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<document id="066285275D6D40775BCD0F1DD25AF310" ID-CLB-Dataset="55535" ID-DOI="10.5281/zenodo.5193661" ID-GBIF-Dataset="c2346a92-3813-4b3d-b139-f3b149b50f4a" ID-ISSN="0253-116X" ID-Zenodo-Dep="5193661" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628802610787" checkinUser="felipe" docAuthor="Jaeger, Bernd" docDate="2015" docId="5C580E35A81CF118FF5EFB8BFF289D6F" docLanguage="en" docName="LinzerbiolBeitr.47.2.1361-1396.pdf" docOrigin="Linzer biologische Beiträge 47 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Anthracus siamensis Jaeger 2015, nov.sp." docType="treatment" docVersion="11" lastPageNumber="1376" masterDocId="A061764DA811F117FFECFFFCFFF39965" masterDocTitle="Revision of the maculate species of the Anthracus annamensis group from the East Palaearctic and Oriental Regions. Part 2. A redescription of Anthracus nesophilus (ANDREWES, 1936) and six new species from Nepal, India and SE Asia (Coleoptera, Carabidae, Harpalini, Stenolophina)" masterLastPageNumber="1396" masterPageNumber="1361" pageNumber="1374" updateTime="1699052547504" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="867DECB53AB30AA53C44085E3D8C05C9">
<mods:title id="0863C90591A0286215CFD76387080CB6">Revision of the maculate species of the Anthracus annamensis group from the East Palaearctic and Oriental Regions. Part 2. A redescription of Anthracus nesophilus (ANDREWES, 1936) and six new species from Nepal, India and SE Asia (Coleoptera, Carabidae, Harpalini, Stenolophina)</mods:title>
</mods:titleInfo>
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<mods:namePart id="25D982937D49495B288DA2135C9FBA91">Jaeger, Bernd</mods:namePart>
</mods:name>
<mods:typeOfResource id="C2988B1D5EA27E181F84935753E40B93">text</mods:typeOfResource>
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<mods:title id="168280E186AA1D3CA1F0B9C4427293D4">Linzer biologische Beiträge</mods:title>
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<mods:part id="FD4F8A8FBE65B073C2F2CBD008B4D721">
<mods:date id="7D81CAA5C5770EEF4F38DDD0B2A6BA43">2015</mods:date>
<mods:detail id="3B35226FC9965D1F503F13E215B209E9" type="pubDate">
<mods:number id="F835ED0847CDF3A8A4B866A8F312E450">2015-12-30</mods:number>
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<mods:detail id="57438602CE0647C32C4C794AAC51BC4C" type="volume">
<mods:number id="25588C67A634C80A2884C92ED648543B">47</mods:number>
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<mods:classification id="618CA2962A813A842360240E934DF6D1">journal article</mods:classification>
<mods:identifier id="E9485D16C17E9852326C8EF171886BBC" type="CLB-Dataset">55535</mods:identifier>
<mods:identifier id="DC83F9E1104C264227209E79E6BD4201" type="DOI">10.5281/zenodo.5193661</mods:identifier>
<mods:identifier id="E0DDD6B87795B99EE4806D8636D05802" type="GBIF-Dataset">c2346a92-3813-4b3d-b139-f3b149b50f4a</mods:identifier>
<mods:identifier id="FDF69DCE02F566CA3024B6D92D000B25" type="ISSN">0253-116X</mods:identifier>
<mods:identifier id="643576E0D5E3CE3ABC2191EE3DB79FB7" type="Zenodo-Dep">5193661</mods:identifier>
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<treatment id="5C580E35A81CF118FF5EFB8BFF289D6F" ID-DOI="http://doi.org/10.5281/zenodo.6546860" ID-GBIF-Taxon="195305591" ID-Zenodo-Dep="6546860" LSID="urn:lsid:plazi:treatment:5C580E35A81CF118FF5EFB8BFF289D6F" httpUri="http://treatment.plazi.org/id/5C580E35A81CF118FF5EFB8BFF289D6F" lastPageId="15" lastPageNumber="1376" pageId="13" pageNumber="1374">
<subSubSection id="9CEBECA8A81CF11AFF5EFB8BFE2D9DE9" box="[178,478,1143,1165]" pageId="13" pageNumber="1374" type="nomenclature">
<paragraph id="D44EBF23A81CF11AFF5EFB8BFE2D9DE9" blockId="13.[178,793,1143,1165]" box="[178,478,1143,1165]" pageId="13" pageNumber="1374">
<heading id="8F06084FA81CF11AFF5EFB8BFE2D9DE9" box="[178,478,1143,1165]" fontSize="8" level="2" pageId="13" pageNumber="1374" reason="2">
<emphasis id="E6856331A81CF11AFF5EFB8BFE2D9DE9" bold="true" box="[178,478,1143,1165]" pageId="13" pageNumber="1374">
<taxonomicName id="13F1C4A0A81CF11AFF5EFB8BFE789DE9" ID-CoL="67GWG" authority="Jaeger, 2015" authorityName="Jaeger" authorityYear="2015" box="[178,395,1143,1165]" class="Insecta" family="Carabidae" genus="Anthracus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1374" phylum="Arthropoda" rank="species" species="siamensis" status="sp. nov.">
<emphasis id="E6856331A81CF11AFF5EFB8BFE789DE9" bold="true" box="[178,395,1143,1165]" italics="true" pageId="13" pageNumber="1374">Anthracus siamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FDB6DE4AA81CF11AFE7DFB8BFE2D9DE9" box="[401,478,1143,1164]" pageId="13" pageNumber="1374" rank="species">nov.sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="9CEBECA8A81CF11AFE08FB8BFCEA9DE8" box="[484,793,1143,1165]" pageId="13" pageNumber="1374" type="description">
<paragraph id="D44EBF23A81CF11AFE08FB8BFCEA9DE8" blockId="13.[178,793,1143,1165]" box="[484,793,1143,1165]" pageId="13" pageNumber="1374">
<heading id="8F06084FA81CF11AFE08FB8BFCEA9DE8" box="[484,793,1143,1165]" fontSize="8" level="2" pageId="13" pageNumber="1374" reason="2">
<emphasis id="E6856331A81CF11AFE08FB8BFCEA9DE8" bold="true" box="[484,793,1143,1165]" pageId="13" pageNumber="1374">
(
<figureCitation id="4CCAA3A6A81CF11AFE07FB8BFDAF9DE9" box="[491,604,1143,1165]" captionStart="Figs 21-23" captionStartId="24.[178,218,1496,1518]" captionTargetBox="[178,1085,212,1573]" captionTargetId="figure-0@24.[178,1085,212,1573]" captionTargetPageId="24" captionText="Figs 21-23: Anthracus siamensis nov.sp. Habitus, head and pronotum. (21) HT, (22, 23) PT, Laos, Pakkading." figureDoi="http://doi.org/10.5281/zenodo.5193675" httpUri="https://zenodo.org/record/5193675/files/figure.png" pageId="13" pageNumber="1374">Figs 21-23</figureCitation>
,
<figureCitation id="4CCAA3A6A81CF11AFD8BFB8BFD579DE9" box="[615,676,1143,1164]" captionStart="Figs 62-67" captionStartId="31.[178,218,1520,1542]" captionTargetBox="[150,1112,174,1495]" captionTargetId="figure-0@31.[150,1112,174,1617]" captionTargetPageId="31" captionText="Figs 62-67: Median lobe of aedoeagus, lateral aspect. Anthracus javaensis nov.sp. (62) HT and A. siamensis nov.sp. (63, 64, 67) PT, Laos, Pakkading, (65) HT, (66) Thailand, Ban Khoun." figureDoi="http://doi.org/10.5281/zenodo.6546849" httpUri="https://zenodo.org/record/6546849/files/figure.png" pageId="13" pageNumber="1374">63-67</figureCitation>
,
<figureCitation id="4CCAA3A6A81CF11AFD43FB8BFD1F9DE9" box="[687,748,1143,1164]" captionStart="Figs 68-73" captionStartId="32.[178,218,1420,1442]" captionTargetBox="[150,1112,159,1602]" captionTargetId="figure-0@32.[150,1112,159,1602]" captionTargetPageId="32" captionText="Figs 68-73: Median lobe of aedoeagus, dorsal aspect. Anthracus javaensis nov.sp. (68) HT and A. siamensis nov.sp. (69) PT, Thailand, Umphang, (70, 73) PT, Laos, Pakkading, (71) HT, (72) Thailand, Ban Khoun." figureDoi="http://doi.org/10.5281/zenodo.5194872" httpUri="https://zenodo.org/record/5194872/files/figure.png" pageId="13" pageNumber="1374">69-73</figureCitation>
,
<figureCitation id="4CCAA3A6A81CF11AFD1BFB8BFCE19DE8" box="[759,786,1143,1165]" captionStart="Fig 80" captionStartId="34.[178,210,909,931]" captionTargetBox="[266,997,212,896]" captionTargetId="figure-65@34.[266,997,212,896]" captionTargetPageId="34" captionText="Fig 80: Distribution: A. nesophilus (ANDREWES) (triangles), A. javaensis nov.sp. (white star), A. sumatraensis nov.sp. (square) A. philippinensis nov.sp. (circle), A. siamensis nov.sp. (black star)." figureDoi="http://doi.org/10.5281/zenodo.5193681" httpUri="https://zenodo.org/record/5193681/files/figure.png" pageId="13" pageNumber="1374">80</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="9CEBECA8A81CF11AFF5EFB58FCF69F2F" pageId="13" pageNumber="1374" type="materials_examined">
<paragraph id="D44EBF23A81CF11AFF5EFB58FD469CAF" blockId="13.[178,1086,1188,1610]" pageId="13" pageNumber="1374">
Type material:
<materialsCitation id="6499B57EA81CF11AFE63FB58FBF69D82" ID-GBIF-Occurrence="3766781316" collectingDate="1991-04-26" collectingDateMax="1991-05-06" collectingDateMin="1991-04-26" collectionCode="NHMB" collectorName="D. Kral &amp; V. Kuban" country="Thailand'" elevation="500" latitude="16.066668" location="UMPHANG" longLatPrecision="1283" longitude="98.88333" pageId="13" pageNumber="1374" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="0B4A0181A81CF11AFE63FB58FE109DDF" box="[399,483,1188,1210]" pageId="13" pageNumber="1374" type="holotype">Holotype</typeStatus>
:
<specimenCount id="C2F774AAA81CF11AFE03FB58FDF19DDC" box="[495,514,1188,1209]" pageId="13" pageNumber="1374" type="male">♂</specimenCount>
(
<collectionCode id="B2E027E6A81CF11AFDF8FB58FDA89DDF" box="[532,603,1188,1210]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="13" pageNumber="1374">NHMB</collectionCode>
) labelled &quot;THAI,
<date id="A04F99E3A81CF11AFCF8FB58FC599DDF" box="[788,938,1188,1210]" pageId="13" pageNumber="1374" value="1991-04-26" valueMax="1991-05-06" valueMin="1991-04-26">
<collectingDate id="B00B600BA81CF11AFCF8FB58FC599DDF" box="[788,938,1188,1210]" pageId="13" pageNumber="1374" value="1991-04-26" valueMax="1991-05-06" valueMin="1991-04-26">26.IV.-6.V.1991</collectingDate>
</date>
/
<location id="D12EE9F8A81CF11AFC26FB58FBCE9DDC" LSID="urn:lsid:plazi:treatment:5C580E35A81CF118FF5EFB8BFF289D6F:D12EE9F8A81CF11AFC26FB58FBCE9DDC" box="[970,1085,1188,1209]" country="Thailand'" latitude="16.066668" longLatPrecision="1283" longitude="98.88333" name="UMPHANG" pageId="13" pageNumber="1374">UMPHANG</location>
<quantity id="130912C6A81CF11AFF5EFB46FF159DB5" box="[178,230,1210,1232]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="1374" unit="m" value="500.0">
<elevation id="5FDC5810A81CF11AFF5EFB46FF159DB5" box="[178,230,1210,1232]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="1374" unit="m" value="500.0">500m</elevation>
</quantity>
/
<geoCoordinate id="B1C5D9E4A81CF11AFF11FB47FEBB9DB4" box="[253,328,1211,1233]" degrees="16" direction="north" minutes="04" orientation="latitude" pageId="13" pageNumber="1374" precision="925" value="16.066668">16°04'N</geoCoordinate>
<geoCoordinate id="B1C5D9E4A81CF11AFEBCFB46FE6A9DB5" box="[336,409,1210,1232]" degrees="98" direction="east" minutes="53" orientation="longitude" pageId="13" pageNumber="1374" precision="925" value="98.88333">98°53'E</geoCoordinate>
/
<collectorName id="7904DAF5A81CF11AFE5CFB47FDE19DB5" box="[432,530,1211,1233]" pageId="13" pageNumber="1374">Vit Kubáň</collectorName>
leg.&quot;, &quot;
<collectingCountry id="ACE6FFB3A81CF11AFDB7FB47FD419DB4" box="[603,690,1211,1233]" pageId="13" pageNumber="1374">Thailand'</collectingCountry>
91/&quot;
<location id="D12EE9F8A81CF11AFD32FB47FC6C9DB4" LSID="urn:lsid:plazi:treatment:5C580E35A81CF118FF5EFB8BFF289D6F:D12EE9F8A81CF11AFD32FB47FC6C9DB4" box="[734,927,1211,1233]" country="Thailand'" latitude="16.066668" longLatPrecision="1283" longitude="98.88333" name="Thanon Thong Chai" pageId="13" pageNumber="1374">Thanon Thong Chai</location>
&quot; /
<collectorName id="7904DAF5A81CF11AFC51FB47FBF79DB4" box="[957,1028,1211,1233]" pageId="13" pageNumber="1374">D. Král</collectorName>
&amp;
<collectorName id="7904DAF5A81CF11AFBCBFB47FF1D9D83" pageId="13" pageNumber="1374">V. Kubáň</collectorName>
&quot; and &quot;HOLOTYPE ♂ /
<originalDetermination id="F7EF07DFA81CF11AFE3CFB2DFD3C9D82" box="[464,719,1233,1255]" pageId="13" pageNumber="1374">
<taxonomicName id="13F1C4A0A81CF11AFE3CFB2DFDDE9D82" authorityName="Motschoulsky" authorityYear="1850" box="[464,557,1233,1255]" class="Insecta" family="Carabidae" genus="Anthracus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1374" phylum="Arthropoda" rank="genus">Anthracus</taxonomicName>
/ siamensis
<taxonomicNameLabel id="FDB6DE4AA81CF11AFD72FB2EFD3C9D82" box="[670,719,1234,1255]" pageId="13" pageNumber="1374" rank="species">sp. n.</taxonomicNameLabel>
</originalDetermination>
/ des.
<collectorName id="7904DAF5A81CF11AFCE6FB2EFCAD9D82" box="[778,862,1234,1255]" pageId="13" pageNumber="1374">
B.
<bibRefCitation id="B060C2D2A81CF11AFCC8FB2EFCAD9D82" author="JAEGER B" box="[804,862,1234,1255]" pageId="13" pageNumber="1374" refId="ref12895" refString="JAEGER B. (2015): Two new maculate species of the Anthracus annamensis group from Nepal and adjacent areas (Insecta: Coleoptera: Carabidae: Harpalini: Stenolophina). Pp. 339 - 345. - In: HARTMANN M. &amp; J. WEIPERT (eds), Biodiversitat und Naturausstattung im Himalaya V. - Verein der Freunde und Forderer des Naturkundemuseums Erfurt e. V. Erfurt: 580 pp." type="book" year="2015">Jaeger</bibRefCitation>
</collectorName>
2015&quot; [red label].
</materialsCitation>
<materialsCitation id="6499B57EA81CF11AFF5EFB14FE7F9C4E" ID-GBIF-Occurrence="3766781328" collectionCode="NHMB" collectorName="D. Kral &amp; V. Kuban" country="Thailand'" elevation="1000" latitude="16.116667" location="UMPHANG river" longLatPrecision="1283" longitude="99.0" pageId="13" pageNumber="1374" specimenCount="1" typeStatus="paratype">
<typeStatus id="0B4A0181A81CF11AFF5EFB14FEF99D98" box="[178,266,1256,1277]" pageId="13" pageNumber="1374" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="C2F774AAA81CF11AFEC8FB14FEA69D98" box="[292,341,1256,1277]" pageId="13" pageNumber="1374">2♂♂</specimenCount>
(
<collectionCode id="B2E027E6A81CF11AFE88FB14FE5E9D9B" box="[356,429,1256,1278]" pageId="13" pageNumber="1374">NHMB</collectionCode>
,) with same labels as the holotype. 1♂ labelled &quot;THAI 28/4-6/5.91 /
<location id="D12EE9F8A81CF11AFF5EFB03FEA59C70" LSID="urn:lsid:plazi:treatment:5C580E35A81CF118FF5EFB8BFF289D6F:D12EE9F8A81CF11AFF5EFB03FEA59C70" box="[178,342,1279,1301]" country="Thailand'" latitude="16.116667" longLatPrecision="1283" longitude="99.0" name="UMPHANG river" pageId="13" pageNumber="1374">UMPHANG river</location>
/
<geoCoordinate id="B1C5D9E4A81CF11AFE84FB03FE409C70" box="[360,435,1279,1301]" degrees="16" direction="north" minutes="07" orientation="latitude" pageId="13" pageNumber="1374" precision="925" value="16.116667">16°07'N</geoCoordinate>
<geoCoordinate id="B1C5D9E4A81CF11AFE55FB03FDF19C71" box="[441,514,1279,1300]" degrees="99" direction="east" minutes="00" orientation="longitude" pageId="13" pageNumber="1374" precision="925" value="99.0">99°00'E</geoCoordinate>
/ lgt.
<collectorName id="7904DAF5A81CF11AFDD4FB03FD849C70" box="[568,631,1279,1301]" pageId="13" pageNumber="1374">D.Král</collectorName>
<quantity id="130912C6A81CF11AFD91FB03FD4D9C71" box="[637,702,1279,1300]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="1374" unit="m" value="1000.0">
<elevation id="5FDC5810A81CF11AFD91FB03FD4D9C71" box="[637,702,1279,1300]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="1374" unit="m" value="1000.0">1000m</elevation>
</quantity>
&quot; and &quot;
<collectingCountry id="ACE6FFB3A81CF11AFD15FB03FCA39C70" box="[761,848,1279,1301]" pageId="13" pageNumber="1374">Thailand'</collectingCountry>
91/&quot;
<location id="D12EE9F8A81CF11AFC95FB03FBC59C70" LSID="urn:lsid:plazi:treatment:5C580E35A81CF118FF5EFB8BFF289D6F:D12EE9F8A81CF11AFC95FB03FBC59C70" box="[889,1078,1279,1301]" country="Thailand'" latitude="16.116667" longLatPrecision="1283" longitude="99.0" name="Thanon Thong Chai" pageId="13" pageNumber="1374">Thanon Thong Chai</location>
&quot; /
<collectorName id="7904DAF5A81CF11AFF2CFAEAFEF69C4E" box="[192,261,1301,1323]" pageId="13" pageNumber="1374">D. Král</collectorName>
&amp;
<collectorName id="7904DAF5A81CF11AFECAFAEAFE729C4E" box="[294,385,1301,1323]" pageId="13" pageNumber="1374">V. Kubáň</collectorName>
&quot;
</materialsCitation>
.
<materialsCitation id="6499B57EA81CF11AFE7BFAEAFC659C24" ID-GBIF-Occurrence="3766781325" latitude="16.066668" longitude="98.88333" pageId="13" pageNumber="1374">
1♂ labelled &quot;THAI 28/4-6/5.91 / UMPHANG
<quantity id="130912C6A81CF11AFCA1FAE9FC729C4E" box="[845,897,1301,1323]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="1374" unit="m" value="500.0">500m</quantity>
/
<geoCoordinate id="B1C5D9E4A81CF11AFC7AFAE9FC2C9C4E" box="[918,991,1301,1323]" degrees="16" direction="north" minutes="04" orientation="latitude" pageId="13" pageNumber="1374" precision="925" value="16.066668">16 04'N</geoCoordinate>
<geoCoordinate id="B1C5D9E4A81CF11AFC0BFAE9FBDC9C4E" box="[999,1071,1301,1323]" degrees="98" direction="east" minutes="53" orientation="longitude" pageId="13" pageNumber="1374" precision="925" value="98.88333">98°53'E</geoCoordinate>
/ David Král lgt.&quot; and &quot;Thailand' 91/&quot;Thanon Thong Chai&quot; / D. Král &amp; V. Kubáň&quot;.
</materialsCitation>
<materialsCitation id="6499B57EA81CF11AFF5EFABFFEB39C15" ID-GBIF-Occurrence="3766781352" collectingDate="1986-11-07" collectionCode="MHNG" collectorName="P. Schwendinger" country="Thailand" elevation="320" location="Chiang Mai" municipality="Chiang Mai" pageId="13" pageNumber="1374" specimenCount="1" stateProvince="Chiang Mai" typeStatus="paratype">
<specimenCount id="C2F774AAA81CF11AFF5EFABFFF239C3D" box="[178,208,1347,1368]" pageId="13" pageNumber="1374">1♂</specimenCount>
(
<collectionCode id="B2E027E6A81CF11AFF0FFABFFED89C3C" box="[227,299,1347,1369]" pageId="13" pageNumber="1374">MHNG</collectionCode>
) labelled &quot;
<collectingCountry id="ACE6FFB3A81CF11AFE77FABFFDE19C3D" box="[411,530,1347,1368]" name="Thailand" pageId="13" pageNumber="1374">THAILAND</collectingCountry>
7.XI.86 / prov.
<collectingRegion id="163571C1A81CF11AFD2EFABFFCC59C3C" box="[706,822,1347,1369]" country="Thailand" name="Chiang Mai" pageId="13" pageNumber="1374">Chiang Mai</collectingRegion>
/
<collectingMunicipality id="342A2559A81CF11AFCBAFABFFC399C3C" box="[854,970,1347,1369]" pageId="13" pageNumber="1374">Chiang Mai</collectingMunicipality>
<quantity id="130912C6A81CF11AFC3AFABFFBF99C3D" box="[982,1034,1347,1368]" metricMagnitude="2" metricUnit="m" metricValue="3.2" pageId="13" pageNumber="1374" unit="m" value="320.0">
<elevation id="5FDC5810A81CF11AFC3AFABFFBF99C3D" box="[982,1034,1347,1368]" metricMagnitude="2" metricUnit="m" metricValue="3.2" pageId="13" pageNumber="1374" unit="m" value="320.0">320m</elevation>
</quantity>
/
<collectorName id="7904DAF5A81CF11AFBC6FABFFECA9C15" pageId="13" pageNumber="1374">P. Schwendinger</collectorName>
&quot;
</materialsCitation>
.
<materialsCitation id="6499B57EA81CF11AFEBDFAA6FC579CE3" ID-GBIF-Occurrence="3766781306" collectingDate="2001-06-01" collectingDateMax="2001-06-02" collectingDateMin="2001-06-01" collectionCode="NHMB, cJAE" collectorName="P. Pacholatko" country="LAOS-C" elevation="300" latitude="18.333334" location="PAKKADING" longLatPrecision="1275" longitude="104.0" municipality="PAKKADING" pageId="13" pageNumber="1374" specimenCount="2" specimenCount-female="1" stateProvince="BOLI KHAM XAI" typeStatus="paratype">
<specimenCount id="C2F774AAA81CF11AFEBDFAA6FE719C0A" box="[337,386,1370,1392]" pageId="13" pageNumber="1374">6♂♂</specimenCount>
,
<specimenCount id="C2F774AAA81CF11AFE7AFAA6FE479C11" box="[406,436,1370,1396]" pageId="13" pageNumber="1374" type="female">1♀</specimenCount>
(
<collectionCode id="B2E027E6A81CF11AFE24FAA6FDE39C0A" box="[456,528,1370,1391]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="13" pageNumber="1374">NHMB</collectionCode>
,
<collectionCode id="B2E027E6A81CF11AFDCDFAA6FDA79C15" box="[545,596,1370,1392]" pageId="13" pageNumber="1374">cJAE</collectionCode>
) labelled &quot;
<collectingCountry id="ACE6FFB3A81CF11AFD29FAA6FCEF9C0A" box="[709,796,1370,1391]" pageId="13" pageNumber="1374">LAOS-C</collectingCountry>
;
<collectingRegion id="163571C1A81CF11AFCC1FAA6FC189C0A" box="[813,1003,1370,1391]" pageId="13" pageNumber="1374">BOLI KHAM XAI</collectingRegion>
prov. /
<collectingMunicipality id="342A2559A81CF11AFF5EFA8CFECF9CE0" box="[178,316,1392,1413]" pageId="13" pageNumber="1374">PAKKADING</collectingMunicipality>
; ~
<quantity id="130912C6A81CF11AFEB8FA8CFE789CE0" box="[340,395,1392,1413]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="13" pageNumber="1374" unit="m" value="300.0">
<elevation id="5FDC5810A81CF11AFEB8FA8CFE789CE0" box="[340,395,1392,1413]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="13" pageNumber="1374" unit="m" value="300.0">300m</elevation>
</quantity>
; /
<geoCoordinate id="B1C5D9E4A81CF11AFE44FA8CFE009CE0" box="[424,499,1392,1413]" degrees="18" direction="north" minutes="20" orientation="latitude" pageId="13" pageNumber="1374" precision="925" value="18.333334">18°20'N</geoCoordinate>
<geoCoordinate id="B1C5D9E4A81CF11AFE10FA8CFDA19CE0" box="[508,594,1392,1413]" degrees="104" direction="east" minutes="00" orientation="longitude" pageId="13" pageNumber="1374" precision="925" value="104.0">104°00'E</geoCoordinate>
; /
<collectorName id="7904DAF5A81CF11AFD83FA8DFD039CE3" box="[623,752,1392,1414]" pageId="13" pageNumber="1374">P. Pacholátko</collectorName>
leg.:
<date id="A04F99E3A81CF11AFCC6FA8CFC689CE3" box="[810,923,1392,1414]" pageId="13" pageNumber="1374" value="2001-06-01" valueMax="2001-06-02" valueMin="2001-06-01">
<collectingDate id="B00B600BA81CF11AFCC6FA8CFC689CE3" box="[810,923,1392,1414]" pageId="13" pageNumber="1374" value="2001-06-01" valueMax="2001-06-02" valueMin="2001-06-01">1.-2.vi.2001</collectingDate>
</date>
&quot;
</materialsCitation>
.
<materialsCitation id="6499B57EA81CF11AFC5DFA8CFE2D9CD1" ID-GBIF-Occurrence="3766781341" collectingDate="2001-04-24" collectingDateMax="2001-04-29" collectingDateMin="2001-04-24" collectionCode="NHMB, cJAE" collectorName="Vit Kuban" country="Laos" elevation="200" latitude="18.116667" location="Ban Khoun Ngeun" longLatPrecision="1276" longitude="104.48333" pageId="13" pageNumber="1374" specimenCount="1" stateProvince="Khammouan prov." typeStatus="paratype">
<specimenCount id="C2F774AAA81CF11AFC5DFA8CFC119CE0" box="[945,994,1392,1413]" pageId="13" pageNumber="1374">2♂♂</specimenCount>
(
<collectionCode id="B2E027E6A81CF11AFC1EFA8CFBC99CE0" box="[1010,1082,1392,1413]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="13" pageNumber="1374">NHMB</collectionCode>
,
<collectionCode id="B2E027E6A81CF11AFF5EFA7BFF169CF8" box="[178,229,1415,1437]" pageId="13" pageNumber="1374">cJAE</collectionCode>
) labelled &quot;
<collectingCountry id="ACE6FFB3A81CF11AFEBCFA7BFE7C9CF9" box="[336,399,1415,1436]" name="Laos" pageId="13" pageNumber="1374">LAOS</collectingCountry>
,
<date id="A04F99E3A81CF11AFE72FA7BFDEE9CF8" box="[414,541,1415,1437]" pageId="13" pageNumber="1374" value="2001-04-24" valueMax="2001-04-29" valueMin="2001-04-24">
<collectingDate id="B00B600BA81CF11AFE72FA7BFDEE9CF8" box="[414,541,1415,1437]" pageId="13" pageNumber="1374" value="2001-04-24" valueMax="2001-04-29" valueMin="2001-04-24">24-29.iv.2001</collectingDate>
</date>
/
<collectingRegion id="163571C1A81CF11AFDD5FA7BFD199CF8" box="[569,746,1415,1437]" country="Laos" name="Khammouan" pageId="13" pageNumber="1374">Khammouan prov.</collectingRegion>
/
<geoCoordinate id="B1C5D9E4A81CF11AFCE9FA7BFCBC9CF9" box="[773,847,1415,1436]" degrees="18" direction="north" minutes="07" orientation="latitude" pageId="13" pageNumber="1374" precision="925" value="18.116667">18°07'N</geoCoordinate>
<geoCoordinate id="B1C5D9E4A81CF11AFCB6FA7BFC439CF9" box="[858,944,1415,1436]" degrees="104" direction="east" minutes="29" orientation="longitude" pageId="13" pageNumber="1374" precision="925" value="104.48333">104°29'E</geoCoordinate>
, /
<location id="D12EE9F8A81CF11AFC3CFA74FF1C9CD6" LSID="urn:lsid:plazi:treatment:5C580E35A81CF118FF5EFB8BFF289D6F:D12EE9F8A81CF11AFC3CFA74FF1C9CD6" country="Laos" latitude="18.116667" longLatPrecision="1276" longitude="104.48333" name="Ban Khoun Ngeun" pageId="13" pageNumber="1374" stateProvince="Khammouan prov.">Ban Khoun Ngeun</location>
/ -
<quantity id="130912C6A81CF11AFEE6FA62FEB29CD6" box="[266,321,1438,1459]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="1374" unit="m" value="200.0">
<elevation id="5FDC5810A81CF11AFEE6FA62FEB29CD6" box="[266,321,1438,1459]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="1374" unit="m" value="200.0">200m</elevation>
</quantity>
,
<collectorName id="7904DAF5A81CF11AFEA0FA62FE5F9CD6" box="[332,428,1438,1460]" pageId="13" pageNumber="1374">Vit Kubáň</collectorName>
leg.&quot;
</materialsCitation>
All
<typeStatus id="0B4A0181A81CF11AFDE6FA62FD929CD6" box="[522,609,1438,1459]" pageId="13" pageNumber="1374" type="paratype">paratypes</typeStatus>
additionally with my label &quot;
<typeStatus id="0B4A0181A81CF11AFC86FA62FC119CD6" box="[874,994,1438,1459]" pageId="13" pageNumber="1374" type="paratype">PARATYPE</typeStatus>
♂ or
<specimenCount id="C2F774AAA81CF11AFBF1FA5FFBC39CDD" box="[1053,1072,1443,1464]" pageId="13" pageNumber="1374" type="female">♀</specimenCount>
/
<taxonomicName id="13F1C4A0A81CF11AFF5EFA48FEFC9CAF" authorityName="Motschoulsky" authorityYear="1850" box="[178,271,1460,1482]" class="Insecta" family="Carabidae" genus="Anthracus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1374" phylum="Arthropoda" rank="genus">Anthracus</taxonomicName>
/ siamensis
<taxonomicNameLabel id="FDB6DE4AA81CF11AFE93FA49FE429CAF" box="[383,433,1461,1482]" pageId="13" pageNumber="1374" rank="species">sp. n.</taxonomicNameLabel>
/ des. B.
<bibRefCitation id="B060C2D2A81CF11AFDEAFA49FD869CAF" author="JAEGER B" box="[518,629,1460,1482]" pageId="13" pageNumber="1374" refId="ref12895" refString="JAEGER B. (2015): Two new maculate species of the Anthracus annamensis group from Nepal and adjacent areas (Insecta: Coleoptera: Carabidae: Harpalini: Stenolophina). Pp. 339 - 345. - In: HARTMANN M. &amp; J. WEIPERT (eds), Biodiversitat und Naturausstattung im Himalaya V. - Verein der Freunde und Forderer des Naturkundemuseums Erfurt e. V. Erfurt: 580 pp." type="book" year="2015">Jaeger 2015</bibRefCitation>
&quot; [red].
</paragraph>
<paragraph id="D44EBF23A81CF11AFF5EFA28FCF69F2F" blockId="13.[178,1086,1188,1610]" pageId="13" pageNumber="1374">
Additional Material examined:
<materialsCitation id="6499B57EA81CF11AFD47FA28FC139C8F" ID-GBIF-Occurrence="3766781340" box="[683,992,1492,1518]" collectingDate="1991-04-26" collectingDateMax="1991-05-06" collectingDateMin="1991-04-26" collectionCode="NHMB" collectorName="Vit Kuban" country="Thailand'" elevation="500" latitude="16.066668" location="UMPHANG" longLatPrecision="1283" longitude="98.88333" pageId="13" pageNumber="1374" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<specimenCount id="C2F774AAA81CF11AFD47FA28FD3D9C8B" box="[683,718,1492,1518]" pageId="13" pageNumber="1374" type="female">1♀</specimenCount>
with same data as the
<typeStatus id="0B4A0181A81CF11AFC7EFA28FC139C8F" box="[914,992,1492,1514]" pageId="13" pageNumber="1374" type="holotype">holotype</typeStatus>
</materialsCitation>
, and
<materialsCitation id="6499B57EA81CF11AFBE1FA28FD289F67" ID-GBIF-Occurrence="3766781333" pageId="13" pageNumber="1374" typeStatus="paratype">
<specimenCount id="C2F774AAA81CF11AFBE1FA28FBCE9C8B" box="[1037,1085,1492,1518]" pageId="13" pageNumber="1374" type="female">4♀♀</specimenCount>
with same data as the male
<typeStatus id="0B4A0181A81CF11AFE07FA11FDCD9F67" box="[491,574,1517,1538]" pageId="13" pageNumber="1374" type="paratype">paratypes</typeStatus>
from Pakkading
</materialsCitation>
. These females probably belong to
<taxonomicName id="13F1C4A0A81CF11AFBC6FA10FEC09F7F" authorityName="Jaeger" authorityYear="2015" class="Insecta" family="Carabidae" genus="Anthracus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1374" phylum="Arthropoda" rank="species" species="siamensis">
<emphasis id="E6856331A81CF11AFBC6FA10FEC09F7F" italics="true" pageId="13" pageNumber="1374">A. siamensis</emphasis>
</taxonomicName>
, but differ externally more or less from the males from the same localities. They were excluded from the type series, because it cannot be ruled out that they belong to another similar species occurring sympatrically in
<collectingCountry id="ACE6FFB3A81CF11AFDE7F9C9FDCB9F2F" box="[523,568,1589,1610]" name="Laos" pageId="13" pageNumber="1374">Laos</collectingCountry>
,
<collectingCountry id="ACE6FFB3A81CF11AFDACF9C8FD7E9F2F" box="[576,653,1588,1610]" name="Thailand" pageId="13" pageNumber="1374">Thailand</collectingCountry>
or
<collectingCountry id="ACE6FFB3A81CF11AFD45F9C9FCF29F2F" box="[681,769,1589,1610]" name="Myanmar" pageId="13" pageNumber="1374">Myanmar</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="9CEBECA8A81FF119FF5EFF27FBFA9994" box="[178,1033,219,241]" pageId="14" pageNumber="1375" type="etymology">
<paragraph id="D44EBF23A81FF119FF5EFF27FBFA9994" blockId="14.[178,1033,219,241]" box="[178,1033,219,241]" pageId="14" pageNumber="1375">Etymology: The species name refers to the known distribution of the species.</paragraph>
</subSubSection>
<subSubSection id="9CEBECA8A81FF118FF5EFEFCFC3998CD" lastPageId="15" lastPageNumber="1376" pageId="14" pageNumber="1375" type="description">
<paragraph id="D44EBF23A81FF119FF5EFEFCFDF19856" blockId="14.[178,1085,256,307]" pageId="14" pageNumber="1375">
Description: General appearance as figured (
<figureCitation id="4CCAA3A6A81FF119FD37FEFCFCDA9873" box="[731,809,256,278]" captionStart="Figs 21-23" captionStartId="24.[178,218,1496,1518]" captionTargetBox="[178,1085,212,1573]" captionTargetId="figure-0@24.[178,1085,212,1573]" captionTargetPageId="24" captionText="Figs 21-23: Anthracus siamensis nov.sp. Habitus, head and pronotum. (21) HT, (22, 23) PT, Laos, Pakkading." figureDoi="http://doi.org/10.5281/zenodo.5193675" httpUri="https://zenodo.org/record/5193675/files/figure.png" pageId="14" pageNumber="1375">Fig. 21</figureCitation>
). Body length
<quantity id="130912C6A81FF119FC2AFEFCFBCE9873" box="[966,1085,256,278]" metricMagnitude="-3" metricUnit="m" metricValue="3.45" metricValueMax="3.7" metricValueMin="3.2" pageId="14" pageNumber="1375" unit="mm" value="3.45" valueMax="3.7" valueMin="3.2">3.2-3.7 mm</quantity>
(HT
<quantity id="130912C6A81FF119FF0FFEE1FEC79856" box="[227,308,285,307]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="14" pageNumber="1375" unit="mm" value="3.7">3.7 mm</quantity>
); width
<quantity id="130912C6A81FF119FE69FEE1FE0D9856" box="[389,510,285,307]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="14" pageNumber="1375" unit="mm" value="1.35" valueMax="1.4" valueMin="1.3">1.3-1.4 mm</quantity>
.
</paragraph>
<paragraph id="D44EBF23A81FF119FF5EFEBFFCC59B26" blockId="14.[178,1086,323,579]" pageId="14" pageNumber="1375">Shiny, pronotum slightly, elytra moderately iridescent. Head and pronotum either dark brown to dark reddish brown, or paler reddish brown, with clypeus, labrum, and mandibles (inner margins and apices blackish) reddish yellow. Elytra paler than head and pronotum, paler to darker reddish yellow, with each elytron having a large blackish or dark brown central macula, expanding laterally to interval 7 or 8, and leaving base, apex and first interval reddish yellow. Legs and palpi pale yellowish brown, antennae with first two antennomeres yellowish brown, remaining ones moderately infuscated. Ventral surface mainly dark brown or dark reddish brown, with prosternum, epipleura, and sometimes medial part of abdominal sternites moderately paler.</paragraph>
<paragraph id="D44EBF23A81FF119FF5EFDAEFCC19A37" blockId="14.[178,1086,594,850]" pageId="14" pageNumber="1375">
Head (
<figureCitation id="4CCAA3A6A81FF119FF1AFDAEFE949B0D" box="[246,359,594,616]" captionStart="Figs 21-23" captionStartId="24.[178,218,1496,1518]" captionTargetBox="[178,1085,212,1573]" captionTargetId="figure-0@24.[178,1085,212,1573]" captionTargetPageId="24" captionText="Figs 21-23: Anthracus siamensis nov.sp. Habitus, head and pronotum. (21) HT, (22, 23) PT, Laos, Pakkading." figureDoi="http://doi.org/10.5281/zenodo.5193675" httpUri="https://zenodo.org/record/5193675/files/figure.png" pageId="14" pageNumber="1375">Figs 21-23</figureCitation>
) including eyes 0.78-0.83 times as wide as pronotum, with eyes moderately to distinctly prominent (head 1.61-1.71 times as wide as head between eyes). Labrum almost rectilinear or weakly rounded at apical margin, often somewhat sloped down to the right side. Mandibles medium sized, left mandible rather sharp at apex, not thickened or truncate. Antennae moderately long, 2.42-2.57 times as long as pronotum and 0.87-0.93 times as long as elytra. Microsculpture on clypeus distinct and almost isodiametric, on labrum weakly transverse, on frons with very lightly impressed (sometimes partly obliterated) and on vertex with moderately impressed isodiametric meshes becoming weakly transverse in front of pronotal apical margin.
</paragraph>
<paragraph id="D44EBF23A81FF119FF5EFC9EFC3D9DB2" blockId="14.[178,1085,865,1239]" pageId="14" pageNumber="1375">
Pronotum (
<figureCitation id="4CCAA3A6A81FF119FEC8FC9EFE679A1D" box="[292,404,866,888]" captionStart="Figs 21-23" captionStartId="24.[178,218,1496,1518]" captionTargetBox="[178,1085,212,1573]" captionTargetId="figure-0@24.[178,1085,212,1573]" captionTargetPageId="24" captionText="Figs 21-23: Anthracus siamensis nov.sp. Habitus, head and pronotum. (21) HT, (22, 23) PT, Laos, Pakkading." figureDoi="http://doi.org/10.5281/zenodo.5193675" httpUri="https://zenodo.org/record/5193675/files/figure.png" pageId="14" pageNumber="1375">Figs 21-23</figureCitation>
) 1.30-1.35 times as wide as long, 1.20-1.28 times as wide as head, widest in second quarter, lateral seta inserted a little posterior to beginning of second quarter. Apical margin almost rectangular or weakly emarginated, anterior angles narrowly rounded at tips, not, or weakly to moderately projecting forward. Sides convex in anterior half and rectilinearly narrowed to posterior angles, which are obtuse and weakly to moderately rounded. Basal margin sometimes rectilinear, sometimes weakly arcuate medially, and moderately to markedly oblique laterally. Lateral furrows moderately wide anteriorly, becoming weakly widened at posterior quarter, where they are fused with the baso-lateral impressions. Baso-lateral impressions medium sized, distinctly delimited from the convex pronotal disc and the weakly depressed medial part of base, flattened to basal and lateral margin. Anterior transverse impression almost obsolete. Microsculpture on disc with weakly impressed, moderately to strongly transverse meshes, and at basolateral impressions and lateral furrows with distinct weakly transverse meshes.
</paragraph>
<paragraph id="D44EBF23A81FF119FF5EFB1AFCC19CEB" blockId="14.[178,1086,1254,1422]" pageId="14" pageNumber="1375">
Elytra (
<figureCitation id="4CCAA3A6A81FF119FF13FB1AFEBA9D99" box="[255,329,1254,1276]" captionStart="Figs 21-23" captionStartId="24.[178,218,1496,1518]" captionTargetBox="[178,1085,212,1573]" captionTargetId="figure-0@24.[178,1085,212,1573]" captionTargetPageId="24" captionText="Figs 21-23: Anthracus siamensis nov.sp. Habitus, head and pronotum. (21) HT, (22, 23) PT, Laos, Pakkading." figureDoi="http://doi.org/10.5281/zenodo.5193675" httpUri="https://zenodo.org/record/5193675/files/figure.png" pageId="14" pageNumber="1375">Fig. 21</figureCitation>
) rather long, 1.53-1.61 times as long as wide, 2.71-2.85 times as long and 1.31-1.38 times as wide as pronotum. Sides weakly widened posteriad, widest at or just posterior to middle. Subapical sinuation weak. Elytral striae distinctly impressed and impunctate, intervals rather flat, becoming weakly narrowed and moderately convex at apex. Microsculpture on scutellum and around basal pore isodiametric, on elytral intervals only with traces of very lightly impressed transverse lines.
</paragraph>
<paragraph id="D44EBF23A81FF119FF5EFA62FD489F6E" blockId="14.[178,1085,1438,1548]" pageId="14" pageNumber="1375">Metepisterna long and distinctly narrowed posteriad, at inner margin about 1.5 times longer than wide at basal margin. Prosternum medially with 6 and close to apical margin with a row of 9-10 medium long setae (often broken and then insertion points difficult to observe). Prosternal process with 1-2 distinct setae.</paragraph>
<paragraph id="D44EBF23A81FF118FF5EF9E7FC0C986B" blockId="14.[178,1085,1563,1585]" lastBlockId="15.[178,1084,219,270]" lastPageId="15" lastPageNumber="1376" pageId="14" pageNumber="1375">Protarsomeres 1-4 of males moderately and mesotarsomeres 2-4 weakly dilated. Protarsomere 4 markedly and mesotarsomere 4 moderately bilobed. Protarsomeres and mesotarsomeres 1-4 of males with biseriately arranged adhesive hairs on ventral surface.</paragraph>
<paragraph id="D44EBF23A81EF118FF5EFEE1FC3998CD" blockId="15.[178,1085,285,424]" pageId="15" pageNumber="1376">
Median lobe of aedeagus (
<figureCitation id="4CCAA3A6A81EF118FE25FEE1FDCE9856" box="[457,573,285,307]" captionStart="Figs 62-67" captionStartId="31.[178,218,1520,1542]" captionTargetBox="[150,1112,174,1495]" captionTargetId="figure-0@31.[150,1112,174,1617]" captionTargetPageId="31" captionText="Figs 62-67: Median lobe of aedoeagus, lateral aspect. Anthracus javaensis nov.sp. (62) HT and A. siamensis nov.sp. (63, 64, 67) PT, Laos, Pakkading, (65) HT, (66) Thailand, Ban Khoun." figureDoi="http://doi.org/10.5281/zenodo.6546849" httpUri="https://zenodo.org/record/6546849/files/figure.png" pageId="15" pageNumber="1376">Figs 63-67</figureCitation>
,
<figureCitation id="4CCAA3A6A81EF118FDA6FEE1FD759856" box="[586,646,285,307]" captionStart="Figs 68-73" captionStartId="32.[178,218,1420,1442]" captionTargetBox="[150,1112,159,1602]" captionTargetId="figure-0@32.[150,1112,159,1602]" captionTargetPageId="32" captionText="Figs 68-73: Median lobe of aedoeagus, dorsal aspect. Anthracus javaensis nov.sp. (68) HT and A. siamensis nov.sp. (69) PT, Thailand, Umphang, (70, 73) PT, Laos, Pakkading, (71) HT, (72) Thailand, Ban Khoun." figureDoi="http://doi.org/10.5281/zenodo.5194872" httpUri="https://zenodo.org/record/5194872/files/figure.png" pageId="15" pageNumber="1376">69-73</figureCitation>
) moderately large, with external shape as figured. Apical lamella rather short, not markedly narrowed (dorsal aspect), the apex abruptly bent upward (lateral aspect). Internal structures composed of one large apical tooth usually orientated to left side (lateral aspect), 0-3 medium-sized subapical teeth, 1- 3 small teeth in medial portion and 1-2 small and tall teeth somewhat beneath.
</paragraph>
</subSubSection>
<subSubSection id="9CEBECA8A81EF118FF5EFE44FC6F9AA2" pageId="15" pageNumber="1376" type="discussion">
<paragraph id="D44EBF23A81EF118FF5EFE44FD469B18" blockId="15.[178,1085,440,637]" pageId="15" pageNumber="1376">
Comparisons
<taxonomicName id="13F1C4A0A81EF118FE72FE44FDEE98A8" authority="Jaeger, 2015" authorityName="Jaeger" authorityYear="2015" box="[414,541,440,461]" class="Insecta" family="Carabidae" genus="Anthracus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="1376" phylum="Arthropoda" rank="species" species="siamensis" status="sp. nov.">
<emphasis id="E6856331A81EF118FE72FE44FDEE98A8" box="[414,541,440,461]" italics="true" pageId="15" pageNumber="1376">A. siamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FDB6DE4AA81EF118FDC9FE44FD9C98A8" box="[549,623,440,461]" pageId="15" pageNumber="1376" rank="species">nov.sp.</taxonomicNameLabel>
is closely related to
<taxonomicName id="13F1C4A0A81EF118FCA9FE44FC3398A8" authority="Jaeger, 2015" authorityName="Jaeger" authorityYear="2015" box="[837,960,440,461]" class="Insecta" family="Carabidae" genus="Anthracus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="1376" phylum="Arthropoda" rank="species" species="javaensis" status="sp. nov.">
<emphasis id="E6856331A81EF118FCA9FE44FC3398A8" box="[837,960,440,461]" italics="true" pageId="15" pageNumber="1376">A. javaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FDB6DE4AA81EF118FC24FE44FBE198A8" box="[968,1042,440,461]" pageId="15" pageNumber="1376" rank="species">nov.sp.</taxonomicNameLabel>
and
<taxonomicName id="13F1C4A0A81EF118FF5EFE29FE6E988E" authority="JAEGER, 2015" authorityName="JAEGER" authorityYear="2015" box="[178,413,469,491]" class="Insecta" family="Carabidae" genus="Anthracus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="1376" phylum="Arthropoda" rank="species" species="skalei">
<emphasis id="E6856331A81EF118FF5EFE29FEF9988F" box="[178,266,469,490]" italics="true" pageId="15" pageNumber="1376">A. skalei</emphasis>
<bibRefCitation id="B060C2D2A81EF118FEFDFE29FE6E988E" author="JAEGER B" box="[273,413,469,491]" pageId="15" pageNumber="1376" refId="ref12895" refString="JAEGER B. (2015): Two new maculate species of the Anthracus annamensis group from Nepal and adjacent areas (Insecta: Coleoptera: Carabidae: Harpalini: Stenolophina). Pp. 339 - 345. - In: HARTMANN M. &amp; J. WEIPERT (eds), Biodiversitat und Naturausstattung im Himalaya V. - Verein der Freunde und Forderer des Naturkundemuseums Erfurt e. V. Erfurt: 580 pp." type="book" year="2015">JAEGER, 2015</bibRefCitation>
</taxonomicName>
and obviously represents a closely related eastern vicariant of the latter. Both species are very similar in general appearance, microsculpture and body proportions which do not provide reliable differences to distinguish them. However, they differ by constant differences in the arrangement of teeth in the internal sac of the aedeagus (see
<figureCitation id="4CCAA3A6A81EF118FEBCFDB6FDE29B05" box="[336,529,586,608]" captionStart-0="Figs 62-67" captionStart-1="Figs 68-73" captionStart-2="Figs 74-79" captionStartId-0="31.[178,218,1520,1542]" captionStartId-1="32.[178,218,1420,1442]" captionStartId-2="33.[178,218,1494,1516]" captionTargetBox-0="[150,1112,174,1495]" captionTargetBox-1="[150,1112,159,1602]" captionTargetBox-2="[150,1107,181,1473]" captionTargetId-0="figure-0@31.[150,1112,174,1617]" captionTargetId-1="figure-0@32.[150,1112,159,1602]" captionTargetId-2="figure-0@33.[150,1112,181,1624]" captionTargetPageId-0="31" captionTargetPageId-1="32" captionTargetPageId-2="33" captionText-0="Figs 62-67: Median lobe of aedoeagus, lateral aspect. Anthracus javaensis nov.sp. (62) HT and A. siamensis nov.sp. (63, 64, 67) PT, Laos, Pakkading, (65) HT, (66) Thailand, Ban Khoun." captionText-1="Figs 68-73: Median lobe of aedoeagus, dorsal aspect. Anthracus javaensis nov.sp. (68) HT and A. siamensis nov.sp. (69) PT, Thailand, Umphang, (70, 73) PT, Laos, Pakkading, (71) HT, (72) Thailand, Ban Khoun." captionText-2="Figs 74-79: Median lobe of aedoeagus, lateral and dorsal aspect. Anthracus skalei JAEGER (74) PT, Bangladesh, Nasirabad, (75) PT, India, Kurseong, A. spec. 1 near A. biplagiatus (BOHEMAN) (76, 77) Bangladesh, Nasirabad, A. spec. 2 near A. annamensis (BATES) (78,79) Myanmar, Pegu." figureDoi-0="http://doi.org/10.5281/zenodo.6546849" figureDoi-1="http://doi.org/10.5281/zenodo.5194872" figureDoi-2="http://doi.org/10.5281/zenodo.6546851" httpUri-0="https://zenodo.org/record/6546849/files/figure.png" httpUri-1="https://zenodo.org/record/5194872/files/figure.png" httpUri-2="https://zenodo.org/record/6546851/files/figure.png" pageId="15" pageNumber="1376">Figs 62-75, 74-75</figureCitation>
) and partly also by the darker colour of head and pronotum (only darker specimens of
<taxonomicName id="13F1C4A0A81EF118FDC4FD94FD549B18" authorityName="Jaeger" authorityYear="2015" box="[552,679,616,637]" class="Insecta" family="Carabidae" genus="Anthracus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="1376" phylum="Arthropoda" rank="species" species="siamensis">
<emphasis id="E6856331A81EF118FDC4FD94FD549B18" box="[552,679,616,637]" italics="true" pageId="15" pageNumber="1376">A. siamensis</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="D44EBF23A81EF118FF5EFD71FC6F9AA2" blockId="15.[178,1086,653,967]" pageId="15" pageNumber="1376">
<taxonomicName id="13F1C4A0A81EF118FF5EFD71FEC29BC7" authorityName="Jaeger" authorityYear="2015" box="[178,305,653,674]" class="Insecta" family="Carabidae" genus="Anthracus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="1376" phylum="Arthropoda" rank="species" species="siamensis">
<emphasis id="E6856331A81EF118FF5EFD71FEC29BC7" box="[178,305,653,674]" italics="true" pageId="15" pageNumber="1376">A. siamensis</emphasis>
</taxonomicName>
is also very similar in general habitus to
<taxonomicName id="13F1C4A0A81EF118FD37FD71FCAC9BC7" authority="Jaeger, 2015" authorityName="Jaeger" authorityYear="2015" box="[731,863,653,674]" class="Insecta" family="Carabidae" genus="Anthracus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="1376" phylum="Arthropoda" rank="species" species="hornburgi" status="sp. nov.">
<emphasis id="E6856331A81EF118FD37FD71FCAC9BC7" box="[731,863,653,674]" italics="true" pageId="15" pageNumber="1376">A. hornburgi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FDB6DE4AA81EF118FC8AFD71FC5C9BC7" box="[870,943,653,674]" pageId="15" pageNumber="1376" rank="species">nov.sp.</taxonomicNameLabel>
which occurs in the adjacent areas of
<collectingCountry id="ACE6FFB3A81EF118FE48FD56FDFA9BDA" box="[420,521,682,703]" name="Myanmar" pageId="15" pageNumber="1376">Myanmar</collectingCountry>
, and to
<taxonomicName id="13F1C4A0A81EF118FDB5FD56FCB59BDA" authority="(BATES)" baseAuthorityName="BATES" box="[601,838,682,704]" class="Insecta" family="Carabidae" genus="Acupalpus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="1376" phylum="Arthropoda" rank="species" species="annamensis">
<emphasis id="E6856331A81EF118FDB5FD56FD1F9BA5" box="[601,748,682,704]" italics="true" pageId="15" pageNumber="1376">A. annamensis</emphasis>
(BATES)
</taxonomicName>
and closely related taxa which occur in
<collectingCountry id="ACE6FFB3A81EF118FEB2FD34FE309BB8" box="[350,451,712,733]" name="Myanmar" pageId="15" pageNumber="1376">Myanmar</collectingCountry>
,
<collectingCountry id="ACE6FFB3A81EF118FE3EFD34FDF09BB8" box="[466,515,712,733]" name="Laos" pageId="15" pageNumber="1376">Laos</collectingCountry>
and
<collectingCountry id="ACE6FFB3A81EF118FDD3FD3BFD689BB8" box="[575,667,711,733]" name="Thailand" pageId="15" pageNumber="1376">Thailand</collectingCountry>
, at
<collectingRegion id="163571C1A81EF118FD24FD3BFCB59BB8" box="[712,838,711,733]" country="Thailand" name="Chiang Mai" pageId="15" pageNumber="1376">Chiang Mai</collectingRegion>
even sympatrically.
<taxonomicName id="13F1C4A0A81EF118FBCBFD34FEE79B9F" authorityName="Jaeger" authorityYear="2015" class="Insecta" family="Carabidae" genus="Anthracus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="1376" phylum="Arthropoda" rank="species" species="siamensis">
<emphasis id="E6856331A81EF118FBCBFD34FEE79B9F" italics="true" pageId="15" pageNumber="1376">A. siamensis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="13F1C4A0A81EF118FE4CFD19FDD69B9C" authority="Jaeger, 2015" authorityName="Jaeger" authorityYear="2015" box="[416,549,740,762]" class="Insecta" family="Carabidae" genus="Anthracus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="1376" phylum="Arthropoda" rank="species" species="hornburgi" status="sp. nov.">
<emphasis id="E6856331A81EF118FE4CFD19FDD69B9C" box="[416,549,740,762]" italics="true" pageId="15" pageNumber="1376">A. hornburgi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FDB6DE4AA81EF118FDC1FD19FD849B9F" box="[557,631,741,762]" pageId="15" pageNumber="1376" rank="species">nov.sp.</taxonomicNameLabel>
by the aedeagal characters given under this species, and from
<taxonomicName id="13F1C4A0A81EF118FE9CFCFEFDF69A72" authority="(BATES)" baseAuthorityName="BATES" box="[368,517,770,791]" class="Insecta" family="Carabidae" genus="Acupalpus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="1376" phylum="Arthropoda" rank="species" sensu="lato" species="annamensis">
<emphasis id="E6856331A81EF118FE9CFCFEFDF69A72" box="[368,517,770,791]" italics="true" pageId="15" pageNumber="1376">A. annamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FDB6DE4AA81EF118FDE2FCFEFDD89A7D" box="[526,555,770,792]" pageId="15" pageNumber="1376" sensu="lato">s.l.</taxonomicNameLabel>
(
<figureCitation id="4CCAA3A6A81EF118FDD7FCFEFD5E9A7D" box="[571,685,770,792]" captionStart="Figs 74-79" captionStartId="33.[178,218,1494,1516]" captionTargetBox="[150,1107,181,1473]" captionTargetId="figure-0@33.[150,1112,181,1624]" captionTargetPageId="33" captionText="Figs 74-79: Median lobe of aedoeagus, lateral and dorsal aspect. Anthracus skalei JAEGER (74) PT, Bangladesh, Nasirabad, (75) PT, India, Kurseong, A. spec. 1 near A. biplagiatus (BOHEMAN) (76, 77) Bangladesh, Nasirabad, A. spec. 2 near A. annamensis (BATES) (78,79) Myanmar, Pegu." figureDoi="http://doi.org/10.5281/zenodo.6546851" httpUri="https://zenodo.org/record/6546851/files/figure.png" pageId="15" pageNumber="1376">Figs 78-79</figureCitation>
) by the different external shape of the median lobe of the aedeagus with the peculiar shape of the apex in lateral aspect and the different internal structures with large apical and subapical teeth (in
<taxonomicName id="13F1C4A0A81EF118FC96FCC1FBE39A37" authority="(BATES)" baseAuthorityName="BATES" box="[890,1040,829,850]" class="Insecta" family="Carabidae" genus="Acupalpus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="1376" phylum="Arthropoda" rank="species" sensu="lato" species="annamensis">
<emphasis id="E6856331A81EF118FC96FCC1FBE39A37" box="[890,1040,829,850]" italics="true" pageId="15" pageNumber="1376">A. annamensis</emphasis>
</taxonomicName>
and closely related taxa apex simple without any hook and teeth markedly smaller and with different arrangement). Externally there are often no reliable differences to differentiate
<taxonomicName id="13F1C4A0A81EF118FF5EFC68FEC29ACF" authorityName="Jaeger" authorityYear="2015" box="[178,305,916,938]" class="Insecta" family="Carabidae" genus="Anthracus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="1376" phylum="Arthropoda" rank="species" species="siamensis">
<emphasis id="E6856331A81EF118FF5EFC68FEC29ACF" box="[178,305,916,938]" italics="true" pageId="15" pageNumber="1376">A. siamensis</emphasis>
</taxonomicName>
from
<taxonomicName id="13F1C4A0A81EF118FE9DFC68FE069ACC" authorityName="Jaeger" authorityYear="2015" box="[369,501,916,937]" class="Insecta" family="Carabidae" genus="Anthracus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="1376" phylum="Arthropoda" rank="species" species="hornburgi">
<emphasis id="E6856331A81EF118FE9DFC68FE069ACC" box="[369,501,916,937]" italics="true" pageId="15" pageNumber="1376">A. hornburgi</emphasis>
</taxonomicName>
and
<taxonomicName id="13F1C4A0A81EF118FDC4FC68FD4F9ACF" authority="(BATES)" baseAuthorityName="BATES" box="[552,700,916,938]" class="Insecta" family="Carabidae" genus="Acupalpus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="1376" phylum="Arthropoda" rank="species" sensu="lato" species="annamensis">
<emphasis id="E6856331A81EF118FDC4FC68FD4F9ACF" box="[552,700,916,938]" italics="true" pageId="15" pageNumber="1376">A. annamensis</emphasis>
</taxonomicName>
(incl. closely related taxa) but darker specimens differ from these taxa by their dark brown head and pronotum.
</paragraph>
</subSubSection>
<subSubSection id="9CEBECA8A81EF118FF5EFC2BFF289D6F" pageId="15" pageNumber="1376" type="distribution">
<paragraph id="D44EBF23A81EF118FF5EFC2BFF289D6F" blockId="15.[178,1086,983,1034]" pageId="15" pageNumber="1376">
Distribution
<taxonomicName id="13F1C4A0A81EF118FE76FC2BFDEA9A89" authority="Jaeger, 2015" authorityName="Jaeger" authorityYear="2015" box="[410,537,983,1004]" class="Insecta" family="Carabidae" genus="Anthracus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="1376" phylum="Arthropoda" rank="species" species="siamensis" status="sp. nov.">
<emphasis id="E6856331A81EF118FE76FC2BFDEA9A89" box="[410,537,983,1004]" italics="true" pageId="15" pageNumber="1376">A. siamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FDB6DE4AA81EF118FDF3FC2BFD9A9A89" box="[543,617,983,1004]" pageId="15" pageNumber="1376" rank="species">nov.sp.</taxonomicNameLabel>
is so far known from
<collectingCountry id="ACE6FFB3A81EF118FCA5FC2BFC509A88" box="[841,931,983,1005]" name="Thailand" pageId="15" pageNumber="1376">Thailand</collectingCountry>
and
<collectingCountry id="ACE6FFB3A81EF118FC39FC2BFBF59A89" box="[981,1030,983,1004]" name="Laos" pageId="15" pageNumber="1376">Laos</collectingCountry>
(
<figureCitation id="4CCAA3A6A81EF118FBF8FC2BFF3C9D6F" captionStart="Fig 80" captionStartId="34.[178,210,909,931]" captionTargetBox="[266,997,212,896]" captionTargetId="figure-65@34.[266,997,212,896]" captionTargetPageId="34" captionText="Fig 80: Distribution: A. nesophilus (ANDREWES) (triangles), A. javaensis nov.sp. (white star), A. sumatraensis nov.sp. (square) A. philippinensis nov.sp. (circle), A. siamensis nov.sp. (black star)." figureDoi="http://doi.org/10.5281/zenodo.5193681" httpUri="https://zenodo.org/record/5193681/files/figure.png" pageId="15" pageNumber="1376">Fig. 80</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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