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<document id="9EC1DFF70214986CD025B302522281C7" ID-CLB-Dataset="26866" ID-DOI="10.11646/zootaxa.4603.3.1" ID-GBIF-Dataset="e27c1a28-901e-4fcf-896b-bf3e3e066518" ID-ISSN="1175-5326" ID-Zenodo-Dep="2702204" ID-ZooBank="3EB0C172-76FF-4F4B-BC4F-1853586D0C25" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1557466952685" checkinUser="plazi" docAuthor="Fajfer, Monika" docDate="2019" docId="F01B878A6662FFCFE791FD2EFFF1F894" docLanguage="en" docName="zootaxa.4603.3.1.pdf.imf" docOrigin="Zootaxa 4603 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gnathosoma Krefft 1871" docType="treatment" docVersion="9" lastPageNumber="407" masterDocId="0C22FFF26664FFC9E706FF89FE68FF89" masterDocTitle="Systematics of reptile-associated scale mites of the genus Pterygosoma (Acariformes: Pterygosomatidae) derived from external morphology" masterLastPageNumber="440" masterPageNumber="401" pageNumber="407" updateTime="1698726866814" updateUser="ExternalLinkService">
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<mods:title id="A05899F1579479F4142E7FE8D6273594">Systematics of reptile-associated scale mites of the genus Pterygosoma (Acariformes: Pterygosomatidae) derived from external morphology</mods:title>
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<mods:namePart id="A60A3CFE1FA55225B549D248EAB30F46">Fajfer, Monika</mods:namePart>
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<treatment id="F01B878A6662FFCFE791FD2EFFF1F894" ID-GBIF-Taxon="157119428" LSID="urn:lsid:plazi:treatment:F01B878A6662FFCFE791FD2EFFF1F894" httpUri="http://treatment.plazi.org/id/F01B878A6662FFCFE791FD2EFFF1F894" lastPageNumber="407" pageId="6" pageNumber="407">
<subSubSection id="30A865176662FFCFE791FD2EFDB0FC81" pageId="6" pageNumber="407" type="nomenclature">
<paragraph id="780D369C6662FFCFE791FD2EFF43FD49" blockId="6.[151,1437,679,1821]" box="[151,299,679,704]" pageId="6" pageNumber="407">
<heading id="234581F06662FFCFE791FD2EFF43FD49" bold="true" box="[151,299,679,704]" fontSize="10" level="2" pageId="6" pageNumber="407" reason="2">
<taxonomicName id="BFB24D1F6662FFCFE791FD2EFF43FD49" authorityName="Krefft" authorityYear="1871" box="[151,299,679,704]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="6" pageNumber="407" phylum="Nematoda" rank="genus">
<emphasis id="4AC6EA8E6662FFCFE791FD2EFF43FD49" bold="true" box="[151,299,679,704]" italics="true" pageId="6" pageNumber="407">Gnathosoma</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="780D369C6662FFCFE7C1FD44FDB0FC81" blockId="6.[151,1437,679,1821]" pageId="6" pageNumber="407">
In the genus
<taxonomicName id="BFB24D1F6662FFCFE658FD44FF9AFD6D" authorityName="Fajfer" authorityYear="2019" box="[350,498,717,740]" class="Arachnida" family="Pterygosomatidae" genus="Pterygosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="407" phylum="Arthropoda" rank="genus">
<emphasis id="4AC6EA8E6662FFCFE658FD44FF9AFD6D" box="[350,498,717,740]" italics="true" pageId="6" pageNumber="407">Pterygosoma</emphasis>
</taxonomicName>
the
<taxonomicName id="BFB24D1F6662FFCFE52EFD45FCC7FD6C" authorityName="Krefft" authorityYear="1871" box="[552,687,716,741]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="6" pageNumber="422" phylum="Nematoda" rank="genus">gnathosoma</taxonomicName>
is relatively well-developed, at least three times shorter than the idiosoma and typically situated terminally or subterminally (
<figureCitation id="E0892A196662FFCFE43AFD66FDEBFC81" box="[828,899,751,776]" captionStart="FIGURE 1" captionStartId="7.[151,250,1898,1920]" captionTargetBox="[167,1418,226,1833]" captionTargetId="figure@7.[151,1436,193,1877]" captionTargetPageId="7" captionText="FIGURE 1. Pterygosoma agamae Peters, 1849, general scheme. A, dorsal view; B, ventral view. Abbreviations: perit, peritreme; ant, anterior setae; lat, lateral setae; post, posterior setae; per, peripheral setae; dm, dorso-median setae; vm, ventromedian setae; ps, pseudanal setae; g, genital setae; sca, subcapitulum; c.fs, coxal fields; Tr, trochanter; F, femur; G, genu; Ti, tibia; Ta, tarsus." httpUri="https://zenodo.org/record/2702206/files/figure.png" pageId="6" pageNumber="407">Figs 1</figureCitation>
and
<figureCitation id="E0892A196662FFCFE4BFFD79FDA2FC81" box="[953,970,752,776]" captionStart="FIGURE 2" captionStartId="8.[151,250,1953,1975]" captionTargetBox="[260,1313,210,1912]" captionTargetId="figure@8.[251,1320,193,1934]" captionTargetPageId="8" captionText="FIGURE 2. Male of Pterygosoma sp., general scheme. A, dorsal view; B, ventral view. Abbreviations: ae, aedeagus; ant, anterior setae; c.fs, coxal fields; g.c, genital cone; dm, dorso-median setae; vm, ventro-median setae; perit, peritreme; post.-lat, postero-lateral setae; sca, subcapitulum." httpUri="https://zenodo.org/record/2702208/files/figure.png" pageId="6" pageNumber="407">2</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="30A865176662FFCFE7C1FC9DFFF1F894" pageId="6" pageNumber="407" type="description">
<paragraph id="780D369C6662FFCFE7C1FC9DFAB5FC35" blockId="6.[151,1437,679,1821]" pageId="6" pageNumber="407">
<emphasis id="4AC6EA8E6662FFCFE7C1FC9DFF2AFCA4" box="[199,322,788,813]" italics="true" pageId="6" pageNumber="407">Chelicerae</emphasis>
. The fixed cheliceral digit (
<emphasis id="4AC6EA8E6662FFCFE582FC9DFD67FCA5" box="[644,783,788,813]" italics="true" pageId="6" pageNumber="407">digitus fixus</emphasis>
) is strongly reduced, and may be rounded, membranous, spiniform or branched with few or numerous tines in the apical part (
<figureCitation id="E0892A196662FFCFE4C2FCBEFA46FCD9" box="[964,1070,823,848]" captionStart="FIGURE 4" captionStartId="10.[151,250,1063,1085]" captionTargetBox="[174,1415,276,1033]" captionTargetId="figure@10.[151,1436,265,1042]" captionTargetPageId="10" captionText="FIGURE 4. Fixed cheliceral digit. A, spiniform; B, elongated; C, branched; D, rounded. Hypostomal apex. E, rounded; F, triangular; G, flattened; H, with depression; I, with teeth." httpUri="https://zenodo.org/record/2702212/files/figure.png" pageId="6" pageNumber="407">Fig. 4 A</figureCitation>
̄D). The movable digit (
<emphasis id="4AC6EA8E6662FFCFE256FCBEFE82FCFC" italics="true" pageId="6" pageNumber="407">digitus mobilis</emphasis>
) in most species has twisted so that the terminal point and the basal spur are directed laterally, thus acting as a barb or holdfast, in a similar way to the chelicerae of ticks. The basal spur is absent only in the subgenus
<emphasis id="4AC6EA8E6662FFCFE791FC2DFF23FC34" box="[151,331,932,957]" italics="true" pageId="6" pageNumber="407">Gerrhosaurobia</emphasis>
and the
<taxonomicName id="BFB24D1F6662FFCFE6ADFC2DFF85FC34" authorityName="Fajfer and Gonzalez-Acuna" authorityYear="2013" box="[427,493,932,957]" class="Arachnida" family="Pterygosomatidae" genus="Pterygosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="ligare">
<emphasis id="4AC6EA8E6662FFCFE6ADFC2DFF85FC34" box="[427,493,932,957]" italics="true" pageId="6" pageNumber="407">ligare</emphasis>
</taxonomicName>
group of
<taxonomicName id="BFB24D1F6662FFCFE55CFC2CFC86FC35" authority="Fajfer, 2019" authorityName="Fajfer" authorityYear="2019" box="[602,750,933,956]" class="Arachnida" family="Pterygosomatidae" genus="Pterygosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="407" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="4AC6EA8E6662FFCFE55CFC2CFC86FC35" box="[602,750,933,956]" italics="true" pageId="6" pageNumber="407">Pterygosoma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="51F557F56662FFCFE5F3FC2CFD58FC35" box="[757,816,933,956]" pageId="6" pageNumber="407" sensu="stricto">s. str.</taxonomicNameLabel>
(see
<bibRefCitation id="1C234B6D6662FFCFE46DFC2DFAB9FC35" author="Fajfer, M. &amp; Gonzalez-Acuna, D." box="[875,1233,932,957]" pageId="6" pageNumber="407" pagination="301 - 319" refId="ref17147" refString="Fajfer, M. &amp; Gonzalez-Acuna, D. (2013) Pterygosomatid mites of a new species group ligare (Acariformes: Pterygosomatidae: Pterygosoma) parasitizing tree iguanas (Squamata: Liolaemidae: Liolaemus). Zootaxa, 3693, 301 - 319." type="journal article" year="2013">Fajfer &amp; Gonzalez-Acuna 2013</bibRefCitation>
).
</paragraph>
<paragraph id="780D369C6662FFCFE7C1FC40FDC3FB1D" blockId="6.[151,1437,679,1821]" pageId="6" pageNumber="407">
<emphasis id="4AC6EA8E6662FFCFE7C1FC40FF2DFC69" box="[199,325,969,992]" italics="true" pageId="6" pageNumber="407">Hypostome</emphasis>
. In all pterygosomatids, the hypostome forms a flat or slightly rolled plate in which the chelicerae slide and is situated ventrally, except for mites of the genus
<taxonomicName id="BFB24D1F6662FFCFE443FC65FDAEFB8C" authorityName="Lawrence" authorityYear="1935" box="[837,966,1004,1029]" class="Arachnida" family="Pterygosomatidae" genus="Ixodiderma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="407" phylum="Arthropoda" rank="genus">
<emphasis id="4AC6EA8E6662FFCFE443FC65FDAEFB8C" box="[837,966,1004,1029]" italics="true" pageId="6" pageNumber="407">Ixodiderma</emphasis>
</taxonomicName>
where the hypostome lies dorsally to the chelicerae and is curved round slightly below them. In the genus
<taxonomicName id="BFB24D1F6662FFCFE477FB98FA6DFBA1" authorityName="Fajfer" authorityYear="2019" box="[881,1029,1041,1064]" class="Arachnida" family="Pterygosomatidae" genus="Pterygosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="407" phylum="Arthropoda" rank="genus">
<emphasis id="4AC6EA8E6662FFCFE477FB98FA6DFBA1" box="[881,1029,1041,1064]" italics="true" pageId="6" pageNumber="407">Pterygosoma</emphasis>
</taxonomicName>
the hypostomal apex takes different shapes in various species (
<figureCitation id="E0892A196662FFCFE6C7FBBDFC73FBC4" box="[449,539,1076,1101]" captionStart="FIGURE 4" captionStartId="10.[151,250,1063,1085]" captionTargetBox="[174,1415,276,1033]" captionTargetId="figure@10.[151,1436,265,1042]" captionTargetPageId="10" captionText="FIGURE 4. Fixed cheliceral digit. A, spiniform; B, elongated; C, branched; D, rounded. Hypostomal apex. E, rounded; F, triangular; G, flattened; H, with depression; I, with teeth." httpUri="https://zenodo.org/record/2702212/files/figure.png" pageId="6" pageNumber="407">Fig. 4E</figureCitation>
̄I). It can be rounded, triangular, flat or with a small depression, and its tip may be smooth or with different-sized teeth. Fused palpal coxae are situated basally to the hypostome which form a subcapitulum equipped with one pair of proximal subcapitular setae
<emphasis id="4AC6EA8E6662FFCFE490FBF4FDCCFB1D" box="[918,932,1149,1172]" italics="true" pageId="6" pageNumber="407">n</emphasis>
.
</paragraph>
<paragraph id="780D369C6662FFCFE7C1FB16FDA3FAE4" blockId="6.[151,1437,679,1821]" pageId="6" pageNumber="407">
<emphasis id="4AC6EA8E6662FFCFE7C1FB16FFDFFB31" box="[199,439,1183,1208]" italics="true" pageId="6" pageNumber="407">Palpal segmentation.</emphasis>
The palpal tibia bears a well-developed claw and strongly reduced tarsus (
<figureCitation id="E0892A196662FFCFE215FB16FB31FB31" box="[1299,1369,1183,1208]" captionStart="FIGURE 3" captionStartId="9.[151,250,1406,1428]" captionTargetBox="[169,1429,209,1373]" captionTargetId="figure@9.[150,1436,193,1385]" captionTargetPageId="9" captionText="FIGURE 3. Gnathosoma of Pterygosoma aqabensis Fajfer and Melnikov, 2014. A, dorsal view; B, ventral view. Palpal tarsi of: C, Pterygosoma persicus Hirst, 1926; D, Gerrhosaurobia bicolor (Lawrence, 1936); E, Pterygosoma levissima Fajfer and González-Acuña, 2013." httpUri="https://zenodo.org/record/2702210/files/figure.png" pageId="6" pageNumber="407">Fig. 3</figureCitation>
). The tibial claw is more or less curved and differs in length among different species, only in three species of
<taxonomicName id="BFB24D1F6662FFCFE791FB60FF43FA89" authorityName="Fajfer" authorityYear="2019" box="[151,299,1257,1280]" class="Arachnida" family="Pterygosomatidae" genus="Pterygosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="407" phylum="Arthropoda" rank="genus">
<emphasis id="4AC6EA8E6662FFCFE791FB60FF43FA89" box="[151,299,1257,1280]" italics="true" pageId="6" pageNumber="407">Pterygosoma</emphasis>
</taxonomicName>
(
<taxonomicName id="BFB24D1F6662FFCFE63DFB60FC00FA89" authority="Jack, 1962" authorityName="Jack" authorityYear="1962" box="[315,616,1255,1280]" class="Arachnida" family="Pterygosomatidae" genus="Pterygosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="gracilipalpis">
<emphasis id="4AC6EA8E6662FFCFE63DFB60FF8EFA89" box="[315,486,1255,1280]" italics="true" pageId="6" pageNumber="407">P. gracilipalpis</emphasis>
Jack, 1962
</taxonomicName>
,
<taxonomicName id="BFB24D1F6662FFCFE572FB60FDE7FA89" authority="Jack, 1962" authorityName="Jack" authorityYear="1962" box="[628,911,1255,1280]" class="Arachnida" family="Pterygosomatidae" genus="Pterygosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="tuberculata">
<emphasis id="4AC6EA8E6662FFCFE572FB60FD67FA89" box="[628,783,1255,1280]" italics="true" pageId="6" pageNumber="407">P. tuberculata</emphasis>
Jack, 1962
</taxonomicName>
and
<taxonomicName id="BFB24D1F6662FFCFE4C0FB60FABBFA89" authority="Jack, 1962" authorityName="Jack" authorityYear="1962" box="[966,1235,1255,1280]" class="Arachnida" family="Pterygosomatidae" genus="Pterygosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="caucasica">
<emphasis id="4AC6EA8E6662FFCFE4C0FB60FA27FA89" box="[966,1103,1257,1280]" italics="true" pageId="6" pageNumber="407">P. caucasica</emphasis>
Jack, 1962
</taxonomicName>
) the tibial claw is absent and the eupathidion (
<emphasis id="4AC6EA8E6662FFCFE6DBFA85FF80FAAC" box="[477,488,1292,1317]" italics="true" pageId="6" pageNumber="407">ζ</emphasis>
) is present. In most
<taxonomicName id="BFB24D1F6662FFCFE5DCFA84FD06FAAD" authorityName="Fajfer" authorityYear="2019" box="[730,878,1293,1316]" class="Arachnida" family="Pterygosomatidae" genus="Pterygosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="407" phylum="Arthropoda" rank="genus">
<emphasis id="4AC6EA8E6662FFCFE5DCFA84FD06FAAD" box="[730,878,1293,1316]" italics="true" pageId="6" pageNumber="407">Pterygosoma</emphasis>
</taxonomicName>
species the palpal tarsus is partly or completely fused with the tibia, whereas in the subgenus
<emphasis id="4AC6EA8E6662FFCFE5B6FAA6FD0DFAC1" box="[688,869,1327,1352]" italics="true" pageId="6" pageNumber="407">Gerrhosaurobia</emphasis>
and the
<taxonomicName id="BFB24D1F6662FFCFE4C8FAA6FA78FAC1" authorityName="Fajfer and Gonzalez-Acuna" authorityYear="2013" box="[974,1040,1327,1352]" class="Arachnida" family="Pterygosomatidae" genus="Pterygosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="ligare">
<emphasis id="4AC6EA8E6662FFCFE4C8FAA6FA78FAC1" box="[974,1040,1327,1352]" italics="true" pageId="6" pageNumber="407">ligare</emphasis>
</taxonomicName>
group of
<taxonomicName id="BFB24D1F6662FFCFE38FFAB8FB75FAC1" authority="Fajfer, 2019" authorityName="Fajfer" authorityYear="2019" box="[1161,1309,1329,1352]" class="Arachnida" family="Pterygosomatidae" genus="Pterygosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="407" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="4AC6EA8E6662FFCFE38FFAB8FB75FAC1" box="[1161,1309,1329,1352]" italics="true" pageId="6" pageNumber="407">Pterygosoma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="51F557F56662FFCFE221FAB8FB0DFAC1" box="[1319,1381,1329,1352]" pageId="6" pageNumber="407" sensu="stricto">s. str.</taxonomicNameLabel>
it is rounded, well-articulated and displaced to the ventral surface of the tibia.
</paragraph>
<paragraph id="780D369C6662FFCFE7C1FAFEFAE9FA74" blockId="6.[151,1437,679,1821]" pageId="6" pageNumber="407">
<emphasis id="4AC6EA8E6662FFCFE7C1FAFEFF11FA19" box="[199,377,1399,1424]" italics="true" pageId="6" pageNumber="407">Palpal setation.</emphasis>
In all
<taxonomicName id="BFB24D1F6662FFCFE6C5FAF0FC3FFA19" authorityName="Fajfer" authorityYear="2019" box="[451,599,1401,1424]" class="Arachnida" family="Pterygosomatidae" genus="Pterygosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="407" phylum="Arthropoda" rank="genus">
<emphasis id="4AC6EA8E6662FFCFE6C5FAF0FC3FFA19" box="[451,599,1401,1424]" italics="true" pageId="6" pageNumber="407">Pterygosoma</emphasis>
</taxonomicName>
species the palpal setae are strongly reduced in number. The palpal femur and genu bears one mid-dorsal seta
<emphasis id="4AC6EA8E6662FFCFE523FA15FC2BFA3C" box="[549,579,1436,1461]" italics="true" pageId="6" pageNumber="407">dF</emphasis>
and
<emphasis id="4AC6EA8E6662FFCFE57CFA15FCF1FA3C" box="[634,665,1436,1461]" italics="true" pageId="6" pageNumber="407">dG</emphasis>
, respectively (
<figureCitation id="E0892A196662FFCFE43AFA15FDFEFA3D" box="[828,918,1436,1461]" captionStart="FIGURE 3" captionStartId="9.[151,250,1406,1428]" captionTargetBox="[169,1429,209,1373]" captionTargetId="figure@9.[150,1436,193,1385]" captionTargetPageId="9" captionText="FIGURE 3. Gnathosoma of Pterygosoma aqabensis Fajfer and Melnikov, 2014. A, dorsal view; B, ventral view. Palpal tarsi of: C, Pterygosoma persicus Hirst, 1926; D, Gerrhosaurobia bicolor (Lawrence, 1936); E, Pterygosoma levissima Fajfer and González-Acuña, 2013." httpUri="https://zenodo.org/record/2702210/files/figure.png" pageId="6" pageNumber="407">Fig. 3A</figureCitation>
). The palpal tibia bears three setae: two lateral setae (
<emphasis id="4AC6EA8E6662FFCFE7E7FA36FF60FA51" box="[225,264,1471,1496]" italics="true" pageId="6" pageNumber="407">lTi</emphasis>
,
<emphasis id="4AC6EA8E6662FFCFE61EFA36FF2CFA51" box="[280,324,1471,1496]" italics="true" pageId="6" pageNumber="407">l”Ti</emphasis>
) and one dorsal (
<emphasis id="4AC6EA8E6662FFCFE517FA36FC5DFA51" box="[529,565,1471,1496]" italics="true" pageId="6" pageNumber="407">dTi</emphasis>
) seta (only in
<taxonomicName id="BFB24D1F6662FFCFE5D8FA48FDE2FA51" authorityName="Jack" authorityYear="1962" box="[734,906,1471,1496]" class="Arachnida" family="Pterygosomatidae" genus="Pterygosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="gracilipalpis">
<emphasis id="4AC6EA8E6662FFCFE5D8FA48FDE2FA51" box="[734,906,1471,1496]" italics="true" pageId="6" pageNumber="407">P. gracilipalpis</emphasis>
</taxonomicName>
,
<taxonomicName id="BFB24D1F6662FFCFE49CFA48FA40FA51" authorityName="Jack" authorityYear="1962" box="[922,1064,1473,1496]" class="Arachnida" family="Pterygosomatidae" genus="Pterygosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="caucasica">
<emphasis id="4AC6EA8E6662FFCFE49CFA48FA40FA51" box="[922,1064,1473,1496]" italics="true" pageId="6" pageNumber="407">P. caucasica</emphasis>
</taxonomicName>
and
<taxonomicName id="BFB24D1F6662FFCFE364FA48FA97FA51" authorityName="Jack" authorityYear="1962" box="[1122,1279,1471,1496]" class="Arachnida" family="Pterygosomatidae" genus="Pterygosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="tuberculata">
<emphasis id="4AC6EA8E6662FFCFE364FA48FA97FA51" box="[1122,1279,1471,1496]" italics="true" pageId="6" pageNumber="407">P. tuberculata</emphasis>
</taxonomicName>
an additional seta,
<emphasis id="4AC6EA8E6662FFCFE7D6FA6CFE9AFA75" box="[208,242,1509,1532]" italics="true" pageId="6" pageNumber="407">vTi</emphasis>
, in the form of a eupathidion may be present as a replacement of the tibial claw).
</paragraph>
<paragraph id="780D369C6662FFCFE7C1F981FC8FF926" blockId="6.[151,1437,679,1821]" pageId="6" pageNumber="407">
Among
<taxonomicName id="BFB24D1F6662FFCFE623F980FFD1F9A9" authorityName="Fajfer" authorityYear="2019" box="[293,441,1545,1568]" class="Arachnida" family="Pterygosomatidae" genus="Pterygosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="407" phylum="Arthropoda" rank="genus">
<emphasis id="4AC6EA8E6662FFCFE623F980FFD1F9A9" box="[293,441,1545,1568]" italics="true" pageId="6" pageNumber="407">Pterygosoma</emphasis>
</taxonomicName>
species, the subgenus
<emphasis id="4AC6EA8E6662FFCFE5C2F98EFD11F9A9" box="[708,889,1543,1568]" italics="true" pageId="6" pageNumber="407">Gerrhosaurobia</emphasis>
has the richest tarsal chaetotaxy and possesses four non-modified tactile setae (
<emphasis id="4AC6EA8E6662FFCFE504F9A5FC76F9CC" box="[514,542,1580,1605]" italics="true" pageId="6" pageNumber="407">ba</emphasis>
,
<emphasis id="4AC6EA8E6662FFCFE52AF9A5FC20F9CC" box="[556,584,1580,1605]" italics="true" pageId="6" pageNumber="407">bp</emphasis>
,
<emphasis id="4AC6EA8E6662FFCFE550F9A5FC04F9CC" box="[598,620,1580,1605]" italics="true" pageId="6" pageNumber="407">lp</emphasis>
and
<emphasis id="4AC6EA8E6662FFCFE5A2F9A4FCD7F9CD" box="[676,703,1581,1604]" italics="true" pageId="6" pageNumber="407">va</emphasis>
), one or two terminal eupathidia (
<emphasis id="4AC6EA8E6662FFCFE346F9A5FA37F9CC" box="[1088,1119,1580,1605]" italics="true" pageId="6" pageNumber="407">ul</emphasis>
,
<emphasis id="4AC6EA8E6662FFCFE368F9A5FAE6F9CC" box="[1134,1166,1580,1605]" italics="true" pageId="6" pageNumber="407">sul</emphasis>
) and solenidion
<emphasis id="4AC6EA8E6662FFCFE24AF9A4FB37F9CD" box="[1356,1375,1581,1604]" italics="true" pageId="6" pageNumber="407">ω</emphasis>
(
<figureCitation id="E0892A196662FFCFE268F9A5FED4F9E1" captionStart="FIGURE 3" captionStartId="9.[151,250,1406,1428]" captionTargetBox="[169,1429,209,1373]" captionTargetId="figure@9.[150,1436,193,1385]" captionTargetPageId="9" captionText="FIGURE 3. Gnathosoma of Pterygosoma aqabensis Fajfer and Melnikov, 2014. A, dorsal view; B, ventral view. Palpal tarsi of: C, Pterygosoma persicus Hirst, 1926; D, Gerrhosaurobia bicolor (Lawrence, 1936); E, Pterygosoma levissima Fajfer and González-Acuña, 2013." httpUri="https://zenodo.org/record/2702210/files/figure.png" pageId="6" pageNumber="407">Fig. 3D</figureCitation>
). In representatives of the
<taxonomicName id="BFB24D1F6662FFCFE6E3F9C6FC4FF9E1" authorityName="Fajfer and Gonzalez-Acuna" authorityYear="2013" box="[485,551,1615,1640]" class="Arachnida" family="Pterygosomatidae" genus="Pterygosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="ligare">
<emphasis id="4AC6EA8E6662FFCFE6E3F9C6FC4FF9E1" box="[485,551,1615,1640]" italics="true" pageId="6" pageNumber="407">ligare</emphasis>
</taxonomicName>
group and
<taxonomicName id="BFB24D1F6662FFCFE5A0F9D8FAE9F9E1" authority="Mostafa, 1974" authorityName="Mostafa" authorityYear="1974" box="[678,1153,1615,1640]" class="Arachnida" family="Pterygosomatidae" genus="Pterygosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="aegyptiaca">
<emphasis id="4AC6EA8E6662FFCFE5A0F9D8FD54F9E1" box="[678,828,1617,1640]" italics="true" pageId="6" pageNumber="407">P. aegyptiaca</emphasis>
Mostafa, 1974 (
<bibRefCitation id="1C234B6D6662FFCFE4F2F9C6FA11F9E1" author="Baker, A. S." box="[1012,1145,1615,1640]" pageId="6" pageNumber="407" pagination="1970 - 1988" refId="ref16286" refString="Baker, A. S. (2017) A redescription of Pterygosoma aegyptiaca Mostafa (Acari: Prostigmata: Pterygosomatidae), a little known ectoparasitic mite of spiny-tailed lizards (Squamata: Agamidae), with new morphological data for the Pterygosomatidae. Systematic &amp; Applied Acarology, 22, 1970 - 1988." type="journal article" year="2017">Baker 2017</bibRefCitation>
)
</taxonomicName>
five setae (
<emphasis id="4AC6EA8E6662FFCFE205F9C6FB77F9E1" box="[1283,1311,1615,1640]" italics="true" pageId="6" pageNumber="407">ba</emphasis>
,
<emphasis id="4AC6EA8E6662FFCFE22AF9C6FB20F9E1" box="[1324,1352,1615,1640]" italics="true" pageId="6" pageNumber="407">bp</emphasis>
,
<emphasis id="4AC6EA8E6662FFCFE251F9C6FB05F9E1" box="[1367,1389,1615,1640]" italics="true" pageId="6" pageNumber="407">lp</emphasis>
,
<emphasis id="4AC6EA8E6662FFCFE27CF9D8FBFDF9E1" box="[1402,1429,1617,1640]" italics="true" pageId="6" pageNumber="407">va</emphasis>
,
<emphasis id="4AC6EA8E6662FFCFE791F9FDFEDEF904" box="[151,182,1652,1677]" italics="true" pageId="6" pageNumber="407">ul</emphasis>
) and solenidion
<emphasis id="4AC6EA8E6662FFCFE672F9FCFFEFF905" box="[372,391,1653,1676]" italics="true" pageId="6" pageNumber="407">ω</emphasis>
(
<figureCitation id="E0892A196662FFCFE699F9FDFF9CF905" box="[415,500,1652,1677]" captionStart="FIGURE 3" captionStartId="9.[151,250,1406,1428]" captionTargetBox="[169,1429,209,1373]" captionTargetId="figure@9.[150,1436,193,1385]" captionTargetPageId="9" captionText="FIGURE 3. Gnathosoma of Pterygosoma aqabensis Fajfer and Melnikov, 2014. A, dorsal view; B, ventral view. Palpal tarsi of: C, Pterygosoma persicus Hirst, 1926; D, Gerrhosaurobia bicolor (Lawrence, 1936); E, Pterygosoma levissima Fajfer and González-Acuña, 2013." httpUri="https://zenodo.org/record/2702210/files/figure.png" pageId="6" pageNumber="407">Fig. 3E</figureCitation>
) are present whereas in the remaining species of
<taxonomicName id="BFB24D1F6662FFCFE325F9FCFADFF905" authorityName="Fajfer" authorityYear="2019" box="[1059,1207,1653,1676]" class="Arachnida" family="Pterygosomatidae" genus="Pterygosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="407" phylum="Arthropoda" rank="genus">
<emphasis id="4AC6EA8E6662FFCFE325F9FCFADFF905" box="[1059,1207,1653,1676]" italics="true" pageId="6" pageNumber="407">Pterygosoma</emphasis>
</taxonomicName>
four (
<emphasis id="4AC6EA8E6662FFCFE3F8F9FDFB72F904" box="[1278,1306,1652,1677]" italics="true" pageId="6" pageNumber="407">ba</emphasis>
,
<emphasis id="4AC6EA8E6662FFCFE22FF9FDFB2DF904" box="[1321,1349,1652,1677]" italics="true" pageId="6" pageNumber="407">bp</emphasis>
,
<emphasis id="4AC6EA8E6662FFCFE252F9FDFB02F904" box="[1364,1386,1652,1677]" italics="true" pageId="6" pageNumber="407">lp</emphasis>
,
<emphasis id="4AC6EA8E6662FFCFE27EF9FCFBFBF905" box="[1400,1427,1653,1676]" italics="true" pageId="6" pageNumber="407">va</emphasis>
) (
<figureCitation id="E0892A196662FFCFE799F91EFE9DF939" box="[159,245,1687,1712]" captionStart="FIGURE 3" captionStartId="9.[151,250,1406,1428]" captionTargetBox="[169,1429,209,1373]" captionTargetId="figure@9.[150,1436,193,1385]" captionTargetPageId="9" captionText="FIGURE 3. Gnathosoma of Pterygosoma aqabensis Fajfer and Melnikov, 2014. A, dorsal view; B, ventral view. Palpal tarsi of: C, Pterygosoma persicus Hirst, 1926; D, Gerrhosaurobia bicolor (Lawrence, 1936); E, Pterygosoma levissima Fajfer and González-Acuña, 2013." httpUri="https://zenodo.org/record/2702210/files/figure.png" pageId="6" pageNumber="407">Fig. 3C</figureCitation>
) or three (
<emphasis id="4AC6EA8E6662FFCFE66DF91EFFEFF939" box="[363,391,1687,1712]" italics="true" pageId="6" pageNumber="407">ba</emphasis>
,
<emphasis id="4AC6EA8E6662FFCFE693F91EFFD9F939" box="[405,433,1687,1712]" italics="true" pageId="6" pageNumber="407">bp</emphasis>
and
<emphasis id="4AC6EA8E6662FFCFE6E1F910FC6AF939" box="[487,514,1689,1712]" italics="true" pageId="6" pageNumber="407">va</emphasis>
) simple setae occur.
</paragraph>
<paragraph id="780D369C6662FFCFE7C1F935FFF1F894" blockId="6.[151,1437,679,1821]" pageId="6" pageNumber="407">The respiratory structures, like stigmae and peritremes are situated dorsally. The stigma openings are located between the swollen cheliceral parts in the midline of the chelicerae and the segmented peritremes run from each stigma antero-laterally.</paragraph>
</subSubSection>
</treatment>
</document>