251 lines
16 KiB
XML
251 lines
16 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1147.93824" ID-Pensoft-Pub="1313-2970-1147-1" ID-Pensoft-UUID="FF2B6B317B505F9EA0E1000BDCD16CE7" ID-ZooBank="9BB9881B0076473D8E53155D37CA1F50" ModsDocID="1313-2970-1147-1" checkinTime="1676378625502" checkinUser="pensoft" docAuthor="Pall-Gergely, Barna, Hunyadi, Andras, Vermeulen, Jaap J., Grego, Jozef, Sutcharit, Chirasak, Reischuetz, Alexander, Dumrongrojwattana, Pongrat, Botta-Dukat, Zoltan, Oerstan, Aydin, Fekete, Judit & Jochum, Adrienne" docDate="2023" docId="94F9FF489F545B339FA83931DBBA34AC" docLanguage="en" docName="ZooKeys 1147: 1-177" docOrigin="ZooKeys 1147" docPubDate="2023-02-13" docSource="http://dx.doi.org/10.3897/zookeys.1147.93824" docTitle="Angustopila papaver Pall-Gergely & Hunyadi 2023, sp. nov." docType="treatment" docUuid="31264F15-F05F-4C41-AA05-6B6C13D46B12" docUuidSource="ZooBank" docVersion="1" id="FF2B6B317B505F9EA0E1000BDCD16CE7" lastPageNumber="1" masterDocId="FF2B6B317B505F9EA0E1000BDCD16CE7" masterDocTitle="Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae)" masterLastPageNumber="177" masterPageNumber="1" pageNumber="1" updateTime="1676378625502" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Pall-Gergely, Barna</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6167-7221</mods:nameIdentifier>
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<mods:affiliation>Centre for Agricultural Research, Plant Protection Institute, Eoetvoes Lorand Research Network, Herman Otto ut 15, H- 1022 Budapest, Hungary</mods:affiliation>
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<mods:nameIdentifier type="email">pallgergely2@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hunyadi, Andras</mods:namePart>
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<mods:affiliation>Adria setany 10 G 2 / 5., H- 1148 Budapest, Hungary</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Vermeulen, Jaap J.</mods:namePart>
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<mods:affiliation>JK Art and Science, Lauwerbes 8, 2318 AT Leiden, Netherlands</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Grego, Jozef</mods:namePart>
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<mods:affiliation>Horna Micina 219, SK- 97401 Banska Bystrica, Slovakia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation>Animal Systematic Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Reischuetz, Alexander</mods:namePart>
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<mods:affiliation>Puechhaimgasse 52, A- 3580 Horn, Austria</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Dumrongrojwattana, Pongrat</mods:namePart>
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<mods:affiliation>Department of Biology, Faculty of Science, Burapha University, 169 Longhardbangsaen Road, Muang District, Chonburi, 20131, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Botta-Dukat, Zoltan</mods:namePart>
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<mods:affiliation>Centre for Ecological Research, Institute of Ecology and Botany, Alkotmany 2 - 4, H- 2600, Vacratot, Hungary</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Oerstan, Aydin</mods:namePart>
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<mods:affiliation>12501 Milestone Manor Lane, Germantown, Maryland, 20876, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fekete, Judit</mods:namePart>
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<mods:affiliation>University of Pannonia, Centre of Natural Science, Research Group of Limnology, Egyetem u. 10, H- 8200 Veszprem, Hungary & Centre for Ecological Research, Institute of Aquatic Ecology, Department of Tisza Research, 18 / c Bem square, H- 4026 Debrecen, Hungary</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jochum, Adrienne</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6624-6412</mods:nameIdentifier>
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<mods:affiliation>Naturhistorisches Museum der Burgergemeinde Bern, CH- 3005 Bern, Switzerland & Institute of Ecology and Evolution, University of Bern, CH- 3012 Bern, Switzerland & Senckenberg Forschungsinstitut und Naturmuseum, 60325 Frankfurt am Main, Germany</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-02-13</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1147</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>177</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1147.93824</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1147.93824</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1147-1</mods:identifier>
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<mods:identifier type="ZooBank">9BB9881B0076473D8E53155D37CA1F50</mods:identifier>
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<mods:identifier type="Pensoft-UUID">FF2B6B317B505F9EA0E1000BDCD16CE7</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:zoobank.org:act:31264F15-F05F-4C41-AA05-6B6C13D46B12" httpUri="http://treatment.plazi.org/id/94F9FF489F545B339FA83931DBBA34AC" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="https://zoobank.org/31264F15-F05F-4C41-AA05-6B6C13D46B12" authority="Páll-Gergely & Hunyadi" authorityName="Pall-Gergely & Hunyadi" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Angustopila papaver" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="papaver" status="sp. nov.">
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Angustopila papaver
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<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
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& Hunyadi
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</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">
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<figureCitation captionStart="Figure 73" captionStartId="F73" captionText="Figure 73. Angustopila papaver Pall-Gergely & Hunyadi, sp. nov. A, C-I holotype (HNHM 105297) B paratype. Apertural (A, B), lateral (C), apical (D) and ventral (E) sides of the shell; sculpture on the protoconch (F); aperture (G); ventral (H) and frontal (I) surface of the body whorl." figureDoi="10.3897/zookeys.1147.93824.figure73" httpUri="https://binary.pensoft.net/fig/808882" pageId="0" pageNumber="1">Fig. 73</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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: Vietnam • 1 empty shell (H: 0.66 mm, D: 0.66 mm); Thanh
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<normalizedToken originalValue="Hóa">Hoa</normalizedToken>
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Province,
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<normalizedToken originalValue="Như">Như</normalizedToken>
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Thanh District, Hải
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<normalizedToken originalValue="Vân">Van</normalizedToken>
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, Hang
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<normalizedToken originalValue="Lò">Lo</normalizedToken>
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Cao
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<normalizedToken originalValue="Kháng">Khang</normalizedToken>
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Chiến, vicinity of the cave (locality code: 2020/41);
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<geoCoordinate degrees="19" direction="north" minutes="37.08" orientation="latitude" precision="9" value="19.618">19°37.08'N</geoCoordinate>
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,
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<geoCoordinate degrees="105" direction="east" minutes="34.63" orientation="longitude" precision="9" value="105.57716">105°34.63'E</geoCoordinate>
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; 20 m a.s.l.; 14 Feb. 2020; A. Hunyadi leg.; HNHM 105297.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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: Vietnam • 1 figured shell + 39 shells + 3 live-collected specimens covered in mud; same data as for holotype; coll. HA.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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A small
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<taxonomicName authorityName="Jochum, Slapnik & Pall-Gergely" authorityYear="2014" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Angustopila</emphasis>
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</taxonomicName>
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species with a conical to conical-globular shell, a shouldered body whorl, a strong parietal tooth reaching the peristome edge, and an upper palatal tooth of ca. 1/2 size.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Shell small for the genus, shell slightly higher than wide or rarely as high as wide; off-white, translucent when fresh, conical to conical-globular; body whorl widest in apertural view; protoconch consists of 1-1.25 whorls with spiral striation preceding the first teleoconch whorl; teleoconch with very weak, regularly spaced radial ribs crossed by stronger spiral striae (ca. 12-15 on body whorl from apertural view); whorls ca. 3.5, slightly shouldered,
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<normalizedToken originalValue="“pushed”">"pushed"</normalizedToken>
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from baso-lateral direction from apertural view; aperture very slightly oblique to shell axis from lateral view; umbilicus moderately wide; aperture pear-shaped, sinulus narrow and strongly isolated due to the near joining of the strong parietal and upper palatal teeth; peristome expanded, not reflected; parietal callus slightly protruding, slightly detached from penultimate whorl (aperture not adnate); parietal tooth elevated, strong, short, reaches parietal callus, slightly curved in direction of palatal tooth; upper palatal tooth ca. 1/2 as high as parietal tooth, situated near peristome edge, just opposite of parietal tooth.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Measurements (in mm).</paragraph>
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<paragraph pageId="0" pageNumber="1">
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H = 0.66-0.75, D = 0.64-0.75, H/D*100 = 100-112.5 (
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<emphasis italics="true" pageId="0" pageNumber="1">n</emphasis>
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= 7), RUD = 26.6-29.2 (
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<emphasis italics="true" pageId="0" pageNumber="1">n</emphasis>
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= 3).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
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<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Similar only to
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<taxonomicName lsidName="A. coprologos" pageId="0" pageNumber="1" rank="species" species="coprologos">
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<emphasis italics="true" pageId="0" pageNumber="1">A. coprologos</emphasis>
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</taxonomicName>
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and
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<taxonomicName lsidName="A. psammion" pageId="0" pageNumber="1" rank="species" species="psammion">
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<emphasis italics="true" pageId="0" pageNumber="1">A. psammion</emphasis>
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</taxonomicName>
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in size. This new species can be distinguished from all congeners based on the combination of the very small, conical to conical-globular shell and the two apertural denticles.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="etymology">
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<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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The tiny shells of this species with approximately equal shell height and width resemble poppy (
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<taxonomicName class="Magnoliopsida" family="Papaveraceae" genus="Papaver" kingdom="Plantae" lsidName="Papaver" order="Ranunculales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Papaver</emphasis>
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</taxonomicName>
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) seeds. The specific epithet is to be used as a noun in apposition.
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</paragraph>
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<caption doi="10.3897/zookeys.1147.93824.figure73" httpUri="https://binary.pensoft.net/fig/808882" pageId="0" pageNumber="1" start="Figure 73" startId="F73">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 73.</emphasis>
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<taxonomicName authorityName="Pall-Gergely & Hunyadi" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila papaver" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="papaver">
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<emphasis italics="true" pageId="0" pageNumber="1">Angustopila papaver</emphasis>
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</taxonomicName>
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<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
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& Hunyadi, sp. nov.
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<emphasis bold="true" pageId="0" pageNumber="1">A, C-I</emphasis>
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holotype (HNHM 105297)
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<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
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paratype. Apertural (
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<emphasis bold="true" pageId="0" pageNumber="1">A, B</emphasis>
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), lateral (
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<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
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), apical (
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<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
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) and ventral (
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<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
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) sides of the shell; sculpture on the protoconch (
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<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
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); aperture (
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<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
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); ventral (
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<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
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) and frontal (
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<emphasis bold="true" pageId="0" pageNumber="1">I</emphasis>
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) surface of the body whorl.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Pall-Gergely & Hunyadi" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila papaver" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="papaver">
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<emphasis italics="true" pageId="0" pageNumber="1">Angustopila papaver</emphasis>
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</taxonomicName>
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sp. nov. is known from the type locality only in Thanh
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<normalizedToken originalValue="Hóa">Hoa</normalizedToken>
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Province of northern Vietnam (Fig.
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<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Distribution of Angustopila species." figureDoi="10.3897/zookeys.1147.93824.figure48" httpUri="https://binary.pensoft.net/fig/808857" pageId="0" pageNumber="1">48</figureCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |