227 lines
29 KiB
XML
227 lines
29 KiB
XML
<document id="DB7DC1241DD98F258F6B997A723AE11F" ID-DOI="10.5281/zenodo.13643699" ID-ISSN="1732-2421" ID-Zenodo-Dep="13643699" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="karina" checkinTime="1723142205826" checkinUser="felipe" docAuthor="Soriano, Carmen & Delclòs, Xavier" docDate="2006" docId="455787BCFF9A2455FC94FC756D81F8A7" docLanguage="en" docName="ActaPalaeontolPol.51.1.185-200.pdf" docOrigin="Acta Palaeontologica Polonica 51 (1)" docStyle="DocumentStyle:93F5927C23D90727DE855278A6F17B37.1:ActaPalaeontolPol.2002-2009.journal_article" docStyleId="93F5927C23D90727DE855278A6F17B37" docStyleName="ActaPalaeontolPol.2002-2009.journal_article" docStyleVersion="1" docTitle="Cionocoleus longicapitis Soriano & Delclòs, 2006, sp. nov." docType="treatment" docVersion="3" lastPageNumber="188" masterDocId="B96EFFC4FF982456FFB8FF876E36FFDB" masterDocTitle="New cupedid beetles from the Lower Cretaceous of Spain and the palaeogeography of the family" masterLastPageNumber="200" masterPageNumber="185" pageNumber="187" updateTime="1725365340826" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="BF5E3BDEE29DE4BEDFF057BBFC4D149C">New cupedid beetles from the Lower Cretaceous of Spain and the palaeogeography of the family</mods:title>
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<mods:namePart id="0F7C8E1F963E22A13A90B1D01310FA00">Soriano, Carmen</mods:namePart>
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<mods:namePart id="CE12208C34D806D7209530F68835A7B4">Delclòs, Xavier</mods:namePart>
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<mods:title id="5BE8BD10F793CC1B0859D25C64A96A9A">Acta Palaeontologica Polonica</mods:title>
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<mods:part id="697D2335734E1253F47A315BCC014E7F">
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<mods:date id="A84784F51D2869B3FAC754D9464C1AAB">2006</mods:date>
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<mods:number id="74D0906F4C5C235D1FF6936BE1B988CD">51</mods:number>
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<mods:identifier id="108751ADD645FC8A73906AB4ABE823B0" type="DOI">10.5281/zenodo.13643699</mods:identifier>
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<mods:identifier id="6256DE19EC768718468C4241B86585E7" type="ISSN">1732-2421</mods:identifier>
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<treatment id="455787BCFF9A2455FC94FC756D81F8A7" ID-DOI="http://doi.org/10.5281/zenodo.13643725" ID-Zenodo-Dep="13643725" LSID="urn:lsid:plazi:treatment:455787BCFF9A2455FC94FC756D81F8A7" httpUri="http://treatment.plazi.org/id/455787BCFF9A2455FC94FC756D81F8A7" lastPageId="3" lastPageNumber="188" pageId="2" pageNumber="187">
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<subSubSection id="85E46521FF9A2454FC94FC756AE1FBD6" box="[812,1239,1009,1038]" pageId="2" pageNumber="187" type="nomenclature">
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<paragraph id="CD4136AAFF9A2454FC94FC756AE1FBD6" blockId="2.[812,1239,1009,1070]" box="[812,1239,1009,1038]" pageId="2" pageNumber="187">
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<heading id="960981C6FF9A2454FC94FC756AE1FBD6" box="[812,1239,1009,1038]" fontSize="12" level="2" pageId="2" pageNumber="187" reason="9">
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<taxonomicName id="0AFE4D29FF9A2454FC94FC756A5DFBD5" authorityName="Soriano & Delclòs" authorityYear="2006" box="[812,1131,1010,1038]" class="Insecta" family="Ommatidae" genus="Cionocoleus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="187" phylum="Arthropoda" rank="species" species="longicapitis" status="sp. nov.">
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<emphasis id="FF8AEAB8FF9A2454FC94FC756A5DFBD5" box="[812,1131,1010,1038]" italics="true" pageId="2" pageNumber="187">Cionocoleus longicapitis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="E4B957C3FF9A2454FBCBFC766AE1FBD6" box="[1139,1239,1009,1037]" pageId="2" pageNumber="187" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="85E46521FF9A2455FC94FB9E6D81F8A7" lastPageId="3" lastPageNumber="188" pageId="2" pageNumber="187" type="description">
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||
<paragraph id="CD4136AAFF9A2454FC94FB9E6D98FBF5" blockId="2.[812,1239,1009,1070]" box="[812,942,1049,1070]" pageId="2" pageNumber="187">
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||
<figureCitation id="55C52A2FFF9A2454FC94FB9E6D49FBF5" box="[812,895,1049,1070]" captionStart="Fig" captionStartId="3.[102,134,1570,1590]" captionTargetBox="[206,1382,225,1543]" captionTargetId="figure-150@3.[206,1382,225,1543]" captionTargetPageId="3" captionText="Fig. 2. Barremian (Early Cretaceous) ommatins Tetraphalerus, Cionocoleus (both Ommatini), Brochocoleus (Brochocoleini), and Zygadenia (Notocupedini) from Spain. A. Tetraphalerus ponomarenkoi sp. nov. Holotype MCCM LH 78a, part, Las Hoyas, Cuenca Province. B. Tetraphalerus penalveri sp. nov. Holotype DEPGM LP 0058b, counterpart, La Pedrera, Lleida Province. C. Cionocoleus longicapitis sp. nov. Holotype IEI LP 0164G, La Pedrera, Lleida Province. D. Brochocoleus indibili sp. nov. Holotype IEI LC 4636a, part, La Cabrúa, Lleida Province. E. Zygadenia viridis sp. nov. Holotype MCCM LH 1805a, part, Las Hoyas, Cuenca Province. F. Zygadenia oculata sp. nov. Holotype IEI LC 3675a, part, La Cabrúa, Lleida Province. Scale bars 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13643703" httpUri="https://zenodo.org/record/13643703/files/figure.png" pageId="2" pageNumber="187">Figs. 2C</figureCitation>
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,
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<figureCitation id="55C52A2FFF9A2454FC32FB9E6D9CFBF5" box="[906,938,1049,1070]" captionStart="Fig" captionStartId="4.[102,135,894,914]" captionTargetBox="[107,1479,229,858]" captionTargetId="figure-460@4.[102,1485,229,864]" captionTargetPageId="4" captionText="Fig. 3. Camera lucida drawings of Barremian (Early Cretaceous) ommatin Cionocoleus and brochocoleinin Brochocoleus from Lleida Province of Spain. A. Cionocoleus longicapitis sp. nov. Holotype IEI LP 0164G, La Pedrera, dorsal (A1) and ventral (A2) aspects. B. Brochocoleus indibili sp. nov. Holotype IEI LC 4636a, part, La Cabrúa, dorsal (B1) and ventral (B2) aspects. Scale bars 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13643707" httpUri="https://zenodo.org/record/13643707/files/figure.png" pageId="2" pageNumber="187">3A</figureCitation>
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.
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</paragraph>
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<paragraph id="CD4136AAFF9A2454FC94FBC66B5EFBA9" blockId="2.[812,1486,1087,1262]" pageId="2" pageNumber="187">
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<materialsCitation id="7D963CF7FF9A2454FC94FBC66B52FBA9" country="Spain" county="Lleida Province" location="Quarry" municipality="Sierra del Montsec" pageId="2" pageNumber="187" specimenCount="1" typeStatus="holotype">
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<emphasis id="FF8AEAB8FF9A2454FC94FBC66D9EFB8D" box="[812,936,1089,1110]" italics="true" pageId="2" pageNumber="187">
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<typeStatus id="12458808FF9A2454FC94FBC66D6DFB8D" box="[812,859,1089,1110]" pageId="2" pageNumber="187">Type</typeStatus>
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locality
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</emphasis>
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:
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<location id="C8216071FF9A2454FC0CFBC66DCEFB8D" LSID="urn:lsid:plazi:treatment:455787BCFF9A2455FC94FC756D81F8A7:C8216071FF9A2454FC0CFBC66DCEFB8D" box="[948,1016,1089,1110]" country="Spain" county="Lleida Province" municipality="Sierra del Montsec" name="Quarry" pageId="2" pageNumber="187">Quarry</location>
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near
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<location id="C8216071FF9A2454FB93FBC66A46FB8D" LSID="urn:lsid:plazi:treatment:455787BCFF9A2455FC94FC756D81F8A7:C8216071FF9A2454FB93FBC66A46FB8D" box="[1067,1136,1089,1110]" country="Spain" county="Lleida Province" municipality="Sierra del Montsec" name="Rubies" pageId="2" pageNumber="187">Rubies</location>
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,
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<location id="C8216071FF9A2454FBC1FBC66B02FB8D" LSID="urn:lsid:plazi:treatment:455787BCFF9A2455FC94FC756D81F8A7:C8216071FF9A2454FBC1FBC66B02FB8D" box="[1145,1332,1087,1110]" country="Spain" county="Lleida Province" municipality="Sierra del Montsec" name="La Pedrera de Meia" pageId="2" pageNumber="187">La Pedrera de Meià</location>
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fossil site,
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<location id="C8216071FF9A2454FA10FBC66D8FFBA9" LSID="urn:lsid:plazi:treatment:455787BCFF9A2455FC94FC756D81F8A7:C8216071FF9A2454FA10FBC66D8FFBA9" country="Spain" county="Lleida Province" municipality="Sierra del Montsec" name="Sta. Maria de Meia" pageId="2" pageNumber="187">Sta. Maria de Meià</location>
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,
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<collectingMunicipality id="2D25ACD0FF9A2454FC7DFBDA6A4AFBA9" box="[965,1148,1117,1138]" pageId="2" pageNumber="187">Sierra del Montsec</collectingMunicipality>
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,
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<collectingCounty id="24204E26FF9A2454FB3EFBDA6B17FBA9" box="[1158,1313,1117,1138]" pageId="2" pageNumber="187">Lleida Province</collectingCounty>
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,
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<collectingCountry id="B5E9763AFF9A2454FA92FBDA6B52FBA9" box="[1322,1380,1117,1138]" name="Spain" pageId="2" pageNumber="187">Spain</collectingCountry>
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="CD4136AAFF9A2454FC94FBF86A53FB17" blockId="2.[812,1486,1087,1262]" pageId="2" pageNumber="187">
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<emphasis id="FF8AEAB8FF9A2454FC94FBF86DCCFB4F" box="[812,1018,1151,1172]" italics="true" pageId="2" pageNumber="187">Stratigraphic horizon</emphasis>
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: Calcaires lithographiques à Plantes et Vértébres de la Pedrera de Rubies Fm., Lower Cretaceous, Barremian (
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<bibRefCitation id="A96F4B5BFF9A2454FA3AFB1C6A6CFB17" author="Martin-Closas, C. & Lopez-Moron, N." pageId="2" pageNumber="187" pagination="29 - 31" refId="ref9705" refString="Martin-Closas, C. and Lopez-Moron, N. 1995. The Charophyte flora. In: X. Martinez-Delclos (ed.), Montsec and Montral-Alcover. Two Konservat- Lagerstatten, Catalonia, Spain. II International Symposium on Lithographic Limestones field trip guide book, 29 - 31. Institut d'Estudis Illerdens, Lleida." type="book chapter" year="1995">Martín−Closas and López−Morón 1995</bibRefCitation>
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).
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</paragraph>
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<paragraph id="CD4136AAFF9A2454FC94FB5E6B53FB35" blockId="2.[812,1486,1087,1262]" box="[812,1381,1241,1262]" pageId="2" pageNumber="187">
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<emphasis id="FF8AEAB8FF9A2454FC94FB5E6A3CFB35" box="[812,1034,1241,1262]" italics="true" pageId="2" pageNumber="187">Derivation of the name</emphasis>
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: After the long head of this species.
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</paragraph>
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<paragraph id="CD4136AAFF9A2454FC94FB786B54FACC" blockId="2.[812,1378,1279,1304]" box="[812,1378,1279,1304]" pageId="2" pageNumber="187">
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<emphasis id="FF8AEAB8FF9A2454FC94FB786DB8FACC" box="[812,910,1279,1303]" italics="true" pageId="2" pageNumber="187">Material</emphasis>
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.—
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<materialsCitation id="7D963CF7FF9A2454FC17FB786B54FACC" box="[943,1378,1279,1304]" collectionCode="IEI" pageId="2" pageNumber="187" specimenCode="LP 0164" specimenCount="1" typeStatus="holotype">
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<typeStatus id="12458808FF9A2454FC17FB786A2FFAC3" box="[943,1049,1279,1304]" pageId="2" pageNumber="187" type="holotype">Holotype</typeStatus>
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<collectionCode id="ABEFAE6FFF9A2454FB98FA876A72FACC" box="[1056,1092,1280,1303]" country="Spain" httpUri="http://grbio.org/cool/1m8n-tipp" name="Institut d'Estudis Ilerdencs" pageId="2" pageNumber="187">IEI</collectionCode>
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<specimenCode id="9D589ED1FF9A2454FBF3FA876A98FAC3" box="[1099,1198,1279,1304]" collectionCode="IEI" country="Spain" httpUri="http://grbio.org/cool/1m8n-tipp" name="Institut d'Estudis Ilerdencs" pageId="2" pageNumber="187">LP 0164</specimenCode>
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G, only the part.
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</materialsCitation>
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</paragraph>
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<paragraph id="CD4136AAFF9A2454FC94FAAE6A1AFA1A" blockId="2.[812,1485,1320,1473]" pageId="2" pageNumber="187">
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<emphasis id="FF8AEAB8FF9A2454FC94FAAE6DA8FA9B" box="[812,926,1321,1344]" italics="true" pageId="2" pageNumber="187">Diagnosis</emphasis>
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.—Distinct from
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<taxonomicName id="0AFE4D29FF9A2454FBE7FAAF6BFBFA9A" authority="Ren, 1995" authorityName="Ren" authorityYear="1995" box="[1119,1485,1320,1345]" class="Insecta" family="Ommatidae" genus="Cionocoleus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="187" phylum="Arthropoda" rank="species" species="magicus">
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<emphasis id="FF8AEAB8FF9A2454FBE7FAAF6B66FA9B" box="[1119,1360,1320,1344]" italics="true" pageId="2" pageNumber="187">Cionocoleus magicus</emphasis>
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<bibRefCitation id="A96F4B5BFF9A2454FAE1FAAE6BFBFA9A" author="Ren, D." box="[1369,1485,1320,1345]" pageId="2" pageNumber="187" refId="ref10644" refString="Ren, D. 1995. Fossil Insects, in Faunae and Stratigraphy of the Jurassic - Cretaceous in Beijing and Adjacent Areas, Beijing [in Chinese with English summary]. 197 pp. Seismic, Beijing." type="book" year="1995">
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Ren,
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<quantity id="0A069B4FFF9A2454FA2DFAAF6BFBFA9A" box="[1429,1485,1320,1345]" metricMagnitude="1" metricUnit="m" metricValue="5.0673" pageId="2" pageNumber="187" unit="in" value="1995.0">1995</quantity>
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</bibRefCitation>
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</taxonomicName>
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in elongate elytra, but without acuminate apex like in
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<taxonomicName id="0AFE4D29FF9A2454FA0BFACF6A5DFA5A" authority="Ponomarenko, 2000" authorityName="Ponomarenko" authorityYear="2000" class="Insecta" family="Ommatidae" genus="Cionocoleus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="187" phylum="Arthropoda" rank="species" species="sibirica">
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<emphasis id="FF8AEAB8FF9A2454FA0BFACF6DB7FA5B" italics="true" pageId="2" pageNumber="187">C. sibirica</emphasis>
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Ponomarenko, 2000
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</taxonomicName>
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. From
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<taxonomicName id="0AFE4D29FF9A2454FB0EFAEF6D84FA7A" authority="Ponomarenko, 1997" authorityName="Ponomarenko" authorityYear="1997" class="Insecta" family="Ommatidae" genus="Cionocoleus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="187" phylum="Arthropoda" rank="species" species="ommamimus">
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<emphasis id="FF8AEAB8FF9A2454FB0EFAEF6B54FA5B" box="[1206,1378,1384,1408]" italics="true" pageId="2" pageNumber="187">C. ommamimus</emphasis>
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<bibRefCitation id="A96F4B5BFF9A2454FAD0FAEE6D84FA7A" author="Ponomarenko, A. G." pageId="2" pageNumber="187" pagination="45 - 55" refId="ref10494" refString="Ponomarenko, A. G. 1997. New beetles of the Family Cupedidae from the Mesozoic of Mongolia. Ommatini, Mesocupedini, Priacmini. Paleontological Journal 31: 45 - 55." type="journal article" year="1997">Ponomarenko, 1997</bibRefCitation>
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</taxonomicName>
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differs in the ratio between abdominal ventrites and length of the head.
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</paragraph>
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<paragraph id="CD4136AAFF9A2455FC94FA556CA8F886" blockId="2.[812,1486,1489,2026]" lastBlockId="3.[102,774,1732,1885]" lastPageId="3" lastPageNumber="188" pageId="2" pageNumber="187">
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<emphasis id="FF8AEAB8FF9A2454FC94FA556D86FA32" box="[812,944,1490,1513]" italics="true" pageId="2" pageNumber="187">Description</emphasis>
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||
.—Measurements: length 19.0 mm; width 7.0 mm; elytra length 13.0 mm. Body rather large and elongate, general shape almost subcylindrical. Head transverse, about two times as wide as long without mandibles; genae as long as eyes, temples rounded and two times shorter than eyes. Mandibles with teeth in vertical plane. Antennae moniliform, antennomeres rounded somewhat wider at posterior edge. Eyes comparatively large and convex. Neck−like constriction not distinct. Pronotum elongate and narrowed towards base and apex, about 1.3 times as long as wide, somewhat wider than head, lateral margins serrate, anterior and posterior angles rounded. Procoxae rounded and contiguous. Mesosternum transverse, about 2.5 times as wide as long. Metasternum 1.3 times as wide as long at its posterior margin, with well expressed transverse and longitudinal sutures. Abdomen with 5 visible ventrites, last ventrite about 1.2 times longer than previous one. Elytra about 3.3 times as long as wide, and almost parallel−sided, epipleura comparatively narrow. Cells on elytra not visible. Profemora somewhat widened posteriorly at its medial length, almost as long as protibiae. Body densely covered with tubercles, larger on head and pronotum, and quite smaller in abdomen.
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</paragraph>
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<caption id="99816622FF9B2455FFDEF9A56B60F97D" ID-DOI="http://doi.org/10.5281/zenodo.13643703" ID-Zenodo-Dep="13643703" httpUri="https://zenodo.org/record/13643703/files/figure.png" pageId="3" pageNumber="188" startId="3.[102,134,1570,1590]" targetBox="[206,1382,225,1543]" targetPageId="3" targetType="figure">
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||
<paragraph id="CD4136AAFF9B2455FFDEF9A56B60F97D" blockId="3.[102,1486,1570,1703]" pageId="3" pageNumber="188">
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Fig. 2. Barremian (Early Cretaceous) ommatins
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<taxonomicName id="0AFE4D29FF9B2455FDB6F9A46CBDF9EC" authorityName="Waterhouse" authorityYear="1901" box="[526,651,1571,1591]" class="Insecta" family="Ommatidae" genus="Tetraphalerus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="188" phylum="Arthropoda" rank="genus">
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||
<emphasis id="FF8AEAB8FF9B2455FDB6F9A46CBDF9EC" box="[526,651,1571,1591]" italics="true" pageId="3" pageNumber="188">Tetraphalerus</emphasis>
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||
</taxonomicName>
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||
,
|
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<taxonomicName id="0AFE4D29FF9B2455FD2DF9A46D32F9EC" authorityName="Ren" authorityYear="1995" box="[661,772,1571,1591]" class="Insecta" family="Ommatidae" genus="Cionocoleus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="188" phylum="Arthropoda" rank="genus">
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||
<emphasis id="FF8AEAB8FF9B2455FD2DF9A46D32F9EC" box="[661,772,1571,1591]" italics="true" pageId="3" pageNumber="188">Cionocoleus</emphasis>
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</taxonomicName>
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(both
|
||
<taxonomicName id="0AFE4D29FF9B2455FC85F9A56DADF9ED" authorityName="Sharp and Muir" authorityYear="1912" box="[829,923,1570,1590]" class="Insecta" family="Ommatidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="188" phylum="Arthropoda" rank="tribe" tribe="Ommatini">Ommatini</taxonomicName>
|
||
),
|
||
<taxonomicName id="0AFE4D29FF9B2455FC11F9A46A15F9EC" authorityName="Hong" authorityYear="1982" box="[937,1059,1571,1591]" class="Insecta" family="Ommatidae" genus="Brochocoleus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="188" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FF8AEAB8FF9B2455FC11F9A46A15F9EC" box="[937,1059,1571,1591]" italics="true" pageId="3" pageNumber="188">Brochocoleus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="0AFE4D29FF9B2455FB96F9A56A86F9ED" authorityName="Hong" authorityYear="1982" box="[1070,1200,1570,1590]" class="Insecta" family="Ommatidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="188" phylum="Arthropoda" rank="tribe" tribe="Brochocoleini">Brochocoleini</taxonomicName>
|
||
), and
|
||
<taxonomicName id="0AFE4D29FF9B2455FB5BF9A46BFBF9ED" authority="(Notocupedini)" baseAuthorityName="Notocupedini" box="[1251,1485,1570,1591]" class="Insecta" family="Ommatidae" genus="Zygadenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="188" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FF8AEAB8FF9B2455FB5BF9A46B77F9EC" box="[1251,1345,1571,1591]" italics="true" pageId="3" pageNumber="188">Zygadenia</emphasis>
|
||
(Notocupedini)
|
||
</taxonomicName>
|
||
from Spain.
|
||
<emphasis id="FF8AEAB8FF9B2455FF6FF9B96ED1F989" bold="true" box="[215,231,1598,1618]" pageId="3" pageNumber="188">A</emphasis>
|
||
.
|
||
<taxonomicName id="0AFE4D29FF9B2455FF4DF9B86FCFF988" authorityName="Soriano & Delclòs" authorityYear="2006" box="[245,505,1599,1619]" class="Insecta" family="Ommatidae" genus="Tetraphalerus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="188" phylum="Arthropoda" rank="species" species="ponomarenkoi" status="sp. nov.">
|
||
<emphasis id="FF8AEAB8FF9B2455FF4DF9B86FCFF988" box="[245,505,1599,1619]" italics="true" pageId="3" pageNumber="188">Tetraphalerus ponomarenkoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E4B957C3FF9B2455FDB9F9B86C7FF989" box="[513,585,1599,1618]" pageId="3" pageNumber="188" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Holotype MCCM LH 78a, part, Las Hoyas, Cuenca Province.
|
||
<emphasis id="FF8AEAB8FF9B2455FB32F9B86AAFF989" bold="true" box="[1162,1177,1599,1618]" pageId="3" pageNumber="188">B</emphasis>
|
||
.
|
||
<taxonomicName id="0AFE4D29FF9B2455FB1EF9B86B49F988" authorityName="Soriano & Delclòs" authorityYear="2006" box="[1190,1407,1599,1619]" class="Insecta" family="Ommatidae" genus="Tetraphalerus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="188" phylum="Arthropoda" rank="species" species="penalveri" status="sp. nov.">
|
||
<emphasis id="FF8AEAB8FF9B2455FB1EF9B86B49F988" box="[1190,1407,1599,1619]" italics="true" pageId="3" pageNumber="188">Tetraphalerus penalveri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E4B957C3FF9B2455FA3EF9B86BF8F989" box="[1414,1486,1599,1618]" pageId="3" pageNumber="188" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Holotype DEPGM LP 0058b, counterpart, La Pedrera, Lleida Province.
|
||
<emphasis id="FF8AEAB8FF9B2455FD62F9DD6CDCF9B5" bold="true" box="[730,746,1626,1646]" pageId="3" pageNumber="188">C</emphasis>
|
||
.
|
||
<taxonomicName id="0AFE4D29FF9B2455FD4CF9DC6DF8F9B4" authorityName="Soriano & Delclòs" authorityYear="2006" box="[756,974,1627,1647]" class="Insecta" family="Ommatidae" genus="Cionocoleus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="188" phylum="Arthropoda" rank="species" species="longicapitis" status="sp. nov.">
|
||
<emphasis id="FF8AEAB8FF9B2455FD4CF9DC6DF8F9B4" box="[756,974,1627,1647]" italics="true" pageId="3" pageNumber="188">Cionocoleus longicapitis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E4B957C3FF9B2455FC6BF9DC6A2EF9B5" box="[979,1048,1627,1646]" pageId="3" pageNumber="188" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Holotype IEI LP 0164G, La Pedrera, Lleida Province.
|
||
<emphasis id="FF8AEAB8FF9B2455FF2DF9F06E93F951" bold="true" box="[149,165,1655,1674]" pageId="3" pageNumber="188">D</emphasis>
|
||
.
|
||
<taxonomicName id="0AFE4D29FF9B2455FF08F9F06F59F950" authorityName="Soriano & Delclòs" authorityYear="2006" box="[176,367,1655,1675]" class="Insecta" family="Ommatidae" genus="Brochocoleus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="188" phylum="Arthropoda" rank="species" species="indibili" status="sp. nov.">
|
||
<emphasis id="FF8AEAB8FF9B2455FF08F9F06F59F950" box="[176,367,1655,1675]" italics="true" pageId="3" pageNumber="188">Brochocoleus indibili</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E4B957C3FF9B2455FECCF9F06F8DF951" box="[372,443,1655,1674]" pageId="3" pageNumber="188" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Holotype IEI LC 4636a, part, La Cabrúa, Lleida Province.
|
||
<emphasis id="FF8AEAB8FF9B2455FC7FF9F06DE0F951" bold="true" box="[967,982,1655,1674]" pageId="3" pageNumber="188">E</emphasis>
|
||
.
|
||
<taxonomicName id="0AFE4D29FF9B2455FC59F9F06A4DF950" authorityName="Soriano & Delclòs" authorityYear="2006" box="[993,1147,1655,1675]" class="Insecta" family="Ommatidae" genus="Zygadenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="188" phylum="Arthropoda" rank="species" species="viridis" status="sp. nov.">
|
||
<emphasis id="FF8AEAB8FF9B2455FC59F9F06A4DF950" box="[993,1147,1655,1675]" italics="true" pageId="3" pageNumber="188">Zygadenia viridis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E4B957C3FF9B2455FB39F9F06AF1F951" box="[1153,1223,1655,1674]" pageId="3" pageNumber="188" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Holotype MCCM LH 1805a, part, Las Hoyas, Cuenca Province.
|
||
<emphasis id="FF8AEAB8FF9B2455FE27F9146F9BF97D" bold="true" box="[415,429,1683,1702]" pageId="3" pageNumber="188">F</emphasis>
|
||
.
|
||
<taxonomicName id="0AFE4D29FF9B2455FE00F9146C69F97C" authorityName="Soriano & Delclòs" authorityYear="2006" box="[440,607,1683,1703]" class="Insecta" family="Ommatidae" genus="Zygadenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="188" phylum="Arthropoda" rank="species" species="oculata" status="sp. nov.">
|
||
<emphasis id="FF8AEAB8FF9B2455FE00F9146C69F97C" box="[440,607,1683,1703]" italics="true" pageId="3" pageNumber="188">Zygadenia oculata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E4B957C3FF9B2455FDDCF9146C9DF97D" box="[612,683,1683,1702]" pageId="3" pageNumber="188" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Holotype IEI LC 3675a, part, La Cabrúa, Lleida Province. Scale bars 2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="CD4136AAFF9B2455FFDEF8F56D81F8A7" blockId="3.[102,775,1905,2026]" lastBlockId="3.[812,1485,1732,1917]" pageId="3" pageNumber="188">
|
||
<emphasis id="FF8AEAB8FF9B2455FFDEF8F56ED7F852" box="[102,225,1906,1929]" italics="true" pageId="3" pageNumber="188">Discussion</emphasis>
|
||
.—This beetle is similar to beetles of the genus
|
||
<taxonomicName id="0AFE4D29FF9B2455FFDEF8156FA3F872" authority="Ponomarenko, 1969" authorityName="Ponomarenko" authorityYear="1969" box="[102,405,1938,1962]" class="Insecta" family="Ommatidae" genus="Omma" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="188" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FF8AEAB8FF9B2455FFDEF8156E87F871" box="[102,177,1938,1962]" italics="true" pageId="3" pageNumber="188">Omma</emphasis>
|
||
<bibRefCitation id="A96F4B5BFF9B2455FF03F8156FA3F872" author="Ponomarenko, A. G." box="[187,405,1938,1961]" pageId="3" pageNumber="188" pagination="1 - 238" refId="ref10112" refString="Ponomarenko, A. G. 1969. Historical development of the Coleoptera-Archostemata [in Russian]. Trudy Paleontologiceskogo Instituta 125: 1 - 238." type="journal article" year="1969">
|
||
Ponomarenko,
|
||
<quantity id="0A069B4FFF9B2455FED8F8156FA3F872" box="[352,405,1938,1961]" metricMagnitude="1" metricUnit="m" metricValue="5.00126" pageId="3" pageNumber="188" unit="in" value="1969.0">1969</quantity>
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
in almost all characters, except for smooth elytra.
|
||
<taxonomicName id="0AFE4D29FF9B2455FEFAF8356FF8F811" authorityName="Ren" authorityYear="1995" box="[322,462,1970,1994]" class="Insecta" family="Ommatidae" genus="Cionocoleus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="188" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FF8AEAB8FF9B2455FEFAF8356FF8F811" box="[322,462,1970,1994]" italics="true" pageId="3" pageNumber="188">Cionocoleus</emphasis>
|
||
</taxonomicName>
|
||
was described by isolated elytra (
|
||
<bibRefCitation id="A96F4B5BFF9B2455FF0DF8566F10F831" author="Ren, D." box="[181,294,2001,2026]" pageId="3" pageNumber="188" refId="ref10644" refString="Ren, D. 1995. Fossil Insects, in Faunae and Stratigraphy of the Jurassic - Cretaceous in Beijing and Adjacent Areas, Beijing [in Chinese with English summary]. 197 pp. Seismic, Beijing." type="book" year="1995">Ren 1995</bibRefCitation>
|
||
), and
|
||
<bibRefCitation id="A96F4B5BFF9B2455FED4F8566C6CF831" author="Ponomarenko, A. G." box="[364,602,2001,2026]" pageId="3" pageNumber="188" pagination="45 - 55" refId="ref10494" refString="Ponomarenko, A. G. 1997. New beetles of the Family Cupedidae from the Mesozoic of Mongolia. Ommatini, Mesocupedini, Priacmini. Paleontological Journal 31: 45 - 55." type="journal article" year="1997">Ponomarenko (1997)</bibRefCitation>
|
||
placed this genus into
|
||
<taxonomicName id="0AFE4D29FF9B2455FC37F9436A3CF906" authorityName="Laporte" authorityYear="1836" box="[911,1034,1732,1757]" class="Insecta" family="Cupedidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="188" phylum="Arthropoda" rank="family">Cupedidae</taxonomicName>
|
||
, but because of the lack of preservation of the fore coxae was not possible to assign it to any subfamily. The new material from the fossil sites of
|
||
<collectingCountry id="B5E9763AFF9B2455FA87F8836B49F8C6" box="[1343,1407,1796,1821]" name="Spain" pageId="3" pageNumber="188">Spain</collectingCountry>
|
||
allows to place this genera in the subfamily
|
||
<taxonomicName id="0AFE4D29FF9B2455FB75F8A36B64F8E6" authorityName="Sharp and Muir" authorityYear="1912" box="[1229,1362,1828,1853]" class="Insecta" family="Ommatidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="188" phylum="Arthropoda" rank="subFamily" subFamily="Ommatinae">Ommatinae</taxonomicName>
|
||
, tribe
|
||
<taxonomicName id="0AFE4D29FF9B2455FA22F8A36D40F886" authorityName="Sharp and Muir" authorityYear="1912" class="Insecta" family="Ommatidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="188" phylum="Arthropoda" rank="tribe" tribe="Ommatini">Ommatini</taxonomicName>
|
||
, by the presence contiguous fore coxae, not divided by any process.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |