272 lines
39 KiB
XML
272 lines
39 KiB
XML
<document id="74E6EF89735601DE113E1326B9ACD06E" ID-CLB-Dataset="32257" ID-DOI="10.11646/zootaxa.4312.3.12" ID-GBIF-Dataset="cad8e5bb-40ef-40b4-b8f5-7d1542d2097d" ID-ISSN="1175-5326" ID-Zenodo-Dep="855675" ID-ZooBank="1F467258-C6D3-4C33-Ba55-A3B34Ff9Acc6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1504174280760" checkinUser="plazi" docAuthor="Novelo-Gutiérrez, Rodolfo" docDate="2017" docId="F417D370DE15FFA4F7B2FCA77CBEB2E9" docLanguage="en" docName="zootaxa.4312.3.12.pdf" docOrigin="Zootaxa 4312 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phyllogomphoides enriquei Novelo-Gutiérrez 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="599" masterDocId="082EAB08DE14FFA0F725FFA2783AB23D" masterDocTitle="Phyllogomphoides enriquei (Odonata: Gomphidae) a new species from Mexico" masterLastPageNumber="600" masterPageNumber="595" pageNumber="596" updateTime="1698467540080" updateUser="ExternalLinkService">
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<mods:title id="38F041D161EA0C3FD2E2DC5EBEC16137">Phyllogomphoides enriquei (Odonata: Gomphidae) a new species from Mexico</mods:title>
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<mods:namePart id="A7E858CFB5F64F37906DD983CEDEFB5C">Novelo-Gutiérrez, Rodolfo</mods:namePart>
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<mods:title id="9A1B7C36877FF0656A2D5AFCF7374B67">Zootaxa</mods:title>
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<mods:part id="1BBD487B905692D4489EB6AA73AF0EC0">
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<mods:date id="D5705D50DECE10C7284DBDA0FAA98803">2017</mods:date>
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<mods:number id="439B057B977DF147D1A7191B67CBA154">2017-08-30</mods:number>
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<mods:number id="7F2EE30AD961E6FCB2B7D27335B53B89">4312</mods:number>
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<mods:identifier id="EC5D9C2149CC922050BE32540DFF3EA8" type="DOI">10.11646/zootaxa.4312.3.12</mods:identifier>
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<treatment id="F417D370DE15FFA4F7B2FCA77CBEB2E9" ID-DOI="http://doi.org/10.5281/zenodo.6043809" ID-GBIF-Taxon="133099979" ID-Zenodo-Dep="6043809" LSID="urn:lsid:plazi:treatment:F417D370DE15FFA4F7B2FCA77CBEB2E9" httpUri="http://treatment.plazi.org/id/F417D370DE15FFA4F7B2FCA77CBEB2E9" lastPageId="4" lastPageNumber="599" pageId="1" pageNumber="596">
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<subSubSection id="34A431EDDE15FFA1F7B2FCA77927B17C" pageId="1" pageNumber="596" type="nomenclature">
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<paragraph id="7C016266DE15FFA1F7B2FCA77A7DB122" blockId="1.[151,583,773,834]" box="[151,583,773,799]" pageId="1" pageNumber="596">
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<heading id="2749D50ADE15FFA1F7B2FCA77A7DB122" bold="true" box="[151,583,773,799]" fontSize="11" level="1" pageId="1" pageNumber="596" reason="1">
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<emphasis id="4ECABE74DE15FFA1F7B2FCA77A7DB122" bold="true" box="[151,583,773,799]" pageId="1" pageNumber="596">
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<taxonomicName id="BBBE19E5DE15FFA1F7B2FCA779D8B122" ID-CoL="9X423" authority="Novelo-Gutiérrez, 2017" authorityName="Novelo-Gutiérrez" authorityYear="2017" box="[151,482,773,799]" class="Insecta" family="Gomphidae" genus="Phyllogomphoides" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="596" phylum="Arthropoda" rank="species" species="enriquei" status="sp. nov.">
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<emphasis id="4ECABE74DE15FFA1F7B2FCA779D8B122" bold="true" box="[151,482,773,799]" italics="true" pageId="1" pageNumber="596">Phyllogomphoides enriquei</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="55F9030FDE15FFA1F6CCFCA77A7DB122" box="[489,583,773,799]" pageId="1" pageNumber="596" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="7C016266DE15FFA1F7B2FC8B7927B17C" blockId="1.[151,583,773,834]" box="[151,285,809,834]" pageId="1" pageNumber="596">
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<figureCitation id="E4857EE3DE15FFA1F7B2FC8B78D8B17F" box="[151,226,809,834]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1854,1877]" captionTargetBox="[251,1269,199,1822]" captionTargetId="figure@2.[250,1279,199,1834]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Phyllogomphoides enriquei sp. nov., holotype ♂. 1) Head and thorax, lateral view (head turned up to the left); 2) S 8 ¯ 10 and caudal appendages, left lateral view, showing inferior margins of S 8 - 9 foliate and scalloped; 3) Accessory genitalia, ventral view: alb, anterior lobe of anterior hamule; Ah, anterior hamuli; Alm, anterior lamina; dt, denticles; fl, flagella; msp, middle spine; n, notch; plb, posterior lobe of anterior hamule; psp, posterior spine of anterior lobe of anterior hamule; V 1, V 4, segments 1 and 4 of vesica spermalis; 4) S 9 ¯ 10 and caudal appendages, dorsal view. Ce, cercus; dt, dorsal tooth; Ep, epiproct." httpUri="https://zenodo.org/record/855679/files/figure.png" pageId="1" pageNumber="596">Figs. 1</figureCitation>
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¯5, 7
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</paragraph>
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</subSubSection>
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<subSubSection id="34A431EDDE15FFA1F7B2FCD27ABFB1F3" pageId="1" pageNumber="596" type="materials_examined">
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<paragraph id="7C016266DE15FFA1F7B2FCD27ABFB1F3" blockId="1.[151,1437,879,2030]" pageId="1" pageNumber="596">
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<materialsCitation id="CCD6683BDE15FFA1F7B2FCD27ABFB1F3" ID-GBIF-Occurrence="1587262052" collectingDate="2005-07-04" collectionCode="IEXA" collectorName="Novelo & J. A. Gomez & Coleccion Entomologica & Instituto de Ecologia, A. C. & Xalapa" country="Mexico" county="Instituto de Ecologia" elevation="616" latitude="19.008734" location="Municipality of Tepalcatepec" longLatPrecision="1" longitude="-103.02427" municipality="Rio Pinolapa" pageId="1" pageNumber="596" specimenCount="1" specimenCount-male="1" stateProvince="Michoacan" typeStatus="holotype">
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<emphasis id="4ECABE74DE15FFA1F7B2FCD27930B1B5" bold="true" box="[151,266,880,904]" pageId="1" pageNumber="596">
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<typeStatus id="A305DCC4DE15FFA1F7B2FCD2793CB1B5" box="[151,262,880,904]" pageId="1" pageNumber="596" type="holotype">Holotype</typeStatus>
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,
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</emphasis>
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<specimenCount id="6AB8A9EFDE15FFA1F630FCD27910B1BA" box="[277,298,880,903]" pageId="1" pageNumber="596" type="male">♂</specimenCount>
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:
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<collectingCountry id="04A922F6DE15FFA1F618FCD2798AB1B5" box="[317,432,880,904]" name="Mexico" pageId="1" pageNumber="596">MEXICO</collectingCountry>
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:
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<collectingRegion id="BE7AAC84DE15FFA1F69BFCCD7A04B1B5" box="[446,574,879,904]" country="Mexico" name="Michoacan de Ocampo" pageId="1" pageNumber="596">Michoacán</collectingRegion>
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;
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<location id="796134BDDE15FFA1F569FCCD7BA7B1B5" LSID="urn:lsid:plazi:treatment:F417D370DE15FFA4F7B2FCA77CBEB2E9:796134BDDE15FFA1F569FCCD7BA7B1B5" box="[588,925,879,904]" country="Mexico" county="Instituto de Ecologia" latitude="19.008734" longLatPrecision="1" longitude="-103.02427" municipality="Rio Pinolapa" name="Municipality of Tepalcatepec" pageId="1" pageNumber="596" stateProvince="Michoacan">Municipality of Tepalcatepec</location>
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,
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<collectingMunicipality id="9C65F81CDE15FFA1F48EFCD27C79B1B5" box="[939,1091,879,904]" pageId="1" pageNumber="596">Río Pinolapa</collectingMunicipality>
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,
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<geoCoordinate id="198A04A1DE15FFA1F377FCCD7CDEB1BA" box="[1106,1252,879,904]" direction="north" orientation="latitude" pageId="1" pageNumber="596" precision="1" value="19.008734">19°00.524 N</geoCoordinate>
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;
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<geoCoordinate id="198A04A1DE15FFA1F3D1FCCD7DA2B1BA" box="[1268,1432,879,904]" direction="west" orientation="longitude" pageId="1" pageNumber="596" precision="1" value="-103.02427">103°01.456 W</geoCoordinate>
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,
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<elevation id="F7938555DE15FFA1F7B2FC307973B196" box="[151,329,914,939]" metricMagnitude="2" metricUnit="m" metricValue="6.16" pageId="1" pageNumber="596" unit="m" value="616.0">
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elevation
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<quantity id="BB46CF83DE15FFA1F623FC307973B196" box="[262,329,914,939]" metricMagnitude="2" metricUnit="m" metricValue="6.16" pageId="1" pageNumber="596" unit="m" value="616.0">616m</quantity>
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</elevation>
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, 0
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<collectingDate id="1844BD4EDE15FFA1F641FC3179D7B196" box="[356,493,914,939]" pageId="1" pageNumber="596" value="2005-07-04">
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<date id="080044A6DE15FFA1F641FC3179D7B196" box="[356,493,914,939]" pageId="1" pageNumber="596" value="2005-07-04">4 July 2005</date>
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</collectingDate>
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, R.
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<collectorName id="D14B07B0DE15FFA1F539FC307A55B196" box="[540,623,914,939]" pageId="1" pageNumber="596">Novelo</collectorName>
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&
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<collectorName id="D14B07B0DE15FFA1F5B0FC317B26B196" box="[661,796,915,939]" pageId="1" pageNumber="596">J.A. Gómez</collectorName>
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leg.
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<typeStatus id="A305DCC4DE15FFA1F472FC307B85B196" box="[855,959,914,939]" pageId="1" pageNumber="596" type="holotype">Holotype</typeStatus>
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deposited in
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<collectorName id="D14B07B0DE15FFA1F37FFC307D4BB196" box="[1114,1393,914,939]" pageId="1" pageNumber="596">Colección Entomológica</collectorName>
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del
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<collectorName id="D14B07B0DE15FFA1F7B2FC1779FFB1F3" box="[151,453,949,974]" pageId="1" pageNumber="596">
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<collectingCounty id="95601AEADE15FFA1F7B2FC1779B9B1F3" box="[151,387,949,974]" pageId="1" pageNumber="596">Instituto de Ecología</collectingCounty>
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, A.C.
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</collectorName>
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,
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<collectorName id="D14B07B0DE15FFA1F6F4FC177A1AB1F3" box="[465,544,949,974]" pageId="1" pageNumber="596">Xalapa</collectorName>
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||
(
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<collectionCode id="1AAFFAA3DE15FFA1F515FC147A43B1F3" box="[560,633,950,974]" pageId="1" pageNumber="596">IEXA</collectionCode>
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||
).
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||
</materialsCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="34A431EDDE15FFA1F7E2FC7B7DBCB628" pageId="1" pageNumber="596" type="etymology">
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<paragraph id="7C016266DE15FFA1F7E2FC7B7DBCB628" blockId="1.[151,1437,879,2030]" pageId="1" pageNumber="596">
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<emphasis id="4ECABE74DE15FFA1F7E2FC7B7974B1CC" bold="true" box="[199,334,985,1009]" pageId="1" pageNumber="596">Etymology.</emphasis>
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We name this species
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<taxonomicName id="BBBE19E5DE15FFA1F57EFC787A82B1CC" authorityName="Novelo-Gutiérrez" authorityYear="2017" box="[603,696,986,1009]" class="Insecta" family="Gomphidae" genus="Phyllogomphoides" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="596" phylum="Arthropoda" rank="species" species="enriquei">
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<emphasis id="4ECABE74DE15FFA1F57EFC787A82B1CC" box="[603,696,986,1009]" italics="true" pageId="1" pageNumber="596">enriquei</emphasis>
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</taxonomicName>
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(a noun in the genitive case) in honor to our friend and fellow odonatologist Enrique González Soriano, for his great contributions to the knowledge of the
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<taxonomicName id="BBBE19E5DE15FFA1F381FC5E7D39B628" authorityName="Fabricius" authorityYear="1793" box="[1188,1283,1020,1045]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="596" phylum="Arthropoda" rank="order">Odonata</taxonomicName>
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of
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<collectingRegion id="BE7AAC84DE15FFA1F20DFC5E7DBBB628" box="[1320,1409,1020,1045]" country="Mexico" name="Mexico" pageId="1" pageNumber="596">
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<collectingCountry id="04A922F6DE15FFA1F20DFC5E7DBBB628" box="[1320,1409,1020,1045]" name="Mexico" pageId="1" pageNumber="596">Mexico</collectingCountry>
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</collectingRegion>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="34A431EDDE15FFA4F7E2FBBD7CBEB2E9" lastPageId="4" lastPageNumber="599" pageId="1" pageNumber="596" type="description">
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<paragraph id="7C016266DE15FFA1F7E2FBBD7A2FB666" blockId="1.[151,1437,879,2030]" pageId="1" pageNumber="596">
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<emphasis id="4ECABE74DE15FFA1F7E2FBBD7A08B605" bold="true" box="[199,562,1055,1080]" pageId="1" pageNumber="596">
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Description of male
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<typeStatus id="A305DCC4DE15FFA1F6E0FB827A14B605" box="[453,558,1056,1080]" pageId="1" pageNumber="596" type="holotype">holotype</typeStatus>
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.
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</emphasis>
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Body mostly dark brown, with five pale stripes on pterothorax, and pale, creamy yellow spots on abdomen.
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</paragraph>
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<paragraph id="7C016266DE15FFA1F7E2FBC779DDB748" blockId="1.[151,1437,879,2030]" pageId="1" pageNumber="596">
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||
<emphasis id="4ECABE74DE15FFA1F7E2FBC7793EB643" box="[199,260,1125,1150]" italics="true" pageId="1" pageNumber="596">Head</emphasis>
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||
(
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<figureCitation id="E4857EE3DE15FFA1F636FBC7796FB643" box="[275,341,1125,1150]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1854,1877]" captionTargetBox="[251,1269,199,1822]" captionTargetId="figure@2.[250,1279,199,1834]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Phyllogomphoides enriquei sp. nov., holotype ♂. 1) Head and thorax, lateral view (head turned up to the left); 2) S 8 ¯ 10 and caudal appendages, left lateral view, showing inferior margins of S 8 - 9 foliate and scalloped; 3) Accessory genitalia, ventral view: alb, anterior lobe of anterior hamule; Ah, anterior hamuli; Alm, anterior lamina; dt, denticles; fl, flagella; msp, middle spine; n, notch; plb, posterior lobe of anterior hamule; psp, posterior spine of anterior lobe of anterior hamule; V 1, V 4, segments 1 and 4 of vesica spermalis; 4) S 9 ¯ 10 and caudal appendages, dorsal view. Ce, cercus; dt, dorsal tooth; Ep, epiproct." httpUri="https://zenodo.org/record/855679/files/figure.png" pageId="1" pageNumber="596">Fig. 1</figureCitation>
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): Face mostly pale covered with numerous long, dark, stiff setae; labrum mostly pale with a wide blackish-brown band on anterior border, posterior border brown, with a small, brown U-shaped median spot; mandibles pale basally, tips reddish-brown; anteclypeus and postclypeus brown inferiorly, creamy pale superiorly; lower surface of frons brown, upper and dorsal surfaces mostly pale, a blackish-brown stripe in the union with vertex; antennal scape dark brown with apical rim creamy pale, pedicel dark brown, flagellum brown; vertex dark brown, depressed area between ocelli with a large, quadrate pale spot, ocelli yellow; occiput dark brown with a large, trapezoid, central pale spot, its posterior border fringed with long, stiff, reddish-brown setae; labium mostly pale, submentum light brown.
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</paragraph>
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<paragraph id="7C016266DE15FFA1F7E2FADD7CEFB5D3" blockId="1.[151,1437,879,2030]" pageId="1" pageNumber="596">
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||
<emphasis id="4ECABE74DE15FFA1F7E2FADD7922B7A5" box="[199,280,1407,1432]" italics="true" pageId="1" pageNumber="596">Thorax</emphasis>
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||
(
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<figureCitation id="E4857EE3DE15FFA1F60CFADD7954B7A5" box="[297,366,1407,1432]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1854,1877]" captionTargetBox="[251,1269,199,1822]" captionTargetId="figure@2.[250,1279,199,1834]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Phyllogomphoides enriquei sp. nov., holotype ♂. 1) Head and thorax, lateral view (head turned up to the left); 2) S 8 ¯ 10 and caudal appendages, left lateral view, showing inferior margins of S 8 - 9 foliate and scalloped; 3) Accessory genitalia, ventral view: alb, anterior lobe of anterior hamule; Ah, anterior hamuli; Alm, anterior lamina; dt, denticles; fl, flagella; msp, middle spine; n, notch; plb, posterior lobe of anterior hamule; psp, posterior spine of anterior lobe of anterior hamule; V 1, V 4, segments 1 and 4 of vesica spermalis; 4) S 9 ¯ 10 and caudal appendages, dorsal view. Ce, cercus; dt, dorsal tooth; Ep, epiproct." httpUri="https://zenodo.org/record/855679/files/figure.png" pageId="1" pageNumber="596">Fig. 1</figureCitation>
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): Pronotum reddish-brown, with a pale middorsal twin-spot on posterior margin of mid- and posterior lobes. Pterothorax reddish-brown, with five complete pale stripes as shown in
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<figureCitation id="E4857EE3DE15FFA1F35DFA007C86B786" box="[1144,1212,1442,1467]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1854,1877]" captionTargetBox="[251,1269,199,1822]" captionTargetId="figure@2.[250,1279,199,1834]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Phyllogomphoides enriquei sp. nov., holotype ♂. 1) Head and thorax, lateral view (head turned up to the left); 2) S 8 ¯ 10 and caudal appendages, left lateral view, showing inferior margins of S 8 - 9 foliate and scalloped; 3) Accessory genitalia, ventral view: alb, anterior lobe of anterior hamule; Ah, anterior hamuli; Alm, anterior lamina; dt, denticles; fl, flagella; msp, middle spine; n, notch; plb, posterior lobe of anterior hamule; psp, posterior spine of anterior lobe of anterior hamule; V 1, V 4, segments 1 and 4 of vesica spermalis; 4) S 9 ¯ 10 and caudal appendages, dorsal view. Ce, cercus; dt, dorsal tooth; Ep, epiproct." httpUri="https://zenodo.org/record/855679/files/figure.png" pageId="1" pageNumber="596">Fig. 1</figureCitation>
|
||
; middorsal thoracic carina pale; antealar crest dark; first antehumeral stripe connected to pale mesothoracic collar forming an inverted "7" reaching border of antealar crest superiorly; second antehumeral stripe the narrowest, divided in a stripe that covers inferior 0.70 of mesepisternum’s length, and a superior circular spot close to antealar crest; mesepimeral stripe two times wider than second antehumeral stripe, covering most of mesepimeron (90%), more or less parallelsided, gradually widening at upper 0.20; metepisternal stripe covering the full-length of metepisternum, almost encircling the spiracle, and forming a circular spot superiorly; metepimeral stripe forming and inverted “L” reaching subalar carina; metaposternum pale; pectus mostly grayish-yellow, with a large, distal, quadrate brown spot.
|
||
<emphasis id="4ECABE74DE15FFA1F7F2F91F7936B4E9" box="[215,268,1725,1748]" italics="true" pageId="1" pageNumber="596">Legs</emphasis>
|
||
: coxae and trocanters ventrally pale, dorsally light brown; femora mostly light brown, distally dark reddish-brown, anterior femora broadly pale ventrally; tibiae, tarsi and pretarsal claws reddish-black; armature black, protibiae with modified spines (scale-like) on distal half of external border, long and slender spines on internal borders.
|
||
<emphasis id="4ECABE74DE15FFA1F670F88579A2B503" box="[341,408,1831,1854]" italics="true" pageId="1" pageNumber="596">Wings</emphasis>
|
||
: hyaline, tinged with brown at extreme base, venation black, anterior margin of costae with a continuous well-defined yellow line in all the wings; basal subcostal crossveins present; second primary antenodal crossvein the 7th (left), the 8th (right) in FW, the 6th in HW; antenodal crossveins FW 19, HW 14; postnodal crossveins FW 13, HW 13; triangles 3-celled; subtriangles 3-celled in FW, 2-celled in right HW, 3-celled in left HW; supratriangles 3-celled; anal loop 3-celled (left), 4-celled (right); anal triangle 4-celled; pterostigma dark brown with brace vein, surmounting six cells (left), five cells (right) in FW, six cells in HW.
|
||
</paragraph>
|
||
<caption id="28C132EEDE16FFA2F7B2F89C78CBB5D3" httpUri="https://zenodo.org/record/855679/files/figure.png" pageId="2" pageNumber="597" targetBox="[251,1269,199,1822]" targetPageId="2">
|
||
<paragraph id="7C016266DE16FFA2F7B2F89C78CBB5D3" blockId="2.[151,1436,1854,2030]" pageId="2" pageNumber="597">
|
||
<emphasis id="4ECABE74DE16FFA2F7B2F89C7900B569" bold="true" box="[151,314,1854,1877]" pageId="2" pageNumber="597">FIGURES 1–4.</emphasis>
|
||
<taxonomicName id="BBBE19E5DE16FFA2F665F89D7A6DB568" authority="Novelo-Gutiérrez, 2017" authorityName="Novelo-Gutiérrez" authorityYear="2017" box="[320,599,1855,1877]" class="Insecta" family="Gomphidae" genus="Phyllogomphoides" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="597" phylum="Arthropoda" rank="species" species="enriquei" status="sp. nov.">
|
||
<emphasis id="4ECABE74DE16FFA2F665F89D7A6DB568" box="[320,599,1855,1877]" italics="true" pageId="2" pageNumber="597">Phyllogomphoides enriquei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="55F9030FDE16FFA2F57BF89D7A94B569" box="[606,686,1855,1876]" pageId="2" pageNumber="597" rank="species">
|
||
<emphasis id="4ECABE74DE16FFA2F57BF89D7A94B569" bold="true" box="[606,686,1855,1876]" pageId="2" pageNumber="597">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype ♂. 1) Head and thorax, lateral view (head turned up to the left); 2) S8¯10 and caudal appendages, left lateral view, showing inferior margins of S8-9 foliate and scalloped; 3) Accessory genitalia, ventral view:
|
||
<emphasis id="4ECABE74DE16FFA2F6A1F8DE799EB5AF" box="[388,420,1916,1938]" italics="true" pageId="2" pageNumber="597">alb</emphasis>
|
||
, anterior lobe of anterior hamule;
|
||
<emphasis id="4ECABE74DE16FFA2F5D8F8DE7B23B5AF" box="[765,793,1916,1938]" italics="true" pageId="2" pageNumber="597">Ah</emphasis>
|
||
, anterior hamuli;
|
||
<emphasis id="4ECABE74DE16FFA2F4E8F8DE7BCCB5AF" box="[973,1014,1916,1938]" italics="true" pageId="2" pageNumber="597">Alm</emphasis>
|
||
, anterior lamina;
|
||
<emphasis id="4ECABE74DE16FFA2F38DF8DE7C86B5AF" box="[1192,1212,1916,1938]" italics="true" pageId="2" pageNumber="597">dt</emphasis>
|
||
, denticles;
|
||
<emphasis id="4ECABE74DE16FFA2F20AF8DE7D07B5AF" box="[1327,1341,1916,1938]" italics="true" pageId="2" pageNumber="597">fl</emphasis>
|
||
, flagella;
|
||
<emphasis id="4ECABE74DE16FFA2F7B2F83E78FAB58C" box="[151,192,1948,1969]" italics="true" pageId="2" pageNumber="597">msp</emphasis>
|
||
, middle spine;
|
||
<emphasis id="4ECABE74DE16FFA2F67EF83E7952B58C" box="[347,360,1948,1969]" italics="true" pageId="2" pageNumber="597">n</emphasis>
|
||
, notch;
|
||
<emphasis id="4ECABE74DE16FFA2F69FF83979E0B58C" box="[442,474,1947,1969]" italics="true" pageId="2" pageNumber="597">plb</emphasis>
|
||
, posterior lobe of anterior hamule;
|
||
<emphasis id="4ECABE74DE16FFA2F464F83E7B5EB58C" box="[833,868,1948,1969]" italics="true" pageId="2" pageNumber="597">psp</emphasis>
|
||
, posterior spine of anterior lobe of anterior hamule; V1, V4, segments 1 and 4 of vesica spermalis; 4) S9¯10 and caudal appendages, dorsal view.
|
||
<emphasis id="4ECABE74DE16FFA2F36BF8187C53B5F2" box="[1102,1129,1978,1999]" italics="true" pageId="2" pageNumber="597">Ce</emphasis>
|
||
, cercus;
|
||
<emphasis id="4ECABE74DE16FFA2F3EEF81B7CE5B5F2" box="[1227,1247,1977,1999]" italics="true" pageId="2" pageNumber="597">dt</emphasis>
|
||
, dorsal tooth;
|
||
<emphasis id="4ECABE74DE16FFA2F25CF8187DAFB5F2" box="[1401,1429,1978,1999]" italics="true" pageId="2" pageNumber="597">Ep</emphasis>
|
||
, epiproct.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="28C132EEDE17FFA3F7B2FBA97A45B662" httpUri="https://zenodo.org/record/855681/files/figure.png" pageId="3" pageNumber="598" targetBox="[173,1436,193,1014]" targetPageId="3">
|
||
<paragraph id="7C016266DE17FFA3F7B2FBA97A45B662" blockId="3.[151,1436,1035,1119]" pageId="3" pageNumber="598">
|
||
<emphasis id="4ECABE74DE17FFA3F7B2FBA97900B61C" bold="true" box="[151,314,1035,1058]" pageId="3" pageNumber="598">FIGURES 5–6.</emphasis>
|
||
Details of
|
||
<taxonomicName id="BBBE19E5DE17FFA3F68DFBAE7A5EB61F" authorityName="Belle" authorityYear="1970" box="[424,612,1036,1058]" class="Insecta" family="Gomphidae" genus="Phyllogomphoides" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="598" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4ECABE74DE17FFA3F68DFBAE7A5EB61F" box="[424,612,1036,1058]" italics="true" pageId="3" pageNumber="598">Phyllogomphoides</emphasis>
|
||
</taxonomicName>
|
||
spp. 5)
|
||
<taxonomicName id="BBBE19E5DE17FFA3F591FBAE7BF0B61F" authority="Novelo-Gutiérrez, 2017" authorityName="Novelo-Gutiérrez" authorityYear="2017" box="[692,970,1036,1058]" class="Insecta" family="Gomphidae" genus="Phyllogomphoides" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="598" phylum="Arthropoda" rank="species" species="enriquei" status="sp. nov.">
|
||
<emphasis id="4ECABE74DE17FFA3F591FBAE7BF0B61F" box="[692,970,1036,1058]" italics="true" pageId="3" pageNumber="598">Phyllogomphoides enriquei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="55F9030FDE17FFA3F4F5FBAE7C1AB61C" box="[976,1056,1036,1057]" pageId="3" pageNumber="598" rank="species">
|
||
<emphasis id="4ECABE74DE17FFA3F4F5FBAE7C1AB61C" bold="true" box="[976,1056,1036,1057]" pageId="3" pageNumber="598">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype ♂. Caudal appendages, left lateral view.
|
||
<emphasis id="4ECABE74DE17FFA3F605FB897906B67D" box="[288,316,1067,1088]" italics="true" pageId="3" pageNumber="598">Ce</emphasis>
|
||
, cercus;
|
||
<emphasis id="4ECABE74DE17FFA3F6B8FB88798AB67D" box="[413,432,1066,1088]" italics="true" pageId="3" pageNumber="598">dt</emphasis>
|
||
, dorsal tooth;
|
||
<emphasis id="4ECABE74DE17FFA3F56BFB887A4BB67D" box="[590,625,1066,1088]" italics="true" pageId="3" pageNumber="598">dsp</emphasis>
|
||
, dorsal spine;
|
||
<emphasis id="4ECABE74DE17FFA3F435FB897B16B67D" box="[784,812,1067,1088]" italics="true" pageId="3" pageNumber="598">Ep</emphasis>
|
||
, epiproct;
|
||
<emphasis id="4ECABE74DE17FFA3F485FB897BF8B67D" box="[928,962,1067,1088]" italics="true" pageId="3" pageNumber="598">vsp</emphasis>
|
||
, ventral spine. 6)
|
||
<taxonomicName id="BBBE19E5DE17FFA3F3A2FB887D42B67D" authorityName="Gonzalez-Soriano & Novelo-Gutierrez" authorityYear="1990" box="[1159,1400,1066,1088]" class="Insecta" family="Gomphidae" genus="Phyllogomphoides" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="598" phylum="Arthropoda" rank="species" species="luisi">
|
||
<emphasis id="4ECABE74DE17FFA3F3A2FB887D42B67D" box="[1159,1400,1066,1088]" italics="true" pageId="3" pageNumber="598">Phyllogomphoides luisi</emphasis>
|
||
</taxonomicName>
|
||
, ♂ (Xochitepec). Accessory genitalia, ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="7C016266DE17FFA3F7E2FB247CB9B5E6" blockId="3.[151,1437,1158,2011]" pageId="3" pageNumber="598">
|
||
<emphasis id="4ECABE74DE17FFA3F7E2FB24790BB6A2" box="[199,305,1158,1183]" italics="true" pageId="3" pageNumber="598">Abdomen</emphasis>
|
||
: Dark color on S1-2 reddish-brown, black on S3¯7, S8¯9 black on dorsum, light brown at sides, S10 light brown. Pale coloration creamy yellow as follows: a middorsal stripe and the ventral 0.5 of tergum on S1; a middorsal stripe, auricles, and a posteroventral, narrow, vertical spot on S2; a middorsal stripe on basal 0.80 constricted at basal 0.85 of its length, and a broad basoventral spot on basal 0.40 of S3; a middorsal stripe on basal 0.80 tapering posteriorly, a slightly broad basoventral spot on basal 0.25 of S4; a middorsal stripe on basal 0.25 tapering posteriorly, a slightly broad basoventral spot on basal 0.25 of S5; a middorsal spot on basal 0.10, a slightly broad basoventral spot on basal 0.20 of S6; basal half of S7; S8 with pale strikes on basoventral 0.10. Inferior margins of S8-9 foliate and scalloped (
|
||
<figureCitation id="E4857EE3DE17FFA3F575FA207AACB7A6" box="[592,662,1410,1435]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1854,1877]" captionTargetBox="[251,1269,199,1822]" captionTargetId="figure@2.[250,1279,199,1834]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Phyllogomphoides enriquei sp. nov., holotype ♂. 1) Head and thorax, lateral view (head turned up to the left); 2) S 8 ¯ 10 and caudal appendages, left lateral view, showing inferior margins of S 8 - 9 foliate and scalloped; 3) Accessory genitalia, ventral view: alb, anterior lobe of anterior hamule; Ah, anterior hamuli; Alm, anterior lamina; dt, denticles; fl, flagella; msp, middle spine; n, notch; plb, posterior lobe of anterior hamule; psp, posterior spine of anterior lobe of anterior hamule; V 1, V 4, segments 1 and 4 of vesica spermalis; 4) S 9 ¯ 10 and caudal appendages, dorsal view. Ce, cercus; dt, dorsal tooth; Ep, epiproct." httpUri="https://zenodo.org/record/855679/files/figure.png" pageId="3" pageNumber="598">Fig. 2</figureCitation>
|
||
), foliation on S8 increasing gradually in width caudally, ending in a wide rounded lobe which surpasses slightly the anterior margin of S9 (
|
||
<quantity id="BB46CF83DE17FFA3F49CFA047C1EB782" box="[953,1060,1446,1471]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="598" unit="mm" value="0.5">0.50 mm</quantity>
|
||
), the edge of this foliation with a row of small spines on apical 0.30 of its length, maximum width of foliation
|
||
<quantity id="BB46CF83DE17FFA3F4DCFA687C5BB7DF" box="[1017,1121,1482,1507]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="3" pageNumber="598" unit="mm" value="0.55">0.55 mm</quantity>
|
||
; foliation on S9 of the same width along the entire margin,
|
||
<quantity id="BB46CF83DE17FFA3F6D7FA4C7A6CB43B" box="[498,598,1518,1543]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="3" pageNumber="598" unit="mm" value="0.45">0.45 mm</quantity>
|
||
wide, inferior margin smooth.
|
||
<emphasis id="4ECABE74DE17FFA3F491FA4D7CBDB43A" box="[948,1159,1518,1543]" italics="true" pageId="3" pageNumber="598">Accesory genitalia</emphasis>
|
||
(
|
||
<figureCitation id="E4857EE3DE17FFA3F3B3FA4C7CE1B43A" box="[1174,1243,1518,1543]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1854,1877]" captionTargetBox="[251,1269,199,1822]" captionTargetId="figure@2.[250,1279,199,1834]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Phyllogomphoides enriquei sp. nov., holotype ♂. 1) Head and thorax, lateral view (head turned up to the left); 2) S 8 ¯ 10 and caudal appendages, left lateral view, showing inferior margins of S 8 - 9 foliate and scalloped; 3) Accessory genitalia, ventral view: alb, anterior lobe of anterior hamule; Ah, anterior hamuli; Alm, anterior lamina; dt, denticles; fl, flagella; msp, middle spine; n, notch; plb, posterior lobe of anterior hamule; psp, posterior spine of anterior lobe of anterior hamule; V 1, V 4, segments 1 and 4 of vesica spermalis; 4) S 9 ¯ 10 and caudal appendages, dorsal view. Ce, cercus; dt, dorsal tooth; Ep, epiproct." httpUri="https://zenodo.org/record/855679/files/figure.png" pageId="3" pageNumber="598">Fig. 3</figureCitation>
|
||
): anterior lamina thick, entire, anterior margin concave and undulate; in ventral view anterior hamuli bilobed, anterior lobe tumid anteriorly, ending in a strong spine posteriorly, followed by a wide, deep cleft between anterior and posterior lobes (
|
||
<figureCitation id="E4857EE3DE17FFA3F7BAF9F878D9B44E" box="[159,227,1626,1651]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1854,1877]" captionTargetBox="[251,1269,199,1822]" captionTargetId="figure@2.[250,1279,199,1834]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Phyllogomphoides enriquei sp. nov., holotype ♂. 1) Head and thorax, lateral view (head turned up to the left); 2) S 8 ¯ 10 and caudal appendages, left lateral view, showing inferior margins of S 8 - 9 foliate and scalloped; 3) Accessory genitalia, ventral view: alb, anterior lobe of anterior hamule; Ah, anterior hamuli; Alm, anterior lamina; dt, denticles; fl, flagella; msp, middle spine; n, notch; plb, posterior lobe of anterior hamule; psp, posterior spine of anterior lobe of anterior hamule; V 1, V 4, segments 1 and 4 of vesica spermalis; 4) S 9 ¯ 10 and caudal appendages, dorsal view. Ce, cercus; dt, dorsal tooth; Ep, epiproct." httpUri="https://zenodo.org/record/855679/files/figure.png" pageId="3" pageNumber="598">Fig. 3</figureCitation>
|
||
); posterior lobe wide, distal end widely rounded, with a small notch between two small denticles on apical third of mesial margin, external surface moderately excavated inferiorly; posterior hamule subcylindrical, with a large, sharply pointed apical spine directed mesally, with abundant long and short whitish hairs. Vesica spermalis:
|
||
<date id="080044A6DE17FFA3F7B2F9657883B4E2" box="[151,185,1735,1759]" pageId="3" pageNumber="598">V1</date>
|
||
bifid with a small tooth on middle (
|
||
<figureCitation id="E4857EE3DE17FFA3F56BF9647AABB4E2" box="[590,657,1734,1759]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1854,1877]" captionTargetBox="[251,1269,199,1822]" captionTargetId="figure@2.[250,1279,199,1834]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Phyllogomphoides enriquei sp. nov., holotype ♂. 1) Head and thorax, lateral view (head turned up to the left); 2) S 8 ¯ 10 and caudal appendages, left lateral view, showing inferior margins of S 8 - 9 foliate and scalloped; 3) Accessory genitalia, ventral view: alb, anterior lobe of anterior hamule; Ah, anterior hamuli; Alm, anterior lamina; dt, denticles; fl, flagella; msp, middle spine; n, notch; plb, posterior lobe of anterior hamule; psp, posterior spine of anterior lobe of anterior hamule; V 1, V 4, segments 1 and 4 of vesica spermalis; 4) S 9 ¯ 10 and caudal appendages, dorsal view. Ce, cercus; dt, dorsal tooth; Ep, epiproct." httpUri="https://zenodo.org/record/855679/files/figure.png" pageId="3" pageNumber="598">Fig. 3</figureCitation>
|
||
),
|
||
<date id="080044A6DE17FFA3F58DF9657AEAB4E2" box="[680,720,1735,1759]" pageId="3" pageNumber="598">V2</date>
|
||
¯3 of usual
|
||
<typeStatus id="A305DCC4DE17FFA3F475F9657BB9B4E2" box="[848,899,1735,1759]" pageId="3" pageNumber="598">type</typeStatus>
|
||
,
|
||
<date id="080044A6DE17FFA3F4ABF9657B8AB4E2" box="[910,944,1735,1759]" pageId="3" pageNumber="598">V4</date>
|
||
with 2 relatively short flagella not reaching posterior margin of
|
||
<date id="080044A6DE17FFA3F65AF9497992B53E" box="[383,424,1771,1795]" pageId="3" pageNumber="598">V1.</date>
|
||
<emphasis id="4ECABE74DE17FFA3F697F9487A4DB53E" box="[434,631,1770,1795]" italics="true" pageId="3" pageNumber="598">Anal appendages</emphasis>
|
||
: Cercus in dorsal view semicircular-forcipate (
|
||
<figureCitation id="E4857EE3DE17FFA3F3BCF9487CDBB53E" box="[1177,1249,1770,1795]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1854,1877]" captionTargetBox="[251,1269,199,1822]" captionTargetId="figure@2.[250,1279,199,1834]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Phyllogomphoides enriquei sp. nov., holotype ♂. 1) Head and thorax, lateral view (head turned up to the left); 2) S 8 ¯ 10 and caudal appendages, left lateral view, showing inferior margins of S 8 - 9 foliate and scalloped; 3) Accessory genitalia, ventral view: alb, anterior lobe of anterior hamule; Ah, anterior hamuli; Alm, anterior lamina; dt, denticles; fl, flagella; msp, middle spine; n, notch; plb, posterior lobe of anterior hamule; psp, posterior spine of anterior lobe of anterior hamule; V 1, V 4, segments 1 and 4 of vesica spermalis; 4) S 9 ¯ 10 and caudal appendages, dorsal view. Ce, cercus; dt, dorsal tooth; Ep, epiproct." httpUri="https://zenodo.org/record/855679/files/figure.png" pageId="3" pageNumber="598">Fig. 4</figureCitation>
|
||
), with abundant long setae, dorsally pale on apical 0.80 its length, basal 0.20, ventral surface, and apices mostly black, a dorsal tooth (
|
||
<quantity id="BB46CF83DE17FFA3F7FAF890797EB577" box="[223,324,1842,1867]" metricMagnitude="-4" metricUnit="m" metricValue="4.6" pageId="3" pageNumber="598" unit="mm" value="0.46">0.46 mm</quantity>
|
||
long) at basal 0.70 directed medially, tips flattened laterally and produced dorsally and ventrally; in ventrolateral view with a short (
|
||
<quantity id="BB46CF83DE17FFA3F525F8F57A5CB553" box="[512,614,1879,1903]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="3" pageNumber="598" unit="mm" value="0.13">0.13 mm</quantity>
|
||
long), sub-basal spine at basal 0.20; in lateral view (
|
||
<figureCitation id="E4857EE3DE17FFA3F399F8F47CC5B552" box="[1212,1279,1878,1903]" captionStart="FIGURES 5 – 6" captionStartId="3.[151,264,1035,1058]" captionTargetBox="[173,1436,193,1014]" captionTargetId="figure@3.[151,1436,193,1014]" captionTargetPageId="3" captionText="FIGURES 5 – 6. Details of Phyllogomphoides spp. 5) Phyllogomphoides enriquei sp. nov., holotype ♂. Caudal appendages, left lateral view. Ce, cercus; dt, dorsal tooth; dsp, dorsal spine; Ep, epiproct; vsp, ventral spine. 6) Phyllogomphoides luisi, ♂ (Xochitepec). Accessory genitalia, ventral view." httpUri="https://zenodo.org/record/855681/files/figure.png" pageId="3" pageNumber="598">Fig. 5</figureCitation>
|
||
) tip of cercus produced dorsally in a large, wide spine, ventrally produced into an acuminate spine,
|
||
<quantity id="BB46CF83DE17FFA3F3A3F8D87CD4B5AF" box="[1158,1262,1914,1939]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="598" unit="mm" value="0.25">0.25 mm</quantity>
|
||
long. Epiproct brown, V-shaped, in lateral view strongly up-curved (
|
||
<figureCitation id="E4857EE3DE17FFA3F5CEF83C7B14B58A" box="[747,814,1950,1975]" captionStart="FIGURES 5 – 6" captionStartId="3.[151,264,1035,1058]" captionTargetBox="[173,1436,193,1014]" captionTargetId="figure@3.[151,1436,193,1014]" captionTargetPageId="3" captionText="FIGURES 5 – 6. Details of Phyllogomphoides spp. 5) Phyllogomphoides enriquei sp. nov., holotype ♂. Caudal appendages, left lateral view. Ce, cercus; dt, dorsal tooth; dsp, dorsal spine; Ep, epiproct; vsp, ventral spine. 6) Phyllogomphoides luisi, ♂ (Xochitepec). Accessory genitalia, ventral view." httpUri="https://zenodo.org/record/855681/files/figure.png" pageId="3" pageNumber="598">Fig. 5</figureCitation>
|
||
), in dorsal view (
|
||
<figureCitation id="E4857EE3DE17FFA3F4D5F83C7C09B58A" box="[1008,1075,1950,1975]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1854,1877]" captionTargetBox="[251,1269,199,1822]" captionTargetId="figure@2.[250,1279,199,1834]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Phyllogomphoides enriquei sp. nov., holotype ♂. 1) Head and thorax, lateral view (head turned up to the left); 2) S 8 ¯ 10 and caudal appendages, left lateral view, showing inferior margins of S 8 - 9 foliate and scalloped; 3) Accessory genitalia, ventral view: alb, anterior lobe of anterior hamule; Ah, anterior hamuli; Alm, anterior lamina; dt, denticles; fl, flagella; msp, middle spine; n, notch; plb, posterior lobe of anterior hamule; psp, posterior spine of anterior lobe of anterior hamule; V 1, V 4, segments 1 and 4 of vesica spermalis; 4) S 9 ¯ 10 and caudal appendages, dorsal view. Ce, cercus; dt, dorsal tooth; Ep, epiproct." httpUri="https://zenodo.org/record/855679/files/figure.png" pageId="3" pageNumber="598">Fig. 4</figureCitation>
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||
) tips sharp, separated from each other by a distance of
|
||
<quantity id="BB46CF83DE17FFA3F6AAF86179CCB5E7" box="[399,502,1987,2011]" metricMagnitude="-4" metricUnit="m" metricValue="7.3" pageId="3" pageNumber="598" unit="mm" value="0.73">0.73 mm</quantity>
|
||
; in ventral and dorsal views with few, short, whitish hairs.
|
||
</paragraph>
|
||
<paragraph id="7C016266DE10FFA4F7E2FF3A7CBEB2E9" blockId="4.[151,1436,151,213]" pageId="4" pageNumber="599">
|
||
<emphasis id="4ECABE74DE10FFA4F7E2FF3A79DDB28D" bold="true" box="[199,487,151,176]" pageId="4" pageNumber="599">Measurements (in mm):</emphasis>
|
||
TL, 61.4; Ab, 46; MWh, 7.7; posterior border of occiput, 2; occiput length, 0.7; Hf, 5.6; FW, 36.5; HW, 34.5; FW width (at level of nodus), 8, HW width, 10; pt, 4.7; cerci, 3.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |