582 lines
70 KiB
XML
582 lines
70 KiB
XML
<document id="14679AB450FD26E75A33E5FEB449A6F5" ID-DOI="10.5281/zenodo.184689" ID-GBIF-Dataset="c0a0fd0e-62ae-4a9e-bf0a-be318473c330" ID-ISSN="1175-5326" ID-Zenodo-Dep="184689" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459997393761" checkinUser="plazi" docAuthor="Arillo, Antonio, Peñalver, Enrique & Delclòs, Xavier" docDate="2008" docId="642F780D0F501335FF0D7EF7FA55FC6A" docLanguage="en" docName="zt01920p040.pdf" docOrigin="Zootaxa 1920" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Microphorites utrillensis Penalver" docType="treatment" docVersion="7" lastPageNumber="35" masterDocId="981600750F521333FF9A7E60FFC6FFFD" masterDocTitle="Microphorites (Diptera: Dolichopodidae) from the Lower Cretaceous amber of San Just (Spain), and the co-occurrence of two ceratopogonid species in Spanish amber deposits" masterLastPageNumber="40" masterPageNumber="29" pageNumber="31" updateTime="1698232985276" updateUser="plazi">
|
||
<mods:mods id="60E9D3CD6123CEC4F9BD5D0FC5E794B7" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="14D8FD8D213A6875528F060B82725ED1">
|
||
<mods:title id="85ACEFB0678B24BC08457068C9B60F66">Microphorites (Diptera: Dolichopodidae) from the Lower Cretaceous amber of San Just (Spain), and the co-occurrence of two ceratopogonid species in Spanish amber deposits</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="59420383B899C8DD5AC328F72736985C" type="personal">
|
||
<mods:role id="4321B8190F952698BFC6FCCFD2C74EE4">
|
||
<mods:roleTerm id="5C82A8BFAD3920163FF5E84C22659A2F">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="0FEEA669827711DF03FB7E16AF48EFA1">Arillo, Antonio</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="129C4D2F4FA9DF48D3535E50E7867CFF" type="personal">
|
||
<mods:role id="4ED1BF373C2047EF03CF6A75E4C3AF56">
|
||
<mods:roleTerm id="F3F646907D539F327E09BE8E27CC1BA6">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="1A277FDC9FFB78BA083E1CAE480B2C88">Peñalver, Enrique</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="886457CA28DF5F7EFE4C63EBBD3D5E57" type="personal">
|
||
<mods:role id="35BE95CC8E2D36E8FAC9E044A8D9C1A4">
|
||
<mods:roleTerm id="0F08E681FD8A77F2BA77BD2FEC09F1EB">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="640358B3FE46165FB07DA8AAE627ACCE">Delclòs, Xavier</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="200607F9C2687C85573BC120EF1EBC62">text</mods:typeOfResource>
|
||
<mods:relatedItem id="9B83404FE460B626468E3C79002FF542" type="host">
|
||
<mods:titleInfo id="6510BE3162AC955F7DE8D31904E39F77">
|
||
<mods:title id="D9B9CF15E5BA56B61D8ADDB2FD32BF68">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="C5C2FE4BBFCF0BB4FFC5412971355A66">
|
||
<mods:date id="9035AA9AA1C1DDAB4018B7F7B5DF0084">2008</mods:date>
|
||
<mods:detail id="A4CE5932B2BE0C87C37DC4B4298E9BCD" type="volume">
|
||
<mods:number id="E91DDAB8948F67D642549DD1DE7DC30D">1920</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="0B075F89E0EDE25529A5AE98206294B5" unit="page">
|
||
<mods:start id="5FF36880D7E4CABDE6BBDDFBE1C06CA3">29</mods:start>
|
||
<mods:end id="DD3919D84A67269CD54CA7D864B59DBD">40</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="6BC0D5EE4783A66E15A639A76258B6B4">journal article</mods:classification>
|
||
<mods:identifier id="79D12E06B5D0AC4A05A72FC78AE0BF6A" type="DOI">10.5281/zenodo.184689</mods:identifier>
|
||
<mods:identifier id="3785275BCE18E8CAF46344D378304AB9" type="GBIF-Dataset">c0a0fd0e-62ae-4a9e-bf0a-be318473c330</mods:identifier>
|
||
<mods:identifier id="5B638B0C5F572414EDD140BD13882E9F" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="A60977BDDF7FE32328DBB998FE62331D" type="Zenodo-Dep">184689</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="642F780D0F501335FF0D7EF7FA55FC6A" ID-DOI="http://doi.org/10.5281/zenodo.6231397" ID-GBIF-Taxon="119380488" ID-Zenodo-Dep="6231397" LSID="urn:lsid:plazi:treatment:642F780D0F501335FF0D7EF7FA55FC6A" httpUri="http://treatment.plazi.org/id/642F780D0F501335FF0D7EF7FA55FC6A" lastPageId="6" lastPageNumber="35" pageId="2" pageNumber="31">
|
||
<subSubSection id="A49C9A900F501331FF0D7EF7FE1FFF29" pageId="2" pageNumber="31" type="nomenclature">
|
||
<paragraph id="EC39C91B0F501331FF0D7EF7FD64FF4C" blockId="2.[151,674,151,212]" box="[151,674,151,177]" pageId="2" pageNumber="31">
|
||
<heading id="B7717E770F501331FF0D7EF7FD64FF4C" bold="true" box="[151,674,151,177]" fontSize="11" level="1" pageId="2" pageNumber="31" reason="1">
|
||
<emphasis id="DEF215090F501331FF0D7EF7FD64FF4C" bold="true" box="[151,674,151,177]" pageId="2" pageNumber="31">
|
||
<taxonomicName id="2B86B2980F501331FF0D7EF7FDF1FF4C" authority="Penalver" authorityName="Penalver" box="[151,567,151,177]" class="Insecta" family="Empididae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="utrillensis">
|
||
<emphasis id="DEF215090F501331FF0D7EF7FE04FF4C" bold="true" box="[151,450,151,177]" italics="true" pageId="2" pageNumber="31">Microphorites utrillensis</emphasis>
|
||
Peñalver
|
||
</taxonomicName>
|
||
, nov. sp.
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="EC39C91B0F501331FF0D7EDAFE1FFF29" blockId="2.[151,674,151,212]" box="[151,473,186,212]" pageId="2" pageNumber="31">
|
||
<figureCitation id="74BDD59E0F501331FF0D7EDAFEC2FF29" box="[151,260,186,212]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="3.[151,254,1302,1326]" captionStartId-1="4.[151,254,1492,1516]" captionStartId-2="5.[151,254,1642,1666]" captionTargetBox-0="[171,1419,203,1267]" captionTargetBox-1="[422,1163,202,1460]" captionTargetBox-2="[166,1421,680,1607]" captionTargetId-0="figure@3.[153,1438,197,1278]" captionTargetId-1="figure@4.[376,1210,197,1467]" captionTargetId-2="figure@5.[151,1436,655,1618]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 1. Camera lucida drawings of Microphorites utrillensis Peñalver nov. sp. (Dolichopodidae), female holotype (CPT- 963), in ventral and dorsal habitus, from San Just Amber (Teruel Province). Bar = 1 mm. See also Plate 1 a." captionText-1="FIGURE 2. Camera lucida drawing of the head of Microphorites utrillensis Peñalver nov. sp. (Dolichopodidae), female holotype (CPT- 963), in ventral view. Bar = 0.1 mm. See also Plate 1 b." captionText-2="FIGURE 3. Camera lucida drawings of the a) wing (crossvein h obscure), b) hind leg and c) distal part of the abdomen in ventral view. Microphorites utrillensis Peñalver nov. sp., female holotype (CPT- 963). cerc = cercus. All to the same scale (bar = 0.1 mm). See also Plate 1 c." httpUri-0="https://zenodo.org/record/184690/files/figure.png" httpUri-1="https://zenodo.org/record/184691/files/figure.png" httpUri-2="https://zenodo.org/record/184692/files/figure.png" pageId="2" pageNumber="31">Figs. 1–3</figureCitation>
|
||
& Pl. 1, figs. a–c.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A49C9A900F501331FF0D7F6AFCEFFE7C" pageId="2" pageNumber="31" type="description">
|
||
<paragraph id="EC39C91B0F501331FF0D7F6AFD45FEDC" blockId="2.[151,809,266,385]" box="[151,643,266,289]" pageId="2" pageNumber="31">
|
||
2006
|
||
<taxonomicName id="2B86B2980F501331FF497F6AFEADFEDC" box="[211,363,266,289]" class="Insecta" family="Empididae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DEF215090F501331FF497F6AFEADFEDC" box="[211,363,266,289]" italics="true" pageId="2" pageNumber="31">Microphorites</emphasis>
|
||
</taxonomicName>
|
||
sp.; Peñalver
|
||
<emphasis id="DEF215090F501331FD9B7F6AFDF1FEDC" box="[513,567,266,289]" italics="true" pageId="2" pageNumber="31">et al.</emphasis>
|
||
; fig. 1.
|
||
</paragraph>
|
||
<paragraph id="EC39C91B0F501331FF0D7F4AFDEDFEBC" blockId="2.[151,809,266,385]" box="[151,555,298,321]" pageId="2" pageNumber="31">
|
||
2007
|
||
<taxonomicName id="2B86B2980F501331FF497F4AFEADFEBC" box="[211,363,298,321]" class="Insecta" family="Empididae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DEF215090F501331FF497F4AFEADFEBC" box="[211,363,298,321]" italics="true" pageId="2" pageNumber="31">Microphorites</emphasis>
|
||
</taxonomicName>
|
||
sp.; Delclòs
|
||
<emphasis id="DEF215090F501331FE6F7F4AFDE3FEBC" box="[501,549,298,321]" italics="true" pageId="2" pageNumber="31">et al</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="EC39C91B0F501331FF0D7F2AFCEFFE7C" blockId="2.[151,809,266,385]" pageId="2" pageNumber="31">
|
||
2007
|
||
<taxonomicName id="2B86B2980F501331FF497F2AFEADFE9C" box="[211,363,330,353]" class="Insecta" family="Empididae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DEF215090F501331FF497F2AFEADFE9C" box="[211,363,330,353]" italics="true" pageId="2" pageNumber="31">Microphorites</emphasis>
|
||
</taxonomicName>
|
||
sp.; Peñalver
|
||
<emphasis id="DEF215090F501331FD9B7F2AFDF6FE9C" box="[513,560,330,353]" italics="true" pageId="2" pageNumber="31">el al</emphasis>
|
||
.; figs. 6D & 7. 2007 Dipteran
|
||
<taxonomicName id="2B86B2980F501331FEAD7F0AFE08FE7C" box="[311,462,362,385]" class="Insecta" family="Empididae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DEF215090F501331FEAD7F0AFE08FE7C" box="[311,462,362,385]" italics="true" pageId="2" pageNumber="31">Microphorites</emphasis>
|
||
</taxonomicName>
|
||
sp.; Delclòs & Soriano: fig. 4H.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A49C9A900F501331FF0D7FD2FA93FD19" pageId="2" pageNumber="31" type="materials_examined">
|
||
<paragraph id="EC39C91B0F501331FF0D7FD2FB02FDB9" blockId="2.[151,1437,434,2038]" pageId="2" pageNumber="31">
|
||
<typeStatus id="333D77B90F501331FF0D7FD2FECFFE31" box="[151,265,434,460]" pageId="2" pageNumber="31" type="holotype">
|
||
<emphasis id="DEF215090F501331FF0D7FD2FECFFE31" bold="true" box="[151,265,434,460]" pageId="2" pageNumber="31">Holotype</emphasis>
|
||
</typeStatus>
|
||
: Adult female. Specimen CPT-963, housed in the Fundación Conjunto Paleontológico de Teruel- Dinópolis (Teruel Province,
|
||
<collectingCountry id="9491898B0F501331FE7E7FBAFDEDFE09" box="[484,555,474,500]" name="Spain" pageId="2" pageNumber="31">Spain</collectingCountry>
|
||
). Complete specimen present in a transparent, clear piece of amber of
|
||
<date id="9838EFDB0F501331FAFB7FBAFA40FE09" box="[1377,1414,474,500]" pageId="2" pageNumber="31">9 x</date>
|
||
<quantity id="2B7E64FE0F501331FA177FBAFF03FDE1" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="31" unit="mm" value="5.0">5 mm</quantity>
|
||
into an epoxy resin preparation of 15 x
|
||
<date id="9838EFDB0F501331FD297C62FD1DFDE1" box="[691,731,514,540]" pageId="2" pageNumber="31">9 x</date>
|
||
<quantity id="2B7E64FE0F501331FD7F7C62FCF6FDE1" box="[741,816,514,540]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="31" unit="mm" value="2.0">2 mm</quantity>
|
||
. The specimen, along with other insects, is trapped within a spider orb web, which is the oldest web in the fossil record (
|
||
<bibRefCitation id="8817B4EA0F501331FC247C4AFB71FDB9" author="Penalver" box="[958,1207,554,580]" pageId="2" pageNumber="40" refString="Penalver, E., Grimaldi, D. A. & Delclos, X. (2006) Early Cretaceous Spider Web with Its Prey. Science, 312, 1761." type="journal article" year="2006">
|
||
Peñalver
|
||
<emphasis id="DEF215090F501331FBB47C4AFBA7FDBE" box="[1070,1121,554,579]" italics="true" pageId="2" pageNumber="31">et al</emphasis>
|
||
., 2006
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="EC39C91B0F501331FF5C7C32FCA6FD41" blockId="2.[151,1437,434,2038]" pageId="2" pageNumber="31">
|
||
<emphasis id="DEF215090F501331FF5C7C32FEB6FD91" bold="true" box="[198,368,594,620]" pageId="2" pageNumber="31">
|
||
<typeStatus id="333D77B90F501331FF5C7C32FEC2FD91" box="[198,260,594,620]" pageId="2" pageNumber="31">Type</typeStatus>
|
||
locality:
|
||
</emphasis>
|
||
Specimen collected in San Just outcrop, in the municipality of Utrillas, near the village of Escucha (Teruel Province,
|
||
<collectingCountry id="9491898B0F501331FE7E7C1AFDEAFD69" box="[484,556,634,660]" name="Spain" pageId="2" pageNumber="31">Spain</collectingCountry>
|
||
). Escucha Fm. (La Orden Member), Lower Cretaceous (Lower-Middle Albian). See Peñalver
|
||
<emphasis id="DEF215090F501331FE047CC2FE14FD46" box="[414,466,674,699]" italics="true" pageId="2" pageNumber="31">et al</emphasis>
|
||
. (2007) and Delclòs
|
||
<emphasis id="DEF215090F501331FD527CC2FD3DFD46" box="[712,763,674,699]" italics="true" pageId="2" pageNumber="31">et al</emphasis>
|
||
. (2007).
|
||
</paragraph>
|
||
<paragraph id="EC39C91B0F501331FF5C7CAAFA93FD19" blockId="2.[151,1437,434,2038]" box="[198,1365,714,740]" pageId="2" pageNumber="31">
|
||
<emphasis id="DEF215090F501331FF5C7CAAFE71FD19" bold="true" box="[198,439,714,740]" pageId="2" pageNumber="31">Derivation of name</emphasis>
|
||
: After Utrillas, the name of the municipality where the
|
||
<typeStatus id="333D77B90F501331FBDE7CAAFBB1FD19" box="[1092,1143,714,740]" pageId="2" pageNumber="31">type</typeStatus>
|
||
locality is located.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A49C9A900F501331FF5C7C92FC77FC4F" pageId="2" pageNumber="31" type="diagnosis">
|
||
<paragraph id="EC39C91B0F501331FF5C7C92FC77FC4F" blockId="2.[151,1437,434,2038]" pageId="2" pageNumber="31">
|
||
<emphasis id="DEF215090F501331FF5C7C92FEF9FCF1" bold="true" box="[198,319,754,780]" pageId="2" pageNumber="31">Diagnosis</emphasis>
|
||
: Distinghished from other female and male
|
||
<taxonomicName id="2B86B2980F501331FCFA7C92FBC1FCF6" box="[864,1031,754,779]" class="Insecta" family="Empididae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DEF215090F501331FCFA7C92FBC1FCF6" box="[864,1031,754,779]" italics="true" pageId="2" pageNumber="31">Microphorites</emphasis>
|
||
</taxonomicName>
|
||
by the following combination of characters: very short arista (shorter than basal flagellomere), scutellum with two pairs of strong apical setae, costal vein ends at apex of M1, R1 ends at a basal position with respect to m-cu crossvein, R2+3 ends in a medial position between the ends of R1 and R4+5, crossvein r-m at 2/5 of the wing length, dm-cu not arched and 3.5 times longer than m-cu, abdomen elongate (not short and compact).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A49C9A900F501330FF5C7DA0FD41F99B" lastPageId="3" lastPageNumber="32" pageId="2" pageNumber="31" type="description">
|
||
<paragraph id="EC39C91B0F501331FF5C7DA0FE84FA6F" blockId="2.[151,1437,434,2038]" pageId="2" pageNumber="31">
|
||
<emphasis id="DEF215090F501331FF5C7DA0FE91FC27" bold="true" box="[198,343,960,986]" pageId="2" pageNumber="31">Description</emphasis>
|
||
: Body length
|
||
<quantity id="2B7E64FE0F501331FE657DA0FDAEFC27" box="[511,616,960,986]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="2" pageNumber="31" unit="mm" value="1.7">1.70 mm</quantity>
|
||
(from head to the evaginated distal part of the abdomen, see
|
||
<figureCitation id="74BDD59E0F501331FAA37DA0FAB8FC27" box="[1337,1406,960,986]" captionStart="FIGURE 1" captionStartId="3.[151,254,1302,1326]" captionTargetBox="[171,1419,203,1267]" captionTargetId="figure@3.[153,1438,197,1278]" captionTargetPageId="3" captionText="FIGURE 1. Camera lucida drawings of Microphorites utrillensis Peñalver nov. sp. (Dolichopodidae), female holotype (CPT- 963), in ventral and dorsal habitus, from San Just Amber (Teruel Province). Bar = 1 mm. See also Plate 1 a." httpUri="https://zenodo.org/record/184690/files/figure.png" pageId="2" pageNumber="31">Fig. 1</figureCitation>
|
||
& Pl. 1, fig. a). Head
|
||
<quantity id="2B7E64FE0F501331FEEE7D88FE1AFBFF" box="[372,476,1000,1026]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="31" unit="mm" value="0.25">0.25 mm</quantity>
|
||
long,
|
||
<quantity id="2B7E64FE0F501331FDBC7D88FD48FBFF" box="[550,654,1000,1026]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="2" pageNumber="31" unit="mm" value="0.32">0.32 mm</quantity>
|
||
greatest width. Eye dichoptic (slightly separate in dorsal position), bare, not emarginated near antennae, diameter
|
||
<quantity id="2B7E64FE0F501331FD257A70FCEDFBD7" box="[703,811,1040,1066]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="31" unit="mm" value="0.2">0.20 mm</quantity>
|
||
, occupying entire lateral surface of head. Ommatidia uniform (
|
||
<figureCitation id="74BDD59E0F501331FE927A58FE96FBAF" box="[264,336,1080,1106]" captionStart="FIGURE 2" captionStartId="4.[151,254,1492,1516]" captionTargetBox="[422,1163,202,1460]" captionTargetId="figure@4.[376,1210,197,1467]" captionTargetPageId="4" captionText="FIGURE 2. Camera lucida drawing of the head of Microphorites utrillensis Peñalver nov. sp. (Dolichopodidae), female holotype (CPT- 963), in ventral view. Bar = 0.1 mm. See also Plate 1 b." httpUri="https://zenodo.org/record/184691/files/figure.png" pageId="2" pageNumber="31">Fig. 2</figureCitation>
|
||
& Pl. 1, fig. b). Postocular setae apparently absent. Three large ocelli in a narrow triangle without ocellar setae. Frons broad and face broader than frons. Clypeus bare. Proboscis relatively long, with abundant fine, long hairs apically. Palp short and one-segmented, flat, slightly longer than broad (
|
||
<figureCitation id="74BDD59E0F501331FA917AE8FA94FB5F" box="[1291,1362,1160,1186]" captionStart="FIGURE 2" captionStartId="4.[151,254,1492,1516]" captionTargetBox="[422,1163,202,1460]" captionTargetId="figure@4.[376,1210,197,1467]" captionTargetPageId="4" captionText="FIGURE 2. Camera lucida drawing of the head of Microphorites utrillensis Peñalver nov. sp. (Dolichopodidae), female holotype (CPT- 963), in ventral view. Bar = 0.1 mm. See also Plate 1 b." httpUri="https://zenodo.org/record/184691/files/figure.png" pageId="2" pageNumber="31">Fig. 2</figureCitation>
|
||
),
|
||
<quantity id="2B7E64FE0F501331FAF27AE8FF03FB37" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="2" pageNumber="31" unit="mm" value="0.07">0.07 mm</quantity>
|
||
long and
|
||
<quantity id="2B7E64FE0F501331FEA27AD0FE66FB37" box="[312,416,1200,1226]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="31" unit="mm" value="0.04">0.04 mm</quantity>
|
||
wide, with short, fine hairs. Strong setae around mouthparts. Aristate antenna
|
||
<quantity id="2B7E64FE0F501331FAAE7AD0FA5AFB37" box="[1332,1436,1200,1226]" metricMagnitude="-4" metricUnit="m" metricValue="2.6" pageId="2" pageNumber="31" unit="mm" value="0.26">0.26 mm</quantity>
|
||
long with fine, dense microtrichia. Scape reduced. Pedicel
|
||
<quantity id="2B7E64FE0F501331FCDF7AB8FC68FB0F" box="[837,942,1240,1266]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="31" unit="mm" value="0.04">0.04 mm</quantity>
|
||
long,
|
||
<quantity id="2B7E64FE0F501331FC6D7AB8FBA6FB0F" box="[1015,1120,1240,1266]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="31" unit="mm" value="0.05">0.05 mm</quantity>
|
||
wide, with a ring of strong setae apically. Basal flagellomere subtriangular, bilaterally symmetrical, laterally compressed, elongate, basally swollen,
|
||
<quantity id="2B7E64FE0F501331FEC07B48FE04FABF" box="[346,450,1320,1346]" metricMagnitude="-4" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="31" unit="mm" value="0.12">0.12 mm</quantity>
|
||
long,
|
||
<quantity id="2B7E64FE0F501331FD917B48FDB5FABF" box="[523,627,1320,1346]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="31" unit="mm" value="0.04">0.04 mm</quantity>
|
||
greatest width,
|
||
<quantity id="2B7E64FE0F501331FCB67B48FC52FABF" box="[812,916,1320,1346]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="31" unit="mm" value="0.05">0.05 mm</quantity>
|
||
greatest height. Arista apical and short,
|
||
<quantity id="2B7E64FE0F501331FAF27B48FF03FA97" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="31" unit="mm" value="0.1">0.10 mm</quantity>
|
||
long, and 2-articulated, basal article very small, slightly longer than width (
|
||
<quantity id="2B7E64FE0F501331FBDF7B30FB76FA97" box="[1093,1200,1360,1386]" metricMagnitude="-5" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="31" unit="mm" value="0.01">0.01 mm</quantity>
|
||
long). Distal article
|
||
<quantity id="2B7E64FE0F501331FF0D7B18FF39FA6F" box="[151,255,1400,1426]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="31" unit="mm" value="0.09">0.09 mm</quantity>
|
||
long.
|
||
</paragraph>
|
||
<paragraph id="EC39C91B0F501331FF5F7BC0FB6BFA1F" blockId="2.[151,1437,434,2038]" pageId="2" pageNumber="31">
|
||
Thorax
|
||
<quantity id="2B7E64FE0F501331FEB97BC0FE4BFA47" box="[291,397,1440,1466]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="2" pageNumber="31" unit="mm" value="0.45">0.45 mm</quantity>
|
||
long. Proepisternum and prosternum bare. Few strong thoracic setae; scutum with two long, strong setae in a dorsal position and scutellum with two pairs of strong apical setae.
|
||
</paragraph>
|
||
<paragraph id="EC39C91B0F501331FF5F7B90FC80F8D3" blockId="2.[151,1437,434,2038]" pageId="2" pageNumber="31">
|
||
Wings
|
||
<quantity id="2B7E64FE0F501331FE8D7B90FEB9F9F7" box="[279,383,1520,1546]" metricMagnitude="-3" metricUnit="m" metricValue="1.18" pageId="2" pageNumber="31" unit="mm" value="1.18">1.18 mm</quantity>
|
||
long,
|
||
<quantity id="2B7E64FE0F501331FE527B90FDF6F9F7" box="[456,560,1520,1546]" metricMagnitude="-4" metricUnit="m" metricValue="4.4" pageId="2" pageNumber="31" unit="mm" value="0.44">0.44 mm</quantity>
|
||
wide, without a costal bristle, and only very slightly pointed at apex. Apart from the diagnostic venational characters the wing venation (
|
||
<figureCitation id="74BDD59E0F501331FCC47878FC64F9CF" box="[862,930,1560,1586]" captionStart="FIGURE 3" captionStartId="5.[151,254,1642,1666]" captionTargetBox="[166,1421,680,1607]" captionTargetId="figure@5.[151,1436,655,1618]" captionTargetPageId="5" captionText="FIGURE 3. Camera lucida drawings of the a) wing (crossvein h obscure), b) hind leg and c) distal part of the abdomen in ventral view. Microphorites utrillensis Peñalver nov. sp., female holotype (CPT- 963). cerc = cercus. All to the same scale (bar = 0.1 mm). See also Plate 1 c." httpUri="https://zenodo.org/record/184692/files/figure.png" pageId="2" pageNumber="31">Fig. 3</figureCitation>
|
||
a) shows a C with one row of spinules, a Sc parallel with R1 except for a short curved stem which apparently touches it apically (the connection with C is not visible; possibly distal part is evanescent, i.e., Sc vein incomplete), all longitudinal veins complete, an R4+5 unbranched and reaching very close to the wing apex, an M1 straight beyond crossvein dm-cu and flexed apically, a crossvein r-m short and near wing base, an m-cu vein tubular and inclined towards wing apex, a bm-cu crossvein complete, and a CuA2 curved. The veins are not setose, except C, and the crossvein h is obscured (not drawn in
|
||
<figureCitation id="74BDD59E0F501331FEA67974FE4AF8D3" box="[316,396,1812,1838]" captionStart="FIGURE 1" captionStartId="3.[151,254,1302,1326]" captionTargetBox="[171,1419,203,1267]" captionTargetId="figure@3.[153,1438,197,1278]" captionTargetPageId="3" captionText="FIGURE 1. Camera lucida drawings of Microphorites utrillensis Peñalver nov. sp. (Dolichopodidae), female holotype (CPT- 963), in ventral and dorsal habitus, from San Just Amber (Teruel Province). Bar = 1 mm. See also Plate 1 a." httpUri="https://zenodo.org/record/184690/files/figure.png" pageId="2" pageNumber="31">Figs. 1</figureCitation>
|
||
&
|
||
<figureCitation id="74BDD59E0F501331FE2B7974FE06F8D3" box="[433,448,1812,1838]" captionStart="FIGURE 3" captionStartId="5.[151,254,1642,1666]" captionTargetBox="[166,1421,680,1607]" captionTargetId="figure@5.[151,1436,655,1618]" captionTargetPageId="5" captionText="FIGURE 3. Camera lucida drawings of the a) wing (crossvein h obscure), b) hind leg and c) distal part of the abdomen in ventral view. Microphorites utrillensis Peñalver nov. sp., female holotype (CPT- 963). cerc = cercus. All to the same scale (bar = 0.1 mm). See also Plate 1 c." httpUri="https://zenodo.org/record/184692/files/figure.png" pageId="2" pageNumber="31">3</figureCitation>
|
||
a). Anal lobe not well developed.
|
||
</paragraph>
|
||
<paragraph id="EC39C91B0F501331FF5F795CFC6EF80B" blockId="2.[151,1437,434,2038]" pageId="2" pageNumber="31">
|
||
Legs not modified and moderately bristly. Fore leg 1.00 mm long (femur
|
||
<quantity id="2B7E64FE0F501331FBDA795CFB69F8AB" box="[1088,1199,1852,1878]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="2" pageNumber="31" unit="mm" value="0.33">0.33 mm</quantity>
|
||
; tibia
|
||
<quantity id="2B7E64FE0F501331FB61795CFAACF8AB" box="[1275,1386,1852,1878]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="31" unit="mm" value="0.35">0.35 mm</quantity>
|
||
; ta1
|
||
<quantity id="2B7E64FE0F501331FF0D7904FEC4F883" box="[151,258,1892,1918]" metricMagnitude="-4" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="31" unit="mm" value="0.12">0.12 mm</quantity>
|
||
, ta2
|
||
<quantity id="2B7E64FE0F501331FEA27904FE62F883" box="[312,420,1892,1918]" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="31" unit="mm" value="0.06">0.06 mm</quantity>
|
||
, ta3 0.05, ta4
|
||
<quantity id="2B7E64FE0F501331FDDD7904FD69F883" box="[583,687,1892,1918]" metricMagnitude="-5" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="31" unit="mm" value="0.03">0.03 mm</quantity>
|
||
and ta5
|
||
<quantity id="2B7E64FE0F501331FC887904FCB9F883" box="[786,895,1892,1918]" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="31" unit="mm" value="0.06">0.06 mm</quantity>
|
||
). Mid legs difficult to measure (tarsus approximately
|
||
<quantity id="2B7E64FE0F501331FF6279ECFEA1F85B" box="[248,359,1932,1958]" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" pageId="2" pageNumber="31" unit="mm" value="0.36">0.36 mm</quantity>
|
||
).
|
||
<collectingCountry id="9491898B0F501331FEE679ECFE71F85B" box="[380,439,1932,1958]" name="India" pageId="2" pageNumber="31">Hind</collectingCountry>
|
||
leg
|
||
<quantity id="2B7E64FE0F501331FE7779ECFD91F85B" box="[493,599,1932,1958]" metricMagnitude="-3" metricUnit="m" metricValue="1.02" pageId="2" pageNumber="31" unit="mm" value="1.02">1.02 mm</quantity>
|
||
long (femur
|
||
<quantity id="2B7E64FE0F501331FD6C79ECFCA5F85B" box="[758,867,1932,1958]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="31" unit="mm" value="0.35">0.35 mm</quantity>
|
||
; tibia
|
||
<quantity id="2B7E64FE0F501331FC3479ECFBDAF85B" box="[942,1052,1932,1958]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="2" pageNumber="31" unit="mm" value="0.32">0.32 mm</quantity>
|
||
; ta1
|
||
<quantity id="2B7E64FE0F501331FBCD79ECFB02F85B" box="[1111,1220,1932,1958]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="2" pageNumber="31" unit="mm" value="0.16">0.16 mm</quantity>
|
||
, ta2
|
||
<quantity id="2B7E64FE0F501331FB6479ECFAADF85B" box="[1278,1387,1932,1958]" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="31" unit="mm" value="0.06">0.06 mm</quantity>
|
||
, ta3
|
||
<quantity id="2B7E64FE0F501331FF0D79D4FEC2F833" box="[151,260,1972,1998]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="31" unit="mm" value="0.04">0.04 mm</quantity>
|
||
, ta4
|
||
<quantity id="2B7E64FE0F501331FEA479D4FE6FF833" box="[318,425,1972,1998]" metricMagnitude="-5" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="31" unit="mm" value="0.03">0.03 mm</quantity>
|
||
and ta5
|
||
<quantity id="2B7E64FE0F501331FD8879D4FD44F833" box="[530,642,1972,1998]" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="31" unit="mm" value="0.06">0.06 mm</quantity>
|
||
); tibia not clavate and first tarsomere of normal shape. Empodium apparently small or setiform, and pulvilli well-developed (
|
||
<figureCitation id="74BDD59E0F501331FCD979BCFC4EF80B" box="[835,904,2012,2038]" captionStart="FIGURE 3" captionStartId="5.[151,254,1642,1666]" captionTargetBox="[166,1421,680,1607]" captionTargetId="figure@5.[151,1436,655,1618]" captionTargetPageId="5" captionText="FIGURE 3. Camera lucida drawings of the a) wing (crossvein h obscure), b) hind leg and c) distal part of the abdomen in ventral view. Microphorites utrillensis Peñalver nov. sp., female holotype (CPT- 963). cerc = cercus. All to the same scale (bar = 0.1 mm). See also Plate 1 c." httpUri="https://zenodo.org/record/184692/files/figure.png" pageId="2" pageNumber="31">Fig. 3</figureCitation>
|
||
b).
|
||
</paragraph>
|
||
<caption id="B8F999930F511330FF0D7B76FA8FFAAD" httpUri="https://zenodo.org/record/184690/files/figure.png" pageId="3" pageNumber="32" targetBox="[171,1419,203,1267]" targetPageId="3">
|
||
<paragraph id="EC39C91B0F511330FF0D7B76FA8FFAAD" blockId="3.[151,1436,1302,1360]" pageId="3" pageNumber="32">
|
||
<emphasis id="DEF215090F511330FF0D7B76FEDCFAD3" bold="true" box="[151,282,1302,1326]" pageId="3" pageNumber="32">FIGURE 1.</emphasis>
|
||
Camera lucida drawings of
|
||
<taxonomicName id="2B86B2980F511330FDD57B77FC79FAD0" authority="Penalver" authorityName="Penalver" box="[591,959,1302,1326]" class="Insecta" family="Empididae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="utrillensis">
|
||
<emphasis id="DEF215090F511330FDD57B77FC9CFAD3" box="[591,858,1303,1326]" italics="true" pageId="3" pageNumber="32">Microphorites utrillensis</emphasis>
|
||
Peñalver
|
||
</taxonomicName>
|
||
nov. sp. (
|
||
<taxonomicName id="2B86B2980F511330FBB07B76FB1DFAD0" box="[1066,1243,1302,1325]" class="Insecta" family="Dolichopodidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="32" phylum="Arthropoda" rank="family">Dolichopodidae</taxonomicName>
|
||
), female holotype (CPT-963), in ventral and dorsal habitus, from San Just Amber (Teruel Province). Bar = 1 mm. See also Plate 1a.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="EC39C91B0F511330FF5F7BE4FD41F99B" blockId="3.[151,1437,1412,2038]" pageId="3" pageNumber="32">
|
||
Abdomen telescoped, elongate,
|
||
<quantity id="2B7E64FE0F511330FDA67BE4FD63FA63" box="[572,677,1412,1438]" metricMagnitude="-3" metricUnit="m" metricValue="1.14" pageId="3" pageNumber="32" unit="mm" value="1.14">1.14 mm</quantity>
|
||
long,
|
||
<quantity id="2B7E64FE0F511330FD757BE4FC9EFA63" box="[751,856,1412,1438]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="32" unit="mm" value="0.3">0.30 mm</quantity>
|
||
wide, with the terminalia extruded (
|
||
<figureCitation id="74BDD59E0F511330FA997BE4FA8EFA63" box="[1283,1352,1412,1438]" captionStart="FIGURE 3" captionStartId="5.[151,254,1642,1666]" captionTargetBox="[166,1421,680,1607]" captionTargetId="figure@5.[151,1436,655,1618]" captionTargetPageId="5" captionText="FIGURE 3. Camera lucida drawings of the a) wing (crossvein h obscure), b) hind leg and c) distal part of the abdomen in ventral view. Microphorites utrillensis Peñalver nov. sp., female holotype (CPT- 963). cerc = cercus. All to the same scale (bar = 0.1 mm). See also Plate 1 c." httpUri="https://zenodo.org/record/184692/files/figure.png" pageId="3" pageNumber="32">Fig. 3</figureCitation>
|
||
c & Pl. 1, fig. c) (abdomen partially destroyed and impregnated with resin). Tergite 10 short (
|
||
<quantity id="2B7E64FE0F511330FB127BCCFB35FA3B" box="[1160,1267,1452,1478]" metricMagnitude="-5" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="32" unit="mm" value="0.03">0.03 mm</quantity>
|
||
long and
|
||
<quantity id="2B7E64FE0F511330FAF37BCCFF03FA13" metricMagnitude="-5" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="32" unit="mm" value="0.03">0.03 mm</quantity>
|
||
wide), with sclerite apparently undivided, and without acanthophorite apical spines. Apical segments of the abdomen with long setulae (segment 10 with setulae only in the cerci). Cerci sclerotized (apparently heavily sclerotized), short (
|
||
<quantity id="2B7E64FE0F511330FE4D7844FD87F9C3" box="[471,577,1572,1598]" metricMagnitude="-5" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="32" unit="mm" value="0.03">0.03 mm</quantity>
|
||
long), held horizontally, lateral in position, with at least four strong bristles each (up to
|
||
<quantity id="2B7E64FE0F511330FECA782CFE7DF99B" box="[336,443,1612,1638]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="32" unit="mm" value="0.05">0.05 mm</quantity>
|
||
) and few setulae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A49C9A900F511330FF5F7813FAC0F8AB" pageId="3" pageNumber="32" type="discussion">
|
||
<paragraph id="EC39C91B0F511330FF5F7813FAC0F8AB" blockId="3.[151,1437,1412,2038]" pageId="3" pageNumber="32">
|
||
<emphasis id="DEF215090F511330FF5F7813FE8FF970" bold="true" box="[197,329,1651,1677]" pageId="3" pageNumber="32">Discussion</emphasis>
|
||
: This female specimen found in San Just Amber clearly belongs to the genus
|
||
<taxonomicName id="2B86B2980F511330FB6F7814FEFCF94B" authority="Hennig, 1971" authorityName="Hennig" authorityYear="1971" class="Insecta" family="Empididae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DEF215090F511330FB6F7814FA5DF970" box="[1269,1435,1652,1677]" italics="true" pageId="3" pageNumber="32">Microphorites</emphasis>
|
||
<bibRefCitation id="8817B4EA0F511330FF0D78FCFEFCF94B" author="Hennig" box="[151,314,1692,1718]" pageId="3" pageNumber="40" refString="Hennig, W. (1971) Insektenfossilien aus der unteren Kreide. III. Empidiformia (" Microphoridae ") aus der unteren Kreide und aus dem Baltischen Bernstein; ein Vertreter der Cyclorrhapha aus der unteren Kreide. Stuttgarter Beitrage zur Naturkunde, 232, 1 - 28." type="journal article" year="1971">Hennig, 1971</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="2B86B2980F511330FED178FCFDCBF94B" box="[331,525,1692,1718]" class="Insecta" family="Dolichopodidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="32" phylum="Arthropoda" rank="family">Dolichopodidae</taxonomicName>
|
||
:
|
||
<taxonomicName id="2B86B2980F511330FD8078FCFD08F94B" box="[538,718,1692,1718]" class="Insecta" family="Dolichopodidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="32" phylum="Arthropoda" rank="subFamily" subFamily="Microphorinae">Microphorinae</taxonomicName>
|
||
). It shows all the characters indicated by
|
||
<bibRefCitation id="8817B4EA0F511330FB5E78FCFEEEF923" author="Grimaldi" pageId="3" pageNumber="40" refString="Grimaldi, D. A. & Cumming, J. (1999) Brachyceran Diptera in Cretaceous ambers and Mesozoic diversification of the Eremoneura. Bulletin of the American Museum of Natural History, 239, 124 pp." type="book" year="1999">Grimaldi & Cumming (1999)</bibRefCitation>
|
||
in the translated and summarized diagnosis of
|
||
<bibRefCitation id="8817B4EA0F511330FCD578A4FC38F923" author="Hennig" box="[847,1022,1732,1758]" pageId="3" pageNumber="40" refString="Hennig, W. (1971) Insektenfossilien aus der unteren Kreide. III. Empidiformia (" Microphoridae ") aus der unteren Kreide und aus dem Baltischen Bernstein; ein Vertreter der Cyclorrhapha aus der unteren Kreide. Stuttgarter Beitrage zur Naturkunde, 232, 1 - 28." type="journal article" year="1971">Hennig (1971)</bibRefCitation>
|
||
. It corresponds to a new microphorine species which differentiates from the four species found in the Early Cretaceous ambers from
|
||
<collectingCountry id="9491898B0F511330FAAC788CFA5BF8FB" box="[1334,1437,1772,1798]" name="Lebanon" pageId="3" pageNumber="32">Lebanon</collectingCountry>
|
||
[
|
||
<taxonomicName id="2B86B2980F511330FF3B7974FEE9F8D0" box="[161,303,1812,1837]" class="Insecta" family="Empididae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="extinctus">
|
||
<emphasis id="DEF215090F511330FF3B7974FEE9F8D0" box="[161,303,1812,1837]" italics="true" pageId="3" pageNumber="32">M. extinctus</emphasis>
|
||
</taxonomicName>
|
||
, female (
|
||
<bibRefCitation id="8817B4EA0F511330FE067974FD86F8D3" author="Hennig" box="[412,576,1812,1838]" pageId="3" pageNumber="40" refString="Hennig, W. (1971) Insektenfossilien aus der unteren Kreide. III. Empidiformia (" Microphoridae ") aus der unteren Kreide und aus dem Baltischen Bernstein; ein Vertreter der Cyclorrhapha aus der unteren Kreide. Stuttgarter Beitrage zur Naturkunde, 232, 1 - 28." type="journal article" year="1971">Hennig, 1971</bibRefCitation>
|
||
),
|
||
<emphasis id="DEF215090F511330FDCE7974FD09F8D0" box="[596,719,1812,1837]" italics="true" pageId="3" pageNumber="32">
|
||
<taxonomicName id="2B86B2980F511330FDCE7974FD0CF8D0" box="[596,714,1812,1837]" class="Insecta" family="Empididae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="similis">M. similis</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
female,
|
||
<emphasis id="DEF215090F511330FCAE7974FC7DF8D0" box="[820,955,1812,1837]" italics="true" pageId="3" pageNumber="32">
|
||
<taxonomicName id="2B86B2980F511330FCAE7974FC70F8D0" box="[820,950,1812,1837]" class="Insecta" family="Empididae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="oculeus">M. oculeus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
male (
|
||
<bibRefCitation id="8817B4EA0F511330FB937974FA9DF8D3" author="Grimaldi" box="[1033,1371,1812,1838]" pageId="3" pageNumber="40" refString="Grimaldi, D. A. & Cumming, J. (1999) Brachyceran Diptera in Cretaceous ambers and Mesozoic diversification of the Eremoneura. Bulletin of the American Museum of Natural History, 239, 124 pp." type="book" year="1999">Grimaldi & Cumming, 1999</bibRefCitation>
|
||
)] and
|
||
<collectingCountry id="9491898B0F511330FF0D795CFF21F8AB" box="[151,231,1852,1878]" name="France" pageId="3" pageNumber="32">France</collectingCountry>
|
||
[
|
||
<taxonomicName id="2B86B2980F511330FF62795CFE42F8A8" box="[248,388,1852,1877]" class="Insecta" family="Empididae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="deploegi">
|
||
<emphasis id="DEF215090F511330FF62795CFE42F8A8" box="[248,388,1852,1877]" italics="true" pageId="3" pageNumber="32">M. deploegi</emphasis>
|
||
</taxonomicName>
|
||
males and females (
|
||
<bibRefCitation id="8817B4EA0F511330FDE9795CFCF7F8AB" author="Nel" box="[627,817,1852,1878]" pageId="3" pageNumber="40" refString="Nel, A., Perrichot, V., Daugeron, Ch. & Neraudeau, D. (2004) A new Microphorites in the Lower Cretaceous amber of the Southwest of France (Diptera: Dolichopodidae, " Microphorinae "). Annales de la Societe Entomologique de France (n. s.), 40 (1), 23 - 29." type="journal article" year="2004">
|
||
Nel
|
||
<emphasis id="DEF215090F511330FD3C795CFD27F8A8" box="[678,737,1852,1877]" italics="true" pageId="3" pageNumber="32">et al.</emphasis>
|
||
, 2004
|
||
</bibRefCitation>
|
||
)]. It is closest to the last one mentioned.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A49C9A900F511336FF5C7904FAA2F870" lastPageId="5" lastPageNumber="34" pageId="3" pageNumber="32" type="reference_group">
|
||
<paragraph id="EC39C91B0F511336FF5C7904FE4EFD8F" blockId="3.[151,1437,1412,2038]" lastBlockId="5.[151,1437,152,626]" lastPageId="5" lastPageNumber="34" pageId="3" pageNumber="32">
|
||
<taxonomicName id="2B86B2980F511330FF5C7904FE1BF880" box="[198,477,1892,1917]" class="Insecta" family="Empididae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="deploegi">
|
||
<emphasis id="DEF215090F511330FF5C7904FE1BF880" box="[198,477,1892,1917]" italics="true" pageId="3" pageNumber="32">Microphorites deploegi</emphasis>
|
||
</taxonomicName>
|
||
from Charente-Maritime (
|
||
<collectingCountry id="9491898B0F511330FCBB7904FCB2F883" box="[801,884,1892,1918]" name="France" pageId="3" pageNumber="32">France</collectingCountry>
|
||
) was described from male and female specimens, but the female was not illustrated. (
|
||
<bibRefCitation id="8817B4EA0F511330FD0779ECFCA0F85B" author="Nel" box="[669,870,1932,1958]" pageId="3" pageNumber="40" refString="Nel, A., Perrichot, V., Daugeron, Ch. & Neraudeau, D. (2004) A new Microphorites in the Lower Cretaceous amber of the Southwest of France (Diptera: Dolichopodidae, " Microphorinae "). Annales de la Societe Entomologique de France (n. s.), 40 (1), 23 - 29." type="journal article" year="2004">
|
||
Nel
|
||
<emphasis id="DEF215090F511330FD4E79ECFCCAF858" box="[724,780,1932,1957]" italics="true" pageId="3" pageNumber="32">et al</emphasis>
|
||
., 2004
|
||
</bibRefCitation>
|
||
). The well preserved new female specimen of
|
||
<taxonomicName id="2B86B2980F511330FF0D79D4FEF8F830" box="[151,318,1972,1997]" class="Insecta" family="Empididae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DEF215090F511330FF0D79D4FEF8F830" box="[151,318,1972,1997]" italics="true" pageId="3" pageNumber="32">Microphorites</emphasis>
|
||
</taxonomicName>
|
||
discovered in San Just has been compared in the MNHN of Paris to two
|
||
<typeStatus id="333D77B90F511330FB1579D4FB32F833" box="[1167,1268,1972,1998]" pageId="3" pageNumber="32" type="paratype">paratype</typeStatus>
|
||
females of
|
||
<taxonomicName id="2B86B2980F511330FAE679D4FF3AF808" class="Insecta" family="Empididae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="deploegi">
|
||
<emphasis id="DEF215090F511330FAE679D4FF3AF808" italics="true" pageId="3" pageNumber="32">M. deploegi</emphasis>
|
||
</taxonomicName>
|
||
(ARC 118-R and ARC 198.4-R). Female
|
||
<taxonomicName id="2B86B2980F511330FD7079BCFBCBF808" box="[746,1037,2012,2037]" class="Insecta" family="Empididae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="utrillensis">
|
||
<emphasis id="DEF215090F511330FD7079BCFBCBF808" box="[746,1037,2012,2037]" italics="true" pageId="3" pageNumber="32">Microphorites utrillensis</emphasis>
|
||
</taxonomicName>
|
||
differs from female
|
||
<taxonomicName id="2B86B2980F511330FB6579BCFA4DF808" box="[1279,1419,2012,2037]" class="Insecta" family="Empididae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="deploegi">
|
||
<emphasis id="DEF215090F511330FB6579BCFA4DF808" box="[1279,1419,2012,2037]" italics="true" pageId="3" pageNumber="32">M. deploegi</emphasis>
|
||
</taxonomicName>
|
||
in having body that is graceful and smaller (
|
||
<quantity id="2B7E64FE0F561337FD3A785CFD3DF9AB" box="[672,763,1596,1622]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="4" pageNumber="33" unit="mm" value="1.7">1.7 mm</quantity>
|
||
long
|
||
<emphasis id="DEF215090F561337FCA5785DFC91F9AB" box="[831,855,1597,1622]" italics="true" pageId="4" pageNumber="33">vs</emphasis>
|
||
. ca. 2.0 mm), arista approximately as long as the flagellomere I (
|
||
<emphasis id="DEF215090F561337FED67802FEA2F986" box="[332,356,1634,1659]" italics="true" pageId="4" pageNumber="33">vs</emphasis>
|
||
. approximately twice as long in
|
||
<taxonomicName id="2B86B2980F561337FD787802FCABF986" box="[738,877,1634,1659]" class="Insecta" family="Empididae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="33" phylum="Arthropoda" rank="species" species="deploegi">
|
||
<emphasis id="DEF215090F561337FD787802FCABF986" box="[738,877,1634,1659]" italics="true" pageId="4" pageNumber="33">M. deploegi</emphasis>
|
||
</taxonomicName>
|
||
), wing approximately half long (
|
||
<quantity id="2B7E64FE0F561337FB757802FA8CF981" box="[1263,1354,1634,1660]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="33" unit="mm" value="1.2">1.2 mm</quantity>
|
||
<emphasis id="DEF215090F561337FACB7802FAAFF986" box="[1361,1385,1634,1659]" italics="true" pageId="4" pageNumber="33">vs</emphasis>
|
||
.
|
||
<quantity id="2B7E64FE0F561337FAED7802FF03F95C" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="4" pageNumber="33" unit="mm" value="2.2">2.2 mm</quantity>
|
||
in French specimen ARC 118-R), costal vein ending at the end of M1, abdomen not globular (1.1 x
|
||
<quantity id="2B7E64FE0F561337FAED78E7FF03F934" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="33" unit="mm" value="0.3">0.3 mm</quantity>
|
||
<emphasis id="DEF215090F561337FF5478D0FF20F934" box="[206,230,1712,1737]" italics="true" pageId="4" pageNumber="33">vs</emphasis>
|
||
. 0.8 x
|
||
<quantity id="2B7E64FE0F561337FEA478CFFE58F934" box="[318,414,1711,1737]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="33" unit="mm" value="0.4">0.4 mm</quantity>
|
||
), and fewer and finer setae on head, thorax and legs, with two pairs (
|
||
<emphasis id="DEF215090F561337FB7F78D0FB3BF934" box="[1253,1277,1712,1737]" italics="true" pageId="4" pageNumber="33">vs</emphasis>
|
||
. one pair) of strong apical setae on the scutellum. Interestingly,
|
||
<taxonomicName id="2B86B2980F561337FD6378B5FBD9F913" box="[761,1055,1749,1774]" class="Insecta" family="Empididae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="33" phylum="Arthropoda" rank="species" species="utrillensis">
|
||
<emphasis id="DEF215090F561337FD6378B5FBD9F913" box="[761,1055,1749,1774]" italics="true" pageId="4" pageNumber="33">Microphorites utrillensis</emphasis>
|
||
</taxonomicName>
|
||
has cerci with very long apical setae as in extant genera
|
||
<taxonomicName id="2B86B2980F561337FE50789AFDBAF8EE" box="[458,636,1786,1811]" class="Insecta" family="Dolichopodidae" genus="Heteropsilopus" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DEF215090F561337FE50789AFDBAF8EE" box="[458,636,1786,1811]" italics="true" pageId="4" pageNumber="33">Heteropsilopus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="DEF215090F561337FD21789AFCFBF8EE" box="[699,829,1786,1811]" italics="true" pageId="4" pageNumber="33">
|
||
<taxonomicName id="2B86B2980F561337FD21789AFCFFF8EE" ID-CoL="5S7S" box="[699,825,1786,1811]" class="Insecta" family="Dolichopodidae" genus="Microphor" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="33" phylum="Arthropoda" rank="genus">Microphor</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
among other dolichopodids (see
|
||
<bibRefCitation id="8817B4EA0F561337FB48789AFEE7F8C4" author="Sinclair" pageId="4" pageNumber="40" refString="Sinclair, B. J. & Cumming, J. M. (2006) The morphology, higher-level phylogeny and classification of the Empidoidea (Diptera). Zootaxa, 1180, 1 - 172." type="journal article" year="2006">Sinclair & Cumming, 2006</bibRefCitation>
|
||
, p. 46), an unlike
|
||
<taxonomicName id="2B86B2980F561337FE6C7940FD42F8C4" box="[502,644,1824,1849]" class="Insecta" family="Empididae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="33" phylum="Arthropoda" rank="species" species="deploegi">
|
||
<emphasis id="DEF215090F561337FE6C7940FD42F8C4" box="[502,644,1824,1849]" italics="true" pageId="4" pageNumber="33">M. deploegi</emphasis>
|
||
</taxonomicName>
|
||
which has short setulae. The male
|
||
<typeStatus id="333D77B90F561337FBBF797FFB4DF8C4" box="[1061,1163,1823,1849]" pageId="4" pageNumber="33" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="2B86B2980F561337FB2E7940FA87F8C4" box="[1204,1345,1824,1849]" class="Insecta" family="Empididae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="33" phylum="Arthropoda" rank="species" species="deploegi">
|
||
<emphasis id="DEF215090F561337FB2E7940FA87F8C4" box="[1204,1345,1824,1849]" italics="true" pageId="4" pageNumber="33">M. deploegi</emphasis>
|
||
</taxonomicName>
|
||
has the crossvein r-m spectral and m-cu evanescent (similarly in the female ARC 118-R), both of which are clearly tubular in the new species (like in the female ARC 198.4-R of
|
||
<taxonomicName id="2B86B2980F561337FCEA790AFC3AF87E" box="[880,1020,1898,1923]" class="Insecta" family="Empididae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="33" phylum="Arthropoda" rank="species" species="deploegi">
|
||
<emphasis id="DEF215090F561337FCEA790AFC3AF87E" box="[880,1020,1898,1923]" italics="true" pageId="4" pageNumber="33">M. deploegi</emphasis>
|
||
</taxonomicName>
|
||
), and its Sc vein is different as well (curved back towards R1, closely parallel with R1 but not touching it, connecting C far from the end of R
|
||
<quantity id="2B7E64FE0F561337FACA79FFFAB0F854" box="[1360,1398,1935,1965]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="4" pageNumber="33" unit="in" value="1.0">1 in</quantity>
|
||
<taxonomicName id="2B86B2980F561337FAE679F0FF3AF82C" class="Insecta" family="Empididae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="33" phylum="Arthropoda" rank="species" species="deploegi">
|
||
<emphasis id="DEF215090F561337FAE679F0FF3AF82C" italics="true" pageId="4" pageNumber="33">M. deploegi</emphasis>
|
||
</taxonomicName>
|
||
). In contrast with the new species, a complete Sc connecting C is also present in fossil
|
||
<taxonomicName id="2B86B2980F561337FB6D79D8FAACF82C" box="[1271,1386,1976,2001]" class="Insecta" family="Empididae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="33" phylum="Arthropoda" rank="species" species="similis">
|
||
<emphasis id="DEF215090F561337FB6D79D8FAACF82C" box="[1271,1386,1976,2001]" italics="true" pageId="4" pageNumber="33">M. similis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="2B86B2980F571336FF0D7EF8FED1FF4C" box="[151,279,152,177]" class="Insecta" family="Empididae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="oculeus">
|
||
<emphasis id="DEF215090F571336FF0D7EF8FED1FF4C" box="[151,279,152,177]" italics="true" pageId="5" pageNumber="34">M. oculeus</emphasis>
|
||
</taxonomicName>
|
||
, and in the extant
|
||
<taxonomicName id="2B86B2980F571336FE767EF8FD41FF4C" ID-CoL="9G2RY" box="[492,647,152,177]" class="Insecta" family="Dolichopodidae" genus="Schistostoma" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DEF215090F571336FE767EF8FD41FF4C" box="[492,647,152,177]" italics="true" pageId="5" pageNumber="34">Schistostoma</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="8817B4EA0F571336FD5E7EF8FBD5FF4F" author="Grimaldi" box="[708,1043,152,178]" pageId="5" pageNumber="40" refString="Grimaldi, D. A. & Cumming, J. (1999) Brachyceran Diptera in Cretaceous ambers and Mesozoic diversification of the Eremoneura. Bulletin of the American Museum of Natural History, 239, 124 pp." type="book" year="1999">Grimaldi & Cumming, 1999</bibRefCitation>
|
||
, figs. 37–38 and
|
||
<bibRefCitation id="8817B4EA0F571336FB437EF8FED9FF2A" author="Sinclair" pageId="5" pageNumber="40" refString="Sinclair, B. J. & Cumming, J. M. (2006) The morphology, higher-level phylogeny and classification of the Empidoidea (Diptera). Zootaxa, 1180, 1 - 172." type="journal article" year="2006">Sinclair & Cumming, 2006</bibRefCitation>
|
||
, p. 35). In
|
||
<taxonomicName id="2B86B2980F571336FE067EDDFD96FF2A" authority="Sc" authorityName="Sc" box="[412,592,189,215]" class="Insecta" family="Empididae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="extinctus">
|
||
<emphasis id="DEF215090F571336FE067EDDFDEDFF2B" box="[412,555,189,214]" italics="true" pageId="5" pageNumber="34">M. extinctus</emphasis>
|
||
Sc
|
||
</taxonomicName>
|
||
connects C but it is evanescent apically (see
|
||
<bibRefCitation id="8817B4EA0F571336FBC57EDDFAC7FF2A" author="Hennig" box="[1119,1281,189,215]" pageId="5" pageNumber="40" refString="Hennig, W. (1971) Insektenfossilien aus der unteren Kreide. III. Empidiformia (" Microphoridae ") aus der unteren Kreide und aus dem Baltischen Bernstein; ein Vertreter der Cyclorrhapha aus der unteren Kreide. Stuttgarter Beitrage zur Naturkunde, 232, 1 - 28." type="journal article" year="1971">Hennig, 1971</bibRefCitation>
|
||
, fig. 21).
|
||
<bibRefCitation id="8817B4EA0F571336FAE87EDDFEE1FF01" pageId="5" pageNumber="40" refString="Nel, A., Perrichot, V., Daugeron, Ch. & Neraudeau, D. (2004) A new Microphorites in the Lower Cretaceous amber of the Southwest of France (Diptera: Dolichopodidae, " Microphorinae "). Annales de la Societe Entomologique de France (n. s.), 40 (1), 23 - 29." type="journal article">
|
||
Nel
|
||
<emphasis id="DEF215090F571336FF0D7E82FF0CFF06" box="[151,202,226,251]" italics="true" pageId="5" pageNumber="34">et al</emphasis>
|
||
. (2004)
|
||
</bibRefCitation>
|
||
did not indicate why males and females were put in the same
|
||
<taxonomicName id="2B86B2980F571336FC647E82FB4CFF06" box="[1022,1162,226,251]" class="Insecta" family="Empididae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="deploegi">
|
||
<emphasis id="DEF215090F571336FC647E82FB4CFF06" box="[1022,1162,226,251]" italics="true" pageId="5" pageNumber="34">M. deploegi</emphasis>
|
||
</taxonomicName>
|
||
species; apparently the specimens were found in different amber masses, but the co-occurrence in the same outcrop of several specimens which exhibit similar morphological characteristics suggests they are of the same species. The French males and females have the same antennal morphology and it is in agreement with the sexually dimorphic variation seen in the
|
||
<taxonomicName id="2B86B2980F571336FE0A7F18FD87FE6F" ID-CoL="9G27N" box="[400,577,376,402]" class="Insecta" family="Dolichopodidae" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="subFamily" subFamily="Microphorinae">Microphorinae</taxonomicName>
|
||
, which affects to other characters as was indicated by
|
||
<bibRefCitation id="8817B4EA0F571336FB5D7F18FE99FE4A" author="Grimaldi" pageId="5" pageNumber="40" refString="Grimaldi, D. A. & Cumming, J. (1999) Brachyceran Diptera in Cretaceous ambers and Mesozoic diversification of the Eremoneura. Bulletin of the American Museum of Natural History, 239, 124 pp." type="book" year="1999">Grimaldi & Cumming (1999: 61)</bibRefCitation>
|
||
.
|
||
<bibRefCitation id="8817B4EA0F571336FEF77FFDFDFEFE4A" box="[365,568,413,439]" pageId="5" pageNumber="40" refString="Nel, A., Perrichot, V., Daugeron, Ch. & Neraudeau, D. (2004) A new Microphorites in the Lower Cretaceous amber of the Southwest of France (Diptera: Dolichopodidae, " Microphorinae "). Annales de la Societe Entomologique de France (n. s.), 40 (1), 23 - 29." type="journal article">
|
||
Nel
|
||
<emphasis id="DEF215090F571336FE387FFDFE1EFE4B" box="[418,472,413,438]" italics="true" pageId="5" pageNumber="34">et al</emphasis>
|
||
. (2004)
|
||
</bibRefCitation>
|
||
added to the diagnosis of the genus that the female tergite 10 bears no “acanthophorite” apical spines and that the female cercus is heavily sclerotized, held horizontally, bearing short setulae. The new specimen from San Just is the best known preserved female
|
||
<taxonomicName id="2B86B2980F571336FBFB7F88FACEFDFC" box="[1121,1288,488,513]" class="Insecta" family="Empididae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DEF215090F571336FBFB7F88FACEFDFC" box="[1121,1288,488,513]" italics="true" pageId="5" pageNumber="34">Microphorites</emphasis>
|
||
</taxonomicName>
|
||
and it is in a very clear amber mass, showing the terminalia extruded. This Spanish specimen confirms all the characters for the genus
|
||
<taxonomicName id="2B86B2980F571336FEAC7C52FE1BFDB6" box="[310,477,562,587]" class="Insecta" family="Empididae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DEF215090F571336FEAC7C52FE1BFDB6" box="[310,477,562,587]" italics="true" pageId="5" pageNumber="34">Microphorites</emphasis>
|
||
</taxonomicName>
|
||
added to the diagnosis by
|
||
<bibRefCitation id="8817B4EA0F571336FC8E7C52FC1EFDB1" box="[788,984,562,588]" pageId="5" pageNumber="40" refString="Nel, A., Perrichot, V., Daugeron, Ch. & Neraudeau, D. (2004) A new Microphorites in the Lower Cretaceous amber of the Southwest of France (Diptera: Dolichopodidae, " Microphorinae "). Annales de la Societe Entomologique de France (n. s.), 40 (1), 23 - 29." type="journal article">
|
||
Nel
|
||
<emphasis id="DEF215090F571336FCDC7C52FCBFFDB6" box="[838,889,562,587]" italics="true" pageId="5" pageNumber="34">et al</emphasis>
|
||
. (2004)
|
||
</bibRefCitation>
|
||
, with the single exception that the cercus has long setulae.
|
||
</paragraph>
|
||
<caption id="B8F999930F561337FF0D7BB4FC46F9F3" httpUri="https://zenodo.org/record/184691/files/figure.png" pageId="4" pageNumber="33" targetBox="[422,1163,202,1460]" targetPageId="4">
|
||
<paragraph id="EC39C91B0F561337FF0D7BB4FC46F9F3" blockId="4.[151,1436,1492,1550]" pageId="4" pageNumber="33">
|
||
<emphasis id="DEF215090F561337FF0D7BB4FEDEFA11" bold="true" box="[151,280,1492,1516]" pageId="4" pageNumber="33">FIGURE 2.</emphasis>
|
||
Camera lucida drawing of the head of
|
||
<taxonomicName id="2B86B2980F561337FD267BB4FBEFFA16" authority="Penalver" authorityName="Penalver" box="[700,1065,1492,1515]" class="Insecta" family="Empididae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="33" phylum="Arthropoda" rank="species" species="utrillensis">
|
||
<emphasis id="DEF215090F561337FD267BB4FC03FA16" box="[700,965,1492,1515]" italics="true" pageId="4" pageNumber="33">Microphorites utrillensis</emphasis>
|
||
Peñalver
|
||
</taxonomicName>
|
||
nov. sp. (
|
||
<taxonomicName id="2B86B2980F561337FB0B7BB4FA84FA16" ID-CoL="623XN" box="[1169,1346,1492,1515]" class="Insecta" family="Dolichopodidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="33" phylum="Arthropoda" rank="family">Dolichopodidae</taxonomicName>
|
||
), female holotype (CPT-963), in ventral view. Bar = 0.1 mm. See also Plate 1b.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="B8F999930F571336FF0D780AFDFAF93B" httpUri="https://zenodo.org/record/184692/files/figure.png" pageId="5" pageNumber="34" targetBox="[166,1421,680,1607]" targetPageId="5">
|
||
<paragraph id="EC39C91B0F571336FF0D780AFDFAF93B" blockId="5.[151,1436,1642,1734]" pageId="5" pageNumber="34">
|
||
<emphasis id="DEF215090F571336FF0D780AFEDFF97F" bold="true" box="[151,281,1642,1666]" pageId="5" pageNumber="34">FIGURE 3.</emphasis>
|
||
Camera lucida drawings of the a) wing (crossvein h obscure), b) hind leg and c) distal part of the abdomen in ventral view.
|
||
<taxonomicName id="2B86B2980F571336FEDD78EDFD7EF959" authority="Penalver" authorityName="Penalver" box="[327,696,1677,1700]" class="Insecta" family="Empididae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="utrillensis">
|
||
<emphasis id="DEF215090F571336FEDD78EDFD94F959" box="[327,594,1677,1700]" italics="true" pageId="5" pageNumber="34">Microphorites utrillensis</emphasis>
|
||
Peñalver
|
||
</taxonomicName>
|
||
nov. sp., female holotype (CPT-963). cerc = cercus. All to the same scale (bar = 0.1 mm). See also Plate 1c.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="EC39C91B0F571336FF5F7895FAA2F870" blockId="5.[151,1436,1781,2016]" pageId="5" pageNumber="34">
|
||
The wing venation of the new species is very similar to those of
|
||
<taxonomicName id="2B86B2980F571336FC7B7895FB3CF8F3" box="[993,1274,1781,1806]" class="Insecta" family="Empididae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="extinctus">
|
||
<emphasis id="DEF215090F571336FC7B7895FB3CF8F3" box="[993,1274,1781,1806]" italics="true" pageId="5" pageNumber="34">Microphorites extinctus</emphasis>
|
||
</taxonomicName>
|
||
described by
|
||
<bibRefCitation id="8817B4EA0F571336FF0D797DFE83F8CA" author="Hennig" box="[151,325,1821,1847]" pageId="5" pageNumber="40" refString="Hennig, W. (1971) Insektenfossilien aus der unteren Kreide. III. Empidiformia (" Microphoridae ") aus der unteren Kreide und aus dem Baltischen Bernstein; ein Vertreter der Cyclorrhapha aus der unteren Kreide. Stuttgarter Beitrage zur Naturkunde, 232, 1 - 28." type="journal article" year="1971">Hennig (1971)</bibRefCitation>
|
||
from Lebanese Amber, but differs mainly in having a different Sc vein, an R1 that ends at a basal position respect to m-cu crossvein, an R2+3 that ends more basally (in a medial position between the ends of R1 and R4+5), an m-cu vein inclined towards wing apex, a dm-cu not arched, and an elongate abdomen.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A49C9A900F571335FF5F79FEFA55FC6A" lastPageId="6" lastPageNumber="35" pageId="5" pageNumber="34" type="description">
|
||
<paragraph id="EC39C91B0F571335FF5F79FEFA4DFEF9" blockId="5.[151,1436,1781,2016]" lastBlockId="6.[151,1436,152,919]" lastPageId="6" lastPageNumber="35" pageId="5" pageNumber="34">
|
||
<taxonomicName id="2B86B2980F571336FF5F79FEFE2CF84A" box="[197,490,1950,1975]" class="Insecta" family="Empididae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="utrillensis">
|
||
<emphasis id="DEF215090F571336FF5F79FEFE2CF84A" box="[197,490,1950,1975]" italics="true" pageId="5" pageNumber="34">Microphorites utrillensis</emphasis>
|
||
</taxonomicName>
|
||
nov. sp. differs from
|
||
<taxonomicName id="2B86B2980F571336FD6A79FEFCA3F84A" box="[752,869,1950,1975]" class="Insecta" family="Empididae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="similis">
|
||
<emphasis id="DEF215090F571336FD6A79FEFCA3F84A" box="[752,869,1950,1975]" italics="true" pageId="5" pageNumber="34">M. similis</emphasis>
|
||
</taxonomicName>
|
||
, described by
|
||
<bibRefCitation id="8817B4EA0F571336FB8C79FEFAB3F845" author="Grimaldi" box="[1046,1397,1950,1976]" pageId="5" pageNumber="40" refString="Grimaldi, D. A. & Cumming, J. (1999) Brachyceran Diptera in Cretaceous ambers and Mesozoic diversification of the Eremoneura. Bulletin of the American Museum of Natural History, 239, 124 pp." type="book" year="1999">Grimaldi & Cumming (1999)</bibRefCitation>
|
||
on the basis of a female specimen from
|
||
<collectingCountry id="9491898B0F571336FDC879A6FD7FF81D" box="[594,697,1990,2016]" name="Lebanon" pageId="5" pageNumber="34">Lebanon</collectingCountry>
|
||
Amber, mainly in having a different morphology of the basal flagellomere and a distal one that is very short in comparison, a shorter proboscis, a costal vein ending at apex of M1, a complete Sc, an R1 ending in a basal position respect to m-cu crossvein, an m-cu vein inclined towards wing apex, a dm-cu 3.5 times longer than m-cu, and an abdomen elongated with distinct terminalia.
|
||
</paragraph>
|
||
<paragraph id="EC39C91B0F541335FF5C7F73FD39FE3F" blockId="6.[151,1436,152,919]" pageId="6" pageNumber="35">
|
||
<taxonomicName id="2B86B2980F541335FF5C7F73FE09FED1" box="[198,463,275,300]" class="Insecta" family="Empididae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="oculeus">
|
||
<emphasis id="DEF215090F541335FF5C7F73FE09FED1" box="[198,463,275,300]" italics="true" pageId="6" pageNumber="35">Microphorites oculeus</emphasis>
|
||
</taxonomicName>
|
||
was described on the basis of a male specimen from
|
||
<collectingCountry id="9491898B0F541335FBDF7F72FB6DFED1" box="[1093,1195,274,300]" name="Lebanon" pageId="6" pageNumber="35">Lebanon</collectingCountry>
|
||
Amber by
|
||
<bibRefCitation id="8817B4EA0F541335FAA87F72FE4EFEAF" author="Grimaldi" pageId="6" pageNumber="40" refString="Grimaldi, D. A. & Cumming, J. (1999) Brachyceran Diptera in Cretaceous ambers and Mesozoic diversification of the Eremoneura. Bulletin of the American Museum of Natural History, 239, 124 pp." type="book" year="1999">Grimaldi & Cumming (1999)</bibRefCitation>
|
||
. This species is easily distinguished from all other known
|
||
<taxonomicName id="2B86B2980F541335FBD37F58FB29FEAC" box="[1097,1263,312,337]" class="Insecta" family="Empididae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="35" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DEF215090F541335FBD37F58FB29FEAC" box="[1097,1263,312,337]" italics="true" pageId="6" pageNumber="35">Microphorites</emphasis>
|
||
</taxonomicName>
|
||
, including the new species, by its longer and very abruptly tapered basal flagellomere with a strongly bulbous base and eyes occupying most of the head. Also differs from the new species in having clearly longer articles of the arista, a complete Sc and a costal vein ending at apex of R4+5.
|
||
</paragraph>
|
||
<paragraph id="EC39C91B0F541335FF5F7FB0FA55FC6A" blockId="6.[151,1436,152,919]" pageId="6" pageNumber="35">
|
||
Several male and female specimens of
|
||
<taxonomicName id="2B86B2980F541335FD547FB0FC42FE11" box="[718,900,464,492]" class="Insecta" family="Empididae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="35" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DEF215090F541335FD547FB0FC42FE11" box="[718,900,464,492]" italics="true" pageId="6" pageNumber="35">Microphorites</emphasis>
|
||
</taxonomicName>
|
||
are known from Álava Amber probably representing at least four different forms not yet described. Thus an important goal for future researches will be to elucidate if the species of San Just Amber is already know in the Álava deposit.
|
||
<bibRefCitation id="8817B4EA0F541335FF0D7C30FEA9FD91" box="[151,367,592,620]" pageId="6" pageNumber="40" refString="Nel, A., Perrichot, V., Daugeron, Ch. & Neraudeau, D. (2004) A new Microphorites in the Lower Cretaceous amber of the Southwest of France (Diptera: Dolichopodidae, " Microphorinae "). Annales de la Societe Entomologique de France (n. s.), 40 (1), 23 - 29." type="journal article">
|
||
Nel
|
||
<emphasis id="DEF215090F541335FF547C30FECEFD91" box="[206,264,592,620]" italics="true" pageId="6" pageNumber="35">et al</emphasis>
|
||
. (2004)
|
||
</bibRefCitation>
|
||
provided an extensive discussion about the relationships of the family
|
||
<taxonomicName id="2B86B2980F541335FA907C30FF1AFD6A" class="Insecta" family="Dolichopodidae" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="35" phylum="Arthropoda" rank="family">Dolichopodidae</taxonomicName>
|
||
and the ¨
|
||
<taxonomicName id="2B86B2980F541335FEC37C1BFDDFFD6A" box="[345,537,635,663]" class="Insecta" family="Dolichopodidae" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="35" phylum="Arthropoda" rank="subFamily" subFamily="Microphorinae">Microphorinae</taxonomicName>
|
||
¨ and ¨
|
||
<taxonomicName id="2B86B2980F541335FDF57C1BFCF9FD6A" box="[623,831,635,663]" class="Insecta" family="Dolichopodidae" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="35" phylum="Arthropoda" rank="subFamily" subFamily="Parathalassiinae">Parathalassiinae</taxonomicName>
|
||
¨ groups, indicating the relevance of the genus
|
||
<taxonomicName id="2B86B2980F541335FF0D7CC6FE8BFD3F" box="[151,333,678,706]" class="Insecta" family="Empididae" genus="Microphorites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="35" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="DEF215090F541335FF0D7CC6FE8BFD3F" box="[151,333,678,706]" italics="true" pageId="6" pageNumber="35">Microphorites</emphasis>
|
||
</taxonomicName>
|
||
for the cladistic analysis. Recently, an extensive cladistic analysis of the Empidoidea and basal lineages of the Cyclorrhapha based on morphological characters has been published by
|
||
<bibRefCitation id="8817B4EA0F541335FF0D7C9BFDC9FCEA" author="Sinclair" box="[151,527,763,791]" pageId="6" pageNumber="40" refString="Sinclair, B. J. & Cumming, J. M. (2006) The morphology, higher-level phylogeny and classification of the Empidoidea (Diptera). Zootaxa, 1180, 1 - 172." type="journal article" year="2006">Sinclair & Cumming (2006)</bibRefCitation>
|
||
. That analysis has confirmed the monophyly of both groups and the authors included into Empidoidea the five families
|
||
<taxonomicName id="2B86B2980F541335FCA27D45FC00FCBC" box="[824,966,805,833]" class="Insecta" family="Empididae" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="35" phylum="Arthropoda" rank="family">Empididae</taxonomicName>
|
||
,
|
||
<taxonomicName id="2B86B2980F541335FC4C7D45FB99FCBC" box="[982,1119,805,833]" class="Insecta" family="Hybotidae" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="35" phylum="Arthropoda" rank="family">Hybotidae</taxonomicName>
|
||
,
|
||
<taxonomicName id="2B86B2980F541335FBF77D45FAC6FCBC" box="[1133,1280,805,833]" class="Insecta" family="Atelestidae" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="35" phylum="Arthropoda" rank="family">Atelestidae</taxonomicName>
|
||
,
|
||
<taxonomicName id="2B86B2980F541335FA8A7D45FF3AFC91" class="Insecta" family="Brachystomatidae" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="35" phylum="Arthropoda" rank="family">Brachystomatidae</taxonomicName>
|
||
and
|
||
<taxonomicName id="2B86B2980F541335FEDD7D30FDD5FC91" box="[327,531,848,876]" class="Insecta" family="Dolichopodidae" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="35" phylum="Arthropoda" rank="family">Dolichopodidae</taxonomicName>
|
||
<emphasis id="DEF215090F541335FDBB7D30FD91FC91" box="[545,599,848,876]" italics="true" pageId="6" pageNumber="35">s.lat</emphasis>
|
||
. Last, the family
|
||
<taxonomicName id="2B86B2980F541335FCC87D30FBD6FC91" box="[850,1040,848,876]" class="Insecta" family="Microphoridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="35" phylum="Arthropoda" rank="family">Microphoridae</taxonomicName>
|
||
was not recognized, and the
|
||
<taxonomicName id="2B86B2980F541335FF0D7D1BFE93FC6A" box="[151,341,891,919]" class="Insecta" family="Dolichopodidae" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="35" phylum="Arthropoda" rank="subFamily" subFamily="Microphorinae">Microphorinae</taxonomicName>
|
||
and
|
||
<taxonomicName id="2B86B2980F541335FE097D1BFDA7FC6A" box="[403,609,891,919]" class="Insecta" family="Dolichopodidae" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="35" phylum="Arthropoda" rank="subFamily" subFamily="Parathalassiinae">Parathalassiinae</taxonomicName>
|
||
were assigned to the
|
||
<taxonomicName id="2B86B2980F541335FCEF7D1BFB87FC6A" box="[885,1089,891,919]" class="Insecta" family="Dolichopodidae" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="35" phylum="Arthropoda" rank="family">Dolichopodidae</taxonomicName>
|
||
<emphasis id="DEF215090F541335FBD07D1BFB46FC6A" box="[1098,1152,891,919]" italics="true" pageId="6" pageNumber="35">s.lat</emphasis>
|
||
. by the authors cited.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |