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<document id="5F432A9F08571ABB371504AF21A83F02" ID-DOI="10.5281/zenodo.13643699" ID-ISSN="1732-2421" ID-Zenodo-Dep="13643699" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="karina" checkinTime="1723142205826" checkinUser="felipe" docAuthor="Soriano, Carmen &amp; Delclòs, Xavier" docDate="2006" docId="455787BCFF9E2451FC94FC2D6AC8F958" docLanguage="en" docName="ActaPalaeontolPol.51.1.185-200.pdf" docOrigin="Acta Palaeontologica Polonica 51 (1)" docStyle="DocumentStyle:93F5927C23D90727DE855278A6F17B37.1:ActaPalaeontolPol.2002-2009.journal_article" docStyleId="93F5927C23D90727DE855278A6F17B37" docStyleName="ActaPalaeontolPol.2002-2009.journal_article" docStyleVersion="1" docTitle="Zygadenia martinclosas Soriano &amp; Delclòs, 2006, sp. nov." docType="treatment" docVersion="3" lastPageNumber="191" masterDocId="B96EFFC4FF982456FFB8FF876E36FFDB" masterDocTitle="New cupedid beetles from the Lower Cretaceous of Spain and the palaeogeography of the family" masterLastPageNumber="200" masterPageNumber="185" pageNumber="191" updateTime="1725365340826" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="D1374CC145534DAC30CED71652CFB016">New cupedid beetles from the Lower Cretaceous of Spain and the palaeogeography of the family</mods:title>
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<mods:namePart id="4125FC3D204E7B94C25AC00238C73E9D">Soriano, Carmen</mods:namePart>
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<mods:namePart id="914834A8DB884C2C264AD838D0908C0E">Delclòs, Xavier</mods:namePart>
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<mods:title id="94875D7E27B4C524BFB776FB23AA3233">Acta Palaeontologica Polonica</mods:title>
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<mods:date id="A3B184DB9BBD151F0AC12EEDBE899203">2006</mods:date>
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<mods:number id="E65E7A8B65868FC9A1E68EA29D0DA1D4">51</mods:number>
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<mods:identifier id="ECCF410E73D35FC371302233A3B82DF7" type="ISSN">1732-2421</mods:identifier>
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<treatment id="455787BCFF9E2451FC94FC2D6AC8F958" LSID="urn:lsid:plazi:treatment:455787BCFF9E2451FC94FC2D6AC8F958" httpUri="http://treatment.plazi.org/id/455787BCFF9E2451FC94FC2D6AC8F958" lastPageId="7" lastPageNumber="191" pageId="6" pageNumber="191">
<subSubSection id="85E46521FF9E2450FC94FC2D6AFBFC1D" box="[812,1229,938,966]" pageId="6" pageNumber="191" type="nomenclature">
<paragraph id="CD4136AAFF9E2450FC94FC2D6AFBFC1D" blockId="6.[812,1229,938,1000]" box="[812,1229,938,966]" pageId="6" pageNumber="191">
<heading id="960981C6FF9E2450FC94FC2D6AFBFC1D" box="[812,1229,938,966]" fontSize="12" level="2" pageId="6" pageNumber="191" reason="9">
<taxonomicName id="0AFE4D29FF9E2450FC94FC2D6A56FC1D" authorityName="Soriano &amp; Delclòs" authorityYear="2006" box="[812,1120,938,966]" class="Insecta" family="Ommatidae" genus="Zygadenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="191" phylum="Arthropoda" rank="species" species="martinclosas" status="sp. nov.">
<emphasis id="FF8AEAB8FF9E2450FC94FC2D6A56FC1D" box="[812,1120,938,966]" italics="true" pageId="6" pageNumber="191">Zygadenia martinclosas</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E4B957C3FF9E2450FBD1FC2D6AFBFC1D" box="[1129,1229,938,966]" pageId="6" pageNumber="191" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="85E46521FF9E2451FC94FC546AC8F958" lastPageId="7" lastPageNumber="192" pageId="6" pageNumber="191" type="description">
<paragraph id="CD4136AAFF9E2450FC94FC546D98FC33" blockId="6.[812,1229,938,1000]" box="[812,942,979,1000]" pageId="6" pageNumber="191">
<figureCitation id="55C52A2FFF9E2450FC94FC546D49FC33" box="[812,895,979,1000]" captionStart="Fig" captionStartId="7.[102,134,1379,1399]" captionTargetBox="[160,1427,225,1352]" captionTargetId="figure-233@7.[160,1427,225,1352]" captionTargetPageId="7" captionText="Fig. 6. Barremian (Early Cretaceous) ommatins Brochocoleus (Brochocoleini) and Zygadenia (Notocupedini), and cupedins Priacma (Priacmini) and Anaglyphites (Mesocupedini) from Spain. A. Zygadenia diazromerali sp. nov. Holotype MCCM LH 12001a, part, Las Hoyas, Cuenca Province. B. Zygadenia martinclosas sp. nov. Holotype IEI LC 2663a, part, La Cabrúa, Lleida Province. C. Zygadenia longicoxa sp. nov. Holotype MCCM LH 908030a, part, Las Hoyas, Cuenca Province, Lower Cretaceous. D. Zygadenia siniestri sp. nov. Holotype MCCM LH 23515a, part, Las Hoyas, Cuenca Province. E. Priacma sanzii sp. nov. Holotype MCCM LH 24524, part, Las Hoyas, Cuenca Province. F. Anaglyphites zherikhini sp. nov. Holotype MCCM LH 15122a, part, Las Hoyas, Cuenca Province. G.?Anaglyphites pluricavus sp. nov. Holotype IEI LP 97Ga, part, La Pedrera, Lleida Province. Scale bars 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13643715" httpUri="https://zenodo.org/record/13643715/files/figure.png" pageId="6" pageNumber="191">Figs. 6B</figureCitation>
,
<figureCitation id="55C52A2FFF9E2450FC32FC546D9CFC33" box="[906,938,979,1000]" captionStart="Fig" captionStartId="8.[102,135,703,723]" captionTargetBox="[131,1460,226,677]" captionTargetId="figure-655@8.[116,1472,226,677]" captionTargetPageId="8" captionText="Fig. 7. Camera lucida drawings of Barremian (Early Cretaceos) ommatin Zygadenia. A. Zygadenia martinclosas sp. nov. Holotype IEI LC 2663a, part, La Cabrúa, Lleida Province, dorsal (A1) and ventral (A2) aspects. B. Zygadenia longicoxa sp. nov. Holotype MCCM LH 928030a, part, Las Hoyas, Cuenca Province, dorsal (B1) and ventral (B2) aspects. Scale bars 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13643717" httpUri="https://zenodo.org/record/13643717/files/figure.png" pageId="6" pageNumber="191">7A</figureCitation>
.
</paragraph>
<paragraph id="CD4136AAFF9E2450FC94FC7B6B3BFBF7" blockId="6.[812,1485,1019,1223]" pageId="6" pageNumber="191">
<materialsCitation id="7D963CF7FF9E2450FC94FC7B6B3FFBF7" collectorName="La Cabrua" country="Spain" county="Lleida Province" location="Sta. Maria de Meia" municipality="Sierra del Montsec" pageId="6" pageNumber="191" specimenCount="1" typeStatus="holotype">
<emphasis id="FF8AEAB8FF9E2450FC94FC7B6D9FFBCA" box="[812,937,1020,1041]" italics="true" pageId="6" pageNumber="191">
<typeStatus id="12458808FF9E2450FC94FC7B6D6DFBCA" box="[812,859,1020,1041]" pageId="6" pageNumber="191">Type</typeStatus>
locality
</emphasis>
: Quarry near Rubies,
<collectorName id="600B537CFF9E2450FBC4FC7C6AD6FBCB" box="[1148,1248,1019,1040]" pageId="6" pageNumber="191">La Cabrúa</collectorName>
fossil site,
<location id="C8216071FF9E2450FAF5FC7C6D68FBF7" LSID="urn:lsid:plazi:treatment:455787BCFF9E2451FC94FC2D6AC8F958:C8216071FF9E2450FAF5FC7C6D68FBF7" country="Spain" county="Lleida Province" municipality="Sierra del Montsec" name="Sta. Maria de Meia" pageId="6" pageNumber="191">Sta. Maria de Meià</location>
,
<collectingMunicipality id="2D25ACD0FF9E2450FCD1FB906A17FBF7" box="[873,1057,1047,1068]" pageId="6" pageNumber="191">Sierra del Montsec</collectingMunicipality>
,
<collectingCounty id="24204E26FF9E2450FB92FB906AF0FBF7" box="[1066,1222,1047,1068]" pageId="6" pageNumber="191">Lleida Province</collectingCounty>
,
<collectingCountry id="B5E9763AFF9E2450FB77FB906B3FFBF7" box="[1231,1289,1047,1068]" name="Spain" pageId="6" pageNumber="191">Spain</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="CD4136AAFF9E2450FC94FBBC6A53FB53" blockId="6.[812,1485,1019,1223]" pageId="6" pageNumber="191">
<emphasis id="FF8AEAB8FF9E2450FC94FBBC6DCCFB8B" box="[812,1018,1083,1104]" italics="true" pageId="6" pageNumber="191">Stratigraphic horizon</emphasis>
: Calcaires lithographiques à Plantes et Vértébres de la Pedrera de Rubies Fm., Barremian, Lower Cretaceous (
<bibRefCitation id="A96F4B5BFF9E2450FA3AFBD06A6CFB53" author="Martin-Closas, C. &amp; Lopez-Moron, N." pageId="6" pageNumber="191" pagination="29 - 31" refId="ref9705" refString="Martin-Closas, C. and Lopez-Moron, N. 1995. The Charophyte flora. In: X. Martinez-Delclos (ed.), Montsec and Montral-Alcover. Two Konservat- Lagerstatten, Catalonia, Spain. II International Symposium on Lithographic Limestones field trip guide book, 29 - 31. Institut d'Estudis Illerdens, Lleida." type="book chapter" year="1995">MartínClosas and LópezMorón 1995</bibRefCitation>
).
</paragraph>
<paragraph id="CD4136AAFF9E2450FC94FB116D8BFB1C" blockId="6.[812,1485,1019,1223]" pageId="6" pageNumber="191">
<emphasis id="FF8AEAB8FF9E2450FC94FB116A31FB70" box="[812,1031,1174,1195]" italics="true" pageId="6" pageNumber="191">Derivation of the name</emphasis>
: Named after the Spanish palaeobotanist Carles MartínClosas.
</paragraph>
<paragraph id="CD4136AAFF9E2450FC94FB5C6A2EFA0F" blockId="6.[812,1485,1243,1492]" pageId="6" pageNumber="191">
<emphasis id="FF8AEAB8FF9E2450FC94FB5C6DB8FB28" box="[812,910,1243,1267]" italics="true" pageId="6" pageNumber="191">Material</emphasis>
.—
<materialsCitation id="7D963CF7FF9E2450FC17FB5C6A2EFA0F" collectionCode="IEI, MCCM" collectorName="La Cabrua &amp; The &amp; Las Hoyas" pageId="6" pageNumber="191" specimenCode="LC 2663, LH 20151" specimenCount="1" typeStatus="holotype">
<typeStatus id="12458808FF9E2450FC17FB5C6A2FFB2F" box="[943,1049,1243,1268]" pageId="6" pageNumber="191" type="holotype">Holotype</typeStatus>
<collectionCode id="ABEFAE6FFF9E2450FB90FB5C6A7AFB29" box="[1064,1100,1243,1266]" country="Spain" httpUri="http://grbio.org/cool/1m8n-tipp" name="Institut d'Estudis Ilerdencs" pageId="6" pageNumber="191">IEI</collectionCode>
<specimenCode id="9D589ED1FF9E2450FBE3FB5C6AF0FB2F" box="[1115,1222,1243,1268]" collectionCode="IEI" country="Spain" httpUri="http://grbio.org/cool/1m8n-tipp" name="Institut d'Estudis Ilerdencs" pageId="6" pageNumber="191">LC 2663</specimenCode>
part and counterpart, from
<collectorName id="600B537CFF9E2450FCD3FB7C6DD5FACF" box="[875,995,1275,1300]" pageId="6" pageNumber="191">La Cabrúa</collectorName>
fossil site.
<collectorName id="600B537CFF9E2450FBDFFB7C6AA5FACF" box="[1127,1171,1275,1300]" pageId="6" pageNumber="191">The</collectorName>
exemplar is an impression, well preserved, lacking part of antennae and legs.
<collectionCode id="ABEFAE6FFF9E2450FACEFA9C6BFBFAEF" box="[1398,1485,1307,1332]" country="India" httpUri="http://grbio.org/cool/stc7-amit" name="Madras Christian College" pageId="6" pageNumber="191">MCCM</collectionCode>
<specimenCode id="9D589ED1FF9E2450FC94FABC6DABFA8F" box="[812,925,1339,1364]" collectionCode="MCCM" country="India" httpUri="http://grbio.org/cool/stc7-amit" name="Madras Christian College" pageId="6" pageNumber="191">LH 20151</specimenCode>
from
<collectorName id="600B537CFF9E2450FC58FABC6A60FA88" box="[992,1110,1339,1363]" pageId="6" pageNumber="191">Las Hoyas</collectorName>
fossil site, part and counterpart of a beetle without head.
<collectorName id="600B537CFF9E2450FB8AFADC6A68FAAF" box="[1074,1118,1371,1396]" pageId="6" pageNumber="191">The</collectorName>
deformation of the fossil, especially on prosternum, metasternum and final part of abdomen, make impossible to assign this specimen without any doubt to this species.
</materialsCitation>
</paragraph>
<paragraph id="CD4136AAFF9E2450FC94FA606DAAF944" blockId="6.[812,1486,1510,1695]" pageId="6" pageNumber="191">
<emphasis id="FF8AEAB8FF9E2450FC94FA606DA8FA25" box="[812,926,1511,1534]" italics="true" pageId="6" pageNumber="191">Diagnosis</emphasis>
.—Differs from other species with similar ratio between ventrites and similar size (
<taxonomicName id="0AFE4D29FF9E2450FB13F9806D52F9E4" authority="Ponomarenko, 2000" authorityName="Ponomarenko" authorityYear="2000" class="Insecta" family="Ommatidae" genus="Zygadenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="191" phylum="Arthropoda" rank="species" species="sinitzae">
<emphasis id="FF8AEAB8FF9E2450FB13F9806B16F9C5" box="[1195,1312,1543,1566]" italics="true" pageId="6" pageNumber="191">Z. sinitzae</emphasis>
Ponomarenko, 2000
</taxonomicName>
from the Lower Cretaceous of Transbaikalia and
<taxonomicName id="0AFE4D29FF9E2450FA0EF9A06A43F984" authority="Ponomarenko, 1985 b" authorityName="Ponomarenko" authorityYear="1985" class="Insecta" family="Ommatidae" genus="Zygadenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="191" phylum="Arthropoda" rank="species" species="patulus">
<emphasis id="FF8AEAB8FF9E2450FA0EF9A06D48F985" italics="true" pageId="6" pageNumber="191">Z. patulus</emphasis>
<bibRefCitation id="A96F4B5BFF9E2450FC3EF9C16A43F984" author="Ponomarenko, A. G." box="[902,1141,1606,1631]" pageId="6" pageNumber="191" pagination="47 - 87" refId="ref10209" refString="Ponomarenko, A. G. 1985 b. Jurassic beetles from Siberia and Mongolia [in Russian]. In: A. P. Rasnitsyn (ed.), Jurskie nasekomye Sibiri i Mongolii, 47 - 87. Nauka, Moskva." type="book chapter" year="1985">Ponomarenko, 1985b</bibRefCitation>
</taxonomicName>
from the Lower Cretaceous of West Siberia) in narrower pronotum and elytra, and wider epipleura.
</paragraph>
<paragraph id="CD4136AAFF9E2451FC94F9346AC8F958" blockId="6.[812,1485,1713,2026]" lastBlockId="7.[812,1485,1578,1667]" lastPageId="7" lastPageNumber="192" pageId="6" pageNumber="191">
<emphasis id="FF8AEAB8FF9E2450FC94F9346D9BF911" box="[812,941,1715,1738]" italics="true" pageId="6" pageNumber="191">Description</emphasis>
.—Measurements: length
<quantity id="0A069B4FFF9E2450FB7BF9366B10F912" box="[1219,1318,1713,1738]" metricMagnitude="-2" metricUnit="m" metricValue="1.61" pageId="6" pageNumber="191" unit="mm" value="16.1">16.1 mm</quantity>
; width
<quantity id="0A069B4FFF9E2450FACBF9366BFFF912" box="[1395,1481,1713,1738]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="6" pageNumber="191" unit="mm" value="9.1">9.1 mm</quantity>
; elytra length
<quantity id="0A069B4FFF9E2450FC06F9566A13F932" box="[958,1061,1745,1770]" metricMagnitude="-2" metricUnit="m" metricValue="1.08" pageId="6" pageNumber="191" unit="mm" value="10.8">10.8 mm</quantity>
. Body rather elongate and large. Head transverse, about 1.5 times as wide as long without mandibles, narrowed roundly before and after eyes; cheeks and temples shorter than eyes, occiput sloped. Scape and pedicel about two times longer as each of antennomeres 38. Cervical constriction weak. Pronotum transverse, and abruptly narrowed anteriorly, about 1.2 times as wide as long at anterior edge, and about 1.8 times as wide as long at posterior edge; anterior angles sharply distinct, with no incised margin; lateral sides of pronotum somewhat flattened; posterior angles distinct. Disc of pronotum with large rectangular elevation divided by a longitudinal furrow. Prosternum 1.7 times shorter than mesosternum, and nearly two times shorter than metasternum. Procoxae rounded and contiguous. Mesosternum with a deep medial fossa in its last half. Mesocoxae rounded and contiguous. Metasternum with wellexpressed longitudinal and transverse sutures; metepisterna comparatively wide. Abdomen with 5 visible ventrites with sharply marked relief, last ventrite about 2.5 times as long as previous one. Elytra about 1.4 times as long as its combined width, with apex not drawn out into “tail”; epipleura comparatively wide with a row of distinct cells extending to apex. Cells of elytra rounded and not elongated at posterior edge of elytra, with approximately 25 cells forming a row. Legs moderately developed; mesofemora somewhat elongate, widened posteriorly from its 1/3 posterior edge. Abdomen with no distinct tubercles.
</paragraph>
</subSubSection>
</treatment>
</document>