treatments-xml/data/03/A4/8E/03A48E76DC67FFE5F56235E4FE20FBD9.xml

<document id="339C6162441047AE3F552B8346C24A91" ID-DOI="10.5281/zenodo.10114370" ID-ISSN="0253-116X" ID-Zenodo-Dep="10114370" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630359721084" checkinUser="valdenar" docAuthor="Assing, V." docDate="2007" docId="03A48E76DC67FFE5F56235E4FE20FBD9" docLanguage="en" docName="LinzerbiolBeitr.2007.39.2.777-790.pdf" docOrigin="Linzer biologische Beiträge 39 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Geostiba spinosula Assing, 2007, sp.nov." docType="treatment" docVersion="4" lastPageNumber="789" masterDocId="FF9DF60EDC6DFFE9F439306EFF90FF83" masterDocTitle="Four new species and additional records of Geostiba from Turkey and Crete, and a new synonymy (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="790" masterPageNumber="777" pageNumber="787" updateTime="1700278669180" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="DFC419F963284614597D236410B4A7A5">Four new species and additional records of Geostiba from Turkey and Crete, and a new synonymy (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<mods:date id="9A1D819E7F2EBA2C5EDC35E1DF96C55F">2007</mods:date>
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<subSubSection id="C3176CEBDC67FFE3F56235E4FCD5FA23" box="[347,837,1418,1440]" pageId="10" pageNumber="787" type="nomenclature">
<paragraph id="8BB23F60DC67FFE3F56235E4FCD5FA23" blockId="10.[347,837,1418,1440]" box="[347,837,1418,1440]" pageId="10" pageNumber="787">
<heading id="D0FA880CDC67FFE3F56235E4FCD5FA23" box="[347,837,1418,1440]" fontSize="8" level="2" pageId="10" pageNumber="787" reason="2">
<emphasis id="B979E372DC67FFE3F56235E4FDE3FA23" bold="true" box="[347,627,1418,1440]" pageId="10" pageNumber="787">
<taxonomicName id="4C0D44E3DC67FFE3F56235E4FDB1FA23" ID-CoL="3FSNQ" authorityName="Assing" authorityYear="2007" box="[347,545,1418,1440]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="787" phylum="Arthropoda" rank="species" species="spinosula" status="sp. nov.">
<emphasis id="B979E372DC67FFE3F56235E4FDB1FA23" bold="true" box="[347,545,1418,1440]" italics="true" pageId="10" pageNumber="787">Geostiba spinosula</emphasis>
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<taxonomicNameLabel id="A24A5E09DC67FFE3F61E35E4FDE3FA23" box="[551,627,1418,1440]" pageId="10" pageNumber="787" rank="species">sp.nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="133623E5DC67FFE3F6B935E4FD61FA23" box="[640,753,1418,1440]" captionStart="Figs 36-42" captionStartId="11.[347,387,1604,1624]" captionTargetBox="[347,1247,771,1587]" captionTargetId="figure-141@11.[347,1260,1116,1593]" captionTargetPageId="11" captionText="Figs 36-42: Geostiba spinosula nov.sp.: (36) male pronotum, elytra, and abdominal segment III (holotype); (37) head in lateral view; (38-39) median lobe of aedeagus in lateral and in ventral view; (40) apical lobe of paramere; (41) female sternite VIII; (42) spermatheca. Scale bars: 36: 0.5 mm; 37: 0.2 mm; 38-42: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10114386" httpUri="https://zenodo.org/record/10114386/files/figure.png" pageId="10" pageNumber="787">Figs 36-42</figureCitation>
, 
<figureCitation id="133623E5DC67FFE3F6C235E5FCABFA23" box="[763,827,1418,1440]" captionStart="Map 1" captionStartId="5.[347,392,897,917]" captionTargetBox="[347,1254,432,897]" captionTargetId="figure-7@5.[347,1253,433,889]" captionTargetPageId="5" captionText="Map 1: Distributions of Geostiba janbellini nov.sp. (filled circle), G. atrioculata nov.sp. (open circle), and G. spinosula nov.sp. (square) in southern Anatolia." figureDoi="http://doi.org/10.5281/zenodo.10114376" httpUri="https://zenodo.org/record/10114376/files/figure.png" pageId="10" pageNumber="787">Map 1</figureCitation>
)
</heading>
</paragraph>
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<subSubSection id="C3176CEBDC67FFE5F56235DFFE20FBD9" lastPageId="12" lastPageNumber="789" pageId="10" pageNumber="787" type="materials_examined">
<paragraph id="8BB23F60DC67FFE3F56235DFFC9CFA71" blockId="10.[347,1255,1457,1522]" pageId="10" pageNumber="787">
<materialsCitation id="3B65353DDC67FFE3F56235DFFDBAFA58" collectingDate="2007-04-21" collectorName="Meybohm &amp; Brachat" elevation="1200" location="Bekdemir" municipality="Bahce" pageId="10" pageNumber="787" specimenCount="1" stateProvince="Osmaniye" typeStatus="holotype">
<typeStatus id="54B681C2DC67FFE3F56235DFFE20FA46" box="[347,432,1457,1477]" pageId="10" pageNumber="787" type="holotype">Holotype</typeStatus>
: N37°15'53&quot;, E36°36' (2), TR 
<collectingRegion id="49C9F182DC67FFE3F6DA35DFFCAAFA46" box="[739,826,1457,1477]" country="Turkey" name="Osmaniye" pageId="10" pageNumber="787">Osmanije</collectingRegion>
n. 
<collectingMunicipality id="6BD6A51ADC67FFE3F76E35DFFC02FA46" box="[855,914,1457,1477]" pageId="10" pageNumber="787">Bahce</collectingMunicipality>
, 
<location id="8ED269BBDC67FFE3F7A235DFFC67FA46" LSID="urn:lsid:plazi:treatment:03A48E76DC67FFE5F56235E4FE20FBD9:8ED269BBDC67FFE3F7A235DFFC67FA46" box="[923,1015,1457,1477]" municipality="Bahce" name="Bekdemir" pageId="10" pageNumber="787" stateProvince="Osmaniye">Bekdemir</location>
, 
<quantity id="4CF59285DC67FFE3F03835DFFBD8FA46" box="[1025,1096,1457,1477]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="10" pageNumber="787" unit="m" value="1200.0">
<elevation id="0020D853DC67FFE3F03835DFFBD8FA46" box="[1025,1096,1457,1477]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="10" pageNumber="787" unit="m" value="1200.0">1200 m</elevation>
</quantity>
, 
<date id="FFB319A0DC67FFE3F06B35DFFB2CFA46" box="[1106,1212,1457,1477]" pageId="10" pageNumber="787" value="2007-04-21">
<collectingDate id="EFF7E048DC67FFE3F06B35DFFB2CFA46" box="[1106,1212,1457,1477]" pageId="10" pageNumber="787" value="2007-04-21">21.IV.2007</collectingDate>
</date>
, leg. 
<collectorName id="26F85AB6DC67FFE3F56235A9FE28FA58" box="[347,440,1479,1499]" pageId="10" pageNumber="787">Meybohm</collectorName>
&amp; 
<collectorName id="26F85AB6DC67FFE3F5DA35A9FDBAFA58" box="[483,554,1479,1499]" pageId="10" pageNumber="787">Brachat</collectorName>
</materialsCitation>
/ 
<typeStatus id="54B681C2DC67FFE3F67035A9FD37FA58" box="[585,679,1479,1499]" pageId="10" pageNumber="787" type="holotype">Holotypus</typeStatus>
<taxonomicName id="4C0D44E3DC67FFE3F6ED35A9FC1AFA58" authorityName="Assing" authorityYear="2007" box="[724,906,1479,1499]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="787" phylum="Arthropoda" rank="species" species="spinosula" status="sp. nov.">
<emphasis id="B979E372DC67FFE3F6ED35A9FC1AFA58" box="[724,906,1479,1499]" italics="true" pageId="10" pageNumber="787">Geostiba spinosula</emphasis>
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<taxonomicNameLabel id="A24A5E09DC67FFE3F7AF35A9FC52FA58" box="[918,962,1479,1499]" pageId="10" pageNumber="787" rank="species">sp.n.</taxonomicNameLabel>
det. 
<collectionCode id="ED1CA7A5DC67FFE3F7C535A9FB9FFA58" box="[1020,1039,1479,1499]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="10" pageNumber="787" type="Museum">V</collectionCode>
. Assing 2007 (cAss). 
<typeStatus id="54B681C2DC67FFE3F56235B0FE27FA71" box="[347,439,1502,1522]" pageId="10" pageNumber="787" type="paratype">Paratypes</typeStatus>
: 2: same data as 
<typeStatus id="54B681C2DC67FFE3F64C35B0FD54FA71" box="[629,708,1502,1522]" pageId="10" pageNumber="787" type="holotype">holotype</typeStatus>
(cAss).
</paragraph>
<paragraph id="8BB23F60DC67FFE3F562366FFBD9F995" blockId="10.[347,1097,1536,1558]" box="[347,1097,1536,1558]" pageId="10" pageNumber="787">
D e s c r i p t i o n: 2.0- 
<quantity id="4CF59285DC67FFE3F65F366EFD27F995" box="[614,695,1536,1558]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="10" pageNumber="787" unit="mm" value="2.7">2.7 mm</quantity>
. Coloration: body uniformly yellowish.
</paragraph>
<paragraph id="8BB23F60DC67FFE3F5623648FD2CF932" blockId="10.[347,1255,1574,1713]" pageId="10" pageNumber="787">
Head approximately as wide as long; puncturation extremely fine, barely noticeable; surface with shallow or very shallow microreticulation and with some shine. Eyes strongly reduced (
<figureCitation id="133623E5DC67FFE3F620360EFDF8F9F5" box="[537,616,1632,1654]" captionStart="Figs 36-42" captionStartId="11.[347,387,1604,1624]" captionTargetBox="[347,1247,771,1587]" captionTargetId="figure-141@11.[347,1260,1116,1593]" captionTargetPageId="11" captionText="Figs 36-42: Geostiba spinosula nov.sp.: (36) male pronotum, elytra, and abdominal segment III (holotype); (37) head in lateral view; (38-39) median lobe of aedeagus in lateral and in ventral view; (40) apical lobe of paramere; (41) female sternite VIII; (42) spermatheca. Scale bars: 36: 0.5 mm; 37: 0.2 mm; 38-42: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10114386" httpUri="https://zenodo.org/record/10114386/files/figure.png" pageId="10" pageNumber="787">Fig. 37</figureCitation>
), smaller than antennomere III in cross-section, without pigmentation and without ommatidia. Antennae distinctly incrassate apically; antennomeres V-X 2.0-2.5 times as wide as long.
</paragraph>
<paragraph id="8BB23F60DC67FFE3F56236AFFDCDF892" blockId="10.[347,1255,1728,1809]" pageId="10" pageNumber="787">
Pronotum small, 1.05-1.10 times as wide as head and 1.05-1.10 times as wide as long (
<figureCitation id="133623E5DC67FFE3F55A36B3FE20F970" box="[355,432,1757,1779]" captionStart="Figs 36-42" captionStartId="11.[347,387,1604,1624]" captionTargetBox="[347,1247,771,1587]" captionTargetId="figure-141@11.[347,1260,1116,1593]" captionTargetPageId="11" captionText="Figs 36-42: Geostiba spinosula nov.sp.: (36) male pronotum, elytra, and abdominal segment III (holotype); (37) head in lateral view; (38-39) median lobe of aedeagus in lateral and in ventral view; (40) apical lobe of paramere; (41) female sternite VIII; (42) spermatheca. Scale bars: 36: 0.5 mm; 37: 0.2 mm; 38-42: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10114386" httpUri="https://zenodo.org/record/10114386/files/figure.png" pageId="10" pageNumber="787">Fig. 36</figureCitation>
); puncturation extremely fine, barely noticeable; microreticulation slightly more distinct than that of head.
</paragraph>
<paragraph id="8BB23F60DC66FFE2F736310BFCA3FEF9" blockId="11.[783,819,357,378]" box="[783,819,357,378]" pageId="11" pageNumber="788">788</paragraph>
<paragraph id="8BB23F60DC66FFE2F56231DEFB54FD83" blockId="11.[347,1255,432,512]" pageId="11" pageNumber="788">
Elytra approximately as wide as pronotum or slightly wider, and about 0.55 times as long as pronotum (
<figureCitation id="133623E5DC66FFE2F5DE31A3FDA4FE60" box="[487,564,461,483]" captionStart="Figs 36-42" captionStartId="11.[347,387,1604,1624]" captionTargetBox="[347,1247,771,1587]" captionTargetId="figure-141@11.[347,1260,1116,1593]" captionTargetPageId="11" captionText="Figs 36-42: Geostiba spinosula nov.sp.: (36) male pronotum, elytra, and abdominal segment III (holotype); (37) head in lateral view; (38-39) median lobe of aedeagus in lateral and in ventral view; (40) apical lobe of paramere; (41) female sternite VIII; (42) spermatheca. Scale bars: 36: 0.5 mm; 37: 0.2 mm; 38-42: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10114386" httpUri="https://zenodo.org/record/10114386/files/figure.png" pageId="11" pageNumber="788">Fig. 36</figureCitation>
); puncturation very fine, but slightly more distinct than that of head and pronotum; microreticulation shallow; sexual dimorphism moderately pronounced.
</paragraph>
<paragraph id="8BB23F60DC66FFE2F5623261FD63FDE3" blockId="11.[347,1255,527,608]" pageId="11" pageNumber="788">Abdomen 1.05-1.10 times as wide as elytra; puncturation very fine and sparse; microreticulation very shallow; posterior margin of tergite VII without palisade fringe; posterior margin of tergite VIII distinctly convex.</paragraph>
<paragraph id="8BB23F60DC66FFE2F549321EFBCEFD79" blockId="11.[347,1255,623,762]" pageId="11" pageNumber="788">
: elytra with lateral margins sharply edged; abdominal tergite III with pronounced smooth median tubercle (
<figureCitation id="133623E5DC66FFE2F65E32E3FD27FD20" box="[615,695,653,675]" captionStart="Figs 36-42" captionStartId="11.[347,387,1604,1624]" captionTargetBox="[347,1247,771,1587]" captionTargetId="figure-141@11.[347,1260,1116,1593]" captionTargetPageId="11" captionText="Figs 36-42: Geostiba spinosula nov.sp.: (36) male pronotum, elytra, and abdominal segment III (holotype); (37) head in lateral view; (38-39) median lobe of aedeagus in lateral and in ventral view; (40) apical lobe of paramere; (41) female sternite VIII; (42) spermatheca. Scale bars: 36: 0.5 mm; 37: 0.2 mm; 38-42: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10114386" httpUri="https://zenodo.org/record/10114386/files/figure.png" pageId="11" pageNumber="788">Fig. 36</figureCitation>
); tergite VII with small spine-like median process at posterior margin, this process flattened dorso-ventrally and of somewhat triangular shape in antero-dorsal view; posterior margin of sternite VIII strongly convex; median lobe of aedeagus shaped as in 
<figureCitation id="133623E5DC66FFE2F678328AFD20FD79" box="[577,688,740,762]" captionStart="Figs 36-42" captionStartId="11.[347,387,1604,1624]" captionTargetBox="[347,1247,771,1587]" captionTargetId="figure-141@11.[347,1260,1116,1593]" captionTargetPageId="11" captionText="Figs 36-42: Geostiba spinosula nov.sp.: (36) male pronotum, elytra, and abdominal segment III (holotype); (37) head in lateral view; (38-39) median lobe of aedeagus in lateral and in ventral view; (40) apical lobe of paramere; (41) female sternite VIII; (42) spermatheca. Scale bars: 36: 0.5 mm; 37: 0.2 mm; 38-42: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10114386" httpUri="https://zenodo.org/record/10114386/files/figure.png" pageId="11" pageNumber="788">Figs 38-39</figureCitation>
; apical lobe of paramere slender (
<figureCitation id="133623E5DC66FFE2F03F328AFBC3FD79" box="[1030,1107,740,762]" captionStart="Figs 36-42" captionStartId="11.[347,387,1604,1624]" captionTargetBox="[347,1247,771,1587]" captionTargetId="figure-141@11.[347,1260,1116,1593]" captionTargetPageId="11" captionText="Figs 36-42: Geostiba spinosula nov.sp.: (36) male pronotum, elytra, and abdominal segment III (holotype); (37) head in lateral view; (38-39) median lobe of aedeagus in lateral and in ventral view; (40) apical lobe of paramere; (41) female sternite VIII; (42) spermatheca. Scale bars: 36: 0.5 mm; 37: 0.2 mm; 38-42: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10114386" httpUri="https://zenodo.org/record/10114386/files/figure.png" pageId="11" pageNumber="788">Fig. 40</figureCitation>
).
</paragraph>
<caption id="DF726FE8DC66FFE2F562362AFD3EF91F" ID-DOI="http://doi.org/10.5281/zenodo.10114386" ID-Zenodo-Dep="10114386" httpUri="https://zenodo.org/record/10114386/files/figure.png" pageId="11" pageNumber="788" startId="11.[347,387,1604,1624]" targetBox="[347,1247,771,1587]" targetPageId="11">
<paragraph id="8BB23F60DC66FFE2F562362AFD3EF91F" blockId="11.[347,1255,1604,1692]" pageId="11" pageNumber="788">
<emphasis id="B979E372DC66FFE2F562362AFE51F9DB" bold="true" box="[347,449,1604,1624]" pageId="11" pageNumber="788">Figs 36-42</emphasis>
: 
<taxonomicName id="4C0D44E3DC66FFE2F5E8362AFD12F9DB" authorityName="Assing" authorityYear="2007" box="[465,642,1604,1624]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="788" phylum="Arthropoda" rank="species" species="spinosula" status="sp. nov.">
<emphasis id="B979E372DC66FFE2F5E8362AFD12F9DB" box="[465,642,1604,1624]" italics="true" pageId="11" pageNumber="788">Geostiba spinosula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24A5E09DC66FFE2F6B2362AFD41F9DB" box="[651,721,1604,1624]" pageId="11" pageNumber="788" rank="species">nov.sp.</taxonomicNameLabel>
: (
<emphasis id="B979E372DC66FFE2F6DD362AFD6BF9DB" bold="true" box="[740,763,1604,1624]" pageId="11" pageNumber="788">36</emphasis>
) male pronotum, elytra, and abdominal segment III (holotype); (
<emphasis id="B979E372DC66FFE2F5E83635FE78F9EC" bold="true" box="[465,488,1627,1647]" pageId="11" pageNumber="788">37</emphasis>
) head in lateral view; (
<emphasis id="B979E372DC66FFE2F6F23635FC90F9EC" bold="true" box="[715,768,1627,1647]" pageId="11" pageNumber="788">38-39</emphasis>
) median lobe of aedeagus in lateral and in ventral view; (
<emphasis id="B979E372DC66FFE2F5A7361CFE25F905" bold="true" box="[414,437,1650,1670]" pageId="11" pageNumber="788">40</emphasis>
) apical lobe of paramere; (
<emphasis id="B979E372DC66FFE2F682361CFD42F905" bold="true" box="[699,722,1650,1670]" pageId="11" pageNumber="788">41</emphasis>
) female sternite VIII; (
<emphasis id="B979E372DC66FFE2F78A361CFC5AF905" bold="true" box="[947,970,1650,1670]" pageId="11" pageNumber="788">42</emphasis>
) spermatheca. Scale bars: 36: 0.5 mm; 37: 0.2 mm; 38-42: 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BB23F60DC66FFE2F54936C2FCB2F95D" blockId="11.[347,1255,1707,1758]" pageId="11" pageNumber="788">
: elytra unmodified; posterior margin of sternite VIII broadly convex, in the middle truncate (
<figureCitation id="133623E5DC66FFE2F58236A6FD96F95D" box="[443,518,1736,1758]" captionStart="Figs 36-42" captionStartId="11.[347,387,1604,1624]" captionTargetBox="[347,1247,771,1587]" captionTargetId="figure-141@11.[347,1260,1116,1593]" captionTargetPageId="11" captionText="Figs 36-42: Geostiba spinosula nov.sp.: (36) male pronotum, elytra, and abdominal segment III (holotype); (37) head in lateral view; (38-39) median lobe of aedeagus in lateral and in ventral view; (40) apical lobe of paramere; (41) female sternite VIII; (42) spermatheca. Scale bars: 36: 0.5 mm; 37: 0.2 mm; 38-42: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10114386" httpUri="https://zenodo.org/record/10114386/files/figure.png" pageId="11" pageNumber="788">Fig. 41</figureCitation>
); spermatheca as in 
<figureCitation id="133623E5DC66FFE2F6EA36A6FCB2F95D" box="[723,802,1736,1758]" captionStart="Figs 36-42" captionStartId="11.[347,387,1604,1624]" captionTargetBox="[347,1247,771,1587]" captionTargetId="figure-141@11.[347,1260,1116,1593]" captionTargetPageId="11" captionText="Figs 36-42: Geostiba spinosula nov.sp.: (36) male pronotum, elytra, and abdominal segment III (holotype); (37) head in lateral view; (38-39) median lobe of aedeagus in lateral and in ventral view; (40) apical lobe of paramere; (41) female sternite VIII; (42) spermatheca. Scale bars: 36: 0.5 mm; 37: 0.2 mm; 38-42: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10114386" httpUri="https://zenodo.org/record/10114386/files/figure.png" pageId="11" pageNumber="788">Fig. 42.</figureCitation>
</paragraph>
<paragraph id="8BB23F60DC66FFE2F5623680FB77F880" blockId="11.[347,1255,1773,1795]" box="[347,1255,1773,1795]" pageId="11" pageNumber="788">E t y m o l o g y: The name, diminutive of spinose (Lat., adj.: thorny), refers to the</paragraph>
<paragraph id="8BB23F60DC61FFE5F736310BFCA3FEF9" blockId="12.[783,819,357,378]" box="[783,819,357,378]" pageId="12" pageNumber="789">789</paragraph>
<paragraph id="8BB23F60DC61FFE5F56231DEFE5DFE60" blockId="12.[347,1254,432,483]" pageId="12" pageNumber="789">minute body size and the presence of a small spine-like process at the posterior margin of tergite VII.</paragraph>
<paragraph id="8BB23F60DC61FFE5F562319DFD01FC5E" blockId="12.[347,1255,499,989]" pageId="12" pageNumber="789">
C o m p a r a t i v e n o t e s: Based on external (pale coloration, strongly reduced eyes), as well as on various derived male primary and secondary sexual characters (elytra with sharply edged lateral margins, presence of a tubercle on tergite III, presence of a spine-like process at the posterior margin of tergite VII, morphology of the median lobe of the aedeagus, shape and chaetotaxy of the apical lobe of the paramere), 
<taxonomicName id="4C0D44E3DC61FFE5F05C3206FB77FDFD" authorityName="Assing" authorityYear="2007" box="[1125,1255,616,638]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="789" phylum="Arthropoda" rank="species" species="spinosula">
<emphasis id="B979E372DC61FFE5F05C3206FB77FDFD" box="[1125,1255,616,638]" italics="true" pageId="12" pageNumber="789">G. spinosula</emphasis>
</taxonomicName>
belongs to the 
<taxonomicName id="4C0D44E3DC61FFE5F5C032E8FDFAFD18" box="[505,618,645,667]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="789" phylum="Arthropoda" rank="species" species="confusa">
<emphasis id="B979E372DC61FFE5F5C032E8FDFAFD18" box="[505,618,645,667]" italics="true" pageId="12" pageNumber="789">G. confusa</emphasis>
</taxonomicName>
group (see 
<bibRefCitation id="EF9C4291DC61FFE5F6D332EBFC12FD18" author="ASSING V" box="[746,898,645,667]" pageId="12" pageNumber="789" pagination="1047 - 1070" refId="ref6682" refString="ASSING V. (2005 b): New species and new records of Eastern Mediterranean Geostiba THOMSON (Coleoptera: Staphylinidae, Aleocharinae). - Linzer biologische Beitrage 37 (2): 1047 - 1070." type="journal article" year="2005">ASSING 2005b</bibRefCitation>
), which previously included three species from central southern 
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: 
<taxonomicName id="4C0D44E3DC61FFE5F6C832CDFC23FD3B" authority="ASSING" authorityName="ASSING" box="[753,947,674,696]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="789" phylum="Arthropoda" rank="species" species="confusa">
<emphasis id="B979E372DC61FFE5F6C832CDFCF0FD3B" box="[753,864,674,696]" italics="true" pageId="12" pageNumber="789">G. confusa</emphasis>
ASSING
</taxonomicName>
, 
<taxonomicName id="4C0D44E3DC61FFE5F7F932CDFB05FD3B" authority="ASSING" authorityName="ASSING" box="[960,1173,674,696]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="789" phylum="Arthropoda" rank="species" species="bigibbera">
<emphasis id="B979E372DC61FFE5F7F932CDFBD1FD3B" box="[960,1089,674,696]" italics="true" pageId="12" pageNumber="789">G. bigibbera</emphasis>
ASSING
</taxonomicName>
, and 
<taxonomicName id="4C0D44E3DC61FFE5F0F732CDFDCCFD56" authority="ASSING. Among" authorityName="ASSING. Among" authorityYear="2005" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="789" phylum="Arthropoda" rank="species" species="gibbera">
<emphasis id="B979E372DC61FFE5F0F732CDFE3AFD56" italics="true" pageId="12" pageNumber="789">G. gibbera</emphasis>
ASSING. Among
</taxonomicName>
these species, 
<taxonomicName id="4C0D44E3DC61FFE5F6C532AEFCEFFD56" authorityName="Assing" authorityYear="2007" box="[764,895,703,725]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="789" phylum="Arthropoda" rank="species" species="spinosula">
<emphasis id="B979E372DC61FFE5F6C532AEFCEFFD56" box="[764,895,703,725]" italics="true" pageId="12" pageNumber="789">G. spinosula</emphasis>
</taxonomicName>
is most similar and related to the geographically close 
<taxonomicName id="4C0D44E3DC61FFE5F60E32B3FD37FD70" authorityName="ASSING. Among" authorityYear="2005" box="[567,679,733,755]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="789" phylum="Arthropoda" rank="species" species="gibbera">
<emphasis id="B979E372DC61FFE5F60E32B3FD37FD70" box="[567,679,733,755]" italics="true" pageId="12" pageNumber="789">G. gibbera</emphasis>
</taxonomicName>
, which is known only from one locality approximately 
<quantity id="4CF59285DC61FFE5F5623294FE0EFC93" box="[347,414,762,784]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="12" pageNumber="789" unit="km" value="30.0">30 km</quantity>
SSW of Kahramanmaraú. It is distinguished from this species by smaller body size, distinctly paler coloration (in 
<taxonomicName id="4C0D44E3DC61FFE5F6C93376FCF8FCAE" authorityName="ASSING. Among" authorityYear="2005" box="[752,872,791,813]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="789" phylum="Arthropoda" rank="species" species="gibbera">
<emphasis id="B979E372DC61FFE5F6C93376FCF8FCAE" box="[752,872,791,813]" italics="true" pageId="12" pageNumber="789">G. gibbera</emphasis>
</taxonomicName>
reddish yellow), less pronounced microsculpture especially of the pronotum, shorter antennae, a more pronounced tubercle on the male tergite III, a differently shaped process of the male tergite VII, the different morphology of the aedeagus (ventral process shorter, not bent in lateral view, and apically less acute in ventral view), and a spermatheca with a more slender capsule and a shorter duct. For illustrations of the habitus and the sexual characters of 
<taxonomicName id="4C0D44E3DC61FFE5F07733C4FB2EFC43" authorityName="ASSING. Among" authorityYear="2005" box="[1102,1214,938,960]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="789" phylum="Arthropoda" rank="species" species="gibbera">
<emphasis id="B979E372DC61FFE5F07733C4FB2EFC43" box="[1102,1214,938,960]" italics="true" pageId="12" pageNumber="789">G. gibbera</emphasis>
</taxonomicName>
see the figures in 
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.
</paragraph>
<paragraph id="8BB23F60DC61FFE5F5623383FE20FBD9" blockId="12.[347,1255,1004,1114]" pageId="12" pageNumber="789">
D i s t r i b u t i o n a n d b i o n o m i c s: The 
<typeStatus id="54B681C2DC61FFE5F7B33383FC3EFB81" box="[906,942,1005,1026]" pageId="12" pageNumber="789">type</typeStatus>
locality is situated some 45 southwest of Kahramanmaraú, central southern 
<collectingCountry id="F31A7FF0DC61FFE5F7753464FC33FBA3" box="[844,931,1034,1056]" name="Turkey" pageId="12" pageNumber="789">Anatolia</collectingCountry>
(
<figureCitation id="133623E5DC61FFE5F78A3464FC69FBA3" box="[947,1017,1034,1056]" captionStart="Map 1" captionStartId="5.[347,392,897,917]" captionTargetBox="[347,1254,432,897]" captionTargetId="figure-7@5.[347,1253,433,889]" captionTargetPageId="5" captionText="Map 1: Distributions of Geostiba janbellini nov.sp. (filled circle), G. atrioculata nov.sp. (open circle), and G. spinosula nov.sp. (square) in southern Anatolia." figureDoi="http://doi.org/10.5281/zenodo.10114376" httpUri="https://zenodo.org/record/10114376/files/figure.png" pageId="12" pageNumber="789">Map 1</figureCitation>
). The specimens were collected under stones on a grassy slope at an altitude of 
<quantity id="4CF59285DC61FFE5F7E63449FBA1FBBE" box="[991,1073,1063,1085]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="12" pageNumber="789" unit="m" value="1200.0">1200 m</quantity>
(BRACHAT pers. comm.).
</paragraph>
</subSubSection>
</treatment>
</document>