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<document id="D6C30B7233F7BED5709CA03C4B5FEDCA" ID-CLB-Dataset="55535" ID-DOI="10.5281/zenodo.5193661" ID-GBIF-Dataset="c2346a92-3813-4b3d-b139-f3b149b50f4a" ID-ISSN="0253-116X" ID-Zenodo-Dep="5193661" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628802610787" checkinUser="felipe" docAuthor="Jaeger, Bernd" docDate="2015" docId="5C580E35A819F11DFF5EFB32FDF69B65" docLanguage="en" docName="LinzerbiolBeitr.47.2.1361-1396.pdf" docOrigin="Linzer biologische Beiträge 47 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Anthracus sumatraensis Jaeger 2015, nov.sp." docType="treatment" docVersion="11" lastPageNumber="1371" masterDocId="A061764DA811F117FFECFFFCFFF39965" masterDocTitle="Revision of the maculate species of the Anthracus annamensis group from the East Palaearctic and Oriental Regions. Part 2. A redescription of Anthracus nesophilus (ANDREWES, 1936) and six new species from Nepal, India and SE Asia (Coleoptera, Carabidae, Harpalini, Stenolophina)" masterLastPageNumber="1396" masterPageNumber="1361" pageNumber="1369" updateTime="1699052547504" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="51BBC0FE0BC5039A6BE99A86317C8EAD">Revision of the maculate species of the Anthracus annamensis group from the East Palaearctic and Oriental Regions. Part 2. A redescription of Anthracus nesophilus (ANDREWES, 1936) and six new species from Nepal, India and SE Asia (Coleoptera, Carabidae, Harpalini, Stenolophina)</mods:title>
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<mods:namePart id="DD04D0B024FD90C4FFA977CF8438AF14">Jaeger, Bernd</mods:namePart>
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<mods:title id="777816CDB755A1022D75F6749B029946">Linzer biologische Beiträge</mods:title>
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<mods:date id="805BC3F75FCD86D99FA885B32FFA1886">2015</mods:date>
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<mods:number id="B614E2BF6C87AE9E1050976F31ECE47C">2015-12-30</mods:number>
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<treatment id="5C580E35A819F11DFF5EFB32FDF69B65" ID-DOI="http://doi.org/10.5281/zenodo.6546856" ID-GBIF-Taxon="195305593" ID-Zenodo-Dep="6546856" LSID="urn:lsid:plazi:treatment:5C580E35A819F11DFF5EFB32FDF69B65" httpUri="http://treatment.plazi.org/id/5C580E35A819F11DFF5EFB32FDF69B65" lastPageId="10" lastPageNumber="1371" pageId="8" pageNumber="1369">
<subSubSection id="9CEBECA8A819F11FFF5EFB32FDF19D81" box="[178,514,1230,1252]" pageId="8" pageNumber="1369" type="nomenclature">
<paragraph id="D44EBF23A819F11FFF5EFB32FDF19D81" blockId="8.[178,757,1230,1253]" box="[178,514,1230,1252]" pageId="8" pageNumber="1369">
<heading id="8F06084FA819F11FFF5EFB32FDF19D81" box="[178,514,1230,1252]" fontSize="8" level="2" pageId="8" pageNumber="1369" reason="2">
<emphasis id="E6856331A819F11FFF5EFB32FDF19D81" bold="true" box="[178,514,1230,1252]" pageId="8" pageNumber="1369">
<taxonomicName id="13F1C4A0A819F11FFF5EFB32FE5C9D81" ID-CoL="5VH8F" authority="Jaeger, 2015" authorityName="Jaeger" authorityYear="2015" box="[178,431,1230,1252]" class="Insecta" family="Carabidae" genus="Anthracus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1369" phylum="Arthropoda" rank="species" species="sumatraensis" status="sp. nov.">
<emphasis id="E6856331A819F11FFF5EFB32FE5C9D81" bold="true" box="[178,431,1230,1252]" italics="true" pageId="8" pageNumber="1369">Anthracus sumatraensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FDB6DE4AA819F11FFE5AFB33FDF19D81" box="[438,514,1231,1252]" pageId="8" pageNumber="1369" rank="species">nov.sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="9CEBECA8A819F11FFDE4FB33FD069D80" box="[520,757,1231,1253]" pageId="8" pageNumber="1369" type="description">
<paragraph id="D44EBF23A819F11FFDE4FB33FD069D80" blockId="8.[178,757,1230,1253]" box="[520,757,1231,1253]" pageId="8" pageNumber="1369">
<heading id="8F06084FA819F11FFDE4FB33FD069D80" box="[520,757,1231,1253]" fontSize="8" level="2" pageId="8" pageNumber="1369" reason="2">
<emphasis id="E6856331A819F11FFDE4FB33FD069D80" bold="true" box="[520,757,1231,1253]" pageId="8" pageNumber="1369">
(
<figureCitation id="4CCAA3A6A819F11FFDE3FB33FD739D81" box="[527,640,1231,1253]" captionStart="Figs 13-15" captionStartId="21.[178,218,1430,1452]" captionTargetBox="[178,1085,212,1476]" captionTargetId="figure-0@21.[178,1085,212,1476]" captionTargetPageId="21" captionText="Figs 13-15: Anthracus sumatraensis nov.sp. Habitus, head and pronotum. (13) HT, (14-15) PT, Sumatra, Sibolga." figureDoi="http://doi.org/10.5281/zenodo.5193669" httpUri="https://zenodo.org/record/5193669/files/figure.png" pageId="8" pageNumber="1369">Figs 13-15</figureCitation>
,
<figureCitation id="4CCAA3A6A819F11FFD60FB33FD3B9D81" box="[652,712,1231,1252]" captionStart="Figs 50-55" captionStartId="29.[178,218,1492,1514]" captionTargetBox="[150,1110,191,1479]" captionTargetId="figure-0@29.[150,1112,189,1632]" captionTargetPageId="29" captionText="Figs 50-55: Anthracus sumatraensis nov.sp. Median lobe of aedoeagus, lateral and dorsal aspect. (50, 52, 53, 55) PT, (51, 54) HT." figureDoi="http://doi.org/10.5281/zenodo.6546843" httpUri="https://zenodo.org/record/6546843/files/figure.png" pageId="8" pageNumber="1369">50-55</figureCitation>
,
<figureCitation id="4CCAA3A6A819F11FFD38FB33FD1C9D80" box="[724,751,1231,1253]" captionStart="Fig 80" captionStartId="34.[178,210,909,931]" captionTargetBox="[266,997,212,896]" captionTargetId="figure-65@34.[266,997,212,896]" captionTargetPageId="34" captionText="Fig 80: Distribution: A. nesophilus (ANDREWES) (triangles), A. javaensis nov.sp. (white star), A. sumatraensis nov.sp. (square) A. philippinensis nov.sp. (circle), A. siamensis nov.sp. (black star)." figureDoi="http://doi.org/10.5281/zenodo.5193681" httpUri="https://zenodo.org/record/5193681/files/figure.png" pageId="8" pageNumber="1369">80</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="9CEBECA8A819F11FFF5EFAFCFCF79CE4" pageId="8" pageNumber="1369" type="materials_examined">
<paragraph id="D44EBF23A819F11FFF5EFAFCFEA09C2E" blockId="8.[178,1085,1280,1410]" pageId="8" pageNumber="1369">
Type material:
<materialsCitation id="6499B57EA819F11FFE65FAFCFEA09C2E" ID-GBIF-Occurrence="3766781319" collectionCode="NHMW" collectorName="Schillhammer &amp; B. Jaeger" country="Indonesia" location="Sumatra" pageId="8" pageNumber="1369" specimenCount="1" specimenCount-male="1" stateProvince="Sumatra" typeStatus="holotype">
<typeStatus id="0B4A0181A819F11FFE65FAFCFE2E9C73" box="[393,477,1280,1302]" pageId="8" pageNumber="1369" type="holotype">Holotype</typeStatus>
:
<specimenCount id="C2F774AAA819F11FFE07FAFCFE0D9C70" box="[491,510,1280,1301]" pageId="8" pageNumber="1369" type="male">♂</specimenCount>
(
<collectionCode id="B2E027E6A819F11FFDE7FAFCFDAB9C73" box="[523,600,1280,1302]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="8" pageNumber="1369" type="Museum">NHMW</collectionCode>
) labelled &quot;
<collectingCountry id="ACE6FFB3A819F11FFD57FAFCFCE69C73" box="[699,789,1280,1302]" name="Indonesia" pageId="8" pageNumber="1369">Indonesia</collectingCountry>
1990 / leg.
<collectorName id="7904DAF5A819F11FFC6AFAFCFBF99C73" box="[902,1034,1280,1302]" pageId="8" pageNumber="1369">Schillhammer</collectorName>
&quot;; &quot;
<collectingRegion id="163571C1A819F11FFBC8FAFCFF0D9C55" country="Indonesia" name="Sumatera Utara" pageId="8" pageNumber="1369">N-Sumatra</collectingRegion>
/ 16. 2. / n-Sibolga&quot; and &quot;HOLOTYPE ♂ /
<originalDetermination id="F7EF07DFA819F11FFD7DFAE7FC5E9C54" box="[657,941,1307,1329]" pageId="8" pageNumber="1369">
<taxonomicName id="13F1C4A0A819F11FFD7DFAE7FD1D9C54" authorityName="Motschoulsky" authorityYear="1850" box="[657,750,1307,1329]" class="Insecta" family="Carabidae" genus="Anthracus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1369" phylum="Arthropoda" rank="genus">Anthracus</taxonomicName>
/ sumatraensis
<taxonomicNameLabel id="FDB6DE4AA819F11FFC97FAE0FC5E9C54" box="[891,941,1308,1329]" pageId="8" pageNumber="1369" rank="species">sp. n.</taxonomicNameLabel>
</originalDetermination>
/ des.
<collectorName id="7904DAF5A819F11FFC04FAE0FBCE9C54" box="[1000,1085,1308,1329]" pageId="8" pageNumber="1369">
B.
<bibRefCitation id="B060C2D2A819F11FFBEFFAE0FBCE9C54" author="JAEGER B" box="[1027,1085,1308,1329]" pageId="8" pageNumber="1369" refId="ref12895" refString="JAEGER B. (2015): Two new maculate species of the Anthracus annamensis group from Nepal and adjacent areas (Insecta: Coleoptera: Carabidae: Harpalini: Stenolophina). Pp. 339 - 345. - In: HARTMANN M. &amp; J. WEIPERT (eds), Biodiversitat und Naturausstattung im Himalaya V. - Verein der Freunde und Forderer des Naturkundemuseums Erfurt e. V. Erfurt: 580 pp." type="book" year="2015">Jaeger</bibRefCitation>
</collectorName>
2015&quot; [red label].
</materialsCitation>
</paragraph>
<paragraph id="D44EBF23A819F11FFF5EFAAEFCF79CE4" blockId="8.[178,1085,1280,1410]" pageId="8" pageNumber="1369">
<materialsCitation id="6499B57EA819F11FFF5EFAAEFD0C9CE4" ID-GBIF-Occurrence="3766781329" pageId="8" pageNumber="1369" typeStatus="paratype">
<typeStatus id="0B4A0181A819F11FFF5EFAAEFEF99C02" box="[178,266,1362,1383]" pageId="8" pageNumber="1369" type="paratype">Paratypes</typeStatus>
: 2♂♂ (NHMW, cJAE) with same locality labels as the holotype and &quot;PARATYPE ♂ /
<taxonomicName id="13F1C4A0A819F11FFF5EFA90FEFC9CE7" authorityName="Motschoulsky" authorityYear="1850" box="[178,271,1388,1410]" class="Insecta" family="Carabidae" genus="Anthracus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1369" phylum="Arthropoda" rank="genus">Anthracus</taxonomicName>
/ sumatraensis
<taxonomicNameLabel id="FDB6DE4AA819F11FFE70FA90FE3D9CE4" box="[412,462,1388,1409]" pageId="8" pageNumber="1369" rank="species">sp. n.</taxonomicNameLabel>
/ des. B.
<bibRefCitation id="B060C2D2A819F11FFDCFFA90FD619CE7" author="JAEGER B" box="[547,658,1388,1410]" pageId="8" pageNumber="1369" refId="ref12895" refString="JAEGER B. (2015): Two new maculate species of the Anthracus annamensis group from Nepal and adjacent areas (Insecta: Coleoptera: Carabidae: Harpalini: Stenolophina). Pp. 339 - 345. - In: HARTMANN M. &amp; J. WEIPERT (eds), Biodiversitat und Naturausstattung im Himalaya V. - Verein der Freunde und Forderer des Naturkundemuseums Erfurt e. V. Erfurt: 580 pp." type="book" year="2015">Jaeger 2015</bibRefCitation>
&quot; [red label]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="9CEBECA8A819F11FFF5EFA6DFBFA9CC2" box="[178,1033,1425,1447]" pageId="8" pageNumber="1369" type="etymology">
<paragraph id="D44EBF23A819F11FFF5EFA6DFBFA9CC2" blockId="8.[178,1033,1425,1447]" box="[178,1033,1425,1447]" pageId="8" pageNumber="1369">Etymology: The species name refers to the known distribution of the species.</paragraph>
</subSubSection>
<subSubSection id="9CEBECA8A819F11EFF5EFA4AFE169F69" lastPageId="9" lastPageNumber="1370" pageId="8" pageNumber="1369" type="description">
<paragraph id="D44EBF23A819F11FFF5EFA4AFE269C8C" blockId="8.[178,1085,1462,1513]" pageId="8" pageNumber="1369">
Description: General appearance as figured (
<figureCitation id="4CCAA3A6A819F11FFD37FA4AFCDA9CA9" box="[731,809,1462,1484]" captionStart="Figs 13-15" captionStartId="21.[178,218,1430,1452]" captionTargetBox="[178,1085,212,1476]" captionTargetId="figure-0@21.[178,1085,212,1476]" captionTargetPageId="21" captionText="Figs 13-15: Anthracus sumatraensis nov.sp. Habitus, head and pronotum. (13) HT, (14-15) PT, Sumatra, Sibolga." figureDoi="http://doi.org/10.5281/zenodo.5193669" httpUri="https://zenodo.org/record/5193669/files/figure.png" pageId="8" pageNumber="1369">Fig. 13</figureCitation>
). Body length
<quantity id="130912C6A819F11FFC2AFA4AFBCE9CAE" box="[966,1085,1462,1484]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" metricValueMax="3.9" metricValueMin="3.6" pageId="8" pageNumber="1369" unit="mm" value="3.75" valueMax="3.9" valueMin="3.6">3.6-3.9 mm</quantity>
(HT 3.9); width
<quantity id="130912C6A819F11FFEB4FA2FFE239C8C" box="[344,464,1491,1513]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="8" pageNumber="1369" unit="mm" value="1.45" valueMax="1.5" valueMin="1.4">1.4-1.5 mm</quantity>
.
</paragraph>
<paragraph id="D44EBF23A819F11EFF5EFA04FE509803" blockId="8.[178,1085,1528,1609]" lastBlockId="9.[178,1086,219,358]" lastPageId="9" lastPageNumber="1370" pageId="8" pageNumber="1369">Shiny, pronotum slightly, elytra moderately iridescent. Head and pronotum reddish yellow, with clypeus, labrum, and mandibles (inner margins and apices blackish) weakly paler. Elytra with same ground colour as head and pronotum, with each elytron having a large, indistinct dark brown central macula, expanding laterally to interval 5, 6 or 7, and leaving base, apex and first interval reddish yellow. Legs and palpi pale yellowish brown, antennae with first two antennomeres yellowish brown, remaining ones weakly infuscated. Ventral surface mainly reddish brown, sometimes abdominal sternites and epipleura slightly paler.</paragraph>
<paragraph id="D44EBF23A818F11EFF5EFE89FC709B3D" blockId="9.[178,1085,373,600]" pageId="9" pageNumber="1370">
Head (
<figureCitation id="4CCAA3A6A818F11EFF1AFE89FE9598EE" box="[246,358,373,395]" captionStart="Figs 13-15" captionStartId="21.[178,218,1430,1452]" captionTargetBox="[178,1085,212,1476]" captionTargetId="figure-0@21.[178,1085,212,1476]" captionTargetPageId="21" captionText="Figs 13-15: Anthracus sumatraensis nov.sp. Habitus, head and pronotum. (13) HT, (14-15) PT, Sumatra, Sibolga." figureDoi="http://doi.org/10.5281/zenodo.5193669" httpUri="https://zenodo.org/record/5193669/files/figure.png" pageId="9" pageNumber="1370">Figs 13-15</figureCitation>
) including eyes 0.81-0.83 times as wide as pronotum, eyes moderately prominent (head 1.59-1.65 times as wide as head between eyes). Labrum with apical margin linear but somewhat sloping down to the right side. Mandibles medium sized, left mandible rather sharp at apex, not thickened or truncate. Antennae moderately long, 2.52-2.62 times as long as pronotum and 0.90-0.92 times as long as elytra. Microsculpture on clypeus almost isodiametric, on labrum weakly transverse, on frons with very fine and lightly impressed and on vertex with moderately impressed isodiametric meshes becoming weakly transverse in front of pronotal apical margin.
</paragraph>
<paragraph id="D44EBF23A818F11EFF5EFD94FC7F9AB8" blockId="9.[178,1086,615,989]" pageId="9" pageNumber="1370">
Pronotum (
<figureCitation id="4CCAA3A6A818F11EFEC8FD9BFE609B18" box="[292,403,615,637]" captionStart="Figs 13-15" captionStartId="21.[178,218,1430,1452]" captionTargetBox="[178,1085,212,1476]" captionTargetId="figure-0@21.[178,1085,212,1476]" captionTargetPageId="21" captionText="Figs 13-15: Anthracus sumatraensis nov.sp. Habitus, head and pronotum. (13) HT, (14-15) PT, Sumatra, Sibolga." figureDoi="http://doi.org/10.5281/zenodo.5193669" httpUri="https://zenodo.org/record/5193669/files/figure.png" pageId="9" pageNumber="1370">Figs 13-15</figureCitation>
) 1.26-1.31 times as wide as long, 1.21-1.24 times as wide as head, widest in second quarter, lateral seta inserted a little posterior to beginning of second quarter. Apical margin almost rectilinear or very faintly emarginated, anterior angles narrowly rounded at tips, not or very weakly projecting forward. Sides convex in anterior half and rectilinearly narrowed to posterior angles, which are obtuse and moderately rounded. Basal margin weakly to moderately arcuate medially and oblique to posterior angles. Lateral furrows moderately wide anteriorly, becoming moderately widened in posterior third, where they are fused with the baso-lateral impressions. The latter medium sized and somewhat oblique, distinctly delimited from the pronotal disc and the somewhat depressed median part of base, fused with basal and lateral margins, the latter weakly reflexed. Anterior transverse impression obsolete, rarely suggested. Microsculpture on disc with lightly impressed, strongly transverse meshes, at baso-lateral impressions and lateral furrows with markedly impressed isodiametric meshes.
</paragraph>
<paragraph id="D44EBF23A818F11EFF5EFC10FD3C9DF1" blockId="9.[178,1086,1004,1172]" pageId="9" pageNumber="1370">
Elytra (
<figureCitation id="4CCAA3A6A818F11EFF12FC10FEBB9D67" box="[254,328,1004,1026]" captionStart="Figs 13-15" captionStartId="21.[178,218,1430,1452]" captionTargetBox="[178,1085,212,1476]" captionTargetId="figure-0@21.[178,1085,212,1476]" captionTargetPageId="21" captionText="Figs 13-15: Anthracus sumatraensis nov.sp. Habitus, head and pronotum. (13) HT, (14-15) PT, Sumatra, Sibolga." figureDoi="http://doi.org/10.5281/zenodo.5193669" httpUri="https://zenodo.org/record/5193669/files/figure.png" pageId="9" pageNumber="1370">Fig. 13</figureCitation>
) rather short, 1.53-1.57 times as long as wide, 2.75-2.91 times as long and 1.41-1.43 times as wide as pronotum. Sides moderately widened posteriorly, widest just posterior to middle. Subapical sinuation weak. Elytral striae distinctly impressed and impunctate, intervals rather flat, becoming weakly narrowed and weakly convex at apex. Microsculpture on scutellum and around basal pore isodiametric, on elytral intervals only with traces of very lightly impressed transverse lines.
</paragraph>
<paragraph id="D44EBF23A818F11EFF5EFB58FD689C74" blockId="9.[178,1085,1187,1297]" pageId="9" pageNumber="1370">Metepisterna long, distinctly narrowed posteriad, at inner margin about 1.5 times longer than wide at the basal margin. Prosternum medially with 6 and close to apical margin with a row of 6 medium long setae (often broken resulting in insertion points difficult to observe). Prosternal process with 1 distinct seta.</paragraph>
<paragraph id="D44EBF23A818F11EFF5EFADDFCF49C14" blockId="9.[178,1085,1313,1393]" pageId="9" pageNumber="1370">Pro- and mesotarsomeres 2-4 of males distinctly dilated. Protarsomere 4 markedly, mesotarsomere 4 weakly bilobed. Protarsomeres 1-4 and mesotarsomeres 2-4 of males with biseriately arranged adhesive hairs on ventral surface.</paragraph>
<paragraph id="D44EBF23A818F11EFF5EFA7CFE169F69" blockId="9.[178,1085,1408,1548]" pageId="9" pageNumber="1370">
Median lobe of aedeagus (
<figureCitation id="4CCAA3A6A818F11EFE52FA7CFDDD9CF3" box="[446,558,1408,1430]" captionStart="Figs 50-55" captionStartId="29.[178,218,1492,1514]" captionTargetBox="[150,1110,191,1479]" captionTargetId="figure-0@29.[150,1112,189,1632]" captionTargetPageId="29" captionText="Figs 50-55: Anthracus sumatraensis nov.sp. Median lobe of aedoeagus, lateral and dorsal aspect. (50, 52, 53, 55) PT, (51, 54) HT." figureDoi="http://doi.org/10.5281/zenodo.6546843" httpUri="https://zenodo.org/record/6546843/files/figure.png" pageId="9" pageNumber="1370">Figs 50-55</figureCitation>
) with general appearance as figured. Apical plateau with shape rather variable, moderately long and moderately narrowed to apex (dorsal aspect), the latter somewhat thickened, with no distinct hook (lateral aspect). Internal structures composed of 3 larger subapical teeth, 1-2 smaller teeth in medial portion and 1-2 smaller teeth in basal half.
</paragraph>
</subSubSection>
<subSubSection id="9CEBECA8A818F11DFF5EF9E7FE9F98D8" lastPageId="10" lastPageNumber="1371" pageId="9" pageNumber="1370" type="discussion">
<paragraph id="D44EBF23A818F11DFF5EF9E7FE9F98D8" blockId="9.[178,1085,1563,1585]" lastBlockId="10.[178,1085,219,446]" lastPageId="10" lastPageNumber="1371" pageId="9" pageNumber="1370">
Comparisons:
<taxonomicName id="13F1C4A0A818F11EFE72F9E7FD669F54" authority="Jaeger, 2015" authorityName="Jaeger" authorityYear="2015" box="[414,661,1563,1585]" class="Insecta" family="Carabidae" genus="Anthracus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1370" phylum="Arthropoda" rank="species" species="sumatraensis" status="sp. nov.">
<emphasis id="E6856331A818F11EFE72F9E7FD669F54" box="[414,661,1563,1585]" italics="true" pageId="9" pageNumber="1370">Anthracus sumatraensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FDB6DE4AA818F11EFD73F9E7FD1A9F55" box="[671,745,1563,1584]" pageId="9" pageNumber="1370" rank="species">nov.sp.</taxonomicNameLabel>
is the only maculate species so far known from
<collectingRegion id="163571C1A81BF11DFEB4FF27FE5C9995" box="[344,431,219,240]" country="Indonesia" name="Sumatera Utara" pageId="10" pageNumber="1371">Sumatra</collectingRegion>
. It differs from
<taxonomicName id="13F1C4A0A81BF11DFDA3FF27FD249995" baseAuthorityName="ANDREWES" baseAuthorityYear="1936" box="[591,727,219,240]" class="Insecta" family="Carabidae" genus="Anthracus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="1371" phylum="Arthropoda" rank="species" species="nesophilus">
<emphasis id="E6856331A81BF11DFDA3FF27FD249995" box="[591,727,219,240]" italics="true" pageId="10" pageNumber="1371">A. nesophilus</emphasis>
</taxonomicName>
and
<taxonomicName id="13F1C4A0A81BF11DFCE5FF27FC709995" authority="Jaeger, 2015" authorityName="Jaeger" authorityYear="2015" box="[777,899,219,240]" class="Insecta" family="Carabidae" genus="Anthracus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="1371" phylum="Arthropoda" rank="species" species="javaensis" status="sp. nov.">
<emphasis id="E6856331A81BF11DFCE5FF27FC709995" box="[777,899,219,240]" italics="true" pageId="10" pageNumber="1371">A. javaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FDB6DE4AA81BF11DFC65FF27FC209995" box="[905,979,219,240]" pageId="10" pageNumber="1371" rank="species">nov.sp.</taxonomicNameLabel>
from
<collectingRegion id="163571C1A81BF11DFBFCFF27FBCE9995" box="[1040,1085,219,240]" country="Indonesia" name="Jawa Timur" pageId="10" pageNumber="1371">Java</collectingRegion>
by characters given under these species. From
<taxonomicName id="13F1C4A0A81BF11DFD4FFF04FCD7986B" authority="Jaeger, 2015" authorityName="Jaeger" authorityYear="2015" box="[675,804,248,270]" class="Insecta" family="Carabidae" genus="Anthracus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="1371" phylum="Arthropoda" rank="species" species="siamensis" status="sp. nov.">
<emphasis id="E6856331A81BF11DFD4FFF04FCD7986B" box="[675,804,248,270]" italics="true" pageId="10" pageNumber="1371">A. siamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FDB6DE4AA81BF11DFCC2FF04FC8B9868" box="[814,888,248,269]" pageId="10" pageNumber="1371" rank="species">nov.sp.</taxonomicNameLabel>
which is the most closely distributed species from SE Asian mainland,
<taxonomicName id="13F1C4A0A81BF11DFD20FEEAFC9D984E" authorityName="Jaeger" authorityYear="2015" box="[716,878,278,299]" class="Insecta" family="Carabidae" genus="Anthracus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="1371" phylum="Arthropoda" rank="species" species="sumatraensis">
<emphasis id="E6856331A81BF11DFD20FEEAFC9D984E" box="[716,878,278,299]" italics="true" pageId="10" pageNumber="1371">A. sumatraensis</emphasis>
</taxonomicName>
can be separated by the different external shape and internal structures of the median lobe of the aedeagus (
<figureCitation id="4CCAA3A6A81BF11DFF55FEACFEDD9803" box="[185,302,336,358]" captionStart="Figs 50-55" captionStartId="29.[178,218,1492,1514]" captionTargetBox="[150,1110,191,1479]" captionTargetId="figure-0@29.[150,1112,189,1632]" captionTargetPageId="29" captionText="Figs 50-55: Anthracus sumatraensis nov.sp. Median lobe of aedoeagus, lateral and dorsal aspect. (50, 52, 53, 55) PT, (51, 54) HT." figureDoi="http://doi.org/10.5281/zenodo.6546843" httpUri="https://zenodo.org/record/6546843/files/figure.png" pageId="10" pageNumber="1371">Figs 50-55</figureCitation>
,
<figureCitation id="4CCAA3A6A81BF11DFED2FEACFE889803" box="[318,379,336,358]" captionStart="Figs 62-67" captionStartId="31.[178,218,1520,1542]" captionTargetBox="[150,1112,174,1495]" captionTargetId="figure-0@31.[150,1112,174,1617]" captionTargetPageId="31" captionText="Figs 62-67: Median lobe of aedoeagus, lateral aspect. Anthracus javaensis nov.sp. (62) HT and A. siamensis nov.sp. (63, 64, 67) PT, Laos, Pakkading, (65) HT, (66) Thailand, Ban Khoun." figureDoi="http://doi.org/10.5281/zenodo.6546849" httpUri="https://zenodo.org/record/6546849/files/figure.png" pageId="10" pageNumber="1371">63-67</figureCitation>
,
<figureCitation id="4CCAA3A6A81BF11DFE67FEACFE3A9803" box="[395,457,336,358]" captionStart="Figs 68-73" captionStartId="32.[178,218,1420,1442]" captionTargetBox="[150,1112,159,1602]" captionTargetId="figure-0@32.[150,1112,159,1602]" captionTargetPageId="32" captionText="Figs 68-73: Median lobe of aedoeagus, dorsal aspect. Anthracus javaensis nov.sp. (68) HT and A. siamensis nov.sp. (69) PT, Thailand, Umphang, (70, 73) PT, Laos, Pakkading, (71) HT, (72) Thailand, Ban Khoun." figureDoi="http://doi.org/10.5281/zenodo.5194872" httpUri="https://zenodo.org/record/5194872/files/figure.png" pageId="10" pageNumber="1371">69-73</figureCitation>
), the elytra, which are broader relative to pronotum (ratio EW/PW&gt;1.40, in
<taxonomicName id="13F1C4A0A81BF11DFE9AFE91FE0598E6" authorityName="Jaeger" authorityYear="2015" box="[374,502,365,387]" class="Insecta" family="Carabidae" genus="Anthracus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="1371" phylum="Arthropoda" rank="species" species="siamensis">
<emphasis id="E6856331A81BF11DFE9AFE91FE0598E6" box="[374,502,365,387]" italics="true" pageId="10" pageNumber="1371">A. siamensis</emphasis>
</taxonomicName>
&lt;1.39), and the colour of head and pronotum which is usually paler and/or the elytral macula which are only indistinct and not clearly marked as in
<taxonomicName id="13F1C4A0A81BF11DFF0BFE54FE9598D8" authorityName="Jaeger" authorityYear="2015" box="[231,358,424,445]" class="Insecta" family="Carabidae" genus="Anthracus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="1371" phylum="Arthropoda" rank="species" species="siamensis">
<emphasis id="E6856331A81BF11DFF0BFE54FE9598D8" box="[231,358,424,445]" italics="true" pageId="10" pageNumber="1371">A. siamensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="9CEBECA8A81BF11DFF5EFE31FDF69B65" pageId="10" pageNumber="1371" type="distribution">
<paragraph id="D44EBF23A81BF11DFF5EFE31FDF69B65" blockId="10.[178,1085,461,512]" pageId="10" pageNumber="1371">
Distribution
<taxonomicName id="13F1C4A0A81BF11DFE70FE31FDCC9886" authority="Jaeger, 2015" authorityName="Jaeger" authorityYear="2015" box="[412,575,461,483]" class="Insecta" family="Carabidae" genus="Anthracus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="1371" phylum="Arthropoda" rank="species" species="sumatraensis" status="sp. nov.">
<emphasis id="E6856331A81BF11DFE70FE31FDCC9886" box="[412,575,461,483]" italics="true" pageId="10" pageNumber="1371">A. sumatraensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FDB6DE4AA81BF11DFDABFE31FD629887" box="[583,657,461,482]" pageId="10" pageNumber="1371" rank="species">nov.sp.</taxonomicNameLabel>
is so far known only from the
<typeStatus id="0B4A0181A81BF11DFC3BFE31FBF09887" box="[983,1027,461,482]" pageId="10" pageNumber="1371">type</typeStatus>
locality in northern
<collectingRegion id="163571C1A81BF11DFEBDFE16FE56989A" box="[337,421,490,511]" country="Indonesia" name="Sumatera Utara" pageId="10" pageNumber="1371">Sumatra</collectingRegion>
(
<figureCitation id="4CCAA3A6A81BF11DFE5EFE16FE0A9B65" box="[434,505,490,512]" captionStart="Fig 80" captionStartId="34.[178,210,909,931]" captionTargetBox="[266,997,212,896]" captionTargetId="figure-65@34.[266,997,212,896]" captionTargetPageId="34" captionText="Fig 80: Distribution: A. nesophilus (ANDREWES) (triangles), A. javaensis nov.sp. (white star), A. sumatraensis nov.sp. (square) A. philippinensis nov.sp. (circle), A. siamensis nov.sp. (black star)." figureDoi="http://doi.org/10.5281/zenodo.5193681" httpUri="https://zenodo.org/record/5193681/files/figure.png" pageId="10" pageNumber="1371">Fig 80</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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