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<document id="1E65F1C854B0B5DCA1915D1C036D45F9" ID-CLB-Dataset="8610" ID-DOI="10.11646/zootaxa.4838.4.9" ID-GBIF-Dataset="d6d2d99d-db64-4025-a012-16c5e122f18b" ID-ISSN="1175-5326" ID-Zenodo-Dep="4405509" ID-ZooBank="FF8E8986-381B-4B08-A8D8-DA07F74496FD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609416766718" checkinUser="plazi" docAuthor="Ceríaco, Luis M. P., Marques, Mariana P. &amp; Bauer, Aaron M." docDate="2020" docId="8E7487A3EC6A9015FF36FD0EFB2F00FA" docLanguage="en" docName="zootaxa.4838.4.9.pdf" docOrigin="Zootaxa 4838 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Atheris squamigera" docType="treatment" docVersion="8" lastPageNumber="585" masterDocId="724DFFDBEC699011FFA1FFBDFFA8050E" masterDocTitle="The Bush Vipers, genus Atheris Cope, 1862 (Squamata: Viperidae) of Bioko Island Gulf of Guinea, with the description of a new species" masterLastPageNumber="593" masterPageNumber="581" pageNumber="584" updateTime="1698850304638" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="A4463E2E8F28067790572CB67719E01D">The Bush Vipers, genus Atheris Cope, 1862 (Squamata: Viperidae) of Bioko Island Gulf of Guinea, with the description of a new species</mods:title>
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<mods:namePart id="85DB3CB6EAC290163153CDA3C8E08C5A">Ceríaco, Luis M. P.</mods:namePart>
<mods:affiliation id="108D20E2462B38CC2D4E65719CCE69E7">Museu de História Natural e da Ciência da Universidade do Porto, Praça Gomes Teixeira, 4099 - 002 Porto, Portugal. &amp; Departamento de Zoologia e Antropologia (Museu Bocage), Museu Nacional de História Natural e da Ciência, Universidade de Lisboa, Rua da Escola Politécnica, 58, 1269 - 102 Lisboa, Portugal</mods:affiliation>
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<mods:namePart id="963FFDF6F30AE07BE422DCAA08FB9C59">Marques, Mariana P.</mods:namePart>
<mods:affiliation id="A71F0B06EC74F84E00E7FC72A01D2565">Centro de Investigação em Biodiversidade e Recursos Genéticos (CIBIO), Rede de Investigação em Biodiversidade e Biologia Evolutiva (InBIO), Universidade do Porto, Rua Padre Armando Quintas 7, 4485 - 661, Vairão, Portugal. &amp; Departamento de Zoologia e Antropologia (Museu Bocage), Museu Nacional de História Natural e da Ciência, Universidade de Lisboa, Rua da Escola Politécnica, 58, 1269 - 102 Lisboa, Portugal</mods:affiliation>
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<mods:namePart id="D78C7A4165B2222B29D8934E2D627FA5">Bauer, Aaron M.</mods:namePart>
<mods:affiliation id="AE78E93F6B14F9D075E8E314246215C3">Department of Biology and Center for Biodiversity and Ecosystem Stewardship, Villanova University, 800 Lancaster Avenue, Villanova, Pennsylvania 19085 - 1699, USA</mods:affiliation>
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<heading id="5D2A81D9EC6A9012FF36FD0EFDF707C3" bold="true" box="[151,607,691,718]" fontSize="11" level="1" pageId="3" pageNumber="584" reason="1">
<taxonomicName id="C1DD4D36EC6A9012FF36FD0EFDF707C3" authority="(Hallowell, 1855)" baseAuthorityName="Hallowell" baseAuthorityYear="1855" box="[151,607,691,718]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="584" phylum="Chordata" rank="species" species="squamigera">
<emphasis id="34A9EAA7EC6A9012FF36FD0EFDF707C3" bold="true" box="[151,607,691,718]" pageId="3" pageNumber="584">
<emphasis id="34A9EAA7EC6A9012FF36FD0EFE2D07C3" bold="true" box="[151,389,691,718]" italics="true" pageId="3" pageNumber="584">Atheris squamigera</emphasis>
(Hallowell, 1855)
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<taxonomicName id="C1DD4D36EC6A9012FF36FD45FD82061E" authority="Hallowell, 1855: 193" authorityName="Hallowell" authorityPageNumber="193" authorityYear="1855" box="[151,554,759,784]" class="Reptilia" family="Viperidae" genus="Echis" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="584" phylum="Chordata" rank="species" species="squamigera">
<emphasis id="34A9EAA7EC6A9012FF36FD45FEE60601" box="[151,334,760,784]" italics="true" pageId="3" pageNumber="584">Echis squamigera</emphasis>
Hallowell, 1855: 193
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<taxonomicName id="C1DD4D36EC6A9012FF36FCABFDEE0620" authority=": Boulenger (1905: 216)" authorityName=": Boulenger" authorityPageNumber="216" authorityYear="1905" box="[151,582,790,814]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="584" phylum="Chordata" rank="species" species="squamiger">
<emphasis id="34A9EAA7EC6A9012FF36FCABFEFA0620" box="[151,338,790,814]" italics="true" pageId="3" pageNumber="584">Atheris squamiger</emphasis>
: Boulenger (1905: 216)
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<treatmentCitationGroup id="26CD119BEC6A9012FF36FC88FCB20643" box="[151,794,821,845]" pageId="3" pageNumber="584">
<taxonomicName id="C1DD4D36EC6A9012FF36FC88FE750643" authority="squamigera" box="[151,477,821,845]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="584" phylum="Chordata" rank="subSpecies" species="squamigera" subSpecies="squamigera">
<emphasis id="34A9EAA7EC6A9012FF36FC88FE750643" box="[151,477,821,845]" italics="true" pageId="3" pageNumber="584">Atheris squamigera squamigera</emphasis>
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[part]:
<treatmentCitation id="877C10A4EC6A9012FD89FC88FCB20643" author="Capocaccia, L." box="[552,794,821,845]" page="304" pageId="3" pageNumber="584" year="1961">
<bibRefCitation id="624C4B44EC6A9012FD89FC88FCB20643" author="Capocaccia, L." box="[552,794,821,845]" pageId="3" pageNumber="584" pagination="285 - 309" refId="ref8868" refString="Capocaccia, L. (1961) Contributo allo studio dei serpenti delle isole del Golfo di Guinea. Annali del Museo Civico di Storia Naturale &quot; Giacomo Doria &quot;, 72, 285 - 309." type="journal article" year="1961">Capocaccia (1961: 304)</bibRefCitation>
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<treatmentCitationGroup id="26CD119BEC6A9012FF36FCE9FD080665" box="[151,672,851,876]" pageId="3" pageNumber="584">
<treatmentCitation id="877C10A4EC6A9012FF36FCE9FD080665" author="Mertens, R." box="[151,672,851,876]" page=")" pageId="3" pageNumber="584" year="1964">
<taxonomicName id="C1DD4D36EC6A9012FF36FCE9FD080665" authority="squamiger: Mertens (1964: 235)" authorityName="Mertens" authorityPageNumber="235" authorityYear="1964" box="[151,672,851,876]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="584" phylum="Chordata" rank="subSpecies" species="squamiger" subSpecies="squamiger">
<emphasis id="34A9EAA7EC6A9012FF36FCE9FE6B0665" box="[151,451,852,876]" italics="true" pageId="3" pageNumber="584">Atheris squamiger squamiger</emphasis>
:
<bibRefCitation id="624C4B44EC6A9012FE71FCE9FD080665" author="Mertens, R." box="[464,672,851,875]" pageId="3" pageNumber="584" pagination="211 - 238" refId="ref10030" refString="Mertens, R. (1964) Die Reptilien von Fernando Poo. Bonner Zoologische Beitrage, 15, 211 - 238." type="journal article" year="1964">Mertens (1964: 235)</bibRefCitation>
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<paragraph id="066236B5EC6A9012FF36FCCFFC100684" blockId="3.[151,952,759,906]" box="[151,952,882,906]" pageId="3" pageNumber="584">
<treatmentCitationGroup id="26CD119BEC6A9012FF36FCCFFC100684" box="[151,952,882,906]" pageId="3" pageNumber="584">
<treatmentCitation id="877C10A4EC6A9012FF36FCCFFDC20684" author="Wallach, V. &amp; Kenneth, I. W. &amp; Boundy, J." box="[151,618,882,906]" page=")" pageId="3" pageNumber="584" year="2014">
<taxonomicName id="C1DD4D36EC6A9012FF36FCCFFDC20684" authority=": Wallach et al. (2014: 64)" authorityName="Wallach" authorityPageNumber="64" authorityYear="2014" box="[151,618,882,906]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="584" phylum="Chordata" rank="species" species="squamigera">
<emphasis id="34A9EAA7EC6A9012FF36FCCFFEF70684" box="[151,351,882,906]" italics="true" pageId="3" pageNumber="584">Atheris squamigera</emphasis>
:
<bibRefCitation id="624C4B44EC6A9012FECAFCCFFDC20684" author="Wallach, V. &amp; Kenneth, I. W. &amp; Boundy, J." box="[363,618,882,906]" pageId="3" pageNumber="584" refId="ref10876" refString="Wallach, V., Kenneth, I. W. &amp; Boundy, J. (2014) Snakes of the World: A Catalogue of Living and Extinct Species. CRC Press, Boca Raton, Florida, xxvi + [2] + 1209 pp." type="book" year="2014">
Wallach
<emphasis id="34A9EAA7EC6A9012FE65FCCEFE5F0684" box="[452,503,882,906]" italics="true" pageId="3" pageNumber="584">et al.</emphasis>
(2014: 64)
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;
<treatmentCitation id="877C10A4EC6A9012FDD4FCCFFC100684" author="Chippaux, J. - P. &amp; Jackson, K." box="[629,952,882,906]" page="78" pageId="3" pageNumber="584" year="2019">
<bibRefCitation id="624C4B44EC6A9012FDD4FCCFFC100684" author="Chippaux, J. - P. &amp; Jackson, K." box="[629,952,882,906]" pageId="3" pageNumber="584" refId="ref9208" refString="Chippaux, J. - P. &amp; Jackson, K. (2019) Snakes of Central and Western Africa. Johns Hopkins University Press, Baltimore, Maryland, 448 pp." type="book" year="2019">Chippaux &amp; Jackson (2019: 78)</bibRefCitation>
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Boulenger (1905) was the first to denote the presence of the species in the island, as
<taxonomicName id="C1DD4D36EC6A9012FB26FC05FAF006DC" authorityName=": Boulenger" authorityYear="1905" box="[1159,1368,952,978]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="584" phylum="Chordata" rank="species" species="squamiger">
<emphasis id="34A9EAA7EC6A9012FB26FC05FAF006DC" box="[1159,1368,952,978]" italics="true" pageId="3" pageNumber="584">Atheris squamiger</emphasis>
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[sic], based on an unnumbered series of specimens collected by the Italian explorer Leonardo Fea (1852–1903) in Moka, southern Bioko Island. These specimens were later studied in more detail by
<bibRefCitation id="624C4B44EC6A9012FC56FBBDFB650115" author="Capocaccia, L." box="[1015,1229,1024,1051]" pageId="3" pageNumber="584" pagination="285 - 309" refId="ref8868" refString="Capocaccia, L. (1961) Contributo allo studio dei serpenti delle isole del Golfo di Guinea. Annali del Museo Civico di Storia Naturale &quot; Giacomo Doria &quot;, 72, 285 - 309." type="journal article" year="1961">Capocaccia (1961)</bibRefCitation>
, who reported one female and one juvenile of the species. For the female (MSNG/CE 30428a), the author provided a detailed description, which has 150 ventral scales, 52 subcaudals, 25 scales rows around the neck, 21 at midbody, and 15 before the cloaca, 10 supralabials and 10 infralabials, three scales between the nasals and eye on the right side of the head, two on the left, nine interoculars, 14 oculars on the right side, 17 on the left side, and only one row of scales between eye and supralabials. For the juvenile (MSNG/CE 30428b),
<bibRefCitation id="624C4B44EC6A9012FCA4FB09FC7E01C1" author="Capocaccia, L." box="[773,982,1204,1231]" pageId="3" pageNumber="584" pagination="285 - 309" refId="ref8868" refString="Capocaccia, L. (1961) Contributo allo studio dei serpenti delle isole del Golfo di Guinea. Annali del Museo Civico di Storia Naturale &quot; Giacomo Doria &quot;, 72, 285 - 309." type="journal article" year="1961">Capocaccia (1961)</bibRefCitation>
reported 150 ventrals, 53 subcaudals, 25 scales rows around the neck, 20 at midbody, and 17 before the cloaca, nine supralabials on the right, but 10 on the left side, 10 infralabials on the right side, nine on the left side, two scales between nasal and the eye, nine interoculars, 15 oculars on the right side, 14 on the left side, and only one row of scales between the eye and supralabials. Both these specimens were reviewed by us (this study).
</paragraph>
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<paragraph id="066236B5EC6A9012FF66FAD4FCB50371" blockId="3.[151,1437,952,2023]" pageId="3" pageNumber="584">
<bibRefCitation id="624C4B44EC6A9012FF66FAD4FEDD008D" author="Mertens, R." box="[199,373,1384,1411]" pageId="3" pageNumber="584" pagination="211 - 238" refId="ref10030" refString="Mertens, R. (1964) Die Reptilien von Fernando Poo. Bonner Zoologische Beitrage, 15, 211 - 238." type="journal article" year="1964">Mertens (1964)</bibRefCitation>
reported a total of seven poorly preserved specimens (see Bibliographic Records below), three males, three females and a juvenile female. All the characters presented by
<bibRefCitation id="624C4B44EC6A9012FC7FFA30FB2300A9" author="Mertens, R." box="[990,1163,1420,1447]" pageId="3" pageNumber="584" pagination="211 - 238" refId="ref10030" refString="Mertens, R. (1964) Die Reptilien von Fernando Poo. Bonner Zoologische Beitrage, 15, 211 - 238." type="journal article" year="1964">Mertens (1964)</bibRefCitation>
unambiguously identify these specimens as the same taxon as those cited by
<bibRefCitation id="624C4B44EC6A9012FD40FA0DFC1B00C5" author="Capocaccia, L." box="[737,947,1456,1483]" pageId="3" pageNumber="584" pagination="285 - 309" refId="ref8868" refString="Capocaccia, L. (1961) Contributo allo studio dei serpenti delle isole del Golfo di Guinea. Annali del Museo Civico di Storia Naturale &quot; Giacomo Doria &quot;, 72, 285 - 309." type="journal article" year="1961">Capocaccia (1961)</bibRefCitation>
and the material present in the CAS collections. According to
<bibRefCitation id="624C4B44EC6A9012FED5FA68FD8B00E1" author="Mertens, R." box="[372,547,1492,1519]" pageId="3" pageNumber="584" pagination="211 - 238" refId="ref10030" refString="Mertens, R. (1964) Die Reptilien von Fernando Poo. Bonner Zoologische Beitrage, 15, 211 - 238." type="journal article" year="1964">Mertens (1964)</bibRefCitation>
, three specimens had seven interoculars, three others had eight, and a single individual had nine; all the specimens had only single row of scales between the eyes and supralabials. In males there were 19–24 midbody scale rows, 148–153 ventrals and 57–60 subcaudals in males, 148–155 ventrals and 46–51 subcaudals in females, and 145 ventrals and 56 subcaudals in the juvenile female. According to
<bibRefCitation id="624C4B44EC6A9012FB4DF9FCFA340355" author="Mertens, R." box="[1260,1436,1600,1627]" pageId="3" pageNumber="584" pagination="211 - 238" refId="ref10030" refString="Mertens, R. (1964) Die Reptilien von Fernando Poo. Bonner Zoologische Beitrage, 15, 211 - 238." type="journal article" year="1964">Mertens (1964)</bibRefCitation>
these specimens did not have dark streaks behind the eye.
</paragraph>
<paragraph id="066236B5EC6A9012FF66F935FB9202E9" blockId="3.[151,1437,952,2023]" pageId="3" pageNumber="584">
The recent material comprises two adult males (CAS 201867, CAS 201869) and two adult females (CAS 201866, CAS 201868 [
<figureCitation id="9EE62A30EC6A9012FE3AF911FE7603C9" box="[411,478,1708,1735]" captionStart="FIGURE 1" captionStartId="4.[151,250,1139,1163]" captionTargetBox="[151,1436,181,1115]" captionTargetId="figure-29@4.[151,1436,181,1115]" captionTargetPageId="4" captionText="FIGURE 1. Specimen of Atheris squamigera (CAS 207868) from Bioko Island." figureDoi="http://doi.org/10.5281/zenodo.4405511" httpUri="https://zenodo.org/record/4405511/files/figure.png" pageId="3" pageNumber="584">Fig. 1</figureCitation>
]). The measurements, meristic and scalation data (
<tableCitation id="4B5F030EEC6A9012FBBFF911FBD903C9" box="[1054,1137,1708,1735]" captionStart="TABLE 1" captionStartId="2.[151,239,587,613]" captionText="TABLE 1. Morphometric and meristic data on Atheris squamigera and Atheris hetfieldi sp. nov. types series from Bioko Island. Measurements in centimeters. All abbreviations follow the Materials and Methods section. Left and right side presented as L/R." pageId="3" pageNumber="584">Table 1</tableCitation>
) of all of these specimens agree entirely with
<taxonomicName id="C1DD4D36EC6A9012FED4F96CFDB103E4" box="[373,537,1745,1771]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="584" phylum="Chordata" rank="species" species="squamigera">
<emphasis id="34A9EAA7EC6A9012FED4F96CFDB103E4" box="[373,537,1745,1771]" italics="true" pageId="3" pageNumber="584">A. squamigera</emphasis>
</taxonomicName>
, and all specimens have a small dark band extending posteriorly from the eye. This dark band is, however, not as large and conspicuous as that of
<taxonomicName id="C1DD4D36EC6A9012FC01F949FBCB0200" authorityName="Lawson" authorityYear="1999" box="[928,1123,1780,1806]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="584" phylum="Chordata" rank="species" species="broadleyi">
<emphasis id="34A9EAA7EC6A9012FC01F949FBCB0200" box="[928,1123,1780,1806]" italics="true" pageId="3" pageNumber="584">Atheris broadleyi</emphasis>
</taxonomicName>
, and is similar to the small bands typical of other
<taxonomicName id="C1DD4D36EC6A9012FE30F8A4FD9C023C" box="[401,564,1817,1843]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="584" phylum="Chordata" rank="species" species="squamigera">
<emphasis id="34A9EAA7EC6A9012FE30F8A4FD9C023C" box="[401,564,1817,1843]" italics="true" pageId="3" pageNumber="584">A. squamigera</emphasis>
</taxonomicName>
from the continent, including the
<typeStatus id="D9668817EC6A9012FC10F8A4FC49023D" box="[945,993,1817,1843]" pageId="3" pageNumber="584">type</typeStatus>
specimen of the species, still present in the collections of the Academy of Natural Sciences of Drexel University (ANSP 6949), in Philadelphia,
<collectingCountry id="7ECA7625EC6A9012FA90F880FACF0259" box="[1329,1383,1853,1879]" name="United States of America" pageId="3" pageNumber="584">USA</collectingCountry>
(see
<bibRefCitation id="624C4B44EC6A9012FF36F8DCFE9D0275" author="Lawson, D. P." box="[151,309,1889,1915]" pageId="3" pageNumber="584" pagination="793 - 803" refId="ref9681" refString="Lawson, D. P. (1999) A new species of arboreal viper (Serpentes: Viperidae: Atheris) from Cameroon, Africa. Proceedings of the Biological Society of Washington, 112 (4), 793 - 803." type="journal article" year="1999">Lawson 1999</bibRefCitation>
). In terms of coloration, all specimens have an olive-green background, with a series of transverse black and yellow bars from the neck region to the tail base. The tip of the tail is black. Ventrally, all specimens are yellowish green, with some ventrolateral yellow spots at the outer limits of the ventrals. The species is currently only known from the central and southern regions of the island, near Moka (
<figureCitation id="9EE62A30EC6A9012FC4FF871FB9902E9" box="[1006,1073,1996,2023]" captionStart="FIGURE 2" captionStartId="5.[151,250,1975,1999]" captionTargetBox="[221,1368,502,1941]" captionTargetId="figure-186@5.[215,1371,489,1951]" captionTargetPageId="5" captionText="FIGURE 2. Distribution map of the Atheris on Bioko Island. Orange circles: Atheris squamigera; Blue stars: Atheris hetfieldi sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4405513" httpUri="https://zenodo.org/record/4405513/files/figure.png" pageId="3" pageNumber="584">Fig. 2</figureCitation>
)
</paragraph>
</subSubSection>
<caption id="52A2663DEC6D9015FF36FBCEFC670185" ID-DOI="http://doi.org/10.5281/zenodo.4405511" ID-Zenodo-Dep="4405511" box="[151,975,1139,1164]" httpUri="https://zenodo.org/record/4405511/files/figure.png" pageId="4" pageNumber="585" startId="4.[151,250,1139,1163]" targetBox="[151,1436,181,1115]" targetPageId="4">
<paragraph id="066236B5EC6D9015FF36FBCEFC670185" blockId="4.[151,975,1139,1164]" box="[151,975,1139,1164]" pageId="4" pageNumber="585">
<emphasis id="34A9EAA7EC6D9015FF36FBCEFEB10185" bold="true" box="[151,281,1139,1163]" pageId="4" pageNumber="585">FIGURE 1.</emphasis>
Specimen of
<taxonomicName id="C1DD4D36EC6D9015FE04FBCEFDC50185" box="[421,621,1139,1163]" class="Reptilia" family="Viperidae" genus="Atheris" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="585" phylum="Chordata" rank="species" species="squamigera">
<emphasis id="34A9EAA7EC6D9015FE04FBCEFDC50185" box="[421,621,1139,1163]" italics="true" pageId="4" pageNumber="585">Atheris squamigera</emphasis>
</taxonomicName>
(CAS 207868) from Bioko Island.
</paragraph>
</caption>
<subSubSection id="4EC7653EEC6D9015FF66FB04FB2F00FA" pageId="4" pageNumber="585" type="materials_examined">
<paragraph id="066236B5EC6D9015FF66FB04FD36006A" blockId="4.[151,1437,1209,1524]" pageId="4" pageNumber="585">
<emphasis id="34A9EAA7EC6D9015FF66FB04FDFF01DA" bold="true" box="[199,599,1209,1236]" pageId="4" pageNumber="585">Material examined. Bioko Island,</emphasis>
<materialsCitation id="B6B53CE8EC6D9015FDFEFB04FF7A0012" ID-GBIF-Occurrence="3026667305" collectingDate="1998-10" collectionCode="CAS" collectorName="Robert C. Drewes &amp; Lindsay G. Henwood &amp; Jens" country="Equatorial Guinea" elevation="1168" location="Moka Malabo" pageId="4" pageNumber="585" specimenCode="CAS 207866-207868" specimenCount="1">
<emphasis id="34A9EAA7EC6D9015FDFEFB04FCE001DD" bold="true" box="[607,840,1209,1235]" pageId="4" pageNumber="585">
<collectingCountry id="7ECA7625EC6D9015FDFEFB04FCEB01DD" box="[607,835,1209,1235]" name="Equatorial Guinea" pageId="4" pageNumber="585">Equatorial Guinea</collectingCountry>
:
</emphasis>
<specimenCode id="567B9ECEEC6D9015FCEEFB07FBEE01DD" box="[847,1094,1209,1236]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="4" pageNumber="585" type="Museum">CAS 207866–207868</specimenCode>
, vic.
<location id="0302606EEC6D9015FB23FB07FA8901DA" LSID="urn:lsid:plazi:treatment:8E7487A3EC6A9015FF36FD0EFB2F00FA:0302606EEC6D9015FB23FB07FA8901DA" box="[1154,1313,1210,1236]" country="Equatorial Guinea" name="Moka Malabo" pageId="4" pageNumber="585">Moka Malabo</location>
(3.52267º, 8.69417º,
<quantity id="C1259B50EC6D9015FEA9FB60FEF601F6" box="[264,350,1245,1272]" metricMagnitude="3" metricUnit="m" metricValue="1.168" pageId="4" pageNumber="585" unit="m" value="1168.0">
<elevation id="8DF0D186EC6D9015FEA9FB60FEF601F6" box="[264,350,1245,1272]" metricMagnitude="3" metricUnit="m" metricValue="1.168" pageId="4" pageNumber="585" unit="m" value="1168.0">1168 m</elevation>
</quantity>
), collected by
<collectorName id="AB285363EC6D9015FE5EFB63FD6101F6" box="[511,713,1246,1272]" pageId="4" pageNumber="585">Robert C. Drewes</collectorName>
,
<collectorName id="AB285363EC6D9015FD75FB63FC6B01F6" box="[724,963,1246,1272]" pageId="4" pageNumber="585">Lindsay G. Henwood</collectorName>
and
<collectorName id="AB285363EC6D9015FC59FB63FB8001F6" box="[1016,1064,1246,1272]" pageId="4" pageNumber="585">Jens</collectorName>
<collectionCode id="60CCAE70EC6D9015FB8FFB63FBEA01F6" box="[1070,1090,1246,1272]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" pageNumber="585" type="Museum">V</collectionCode>
.
<location id="0302606EEC6D9015FBEAFB63FB0D01F6" LSID="urn:lsid:plazi:treatment:8E7487A3EC6A9015FF36FD0EFB2F00FA:0302606EEC6D9015FBEAFB63FB0D01F6" box="[1099,1189,1246,1272]" country="Equatorial Guinea" name="Vindum" pageId="4" pageNumber="585">Vindum</location>
on 2 and 3 of
<date id="72631075EC6D9015FAE3FB63FF7A0012" pageId="4" pageNumber="585" value="1998-10">
<collectingDate id="6227E99DEC6D9015FAE3FB63FF7A0012" pageId="4" pageNumber="585" value="1998-10">October 1998</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="B6B53CE8EC6D9015FF7FFABFFCF4004E" ID-GBIF-Occurrence="3026667304" collectingDate="1998-10" collectionCode="CAS" collectorName="Robert C. Drewes &amp; Lindsay G. Henwood &amp; Jens V. Vindum" country="Equatorial Guinea" elevation="1553" location="Moka" pageId="4" pageNumber="585" specimenCode="CAS 207869" specimenCount="1">
<specimenCode id="567B9ECEEC6D9015FF7FFABFFEDB0015" box="[222,371,1281,1308]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="4" pageNumber="585" type="Museum">CAS 207869</specimenCode>
, small creek on E side of
<location id="0302606EEC6D9015FD36FABFFD700012" LSID="urn:lsid:plazi:treatment:8E7487A3EC6A9015FF36FD0EFB2F00FA:0302606EEC6D9015FD36FABFFD700012" box="[663,728,1282,1308]" country="Equatorial Guinea" name="Moka" pageId="4" pageNumber="585">Moka</location>
(3.36672º, 8.66492º,
<quantity id="C1259B50EC6D9015FC6FFABCFB8E0012" box="[974,1062,1281,1308]" metricMagnitude="3" metricUnit="m" metricValue="1.553" pageId="4" pageNumber="585" unit="m" value="1553.0">
<elevation id="8DF0D186EC6D9015FC6FFABCFB8E0012" box="[974,1062,1281,1308]" metricMagnitude="3" metricUnit="m" metricValue="1.553" pageId="4" pageNumber="585" unit="m" value="1553.0">1553 m</elevation>
</quantity>
), collected by
<collectorName id="AB285363EC6D9015FB6AFABFFA300012" box="[1227,1432,1282,1308]" pageId="4" pageNumber="585">Robert C. Drewes</collectorName>
,
<collectorName id="AB285363EC6D9015FF36FA9BFE2E004E" box="[151,390,1318,1344]" pageId="4" pageNumber="585">Lindsay G. Henwood</collectorName>
and
<collectorName id="AB285363EC6D9015FE1BFA9BFE42004E" box="[442,490,1318,1344]" pageId="4" pageNumber="585">Jens</collectorName>
<collectionCode id="60CCAE70EC6D9015FE51FA9BFDAC004E" box="[496,516,1318,1344]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" pageNumber="585" type="Museum">V</collectionCode>
.
<location id="0302606EEC6D9015FDACFA9BFDCF004E" LSID="urn:lsid:plazi:treatment:8E7487A3EC6A9015FF36FD0EFB2F00FA:0302606EEC6D9015FDACFA9BFDCF004E" box="[525,615,1318,1344]" country="Equatorial Guinea" name="Vindum" pageId="4" pageNumber="585">Vindum</location>
on 3 of
<date id="72631075EC6D9015FD60FA9BFCF4004E" box="[705,860,1318,1344]" pageId="4" pageNumber="585" value="1998-10">
<collectingDate id="6227E99DEC6D9015FD60FA9BFCF4004E" box="[705,860,1318,1344]" pageId="4" pageNumber="585" value="1998-10">October 1998</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="B6B53CE8EC6D9015FCC6FA9BFD31006A" ID-GBIF-Occurrence="3026667306" collectionCode="MSNG" collectorName="Leonardo Fea" country="Equatorial Guinea" elevation="1369" location="Moka" pageId="4" pageNumber="585" specimenCode="MSNG 30428" specimenCount="1">
<specimenCode id="567B9ECEEC6D9015FCC6FA9BFBA1004E" box="[871,1033,1318,1344]" collectionCode="MSNG" country="Italy" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" pageId="4" pageNumber="585">MSNG 30428</specimenCode>
b,c,
<location id="0302606EEC6D9015FB93FA9BFBDB004E" LSID="urn:lsid:plazi:treatment:8E7487A3EC6A9015FF36FD0EFB2F00FA:0302606EEC6D9015FB93FA9BFBDB004E" box="[1074,1139,1318,1344]" country="Equatorial Guinea" name="Moka" pageId="4" pageNumber="585">Moka</location>
(3.35081º, 8.66254º,
<quantity id="C1259B50EC6D9015FAC5FA98FF18006A" metricMagnitude="3" metricUnit="m" metricValue="1.369" pageId="4" pageNumber="585" unit="m" value="1369.0">
<elevation id="8DF0D186EC6D9015FAC5FA98FF18006A" metricMagnitude="3" metricUnit="m" metricValue="1.369" pageId="4" pageNumber="585" unit="m" value="1369.0">1369 m</elevation>
</quantity>
), collected by
<collectorName id="AB285363EC6D9015FEF2FAF7FE46006A" box="[339,494,1354,1380]" pageId="4" pageNumber="585">Leonardo Fea</collectorName>
on 1901–1902
</materialsCitation>
.
</paragraph>
<paragraph id="066236B5EC6D9015FF66FAD0FB2F00FA" blockId="4.[151,1437,1209,1524]" pageId="4" pageNumber="585">
<emphasis id="34A9EAA7EC6D9015FF66FAD0FD380089" bold="true" box="[199,656,1389,1416]" pageId="4" pageNumber="585">Bibliographic records (not examined).</emphasis>
<materialsCitation id="B6B53CE8EC6D9015FD01FAD3FB2800FA" ID-GBIF-Occurrence="3026667302" collectingDate="1962-11" collectingDateMax="1963-02" collectingDateMin="1962-11" collectionCode="ZFMK" collectorName="Martin Eisentraut" elevation="1369" pageId="4" pageNumber="585" specimenCode="REP 5452-5456" specimenCount="1">
Seven specimens (still in the collections of the Zoologisches Forschungsinstitut und Museum Alexander Koenig, under the accession numbers
<collectionCode id="60CCAE70EC6D9015FC5CFA2FFBE300A2" box="[1021,1099,1426,1452]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="4" pageNumber="585" type="Museum">ZFMK</collectionCode>
/
<specimenCode id="567B9ECEEC6D9015FBF2FA2FFAB900A2" box="[1107,1297,1425,1452]" collectionCode="ZFMK" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="4" pageNumber="585" type="Museum">REP 5452–5456</specimenCode>
), three male adults, three female adults and one juvenile from “Moca” [=Moka] (3.35081º, 8.66254º,
<quantity id="C1259B50EC6D9015FB3EFA08FB5200DE" box="[1183,1274,1461,1488]" metricMagnitude="3" metricUnit="m" metricValue="1.369" pageId="4" pageNumber="585" unit="m" value="1369.0">
<elevation id="8DF0D186EC6D9015FB3EFA08FB5200DE" box="[1183,1274,1461,1488]" metricMagnitude="3" metricUnit="m" metricValue="1.369" pageId="4" pageNumber="585" unit="m" value="1369.0">1369 m</elevation>
</quantity>
), collected by
<collectorName id="AB285363EC6D9015FF36FA67FEF400FA" box="[151,348,1498,1524]" pageId="4" pageNumber="585">Martin Eisentraut</collectorName>
between 7 of
<date id="72631075EC6D9015FE5AFA67FD1800FD" box="[507,688,1497,1524]" pageId="4" pageNumber="585" value="1962-11">
<collectingDate id="6227E99DEC6D9015FE5AFA67FD1800FD" box="[507,688,1497,1524]" pageId="4" pageNumber="585" value="1962-11">November 1962</collectingDate>
</date>
and 19 of
<date id="72631075EC6D9015FC89FA67FC6400FD" box="[808,972,1497,1524]" pageId="4" pageNumber="585" value="1963-02">
<collectingDate id="6227E99DEC6D9015FC89FA67FC6400FD" box="[808,972,1497,1524]" pageId="4" pageNumber="585" value="1963-02">February 1963</collectingDate>
</date>
(
<bibRefCitation id="624C4B44EC6D9015FC7BFA67FBD000FA" author="Mertens, R." box="[986,1144,1497,1524]" pageId="4" pageNumber="585" pagination="211 - 238" refId="ref10030" refString="Mertens, R. (1964) Die Reptilien von Fernando Poo. Bonner Zoologische Beitrage, 15, 211 - 238." type="journal article" year="1964">Mertens 1964</bibRefCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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