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<mods:title id="9D244C291EC6774995F8D5B8A0D9478F">The freshwater crabs of Macau, with the description of a new species of Nanhaipotamon Bott, 1968 and the redescription of Nanhaipotamonwupingense Cheng, Yang, Zhong &amp; Li, 2003 (Crustacea, Decapoda, Potamidae)</mods:title>
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<mods:namePart id="4FB0057D8507CAFBD7482ACD1DFD59DD">Huang, Chao</mods:namePart>
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<mods:namePart id="D1A8815A22B176CA6FEB5CBEB484FD49">Wong, Kai Chin</mods:namePart>
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<mods:namePart id="CCE40A3196726F4CE78C327FF79453E9">Ahyong, Shane T.</mods:namePart>
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<treatment id="23BECC077943CC8DD88514A15523660B" ID-GBIF-Taxon="154125118" LSID="urn:lsid:plazi:treatment:23BECC077943CC8DD88514A15523660B" httpUri="http://treatment.plazi.org/id/23BECC077943CC8DD88514A15523660B" lastPageId="16" lastPageNumber="107" pageId="14" pageNumber="105">
<subSubSection id="D3B4A847CB950E9EFCF85CD6515761B3" pageId="14" pageNumber="105" type="nomenclature">
<paragraph id="96C18445D98B4B7FE77FC2C69263CC66" pageId="14" pageNumber="105">
<taxonomicName id="C3F0A17903A4B39E5726989FA2ECDC09" ID-CoL="4Y8F8" authority="Ng &amp; Dudgeon, 1992" authorityName="Ng &amp; Dudgeon" authorityYear="1992" class="Malacostraca" family="Gecarcinucidae" genus="Somanniathelphusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Somanniathelphusa zanklon" order="Decapoda" pageId="14" pageNumber="105" phylum="Arthropoda" rank="species" species="zanklon">Somanniathelphusa zanklon Ng &amp; Dudgeon, 1992</taxonomicName>
Figs 2D, 9
</paragraph>
</subSubSection>
<subSubSection id="B8A1BED4E053DF744E2EB3770AB1695A" pageId="14" pageNumber="105" type="reference_group">
<paragraph id="7C6287CA6B5AD54056724D50FAE495B2" pageId="14" pageNumber="105">
<taxonomicName id="907E1DA6BA278A51CEDFA6B5AA86AFD7" class="Malacostraca" family="Gecarcinucidae" genus="Somanniathelphusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Somanniathelphusa zanklon" order="Decapoda" pageId="14" pageNumber="105" phylum="Arthropoda" rank="species" species="zanklon">Somanniathelphusa zanklon</taxonomicName>
Ng &amp; Dudgeon, 1992: figs 11-13.
</paragraph>
<paragraph id="66F5F71AD5094ECC0CFD86BA8DB444FF" pageId="14" pageNumber="105">
<taxonomicName id="9300A5E17F5074C5DED275333DF5B306" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parathelphusa sinensis" order="Decapoda" pageId="14" pageNumber="105" phylum="Arthropoda" rank="species" species="sinensis">Parathelphusa sinensis</taxonomicName>
:
<bibRefCitation id="11A12CB20102395E4622CBBEA491EBE1" author="Doflein, F" journalOrPublisher="Raffles Bulletin of Zoology" pageId="17" pageNumber="108" url="https://doi.org/10.1163/9789004190832_021" year="1902">Doflein 1902</bibRefCitation>
: 662.
</paragraph>
<paragraph id="5F329013CC3E58D223097A5C7BECBDD1" pageId="14" pageNumber="105">
<taxonomicName id="4A17A6CACF3B019F455716E24AEEBBD0" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parathelphusa (Parathelphusa) sinensis" order="Decapoda" pageId="14" pageNumber="105" phylum="Arthropoda" rank="species" species="sinensis" subGenus="Parathelphusa">Parathelphusa (Parathelphusa) sinensis</taxonomicName>
:
<bibRefCitation id="8C9B6CDA1C1FD5BAD8E430A164EC77DD" author="Gee, NG" journalOrPublisher="Lignan Agricultural Review" pageId="17" pageNumber="108" pagination="156 - 166" title="Tentative list of Chinese decapod Crustacea, including those represented in the collections of the United States National Museum (marked with an *) with localities at which collected." volume="3" year="1925">Gee 1925</bibRefCitation>
: 159; Wu 1934: 339.
</paragraph>
<paragraph id="11EB66BCF0D1A5905A0E7C11D657CDA4" pageId="14" pageNumber="105">
<taxonomicName id="76D690E759E443E849909DE583250F41" class="Malacostraca" family="Gecarcinucidae" genus="Somanniathelphusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Somanniathelphusa sinensis" order="Decapoda" pageId="14" pageNumber="105" phylum="Arthropoda" rank="subspecies" species="sinensis" subspecies="sinensis">Somanniathelphusa sinensis sinensis</taxonomicName>
:
<bibRefCitation id="89C5DBE24E13D5C634873AFAC519F42F" author="Bott, R" journalOrPublisher="Senckenbergiana biologica" pageId="17" pageNumber="108" pagination="403 - 422" title="Parathelphusiden aus Hinterindien (Crustacea, Decapoda, Parathelphusidae)." volume="49" year="1968 b">Bott 1968b</bibRefCitation>
: 409, figs 11, 12, 30;
<bibRefCitation id="4AE295FC5670739E160D1FC0F026154B" author="Bott, R" journalOrPublisher="Revue Suisse (Zoologie)" pageId="17" pageNumber="108" pagination="327 - 344" title="Betrachtungen iiber entwicklungsgeschichte der Siisswasserkrabben nach der Sammlung des Naturhistorischen Museums in Genf / Schweiz." url="https://doi.org/10.5962/bhl.part.75900" volume="77" year="1970 a">Bott 1970a</bibRefCitation>
: 338;
</paragraph>
<paragraph id="1B0BCB02612AADC1B51CD336FC7CAC0A" pageId="14" pageNumber="105">
<bibRefCitation id="8ED164C0460FEA97D3C7E991F7F6647F" author="Bott, R" journalOrPublisher="Abhandlungen Senckenbergischen Naturforschenden Gesellschaft" pageId="17" pageNumber="108" pagination="1 - 338" title="Dis Siisswasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Eine Revision der Potamoidea und Parathelphusoidea (Crustacea, Decapoda)." volume="526" year="1970 b">Bott 1970b</bibRefCitation>
: 111, pl. 20, figs 42-44; Ng, 1988: 105.
</paragraph>
<paragraph id="88156463359B62F62DEF9210F210980F" pageId="14" pageNumber="105">
<taxonomicName id="C51B3DB365C965253B2851A2EF383C85" class="Malacostraca" family="Gecarcinucidae" genus="Somanniathelphusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Somanniathelphusa sinensis" order="Decapoda" pageId="14" pageNumber="105" phylum="Arthropoda" rank="species" species="sinensis">Somanniathelphusa sinensis</taxonomicName>
:
<bibRefCitation id="D2377E83BBFC4BC679BF990E20AD7AB8" pageId="14" pageNumber="105">Dai 1999</bibRefCitation>
: 67, fig. 29, pl. 2.
</paragraph>
</subSubSection>
<subSubSection id="E55111B76180613689E028BE9084D64A" pageId="14" pageNumber="105" type="materials_examined">
<paragraph id="2B575AF23BC77C7203921251B8E2F8EB" pageId="14" pageNumber="105">Material examined.</paragraph>
<paragraph id="ECA5BBBA48D4745F5C5DDAA096D25389" pageId="14" pageNumber="105">
SYSBM 101001, 1 male (27.2
<normalizedToken id="4E56EFC2270D0B2C575C5A52D497BDDD" originalValue="×">x</normalizedToken>
23.1 mm), Nanping (
<geoCoordinate id="EC28317A135E8655C7726A36EDB13904" direction="north" orientation="latitude" precision="555" value="22.19">22.19N</geoCoordinate>
,
<geoCoordinate id="663FA81A8614EBCA7CF8B70AD8758964" direction="east" orientation="longitude" precision="5555" value="113.5">113.5E</geoCoordinate>
), Zhuhai City, Guangdong, reservoir, coll. C. Huang, April 2015. SYSBM 101002-101003, 2 males (38.8
<normalizedToken id="A8F026925BF82A5913D5437E26DB284A" originalValue="×">x</normalizedToken>
31.9 mm, 30.4
<normalizedToken id="65903000EE3E7A8060916C6A9981F1DD" originalValue="×">x</normalizedToken>
25.6 mm), Jinding (
<geoCoordinate id="0392B457A3EF9BA8B2CEB0924B10D80A" direction="north" orientation="latitude" precision="555" value="22.38">22.38N</geoCoordinate>
,
<geoCoordinate id="2F507517CC476B70EF11A8D1FB913E77" direction="east" orientation="longitude" precision="555" value="113.54">113.54E</geoCoordinate>
), Zhuhai City, Guangdong, coll. local, May 2014. SYSBM 101004-101005, 2 males (42.0
<normalizedToken id="F7ABCC3389D46F8B98C215F42EF73881" originalValue="×">x</normalizedToken>
33.5 mm, 34.2
<normalizedToken id="424B08588C4113172A84B3485717F466" originalValue="×">x</normalizedToken>
28.1 mm), Sun Yat-sen University (
<geoCoordinate id="48B4C4F67DA7B47BA7CDE7A704379907" direction="north" orientation="latitude" precision="555" value="23.1">23.10N</geoCoordinate>
,
<geoCoordinate id="942797B9F1F60875157EFA26DC9621F3" direction="east" orientation="longitude" precision="555" value="113.3">113.30E</geoCoordinate>
), Guangzhou City, Guangdong, fish pond, coll. C. Huang, June 2013. SYSBM 101006, 1 female (37.1
<normalizedToken id="EEA9A5C2A551E94B7215E85D0EF4627F" originalValue="×">x</normalizedToken>
29.5 mm), same data as above. SYSBM 101007, 1 male (30.6
<normalizedToken id="519435626EAB941DFBBC829036730247" originalValue="×">x</normalizedToken>
24.5 mm), Coloane (
<geoCoordinate id="005DCECA59DB3A28E3FEC7812473172D" direction="north" orientation="latitude" precision="555" value="22.12">22.12N</geoCoordinate>
,
<geoCoordinate id="A46071CFEC4677A346F3F88A154DC80A" direction="east" orientation="longitude" precision="555" value="113.56">113.56E</geoCoordinate>
), Macau, reservoir, coll. K.C. Wong, July 2008. SYSBM 101008, 1 male (28.2
<normalizedToken id="F283B5ADB5532DA5557A8A6CBA2613FE" originalValue="×">x</normalizedToken>
22.5 mm), Coloane, Macau, reservoir, coll. K.C. Wong, July 2009. IACM, 1 male (24.2
<normalizedToken id="97574557B6B9E21A2CB0CCC6E5D7C5B2" originalValue="×">x</normalizedToken>
19.8 mm), Coloane, Macau, reservoir, coll. K.C. Wong, February 2013. SYSBM 101009-101010, 2 males (19.5
<normalizedToken id="0AC93C6056636D86A8D9436FE7931C54" originalValue="×">x</normalizedToken>
16.8 mm, 19.2
<normalizedToken id="EABF799B87FFEA14467932ED1BE95B4C" originalValue="×">x</normalizedToken>
16.4 mm), Shenzhen City (
<geoCoordinate id="1B9378553F4072AE36A67EC5AF9DA7C4" direction="north" orientation="latitude" precision="5555" value="22.6">22.6N</geoCoordinate>
,
<geoCoordinate id="24A4918D9EDCD497DA534811F8CD41D1" direction="east" orientation="longitude" precision="5555" value="114.0">114.0E</geoCoordinate>
), Guangdong, coll. local, August 2015. SYSBM 101011, 1 female (19.6
<normalizedToken id="F88ADCDBD3935307475C908BF9D29834" originalValue="×">x</normalizedToken>
16.4 mm), same data as above. SYSBM 101015-101016, 2 males (35.7
<normalizedToken id="C690265420B3900D6238FACEB666ED91" originalValue="×">x</normalizedToken>
28.4 mm, 37.2
<normalizedToken id="0921DFD7B2D0DFD9626C84231CFCA03A" originalValue="×">x</normalizedToken>
29.9 mm), Sihui City (
<geoCoordinate id="287B972157707A4F64EE6BE3DEDC2D06" direction="north" orientation="latitude" precision="555" value="23.12">23.12N</geoCoordinate>
,
<geoCoordinate id="8E12981327EB35DEE9575F1996C15A9B" direction="east" orientation="longitude" precision="555" value="113.56">113.56E</geoCoordinate>
), Guangdong, coll. local, August 2013. SYSBM 101017, 1 female (31.6
<normalizedToken id="F511D98082318C674A6DAEDDDF9455BA" originalValue="×">x</normalizedToken>
25.9 mm), same data as above. SYSBM 101018-101020, 3 males (38.2
<normalizedToken id="DC6E8138010D78E219272D86D9DD66E8" originalValue="×">x</normalizedToken>
30.8 mm, 35.4
<normalizedToken id="011740F7A8DC21451CD565E034DEECCF" originalValue="×">x</normalizedToken>
28.1 mm, 28.3
<normalizedToken id="A819806FEA6202E15DEB5EF9D2C4E11A" originalValue="×">x</normalizedToken>
21.3 mm), Renhua, Shaoguan City, Guangdong, coll. local, August 2013. SYSBM 101021, 1 male (31.5
<normalizedToken id="0A298B00C21434616ACB24584288263E" originalValue="×">x</normalizedToken>
26.6 mm), Lianhua Mountain, Shanwei City, Guangdong, coll. local, October 2013. SYSBM 101022, 1 female (27.3
<normalizedToken id="0E83A532DBF0DCF629B9CB0614A0EFED" originalValue="×">x</normalizedToken>
23.2 mm), same data as above. SYSBM 101030, 1 male (39.4
<normalizedToken id="CDFAA58FF09B6B3D8B49FA5AC31ABE8B" originalValue="×">x</normalizedToken>
33.0 mm), Heyuan City, Guangdong, coll. Z.C. Zhou, January 2014. SYSBM 101031, 1 female (32.4
<normalizedToken id="4605C09CDDA79C7FF7417651075E6177" originalValue="×">x</normalizedToken>
27.3 mm), same data as above. SYSBM 101032-101035, 4 males (35.0
<normalizedToken id="B6CAC3D6082E6C3DF003E4925E50521E" originalValue="×">x</normalizedToken>
28.3 mm, 32.7
<normalizedToken id="B04E2572B5213875CAB0A56C887F41A7" originalValue="×">x</normalizedToken>
26.9 mm, 30.8
<normalizedToken id="E9B8E20A387013EE0E7FBFCCF2F5CD78" originalValue="×">x</normalizedToken>
25.0 mm, 25.9
<normalizedToken id="8198C0DBAB18DB76B463678FD842CF17" originalValue="×">x</normalizedToken>
21.7 mm), Raoping, Chaozhou City, Guangdong, coll. Z.C. Zhou, January 2014. SYSBM 101036, 1 female (24.6
<normalizedToken id="345B37655025D4E99D5CA485CCC5E420" originalValue="×">x</normalizedToken>
20.8 mm), same data as above. SYSBM 101037-101040, 4 males (26.3
<normalizedToken id="A1CE750A835B9BB893969305D5F5B692" originalValue="×">x</normalizedToken>
20.4 mm, 30.9
<normalizedToken id="0BD78C53A762EA7B7EDB2172C559D975" originalValue="×">x</normalizedToken>
25.8 mm, 26.6
<normalizedToken id="98A6120F330206B04DA2858CBD4BF504" originalValue="×">x</normalizedToken>
23.0 mm, 27.0
<normalizedToken id="C0B21E0CBF7730117418028239E79A07" originalValue="×">x</normalizedToken>
22.5 mm), Wenzhou City, Zhejiang, coll. local, October 2013. SYSBM 101041, 1 female (29.8
<normalizedToken id="BDDD8178D475C18C239DAB8E84E05441" originalValue="×">x</normalizedToken>
24.6 mm), same data as above.
</paragraph>
</subSubSection>
<subSubSection id="E50C901713BE5788B1525B105817CFBF" pageId="14" pageNumber="105" type="colour in life">
<paragraph id="D6995F63E8748314C2D20B77EBC42245" pageId="14" pageNumber="105">Colour in life.</paragraph>
<paragraph id="7A76CA4483BDB65E96414B4AB2EA62C7" pageId="14" pageNumber="105">Generally brown overall; larger individuals may have dark markings near the cardiac region (Fig. 2D).</paragraph>
</subSubSection>
<subSubSection id="C3FE68A7CF5A3232C95C5D3145798615" pageId="15" pageNumber="106" type="distribution">
<paragraph id="3AE681DC95E25975748DB6DA48360933" pageId="15" pageNumber="106">
<pageBreakToken id="E47AF23C0438BD1763FB2B0221BEDE56" pageId="15" pageNumber="106" start="start">Distribution</pageBreakToken>
.
</paragraph>
<paragraph id="2DCBFCDCC7454150FC11EE141E2AC9B0" pageId="15" pageNumber="106">Coloane, Macau; Guangdong: Guangzhou City, Shenzhen City, Zhuhai City, Sihui City, Shaoguan City, Shanwei City, Heyuan City, Chaozhou City; Zhejiang: Wenzhou City.</paragraph>
</subSubSection>
<subSubSection id="FBCB36593AD959408E7D42621675CB11" lastPageId="16" lastPageNumber="107" pageId="15" pageNumber="106" type="remarks">
<paragraph id="4668F5B45EC995CF249BD8E7F08B7159" pageId="15" pageNumber="106">Remarks.</paragraph>
<paragraph id="E394F846873904F6045C33A68D1834C7" lastPageId="16" lastPageNumber="107" pageId="15" pageNumber="106">
<bibRefCitation id="674025C3FD241E4A749A9697F7FC350E" author="Ng, PKL" journalOrPublisher="Invertebrate Systematics" pageId="18" pageNumber="109" pagination="741 - 768" title="The Potmidae and Parathelphusidae (Crustacea: Decapoda: Brachyura) of Hong Kong." url="https://doi.org/10.1071/IT9920741" volume="6" year="1992">Ng and Dudgeon (1992)</bibRefCitation>
showed former records of
<taxonomicName id="3D4DA157040C39E785E5542EF084F84C" class="Malacostraca" family="Gecarcinucidae" genus="Somanniathelphusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Somanniathelphusa sinensis" order="Decapoda" pageId="15" pageNumber="106" phylum="Arthropoda" rank="species" species="sinensis">Somanniathelphusa sinensis</taxonomicName>
(H. Milne Edwards, 1853) from southern China (Bott 1968, 1970;
<bibRefCitation id="77472D830D96E982E0C41D981FAA260F" pageId="15" pageNumber="106">Dai 1999</bibRefCitation>
) to represent a new species,
<taxonomicName id="09FCDD44DDB091BDEF4C624F7DDCFFCE" lsidName="S. zanklon." pageId="15" pageNumber="106" rank="species" species="zanklon.">S. zanklon.</taxonomicName>
<bibRefCitation id="675F52C05758D1DE38ACE06203A95D7B" author="Shih, HT" journalOrPublisher="Invertebrate Systematics" pageId="18" pageNumber="109" pagination="29 - 37" title="Phylogeny of the freshwater crab genus Somanniathelphusa Bott, (Decapoda: Parathelphusidae) from Taiwan and the coastal regions of China, with notes on their biogeography." url="https://doi.org/10.1071/is06007" volume="21" year="2007">Shih et al. (2007)</bibRefCitation>
included
<taxonomicName id="F5A935AD8F55BFD654EFECD2102C2C06" lsidName="S. zanklon" pageId="15" pageNumber="106" rank="species" species="zanklon">S. zanklon</taxonomicName>
from Hong Kong and Guangdong (Dongguan City and Nanhai City) and showed that specimens
<pageBreakToken id="174EA97C60E5CC3610E3E8083AD136F2" pageId="16" pageNumber="107" start="start">of</pageBreakToken>
<taxonomicName id="93C2DD4E76974809E0CC9B5C09DACB0D" class="Malacostraca" family="Gecarcinucidae" genus="Somanniathelphusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Somanniathelphusa" order="Decapoda" pageId="16" pageNumber="107" phylum="Arthropoda" rank="genus">Somanniathelphusa</taxonomicName>
from eastern Guangdong, Fujian and west-central Taiwan were all very closely related genetically, probably even conspecific. Specimens examined here from different localities were all very similar morphologically (Fig. 9), with those from Zhejiang Province having a slightly longer G1 distal part (Fig. 9H), though the overall shape is very much like the others. We tentatively treat all of these as the same species pending a full revision. Interestingly,
<bibRefCitation id="4CC0A4167A1BF1A9E48AA0E3D15D745A" pageId="16" pageNumber="107">Dai (1999)</bibRefCitation>
also reported the genus from Zhejiang, but none of the species of
<taxonomicName id="9A69DA59C9B906945FAD509E5DA4442E" class="Malacostraca" family="Gecarcinucidae" genus="Somanniathelphusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Somanniathelphusa" order="Decapoda" pageId="16" pageNumber="107" phylum="Arthropoda" rank="genus">Somanniathelphusa</taxonomicName>
in her monograph lists Zhejiang among its localities. The Chinese
<taxonomicName id="8CEC0528E99802598E6F3BD605B021F0" class="Malacostraca" family="Gecarcinucidae" genus="Somanniathelphusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Somanniathelphusa" order="Decapoda" pageId="16" pageNumber="107" phylum="Arthropoda" rank="genus">Somanniathelphusa</taxonomicName>
are particularly problematic as many species look identical externally and were described based on minute differences that we find difficult to detect. Furthermore, being lowland species, they are readily dispersed by floods and are commonly found in aquaculture ponds where their newly hatched crablings are easily translocated. Preliminary genetic evidence suggests that the species diversity of
<taxonomicName id="D4CF21D17B8D9B72823FF07F1F8A2843" class="Malacostraca" family="Gecarcinucidae" genus="Somanniathelphusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Somanniathelphusa" order="Decapoda" pageId="16" pageNumber="107" phylum="Arthropoda" rank="genus">Somanniathelphusa</taxonomicName>
may have been overestimated (
<bibRefCitation id="A381275FA4FD600C00367271E87750B7" author="Shih, HT" journalOrPublisher="Invertebrate Systematics" pageId="18" pageNumber="109" pagination="29 - 37" title="Phylogeny of the freshwater crab genus Somanniathelphusa Bott, (Decapoda: Parathelphusidae) from Taiwan and the coastal regions of China, with notes on their biogeography." url="https://doi.org/10.1071/is06007" volume="21" year="2007">Shih et al. 2007</bibRefCitation>
).
</paragraph>
<caption id="5942B840398F475E4C9F65B3DC2B607B" pageId="16" pageNumber="107">
<paragraph id="58A88F811B9A84FC82053F8DA814B2DC" pageId="16" pageNumber="107">
Figure 9. Comparison of G1 of
<taxonomicName id="CF2B36C37B5B0F2D6868258DD80FA91E" class="Malacostraca" family="Gecarcinucidae" genus="Somanniathelphusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Somanniathelphusa zanklon" order="Decapoda" pageId="16" pageNumber="107" phylum="Arthropoda" rank="species" species="zanklon">Somanniathelphusa zanklon</taxonomicName>
Ng &amp; Dudgeon, 1992, from different localities. Male (30.6
<normalizedToken id="14E469DF5916FBCC768B21D9546D1483" originalValue="×">x</normalizedToken>
24.5 mm), SYSBM 101007, Coloane, Macau (A); male (38.8
<normalizedToken id="94B9ED2F6604B6085C73C9EC3DEB7783" originalValue="×">x</normalizedToken>
31.9 mm), SYSBM 101002, Jinding, Zhuhai City, Guangdong (B); male (42.0
<normalizedToken id="30E2EBD92DCE32344BC985B6512FC69B" originalValue="×">x</normalizedToken>
33.5 mm), SYSBM 101004, Sun Yat-sen University, Guangzhou City, Guangdong (C); male (35.7
<normalizedToken id="5BB9F5FFF07C917AFE0B402E718FCF4B" originalValue="×">x</normalizedToken>
28.4 mm), SYSBM 101015, Sihui City, Guangdong (D); male (28.3
<normalizedToken id="C5B020C12F4FA613BB56D77B419EEE98" originalValue="×">x</normalizedToken>
21.3 mm), SYSBM 101020, Renhua, Shaoguan City, Guangdong (E); male (31.5
<normalizedToken id="A67DF88F95B71F6AFA7132BEB6A0ED3E" originalValue="×">x</normalizedToken>
26.6 mm), SYSBM 101021, Lianhua Mountain, Shanwei City, Guangdong (F); male (35.0
<normalizedToken id="F545FD9A4C7B4D975A4E9067A8A789B2" originalValue="×">x</normalizedToken>
28.3 mm), SYSBM 101032, Raoping, Chaozhou City, Guangdong (G); male (26.3
<normalizedToken id="34B05757AC1DF93039FD5A20F472CCFE" originalValue="×">x</normalizedToken>
20.4 mm), SYSBM 101037, Wenzhou City, Zhejiang (H). Scale bars: 1.0 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="836C855FBBDBD04F99DA7E6D2C383B80" pageId="16" pageNumber="107" type="conservation status">
<paragraph id="61157BE6462CF6FE54255605F940D353" pageId="16" pageNumber="107">Conservation status.</paragraph>
<paragraph id="50135403E4B6DCB8F8300DBEA742153A" pageId="16" pageNumber="107">
<taxonomicName id="396E94211ED6B60A72A8F1595317DE82" class="Malacostraca" family="Gecarcinucidae" genus="Somanniathelphusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Somanniathelphusa zanklon" order="Decapoda" pageId="16" pageNumber="107" phylum="Arthropoda" rank="species" species="zanklon">Somanniathelphusa zanklon</taxonomicName>
is currently assessed as Endangered (
<bibRefCitation id="9961056B154A0F1E22D43D96C0D35702" pageId="16" pageNumber="107">Esser and Cumberlidge 2008</bibRefCitation>
) as it was known from fewer than five locations in Hong Kong with a extant of occurrence less than 5,000 km2, with degrading habitat quality. Our study, however, finds this species to have a widespread occurrence in south-east China with an area of occupancy estimated at over 120,000 km2. Though habitat quality in some of these locations is declining, this resilient lowland species seems to be able to thrive in most water bodies that are not heavily polluted. As such, we find that
<taxonomicName id="2638BBF568172ABC1E85FFD39A06A6E4" lsidName="S. zanklon" pageId="16" pageNumber="107" rank="species" species="zanklon">S. zanklon</taxonomicName>
does not satisfy any IUCN Red List threat categories and thus we suggest Least Concern would be a more appropriate determination at present. In Macau, all known occurrences of
<taxonomicName id="281B86D82D7676C8C6FD9652763F4A4E" lsidName="S. zanklon" pageId="16" pageNumber="107" rank="species" species="zanklon">S. zanklon</taxonomicName>
are from Hac-Sa Reservoir, Coloane, although it most likely also occurs in other water bodies in Coloane. There was an unconfirmed sighting of
<taxonomicName id="A0D26B2AB8522291919001AC1D1E6164" lsidName="S. zanklon" pageId="16" pageNumber="107" rank="species" species="zanklon">S. zanklon</taxonomicName>
in Taipa a few years ago by the second author, though more recent surveys have failed to locate any specimens. There are three large water bodies on the Macau peninsula, of which only one reservoir on the east, next to Parque Municipal do Monte da Guia, which sources freshwater from the mainland, is suitable for lowland freshwater crabs. Although we did not survey this reservoir, it very likely also holds
<taxonomicName id="320D6EBF4A65A2B36BDAEE098F3BA99C" lsidName="S. zanklon" pageId="16" pageNumber="107" rank="species" species="zanklon">S. zanklon</taxonomicName>
as this species was found in one of its water source reservoirs in Zhuhai. The other two water bodies, Sai Van Lake and Nam Van Lake, were once bays that have mostly been artificially closed off by landfill and currently hold sea water.
</paragraph>
</subSubSection>
</treatment>
</document>