considerably (ratio 0.66) at sides distinctly (ratio 0.82) shorter than pronotum at midline; punctation fine and dense, finely asperate, interspaces between punctures with fine, dense, granulose microsculpture, elytra therefore appearing dull; pubescence piceous­black, dense. Wings each reduced to minute, nonfunctional stump. Abdomen with fifth visible tergite lacking pale apical seam of palisade setae; tergite 2 (in front of first visible tergite) entirely, finely and densely punctate and pubescent; posterior basal line on first three visible abdominal tergites simple, almost straight; all tergites very finely and very densely punctate, punctation finer than that on elytra, evenly covering each tergite and in general becoming hardly sparser toward apex of abdomen; interspaces with very fine, rudimentary meshed microsculpture that becomes almost obsolete toward middle of each tergite; pubescence black, dense.
</paragraph>
<paragraph id="8B8536C5FFFB9C28F737FC7A8E447CAC" blockId="3.[264,1325,284,1870]" pageId="3" pageNumber="44">
Male. Sternite 8 with moderately wide and deep, obtusely triangular medioapical emargination. Male genital segment with sternite 9 with rather deep and wide apical emargination (
<figureCitation id="13012A40FFFB9C28F7BBFBAA8AB77C74" box="[436,501,1044,1070]" captionStart="FIGURES 1 – 11" captionStartId="1.[264,382,1771,1795]" captionTargetBox="[282,1312,334,1749]" captionTargetId="figure@1.[274,1315,329,1751]" captionTargetPageId="1" captionText="FIGURES 1 – 11. Details of Sphaerobulbus biplagiatus (1 – 5) and S. ornatus (6 – 11). 1, 6 — sternite 9 of male genital segment; 2, 7 — tergite 10 of male genital segment; 3, 8 — aedoeagus, ventral view; 4, 9 — apical portion of median lobe, ventral view, paramere removed; 5, 10 — underside of apical portion of paramere; 11 — tergite 10 of female genital segment." httpUri="https://zenodo.org/record/173980/files/figure.png" pageId="3" pageNumber="44">Fig.1</figureCitation>
). Tergite 10 as in
<figureCitation id="13012A40FFFB9C28F4C3FBAA88567C74" box="[716,788,1044,1070]" captionStart="FIGURES 1 – 11" captionStartId="1.[264,382,1771,1795]" captionTargetBox="[282,1312,334,1749]" captionTargetId="figure@1.[274,1315,329,1751]" captionTargetPageId="1" captionText="FIGURES 1 – 11. Details of Sphaerobulbus biplagiatus (1 – 5) and S. ornatus (6 – 11). 1, 6 — sternite 9 of male genital segment; 2, 7 — tergite 10 of male genital segment; 3, 8 — aedoeagus, ventral view; 4, 9 — apical portion of median lobe, ventral view, paramere removed; 5, 10 — underside of apical portion of paramere; 11 — tergite 10 of female genital segment." httpUri="https://zenodo.org/record/173980/files/figure.png" pageId="3" pageNumber="44">Fig. 2</figureCitation>
, moderately large, with arcuate apex bearing long apical setae, otherwise moderately densely setose. Aedoeagus as in
<figureCitation id="13012A40FFFB9C28F256FB828F8B7C0C" box="[1113,1225,1084,1110]" captionStart="FIGURES 1 – 11" captionStartId="1.[264,382,1771,1795]" captionTargetBox="[282,1312,334,1749]" captionTargetId="figure@1.[274,1315,329,1751]" captionTargetPageId="1" captionText="FIGURES 1 – 11. Details of Sphaerobulbus biplagiatus (1 – 5) and S. ornatus (6 – 11). 1, 6 — sternite 9 of male genital segment; 2, 7 — tergite 10 of male genital segment; 3, 8 — aedoeagus, ventral view; 4, 9 — apical portion of median lobe, ventral view, paramere removed; 5, 10 — underside of apical portion of paramere; 11 — tergite 10 of female genital segment." httpUri="https://zenodo.org/record/173980/files/figure.png" pageId="3" pageNumber="44">Figs. 3–5</figureCitation>
; median lobe with subparallel­sided middle portion, with rather short, triangular apical portion with acute apex; paramere situated on median lobe asymmetrically, with arcuate apex not quite reaching apex of median lobe; underside of paramere without black sensory peg setae; with two subapical setae at right margin, and with three curved spine­like setae (
<figureCitation id="13012A40FFFB9C28F2B9FB628FB57CAC" box="[1206,1271,1244,1270]" captionStart="FIGURES 1 – 11" captionStartId="1.[264,382,1771,1795]" captionTargetBox="[282,1312,334,1749]" captionTargetId="figure@1.[274,1315,329,1751]" captionTargetPageId="1" captionText="FIGURES 1 – 11. Details of Sphaerobulbus biplagiatus (1 – 5) and S. ornatus (6 – 11). 1, 6 — sternite 9 of male genital segment; 2, 7 — tergite 10 of male genital segment; 3, 8 — aedoeagus, ventral view; 4, 9 — apical portion of median lobe, ventral view, paramere removed; 5, 10 — underside of apical portion of paramere; 11 — tergite 10 of female genital segment." httpUri="https://zenodo.org/record/173980/files/figure.png" pageId="3" pageNumber="44">Fig 5</figureCitation>
).
</paragraph>
<paragraph id="8B8536C5FFFB9C28F737FABA895B7D44" blockId="3.[264,1325,284,1870]" box="[312,537,1284,1310]" pageId="3" pageNumber="44">Female. Unknown.</paragraph>
<paragraph id="8B8536C5FFFB9C28F737FA9288187D1C" blockId="3.[264,1325,284,1870]" box="[312,858,1324,1350]" pageId="3" pageNumber="44">Length 14.0 mm (abdomen slightly extended).</paragraph>
<paragraph id="8B8536C5FFFB9C28F737FAEA89B47DCC" blockId="3.[264,1325,284,1870]" pageId="3" pageNumber="44">
<emphasis id="B94EEAD7FFFB9C28F737FAEA89DB7D34" bold="true" box="[312,665,1364,1390]" pageId="3" pageNumber="44">Geographgical distribution.</emphasis>
<taxonomicName id="4C3A4D46FFFB9C28F4A9FAEA88A47D37" box="[678,998,1364,1389]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="biplagiatus">
<emphasis id="B94EEAD7FFFB9C28F4A9FAEA88A47D37" box="[678,998,1364,1389]" italics="true" pageId="3" pageNumber="44">Sphaerobulbus biplagiatus</emphasis>
</taxonomicName>
is at present known only from the
<typeStatus id="54818867FFFB9C28F77CFAC28AE47DCC" box="[371,422,1404,1430]" pageId="3" pageNumber="44">type</typeStatus>
locality in northern Yunnan.
</paragraph>
<paragraph id="8B8536C5FFFB9C28F737FA1A8FF17DE4" blockId="3.[264,1325,284,1870]" box="[312,1203,1444,1470]" pageId="3" pageNumber="44">
<emphasis id="B94EEAD7FFFB9C28F737FA1A8A807DE4" bold="true" box="[312,450,1444,1470]" pageId="3" pageNumber="44">Bionomics.</emphasis>
Nothing is known about the habitat requirements of this species
</paragraph>
<paragraph id="8B8536C5FFFB9C2FF737FA728E4A7904" blockId="3.[264,1325,284,1870]" lastBlockId="4.[264,1323,284,470]" lastPageId="4" lastPageNumber="45" pageId="3" pageNumber="44">
<emphasis id="B94EEAD7FFFB9C28F737FA7289D77DBC" bold="true" box="[312,661,1484,1510]" pageId="3" pageNumber="44">Recognition and comments.</emphasis>
<taxonomicName id="4C3A4D46FFFB9C28F492FA72889B7DBF" box="[669,985,1484,1509]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="biplagiatus">
<emphasis id="B94EEAD7FFFB9C28F492FA72889B7DBF" box="[669,985,1484,1509]" italics="true" pageId="3" pageNumber="44">Sphaerobulbus biplagiatus</emphasis>
</taxonomicName>
shares the character state of a patch of golden tomentose pubescence in the middle of the visible abdominal tergites 4 and 5 with two species, but it differs from them by other morphological characters, as well as geographically. It is most similar to
<taxonomicName id="4C3A4D46FFFB9C28F4D9F9FA8F4D7E04" authority="Smetana, 2003" authorityName="Smetana" authorityYear="2003" box="[726,1039,1604,1630]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="brezinai">
<emphasis id="B94EEAD7FFFB9C28F4D9F9FA88167E07" box="[726,852,1604,1629]" italics="true" pageId="3" pageNumber="44">S. brezinai</emphasis>
<bibRefCitation id="EFAB4B34FFFB9C28F553F9FA8F4D7E04" author="Smetana" box="[860,1039,1604,1630]" pageId="3" pageNumber="47" refString="Smetana, A. (2003) Contributions to the knowledge of the genera of the &quot; Staphylinus - complex &quot; (Coleoptera: Staphylinidae) of China. Part 4. Key to Chinese genera, treatment of the genera Collocypus gen. n., Ocychinus gen. n., Sphaerobulbus gen. n., Aulacocypus and Apecholinus, and comments on the genus Protocypus. Folia Heyrovskyana, 11, 57 - 135." type="journal article" year="2003">Smetana, 2003</bibRefCitation>
</taxonomicName>
, but the latter species is larger with longer antennae and not quite as short elytra, and the aedoeagus is shaped differently (see Figs.
<quantity id="4CC29B20FFFB9C28F41DF92A893D7EF4" box="[530,639,1684,1710]" metricMagnitude="-1" metricUnit="m" metricValue="8.382" metricValueMax="8.636" metricValueMin="8.128" pageId="3" pageNumber="44" unit="in" value="33.0" valueMax="34.0" valueMin="32.0">32–34 in</quantity>
<bibRefCitation id="EFAB4B34FFFB9C28F483F92A88067EF4" author="Smetana" box="[652,836,1684,1710]" pageId="3" pageNumber="47" refString="Smetana, A. (2003) Contributions to the knowledge of the genera of the &quot; Staphylinus - complex &quot; (Coleoptera: Staphylinidae) of China. Part 4. Key to Chinese genera, treatment of the genera Collocypus gen. n., Ocychinus gen. n., Sphaerobulbus gen. n., Aulacocypus and Apecholinus, and comments on the genus Protocypus. Folia Heyrovskyana, 11, 57 - 135." type="journal article" year="2003">Smetana, 2003</bibRefCitation>
); in addition,
<taxonomicName id="4C3A4D46FFFB9C28F5FAF92A8F357EF7" box="[1013,1143,1684,1709]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="brezinai">
<emphasis id="B94EEAD7FFFB9C28F5FAF92A8F357EF7" box="[1013,1143,1684,1709]" italics="true" pageId="3" pageNumber="44">S. brezinai</emphasis>
</taxonomicName>
is endemic to Yulongshan. The other species,
<taxonomicName id="4C3A4D46FFFB9C28F473F90288AA7E8C" authority="Smetana, 2005" authorityName="Smetana" authorityYear="2005" box="[636,1000,1724,1750]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="businskorum">
<emphasis id="B94EEAD7FFFB9C28F473F90288727E8F" box="[636,816,1724,1749]" italics="true" pageId="3" pageNumber="44">S. businskorum</emphasis>
Smetana, 2005
</taxonomicName>
, is on average considerably larger (14.0–19.0 mm) with forebody distinctly shinier (due to the lack of microsculpture on interspaces between punctures on head and pronotum), with scutellum finely and densely punctate and setose, and with the differently shaped aedoeagus (see Figs.
<quantity id="4CC29B20FFFB9C28F2CBF88A8E697F14" box="[1220,1323,1844,1870]" metricMagnitude="-1" metricUnit="m" metricValue="4.064" metricValueMax="4.318" metricValueMin="3.81" pageId="3" pageNumber="44" unit="in" value="16.0" valueMax="17.0" valueMin="15.0">15–17 in</quantity>
<bibRefCitation id="EFAB4B34FFFC9C2FF707FEA28A8C796C" author="Smetana" box="[264,462,284,310]" pageId="4" pageNumber="47" refString="Smetana, A. (2005 b) Contributions to the knowledge of the genera of the &quot; Staphylinus - complex &quot; (Coleoptera: Staphylinidae) of China. Part 7. The genus Sphaerobulbus Smetana 2003. Section 2. Zootaxa, 1006, 53 - 64." type="journal article" year="2005" yearSuffix="b">Smetana, 2005b</bibRefCitation>
); in addition,
<taxonomicName id="4C3A4D46FFFC9C2FF470FEA28875796F" box="[639,823,284,309]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="businskorum">
<emphasis id="B94EEAD7FFFC9C2FF470FEA28875796F" box="[639,823,284,309]" italics="true" pageId="4" pageNumber="45">S. businskorum</emphasis>
</taxonomicName>
is at present known only from the high elevations of mountain ranges of Daxue Shan and Shalui Shan in west­central Sichuan.
</paragraph>
</subSubSection>
<subSubSection id="C320654EFFFC9C2FF737FED5883E798C" pageId="4" pageNumber="45" type="etymology">
<paragraph id="8B8536C5FFFC9C2FF737FED5883E798C" blockId="4.[264,1323,284,470]" pageId="4" pageNumber="45">
<emphasis id="B94EEAD7FFFC9C2FF737FED58A8679DF" bold="true" box="[312,452,363,389]" pageId="4" pageNumber="45">Etymology.</emphasis>
The specific epithet is a combination of the Latin adverb
<emphasis id="B94EEAD7FFFC9C2FF275FED28FDE79DF" box="[1146,1180,364,389]" italics="true" pageId="4" pageNumber="45">bis</emphasis>
(twice) and the Latin adjective
<emphasis id="B94EEAD7FFFC9C2FF7FEFE2A891C79F7" box="[497,606,404,429]" italics="true" pageId="4" pageNumber="45">plagiatus</emphasis>
, ­
<emphasis id="B94EEAD7FFFC9C2FF476FE2A89CA79F7" box="[633,648,404,429]" italics="true" pageId="4" pageNumber="45">a</emphasis>
, ­
<emphasis id="B94EEAD7FFFC9C2FF4ACFE2A898479F7" box="[675,710,404,429]" italics="true" pageId="4" pageNumber="45">um</emphasis>
(crooked, marked). It refers to the presence of the pair of yellow tomentose spots on abdominal tergites.
</paragraph>
</subSubSection>
</treatment>
</document>
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