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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1135.93843" ID-Pensoft-Pub="1313-2970-1135-61" ID-Pensoft-UUID="FC0EE843319C574DB0BA3E1125784BD4" ID-ZooBank="7E562C71BD10446E9E102AA52A9888FB" ModsDocID="1313-2970-1135-61" checkinTime="1670952533187" checkinUser="pensoft" docAuthor="Qin, Yuyao, Germann, Christoph &amp; Liang, Hongbin" docDate="2022" docId="148A3BC98D235200AD3F0D3C105A4752" docLanguage="en" docName="ZooKeys 1135: 61-91" docOrigin="ZooKeys 1135" docPubDate="2022-12-12" docSource="http://dx.doi.org/10.3897/zookeys.1135.93843" docTitle="Chlaenius (Sphodromimus) pilosus , new record from China" docType="treatment" docVersion="1" id="FC0EE843319C574DB0BA3E1125784BD4" lastPageNumber="61" masterDocId="FC0EE843319C574DB0BA3E1125784BD4" masterDocTitle="Key to the Chinese species of the subgenus Sphodromimus Casale, 1984 (Carabidae, Chlaeniini, Chlaenius) with descriptions of two new species" masterLastPageNumber="91" masterPageNumber="61" pageNumber="61" updateTime="1670952533187" updateUser="pensoft">
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<mods:title>Key to the Chinese species of the subgenus Sphodromimus Casale, 1984 (Carabidae, Chlaeniini, Chlaenius) with descriptions of two new species</mods:title>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Qin, Yuyao</mods:namePart>
<mods:affiliation>College of Life Sciences, Hebei University, Baoding 071002, China &amp; Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Germann, Christoph</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8317-3799</mods:nameIdentifier>
<mods:affiliation>Naturhistorisches Museum Basel, Augustinergasse 2, CH- 4051 Basel, Switzerland</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Liang, Hongbin</mods:namePart>
<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
<mods:nameIdentifier type="email">lianghb@ioz.ac.cn</mods:nameIdentifier>
</mods:name>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-12-12</mods:number>
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<mods:detail type="volume">
<mods:number>1135</mods:number>
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<mods:extent unit="page">
<mods:start>61</mods:start>
<mods:end>91</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1135.93843</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1135.93843</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1135-61</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:148A3BC98D235200AD3F0D3C105A4752" httpUri="http://treatment.plazi.org/id/148A3BC98D235200AD3F0D3C105A4752" lastPageNumber="61" pageId="0" pageNumber="61">
<subSubSection pageId="0" pageNumber="61" type="nomenclature">
<paragraph pageId="0" pageNumber="61">
<taxonomicName LSID="148A3BC9-8D23-5200-AD3F-0D3C105A4752" authority="(Casale, 1984), new record from China" authorityName=", new record from China" baseAuthorityName="Casale" baseAuthorityYear="1984" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius (Sphodromimus) pilosus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="pilosus" subGenus="Sphodromimus">Chlaenius (Sphodromimus) pilosus (Casale, 1984), new record from China</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="description">
<paragraph pageId="0" pageNumber="61">
<figureCitation captionStart="Figures 5, 6" captionStartId="F4" captionText="Figures 5, 6. 5 A, B Chlaenius (Sphodromimus) peterseni (Louwerens, 1967) (female paratype, Philippine, photos by Shi Hongliang) 6 A, B C. (Sphodromimus) pilosus (Casale, 1984) (male, Yunnan, Pingbian). Scale bar: 5.0 mm." figureDoi="10.3897/zookeys.1135.93843.figures5-6" httpUri="https://binary.pensoft.net/fig/780978" pageId="0" pageNumber="61">Figs 6A, B</figureCitation>
<figureCitation captionStart="Figures 1116" captionStartId="F7" captionText="Figures 11 - 16. Metepisternum features of Sphodromimus spp. 11 Chlaenius (Sphodromimus) caperatus sp. nov., holotype 12 C. (S.) yinggelingensis sp. nov., holotype 13 C. (S.) pilosus (Casale, 1984), male 14 C. (S.) flavofemoratus Laporte, 1834, male 15 C. (S.) davidi nom. nov.; male 16 C. (S.) enleensis Mandl, 1992, male." figureDoi="10.3897/zookeys.1135.93843.figures11-16" httpUri="https://binary.pensoft.net/fig/780981" pageId="0" pageNumber="61">, 13</figureCitation>
<figureCitation captionStart="Figures 1724" captionStartId="F8" captionText="Figures 17 - 24. Pronotum features of Sphodromimus spp. 17 Chlaenius (Sphodromimus) caperatus sp. nov.; holotype 18 C. (S.) hunanus (Morvan, 1997) male 19 C. (S.) deuvei (Morvan, 1997); male 20 C. (S.) pilosus (Casale, 1984), male 21 C. (S.) yinggelingensis sp. nov.; holotype 22 C. (S.) flavofemoratus Laporte, 1834, male 23 C. (S.) davidi nom. nov., male 24 C. (S.) enleensis Mandl, 1992, male. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1135.93843.figures17-24" httpUri="https://binary.pensoft.net/fig/780982" pageId="0" pageNumber="61">, 20</figureCitation>
<figureCitation captionStart="Figures 2830" captionStartId="F10" captionText="Figures 28 - 30. Aedeagus of Sphodromimus spp. 28 Chlaenius (Sphodromimus) pilosus (Casale, 1984) (Yunnan, Pingbian) 29 C. (Sphodromimus) yinggelingensis sp. nov, holotype 30 C. (Sphodromimus) flavofemoratus Laporte, 1834 (Yunnan, Menglun) A apical lamella B left lateral view C dorsal view D ventral view E right lateral view. Left scale bars: 0.5 mm (A); right scale bars: 1.0 mm (B-E)." figureDoi="10.3897/zookeys.1135.93843.figures28-30" httpUri="https://binary.pensoft.net/fig/780984" pageId="0" pageNumber="61">, 28A-E</figureCitation>
<figureCitation captionStart="Figure 40" captionStartId="F14" captionText="Figure 40. Distribution of Chlaenius (Sphodromimus) species in China and adjacent areas (a possible distribution of C. enleensis, En-le, Yunnan, represented by a question mark)." figureDoi="10.3897/zookeys.1135.93843.figure40" httpUri="https://binary.pensoft.net/fig/780988" pageId="0" pageNumber="61">, 40</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="reference_group">
<paragraph pageId="0" pageNumber="61">
<taxonomicName authorityName="Casale" authorityYear="1984" class="Insecta" family="Carabidae" genus="Vachinius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vachinius (Sphodromimus) pilosus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="pilosus" subGenus="Sphodromimus">Vachinius (Sphodromimus) pilosus</taxonomicName>
Casale, 1984: 379;
<bibRefCitation author="Lorenz, W" journalOrPublisher="Wolfgang Lorenz, Tutzing" pageId="0" pageNumber="61" refId="B19" refString="Lorenz, W, 1998. Systematic List of Extant Ground Beetles of the World. Wolfgang Lorenz, Tutzing" title="Systematic List of Extant Ground Beetles of the World." year="1998">Lorenz 1998</bibRefCitation>
: 320 (catalogue);
<bibRefCitation author="Lorenz, W" journalOrPublisher="Wolfgang Lorenz, Tutzing" pageId="0" pageNumber="61" refId="B20" refString="Lorenz, W, 2005. Systematic List of Extant Ground Beetles of the World (InsectaColeopteraGeadephaga: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae), second edition. Wolfgang Lorenz, Tutzing" title="Systematic List of Extant Ground Beetles of the World (InsectaColeopteraGeadephaga: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae), second edition." year="2005">Lorenz 2005</bibRefCitation>
: 341 (catalogue);
<bibRefCitation author="Brunk, I" journalOrPublisher="Zeitschrift der Arbeitsgemeinschaft Oesterreichischer Entomologen" pageId="0" pageNumber="61" pagination="51 - 55" refId="B6" refString="Brunk, I, Kirschenhofer, E, 2016. Vachinius (Vachinius) arakanensis sp.n. (Coleoptera: Carabidae: Chlaeniini) from Myanmar. Zeitschrift der Arbeitsgemeinschaft Oesterreichischer Entomologen 68: 51 - 55" title="Vachinius (Vachinius) arakanensis sp. n. (Coleoptera: Carabidae: Chlaeniini) from Myanmar." volume="68" year="2016">Brunk and Kirschenhofer 2016</bibRefCitation>
: 54 (record);
<bibRefCitation DOI="https://doi.org/10.1080/00305316.2018.1561539" author="Azadbakhsh, S" journalOrPublisher="Oriental Insects" pageId="0" pageNumber="61" pagination="566 - 587" refId="B3" refString="Azadbakhsh, S, Kirschenhofer, E, 2019. A new species and subspecies of genus Chlaenius Bonelli, 1810 with remark on the taxonomic position of Haplochlaenius Lutshnik, 1933 and Vachinius Casale, 1984 and a new synonym of subgenus Macrochlaenites Kuntzen, 1919 (Coleoptera: Carabidae: Chlaeniini). Oriental Insects 53 (4): 566 - 587, DOI: https://doi.org/10.1080/00305316.2018.1561539" title="A new species and subspecies of genus Chlaenius Bonelli, 1810 with remark on the taxonomic position of Haplochlaenius Lutshnik, 1933 and Vachinius Casale, 1984 and a new synonym of subgenus Macrochlaenites Kuntzen, 1919 (Coleoptera: Carabidae: Chlaeniini)." url="https://doi.org/10.1080/00305316.2018.1561539" volume="53" year="2019">Azadbakhsh and Kirschenhofer 2019</bibRefCitation>
: 1 (transferred from genus
<taxonomicName authorityName="Casale" authorityYear="1984" class="Insecta" family="Carabidae" genus="Vachinius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vachinius" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">Vachinius</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="type locality">
<paragraph pageId="0" pageNumber="61">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="61">Vietnam, Chapa, Tonkin, Coll. J. Clermont.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="materials_examined">
<paragraph pageId="0" pageNumber="61">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="61">
<materialsCitation collectingDate="2010-05-23" collectorName="X. D. Yang, X. Y. Zhu" country="China" elevation="2100" location="Dawei Shan" municipality="Pingbian" specimenCount="4" specimenCount-male="4" stateProvince="Yunnan Prov.">
<collectingCountry name="China">China</collectingCountry>
-
<collectingRegion country="China" name="Yunnan">Yunnan Prov.</collectingRegion>
:
<specimenCount type="male">4 ♂♂</specimenCount>
(IZAS),
<collectingMunicipality>Pingbian</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:148A3BC98D235200AD3F0D3C105A4752:7338D258EF49598144351675FAA40F91" country="China" municipality="Pingbian" name="Dawei Shan" stateProvince="Yunnan Prov.">Dawei Shan</location>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.1" unit="m" value="2100.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="2.1" unit="m" value="2100.0">2100 m</elevation>
</quantity>
,
<collectingDate value="2010-05-23">2010.V.23</collectingDate>
,
<collectorName>X.D. Yang</collectorName>
,
<collectorName>X.Y. Zhu</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="diagnosis">
<paragraph pageId="0" pageNumber="61">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="61">
Dorsum black. PW/PL = 1.02-1.07; PAW/PBW = 0.98-1.00 (Fig.
<figureCitation captionStart="Figures 1724" captionStartId="F8" captionText="Figures 17 - 24. Pronotum features of Sphodromimus spp. 17 Chlaenius (Sphodromimus) caperatus sp. nov.; holotype 18 C. (S.) hunanus (Morvan, 1997) male 19 C. (S.) deuvei (Morvan, 1997); male 20 C. (S.) pilosus (Casale, 1984), male 21 C. (S.) yinggelingensis sp. nov.; holotype 22 C. (S.) flavofemoratus Laporte, 1834, male 23 C. (S.) davidi nom. nov., male 24 C. (S.) enleensis Mandl, 1992, male. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1135.93843.figures17-24" httpUri="https://binary.pensoft.net/fig/780982" pageId="0" pageNumber="61">20</figureCitation>
); pronotum subquadrate with anterior angles rounded, slightly projected forward; disc densely and completely rugose, punctate. Elytral intervals flat and densely punctate and pubescent. Hind wings reduced. Metepisterna short, MW/ML = 1.3-1.4 (Fig.
<figureCitation captionStart="Figures 1116" captionStartId="F7" captionText="Figures 11 - 16. Metepisternum features of Sphodromimus spp. 11 Chlaenius (Sphodromimus) caperatus sp. nov., holotype 12 C. (S.) yinggelingensis sp. nov., holotype 13 C. (S.) pilosus (Casale, 1984), male 14 C. (S.) flavofemoratus Laporte, 1834, male 15 C. (S.) davidi nom. nov.; male 16 C. (S.) enleensis Mandl, 1992, male." figureDoi="10.3897/zookeys.1135.93843.figures11-16" httpUri="https://binary.pensoft.net/fig/780981" pageId="0" pageNumber="61">13</figureCitation>
). Apical lamella of median lobe truncated, right side with a large denticulation, left side with a small denticulation (Fig.
<figureCitation captionStart="Figures 2830" captionStartId="F10" captionText="Figures 28 - 30. Aedeagus of Sphodromimus spp. 28 Chlaenius (Sphodromimus) pilosus (Casale, 1984) (Yunnan, Pingbian) 29 C. (Sphodromimus) yinggelingensis sp. nov, holotype 30 C. (Sphodromimus) flavofemoratus Laporte, 1834 (Yunnan, Menglun) A apical lamella B left lateral view C dorsal view D ventral view E right lateral view. Left scale bars: 0.5 mm (A); right scale bars: 1.0 mm (B-E)." figureDoi="10.3897/zookeys.1135.93843.figures28-30" httpUri="https://binary.pensoft.net/fig/780984" pageId="0" pageNumber="61">28A-E</figureCitation>
), sometimes, such denticulation absent.
</paragraph>
<caption doi="10.3897/zookeys.1135.93843.figures7-8" httpUri="https://binary.pensoft.net/fig/780979" pageId="0" pageNumber="61" start="Figures 7, 8" startId="F5">
<paragraph pageId="0" pageNumber="61">
<emphasis bold="true" pageId="0" pageNumber="61">Figures 7, 8.</emphasis>
<emphasis bold="true" pageId="0" pageNumber="61">7A, B</emphasis>
<taxonomicName authorityName="Laporte" authorityYear="1834" class="Insecta" family="Carabidae" genus="Chlaenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlaenius (Sphodromimus) flavofemoratus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="flavofemoratus" subGenus="Sphodromimus">Chlaenius (Sphodromimus) flavofemoratus</taxonomicName>
Laporte, 1834 (male, Yunnan, Menglun)
<emphasis bold="true" pageId="0" pageNumber="61">C</emphasis>
<taxonomicName authorityName="Qin &amp; Germann &amp; Liang" authorityYear="2022" family="Carabidae" kingdom="Animalia" lsidName="C. (Sphodromimus) flavofemoratus" order="Coleoptera" pageId="0" pageNumber="61" rank="species" species="flavofemoratus" subGenus="Sphodromimus">C. (Sphodromimus) flavofemoratus</taxonomicName>
Laporte, 1834 (female, Yunnan, Menglun)
<emphasis bold="true" pageId="0" pageNumber="61">8</emphasis>
<taxonomicName family="Carabidae" kingdom="Animalia" lsidName="C. (Sphodromimus) enleensis" order="Coleoptera" pageId="0" pageNumber="61" rank="species" species="enleensis" subGenus="Sphodromimus">C. (Sphodromimus) enleensis</taxonomicName>
Mandl, 1992 (holotype, Indo Chine) Scale bars: 5.0 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="description">
<paragraph pageId="0" pageNumber="61">Description.</paragraph>
<paragraph pageId="0" pageNumber="61">BL = 19.6-21.9 mm, BW = 7.3-7.6 mm, PAW = 3.33-3.55 mm, PBW = 3.40-3.55 mm, PW = 4.35-4.55 mm, PL = 4.05-4.45 mm, MW = 1.65-1.80 mm, ML = 1.20-1.35 mm. Head, pronotum, elytra, legs, and venter black; antennae, labial and maxillary palpi, apex of mouthparts and tarsomeres dark brown.</paragraph>
<paragraph pageId="0" pageNumber="61">
<emphasis bold="true" pageId="0" pageNumber="61">
<emphasis italics="true" pageId="0" pageNumber="61">Head</emphasis>
.
</emphasis>
Vertex punctate and pubescent with a rugose area; antennae long, reaching middle of elytra; antennomere 3 ~ 1.5
<normalizedToken originalValue="×">x</normalizedToken>
longer than antennomere 4.
</paragraph>
<paragraph pageId="0" pageNumber="61">
<emphasis bold="true" italics="true" pageId="0" pageNumber="61">Pronotum</emphasis>
subquadrate, PW/PL = 1.02-1.07 (Fig.
<figureCitation captionStart="Figures 1724" captionStartId="F8" captionText="Figures 17 - 24. Pronotum features of Sphodromimus spp. 17 Chlaenius (Sphodromimus) caperatus sp. nov.; holotype 18 C. (S.) hunanus (Morvan, 1997) male 19 C. (S.) deuvei (Morvan, 1997); male 20 C. (S.) pilosus (Casale, 1984), male 21 C. (S.) yinggelingensis sp. nov.; holotype 22 C. (S.) flavofemoratus Laporte, 1834, male 23 C. (S.) davidi nom. nov., male 24 C. (S.) enleensis Mandl, 1992, male. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1135.93843.figures17-24" httpUri="https://binary.pensoft.net/fig/780982" pageId="0" pageNumber="61">20</figureCitation>
), widest at apical third; anterior margin slightly concave, its width equal to width of basal margin, PAW/PBW = 0.98-1.00; lateral margins slightly bent before middle, then gently narrowed to base, straight before posterior angles; anterior angles rounded, slightly projected forward; posterior angles right angled, slightly rounded at tips; disc gently convex, sparsely punctate, pubescent, with a few transverse rugosities, without glabrous area in the middle; median line distinct, deep, reaching anterior margin and base; basal foveae deep, short, broad, punctate, and pubescent.
</paragraph>
<paragraph pageId="0" pageNumber="61">
<emphasis bold="true" italics="true" pageId="0" pageNumber="61">Elytra</emphasis>
elongate, EL/BW = 1.64-1.77; gently convex near anterior third, widest near posterior third, rounded at apex in males; striae with deep punctures; parascutellar striae well developed; parascutellar pores present; intervals flat, densely punctate and pubescent; sutural angles sharp; hind wings reduced.
</paragraph>
<paragraph pageId="0" pageNumber="61">
<emphasis bold="true" italics="true" pageId="0" pageNumber="61">Venter</emphasis>
densely punctate, pubescent, metepisterna (Fig.
<figureCitation captionStart="Figures 1116" captionStartId="F7" captionText="Figures 11 - 16. Metepisternum features of Sphodromimus spp. 11 Chlaenius (Sphodromimus) caperatus sp. nov., holotype 12 C. (S.) yinggelingensis sp. nov., holotype 13 C. (S.) pilosus (Casale, 1984), male 14 C. (S.) flavofemoratus Laporte, 1834, male 15 C. (S.) davidi nom. nov.; male 16 C. (S.) enleensis Mandl, 1992, male." figureDoi="10.3897/zookeys.1135.93843.figures11-16" httpUri="https://binary.pensoft.net/fig/780981" pageId="0" pageNumber="61">13</figureCitation>
) short, MW/ML = 1.30-1.38; abdominal sternites III-VI with one setiferous puncture each side, sternite VII with one pair of setiferous punctures in males, two pairs in females; all abdominal sternites with distinct impressions laterally.
</paragraph>
<paragraph pageId="0" pageNumber="61">
<emphasis bold="true" italics="true" pageId="0" pageNumber="61">Legs</emphasis>
long and slender; tarsi nearly smooth dorsally.
</paragraph>
<paragraph pageId="0" pageNumber="61">
<emphasis bold="true" pageId="0" pageNumber="61">
<emphasis italics="true" pageId="0" pageNumber="61">Male genitalia</emphasis>
.
</emphasis>
Median lobe (Fig.
<figureCitation captionStart="Figures 2830" captionStartId="F10" captionText="Figures 28 - 30. Aedeagus of Sphodromimus spp. 28 Chlaenius (Sphodromimus) pilosus (Casale, 1984) (Yunnan, Pingbian) 29 C. (Sphodromimus) yinggelingensis sp. nov, holotype 30 C. (Sphodromimus) flavofemoratus Laporte, 1834 (Yunnan, Menglun) A apical lamella B left lateral view C dorsal view D ventral view E right lateral view. Left scale bars: 0.5 mm (A); right scale bars: 1.0 mm (B-E)." figureDoi="10.3897/zookeys.1135.93843.figures28-30" httpUri="https://binary.pensoft.net/fig/780984" pageId="0" pageNumber="61">28B-E</figureCitation>
) stout, strongly bent to ventral side; apical orifice opening dorsally, long and wide, not reaching basal bulb; in dorsal view, apical lamella (Fig.
<figureCitation captionStart="Figures 2830" captionStartId="F10" captionText="Figures 28 - 30. Aedeagus of Sphodromimus spp. 28 Chlaenius (Sphodromimus) pilosus (Casale, 1984) (Yunnan, Pingbian) 29 C. (Sphodromimus) yinggelingensis sp. nov, holotype 30 C. (Sphodromimus) flavofemoratus Laporte, 1834 (Yunnan, Menglun) A apical lamella B left lateral view C dorsal view D ventral view E right lateral view. Left scale bars: 0.5 mm (A); right scale bars: 1.0 mm (B-E)." figureDoi="10.3897/zookeys.1135.93843.figures28-30" httpUri="https://binary.pensoft.net/fig/780984" pageId="0" pageNumber="61">28A</figureCitation>
) wide and short, wider than long, apex truncated; in left lateral view, apical right side with a large denticulation towards the base and outside, basal left side only thickened and convex or convex into a small denticulation, basal orifice ~ 90 ° relative to preapical shaft; left paramere large and round; right paramere helically curved; endophallus with flagellum coiled; basal part of flagellum with a disciform sclerite facing the right; apical part of flagellum with strip of sclerite.
</paragraph>
<paragraph pageId="0" pageNumber="61">
<emphasis bold="true" italics="true" pageId="0" pageNumber="61">Female genitalia</emphasis>
unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="distribution">
<paragraph pageId="0" pageNumber="61">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="61">China (Yunnan), Vietnam.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="remarks">
<paragraph pageId="0" pageNumber="61">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="61">
The type locality is situated in the mountains of the Black River, northern Vietnam, not far from the Chinese frontier. Our identification is based on the original description and illustration of the male genitalia of the holotype by
<bibRefCitation author="Casale, A" journalOrPublisher="Bollettino del Museo Regionale di Scienze Naturali, Torino" pageId="0" pageNumber="61" pagination="371 - 382" refId="B8" refString="Casale, A, 1984. The new Asiatic genus Vachinius (Carabidae: Callistinae), with three new species. Bollettino del Museo Regionale di Scienze Naturali, Torino 2: 371 - 382" title="The new Asiatic genus Vachinius (Carabidae: Callistinae), with three new species." volume="2" year="1984">Casale (1984</bibRefCitation>
: 379, figs 11, 13).
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