192 lines
21 KiB
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192 lines
21 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/asp.82.e111148" ID-Pensoft-Pub="1864-8312-82-151" ID-Pensoft-UUID="8D187145BFCA55A1A73960CDE5BD124B" ID-ZooBank="43AC036E93CC4D79939A07DF54BE1A2D" ModsDocID="1864-8312-82-151" checkinTime="1711188240520" checkinUser="pensoft" docAuthor="Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo & Perrichot, Vincent" docDate="2024" docId="3F5C11CCF27B5F0391E4012059EA0EB3" docLanguage="en" docName="ArthropodSystPhyl 82: 151-181" docOrigin="Arthropod Systematics & amp; Phylogeny 82" docPubDate="2024-03-22" docSource="http://dx.doi.org/10.3897/asp.82.e111148" docTitle="Cretolyra Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot 2024, gen. nov." docType="treatment" docUuid="0190F601-F697-428D-BA66-B522AF1406E4" docUuidSource="ZooBank" docVersion="1" id="8D187145BFCA55A1A73960CDE5BD124B" lastPageNumber="151" masterDocId="8D187145BFCA55A1A73960CDE5BD124B" masterDocTitle="Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities" masterLastPageNumber="181" masterPageNumber="151" pageNumber="151" updateTime="1711188240520" updateUser="pensoft">
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<mods:title id="2B3500B200AED66EEC16FFA6C115F43A">Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities</mods:title>
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<mods:namePart id="147E8E072FAB0C19364702BFBE72605B">Brazidec, Manuel</mods:namePart>
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<mods:nameIdentifier id="2E63401EA0E48275C6B0EE9845487B78" type="ORCID">https://orcid.org/0000-0002-0860-8972</mods:nameIdentifier>
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<mods:affiliation id="175CBE0A71412049CEF9A72CC3974D8D">Univ Rennes, CNRS, Geosciences Rennes, UMR 6118, 35000, Rennes, France & Institut de Syste ́ matique, E ́ volution, Biodiversite ́ (ISYEB), Muse ́ um national d'Histoire naturelle, CNRS, Sorbonne Universite ́, EPHE, Universite ́ des Antilles, CP 50, 57 rue Cuvier, F- 75005 Paris, France & State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, China</mods:affiliation>
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<mods:nameIdentifier id="489F62CA7CA35B3E6742038193E47692" type="email">manuel.brazidec@gmail.com</mods:nameIdentifier>
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<mods:namePart id="A74174FE0094D5B53F20DC4005CF1B18">Vilhelmsen, Lars</mods:namePart>
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<mods:affiliation id="809CC4666FFC8644A22EEDFC82F2ECB2">Natural History Museum of Denmark, SCIENCE, University of Copenhagen, Universitetsparken 15, DK- 2100, Denmark</mods:affiliation>
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<mods:namePart id="9CAFDF1BAADF295E73B49D4280D04986">Boudinot, Brendon E.</mods:namePart>
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<mods:nameIdentifier id="A1674C0F40F3B02D73AD2F83CF93101E" type="ORCID">https://orcid.org/0000-0002-4588-0430</mods:nameIdentifier>
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<mods:affiliation id="62A9CCCB85B76E005246057F5D04845C">Institut fuer Zoologie und Evolutionsforschung, Friedrich-Schiller-Universitaet Jena, Vor dem Neutor 1, 07743 Jena, Germany & National Museum of Natural History, Smithsonian Institution, 10 th & Constitution Ave. NW, Washington, DC, USA & Senckenberg Gesellschaft fuer Naturforschung und Naturmuseum, Frankfurt am Main, 60325 Germany</mods:affiliation>
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<mods:namePart id="22EBB747F0B38B587CC1FDD969BEFF9F">Richter, Adrian</mods:namePart>
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<mods:affiliation id="CDF970A0F7A2E48B03B41DD631DBE28F">Institut fuer Zoologie und Evolutionsforschung, Friedrich-Schiller-Universitaet Jena, Vor dem Neutor 1, 07743 Jena, Germany & Biodiversity and Biocomplexity Unit, Okinawa Institute of Science and Technology Graduate University, Onna-son, Okinawa, Japan</mods:affiliation>
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<mods:namePart id="F35F3AF3C9D05518C27C22B7A973DF4E">Hammel, Joerg U.</mods:namePart>
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<mods:affiliation id="AC1086437B269F39DD3F788041316638">Institute of Materials Physics, Helmholtz-Zentrum Hereon, Max-Planck-Str. 1, 21502 Geesthacht, Germany</mods:affiliation>
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<mods:namePart id="FE5965D29C55A264983D34D67DC5D5A1">Perkovsky, Evgeny E.</mods:namePart>
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<mods:nameIdentifier id="9D4B8E39DE5A33ACDEBF43909878C327" type="ORCID">https://orcid.org/0000-0002-7959-4379</mods:nameIdentifier>
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<mods:affiliation id="16C5EBC4D406B71F4896C69CEC3233BB">Natural History Museum of Denmark, SCIENCE, University of Copenhagen, Universitetsparken 15, DK- 2100, Denmark & I. I. Schmalhausen Institut of Zoology, National Academy of Sciences of Ukraine, B. Khmelnitskogo 15, Kiev 01030, Ukraine</mods:affiliation>
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<mods:namePart id="7AA01D8A8EDA5A5ACFE10C9DC7E0BE4B">Fan, Yong</mods:namePart>
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<mods:affiliation id="544640A2A9040E406F6BE187583DDF0A">Fushun Amber Institute, Fushun 113005, China</mods:affiliation>
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<mods:namePart id="D6AB47793915456EE482EA82E3DF5919">Wang, Zhen</mods:namePart>
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<mods:affiliation id="F5F19685200C69A0F84D1359F465979F">Natural History Museum of Denmark, SCIENCE, University of Copenhagen, Universitetsparken 15, DK- 2100, Denmark & College of Life Sciences, Capital Normal University, 105 Xisanhuanbeilu, Haidian District, Beijing 100048, China</mods:affiliation>
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<mods:namePart id="8965EA8DEF115CBB31F89BA4F4DA4B95">Wu, Qiong</mods:namePart>
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<mods:affiliation id="DBE08B2E3679074336258345FDE7B1B6">Natural History Museum of Denmark, SCIENCE, University of Copenhagen, Universitetsparken 15, DK- 2100, Denmark & College of Life Sciences, Capital Normal University, 105 Xisanhuanbeilu, Haidian District, Beijing 100048, China</mods:affiliation>
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<mods:namePart id="8E94345BF294B91529D682787F47DC91">Wang, Bo</mods:namePart>
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<mods:affiliation id="F4473DA22D118193B271CC70C974FC80">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, China</mods:affiliation>
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<mods:namePart id="7E26C57F5BE36A994BA7CE54047C250B">Perrichot, Vincent</mods:namePart>
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<mods:nameIdentifier id="4D74564B2BC23055B6D88C457D5BFAEA" type="ORCID">https://orcid.org/0000-0002-7973-0430</mods:nameIdentifier>
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<mods:affiliation id="BC4998BAFF088F25DBFC54B7A20CDAA8">Univ Rennes, CNRS, Geosciences Rennes, UMR 6118, 35000, Rennes, France</mods:affiliation>
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<mods:typeOfResource id="2B4F5485EAE25C387B8CB3967B8F72DC">text</mods:typeOfResource>
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<mods:title id="1A90C62F259E4CD7A742459836C64765">Arthropod Systematics & amp; Phylogeny</mods:title>
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<mods:part id="42E03557B34234398847CFF89A9DE010">
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<mods:date id="16471D6CEC5D04852F6E22B63F91BD2E">2024</mods:date>
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<mods:number id="5DBC321768EAF6B8018A3B5AAF070B32">2024-03-22</mods:number>
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<mods:number id="1358FA43877FBCDDF5F12E641B77D72D">82</mods:number>
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<mods:start id="A0187CA98884ED5E894AC96E7B2C889B">151</mods:start>
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<mods:end id="CC0F909B0C13BAEBD62F6824EF158ABE">181</mods:end>
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<mods:url id="9B1575053B93D5152B792268490EE8D1">http://dx.doi.org/10.3897/asp.82.e111148</mods:url>
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<treatment id="3F5C11CCF27B5F0391E4012059EA0EB3" LSID="urn:lsid:zoobank.org:act:0190F601-F697-428D-BA66-B522AF1406E4" httpUri="http://treatment.plazi.org/id/3F5C11CCF27B5F0391E4012059EA0EB3" lastPageNumber="151" pageId="0" pageNumber="151">
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<subSubSection id="747845ED02A7DEB9AE48861B015BDFF9" pageId="0" pageNumber="151" type="nomenclature">
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<paragraph id="D2510F4646735A47AD1B58DDAB412AA0" pageId="0" pageNumber="151">
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3.1.1.1. Genus †
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<taxonomicName id="4F637F3EDB2B19890CA2F523CE908FB8" LSID="https://zoobank.org/0190F601-F697-428D-BA66-B522AF1406E4" authority="Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot, 2024" authorityName="Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot" authorityYear="2024" family="Megalyridae" genus="Cretolyra" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Cretolyra" order="Hymenoptera" pageId="0" pageNumber="151" rank="genus" status="gen. nov.">Cretolyra</taxonomicName>
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<taxonomicNameLabel id="BF5E6CE21BEF242B76824136D6FB14FC" pageId="0" pageNumber="151">gen. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="BBC8B6E89A505CA1A9641F21C23D2123" pageId="0" pageNumber="151" type="type species">
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<paragraph id="73ABDAE209BF505FC2F185B6607EA10D" pageId="0" pageNumber="151">Type species.</paragraph>
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<paragraph id="E887F61A51AB223AD6F862F1A076AE70" pageId="0" pageNumber="151">
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†
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<taxonomicName id="A8D9D9257877F9F185D00776111AECC6" authorityName="Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot" authorityYear="2024" family="Megalyridae" genus="Cretolyra" kingdom="Animalia" lsidName="Cretolyra noijebumensis" order="Hymenoptera" pageId="0" pageNumber="151" rank="species" species="noijebumensis">
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<emphasis id="F86552E6120A2AC0B454823863B7463B" italics="true" pageId="0" pageNumber="151">Cretolyra noijebumensis</emphasis>
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</taxonomicName>
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<emphasis id="B137A74B8281FF7E1C88DF22A0A7E44A" bold="true" pageId="0" pageNumber="151">sp. nov.</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="A8CEDA95B41016E5A3D195164CB05951" pageId="0" pageNumber="151" type="etymology">
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<paragraph id="6407C60950E0EEFFCE4E1D953FE56ADE" pageId="0" pageNumber="151">Etymology.</paragraph>
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<paragraph id="6633496FCB38C5513CBDD92A93A9E78E" pageId="0" pageNumber="151">
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Combination of
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<emphasis id="9EC05C28AF1F3BECD86F1C492AA52280" italics="true" pageId="0" pageNumber="151">Creto</emphasis>
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- referring to the Cretaceous age of the Burmese amber deposit and the suffix -
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<emphasis id="F8C01B6395292C03B033DD0737BD8B88" italics="true" pageId="0" pageNumber="151">lyra</emphasis>
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often used in megalyrid genus names. Gender feminine.
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</paragraph>
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</subSubSection>
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<subSubSection id="341AA64492FB77A73DEC483D0411F63C" pageId="0" pageNumber="151" type="diagnosis">
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<paragraph id="956304ABB9D2605120E85B4BE8A17728" pageId="0" pageNumber="151">Diagnosis.</paragraph>
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<paragraph id="C9C380811DD2254BED953F20BE4B8FCE" pageId="0" pageNumber="151">
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Compound eye oval, higher than long; frons longitudinally divided by sulcus anterior to median ocellus; flagellomere 1 shorter than individual length of flagellomeres 2-5; occipital carina crenulate. Fore wing with M+Cu, Sc+R, A, M and Cu fully pigmented; cu-a almost branching with A, absent for short distance; medial cell rectangular; marginal cell narrow, closed in straight line (Figs
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<figureCitation id="F4702CBBD2D352248F15682ADD2B1B4F" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Drawings of fore wing venation for the taxa described in the present paper. A † Cretolyra noijebumensis gen. et sp. nov.; B † Cretolyra shawi gen. et sp. nov.; C † Genkyhag innebula gen. et sp. nov.; D † Megacoxa chandrahrasa gen. et sp. nov.; E † Megacoxa janzeni gen. et sp. nov.; F † Megacoxa synchrotron gen. et sp. nov.; G † Kamyristi exfrigore gen. et sp. nov.; H † Kamyristi yantardakhensis gen. et sp. nov. Scale bars: 0.5 mm (A - H)." figureDoi="10.3897/asp.82.e111148.figure2" httpUri="https://binary.pensoft.net/fig/1007131" pageId="0" pageNumber="151">2A, B</figureCitation>
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,
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<figureCitation id="3B8ED07245FC32A8DBD26E6522910EC9" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. † Cretolyra noijebumensis gen. et sp. nov., holotype male IGR. BU- 067, mid-Cretaceous, Kachin amber. A habitus in lateral view; B head and mesosoma in dorsal view; C fore wing; D anterior part of mesosoma in lateral view with structures outlined; E metasoma and posterior part of the mesosoma in lateral view. Scale bars: 0.5 mm (A, B, C, E); 0.25 mm (D). Abbreviations: AG = inner axillar groove; MeS = mesometapectal sulcus; MMS = median mesoscutal sulcus; OC = occipital carina; PpL = parapsidal line; sp = anterior thoracic spiracle." figureDoi="10.3897/asp.82.e111148.figure3" httpUri="https://binary.pensoft.net/fig/1007132" pageId="0" pageNumber="151">3C</figureCitation>
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and
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<figureCitation id="52930FC75D6857226A2027AF5B29A9D1" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. † Cretolyra shawi gen. et sp. nov., holotype female NIGP 203545, mid-Cretaceous, Kachin amber. A habitus in dorsal view; B habitus in latero-ventral view; C head and mesosoma in lateral view; D head in frontal view; E line drawing of left scape, pedicel and flagellomeres 1 and 2. Scale bars: 1 mm (A, B); 0.5 mm (C, D, E). Abbreviations: F 1 = flagellomere 1; F 2 = flagellomere 2; MeS = mesometapectal sulcus; pe = pedicel; sc = scape; sp = anterior thoracic spiracle." figureDoi="10.3897/asp.82.e111148.figure4" httpUri="https://binary.pensoft.net/fig/1007133" pageId="0" pageNumber="151">4A</figureCitation>
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). Median mesoscutal sulcus smooth; parapsidal lines present, diverging anteriorly (Figs
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<figureCitation id="706D7774D18FF0D46673CDFEDCF9249D" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. † Cretolyra noijebumensis gen. et sp. nov., holotype male IGR. BU- 067, mid-Cretaceous, Kachin amber. A habitus in lateral view; B head and mesosoma in dorsal view; C fore wing; D anterior part of mesosoma in lateral view with structures outlined; E metasoma and posterior part of the mesosoma in lateral view. Scale bars: 0.5 mm (A, B, C, E); 0.25 mm (D). Abbreviations: AG = inner axillar groove; MeS = mesometapectal sulcus; MMS = median mesoscutal sulcus; OC = occipital carina; PpL = parapsidal line; sp = anterior thoracic spiracle." figureDoi="10.3897/asp.82.e111148.figure3" httpUri="https://binary.pensoft.net/fig/1007132" pageId="0" pageNumber="151">3B</figureCitation>
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and
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<figureCitation id="D80E0335464413EB28D3E23D135042BB" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. † Cretolyra shawi gen. et sp. nov., holotype female NIGP 203545, mid-Cretaceous, Kachin amber. A habitus in dorsal view; B habitus in latero-ventral view; C head and mesosoma in lateral view; D head in frontal view; E line drawing of left scape, pedicel and flagellomeres 1 and 2. Scale bars: 1 mm (A, B); 0.5 mm (C, D, E). Abbreviations: F 1 = flagellomere 1; F 2 = flagellomere 2; MeS = mesometapectal sulcus; pe = pedicel; sc = scape; sp = anterior thoracic spiracle." figureDoi="10.3897/asp.82.e111148.figure4" httpUri="https://binary.pensoft.net/fig/1007133" pageId="0" pageNumber="151">4A</figureCitation>
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). Metacoxa long; metafemur and metatibia swollen; two mesotibial and one metatibial spurs; row of comb-like setae along ventral surface of metatibia and metabasitarsus. Metasoma elongate and narrowed at apex, longer than mesosoma.
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</paragraph>
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<caption id="7F2F02085041706C910E6C3AA25849F6" doi="10.3897/asp.82.e111148.figure4" httpUri="https://binary.pensoft.net/fig/1007133" pageId="0" pageNumber="151" start="Figure 4" startId="F4">
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<paragraph id="9A29D27CC381995A8B840BD63CD9726A" pageId="0" pageNumber="151">
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<emphasis id="18E40C5E9C33ACEEBC22BCF01AD445A8" bold="true" pageId="0" pageNumber="151">Figure 4.</emphasis>
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†
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<taxonomicName id="7B56C7810E5FB8C2545A694F958210A4" authorityName="Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot" authorityYear="2024" family="Megalyridae" genus="Cretolyra" kingdom="Animalia" lsidName="Cretolyra shawi" order="Hymenoptera" pageId="0" pageNumber="151" rank="species" species="shawi">
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<emphasis id="D316A63DD6FF8442980015EFC833A233" italics="true" pageId="0" pageNumber="151">Cretolyra shawi</emphasis>
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</taxonomicName>
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gen. et sp. nov., holotype female NIGP203545, mid-Cretaceous, Kachin amber.
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<emphasis id="EC1ADC7D1FED19D43851B5E296C91986" bold="true" pageId="0" pageNumber="151">A</emphasis>
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habitus in dorsal view;
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<emphasis id="E23567833A08424B9CBABCD14BCF2CF1" bold="true" pageId="0" pageNumber="151">B</emphasis>
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habitus in latero-ventral view;
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<emphasis id="D8009B04471FDDF1D5E07DFD6AB19923" bold="true" pageId="0" pageNumber="151">C</emphasis>
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head and mesosoma in lateral view;
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<emphasis id="B623CADE627B68B74C94F76E1593A105" bold="true" pageId="0" pageNumber="151">D</emphasis>
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head in frontal view;
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<emphasis id="3598554A4D86CF82169572BA253C3605" bold="true" pageId="0" pageNumber="151">E</emphasis>
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line drawing of left scape, pedicel and flagellomeres 1 and 2. Scale bars: 1 mm (
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<emphasis id="E6AD340D2AE9817FE64336E6802FB61F" bold="true" pageId="0" pageNumber="151">A</emphasis>
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,
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<emphasis id="F8A4A00147DB07520F9DFB3D80004996" bold="true" pageId="0" pageNumber="151">B</emphasis>
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); 0.5 mm (
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<emphasis id="743F5FF64B53FC2BE7E418A054B54206" bold="true" pageId="0" pageNumber="151">C</emphasis>
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,
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<emphasis id="D3CCCF09B4F209E541A2DE21CC2BB5A0" bold="true" pageId="0" pageNumber="151">D, E</emphasis>
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). Abbreviations: F1 = flagellomere 1; F2 = flagellomere 2; MeS = mesometapectal sulcus; pe = pedicel; sc = scape; sp = anterior thoracic spiracle.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |