654 lines
76 KiB
XML
654 lines
76 KiB
XML
<document id="4B692B2631FADBB2E04A53F228C5FE45" ID-DOI="10.1080/00222930802124057" ID-ISSN="1464-5262" ID-Zenodo-Dep="4593937" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615397885932" checkinUser="carolina" docAuthor="Chatzimanolis, Stylianos" docDate="2008" docId="5C09879C77691C067617FDC85BD56BC8" docLanguage="en" docName="JNATHIST.42.25-26.1765-1792.pdf" docOrigin="Journal of Natural History 42 (25 - 26)" docSource="http://dx.doi.org/10.1080/00222930802124057" docStyle="DocumentStyle:FCA295E821436EF9307F50C7AC261B53.5:JNatHist.2008-2012.journal_article.1cover" docStyleId="FCA295E821436EF9307F50C7AC261B53" docStyleName="JNatHist.2008-2012.journal_article.1cover" docStyleVersion="5" docTitle="Isanopus ashei Chatzimanolis 2008, sp. n." docType="treatment" docVersion="3" lastPageNumber="1779" masterDocId="A030FFE477601C09744FFFB0586B6958" masterDocTitle="A revision of the neotropical beetle genus Isanopus Sharp, 1876 (Coleoptera: Staphylinidae: Staphylinini)" masterLastPageNumber="1792" masterPageNumber="1765" pageNumber="1773" updateTime="1698882181256" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="1D81CC265C2C7EB4F009BF4DC0E13665">A revision of the neotropical beetle genus Isanopus Sharp, 1876 (Coleoptera: Staphylinidae: Staphylinini)</mods:title>
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<mods:namePart id="EF0C51499915611728C3E0D7739C333E">Chatzimanolis, Stylianos</mods:namePart>
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<mods:title id="CF50532BB097ACBBBEC0D196F1852ED9">Journal of Natural History</mods:title>
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<mods:part id="E723677A1308008F7F3D38D7A96DA2B8">
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<mods:date id="C053821DB62DA37F9C5B8E9B905C0882">2008</mods:date>
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<mods:number id="53273EFC2E224F98F5A9C408ADE63BA4">2008-07-31</mods:number>
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<mods:number id="DDF05509D8CCEA4E3A11C6071563309E">42</mods:number>
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<mods:number id="F11B78D7D1830676B3515E26C02B0958">25 - 26</mods:number>
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<mods:start id="BDEA5BA142F4B1C0A2C647B38C3D8486">1765</mods:start>
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<mods:end id="0951AF082D3CCC3720DBEC5443EB519C">1792</mods:end>
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<mods:url id="02FF0BFC417FE1A02625FFC04982AE46">http://dx.doi.org/10.1080/00222930802124057</mods:url>
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<mods:identifier id="22ACAF0FA96900D4D44079BFC3247DC2" type="DOI">10.1080/00222930802124057</mods:identifier>
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<mods:identifier id="ACDEC79FD1ED684F237C4579519BB689" type="ISSN">1464-5262</mods:identifier>
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<mods:identifier id="38E0348EC854571067DEA2988A2E30AB" type="Zenodo-Dep">4593937</mods:identifier>
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<treatment id="5C09879C77691C067617FDC85BD56BC8" LSID="urn:lsid:plazi:treatment:5C09879C77691C067617FDC85BD56BC8" httpUri="http://treatment.plazi.org/id/5C09879C77691C067617FDC85BD56BC8" lastPageId="15" lastPageNumber="1779" pageId="9" pageNumber="1773">
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<subSubSection id="9CBA650177691C007617FDC85B296BC8" box="[600,834,632,656]" pageId="9" pageNumber="1773" type="nomenclature">
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<paragraph id="D41F368A77691C007617FDC85B296BC8" blockId="9.[582,855,632,689]" box="[600,834,632,656]" pageId="9" pageNumber="1773">
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<heading id="8F5781E677691C007617FDC85B296BC8" box="[600,834,632,656]" centered="true" fontSize="10" level="2" pageId="9" pageNumber="1773" reason="3">
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<taxonomicName id="13A04D0977691C007617FDC85A906BC8" ID-CoL="3Q34P" authority="Chatzimanolis, 2008" authorityName="Chatzimanolis" authorityYear="2008" box="[600,763,632,656]" class="Insecta" family="Staphylinidae" genus="Isanopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1773" phylum="Arthropoda" rank="species" species="ashei" status="sp. nov.">
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<emphasis id="E6D4EA9877691C007617FDC85A906BC8" bold="true" box="[600,763,632,656]" italics="true" pageId="9" pageNumber="1773">Isanopus ashei</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="FDE757E377691C00774CFDC85B296BC8" box="[771,834,632,656]" pageId="9" pageNumber="1773" rank="species">sp. n.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="9CBA650177691C067609FD295BD56BC8" lastPageId="15" lastPageNumber="1779" pageId="9" pageNumber="1773" type="description">
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<paragraph id="D41F368A77691C007609FD295B3C6BE9" blockId="9.[582,855,632,689]" box="[582,855,665,689]" pageId="9" pageNumber="1773">
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(
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<figureCitation id="4C9B2A0F77691C007602FD295AA66BE9" box="[589,717,665,689]" captionStart="Figures 2" captionStartId="6.[146,222,1349,1371]" captionTargetBox="[149,1102,175,1302]" captionTargetId="figure-63@6.[146,1104,165,1312]" captionTargetPageId="6" captionText="Figures 2. Isanopus species, dorsum of head and pronotum. (A) I. ashei. (B) I. sahlbergi. (C) I. hinojosai. (D) I. eptaskouros." figureDoi="http://doi.org/10.5281/zenodo.4593941" httpUri="https://zenodo.org/record/4593941/files/figure.png" pageId="9" pageNumber="1773">Figures 2A</figureCitation>
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, 4A, 5A–C)
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</paragraph>
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<paragraph id="D41F368A77691C00749AFD5A59296A5A" blockId="9.[213,322,746,770]" box="[213,322,746,770]" pageId="9" pageNumber="1773">
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<emphasis id="E6D4EA9877691C00749AFD5A59296A5A" box="[213,322,746,770]" italics="true" pageId="9" pageNumber="1773">Diagnosis</emphasis>
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</paragraph>
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<paragraph id="D41F368A77691C00749AFCA259926AF3" blockId="9.[213,1169,786,939]" pageId="9" pageNumber="1773">
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This species along with
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<taxonomicName id="13A04D0977691C0075AAFCA25A0A6A72" authorityName="Chatzimanolis" authorityYear="2008" box="[485,609,786,810]" class="Insecta" family="Staphylinidae" genus="Isanopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1773" phylum="Arthropoda" rank="species" species="hinojosai">
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<emphasis id="E6D4EA9877691C0075AAFCA25A0A6A72" box="[485,609,786,810]" italics="true" pageId="9" pageNumber="1773">I. hinojosai</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="13A04D0977691C0076D3FCA25A976A72" authorityName="Sharp" authorityYear="1884" box="[668,764,786,810]" class="Insecta" family="Staphylinidae" genus="Isanopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1773" phylum="Arthropoda" rank="species" species="sallaei">
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<emphasis id="E6D4EA9877691C0076D3FCA25A976A72" box="[668,764,786,810]" italics="true" pageId="9" pageNumber="1773">I. sallaei</emphasis>
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</taxonomicName>
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has fewer and less dense punctures on the head and pronotum than other species of
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<taxonomicName id="13A04D0977691C00776EFC835BE96A13" authorityName="Sharp" authorityYear="1876" box="[801,898,819,843]" class="Insecta" family="Staphylinidae" genus="Isanopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1773" phylum="Arthropoda" rank="genus">
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<emphasis id="E6D4EA9877691C00776EFC835BE96A13" box="[801,898,819,843]" italics="true" pageId="9" pageNumber="1773">Isanopus</emphasis>
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</taxonomicName>
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.
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<taxonomicName id="13A04D0977691C0077DAFC835C526A13" authorityName="Chatzimanolis" authorityYear="2008" box="[917,1081,819,843]" class="Insecta" family="Staphylinidae" genus="Isanopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1773" phylum="Arthropoda" rank="species" species="ashei">
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<emphasis id="E6D4EA9877691C0077DAFC835C526A13" box="[917,1081,819,843]" italics="true" pageId="9" pageNumber="1773">Isanopus ashei</emphasis>
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</taxonomicName>
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can be distinguished from all other species of
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<taxonomicName id="13A04D0977691C0076F8FCE35B736A33" authorityName="Sharp" authorityYear="1876" box="[695,792,851,875]" class="Insecta" family="Staphylinidae" genus="Isanopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1773" phylum="Arthropoda" rank="genus">
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<emphasis id="E6D4EA9877691C0076F8FCE35B736A33" box="[695,792,851,875]" italics="true" pageId="9" pageNumber="1773">Isanopus</emphasis>
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</taxonomicName>
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by the uniform colour of the abdomen (dark orange to rufous) in all visible segments and the shape of the aedeagus (
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<figureCitation id="4C9B2A0F77691C007503FC2359DF6AF3" box="[332,436,915,939]" captionStart="Figure 5" captionStartId="11.[213,280,680,702]" captionTargetBox="[213,1171,165,638]" captionTargetId="figure-227@11.[213,1171,165,643]" captionTargetPageId="11" captionText="Figure 5. Aedeagus of I. ashei. (A) Dorsal view. (B) Lateral view. (C) Detail of paramere, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4593945" httpUri="https://zenodo.org/record/4593945/files/figure.png" pageId="9" pageNumber="1773">Figures 5</figureCitation>
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A–C).
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</paragraph>
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<paragraph id="D41F368A77691C00749AFC54593F6AA4" blockId="9.[213,340,996,1020]" box="[213,340,996,1020]" pageId="9" pageNumber="1773">
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<emphasis id="E6D4EA9877691C00749AFC54593F6AA4" box="[213,340,996,1020]" italics="true" pageId="9" pageNumber="1773">Description</emphasis>
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</paragraph>
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<paragraph id="D41F368A77691C00749AFBBD59D66D7D" blockId="9.[213,445,1037,1061]" box="[213,445,1037,1061]" pageId="9" pageNumber="1773">
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Length
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<quantity id="13589B6F77691C007561FBBD59D26D7D" box="[302,441,1037,1061]" metricMagnitude="-2" metricUnit="m" metricValue="1.225" metricValueMax="1.3" metricValueMin="1.15" pageId="9" pageNumber="1773" unit="mm" value="12.25" valueMax="13.0" valueMin="11.5">11.5–13 mm</quantity>
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.
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</paragraph>
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<paragraph id="D41F368A77691C00749AFBEE5B536D8E" blockId="9.[213,1169,1118,1238]" pageId="9" pageNumber="1773">
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<emphasis id="E6D4EA9877691C00749AFBEE59326D2E" box="[213,345,1118,1142]" italics="true" pageId="9" pageNumber="1773">Colouration</emphasis>
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||
. Colour of head and pronotum brown to black. Antenna brown. Scutellum dark brown to black. Elytra brilliant metallic blue, greenish blue or purplish blue. Abdomen dark orange to rufous, anterior half of every segment slightly darker than posterior half. Legs dark brown.
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</paragraph>
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<paragraph id="D41F368A77691C03749AFAA05A4E6DE2" blockId="9.[213,1169,1295,1706]" lastBlockId="10.[146,1103,1158,1210]" lastPageId="10" lastPageNumber="1774" pageId="9" pageNumber="1773">
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<emphasis id="E6D4EA9877691C00749AFAA0597A6C70" box="[213,273,1296,1320]" italics="true" pageId="9" pageNumber="1773">Head</emphasis>
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. Head with relatively small brown microsetae and longer black macrosetae along borders of head; head subquadrate, width: length ratio51.18. Surface of epicranium glossy, with no micropunctures. Epicranium flat, with sparse mediumsized umbilicate setose punctures laterally, no punctures medially, setose punctures irregularly arranged in 2–3 rows anterior-posteriorly, punctures smaller around eyes, distance between punctures various, from 0.5–1 times width of punctures. Eyes medium, length of eyes/length of head50.46, distance between eyes as wide as 1.8 times length of eye. Postgena with large, longitudinally elongate punctures (about 3– 4 punctures/
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<quantity id="13589B6F77691C00752CF9A159D26F71" box="[355,441,1553,1577]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="1773" unit="mm" value="0.5">0.5 mm</quantity>
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), punctures not uniformly distributed. Antennomeres 1–3 with long macrosetae, antennomere 1 gradually club-like thickened, antennomeres 1–11 longer than wide, antennomere 1 about twice as long as antennomere 2, antennomere 3 1.5 times longer than antennomere 2, antennomere 4 slightly shorter than antennomere 3, antennomeres 5–10 almost identical, 3.5 times longer Figure 4. Antennae. (A)
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<taxonomicName id="13A04D09776A1C0375D3FB37598C6DC4" authorityName="Chatzimanolis" authorityYear="2008" box="[412,487,1159,1180]" class="Insecta" family="Staphylinidae" genus="Isanopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="1774" phylum="Arthropoda" rank="species" species="ashei">
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<emphasis id="E6D4EA98776A1C0375D3FB37598C6DC4" box="[412,487,1159,1180]" italics="true" pageId="10" pageNumber="1774">I. ashei</emphasis>
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</taxonomicName>
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. (B)
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<taxonomicName id="13A04D09776A1C03766EFB375A126DC4" authorityName="Sharp" authorityYear="1884" box="[545,633,1159,1180]" class="Insecta" family="Staphylinidae" genus="Isanopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="1774" phylum="Arthropoda" rank="species" species="sallaei">
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<emphasis id="E6D4EA98776A1C03766EFB375A126DC4" box="[545,633,1159,1180]" italics="true" pageId="10" pageNumber="1774">I. sallaei</emphasis>
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</taxonomicName>
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. (C)
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<taxonomicName id="13A04D09776A1C0376FCFB375B286DC4" authorityName="Chatzimanolis" authorityYear="2008" box="[691,835,1159,1180]" class="Insecta" family="Staphylinidae" genus="Isanopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="1774" phylum="Arthropoda" rank="species" species="eptaskouros">
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<emphasis id="E6D4EA98776A1C0376FCFB375B286DC4" box="[691,835,1159,1180]" italics="true" pageId="10" pageNumber="1774">I. eptaskouros</emphasis>
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</taxonomicName>
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. (D)
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<taxonomicName id="13A04D09776A1C0377CFFB375C6E6DC4" authorityName="Sharp" authorityYear="1876" box="[896,1029,1159,1180]" class="Insecta" family="Staphylinidae" genus="Isanopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="1774" phylum="Arthropoda" rank="species" species="tenuicornis">
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<emphasis id="E6D4EA98776A1C0377CFFB375C6E6DC4" box="[896,1029,1159,1180]" italics="true" pageId="10" pageNumber="1774">I. tenuicornis</emphasis>
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</taxonomicName>
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. (E)
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<taxonomicName id="13A04D09776A1C037070FB3758816DE1" authorityName="Chatzimanolis" authorityYear="2008" class="Insecta" family="Staphylinidae" genus="Isanopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="1774" phylum="Arthropoda" rank="species" species="hinojosai">
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<emphasis id="E6D4EA98776A1C037070FB3758816DE1" italics="true" pageId="10" pageNumber="1774">I. hinojosai</emphasis>
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</taxonomicName>
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. Scale in Figure 4E
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<quantity id="13589B6F776A1C0375FBFB145A496DE2" box="[436,546,1188,1210]" metricMagnitude="-2" metricUnit="m" metricValue="5.138" pageId="10" pageNumber="1774" unit="mm" value="51.38">51.38 mm</quantity>
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.
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</paragraph>
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<paragraph id="D41F368A776A1C0374DDFB545BCB6C44" blockId="10.[146,1103,1252,1308]" pageId="10" pageNumber="1774">than wide, antennomere 11 subequal to previous antennomeres. Neck dorsally with no punctures or micropunctures, laterally with 1–2 small punctures.</paragraph>
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<paragraph id="D41F368A776A1C0374DDFAE45A006FF2" blockId="10.[146,1103,1363,1706]" pageId="10" pageNumber="1774">
|
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<emphasis id="E6D4EA98776A1C0374DDFAE4588F6C34" box="[146,228,1364,1388]" italics="true" pageId="10" pageNumber="1774">Thorax</emphasis>
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. Pronotum longer than wide, width: length ratio50.84; antero-lateral angles obtusely rounded, curved. Lateral margins of pronotum concave in dorsal aspect, pronotum broadest in apical 1/4 and narrower at basal angles. Surface of pronotum flat, glossy, with no micropunctures. Setose punctures absent on disc of pronotum, laterally present in 3–4 rows, distance between rows smaller towards lateral ends; punctures of medium size (about four punctures/
|
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<quantity id="13589B6F776A1C037694FA435B5A6F53" box="[731,817,1523,1547]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="1774" unit="mm" value="0.5">0.5 mm</quantity>
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||
); least distance between punctures for each row various, 1–2 times width of puncture for most. Pronotum with brown microsetae directed postero-medially in lateral and anterior areas and posteriorly in basal half, and few longer black macrosetae along borders. Scutellum prominent, with brown microsetae and sparse punctures, least distance between punctures 0.5–1 times width of puncture.
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||
</paragraph>
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<caption id="80DF6602776B1C02749AFD18593A6B83" ID-DOI="http://doi.org/10.5281/zenodo.4593945" ID-Zenodo-Dep="4593945" httpUri="https://zenodo.org/record/4593945/files/figure.png" pageId="11" pageNumber="1775" startId="11.[213,280,680,702]" targetBox="[213,1171,165,638]" targetPageId="11">
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<paragraph id="D41F368A776B1C02749AFD18593A6B83" blockId="11.[213,1169,680,731]" pageId="11" pageNumber="1775">
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||
Figure 5. Aedeagus of
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<taxonomicName id="13A04D09776B1C027585FD185A7E6BE5" authorityName="Chatzimanolis" authorityYear="2008" box="[458,533,680,701]" class="Insecta" family="Staphylinidae" genus="Isanopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1775" phylum="Arthropoda" rank="species" species="ashei">
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<emphasis id="E6D4EA98776B1C027585FD185A7E6BE5" box="[458,533,680,701]" italics="true" pageId="11" pageNumber="1775">I. ashei</emphasis>
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</taxonomicName>
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. (A) Dorsal view. (B) Lateral view. (C) Detail of paramere, dorsal view.
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</paragraph>
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</caption>
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<paragraph id="D41F368A776B1C0274B0FCA35A066A33" blockId="11.[213,1169,787,875]" pageId="11" pageNumber="1775">
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Elytra subequal in length to pronotum, with medium setose uniform punctures (5–6 punctures/
|
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), covered with yellowish brown microsetae and longer black macrosetae along borders of elytra.
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||
</paragraph>
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<paragraph id="D41F368A776B1C02749AFC13590D6AA3" blockId="11.[213,1169,931,1019]" pageId="11" pageNumber="1775">
|
||
<emphasis id="E6D4EA98776B1C02749AFC1359616AE3" box="[213,266,931,955]" italics="true" pageId="11" pageNumber="1775">Legs</emphasis>
|
||
. Protarsus enlarged; meso and metatarsi elongate, enlarged, tarsomeres 1–4, asymmetrical, dilated distally. Legs almost completely covered with long yellow and brown setae.
|
||
</paragraph>
|
||
<paragraph id="D41F368A776B1C02749AFB835A126C12" blockId="11.[213,1169,1075,1354]" pageId="11" pageNumber="1775">
|
||
<emphasis id="E6D4EA98776B1C02749AFB8359566D13" box="[213,317,1075,1099]" italics="true" pageId="11" pageNumber="1775">Abdomen</emphasis>
|
||
. Abdominal terga III–V with tergal basal and no subbasal (arch-like) carina. Terga III–VII with uniform punctuation patterns, densely covered with punctures and yellowish brown setae, punctures decreasing in size gradually from III–VII. Abdominal sterna III–IV with moderate transverse impressions, V with faint transverse impression, shining. Integument surface appearing iridescent in strong direct light at moderate to high magnification. Sterna III–VII densely punctuated, punctures decreasing in size gradually from III–VII, with yellowish brown setae. Lateral tergal sclerites of the abdominal segment IX long and straight, covered with long black macrosetae.
|
||
</paragraph>
|
||
<paragraph id="D41F368A776B1C02749AFA3359276CA2" blockId="11.[213,1169,1410,1530]" pageId="11" pageNumber="1775">
|
||
<emphasis id="E6D4EA98776B1C02749AFA335A4B6CC3" box="[213,544,1411,1435]" italics="true" pageId="11" pageNumber="1775">Secondary sexual structures</emphasis>
|
||
. Male: distal margin of sternum VII with slight emargination medially; sternum VIII with deep V-shaped emargination medially; sternum IX with small emargination medially. Females: with no obvious sexual structures.
|
||
</paragraph>
|
||
<paragraph id="D41F368A776B1C02749AF9835CFA6FF2" blockId="11.[213,1169,1586,1706]" pageId="11" pageNumber="1775">
|
||
<emphasis id="E6D4EA98776B1C02749AF98359546F13" box="[213,319,1587,1611]" italics="true" pageId="11" pageNumber="1775">Aedeagus</emphasis>
|
||
. As in
|
||
<figureCitation id="4C9B2A0F776B1C0275DEF982599C6F12" box="[401,503,1586,1610]" captionStart="Figure 5" captionStartId="11.[213,280,680,702]" captionTargetBox="[213,1171,165,638]" captionTargetId="figure-227@11.[213,1171,165,643]" captionTargetPageId="11" captionText="Figure 5. Aedeagus of I. ashei. (A) Dorsal view. (B) Lateral view. (C) Detail of paramere, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4593945" httpUri="https://zenodo.org/record/4593945/files/figure.png" pageId="11" pageNumber="1775">Figures 5</figureCitation>
|
||
A–C; paramere extremely reduced; paramere: median lobe ratio50.23. Paramere in dorsal view fairly wide but expanding to two rounded apical processes, with a wide U-shaped emargination between them; in lateral outline almost straight; with no apparent sensory spinules. Median lobe in dorsal view wide,
|
||
</paragraph>
|
||
<paragraph id="D41F368A776C1C0574DDFF185AFC69B9" blockId="12.[146,1103,168,225]" pageId="12" pageNumber="1776">converging to narrow apex; in lateral view becoming narrower and curved from middle to apex, apex with hook-like process.</paragraph>
|
||
<paragraph id="D41F368A776C1C0574DDFE9459E16D7D" blockId="12.[146,1103,292,1061]" pageId="12" pageNumber="1776">
|
||
<materialsCitation id="64C83CD7776C1C0574DDFE945B896825" collectionCode="R" collectorName="J. S. Ashe & Brooks" county="La Culebra" elevation="1490" location="Male" municipality="Chiriqui" pageId="12" pageNumber="1776" specimenCode="SM0016597" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="E6D4EA98776C1C0574DDFE9458926864" box="[146,249,292,316]" italics="true" pageId="12" pageNumber="1776">
|
||
<typeStatus id="0B1B8828776C1C0574DDFE9458926864" box="[146,249,292,316]" pageId="12" pageNumber="1776" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
.
|
||
<location id="D17F6051776C1C057545FE94592C6864" LSID="urn:lsid:plazi:treatment:5C09879C77691C067617FDC85BD56BC8:D17F6051776C1C057545FE94592C6864" box="[266,327,292,316]" county="La Culebra" municipality="Chiriqui" name="Male" pageId="12" pageNumber="1776">Male</location>
|
||
, with labels as follows: ‘‘
|
||
<location id="D17F6051776C1C057624FE945AA56864" LSID="urn:lsid:plazi:treatment:5C09879C77691C067617FDC85BD56BC8:D17F6051776C1C057624FE945AA56864" box="[619,718,292,316]" county="La Culebra" municipality="Chiriqui" name="Panama" pageId="12" pageNumber="1776">Panama</location>
|
||
,
|
||
<collectingMunicipality id="347BACF0776C1C057693FE945B556864" box="[732,830,292,316]" pageId="12" pageNumber="1776">Chiriqui</collectingMunicipality>
|
||
,
|
||
<locationDeviation id="C5AD2FA8776C1C057703FE945C4A6864" box="[844,1057,292,316]" location="Boquete" pageId="12" pageNumber="1776">
|
||
<quantity id="13589B6F776C1C057703FE945BFC6864" box="[844,919,292,316]" metricMagnitude="3" metricUnit="m" metricValue="5.6" pageId="12" pageNumber="1776" unit="km" value="5.6">5.6 km</quantity>
|
||
N Boquete
|
||
</locationDeviation>
|
||
,
|
||
<collectingCounty id="3D7E4E06776C1C057060FE9458866804" pageId="12" pageNumber="1776">La Culebra</collectingCounty>
|
||
trail,
|
||
<quantity id="13589B6F776C1C05757CFEF459ED6804" box="[307,390,324,348]" metricMagnitude="3" metricUnit="m" metricValue="1.49" pageId="12" pageNumber="1776" unit="m" value="1490.0">
|
||
<elevation id="5F8DD1B9776C1C05757CFEF459ED6804" box="[307,390,324,348]" metricMagnitude="3" metricUnit="m" metricValue="1.49" pageId="12" pageNumber="1776" unit="m" value="1490.0">1490 m</elevation>
|
||
</quantity>
|
||
, 8
|
||
<emphasis id="E6D4EA98776C1C0575D0FEF559C26805" bold="true" box="[415,425,325,349]" italics="true" pageId="12" pageNumber="1776">°</emphasis>
|
||
499230N, 82
|
||
<emphasis id="E6D4EA98776C1C05767FFEF55A516805" bold="true" box="[560,570,325,349]" italics="true" pageId="12" pageNumber="1776">°</emphasis>
|
||
259180W, vi.15.1996,
|
||
<collectorName id="7955535C776C1C057762FEF45BFC6804" box="[813,919,324,348]" pageId="12" pageNumber="1776">J.S. Ashe</collectorName>
|
||
and
|
||
<collectionCode id="B2B1AE4F776C1C05779EFEF45B826804" box="[977,1001,324,348]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="1776">R</collectionCode>
|
||
.
|
||
<collectorName id="7955535C776C1C0577BAFEF45C216804" box="[1013,1098,324,348]" pageId="12" pageNumber="1776">Brooks</collectorName>
|
||
, PAN1AB96 159, ex: treefall/slash litter’’/[barcode label] ‘‘
|
||
<specimenCode id="84069EF1776C1C05777AFED55BAE6825" box="[821,965,357,381]" collectionCode="SM" country="Malaysia" name="Sarawak Museum" pageId="12" pageNumber="1776">SM0016597</specimenCode>
|
||
’’/‘‘
|
||
</materialsCitation>
|
||
<materialsCitation id="64C83CD7776C1C0577ADFED55A3C6886" county="Lawrence" location="Chatzimanolis" municipality="University of Kansas" pageId="12" pageNumber="1776" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus id="0B1B8828776C1C0577ADFED55C246825" box="[994,1103,357,381]" pageId="12" pageNumber="1776" type="holotype">Holotype</typeStatus>
|
||
<taxonomicName id="13A04D09776C1C0574DDFE36598668C5" authority="Chatzimanolis" authorityName="Chatzimanolis" authorityYear="2008" box="[146,493,389,414]" class="Insecta" family="Staphylinidae" genus="Isanopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="1776" phylum="Arthropoda" rank="species" species="ashei">
|
||
<emphasis id="E6D4EA98776C1C0574DDFE36595268C6" box="[146,313,390,414]" italics="true" pageId="12" pageNumber="1776">Isanopus ashei</emphasis>
|
||
Chatzimanolis
|
||
</taxonomicName>
|
||
, design.
|
||
<location id="D17F6051776C1C057614FE355A9468C5" LSID="urn:lsid:plazi:treatment:5C09879C77691C067617FDC85BD56BC8:D17F6051776C1C057614FE355A9468C5" box="[603,767,389,413]" county="Lawrence" municipality="University of Kansas" name="Chatzimanolis" pageId="12" pageNumber="1776">Chatzimanolis</location>
|
||
2007’’.
|
||
<location id="D17F6051776C1C05772AFE355B1568C5" LSID="urn:lsid:plazi:treatment:5C09879C77691C067617FDC85BD56BC8:D17F6051776C1C05772AFE355B1568C5" box="[869,894,389,413]" county="Lawrence" municipality="University of Kansas" name="In" pageId="12" pageNumber="1776">In</location>
|
||
the collection of
|
||
<location id="D17F6051776C1C0574DDFE16599668E6" LSID="urn:lsid:plazi:treatment:5C09879C77691C067617FDC85BD56BC8:D17F6051776C1C0574DDFE16599668E6" box="[146,509,422,446]" county="Lawrence" municipality="University of Kansas" name="Snow Entomological Collection" pageId="12" pageNumber="1776">Snow Entomological Collection</location>
|
||
,
|
||
<location id="D17F6051776C1C057647FE165B7768E6" LSID="urn:lsid:plazi:treatment:5C09879C77691C067617FDC85BD56BC8:D17F6051776C1C057647FE165B7768E6" box="[520,796,422,446]" county="Lawrence" municipality="University of Kansas" name="Division of Entomology" pageId="12" pageNumber="1776">Division of Entomology</location>
|
||
,
|
||
<location id="D17F6051776C1C057768FE165C2D68E6" LSID="urn:lsid:plazi:treatment:5C09879C77691C067617FDC85BD56BC8:D17F6051776C1C057768FE165C2D68E6" box="[807,1094,422,446]" county="Lawrence" municipality="University of Kansas" name="Natural History Museum" pageId="12" pageNumber="1776">Natural History Museum</location>
|
||
/ BRC,
|
||
<collectingMunicipality id="347BACF0776C1C057492FE7659BC6886" box="[221,471,454,478]" pageId="12" pageNumber="1776">University of Kansas</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="3D7E4E06776C1C0575AAFE765A3C6886" box="[485,599,454,478]" pageId="12" pageNumber="1776">Lawrence</collectingCounty>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="64C83CD7776C1C05762AFE765B8968A7" collectionCode="SEMC" pageId="12" pageNumber="1776" specimenCode="SM0016596, SM0016598, SM0016599" specimenCount="1" typeStatus="paratype">
|
||
<typeStatus id="0B1B8828776C1C05762AFE765AB36886" box="[613,728,454,478]" pageId="12" pageNumber="1776" type="paratype">Paratypes</typeStatus>
|
||
, 13; with locality data as above and barcode labels:
|
||
<specimenCode id="84069EF1776C1C0575CDFE575A6068A7" box="[386,523,487,511]" collectionCode="SM" country="Malaysia" name="Sarawak Museum" pageId="12" pageNumber="1776">SM0016596</specimenCode>
|
||
,
|
||
<specimenCode id="84069EF1776C1C057654FE575ACF68A7" box="[539,676,487,511]" collectionCode="SM" country="Malaysia" name="Sarawak Museum" pageId="12" pageNumber="1776">SM0016598</specimenCode>
|
||
,
|
||
<specimenCode id="84069EF1776C1C0576FBFE575B5168A7" box="[692,826,487,511]" collectionCode="SM" country="Malaysia" name="Sarawak Museum" pageId="12" pageNumber="1776">SM0016599</specimenCode>
|
||
(
|
||
<quantity id="13589B6F776C1C057701FE575B1668A7" box="[846,893,487,511]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="12" pageNumber="1776" unit="in" value="3.0">3 in</quantity>
|
||
<collectionCode id="B2B1AE4F776C1C0577C7FE575BB168A7" box="[904,986,487,511]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="12" pageNumber="1776" type="Museum">SEMC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="64C83CD7776C1C0577BDFE575C216B38" collectionCode="SEMC" collectorName="J. S. Ashe & Z. Falin & Hinojosa" country="Costa Rica" elevation="2070" latitude="9.46565" location="Chirripo Natl. Park" longLatPrecision="1" longitude="-83.567314" pageId="12" pageNumber="1776" specimenCode="SM0607677, SM0607675, SM0607676" specimenCount="1" stateProvince="San Jose" typeStatus="paratype">
|
||
<collectingCountry id="ACB7761A776C1C0577BDFE5758B76B47" name="Costa Rica" pageId="12" pageNumber="1776">COSTA RICA</collectingCountry>
|
||
:
|
||
<collectingRegion id="1664F868776C1C0574A5FDB759F16B47" box="[234,410,519,543]" country="Costa Rica" name="San Jose" pageId="12" pageNumber="1776">San Jose Prov.</collectingRegion>
|
||
,
|
||
<locationDeviation id="C5AD2FA8776C1C0575E6FDB75B566B47" box="[425,829,519,543]" location="Sn Gerardo de Rivas" pageId="12" pageNumber="1776">
|
||
<quantity id="13589B6F776C1C0575E6FDB7599F6B47" box="[425,500,519,543]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="12" pageNumber="1776" unit="km" value="2.4">2.4 km</quantity>
|
||
ENE Sn Gerardo de Rivas
|
||
</locationDeviation>
|
||
, trail to
|
||
<location id="D17F6051776C1C0577E7FDB758A06B67" LSID="urn:lsid:plazi:treatment:5C09879C77691C067617FDC85BD56BC8:D17F6051776C1C0577E7FDB758A06B67" country="Costa Rica" latitude="9.46565" longLatPrecision="1" longitude="-83.567314" name="Chirripo Natl. Park" pageId="12" pageNumber="1776" stateProvince="San Jose">Chirripo Natl. Park</location>
|
||
,
|
||
<quantity id="13589B6F776C1C057499FD9759426B67" box="[214,297,551,575]" metricMagnitude="3" metricUnit="m" metricValue="2.07" pageId="12" pageNumber="1776" unit="m" value="2070.0">
|
||
<elevation id="5F8DD1B9776C1C057499FD9759426B67" box="[214,297,551,575]" metricMagnitude="3" metricUnit="m" metricValue="2.07" pageId="12" pageNumber="1776" unit="m" value="2070.0">2070 m</elevation>
|
||
</quantity>
|
||
,
|
||
<geoCoordinate id="B194504D776C1C05757CFD9759C36B67" box="[307,424,551,575]" degrees="9" direction="north" minutes="27.939" orientation="latitude" pageId="12" pageNumber="1776" precision="1" value="9.46565">
|
||
9
|
||
<emphasis id="E6D4EA98776C1C05750FFD9759216B67" bold="true" box="[320,330,551,575]" italics="true" pageId="12" pageNumber="1776">°</emphasis>
|
||
27.939N
|
||
</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="B194504D776C1C0575FCFD975A506B67" box="[435,571,551,575]" degrees="83" direction="west" minutes="34.039" orientation="longitude" pageId="12" pageNumber="1776" precision="1" value="-83.567314">
|
||
83
|
||
<emphasis id="E6D4EA98776C1C057581FD9759B36B67" bold="true" box="[462,472,551,575]" italics="true" pageId="12" pageNumber="1776">°</emphasis>
|
||
34.039W
|
||
</geoCoordinate>
|
||
, vi.10.2004,
|
||
<collectorName id="7955535C776C1C057689FD975B416B67" box="[710,810,551,575]" pageId="12" pageNumber="1776">J.S.Ashe</collectorName>
|
||
,
|
||
<collectorName id="7955535C776C1C05777AFD975BFB6B67" box="[821,912,551,575]" pageId="12" pageNumber="1776">Z. Falin</collectorName>
|
||
and I.
|
||
<collectorName id="7955535C776C1C057790FD975C216B67" box="[991,1098,551,575]" pageId="12" pageNumber="1776">Hinojosa</collectorName>
|
||
, ex: treefall litter, barcode labels:
|
||
<specimenCode id="84069EF1776C1C05764DFDF85AE06B38" box="[514,651,584,608]" collectionCode="SM" country="Malaysia" name="Sarawak Museum" pageId="12" pageNumber="1776">SM0607677</specimenCode>
|
||
,
|
||
<specimenCode id="84069EF1776C1C0576D9FDF85B4B6B38" box="[662,800,584,608]" collectionCode="SM" country="Malaysia" name="Sarawak Museum" pageId="12" pageNumber="1776">SM0607675</specimenCode>
|
||
,
|
||
<specimenCode id="84069EF1776C1C057765FDF85BDB6B38" box="[810,944,584,608]" collectionCode="SM" country="Malaysia" name="Sarawak Museum" pageId="12" pageNumber="1776">SM0607676</specimenCode>
|
||
(
|
||
<quantity id="13589B6F776C1C05778FFDF85B816B38" box="[960,1002,584,608]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="12" pageNumber="1776" unit="in" value="3.0">3 in</quantity>
|
||
<collectionCode id="B2B1AE4F776C1C0577BEFDF85C286B38" box="[1009,1091,584,608]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="12" pageNumber="1776" type="Museum">SEMC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="64C83CD7776C1C0574DDFDD85BEC6BF9" collectionCode="SEMC, UTCI" collectorName="P. Hanson" country="Costa Rica" elevation="1600" latitude="10.65" location="Zurqui de Moravia" longLatPrecision="1297" longitude="-84.316666" pageId="12" pageNumber="1776" specimenCode="SM0133116" specimenCount="1" stateProvince="San Jose" typeStatus="paratype">
|
||
<collectingRegion id="1664F868776C1C0574DDFDD858936BD8" box="[146,248,616,640]" country="Costa Rica" name="San Jose" pageId="12" pageNumber="1776">San Jose</collectingRegion>
|
||
,
|
||
<location id="D17F6051776C1C05754AFDD8598F6BD8" LSID="urn:lsid:plazi:treatment:5C09879C77691C067617FDC85BD56BC8:D17F6051776C1C05754AFDD8598F6BD8" box="[261,484,616,640]" country="Costa Rica" latitude="10.65" longLatPrecision="1297" longitude="-84.316666" name="Zurqui de Moravia" pageId="12" pageNumber="1776" stateProvince="San Jose">Zurqui de Moravia</location>
|
||
,
|
||
<quantity id="13589B6F776C1C0575BEFDD85A2F6BD8" box="[497,580,616,640]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="12" pageNumber="1776" unit="m" value="1600.0">
|
||
<elevation id="5F8DD1B9776C1C0575BEFDD85A2F6BD8" box="[497,580,616,640]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="12" pageNumber="1776" unit="m" value="1600.0">1600 m</elevation>
|
||
</quantity>
|
||
,
|
||
<geoCoordinate id="B194504D776C1C05761FFDD85ACF6BD8" box="[592,676,616,640]" degrees="10" direction="north" minutes="39" orientation="latitude" pageId="12" pageNumber="1776" precision="925" value="10.65">
|
||
10
|
||
<emphasis id="E6D4EA98776C1C057624FDD85A1E6BD8" bold="true" box="[619,629,616,640]" italics="true" pageId="12" pageNumber="1776">°</emphasis>
|
||
39N
|
||
</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="B194504D776C1C0576FEFDD85B626BD8" box="[689,777,616,640]" degrees="84" direction="west" minutes="19" orientation="longitude" pageId="12" pageNumber="1776" precision="925" value="-84.316666">
|
||
84
|
||
<emphasis id="E6D4EA98776C1C057683FDD85ABD6BD8" bold="true" box="[716,726,616,640]" italics="true" pageId="12" pageNumber="1776">°</emphasis>
|
||
19W
|
||
</geoCoordinate>
|
||
, i.1–30.1996,
|
||
<collectorName id="7955535C776C1C0577E9FDD85C4A6BD8" box="[934,1057,616,640]" pageId="12" pageNumber="1776">P. Hanson</collectorName>
|
||
, ex: malaise, barcode labels:
|
||
<specimenCode id="84069EF1776C1C0575FBFD395A516BF9" box="[436,570,649,673]" collectionCode="SM" country="Malaysia" name="Sarawak Museum" pageId="12" pageNumber="1776">SM0133116</specimenCode>
|
||
(
|
||
<quantity id="13589B6F776C1C05761FFD395A156BF9" box="[592,638,649,673]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="12" pageNumber="1776" unit="in" value="1.0">1 in</quantity>
|
||
<collectionCode id="B2B1AE4F776C1C0576C6FD395AB06BF9" box="[649,731,649,673]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="12" pageNumber="1776" type="Museum">SEMC</collectionCode>
|
||
), (
|
||
<quantity id="13589B6F776C1C0576B1FD395B406BF9" box="[766,811,649,673]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="12" pageNumber="1776" unit="in" value="1.0">1 in</quantity>
|
||
<collectionCode id="B2B1AE4F776C1C057778FD395B146BF9" box="[823,895,649,673]" pageId="12" pageNumber="1776">UTCI</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="64C83CD7776C1C0577D8FD395B666A5A" collectionCode="SEMC" collectorName="J. S. Ashe & Z. Falin & Hinojosa" country="Costa Rica" elevation="1860" latitude="9.47815" location="Cloudbridge Reserve" longLatPrecision="1" longitude="-83.56682" pageId="12" pageNumber="1776" specimenCode="SM0607686" specimenCount="1" stateProvince="San Jose" typeStatus="paratype">
|
||
<collectingRegion id="1664F868776C1C0577D8FD395C216BF9" box="[919,1098,649,673]" country="Costa Rica" name="San Jose" pageId="12" pageNumber="1776">San Jose Prov.</collectingRegion>
|
||
,
|
||
<locationDeviation id="C5AD2FA8776C1C0574DDFD195A7A6B99" box="[146,529,681,705]" location="Sn Gerardo de Rivas" pageId="12" pageNumber="1776">
|
||
<quantity id="13589B6F776C1C0574DDFD1958B66B99" box="[146,221,681,705]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="12" pageNumber="1776" unit="km" value="2.4">2.4 km</quantity>
|
||
ENE Sn Gerardo de Rivas
|
||
</locationDeviation>
|
||
,
|
||
<location id="D17F6051776C1C057652FD195B666B99" LSID="urn:lsid:plazi:treatment:5C09879C77691C067617FDC85BD56BC8:D17F6051776C1C057652FD195B666B99" box="[541,781,681,705]" country="Costa Rica" latitude="9.47815" longLatPrecision="1" longitude="-83.56682" name="Cloudbridge Reserve" pageId="12" pageNumber="1776" stateProvince="San Jose">Cloudbridge Reserve</location>
|
||
, ridge above covered bridge,
|
||
<quantity id="13589B6F776C1C0574DDFD79588E6BB9" box="[146,229,713,737]" metricMagnitude="3" metricUnit="m" metricValue="1.86" pageId="12" pageNumber="1776" unit="m" value="1860.0">
|
||
<elevation id="5F8DD1B9776C1C0574DDFD79588E6BB9" box="[146,229,713,737]" metricMagnitude="3" metricUnit="m" metricValue="1.86" pageId="12" pageNumber="1776" unit="m" value="1860.0">1860 m</elevation>
|
||
</quantity>
|
||
,
|
||
<geoCoordinate id="B194504D776C1C0574BDFD79590C6BB9" box="[242,359,713,738]" degrees="9" direction="north" minutes="28.689" orientation="latitude" pageId="12" pageNumber="1776" precision="1" value="9.47815">
|
||
9
|
||
<emphasis id="E6D4EA98776C1C0574B0FD7A59626BBA" bold="true" box="[255,265,714,738]" italics="true" pageId="12" pageNumber="1776">°</emphasis>
|
||
28.689N
|
||
</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="B194504D776C1C05753AFD7959966BB9" box="[373,509,713,738]" degrees="83" direction="west" minutes="34.009" orientation="longitude" pageId="12" pageNumber="1776" precision="1" value="-83.56682">
|
||
83
|
||
<emphasis id="E6D4EA98776C1C0575DFFD7A59F16BBA" bold="true" box="[400,410,714,738]" italics="true" pageId="12" pageNumber="1776">°</emphasis>
|
||
34.009W
|
||
</geoCoordinate>
|
||
, vi.11.2004,
|
||
<collectorName id="7955535C776C1C0576C1FD795A996BB9" box="[654,754,713,737]" pageId="12" pageNumber="1776">J.S.Ashe</collectorName>
|
||
,
|
||
<collectorName id="7955535C776C1C05774FFD795B356BB9" box="[768,862,713,737]" pageId="12" pageNumber="1776">Z. Falin</collectorName>
|
||
and I.
|
||
<collectorName id="7955535C776C1C0577F9FD795C4A6BB9" box="[950,1057,713,737]" pageId="12" pageNumber="1776">Hinojosa</collectorName>
|
||
, ex: treefall litter, barcode label:
|
||
<specimenCode id="84069EF1776C1C0575AEFD5A5A0C6A5A" box="[481,615,746,770]" collectionCode="SM" country="Malaysia" name="Sarawak Museum" pageId="12" pageNumber="1776">SM0607686</specimenCode>
|
||
(
|
||
<quantity id="13589B6F776C1C057632FD5A5AC26A5A" box="[637,681,746,770]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="12" pageNumber="1776" unit="in" value="1.0">1 in</quantity>
|
||
<collectionCode id="B2B1AE4F776C1C0576FCFD5A5B6E6A5A" box="[691,773,746,770]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="12" pageNumber="1776" type="Museum">SEMC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="64C83CD7776C1C057753FD5A5C206A7A" collectionCode="R" collectorName="J. S. Ashe" country="Costa Rica" elevation="1490" location="San Juan" pageId="12" pageNumber="1776" specimenCount="1" stateProvince="San Jose" typeStatus="paratype">
|
||
<collectingRegion id="1664F868776C1C057753FD5A5BA66A5A" box="[796,973,746,770]" country="Costa Rica" name="San Jose" pageId="12" pageNumber="1776">San Jose Prov.</collectingRegion>
|
||
,
|
||
<location id="D17F6051776C1C057793FD5A5C216A5A" LSID="urn:lsid:plazi:treatment:5C09879C77691C067617FDC85BD56BC8:D17F6051776C1C057793FD5A5C216A5A" box="[988,1098,746,770]" country="Costa Rica" name="San Juan" pageId="12" pageNumber="1776" stateProvince="San Jose">San Juan</location>
|
||
,
|
||
<locationDeviation id="C5AD2FA8776C1C0574DDFCBA59C76A7A" box="[146,428,778,802]" location="Finca Zurqui" pageId="12" pageNumber="1776">
|
||
<quantity id="13589B6F776C1C0574DDFCBA58BD6A7A" box="[146,214,778,802]" metricMagnitude="4" metricUnit="m" metricValue="1.4" pageId="12" pageNumber="1776" unit="km" value="14.0">14 km</quantity>
|
||
NE Finca Zurqui
|
||
</locationDeviation>
|
||
, 10
|
||
<emphasis id="E6D4EA98776C1C05759AFCBA59B46A7A" bold="true" box="[469,479,778,802]" italics="true" pageId="12" pageNumber="1776">°</emphasis>
|
||
29570N, 84
|
||
<emphasis id="E6D4EA98776C1C057615FCBA5A0F6A7A" bold="true" box="[602,612,778,802]" italics="true" pageId="12" pageNumber="1776">°</emphasis>
|
||
09220W,
|
||
<quantity id="13589B6F776C1C057687FCBA5B706A7A" box="[712,795,778,802]" metricMagnitude="3" metricUnit="m" metricValue="1.49" pageId="12" pageNumber="1776" unit="m" value="1490.0">
|
||
<elevation id="5F8DD1B9776C1C057687FCBA5B706A7A" box="[712,795,778,802]" metricMagnitude="3" metricUnit="m" metricValue="1.49" pageId="12" pageNumber="1776" unit="m" value="1490.0">1490 m</elevation>
|
||
</quantity>
|
||
, vii.4–6.2000,
|
||
<collectorName id="7955535C776C1C05778FFCBA5C4E6A7A" box="[960,1061,778,802]" pageId="12" pageNumber="1776">J.S.Ashe</collectorName>
|
||
,
|
||
<collectionCode id="B2B1AE4F776C1C05707CFCBA5C206A7A" box="[1075,1099,778,802]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="1776">R</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="64C83CD7776C1C0574DDFC9B58876A3B" collectionCode="SEMC" collectorName="Z. Falin" country="Costa Rica" location="Brooks" pageId="12" pageNumber="1776" specimenCode="SM0203734" specimenCount="1" stateProvince="San Jose" typeStatus="paratype">
|
||
<location id="D17F6051776C1C0574DDFC9B588F6A1B" LSID="urn:lsid:plazi:treatment:5C09879C77691C067617FDC85BD56BC8:D17F6051776C1C0574DDFC9B588F6A1B" box="[146,228,811,835]" country="Costa Rica" name="Brooks" pageId="12" pageNumber="1776" stateProvince="San Jose">Brooks</location>
|
||
and
|
||
<collectorName id="7955535C776C1C057564FC9B59FA6A1B" box="[299,401,811,835]" pageId="12" pageNumber="1776">Z. Falin</collectorName>
|
||
, ex:
|
||
<collectingMethod id="0DE14E9D776C1C05759DFC9B5AAB6A1B" box="[466,704,811,835]" pageId="12" pageNumber="1776">flight intercept trap</collectingMethod>
|
||
, barcode label
|
||
<specimenCode id="84069EF1776C1C0577CFFC9B5C6D6A1B" box="[896,1030,811,835]" collectionCode="SM" country="Malaysia" name="Sarawak Museum" pageId="12" pageNumber="1776">SM0203734</specimenCode>
|
||
(
|
||
<quantity id="13589B6F776C1C057050FC9B5C246A1B" box="[1055,1103,811,835]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="12" pageNumber="1776" unit="in" value="1.0">1 in</quantity>
|
||
<collectionCode id="B2B1AE4F776C1C0574DDFCFB588F6A3B" box="[146,228,843,867]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="12" pageNumber="1776" type="Museum">SEMC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="64C83CD7776C1C05754CFCFB5C206A3B" box="[259,1099,843,867]" collectionCode="R" collectorName="P. N. Tapanti." country="Costa Rica" elevation="1740" location="Cartago" pageId="12" pageNumber="1776" specimenCount="1" stateProvince="Cartago" typeStatus="paratype">
|
||
<collectingRegion id="1664F868776C1C05754CFCFB59086A3B" box="[259,355,843,867]" country="Costa Rica" name="Cartago" pageId="12" pageNumber="1776">Cartago</collectingRegion>
|
||
,
|
||
<collectorName id="7955535C776C1C057536FCFB5A646A3B" box="[377,527,843,867]" pageId="12" pageNumber="1776">P.N.Tapanti.</collectorName>
|
||
<quantity id="13589B6F776C1C05766EFCFB5A046A3B" box="[545,623,843,867]" metricMagnitude="3" metricUnit="m" metricValue="1.7399999999999998" pageId="12" pageNumber="1776" unit="m" value="1740.0">
|
||
<elevation id="5F8DD1B9776C1C05766EFCFB5A046A3B" box="[545,623,843,867]" metricMagnitude="3" metricUnit="m" metricValue="1.7399999999999998" pageId="12" pageNumber="1776" unit="m" value="1740.0">1740m</elevation>
|
||
</quantity>
|
||
, 9
|
||
<emphasis id="E6D4EA98776C1C0576DCFCFB5AF66A3B" bold="true" box="[659,669,843,867]" italics="true" pageId="12" pageNumber="1776">°</emphasis>
|
||
449300N, 83
|
||
<emphasis id="E6D4EA98776C1C057761FCFB5B536A3B" bold="true" box="[814,824,843,867]" italics="true" pageId="12" pageNumber="1776">°</emphasis>
|
||
489300W, vi.4.1997,
|
||
<collectionCode id="B2B1AE4F776C1C05707CFCFB5C206A3B" box="[1075,1099,843,867]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="1776">R</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="64C83CD7776C1C0574DDFCDC5C216ADC" box="[146,1098,876,900]" collectionCode="SEMC" country="Costa Rica" location="Anderson" pageId="12" pageNumber="1776" specimenCode="SM0120360" specimenCount="1" stateProvince="Cartago" typeStatus="paratype">
|
||
<location id="D17F6051776C1C0574DDFCDC596F6ADC" LSID="urn:lsid:plazi:treatment:5C09879C77691C067617FDC85BD56BC8:D17F6051776C1C0574DDFCDC596F6ADC" box="[146,260,876,900]" country="Costa Rica" name="Anderson" pageId="12" pageNumber="1776" stateProvince="Cartago">Anderson</location>
|
||
, ex: berlese forest litter, barcode label:
|
||
<specimenCode id="84069EF1776C1C057746FCDC5BE46ADC" box="[777,911,876,900]" collectionCode="SM" country="Malaysia" name="Sarawak Museum" pageId="12" pageNumber="1776">SM0120360</specimenCode>
|
||
(
|
||
<quantity id="13589B6F776C1C0577E4FCDC5BB46ADC" box="[939,991,876,900]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="12" pageNumber="1776" unit="in" value="1.0">1 in</quantity>
|
||
<collectionCode id="B2B1AE4F776C1C0577BEFCDC5C286ADC" box="[1009,1091,876,900]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="12" pageNumber="1776" type="Museum">SEMC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="64C83CD7776C1C0574DDFC3C58B46A9C" collectionCode="R" collectorName="P. N. Tapanti. & J. S. Ashe" country="Costa Rica" elevation="1150" location="Cartago" pageId="12" pageNumber="1776" specimenCount="1" stateProvince="Cartago" typeStatus="paratype">
|
||
<collectingRegion id="1664F868776C1C0574DDFC3C58996AFC" box="[146,242,908,932]" country="Costa Rica" name="Cartago" pageId="12" pageNumber="1776">Cartago</collectingRegion>
|
||
,
|
||
<collectorName id="7955535C776C1C057549FC3C59F06AFC" box="[262,411,908,932]" pageId="12" pageNumber="1776">P.N.Tapanti.</collectorName>
|
||
<quantity id="13589B6F776C1C0575E4FC3C59966AFC" box="[427,509,908,932]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="12" pageNumber="1776" unit="m" value="1150.0">
|
||
<elevation id="5F8DD1B9776C1C0575E4FC3C59966AFC" box="[427,509,908,932]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="12" pageNumber="1776" unit="m" value="1150.0">1150 m</elevation>
|
||
</quantity>
|
||
, 9
|
||
<emphasis id="E6D4EA98776C1C057651FC3C5A436AFC" bold="true" box="[542,552,908,932]" italics="true" pageId="12" pageNumber="1776">°</emphasis>
|
||
459410N, 83
|
||
<emphasis id="E6D4EA98776C1C0576F8FC3C5AAA6AFC" bold="true" box="[695,705,908,932]" italics="true" pageId="12" pageNumber="1776">°</emphasis>
|
||
47950W, vi.17–20.2000,
|
||
<collectorName id="7955535C776C1C057792FC3C5C256AFC" box="[989,1102,908,932]" pageId="12" pageNumber="1776">J.S. Ashe</collectorName>
|
||
and
|
||
<collectionCode id="B2B1AE4F776C1C057488FC1C58B46A9C" box="[199,223,940,964]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="1776">R</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="64C83CD7776C1C0574A2FC1C5C216A9C" box="[237,1098,940,964]" collectionCode="SEMC" country="Costa Rica" location="Brooks" pageId="12" pageNumber="1776" specimenCode="SM0203331" specimenCount="1" stateProvince="Cartago" typeStatus="paratype">
|
||
<location id="D17F6051776C1C0574A2FC1C59296A9C" LSID="urn:lsid:plazi:treatment:5C09879C77691C067617FDC85BD56BC8:D17F6051776C1C0574A2FC1C59296A9C" box="[237,322,940,964]" country="Costa Rica" name="Brooks" pageId="12" pageNumber="1776" stateProvince="Cartago">Brooks</location>
|
||
, ex:
|
||
<collectingMethod id="0DE14E9D776C1C057533FC1C5A086A9C" box="[380,611,940,964]" pageId="12" pageNumber="1776">flight intercept trap</collectingMethod>
|
||
, barcode label:
|
||
<specimenCode id="84069EF1776C1C057750FC1C5BCD6A9C" box="[799,934,940,964]" collectionCode="SM" country="Malaysia" name="Sarawak Museum" pageId="12" pageNumber="1776">SM0203331</specimenCode>
|
||
(
|
||
<quantity id="13589B6F776C1C0577F5FC1C5B8D6A9C" box="[954,998,940,964]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="12" pageNumber="1776" unit="in" value="1.0">1 in</quantity>
|
||
<collectionCode id="B2B1AE4F776C1C0577BEFC1C5C286A9C" box="[1009,1091,940,964]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="12" pageNumber="1776" type="Museum">SEMC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="64C83CD7776C1C0574DDFC7D5AEB6ABD" box="[146,640,973,997]" collectionCode="FIT" country="Costa Rica" elevation="1600" location="Zurqui" pageId="12" pageNumber="1776" specimenCount="1" stateProvince="Heredia" typeStatus="paratype">
|
||
<collectingRegion id="1664F868776C1C0574DDFC7D58846ABD" box="[146,239,973,997]" country="Costa Rica" name="Heredia" pageId="12" pageNumber="1776">Heredia</collectingRegion>
|
||
,
|
||
<location id="D17F6051776C1C0574B1FC7D593B6ABD" LSID="urn:lsid:plazi:treatment:5C09879C77691C067617FDC85BD56BC8:D17F6051776C1C0574B1FC7D593B6ABD" box="[254,336,973,997]" country="Costa Rica" name="Zurqui" pageId="12" pageNumber="1776" stateProvince="Heredia">Zurqui</location>
|
||
,
|
||
<quantity id="13589B6F776C1C057511FC7D59DA6ABD" box="[350,433,973,997]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="12" pageNumber="1776" unit="m" value="1600.0">
|
||
<elevation id="5F8DD1B9776C1C057511FC7D59DA6ABD" box="[350,433,973,997]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="12" pageNumber="1776" unit="m" value="1600.0">1600 m</elevation>
|
||
</quantity>
|
||
, ix.1–9.1998,
|
||
<collectionCode id="B2B1AE4F776C1C05761FFC7D5AEB6ABD" box="[592,640,973,997]" pageId="12" pageNumber="1776">FIT</collectionCode>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="64C83CD7776C1C0576C1FC7D599C6D5D" collectionCode="FMNH" collectorName="C. W. and L. B. O'Brien & All" country="Costa Rica" pageId="12" pageNumber="1776" specimenCount="1" stateProvince="Heredia" typeStatus="paratype">
|
||
<collectorName id="7955535C776C1C0576C1FC7D5BF66ABD" box="[654,925,973,997]" pageId="12" pageNumber="1776">C.W. and L.B. O’Brien</collectorName>
|
||
(
|
||
<quantity id="13589B6F776C1C0577FDFC7D5BB66ABD" box="[946,989,973,997]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="12" pageNumber="1776" unit="in" value="1.0">1 in</quantity>
|
||
<collectionCode id="B2B1AE4F776C1C0577A8FC7D5C286ABD" box="[999,1091,973,997]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="12" pageNumber="1776" type="Museum">FMNH</collectionCode>
|
||
).
|
||
<collectorName id="7955535C776C1C0574DDFC5D58DE6D5D" box="[146,181,1005,1029]" pageId="12" pageNumber="1776">All</collectorName>
|
||
<typeStatus id="0B1B8828776C1C057482FC5D59506D5D" box="[205,315,1005,1029]" pageId="12" pageNumber="1776" type="paratype">paratypes</typeStatus>
|
||
with label ‘‘
|
||
</materialsCitation>
|
||
<typeStatus id="0B1B8828776C1C0575B8FC5D5A0B6D5D" box="[503,608,1005,1029]" pageId="12" pageNumber="1776" type="paratype">Paratype</typeStatus>
|
||
<taxonomicName id="13A04D09776C1C057636FC5D5B8C6D5D" authority="Chatzimanolis" authorityName="Chatzimanolis" authorityYear="2008" box="[633,999,1005,1029]" class="Insecta" family="Staphylinidae" genus="Isanopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="1776" phylum="Arthropoda" rank="species" species="ashei">
|
||
<emphasis id="E6D4EA98776C1C057636FC5D5B406D5D" box="[633,811,1005,1029]" italics="true" pageId="12" pageNumber="1776">Isanopus ashei</emphasis>
|
||
Chatzimanolis
|
||
</taxonomicName>
|
||
design. Chatzimanolis 2007’’.
|
||
</paragraph>
|
||
<paragraph id="D41F368A776C1C0574DDFBD95B9C6DF9" blockId="12.[146,1103,1129,1185]" pageId="12" pageNumber="1776">
|
||
<emphasis id="E6D4EA98776C1C0574DDFBD959606DD9" box="[146,267,1129,1153]" italics="true" pageId="12" pageNumber="1776">Etymology</emphasis>
|
||
. This species is named in honor of the late J. Steve Ashe, friend and mentor, who inspired my
|
||
<taxonomicName id="13A04D09776C1C0575F2FB395A0B6DF9" authorityName="Sharp" authorityYear="1884" box="[445,608,1161,1185]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="1776" phylum="Arthropoda" rank="subTribe" subTribe="Xanthopygina">Xanthopygina</taxonomicName>
|
||
studies and collected the
|
||
<typeStatus id="0B1B8828776C1C0577C3FB395B9F6DF9" box="[908,1012,1161,1185]" pageId="12" pageNumber="1776" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="D41F368A776C1C0574DDFB6A597E6C4A" blockId="12.[146,1103,1242,1298]" pageId="12" pageNumber="1776">
|
||
<emphasis id="E6D4EA98776C1C0574DDFB6A59736DAA" box="[146,280,1242,1266]" italics="true" pageId="12" pageNumber="1776">Distribution</emphasis>
|
||
. Known from elevations of
|
||
<quantity id="13589B6F776C1C057621FB6A5B6A6DAA" box="[622,769,1242,1266]" metricMagnitude="3" metricUnit="m" metricValue="1.61" metricValueMax="2.07" metricValueMin="1.15" pageId="12" pageNumber="1776" unit="m" value="1610.0" valueMax="2070.0" valueMin="1150.0">1150–2070 m</quantity>
|
||
in
|
||
<collectingCountry id="ACB7761A776C1C057762FB6A5BDB6DAA" box="[813,944,1242,1266]" name="Costa Rica" pageId="12" pageNumber="1776">Costa Rica</collectingCountry>
|
||
and
|
||
<collectingCountry id="ACB7761A776C1C0577BDFB6A5C246DAA" box="[1010,1103,1242,1266]" name="Panama" pageId="12" pageNumber="1776">Panama</collectingCountry>
|
||
(
|
||
<figureCitation id="4C9B2A0F776C1C0574D5FB4A59636C4A" box="[154,264,1274,1298]" captionStart="Figure 10" captionStartId="23.[213,280,856,878]" captionTargetBox="[213,1171,165,820]" captionTargetId="figure-204@23.[213,1171,165,820]" captionTargetPageId="23" captionText="Figure 10. Distribution map of I. ashei, I. hinojosai and I. sallaei." figureDoi="http://doi.org/10.5281/zenodo.4593955" httpUri="https://zenodo.org/record/4593955/files/figure.png" pageId="12" pageNumber="1776">Figure 10</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="D41F368A776C1C0574DDFAFC5B846C3B" blockId="12.[146,1007,1355,1380]" box="[146,1007,1355,1380]" pageId="12" pageNumber="1776">
|
||
<emphasis id="E6D4EA98776C1C0574DDFAFC58826C3C" box="[146,233,1356,1380]" italics="true" pageId="12" pageNumber="1776">Habitat</emphasis>
|
||
. Collected in treefall litter, malaise traps and flight intercept traps.
|
||
</paragraph>
|
||
<paragraph id="D41F368A776C1C0575B4FA245B596CF4" blockId="12.[507,818,1428,1485]" box="[507,818,1428,1452]" pageId="12" pageNumber="1776">
|
||
<taxonomicName id="13A04D09776C1C0575B4FA245A816CF4" authority="Chatzimanolis, 2008" authorityName="Chatzimanolis" authorityYear="2008" box="[507,746,1428,1452]" class="Insecta" family="Staphylinidae" genus="Isanopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="1776" phylum="Arthropoda" rank="species" species="eptaskouros" status="sp. nov.">
|
||
<emphasis id="E6D4EA98776C1C0575B4FA245A816CF4" bold="true" box="[507,746,1428,1452]" italics="true" pageId="12" pageNumber="1776">Isanopus eptaskouros</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="FDE757E3776C1C0576BCFA245B596CF4" box="[755,818,1428,1452]" pageId="12" pageNumber="1776" rank="species">sp. n.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="D41F368A776C1C057649FA055B4C6C95" blockId="12.[507,818,1428,1485]" box="[518,807,1461,1485]" pageId="12" pageNumber="1776">
|
||
(
|
||
<figureCitation id="4C9B2A0F776C1C057641FA055AE46C95" box="[526,655,1461,1485]" captionStart="Figures 2" captionStartId="6.[146,222,1349,1371]" captionTargetBox="[149,1102,175,1302]" captionTargetId="figure-63@6.[146,1104,165,1312]" captionTargetPageId="6" captionText="Figures 2. Isanopus species, dorsum of head and pronotum. (A) I. ashei. (B) I. sahlbergi. (C) I. hinojosai. (D) I. eptaskouros." figureDoi="http://doi.org/10.5281/zenodo.4593941" httpUri="https://zenodo.org/record/4593941/files/figure.png" pageId="12" pageNumber="1776">Figures 2D</figureCitation>
|
||
, 4C, 6A, 6C)
|
||
</paragraph>
|
||
<paragraph id="D41F368A776C1C0574DDF9B658946F46" blockId="12.[146,255,1542,1566]" box="[146,255,1542,1566]" pageId="12" pageNumber="1776">
|
||
<emphasis id="E6D4EA98776C1C0574DDF9B658946F46" box="[146,255,1542,1566]" italics="true" pageId="12" pageNumber="1776">Diagnosis</emphasis>
|
||
</paragraph>
|
||
<paragraph id="D41F368A776C1C0574DDF99F5B9F6FF0" blockId="12.[146,1103,1583,1704]" pageId="12" pageNumber="1776">
|
||
<taxonomicName id="13A04D09776C1C0574DDF99F59EB6F1F" authorityName="Chatzimanolis" authorityYear="2008" box="[146,384,1583,1607]" class="Insecta" family="Staphylinidae" genus="Isanopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="1776" phylum="Arthropoda" rank="species" species="eptaskouros">
|
||
<emphasis id="E6D4EA98776C1C0574DDF99F59EB6F1F" box="[146,384,1583,1607]" italics="true" pageId="12" pageNumber="1776">Isanopus eptaskouros</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from all other species in
|
||
<taxonomicName id="13A04D09776C1C0577D3F99F5B966F1F" authorityName="Sharp" authorityYear="1876" box="[924,1021,1583,1607]" class="Insecta" family="Staphylinidae" genus="Isanopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="1776" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E6D4EA98776C1C0577D3F99F5B966F1F" box="[924,1021,1583,1607]" italics="true" pageId="12" pageNumber="1776">Isanopus</emphasis>
|
||
</taxonomicName>
|
||
by the colouration of abdominal segment VII. In
|
||
<taxonomicName id="13A04D09776C1C0576EEF9FF5B2F6F3F" authorityName="Chatzimanolis" authorityYear="2008" box="[673,836,1615,1639]" class="Insecta" family="Staphylinidae" genus="Isanopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="1776" phylum="Arthropoda" rank="species" species="eptaskouros">
|
||
<emphasis id="E6D4EA98776C1C0576EEF9FF5B2F6F3F" box="[673,836,1615,1639]" italics="true" pageId="12" pageNumber="1776">I. eptaskouros</emphasis>
|
||
</taxonomicName>
|
||
it is completely dark brown-black, while in other species it is either yellowish-orange or a combination of dark brown-black and orange. Aedeagus (
|
||
<figureCitation id="4C9B2A0F776C1C057636F9205A8A6FF0" box="[633,737,1680,1704]" captionStart="Figure 6" captionStartId="13.[213,280,1654,1676]" captionTargetBox="[213,1164,1116,1611]" captionTargetId="figure-238@13.[213,1171,1116,1617]" captionTargetPageId="13" captionText="Figure 6. Aedeagus of I. eptaskouros. (A) Dorsal view. (B) Lateral view. (C) Detail of paramere, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4593947" httpUri="https://zenodo.org/record/4593947/files/figure.png" pageId="12" pageNumber="1776">Figures 6</figureCitation>
|
||
A–C) is also distinctive.
|
||
</paragraph>
|
||
<paragraph id="D41F368A776D1C04749AFF19593F6999" blockId="13.[213,340,169,193]" box="[213,340,169,193]" pageId="13" pageNumber="1777">
|
||
<emphasis id="E6D4EA98776D1C04749AFF19593F6999" box="[213,340,169,193]" italics="true" pageId="13" pageNumber="1777">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="D41F368A776D1C04749AFF6159D669B1" blockId="13.[213,445,209,233]" box="[213,445,209,233]" pageId="13" pageNumber="1777">
|
||
Length
|
||
<quantity id="13589B6F776D1C047561FF6159D269B1" box="[302,441,209,233]" metricMagnitude="-2" metricUnit="m" metricValue="1.175" metricValueMax="1.25" metricValueMin="1.1" pageId="13" pageNumber="1777" unit="mm" value="11.75" valueMax="12.5" valueMin="11.0">11–12.5 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="D41F368A776D1C04749AFE92596668E4" blockId="13.[213,1170,290,444]" pageId="13" pageNumber="1777">
|
||
<emphasis id="E6D4EA98776D1C04749AFE9259326862" box="[213,345,290,314]" italics="true" pageId="13" pageNumber="1777">Colouration</emphasis>
|
||
. Colour of head and pronotum dark brown-black. Antenna brown. Scutellum dark brown-black. Elytra bright metallic blue, purplish blue, sometimes with green overtones. Abdomen dark brown, except segments VIII–IX yellow. Legs dark brown. Head, pronotum and abdomen integument appearing iridescent under direct light.
|
||
</paragraph>
|
||
<paragraph id="D41F368A776D1C04749AFE455B436D4B" blockId="13.[213,1169,500,1043]" pageId="13" pageNumber="1777">
|
||
<emphasis id="E6D4EA98776D1C04749AFE45597A6B55" box="[213,273,501,525]" italics="true" pageId="13" pageNumber="1777">Head</emphasis>
|
||
. Head with many relatively small brown microsetae and longer black macrosetae along borders of head; head subquadrate, width: length ratio50.91. Surface of epicranium glossy, with few micropunctures distributed at centre. Epicranium slightly convex, with small-sized umbilicate setose punctures covering head except medially, punctures only slighter smaller around eyes and at base of head, distance between punctures various, from 0.5–1 times width of punctures at centre, but punctures at antero-lateral and posterior part of head almost confluent. Eyes medium, length of eyes/length of head50.60, distance between eyes as wide as 1.6 times length of eye. Postgena with medium to large, not longitudinally elongate punctures (about eight punctures/
|
||
<quantity id="13589B6F776D1C0476CAFCA85AB06A68" box="[645,731,792,816]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="1777" unit="mm" value="0.5">0.5 mm</quantity>
|
||
), punctures uniformly distributed. Antennomeres with long macrosetae, antennomere 1 gradually club-like thickened, antennomeres 1–11 longer than wide, antennomere 1 about twice as long as antennomere 2, antennomere 3 about 1.5 times as long as antennomere 2, antennomere 4 slightly shorter than antennomere 3, antennomeres 5–7 almost identical, twice as long as wide, antennomeres 8–10 almost identical, shorter than 5– 7, antennomere 11 longer than previous antennomeres. Neck with multiple small irregular punctures and micropunctures posteriorly.
|
||
</paragraph>
|
||
<caption id="80DF6602776D1C04749AF9C659A36FF1" ID-DOI="http://doi.org/10.5281/zenodo.4593947" ID-Zenodo-Dep="4593947" httpUri="https://zenodo.org/record/4593947/files/figure.png" pageId="13" pageNumber="1777" startId="13.[213,280,1654,1676]" targetBox="[213,1164,1116,1611]" targetPageId="13">
|
||
<paragraph id="D41F368A776D1C04749AF9C659A36FF1" blockId="13.[213,1169,1654,1705]" pageId="13" pageNumber="1777">
|
||
Figure 6. Aedeagus of
|
||
<taxonomicName id="13A04D09776D1C04759AF9C65A0C6FD3" authorityName="Chatzimanolis" authorityYear="2008" box="[469,615,1654,1675]" class="Insecta" family="Staphylinidae" genus="Isanopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1777" phylum="Arthropoda" rank="species" species="eptaskouros">
|
||
<emphasis id="E6D4EA98776D1C04759AF9C65A0C6FD3" box="[469,615,1654,1675]" italics="true" pageId="13" pageNumber="1777">I. eptaskouros</emphasis>
|
||
</taxonomicName>
|
||
. (A) Dorsal view. (B) Lateral view. (C) Detail of paramere, ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D41F368A776E1C0774DDFF1859966B46" blockId="14.[146,1103,167,638]" pageId="14" pageNumber="1778">
|
||
<emphasis id="E6D4EA98776E1C0774DDFF18588F6998" box="[146,228,168,192]" italics="true" pageId="14" pageNumber="1778">Thorax</emphasis>
|
||
. Pronotum longer than wide, width: length ratio50.90; antero-lateral angles obtusely rounded, slightly curved. Lateral margins of pronotum concave in dorsal aspect, pronotum broadest in apical 1/3 and narrower at basal angles. Surface of pronotum flat, glossy, with micropunctures. Setose punctures absent on disc of pronotum, laterally present in 4–5 rows, distance between rows smaller towards lateral ends; punctures of medium size (about four punctures/
|
||
<quantity id="13589B6F776E1C077733FEF75BB96807" box="[892,978,327,351]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="14" pageNumber="1778" unit="mm" value="0.5">0.5 mm</quantity>
|
||
); the least distance between punctures for each row various, 1–2 times width of puncture for most; antero-lateral angles densely punctuated, distance between punctures 0.5–1 times width of puncture. Pronotum with brown microsetae directed postero-medially in lateral and anterior areas and posteriorly in basal half; and few longer black macrosetae along borders. Scutellum prominent, with brown microsetae and dense punctures, punctures confluent.
|
||
</paragraph>
|
||
<paragraph id="D41F368A776E1C0774F2FD965AA96B26" blockId="14.[146,1103,167,638]" pageId="14" pageNumber="1778">
|
||
Elytra subequal in length to pronotum, with medium setose uniform punctures (about seven punctures/
|
||
<quantity id="13589B6F776E1C0775FFFDF65A6D6B06" box="[432,518,582,606]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="14" pageNumber="1778" unit="mm" value="0.5">0.5 mm</quantity>
|
||
), covered with yellowish brown microsetae and longer black macrosetae along borders of elytra.
|
||
</paragraph>
|
||
<paragraph id="D41F368A776E1C0774DDFD01598F6A50" blockId="14.[146,1103,688,776]" pageId="14" pageNumber="1778">
|
||
<emphasis id="E6D4EA98776E1C0774DDFD0158AC6B91" box="[146,199,689,713]" italics="true" pageId="14" pageNumber="1778">Legs</emphasis>
|
||
. Protarsus enlarged; meso and metatarsi slightly elongate and enlarged, tarsomeres 1–4 asymmetrical, dilated distally. Legs almost completely covered with long yellow and brown setae.
|
||
</paragraph>
|
||
<paragraph id="D41F368A776E1C0774DDFC8B5AF66D0A" blockId="14.[146,1103,827,1106]" pageId="14" pageNumber="1778">
|
||
<emphasis id="E6D4EA98776E1C0774DDFC8B58916A0B" box="[146,250,827,851]" italics="true" pageId="14" pageNumber="1778">Abdomen</emphasis>
|
||
. Abdominal terga III–V with tergal basal and no subbasal (arch-like) carina. Terga III–VII with uniform punctuation patterns, densely covered with punctures and yellowish brown setae, punctures not decreasing in size gradually from III–VII. Abdominal sterna III–IV with moderate transverse impressions, V with faint transverse impression, shining. Integument surface appearing strongly iridescent in strong direct light at moderate to high magnification. Sterna III–VII densely punctuated, punctures not decreasing in size gradually from III–VII, with yellowish brown setae. Lateral tergal sclerites of the abdominal segment IX long and straight, covered with long black macrosetae.
|
||
</paragraph>
|
||
<paragraph id="D41F368A776E1C0774DDFB3459626DA4" blockId="14.[146,1103,1156,1276]" pageId="14" pageNumber="1778">
|
||
<emphasis id="E6D4EA98776E1C0774DDFB3459B66DC4" box="[146,477,1156,1180]" italics="true" pageId="14" pageNumber="1778">Secondary sexual structures</emphasis>
|
||
. Male: distal margin of sternum VII with slight emargination medially; sternum VIII with deep V-shaped emargination medially; sternum IX with small emargination medially. Females: with no obvious sexual structures.
|
||
</paragraph>
|
||
<paragraph id="D41F368A776E1C0674DDFA845BA86818" blockId="14.[146,1103,1331,1706]" lastBlockId="15.[213,1169,168,320]" lastPageId="15" lastPageNumber="1779" pageId="14" pageNumber="1778">
|
||
<emphasis id="E6D4EA98776E1C0774DDFA8458976C14" box="[146,252,1332,1356]" italics="true" pageId="14" pageNumber="1778">Aedeagus</emphasis>
|
||
. As in
|
||
<figureCitation id="4C9B2A0F776E1C077517FA8359D56C13" box="[344,446,1331,1355]" captionStart="Figure 6" captionStartId="13.[213,280,1654,1676]" captionTargetBox="[213,1164,1116,1611]" captionTargetId="figure-238@13.[213,1171,1116,1617]" captionTargetPageId="13" captionText="Figure 6. Aedeagus of I. eptaskouros. (A) Dorsal view. (B) Lateral view. (C) Detail of paramere, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4593947" httpUri="https://zenodo.org/record/4593947/files/figure.png" pageId="14" pageNumber="1778">Figures 6</figureCitation>
|
||
A–C; paramere slightly reduced; paramere: median lobe ratio50.74. Paramere in dorsal view almost parallel-sided converging to rounded tip; in lateral outline almost straight, with two rows of spinules. Median lobe in dorsal view wide, almost parallel-sided, constricted to a narrower rounded apical lobe; in lateral view becoming narrower from middle to apex, apex with a small dorsal tooth, rounded.
|
||
<typeStatus id="0B1B8828776E1C07754AFA6359186CB3" box="[261,371,1491,1515]" pageId="14" pageNumber="1778" type="holotype">Holotype</typeStatus>
|
||
, male, with labels as follows: ‘‘
|
||
<collectingCountry id="ACB7761A776E1C0776A5FA635B4E6CB3" box="[746,805,1491,1515]" name="Peru" pageId="14" pageNumber="1778">Peru</collectingCountry>
|
||
:
|
||
<collectingRegion id="1664F868776E1C07777AFA635B166CB3" box="[821,893,1491,1515]" country="Peru" name="Cusco" pageId="14" pageNumber="1778">Cuzco</collectingRegion>
|
||
Dept., Consuelo, Manu Rd. km 165, x.7.1982’’/‘‘FMHD #82-350, ex litter under crown of felled tree. L. E. Watrous and A. C. Mazurek’’/‘‘
|
||
<taxonomicName id="13A04D09776E1C07763CF9A35ABF6F73" authorityName="Sharp" authorityYear="1876" box="[627,724,1555,1579]" class="Insecta" family="Staphylinidae" genus="Isanopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1778" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E6D4EA98776E1C07763CF9A35ABF6F73" box="[627,724,1555,1579]" italics="true" pageId="14" pageNumber="1778">Isanopus</emphasis>
|
||
</taxonomicName>
|
||
det. Newton 1992’’/‘‘
|
||
<typeStatus id="0B1B8828776E1C0777AFF9A25C256F72" box="[992,1102,1554,1578]" pageId="14" pageNumber="1778" type="holotype">Holotype</typeStatus>
|
||
<taxonomicName id="13A04D09776E1C0774DDF9835A446F12" authority="Chatzimanolis" authorityName="Chatzimanolis" authorityYear="2008" box="[146,559,1586,1611]" class="Insecta" family="Staphylinidae" genus="Isanopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1778" phylum="Arthropoda" rank="species" species="eptaskouros">
|
||
<emphasis id="E6D4EA98776E1C0774DDF98359EB6F13" box="[146,384,1587,1611]" italics="true" pageId="14" pageNumber="1778">Isanopus eptaskouros</emphasis>
|
||
Chatzimanolis
|
||
</taxonomicName>
|
||
, design. Chatzimanolis 2007’’. In the collection of the Field Museum of Natural History, Chicago. The type locality is also known as ‘‘
|
||
<quantity id="13589B6F776E1C0774D2F9C258926FD2" box="[157,249,1650,1674]" metricMagnitude="4" metricUnit="m" metricValue="1.59" pageId="14" pageNumber="1778" unit="km" value="15.9">15.9 km</quantity>
|
||
SW Pilcopata,
|
||
<geoCoordinate id="B194504D776E1C0775E2F9C25A756FD2" box="[429,542,1650,1674]" degrees="13.02369" direction="south" orientation="latitude" pageId="14" pageNumber="1778" precision="1" value="-13.02369">13.02369S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="B194504D776E1C077663F9C25AD66FD2" box="[556,701,1650,1674]" degrees="71.491859" direction="west" orientation="longitude" pageId="14" pageNumber="1778" precision="1" value="-71.49186">71.491859W’</geoCoordinate>
|
||
’ (
|
||
<bibRefCitation id="B0314B7B776E1C07769DF9C25BAF6FD2" author="Patterson BD & Stotz DF & Solari S" box="[722,964,1650,1674]" pageId="14" pageNumber="1778" pagination="3 - 12" refId="ref12224" refString="Patterson BD, Stotz DF, Solari S. 2006. Biological surveys and inventories in Manu. In: Patterson BD, Stotz DF, Solari S, editors. Mammals and Birds of the Manu Biosphere Reserve. Fieldiana: Zoology, n. s., no. 110. p. 3 - 12." type="journal article" year="2006">Patterson et al. 2006</bibRefCitation>
|
||
).
|
||
<typeStatus id="0B1B8828776E1C077797F9C25C206FD2" box="[984,1099,1650,1674]" pageId="14" pageNumber="1778" type="paratype">Paratypes</typeStatus>
|
||
: 3, with same first locality label as the
|
||
<typeStatus id="0B1B8828776E1C07762CF9225AA16FF2" box="[611,714,1682,1706]" pageId="14" pageNumber="1778" type="holotype">holotype</typeStatus>
|
||
, and subsequent labels for each: ‘‘FMHD #82-319, ex leaf litter. L. E. Watrous and A. C. Mazurek’’; ‘‘FMHD #82- 356, ex beating palm branches. L. E. Watrous and A. C. Mazurek’’; and ‘‘FMHD #82-332, ex rotten palm. L. E. Watrous and A. C. Mazurek’’. All
|
||
<typeStatus id="0B1B8828776F1C0677A8FF585C3E6858" box="[999,1109,232,256]" pageId="15" pageNumber="1779" type="paratype">paratypes</typeStatus>
|
||
with label ‘‘
|
||
<typeStatus id="0B1B8828776F1C067554FEB859EF6878" box="[283,388,264,288]" pageId="15" pageNumber="1779" type="paratype">Paratype</typeStatus>
|
||
<taxonomicName id="13A04D09776F1C0675C4FEB85B4A6878" authority="Chatzimanolis" authorityName="Chatzimanolis" authorityYear="2008" box="[395,801,264,288]" class="Insecta" family="Staphylinidae" genus="Isanopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="1779" phylum="Arthropoda" rank="species" species="eptaskouros">
|
||
<emphasis id="E6D4EA98776F1C0675C4FEB85A1C6878" box="[395,631,264,288]" italics="true" pageId="15" pageNumber="1779">Isanopus eptaskouros</emphasis>
|
||
Chatzimanolis
|
||
</taxonomicName>
|
||
design. Chatzimanolis 2007’’. In the collection of the Field Museum of Natural History, Chicago.
|
||
</paragraph>
|
||
<paragraph id="D41F368A776F1C06749AFEC85A5568A8" blockId="15.[213,1169,376,496]" pageId="15" pageNumber="1779">
|
||
<emphasis id="E6D4EA98776F1C06749AFEC8592568C8" box="[213,334,376,400]" italics="true" pageId="15" pageNumber="1779">Etymology</emphasis>
|
||
. The specific epithet is the combination of two Greek words, the word
|
||
<emphasis id="E6D4EA98776F1C06749AFE28596F68E8" box="[213,260,408,432]" italics="true" pageId="15" pageNumber="1779">epta</emphasis>
|
||
(
|
||
<emphasis id="E6D4EA98776F1C06755AFE28592C68E8" box="[277,327,408,432]" italics="true" pageId="15" pageNumber="1779">EPTA</emphasis>
|
||
) that means seven and the word
|
||
<emphasis id="E6D4EA98776F1C06768DFE285B7C68E8" box="[706,791,408,432]" italics="true" pageId="15" pageNumber="1779">skouros</emphasis>
|
||
(
|
||
<emphasis id="E6D4EA98776F1C067767FE285B2E68E8" box="[808,837,408,432]" italics="true" pageId="15" pageNumber="1779">SK</emphasis>
|
||
o
|
||
<emphasis id="E6D4EA98776F1C06771BFE285B1A68E8" box="[852,881,408,432]" italics="true" pageId="15" pageNumber="1779">MR</emphasis>
|
||
o
|
||
<emphasis id="E6D4EA98776F1C0677CFFE285BE768E8" box="[896,908,408,432]" italics="true" pageId="15" pageNumber="1779">z</emphasis>
|
||
) that means dark, and refers to the abdominal segment VII that is completely dark brown to back. The epithet is a noun in apposition.
|
||
</paragraph>
|
||
<paragraph id="D41F368A776F1C06749AFD985BE96B18" blockId="15.[213,898,552,576]" box="[213,898,552,576]" pageId="15" pageNumber="1779">
|
||
<emphasis id="E6D4EA98776F1C06749AFD9859306B18" box="[213,347,552,576]" italics="true" pageId="15" pageNumber="1779">Distribution</emphasis>
|
||
. Known from
|
||
<collectingRegion id="1664F868776F1C067646FD985A3A6B18" box="[521,593,552,576]" country="Peru" name="Cusco" pageId="15" pageNumber="1779">Cuzco</collectingRegion>
|
||
Dept. in
|
||
<collectingCountry id="ACB7761A776F1C06768EFD985A9D6B18" box="[705,758,552,576]" name="Peru" pageId="15" pageNumber="1779">Peru</collectingCountry>
|
||
(
|
||
<figureCitation id="4C9B2A0F776F1C067748FD985B1F6B18" box="[775,884,552,576]" captionStart="Figure 11" captionStartId="26.[146,213,1683,1705]" captionTargetBox="[146,1104,906,1646]" captionTargetId="figure-309@26.[146,1104,906,1646]" captionTargetPageId="26" captionText="Figure 11. Distribution map of I. eptaskouros, I. sahlbergi and I. tenuicornis." figureDoi="http://doi.org/10.5281/zenodo.4593959" httpUri="https://zenodo.org/record/4593959/files/figure.png" pageId="15" pageNumber="1779">Figure 11</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="D41F368A776F1C06749AFDC95BD56BC8" blockId="15.[213,958,632,657]" box="[213,958,632,657]" pageId="15" pageNumber="1779">
|
||
<emphasis id="E6D4EA98776F1C06749AFDC959476BC9" box="[213,300,633,657]" italics="true" pageId="15" pageNumber="1779">Habitat</emphasis>
|
||
. Collected in rotten palms, and leaf litter of rotten trees.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |