256 lines
29 KiB
XML
256 lines
29 KiB
XML
<document ID-DOI="10.5281/zenodo.163451" ID-GBIF-Dataset="f9566778-1df4-44d6-9a4c-849de39a4b6c" ID-GBIF-Taxon="124955158" ID-Zenodo-Dep="163451" checkinTime="1480975261095" checkinUser="plazi" docAuthor="Vincent J. Kalkman" docDate="2016" docId="03938796FFD2FFFE26B68B28FE6DFA9E" docLanguage="en" docName="10.5281zenodo.163451_Kalkman.pdf" docOrigin="Odonatologica 45 (3)" docStyle="DocumentStyle{}" docTitle="Celebargiolestes askewi Vincent J. Kalkman 2016, sp. nov." docType="treatment" docVersion="11" lastPageId="16" lastPageNumber="251" masterDocId="FFAAFFEEFFDEFFEE27258934FFCBFF99" masterDocTitle="Revision of the genus Celebargiolestes Kennedy, 1925 (Odonata: Argiolestidae)" masterLastPageNumber="269" masterPageNumber="235" pageId="12" pageNumber="247" updateTime="1643502553511" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of the genus Celebargiolestes Kennedy, 1925 (Odonata: Argiolestidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Vincent J. Kalkman</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Odonatologica</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2016</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2016-12-01</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>45</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>3</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>235</mods:start>
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<mods:end>269</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.5281/zenodo.163451</mods:identifier>
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<mods:identifier type="GBIF-Dataset">f9566778-1df4-44d6-9a4c-849de39a4b6c</mods:identifier>
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<mods:identifier type="Zenodo-Dep">163451</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="124955158" LSID="urn:lsid:plazi:treatment:03938796FFD2FFFE26B68B28FE6DFA9E" httpUri="http://treatment.plazi.org/id/03938796FFD2FFFE26B68B28FE6DFA9E" lastPageId="16" lastPageNumber="251" pageId="12" pageNumber="247">
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<subSubSection box="[403,792,540,568]" pageId="12" pageNumber="247" type="nomenclature">
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<paragraph blockId="12.[159,1036,540,769]" box="[403,792,540,568]" pageId="12" pageNumber="247">
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<heading bold="true" box="[403,792,540,568]" centered="true" fontSize="11" level="2" pageId="12" pageNumber="247" reason="0">
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<emphasis bold="true" box="[403,792,540,568]" pageId="12" pageNumber="247">
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<taxonomicName authority="Vincent J. Kalkman, 2016" authorityName="Vincent J. Kalkman" authorityYear="2016" box="[403,693,540,568]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="247" phylum="Arthropoda" rank="species" species="askewi" status="sp. nov.">
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<emphasis bold="true" box="[403,693,540,568]" italics="true" pageId="12" pageNumber="247">Celebargiolestes askewi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[699,792,540,568]" pageId="12" pageNumber="247" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[437,757,574,602]" pageId="12" pageNumber="247" type="description">
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<paragraph blockId="12.[159,1036,540,769]" box="[437,757,574,602]" pageId="12" pageNumber="247">
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<heading box="[437,757,574,602]" centered="true" fontSize="11" level="4" pageId="12" pageNumber="247" reason="1">
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(
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<figureCitation box="[444,531,574,602]" captionStart="Figures 7 – 14" captionStartId="13.[159,238,1384,1409]" captionTargetBox="[196,998,249,1341]" captionTargetId="figure@13.[196,998,249,1342]" captionTargetPageId="13" captionText="Figures 7 – 14. Head of Celebargiolestes males. 7 – C. cinctus (immature), Ranotongkor, 02 - viii- 1940, JvT 6476; 8 – C. cinctus (old adult), Mapanget, 23 - vi- 1940, JvT 6638; 9 – C. cinctus (‘ red legs’ form), 25 km NE of Luwuk, 30 - i- 1989, JvT 6670; 10 – C. askewi sp. nov., holotype; 11 – C. toli sp. nov., holotype; 12 – C. toli sp. nov., Peleng Island, 05 - viii- 1941, JvT 6453; 13 – C. orri sp. nov., holotype; 14 – C. orri sp. nov., 10 km WNW Palopo, 29 - x- 1993, JvT 11910." httpUri="https://zenodo.org/record/192855/files/figure.png" pageId="12" pageNumber="247">Figs 10</figureCitation>
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,
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<figureCitation box="[542,575,574,602]" captionStart="Figures 15 – 18" captionStartId="14.[159,238,1441,1466]" captionTargetBox="[203,991,248,1401]" captionTargetId="figure@14.[203,991,248,1401]" captionTargetPageId="14" captionText="Figures 15 – 18. Body of Celebargiolestes males. 15 – C. cinctus (immature), Ranotongkor, 02 - viii- 1940, JvT 6476; 16 – C. cinctus (old adult), Mapanget, 23 - vi- 1940, JvT 6638; 17 – C. cinctus (‘ red legs’ form), 25 km NE of Luwuk, 30 - i- 1989, JvT 6670; 18 – C. askewi sp. nov., holotype." httpUri="https://zenodo.org/record/192856/files/figure.png" pageId="12" pageNumber="247">18</figureCitation>
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,
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<figureCitation box="[586,664,574,602]" captionStart="Figures 23 – 27" captionStartId="25.[159,237,1412,1437]" captionTargetBox="[208,986,249,1370]" captionTargetId="figure@25.[208,986,249,1370]" captionTargetPageId="25" captionText="Figures 23 – 27. Genital ligula of Celebargiolestes males. 23 – C. cinctus, Dumoga Bone NP, 04 - vi- 1985, JvT 6486, lateral view; 24 – C. cinctus, Dumoga Bone NP, 04 - vi- 1985, JvT 6486, ventral view; 25 – C. cinctus ‘ red leg’, Banggai Peninsula, 08 - x- 1989, latero-ventral view; 26 – C. askewi sp. nov., Polewali, 23 - x- 1940, JvT 6636, lateral view; 27 – C. askewi sp. nov., Polewali, 23 - x- 1940, JvT 6636, ventral view." httpUri="https://zenodo.org/record/192858/files/figure.png" pageId="12" pageNumber="247">26–27</figureCitation>
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,
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<figureCitation box="[674,751,574,602]" captionStart="Figures 34 – 39" captionStartId="27.[159,238,1413,1438]" captionTargetBox="[156,1038,480,1372]" captionTargetId="figure@27.[156,1038,480,1372]" captionTargetPageId="27" captionText="Figures 34 – 39. Appendages of Celebargiolestes males. 34 – C. cinctus, Ranotongkor, 02 - viii- 1940, JvT 6476, dorsal view; 35 – C. cinctus, Ranotongkor, 02 - viii- 1940, JvT 6476, lateral view; 36 – Ranotongkor, 02 - viii- 1940, JvT 6476, ventral view; 37 – C. askewi sp. nov., holotype, dorsal view; 38 – C. askewi sp. nov., holotype, lateral view; 39 – C. askewi sp. nov., holotype, ventral view." httpUri="https://zenodo.org/record/192860/files/figure.png" pageId="12" pageNumber="247">37–39</figureCitation>
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)
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="12" pageNumber="247" type="materials_examined">
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<paragraph blockId="12.[159,1036,540,769]" box="[159,360,620,646]" pageId="12" pageNumber="247">
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<heading bold="true" box="[159,360,620,646]" fontSize="11" level="3" pageId="12" pageNumber="247" reason="0">
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<emphasis bold="true" box="[159,360,620,646]" pageId="12" pageNumber="247">Material studied</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="12.[159,1036,540,769]" pageId="12" pageNumber="247">
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<materialsCitation ID-GBIF-Occurrence="1413786764" collectingDate="1940-10-23" collectionCode="RMNH" collectorName="J. J. van der Starre" country="Indonesia" location="Polewali" municipality="Polewali" pageId="12" pageNumber="247" specimenCode="JvT6492" specimenCount="1" specimenCount-male="1" stateProvince="Sulawesi" typeStatus="holotype">
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<typeStatus box="[159,263,653,677]" pageId="12" pageNumber="247">
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<emphasis bold="true" box="[159,263,653,677]" pageId="12" pageNumber="247">Holotype</emphasis>
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</typeStatus>
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<specimenCount box="[269,289,653,676]" pageId="12" pageNumber="247" type="male">♂</specimenCount>
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.
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<collectingCountry box="[303,410,653,677]" name="Indonesia" pageId="12" pageNumber="247">Indonesia</collectingCountry>
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,
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<collectingRegion box="[421,514,653,677]" country="Indonesia" name="Sulawesi Utara" pageId="12" pageNumber="247">Sulawesi</collectingRegion>
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,
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<collectingMunicipality box="[525,617,653,677]" pageId="12" pageNumber="247">Polewali</collectingMunicipality>
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,
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<specimenCode box="[627,718,653,677]" pageId="12" pageNumber="247">JvT6492</specimenCode>
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,
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<collectingDate box="[729,840,653,677]" pageId="12" pageNumber="247" value="1940-10-23">
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<date box="[729,840,653,677]" pageId="12" pageNumber="247" value="1940-10-23">23-x-1940</date>
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</collectingDate>
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,
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<collectorName box="[850,1036,653,677]" pageId="12" pageNumber="247">J.J. van der Starre</collectorName>
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leg., (
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<collectionCode box="[214,301,684,708]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum ("Naturalis") [formerly Rijksmuseum van Natuurlijke Historie]" pageId="12" pageNumber="247">RMNH</collectionCode>
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). Handwritten text on envelope: »Celebes, Polewali,
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<date box="[847,975,684,708]" pageId="12" pageNumber="247" value="1940-10-23">23 Oct 1940</date>
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«, text made by stamp: »J.J. van der Starre«.
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</materialsCitation>
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</paragraph>
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<paragraph blockId="12.[159,1036,540,769]" box="[159,1036,745,769]" pageId="12" pageNumber="247">
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<emphasis bold="true" box="[159,351,745,769]" pageId="12" pageNumber="247">Further material.</emphasis>
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Sulawesi Barat 5♂ 3♀, Polewali, 23-x-1940, J.J van der Starre leg.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="12" pageNumber="247" type="etymology">
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<paragraph blockId="12.[159,1036,818,919]" box="[159,292,818,844]" pageId="12" pageNumber="247">
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<heading bold="true" box="[159,292,818,844]" fontSize="11" level="3" pageId="12" pageNumber="247" reason="0">
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<emphasis bold="true" box="[159,292,818,844]" pageId="12" pageNumber="247">Etymology</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="12.[159,1036,818,919]" pageId="12" pageNumber="247">The species is dedicated to Richard R. Askew (masculine singular genitive noun).</paragraph>
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</subSubSection>
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<subSubSection lastPageId="16" lastPageNumber="251" pageId="12" pageNumber="247" type="description">
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<paragraph blockId="12.[159,1036,967,1292]" box="[159,349,967,994]" pageId="12" pageNumber="247">
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<heading box="[159,349,967,994]" fontSize="11" level="5" pageId="12" pageNumber="247" reason="1">
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<emphasis bold="true" box="[159,220,968,994]" pageId="12" pageNumber="247">Male</emphasis>
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(holotype)
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</heading>
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</paragraph>
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<paragraph blockId="12.[159,1036,967,1292]" pageId="12" pageNumber="247">
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Head − Anterior ⅔ of labium dark brown, posterior third yellow-brown. Labrum dull brown-black; front of head, including mandibles and sockets of antennae dull yellow-orange (ochre), anteclypeus pale brown, postclypeus dull orange with a median black mark, which is broad in front and becomes smaller towards frons, area in front of antennae black, more constricted smaller towards the clypeus and there connected with the black on the postclypeus (
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<figureCitation box="[312,396,1228,1255]" captionStart="Figures 7 – 14" captionStartId="13.[159,238,1384,1409]" captionTargetBox="[196,998,249,1341]" captionTargetId="figure@13.[196,998,249,1342]" captionTargetPageId="13" captionText="Figures 7 – 14. Head of Celebargiolestes males. 7 – C. cinctus (immature), Ranotongkor, 02 - viii- 1940, JvT 6476; 8 – C. cinctus (old adult), Mapanget, 23 - vi- 1940, JvT 6638; 9 – C. cinctus (‘ red legs’ form), 25 km NE of Luwuk, 30 - i- 1989, JvT 6670; 10 – C. askewi sp. nov., holotype; 11 – C. toli sp. nov., holotype; 12 – C. toli sp. nov., Peleng Island, 05 - viii- 1941, JvT 6453; 13 – C. orri sp. nov., holotype; 14 – C. orri sp. nov., 10 km WNW Palopo, 29 - x- 1993, JvT 11910." httpUri="https://zenodo.org/record/192855/files/figure.png" pageId="12" pageNumber="247">Fig. 10</figureCitation>
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). Antennae orange-brown, top of head black with pale oval spot between antennae and lateral ocelli.
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</paragraph>
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<paragraph blockId="12.[159,1036,1340,1554]" pageId="12" pageNumber="247">
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Thorax − Sides of prothorax dull orange with a posterior black mark; anteri- or and middle lobe largely black but with the sides dull yellow-orange. Posterior lobe of pronotum black with outer border dull yellow-orange, seen from lateral gently sloping and with the outer border steeply raised, lateral corners rounded. Pattern of thorax as in
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<figureCitation box="[636,749,1490,1516]" captionStart="Figures 15 – 18" captionStartId="14.[159,238,1441,1466]" captionTargetBox="[203,991,248,1401]" captionTargetId="figure@14.[203,991,248,1401]" captionTargetPageId="14" captionText="Figures 15 – 18. Body of Celebargiolestes males. 15 – C. cinctus (immature), Ranotongkor, 02 - viii- 1940, JvT 6476; 16 – C. cinctus (old adult), Mapanget, 23 - vi- 1940, JvT 6638; 17 – C. cinctus (‘ red legs’ form), 25 km NE of Luwuk, 30 - i- 1989, JvT 6670; 18 – C. askewi sp. nov., holotype." httpUri="https://zenodo.org/record/192856/files/figure.png" pageId="12" pageNumber="247">Figure 18</figureCitation>
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; darker areas black, pale areas yellow-orange.
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</paragraph>
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<caption httpUri="https://zenodo.org/record/192855/files/figure.png" pageId="13" pageNumber="248" targetBox="[196,998,249,1341]" targetPageId="13">
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<paragraph blockId="13.[159,1037,1383,1555]" pageId="13" pageNumber="248">
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<emphasis bold="true" box="[159,307,1384,1409]" pageId="13" pageNumber="248">Figures 7–14.</emphasis>
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Head of
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<taxonomicName authorityName="Kennedy" authorityYear="1925" box="[412,588,1383,1407]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="248" phylum="Arthropoda" rank="genus">
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<emphasis box="[412,588,1383,1407]" italics="true" pageId="13" pageNumber="248">Celebargiolestes</emphasis>
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</taxonomicName>
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males. 7 –
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<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1886" box="[721,825,1383,1407]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="248" phylum="Arthropoda" rank="species" species="cinctus">
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<emphasis box="[721,825,1383,1407]" italics="true" pageId="13" pageNumber="248">C. cinctus</emphasis>
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</taxonomicName>
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(immature), Ranotongkor, 02-viii-1940, JvT6476; 8 –
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<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1886" box="[545,649,1413,1437]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="248" phylum="Arthropoda" rank="species" species="cinctus">
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<emphasis box="[545,649,1413,1437]" italics="true" pageId="13" pageNumber="248">C. cinctus</emphasis>
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</taxonomicName>
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(old adult), Mapanget, 23-vi-1940, JvT6638; 9 –
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<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1886" box="[301,404,1442,1466]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="248" phylum="Arthropoda" rank="species" species="cinctus">
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<emphasis box="[301,404,1442,1466]" italics="true" pageId="13" pageNumber="248">C. cinctus</emphasis>
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</taxonomicName>
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(‘red legs’ form), 25 km NE of Luwuk, 30-i-1989, JvT6670; 10 –
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<taxonomicName authority="Vincent J. Kalkman, 2016" authorityName="Vincent J. Kalkman" authorityYear="2016" box="[209,308,1471,1495]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="248" phylum="Arthropoda" rank="species" species="askewi" status="sp. nov.">
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<emphasis box="[209,308,1471,1495]" italics="true" pageId="13" pageNumber="248">C. askewi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[313,398,1471,1496]" pageId="13" pageNumber="248" rank="species">sp. nov.</taxonomicNameLabel>
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, holotype; 11 –
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<taxonomicName authority="Vincent J. Kalkman, 2016" authorityName="Vincent J. Kalkman" authorityYear="2016" box="[565,626,1471,1495]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="248" phylum="Arthropoda" rank="species" species="toli" status="sp. nov.">
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<emphasis box="[565,626,1471,1495]" italics="true" pageId="13" pageNumber="248">C. toli</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[631,716,1471,1496]" pageId="13" pageNumber="248" rank="species">sp. nov.</taxonomicNameLabel>
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, holotype; 12 –
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<taxonomicName authority="Vincent J. Kalkman, 2016" authorityName="Vincent J. Kalkman" authorityYear="2016" box="[883,943,1471,1495]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="248" phylum="Arthropoda" rank="species" species="toli" status="sp. nov.">
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<emphasis box="[883,943,1471,1495]" italics="true" pageId="13" pageNumber="248">C. toli</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[948,1033,1471,1496]" pageId="13" pageNumber="248" rank="species">sp. nov.</taxonomicNameLabel>
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, Peleng Island, 05-viii-1941, JvT6453; 13 –
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<taxonomicName authority="Vincent J. Kalkman, 2016" authorityName="Vincent J. Kalkman" authorityYear="2016" box="[607,672,1501,1525]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="248" phylum="Arthropoda" rank="species" species="orri" status="sp. nov.">
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<emphasis box="[607,672,1501,1525]" italics="true" pageId="13" pageNumber="248">C. orri</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[678,764,1501,1526]" pageId="13" pageNumber="248" rank="species">sp. nov.</taxonomicNameLabel>
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, holotype; 14 –
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<taxonomicName authority="Vincent J. Kalkman, 2016" authorityName="Vincent J. Kalkman" authorityYear="2016" box="[934,998,1501,1525]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="248" phylum="Arthropoda" rank="species" species="orri" status="sp. nov.">
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<emphasis box="[934,998,1501,1525]" italics="true" pageId="13" pageNumber="248">C. orri</emphasis>
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</taxonomicName>
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<taxonomicNameLabel pageId="13" pageNumber="248" rank="species">sp. nov.</taxonomicNameLabel>
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, 10 km WNW Palopo, 29-x-1993, JvT11910.
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</paragraph>
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</caption>
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<caption httpUri="https://zenodo.org/record/192856/files/figure.png" pageId="14" pageNumber="249" targetBox="[203,991,248,1401]" targetPageId="14">
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<paragraph blockId="14.[159,1037,1441,1554]" pageId="14" pageNumber="249">
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<emphasis bold="true" box="[159,319,1441,1466]" pageId="14" pageNumber="249">Figures 15–18.</emphasis>
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Body of
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<taxonomicName authorityName="Kennedy" authorityYear="1925" box="[415,591,1441,1465]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="249" phylum="Arthropoda" rank="genus">
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<emphasis box="[415,591,1441,1465]" italics="true" pageId="14" pageNumber="249">Celebargiolestes</emphasis>
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</taxonomicName>
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males. 15 –
|
||
<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1886" box="[728,830,1441,1465]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="249" phylum="Arthropoda" rank="species" species="cinctus">
|
||
<emphasis box="[728,830,1441,1465]" italics="true" pageId="14" pageNumber="249">C. cinctus</emphasis>
|
||
</taxonomicName>
|
||
(immature), Ranotongkor, 02-viii-1940, JvT6476; 16 –
|
||
<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1886" box="[552,655,1471,1495]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="249" phylum="Arthropoda" rank="species" species="cinctus">
|
||
<emphasis box="[552,655,1471,1495]" italics="true" pageId="14" pageNumber="249">C. cinctus</emphasis>
|
||
</taxonomicName>
|
||
(old adult), Mapanget, 23-vi-1940, JvT6638; 17 –
|
||
<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1886" box="[312,414,1500,1524]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="249" phylum="Arthropoda" rank="species" species="cinctus">
|
||
<emphasis box="[312,414,1500,1524]" italics="true" pageId="14" pageNumber="249">C. cinctus</emphasis>
|
||
</taxonomicName>
|
||
(‘red legs’ form), 25 km NE of Luwuk, 30-i-1989, JvT6670; 18 –
|
||
<taxonomicName authority="Vincent J. Kalkman, 2016" authorityName="Vincent J. Kalkman" authorityYear="2016" box="[211,311,1529,1553]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="249" phylum="Arthropoda" rank="species" species="askewi" status="sp. nov.">
|
||
<emphasis box="[211,311,1529,1553]" italics="true" pageId="14" pageNumber="249">C. askewi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[317,403,1529,1554]" pageId="14" pageNumber="249" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="15.[159,1036,253,430]" pageId="15" pageNumber="250">Legs − Coxae and trochanters dull yellow-orange, femora yellow-orange with two broad but poorly defined black rings, knees black, tibiae and tarsi pale brown, spines pale brown. Femora with respectively 9, 10, 17 spines laterally; tibiae of fore legs with 12 smaller and 6 larger spines laterally; tibiae of mid and hind legs with 12 and 15 spines, respectively laterally.</paragraph>
|
||
<paragraph blockId="15.[159,1036,477,691]" pageId="15" pageNumber="250">Wings − Venation black. Membranes slightly suffused and with a smoky out- er border. Fw and Hw with 2 Ax; Fw with 34–36 Px and Hw with 31–32Px. Ac at about ¼ to ⅓ the distance between Ax1 and Ax2. Brace vein absent. Pterostigma dark brown. Up to three rows of cells between anal vein and bor- der of Fw and up to four in Hw. Fields between Ir2 and R3, between R3 and IR3, and between IR3 and R4 at distal end containing 3 or more rows of cells.</paragraph>
|
||
<paragraph blockId="15.[159,1036,739,1213]" pageId="15" pageNumber="250">
|
||
Abdomen − Pattern as in
|
||
<figureCitation box="[469,583,739,765]" captionStart="Figures 15 – 18" captionStartId="14.[159,238,1441,1466]" captionTargetBox="[203,991,248,1401]" captionTargetId="figure@14.[203,991,248,1401]" captionTargetPageId="14" captionText="Figures 15 – 18. Body of Celebargiolestes males. 15 – C. cinctus (immature), Ranotongkor, 02 - viii- 1940, JvT 6476; 16 – C. cinctus (old adult), Mapanget, 23 - vi- 1940, JvT 6638; 17 – C. cinctus (‘ red legs’ form), 25 km NE of Luwuk, 30 - i- 1989, JvT 6670; 18 – C. askewi sp. nov., holotype." httpUri="https://zenodo.org/record/192856/files/figure.png" pageId="15" pageNumber="250">Figure 18</figureCitation>
|
||
; dark pattern elements dark brown to black, pale pattern elements yellow-brown. Anterior half of S1 yellow, posterior half brown to black, S2 with dorsum black and laterally pale brown, S3–6 dorsally largely dark, ventral and lower lateral areas pale brown, anteriorly on each segment the pale brown reaches to the dorsum forming an inconspicuous brown ring covering approximately ¹/₁0 of the segments. S7–10 black, the latter strongly depressed in the middle, hind border of S10 without modifications. Appendages as shown in
|
||
<figureCitation box="[726,893,1000,1026]" captionStart="Figures 34 – 39" captionStartId="27.[159,238,1413,1438]" captionTargetBox="[156,1038,480,1372]" captionTargetId="figure@27.[156,1038,480,1372]" captionTargetPageId="27" captionText="Figures 34 – 39. Appendages of Celebargiolestes males. 34 – C. cinctus, Ranotongkor, 02 - viii- 1940, JvT 6476, dorsal view; 35 – C. cinctus, Ranotongkor, 02 - viii- 1940, JvT 6476, lateral view; 36 – Ranotongkor, 02 - viii- 1940, JvT 6476, ventral view; 37 – C. askewi sp. nov., holotype, dorsal view; 38 – C. askewi sp. nov., holotype, lateral view; 39 – C. askewi sp. nov., holotype, ventral view." httpUri="https://zenodo.org/record/192860/files/figure.png" pageId="15" pageNumber="250">Figures 37–39</figureCitation>
|
||
; both upper and lower appendages dark brown to black. Lower appendages short, about ¼ the length of upper appendages, slightly down-curved and, when seen from lateral, with tip not reaching level of the upper appendages. Genital ligula as in
|
||
<figureCitation box="[290,457,1149,1175]" captionStart="Figures 23 – 27" captionStartId="25.[159,237,1412,1437]" captionTargetBox="[208,986,249,1370]" captionTargetId="figure@25.[208,986,249,1370]" captionTargetPageId="25" captionText="Figures 23 – 27. Genital ligula of Celebargiolestes males. 23 – C. cinctus, Dumoga Bone NP, 04 - vi- 1985, JvT 6486, lateral view; 24 – C. cinctus, Dumoga Bone NP, 04 - vi- 1985, JvT 6486, ventral view; 25 – C. cinctus ‘ red leg’, Banggai Peninsula, 08 - x- 1989, latero-ventral view; 26 – C. askewi sp. nov., Polewali, 23 - x- 1940, JvT 6636, lateral view; 27 – C. askewi sp. nov., Polewali, 23 - x- 1940, JvT 6636, ventral view." httpUri="https://zenodo.org/record/192858/files/figure.png" pageId="15" pageNumber="250">Figures 26–27</figureCitation>
|
||
, with a broad but shallow apical cleft; lateral lobes large, halfway constricted and with a broader rounded tip.
|
||
</paragraph>
|
||
<paragraph blockId="15.[159,1036,1261,1363]" pageId="15" pageNumber="250">Measurements [mm] − Total length 50, abdomen 40, Fw 35; Fw pterostigma 1.6 (costal length), 2.3 (longest length); Hw pterostigma 1.8 (costal length), 2.6 (longest length).</paragraph>
|
||
<paragraph blockId="15.[159,1036,1411,1549]" box="[159,289,1411,1437]" pageId="15" pageNumber="250">
|
||
<heading bold="true" box="[159,289,1411,1437]" fontSize="11" level="3" pageId="15" pageNumber="250" reason="0">
|
||
<emphasis bold="true" box="[159,289,1411,1437]" pageId="15" pageNumber="250">Variability</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="15.[159,1036,1411,1549]" pageId="15" pageNumber="250">In the type the labrum is dull brown black but in some of the other specimens it is shining metallic black. Fw and Hw with 2 Ax; Fw with 34–38 Px and Hw with 32–34 Px.</paragraph>
|
||
<paragraph blockId="16.[159,1036,253,318]" pageId="16" pageNumber="251">Measurements [mm] − Total length 49–52, abdomen 39–42, Fw 34–36, Hw 35–36.</paragraph>
|
||
<paragraph blockId="16.[159,468,366,429]" box="[159,247,366,392]" pageId="16" pageNumber="251">
|
||
<heading bold="true" box="[159,247,366,392]" fontSize="11" level="3" pageId="16" pageNumber="251" reason="0">
|
||
<emphasis bold="true" box="[159,247,366,392]" pageId="16" pageNumber="251">Female</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="16.[159,468,366,429]" box="[159,468,403,429]" pageId="16" pageNumber="251">
|
||
<heading box="[159,468,403,429]" fontSize="11" level="5" pageId="16" pageNumber="251" reason="1">Head − Pattern as in male.</heading>
|
||
</paragraph>
|
||
<paragraph blockId="16.[159,1036,477,579]" pageId="16" pageNumber="251">Thorax − Pattern on prothorax, synthorax and legs as in male. Fw and Hw with 2 Ax; Fw with 28–33 Px and Hw with 26–29 Px. Pt yellowish, outer border often mottled greyish-black.</paragraph>
|
||
<paragraph blockId="16.[159,1036,627,877]" pageId="16" pageNumber="251">
|
||
Abdomen − Pattern on abdomen largely as in male, dark areas are dark brown to black, pale areas are yellow-brown. S3–9 dorsally largely dark, ventral and the lower lateral areas yellow, anteriorly on each segment the yellow reaches to the dorsum forming a yellow ring covering approximately a fifth to a tenth of the segment. S10 black, cerci brown, base of valvifer yellow, becoming brown to black towards the tip, shaped as in
|
||
<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1886" box="[159,274,851,877]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="251" phylum="Arthropoda" rank="species" species="cinctus">
|
||
<emphasis box="[159,274,851,877]" italics="true" pageId="16" pageNumber="251">C.cinctus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="16.[159,1036,925,989]" pageId="16" pageNumber="251">Measurements [mm] − Total length 41–44, abdomen 32–34, Fw 32–34; Hw 32–34.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="16" pageNumber="251" type="diagnosis">
|
||
<paragraph blockId="16.[159,1036,1037,1288]" box="[159,424,1037,1063]" pageId="16" pageNumber="251">
|
||
<heading bold="true" box="[159,424,1037,1063]" fontSize="11" level="3" pageId="16" pageNumber="251" reason="0">
|
||
<emphasis bold="true" box="[159,424,1037,1063]" pageId="16" pageNumber="251">Differential diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="16.[159,1036,1037,1288]" pageId="16" pageNumber="251">
|
||
Males are distinguished from other species of
|
||
<taxonomicName authorityName="Kennedy" authorityYear="1925" box="[705,884,1075,1101]" class="Insecta" family="Megapodagrionidae" genus="Celebargiolestes" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="251" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[705,884,1075,1101]" italics="true" pageId="16" pageNumber="251">Celebargiolestes</emphasis>
|
||
</taxonomicName>
|
||
by the combination of the following characters: (i) labrum black; (ii) postclypeus and frons largely black; (iii) ligula with a shallow apical cleft; (iv) lower appendages short, about ¼ the length of the upper appendages, slightly downcurved and, when seen from lateral, with the tip not reaching the level of the upper appendages.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |