827 lines
174 KiB
XML
827 lines
174 KiB
XML
<document id="4EACD80000EA7700771223F1118BE2F2" ID-CLB-Dataset="298715" ID-DOI="10.11646/zootaxa.5471.2.1" ID-GBIF-Dataset="3a469638-dc21-45a3-80f6-cb307fbf5941" ID-ISSN="1175-5326" ID-Zenodo-Dep="12189904" ID-ZooBank="BAA047F4-2D65-4E52-8798-D426E3B26DA1" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1718892848818" checkinUser="plazi" docAuthor="Kosterin, Oleg E., Onishko, Vladimir V., Ilyina, Elena V., Chepurnov, Grigory Yu. & Blinov, Alexander G." docDate="2024" docId="DB0F87E19119F10EA7D3FF1F7F207CEF" docLanguage="en" docName="zootaxa.5471.2.1.pdf" docOrigin="Zootaxa 5471 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5471.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Coenagrion australocaspicum Dumont & Heidari 1996" docType="treatment" docVersion="5" lastPageNumber="167" masterDocId="2736FF999111F11EA744F92E7A657E46" masterDocTitle="The genus Coenagrion Kirby, 1890 (Odonata: Coenagrionidae) in the Russian part of the Caucasus" masterLastPageNumber="190" masterPageNumber="151" pageNumber="159" updateTime="1718988639371" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="C5652CE4152749B229D91C72BFCF9279">The genus Coenagrion Kirby, 1890 (Odonata: Coenagrionidae) in the Russian part of the Caucasus</mods:title>
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<mods:namePart id="FFE92F9DAF09F25DD6EE401867BA3323">Kosterin, Oleg E.</mods:namePart>
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<mods:affiliation id="F4481E2A207EF61458BEB4E06436F42B">Institute of Cytology & Genetics SB RAS, Acad. Lavrentyev ave. 10, Novosibirsk, 630090, Russia.</mods:affiliation>
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<mods:nameIdentifier id="62DE252FBCB7B7738264C8D3757EB03F" type="email">kosterin@bionet.nsc.ru</mods:nameIdentifier>
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<mods:namePart id="4DFDED59B3BD48C5615CE8B66709EBE6">Onishko, Vladimir V.</mods:namePart>
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<mods:affiliation id="46C9FBDC1BF3DB432DC02DC5843A8CA8">Moscow Zoo, Department of Herpetology, Bolshaya Gruzinskaya Str. 1, Moscow 123242 Russia.</mods:affiliation>
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<mods:nameIdentifier id="8A54FBD2526E357A163BB5F41559C957" type="email">wervolf999@yandex.ru</mods:nameIdentifier>
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<mods:namePart id="A89BC6A71E7178D3447BB25CC3794B7C">Ilyina, Elena V.</mods:namePart>
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<mods:affiliation id="2FAC62FE715118BDA0AB12963B103DCA">Precaspian Institute of Biological Resources of the Daghestan Federal Research Centre of the Russian Academy of Sciences, Dagestanskiy State Nature Reserve, Gagarina Street 120, Makhachkala, Republic of Dagestan, Russia.</mods:affiliation>
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<mods:nameIdentifier id="C530E27D047983F56A2D2C30D08B36DE" type="email">carabus@list.ru</mods:nameIdentifier>
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<mods:namePart id="90EBA383472C4758B3325CB4C4A31F7D">Chepurnov, Grigory Yu.</mods:namePart>
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<mods:affiliation id="24F1199537E05F37E919B868EB4DEB1C">Institute of Cytology & Genetics SB RAS, Acad. Lavrentyev ave. 10, Novosibirsk, 630090, Russia. & Institute of Molecular and Cellular Biology SB RAS, Acad. Lavrentyev ave. 8 / 2, Novosibirsk, 630090, Russia.</mods:affiliation>
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<mods:affiliation id="AD883BEE17E501D9FE3A2C17B90F3829">Institute of Cytology & Genetics SB RAS, Acad. Lavrentyev ave. 10, Novosibirsk, 630090, Russia. & Institute of Molecular and Cellular Biology SB RAS, Acad. Lavrentyev ave. 8 / 2, Novosibirsk, 630090, Russia.</mods:affiliation>
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<treatment id="DB0F87E19119F10EA7D3FF1F7F207CEF" ID-DOI="http://doi.org/10.5281/zenodo.12190715" ID-GBIF-Taxon="233309481" ID-Zenodo-Dep="12190715" LSID="urn:lsid:plazi:treatment:DB0F87E19119F10EA7D3FF1F7F207CEF" httpUri="http://treatment.plazi.org/id/DB0F87E19119F10EA7D3FF1F7F207CEF" lastPageId="16" lastPageNumber="167" pageId="8" pageNumber="159">
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1.
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<taxonomicName id="94A64D749119F116A7F1FF1F7905780D" ID-CoL="WTWH" authority="Dumont & Heidari, 1996" authorityName="Dumont & Heidari" authorityYear="1996" box="[181,864,1585,1612]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="australocaspicum">
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<emphasis id="61D2EAE59119F116A7F1FF1F7844780A" bold="true" box="[181,545,1585,1612]" italics="true" pageId="8" pageNumber="159">Coenagrion australocaspicum</emphasis>
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<bibRefCitation id="37374B069119F116A56CFF1F7905780D" author="Dumont, H. J. & Heidari, H." box="[552,864,1585,1611]" pageId="8" pageNumber="159" pagination="63 - 78" refId="ref27800" refString="Dumont, H. J. & Heidari, H. (1996) On a collection of spring Odonata from Iran, with the description of Coenagrion australocaspicum n. sp. Bulletin et Annales de la Societe Royale Belge d'Entomologie, 132, 63 - 78." type="journal article" year="1996">Dumont & Heidari, 1996</bibRefCitation>
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<paragraph id="531936F79119F116A7D3FF7B7B907836" blockId="8.[151,864,1585,1648]" box="[151,501,1621,1648]" pageId="8" pageNumber="159">
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<figureCitation id="CB9D2A729119F116A7D3FF7B7B637829" box="[151,262,1621,1647]" captionStart="FIGURE 2" captionStartId="9.[151,250,1421,1445]" captionTargetBox="[381,1200,187,1391]" captionTargetId="figure-19@9.[381,1206,181,1397]" captionTargetPageId="9" captionText="FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189910" httpUri="https://zenodo.org/record/12189910/files/figure.png" pageId="8" pageNumber="159">Figs 2a–c</figureCitation>
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,
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<figureCitation id="CB9D2A729119F116A655FF7B7B2C7829" box="[273,329,1621,1647]" captionStart="FIGURE3" captionStartId="10.[151,250,1432,1456]" captionTargetBox="[393,1180,188,1398]" captionTargetId="figure-24@10.[393,1194,181,1408]" captionTargetPageId="10" captionText="FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189912" httpUri="https://zenodo.org/record/12189912/files/figure.png" pageId="8" pageNumber="159">3a–c</figureCitation>
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,
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<figureCitation id="CB9D2A729119F116A610FF7B7BE87829" box="[340,397,1621,1647]" captionStart="FIGURE 4" captionStartId="11.[151,250,1054,1078]" captionTargetBox="[189,1398,181,1021]" captionTargetId="figure-19@11.[189,1398,181,1030]" captionTargetPageId="11" captionText="FIGURE 4. Male appendages of Coenagrion spp. of the puella-group in the broad sense in posterior view: a–b—C. australocaspicum (b—with the appendages expanded); c–d—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); e-f—C. pulchellum; g–k—C. puella; l—C. persicum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c–d—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; e—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; f—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; g–h—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; i—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, a brook and pond, 25.06.2021; j—Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189914" httpUri="https://zenodo.org/record/12189914/files/figure.png" pageId="8" pageNumber="159">4a–b</figureCitation>
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,
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<figureCitation id="CB9D2A729119F116A6DDFF7B7BD77836" box="[409,434,1621,1648]" captionStart="FIGURE 5" captionStartId="13.[151,250,1570,1594]" captionTargetBox="[151,1436,181,1541]" captionTargetId="figure-19@13.[151,1436,181,1546]" captionTargetPageId="13" captionText="FIGURE 5. Male pronotum of Coenagrion spp. of the puella-group in the broad sense in dorsal view: a—C. persicum; b–c—C. pulchellum; d–e—C. ponticum (males with a more (d) and less (e) expanded black abdominal maculation); f—C. australocaspicum; g–l—C. puella; a—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; b—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; c—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; d–e—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; f—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; g —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; h–i—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021; j—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; k—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; l—Russia, Karachay- Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189916" httpUri="https://zenodo.org/record/12189916/files/figure.png" pageId="8" pageNumber="159">5f</figureCitation>
|
||
,
|
||
<figureCitation id="CB9D2A729119F116A6FAFF7B7BAB7829" box="[446,462,1621,1647]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="8" pageNumber="159">6</figureCitation>
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||
,
|
||
<figureCitation id="CB9D2A729119F116A69EFF787B907836" box="[474,501,1622,1648]" captionStart="FIGURE 7" captionStartId="15.[151,250,1565,1589]" captionTargetBox="[447,1140,184,1541]" captionTargetId="figure-19@15.[447,1140,181,1541]" captionTargetPageId="15" captionText="FIGURE 7. Female pronotum of Coenagrion spp. of the puella-group in the broad sense, a–h—in dorsal, with respect to the body main axis (above), and anteriodorsal, perpendicular to the posterior lobe surface (below), views; i–j—in dorsal view only; a—C. australocaspicum; b–g—C. puella; h—C. ponticum; i—C. persicum; j—C. pulchellum; a—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; b—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; c—Sergokala District, 3 km SW of Myurego village, the Inchkheozen’ River upper valley, 19.06.2021; d–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; h—Russia, Chelyabinsk Province, Nagaybakskiy District, small oxbow lakes at the Gumbeyka River left bank at Ostrolenskiy Settlement SE margin, 6.07.2021; i—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; j—Iran, Markazi Ostan, Shazand Shakhrestan, the Garehchay River at Gavigodar village, 34.1078-1111° N 49.3627-3657° E, 1803–1805 m a.s.l., 19.05.2017; k—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189924" httpUri="https://zenodo.org/record/12189924/files/figure.png" pageId="8" pageNumber="159">7a</figureCitation>
|
||
</paragraph>
|
||
<paragraph id="531936F79119F116A7D3FFB37B9B789D" blockId="8.[151,1436,1693,2008]" pageId="8" pageNumber="159">
|
||
<emphasis id="61D2EAE59119F116A7D3FFB37B6278F1" bold="true" box="[151,263,1693,1719]" pageId="8" pageNumber="159">Reported</emphasis>
|
||
(as
|
||
<emphasis id="61D2EAE59119F116A672FFB0787E78F1" box="[310,539,1693,1720]" italics="true" pageId="8" pageNumber="159">
|
||
<taxonomicName id="94A64D749119F116A672FFB07BD978FE" authorityName="Kirby" authorityYear="1890" box="[310,444,1694,1720]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="159" phylum="Arthropoda" rank="genus">Coenagrion</taxonomicName>
|
||
(
|
||
<taxonomicName id="94A64D749119F116A68FFFB3787678F1" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[459,531,1693,1719]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="puella">puella</taxonomicName>
|
||
)
|
||
</emphasis>
|
||
sp2)
|
||
<emphasis id="61D2EAE59119F116A51EFFB378FB78F1" bold="true" box="[602,670,1693,1719]" pageId="8" pageNumber="159">from:</emphasis>
|
||
the Samurskiy Zakaznik Stationary environs, Magaramkent District of
|
||
<collectingRegion id="9162F8159119F116A7F2FFEC7B7B789A" box="[182,286,1730,1756]" country="Russia" name="Dagestan" pageId="8" pageNumber="159">Dagestan</collectingRegion>
|
||
(
|
||
<bibRefCitation id="37374B069119F116A668FFEF7B97789D" author="Ilyina, E. V. & Onishko, V. V. & Kosterin, O. E." box="[300,498,1729,1755]" pageId="8" pageNumber="159" pagination="31 - 50" refId="ref28161" refString="Ilyina, E. V., Onishko, V. V. & Kosterin, O. E. (2022) Strekozy (Odonata) klastera " Del'ta Samura " natsional'nogo parka " Samurskiy " [Dragonflies (Odonata) of the Samur Delta Cluster of Samurskiy National Park']. Trydy Gosudarstvennogo Prirodnogo Zapvednika " Dagestanskiy ", 18, 31 - 50. [in Russian]" type="journal article" year="2022">
|
||
Ilyina
|
||
<emphasis id="61D2EAE59119F116A632FFEC7BCA789D" box="[374,431,1729,1755]" italics="true" pageId="8" pageNumber="159">et al.</emphasis>
|
||
2022
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="531936F79119F112A783FFCB7BD17E93" blockId="8.[151,1436,1693,2008]" lastBlockId="12.[151,1437,150,2013]" lastPageId="12" lastPageNumber="163" pageId="8" pageNumber="159">
|
||
<emphasis id="61D2EAE59119F116A783FFCB7B8F78B9" bold="true" box="[199,490,1765,1791]" pageId="8" pageNumber="159">
|
||
Specimens.
|
||
<materialsCitation id="E3CE3CAA9119F116A61EFFCB791B7965" ID-GBIF-Occurrence="4897992383" collectingDate="2023-06-08" collectorName="E. Ilyina" county="Babayurt District" elevation="10" latitude="43.612" location="Khamamatyurt village" longLatPrecision="68" longitude="46.455" municipality="Terek River" pageId="8" pageNumber="159" specimenCount="1" specimenCount-male="1" stateProvince="Dagestan">
|
||
<collectingRegion id="9162F8159119F116A61EFFCB780378B9" box="[346,614,1765,1791]" country="Russia" name="Dagestan" pageId="8" pageNumber="159">
|
||
Republic of
|
||
<emphasis id="61D2EAE59119F116A6BCFFCB780378B9" bold="true" box="[504,614,1765,1791]" pageId="8" pageNumber="159">Dagestan</emphasis>
|
||
</collectingRegion>
|
||
:
|
||
<specimenCount id="45A0FD7E9119F116A538FFC878F67946" box="[636,659,1766,1792]" count="1" pageId="8" pageNumber="159" type="male">♂</specimenCount>
|
||
,
|
||
<collectingCounty id="BA784E7B9119F116A5E2FFCB791478B9" box="[678,881,1765,1791]" pageId="8" pageNumber="159">Babayurt District</collectingCounty>
|
||
, the
|
||
<collectingMunicipality id="B37DAC8D9119F116A4F6FFCB7E5B78B9" box="[946,1086,1765,1791]" pageId="8" pageNumber="159">Terek River</collectingMunicipality>
|
||
right bank at
|
||
<location id="5679602C9119F116A3B0FFCB7A867965" LSID="urn:lsid:plazi:treatment:DB0F87E19119F10EA7D3FF1F7F207CEF:5679602C9119F116A3B0FFCB7A867965" county="Babayurt District" latitude="43.612" longLatPrecision="68" longitude="46.455" municipality="Terek River" name="Khamamatyurt village" pageId="8" pageNumber="159" stateProvince="Dagestan">Khamamatyurt village</location>
|
||
,
|
||
<geoCoordinate id="369250309119F116A7A3FE277B387962" box="[231,349,1801,1828]" degrees="43.612" direction="north" orientation="latitude" pageId="8" pageNumber="159" precision="55" value="43.612">43.612 N</geoCoordinate>
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||
,
|
||
<geoCoordinate id="369250309119F116A62DFE277BBD7962" box="[361,472,1801,1828]" degrees="46.455" direction="east" orientation="longitude" pageId="8" pageNumber="159" precision="55" value="46.455">46.455 E</geoCoordinate>
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||
,
|
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<elevation id="D88BD1C49119F116A6A1FE2778397965" box="[485,604,1801,1828]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="159" unit="m" value="10.0">
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<quantity id="945E9B129119F116A6A1FE27787A7962" box="[485,543,1801,1828]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="159" unit="m" value="10.0">10 m</quantity>
|
||
a.s.l.
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||
</elevation>
|
||
,
|
||
<date id="271810379119F116A52EFE2778B97965" box="[618,732,1801,1827]" pageId="8" pageNumber="159" value="2023-06-08">
|
||
<collectingDate id="375CE9DF9119F116A52EFE2778B97965" box="[618,732,1801,1827]" pageId="8" pageNumber="159" value="2023-06-08">8.06.2023</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="FE5353219119F116A5ADFE24792F7965" box="[745,842,1801,1828]" pageId="8" pageNumber="159">E. Ilyina</collectorName>
|
||
leg.
|
||
</materialsCitation>
|
||
</emphasis>
|
||
;
|
||
<materialsCitation id="E3CE3CAA9119F116A4CFFE247B57792D" ID-GBIF-Occurrence="4897992349" collectingDate="2021-05-03" collectorName="E. Ilyina" county="Kumtorkala District" elevation="73" latitude="47.223" location="Kumtorkala Station" longLatPrecision="66" longitude="47.246" municipality="Shura-Ozen' River" pageId="8" pageNumber="159" specimenCount="1" specimenCount-male="1" stateProvince="Dagestan">
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||
<specimenCount id="45A0FD7E9119F116A4CFFE2479C77962" box="[907,930,1802,1828]" count="1" pageId="8" pageNumber="159" type="male">♂</specimenCount>
|
||
,
|
||
<collectingCounty id="BA784E7B9119F116A4EBFE277EF67965" box="[943,1171,1801,1827]" pageId="8" pageNumber="159">Kumtorkala District</collectingCounty>
|
||
, the
|
||
<collectingMunicipality id="B37DAC8D9119F116A38DFE277FF97965" box="[1225,1436,1801,1827]" pageId="8" pageNumber="159">Shura-Ozen’ River</collectingMunicipality>
|
||
at
|
||
<location id="5679602C9119F116A7F7FE037BF77901" LSID="urn:lsid:plazi:treatment:DB0F87E19119F10EA7D3FF1F7F207CEF:5679602C9119F116A7F7FE037BF77901" box="[179,402,1837,1863]" county="Kumtorkala District" latitude="47.223" longLatPrecision="66" longitude="47.246" municipality="Shura-Ozen' River" name="Kumtorkala Station" pageId="8" pageNumber="159" stateProvince="Dagestan">Kumtorkala Station</location>
|
||
, next to
|
||
<location id="5679602C9119F116A6B5FE0378A57901" LSID="urn:lsid:plazi:treatment:DB0F87E19119F10EA7D3FF1F7F207CEF:5679602C9119F116A6B5FE0378A57901" box="[497,704,1837,1863]" county="Kumtorkala District" latitude="47.223" longLatPrecision="66" longitude="47.246" municipality="Shura-Ozen' River" name="Sarykum Barkhan" pageId="8" pageNumber="159" stateProvince="Dagestan">Sarykum Barkhan</location>
|
||
,
|
||
<geoCoordinate id="369250309119F116A58FFE03797D7901" box="[715,792,1837,1863]" degrees="42.997" direction="north" orientation="latitude" pageId="8" pageNumber="159" precision="55" value="42.997">42.997</geoCoordinate>
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||
–43.006 N, 47.223–
|
||
<geoCoordinate id="369250309119F116A4BAFE037E09790E" box="[1022,1132,1837,1864]" degrees="47.246" direction="east" orientation="longitude" pageId="8" pageNumber="159" precision="55" value="47.246">47.246 E</geoCoordinate>
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,
|
||
<elevation id="D88BD1C49119F116A33DFE037F7C7901" box="[1145,1305,1837,1864]" metricMagnitude="1" metricUnit="m" metricValue="7.25" metricValueMax="8.0" metricValueMin="6.5" pageId="8" pageNumber="159" unit="m" value="72.5" valueMax="80.0" valueMin="65.0">
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<quantity id="945E9B129119F116A33DFE037EB9790E" box="[1145,1244,1837,1864]" metricMagnitude="1" metricUnit="m" metricValue="7.25" metricValueMax="8.0" metricValueMin="6.5" pageId="8" pageNumber="159" unit="m" value="72.5" valueMax="80.0" valueMin="65.0">65–80 m</quantity>
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a.s.l.
|
||
</elevation>
|
||
,
|
||
<date id="271810379119F116A261FE037FFD790E" box="[1317,1432,1837,1864]" pageId="8" pageNumber="159" value="2021-05-03">
|
||
<collectingDate id="375CE9DF9119F116A261FE037FFD790E" box="[1317,1432,1837,1864]" pageId="8" pageNumber="159" value="2021-05-03">3.05.2021</collectingDate>
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</date>
|
||
,
|
||
<collectorName id="FE5353219119F116A7D3FE7C7A9E792D" box="[151,251,1873,1900]" pageId="8" pageNumber="159">E. Ilyina</collectorName>
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||
leg.
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||
</materialsCitation>
|
||
;
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||
<specimenCount id="45A0FD7E9119F116A605FE7C7B33792A" box="[321,342,1874,1900]" count="1" pageId="8" pageNumber="159" type="male">♂</specimenCount>
|
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(
|
||
<figureCitation id="CB9D2A729119F116A62CFE7F7BA5792D" box="[360,448,1873,1899]" captionStart="FIGURE 2" captionStartId="9.[151,250,1421,1445]" captionTargetBox="[381,1200,187,1391]" captionTargetId="figure-19@9.[381,1206,181,1397]" captionTargetPageId="9" captionText="FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189910" httpUri="https://zenodo.org/record/12189910/files/figure.png" pageId="8" pageNumber="159">Figs 2c</figureCitation>
|
||
,
|
||
<figureCitation id="CB9D2A729119F116A694FE7F7B8F792D" box="[464,490,1873,1899]" captionStart="FIGURE3" captionStartId="10.[151,250,1432,1456]" captionTargetBox="[393,1180,188,1398]" captionTargetId="figure-24@10.[393,1194,181,1408]" captionTargetPageId="10" captionText="FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189912" httpUri="https://zenodo.org/record/12189912/files/figure.png" pageId="8" pageNumber="159">3c</figureCitation>
|
||
) (196433669, 196426386), the same place,
|
||
<date id="271810379119F116A4A8FE7F7E0B792D" box="[1004,1134,1873,1899]" pageId="8" pageNumber="159" value="2021-06-29">29.06.2021</date>
|
||
;
|
||
<materialsCitation id="E3CE3CAA9119F116A338FE7C7B7C79F2" ID-GBIF-Occurrence="4897992379" collectingDate="2023-05-25" elevation="17" latitude="41.9048" location="Derbent District" longLatPrecision="6" longitude="48.4452" pageId="8" pageNumber="159" specimenCount="1" specimenCount-male="1" stateProvince="Dagestan">
|
||
<specimenCount id="45A0FD7E9119F116A338FE7C7EF4792A" box="[1148,1169,1874,1900]" count="1" pageId="8" pageNumber="159" type="male">♂</specimenCount>
|
||
(teneral) (196427366),
|
||
<location id="5679602C9119F116A7D3FE5B7B3579C9" LSID="urn:lsid:plazi:treatment:DB0F87E19119F10EA7D3FF1F7F207CEF:5679602C9119F116A7D3FE5B7B3579C9" box="[151,336,1909,1935]" latitude="41.9048" longLatPrecision="6" longitude="48.4452" name="Derbent District" pageId="8" pageNumber="159" stateProvince="Dagestan">Derbent District</location>
|
||
,
|
||
<locationDeviation id="42AB2FD59119F116A61FFE5B78D779C9" box="[347,690,1909,1936]" location="Belidzhi village" pageId="8" pageNumber="159">
|
||
<quantity id="945E9B129119F116A61FFE5B7BCF79C9" box="[347,426,1909,1936]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="8" pageNumber="159" unit="km" value="3.5">3.5 km</quantity>
|
||
NE of Belidzhi village
|
||
</locationDeviation>
|
||
, thickets at a swamp,
|
||
<geoCoordinate id="369250309119F116A4EDFE5B7E4C79D6" box="[937,1065,1909,1936]" degrees="41.9048" direction="north" orientation="latitude" pageId="8" pageNumber="159" precision="5" value="41.9048">41.9048 N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="369250309119F116A371FE5B7ED479D6" box="[1077,1201,1909,1936]" degrees="48.4452" direction="east" orientation="longitude" pageId="8" pageNumber="159" precision="5" value="48.4452">48.4452 E</geoCoordinate>
|
||
,
|
||
<quantity id="945E9B129119F116A3FAFE5B7F4479D6" box="[1214,1313,1909,1936]" metricMagnitude="1" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="8" pageNumber="159" unit="m" value="17.0" valueMax="18.0" valueMin="16.0">
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<elevation id="D88BD1C49119F116A3FAFE5B7F4479D6" box="[1214,1313,1909,1936]" metricMagnitude="1" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="8" pageNumber="159" unit="m" value="17.0" valueMax="18.0" valueMin="16.0">16–18 m</elevation>
|
||
</quantity>
|
||
below s.l.,
|
||
<date id="271810379119F116A7D3FEB77B7C79F2" box="[151,281,1945,1972]" pageId="8" pageNumber="159" value="2023-05-25">
|
||
<collectingDate id="375CE9DF9119F116A7D3FEB77B7C79F2" box="[151,281,1945,1972]" pageId="8" pageNumber="159" value="2023-05-25">25.05.2023</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="E3CE3CAA9119F115A661FEB77F2278DC" ID-GBIF-Occurrence="4897992339" collectingDate="2021-06-24" county="Magaramkent District" elevation="16" lastPageId="11" lastPageNumber="162" latitude="48.5539" location="Samurskiy Zakaznik Stationary" longLatPrecision="6" longitude="48.5592" municipality="Samur Delta Forest" pageId="8" pageNumber="159" specimenCount="13" specimenCount-female="3" specimenCount-male="10" stateProvince="Primorskiy">
|
||
<specimenCount id="45A0FD7E9119F116A661FEB77B1679F2" box="[293,371,1945,1972]" count="10" pageId="8" pageNumber="159" type="male">10 ♂♂</specimenCount>
|
||
(
|
||
<figureCitation id="CB9D2A729119F116A6C7FEB77B9379F5" box="[387,502,1945,1971]" captionStart="FIGURE 2" captionStartId="9.[151,250,1421,1445]" captionTargetBox="[381,1200,187,1391]" captionTargetId="figure-19@9.[381,1206,181,1397]" captionTargetPageId="9" captionText="FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189910" httpUri="https://zenodo.org/record/12189910/files/figure.png" pageId="8" pageNumber="159">Figs 2a–b</figureCitation>
|
||
,
|
||
<figureCitation id="CB9D2A729119F116A546FEB7785E79F5" box="[514,571,1945,1971]" captionStart="FIGURE3" captionStartId="10.[151,250,1432,1456]" captionTargetBox="[393,1180,188,1398]" captionTargetId="figure-24@10.[393,1194,181,1408]" captionTargetPageId="10" captionText="FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189912" httpUri="https://zenodo.org/record/12189912/files/figure.png" pageId="8" pageNumber="159">3a–b</figureCitation>
|
||
,
|
||
<figureCitation id="CB9D2A729119F116A503FEB778E579F5" box="[583,640,1945,1971]" captionStart="FIGURE 4" captionStartId="11.[151,250,1054,1078]" captionTargetBox="[189,1398,181,1021]" captionTargetId="figure-19@11.[189,1398,181,1030]" captionTargetPageId="11" captionText="FIGURE 4. Male appendages of Coenagrion spp. of the puella-group in the broad sense in posterior view: a–b—C. australocaspicum (b—with the appendages expanded); c–d—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); e-f—C. pulchellum; g–k—C. puella; l—C. persicum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c–d—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; e—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; f—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; g–h—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; i—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, a brook and pond, 25.06.2021; j—Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189914" httpUri="https://zenodo.org/record/12189914/files/figure.png" pageId="8" pageNumber="159">4a–b</figureCitation>
|
||
,
|
||
<figureCitation id="CB9D2A729119F116A5C9FEB778C279F2" box="[653,679,1945,1972]" captionStart="FIGURE 5" captionStartId="13.[151,250,1570,1594]" captionTargetBox="[151,1436,181,1541]" captionTargetId="figure-19@13.[151,1436,181,1546]" captionTargetPageId="13" captionText="FIGURE 5. Male pronotum of Coenagrion spp. of the puella-group in the broad sense in dorsal view: a—C. persicum; b–c—C. pulchellum; d–e—C. ponticum (males with a more (d) and less (e) expanded black abdominal maculation); f—C. australocaspicum; g–l—C. puella; a—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; b—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; c—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; d–e—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; f—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; g —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; h–i—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021; j—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; k—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; l—Russia, Karachay- Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189916" httpUri="https://zenodo.org/record/12189916/files/figure.png" pageId="8" pageNumber="159">5f</figureCitation>
|
||
),
|
||
<specimenCount id="45A0FD7E9119F116A5F9FEB7789879F2" box="[701,765,1945,1972]" count="3" pageId="8" pageNumber="159" type="female">3 ♀♀</specimenCount>
|
||
(
|
||
<figureCitation id="CB9D2A729119F116A449FEB7793879F5" box="[781,861,1945,1971]" captionStart="FIGURE 7" captionStartId="15.[151,250,1565,1589]" captionTargetBox="[447,1140,184,1541]" captionTargetId="figure-19@15.[447,1140,181,1541]" captionTargetPageId="15" captionText="FIGURE 7. Female pronotum of Coenagrion spp. of the puella-group in the broad sense, a–h—in dorsal, with respect to the body main axis (above), and anteriodorsal, perpendicular to the posterior lobe surface (below), views; i–j—in dorsal view only; a—C. australocaspicum; b–g—C. puella; h—C. ponticum; i—C. persicum; j—C. pulchellum; a—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; b—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; c—Sergokala District, 3 km SW of Myurego village, the Inchkheozen’ River upper valley, 19.06.2021; d–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; h—Russia, Chelyabinsk Province, Nagaybakskiy District, small oxbow lakes at the Gumbeyka River left bank at Ostrolenskiy Settlement SE margin, 6.07.2021; i—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; j—Iran, Markazi Ostan, Shazand Shakhrestan, the Garehchay River at Gavigodar village, 34.1078-1111° N 49.3627-3657° E, 1803–1805 m a.s.l., 19.05.2017; k—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189924" httpUri="https://zenodo.org/record/12189924/files/figure.png" pageId="8" pageNumber="159">Fig. 7a</figureCitation>
|
||
) (196425390, 196425415, 196425436, 196425479 (
|
||
<figureCitation id="CB9D2A729119F116A7DBFE937A9C7991" box="[159,249,1981,2007]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="8" pageNumber="159">
|
||
Fig.
|
||
<quantity id="945E9B129119F116A79FFE937A9C7991" box="[219,249,1981,2007]" metricMagnitude="-3" metricUnit="kg" metricValue="6.0" pageId="8" pageNumber="159" unit="g" value="6.0">6g</quantity>
|
||
</figureCitation>
|
||
), 196425508, 196425543, 196425590, 196425634, 196425653, 196425691, 196425722, 196425755, 196425780, 196425808, 196425854, 196425868, 196425900),
|
||
<collectingCounty id="BA784E7B911AF115A42DFF167E007814" box="[873,1125,1592,1618]" pageId="11" pageNumber="162">Magaramkent District</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="B37DAC8D911AF115A336FF177F317815" box="[1138,1364,1592,1619]" pageId="11" pageNumber="162">Samur Delta Forest</collectingMunicipality>
|
||
,
|
||
<locationDeviation id="42AB2FD5911AF115A225FF177B0E7830" location="Primorskiy" pageId="11" pageNumber="162">
|
||
2 km NW of
|
||
<collectingRegion id="9162F815911AF115A7ABFF727B0E7830" box="[239,363,1628,1654]" country="Russia" name="Primorskiy" pageId="11" pageNumber="162">Primorskiy</collectingRegion>
|
||
</locationDeviation>
|
||
village, environs of
|
||
<location id="5679602C911AF115A51FFF7279A47830" LSID="urn:lsid:plazi:treatment:DB0F87E19119F10EA7D3FF1F7F207CEF:5679602C911AF115A51FFF7279A47830" box="[603,961,1628,1654]" county="Magaramkent District" latitude="48.5539" longLatPrecision="6" longitude="48.5592" municipality="Samur Delta Forest" name="Samurskiy Zakaznik Stationary" pageId="11" pageNumber="162" stateProvince="Primorskiy">Samurskiy Zakaznik Stationary</location>
|
||
, a swampy overgrown estuary of a forest rivulet and a nearby forest road,
|
||
<geoCoordinate id="36925030911AF115A556FFAE780878DD" box="[530,621,1664,1691]" degrees="41.8619" direction="north" orientation="latitude" pageId="11" pageNumber="162" precision="5" value="41.8619">41.8619</geoCoordinate>
|
||
–41.8650 N, 48.5539–
|
||
<geoCoordinate id="36925030911AF115A430FFAE799678DD" box="[884,1011,1664,1691]" degrees="48.5592" direction="east" orientation="longitude" pageId="11" pageNumber="162" precision="5" value="48.5592">48.5592 E</geoCoordinate>
|
||
,
|
||
<quantity id="945E9B12911AF115A345FFAE7E5878DD" box="[1025,1085,1664,1691]" metricMagnitude="1" metricUnit="m" metricValue="1.6" pageId="11" pageNumber="162" unit="m" value="16.0">
|
||
<elevation id="D88BD1C4911AF115A345FFAE7E5878DD" box="[1025,1085,1664,1691]" metricMagnitude="1" metricUnit="m" metricValue="1.6" pageId="11" pageNumber="162" unit="m" value="16.0">16 m</elevation>
|
||
</quantity>
|
||
below s.l.,
|
||
<date id="27181037911AF115A381FFAE7F2278DC" box="[1221,1351,1664,1690]" pageId="11" pageNumber="162" value="2021-06-24">
|
||
<collectingDate id="375CE9DF911AF115A381FFAE7F2278DC" box="[1221,1351,1664,1690]" pageId="11" pageNumber="162" value="2021-06-24">24.06.2021</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<specimenCount id="45A0FD7E911AF115A210FFAF7FF278DD" box="[1364,1431,1665,1691]" count="8" pageId="11" pageNumber="162" type="male">8 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="45A0FD7E911AF115A7D3FF8B7ABC78F9" box="[151,217,1701,1727]" count="7" pageId="11" pageNumber="162" type="female">7 ♀♀</specimenCount>
|
||
(196427413, 196427432, 196427528, 196427547 (
|
||
<figureCitation id="CB9D2A72911AF115A46DFF8A791B78F8" box="[809,894,1700,1726]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="11" pageNumber="162">Fig. 6h</figureCitation>
|
||
), 196427578, 196432921 (
|
||
<figureCitation id="CB9D2A72911AF115A3F1FF8A7F6278F8" box="[1205,1287,1700,1726]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="11" pageNumber="162">Fig. 6f</figureCitation>
|
||
); 196503424 (
|
||
<figureCitation id="CB9D2A72911AF115A7DBFFE67A9178A4" box="[159,244,1736,1762]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="11" pageNumber="162">Fig. 6c</figureCitation>
|
||
)), the same place,
|
||
<geoCoordinate id="36925030911AF115A693FFE6785778A5" box="[471,562,1736,1763]" degrees="41.8634" direction="north" orientation="latitude" pageId="11" pageNumber="162" precision="5" value="41.8634">41.8634</geoCoordinate>
|
||
<geoCoordinate id="36925030911AF115A576FFE678FF78A5" box="[562,666,1736,1763]" degrees="41.8653" direction="west" orientation="longitude" pageId="11" pageNumber="162" precision="5" value="-41.8653">
|
||
–
|
||
<geoCoordinate id="36925030911AF115A504FFE678FF78A5" box="[576,666,1736,1763]" degrees="41.8653" direction="north" orientation="latitude" pageId="11" pageNumber="162" precision="5" value="41.8653">41.8653</geoCoordinate>
|
||
</geoCoordinate>
|
||
N,
|
||
<geoCoordinate id="36925030911AF115A597FFE6794A78A5" box="[723,815,1736,1763]" degrees="48.5530" direction="north" orientation="latitude" pageId="11" pageNumber="162" precision="5" value="48.553">48.5530</geoCoordinate>
|
||
<geoCoordinate id="36925030911AF115A46BFFE679F278A5" box="[815,919,1736,1763]" degrees="48.5563" direction="west" orientation="longitude" pageId="11" pageNumber="162" precision="5" value="-48.5563">
|
||
–
|
||
<geoCoordinate id="36925030911AF115A479FFE679F278A5" box="[829,919,1736,1763]" degrees="48.5563" direction="east" orientation="longitude" pageId="11" pageNumber="162" precision="5" value="48.5563">48.5563</geoCoordinate>
|
||
</geoCoordinate>
|
||
E,
|
||
<quantity id="945E9B12911AF115A489FFE77E5178A5" box="[973,1076,1737,1763]" metricMagnitude="1" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="11" pageNumber="162" unit="m" value="19.0" valueMax="20.0" valueMin="18.0">18–20 m</quantity>
|
||
below s.l.,
|
||
<date id="27181037911AF115A385FFE67F2678A5" box="[1217,1347,1736,1763]" pageId="11" pageNumber="162" value="2023-05-26">26.05.2023</date>
|
||
;
|
||
<specimenCount id="45A0FD7E911AF115A216FFE77FF278A5" box="[1362,1431,1737,1763]" count="4" pageId="11" pageNumber="162" type="male">4 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="45A0FD7E911AF115A7D3FFC37ABC7941" box="[151,217,1773,1799]" count="2" pageId="11" pageNumber="162" type="female">2 ♀♀</specimenCount>
|
||
(196430322 (
|
||
<figureCitation id="CB9D2A72911AF115A63FFFC27BAC7940" box="[379,457,1772,1798]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="11" pageNumber="162">Fig. 6i</figureCitation>
|
||
), 196430351, 196430424, 196430513, 196430593, 196430622 (
|
||
<figureCitation id="CB9D2A72911AF115A3EFFFC27F657940" box="[1195,1280,1772,1798]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="11" pageNumber="162">Fig. 6d</figureCitation>
|
||
), 196430665, 196430702, 196430744, 196430788, 196430823, 196430891, 196431009, 196431069, 196431104, 196431208, 196431263, 196431305, 196431337, 196431381, 196433323 (
|
||
<figureCitation id="CB9D2A72911AF115A43FFE1A79B17908" box="[891,980,1844,1870]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="11" pageNumber="162">Fig. 6a</figureCitation>
|
||
), 196431926, 196431963, 196432017, 196432083, 196432115, 196432155), the same place,
|
||
<date id="27181037911AF115A5AFFE7679087935" box="[747,877,1880,1907]" pageId="11" pageNumber="162" value="2023-05-27">27.05.2023</date>
|
||
;
|
||
<materialsCitation id="E3CE3CAA911AF115A432FE777B517998" ID-GBIF-Occurrence="4897992344" collectingDate="2022-06-28" collectorName="E. Ilyina" county="Magaramkent District" elevation="16" latitude="48.5494" location="Samurskiy Zakaznik Stationary" longLatPrecision="6" longitude="48.5674" municipality="Samur Delta Forest" pageId="11" pageNumber="162" specimenCount="2" specimenCount-male="2" stateProvince="Primorskiy">
|
||
<specimenCount id="45A0FD7E911AF115A432FE7779D07935" box="[886,949,1881,1907]" count="2" pageId="11" pageNumber="162" type="male">2 ♂♂</specimenCount>
|
||
,
|
||
<collectingCounty id="BA784E7B911AF115A4FBFE767ED37934" box="[959,1206,1880,1906]" pageId="11" pageNumber="162">Magaramkent District</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="B37DAC8D911AF115A3FBFE777FFD7935" box="[1215,1432,1880,1907]" pageId="11" pageNumber="162">Samur Delta Forest</collectingMunicipality>
|
||
,
|
||
<locationDeviation id="42AB2FD5911AF115A7D3FE537BCA79D0" box="[151,431,1916,1943]" location="Primorskiy" pageId="11" pageNumber="162">
|
||
<quantity id="945E9B12911AF115A7D3FE537AB779D0" box="[151,210,1916,1943]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="11" pageNumber="162" unit="km" value="3.0">3 km</quantity>
|
||
NW of
|
||
<collectingRegion id="9162F815911AF115A677FE527BCA79D0" box="[307,431,1916,1942]" country="Russia" name="Primorskiy" pageId="11" pageNumber="162">Primorskiy</collectingRegion>
|
||
</locationDeviation>
|
||
village, environs of
|
||
<location id="5679602C911AF115A5DBFE527E6079D0" LSID="urn:lsid:plazi:treatment:DB0F87E19119F10EA7D3FF1F7F207CEF:5679602C911AF115A5DBFE527E6079D0" box="[671,1029,1916,1942]" county="Magaramkent District" latitude="48.5494" longLatPrecision="6" longitude="48.5674" municipality="Samur Delta Forest" name="Samurskiy Zakaznik Stationary" pageId="11" pageNumber="162" stateProvince="Primorskiy">Samurskiy Zakaznik Stationary</location>
|
||
(broadly the same place as above), a coastal lake and nearby meadows,
|
||
<geoCoordinate id="36925030911AF115A506FE8E78F879FD" box="[578,669,1952,1979]" degrees="41.8524" direction="north" orientation="latitude" pageId="11" pageNumber="162" precision="5" value="41.8524">41.8524</geoCoordinate>
|
||
–41.8669 N, 48.5494–
|
||
<geoCoordinate id="36925030911AF115A4E2FE8E7E4079FD" box="[934,1061,1952,1979]" degrees="48.5674" direction="east" orientation="longitude" pageId="11" pageNumber="162" precision="5" value="48.5674">48.5674 E</geoCoordinate>
|
||
,
|
||
<quantity id="945E9B12911AF115A371FE8F7EE879FD" box="[1077,1165,1953,1979]" metricMagnitude="1" metricUnit="m" metricValue="1.55" metricValueMax="2.9" metricValueMin="0.2" pageId="11" pageNumber="162" unit="m" value="15.5" valueMax="29.0" valueMin="2.0">
|
||
<elevation id="D88BD1C4911AF115A371FE8F7EE879FD" box="[1077,1165,1953,1979]" metricMagnitude="1" metricUnit="m" metricValue="1.55" metricValueMax="2.9" metricValueMin="0.2" pageId="11" pageNumber="162" unit="m" value="15.5" valueMax="29.0" valueMin="2.0">2–29 m</elevation>
|
||
</quantity>
|
||
below s.l.,
|
||
<date id="27181037911AF115A253FE8E7FFD79FC" box="[1303,1432,1952,1978]" pageId="11" pageNumber="162" value="2022-06-28">
|
||
<collectingDate id="375CE9DF911AF115A253FE8E7FFD79FC" box="[1303,1432,1952,1978]" pageId="11" pageNumber="162" value="2022-06-28">28.06.2022</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="FE535321911AF115A7D3FEEB7A997998" box="[151,252,1988,2015]" pageId="11" pageNumber="162">E. Ilyina</collectorName>
|
||
leg.
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="E3CE3CAA911AF112A600FEEB7BD57E93" ID-GBIF-Occurrence="4897992324" collectingDate="2023-05-26" elevation="15" lastPageId="12" lastPageNumber="163" latitude="41.871" location="Samur Delta forest" longLatPrecision="6" longitude="48.5382" municipality="Magaramkent District" pageId="11" pageNumber="162" specimenCount="15" specimenCount-female="1" specimenCount-male="14" stateProvince="Primorskiy">
|
||
<specimenCount id="45A0FD7E911AF115A600FEEB7BFD7999" box="[324,408,1989,2015]" count="14" pageId="11" pageNumber="162" type="male">14 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="45A0FD7E911AF115A6ECFEEB7BD87999" box="[424,445,1989,2015]" count="1" pageId="11" pageNumber="162" type="female">♀</specimenCount>
|
||
(196432912, 196432913, 196432914, 196432915, 196432917 (
|
||
<figureCitation id="CB9D2A72911AF115A3E2FEEA7E9B7998" box="[1190,1278,1988,2014]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="11" pageNumber="162">Fig. 6b</figureCitation>
|
||
), 196432918, 196432920),
|
||
<collectingMunicipality id="B37DAC8D911DF112A668F9B978437EF7" box="[300,550,151,177]" pageId="12" pageNumber="163">Magaramkent District</collectingMunicipality>
|
||
,
|
||
<location id="5679602C911DF112A574F9B979627EF7" LSID="urn:lsid:plazi:treatment:DB0F87E19119F10EA7D3FF1F7F207CEF:5679602C911DF112A574F9B979627EF7" box="[560,775,151,177]" latitude="41.871" longLatPrecision="6" longitude="48.5382" municipality="Magaramkent District" name="Samur Delta forest" pageId="12" pageNumber="163" stateProvince="Primorskiy">Samur Delta forest</location>
|
||
, at the 700 years old plane tree,
|
||
<geoCoordinate id="36925030911DF112A32AF9B97E8B7EF7" box="[1134,1262,151,177]" degrees="41.8710" direction="north" orientation="latitude" pageId="12" pageNumber="163" precision="5" value="41.871">41.8710 N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="36925030911DF112A3BDF9B87F117EF7" box="[1273,1396,150,177]" degrees="48.5382" direction="east" orientation="longitude" pageId="12" pageNumber="163" precision="5" value="48.5382">48.5382 E</geoCoordinate>
|
||
,
|
||
<quantity id="945E9B12911DF112A2C4F9B87AC87E93" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="163" unit="m" value="15.0">
|
||
<elevation id="D88BD1C4911DF112A2C4F9B87AC87E93" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="163" unit="m" value="15.0">15 m</elevation>
|
||
</quantity>
|
||
below s.l.,
|
||
<date id="27181037911DF112A66AF9947BD57E93" box="[302,432,186,213]" pageId="12" pageNumber="163" value="2023-05-26">
|
||
<collectingDate id="375CE9DF911DF112A66AF9947BD57E93" box="[302,432,186,213]" pageId="12" pageNumber="163" value="2023-05-26">26.05.2023</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="07D9667F9118F117A7D3FCA379CE79A2" ID-DOI="http://doi.org/10.5281/zenodo.12189910" ID-Zenodo-Dep="12189910" httpUri="https://zenodo.org/record/12189910/files/figure.png" pageId="9" pageNumber="160" startId="9.[151,250,1421,1445]" targetBox="[381,1200,187,1391]" targetPageId="9" targetType="figure">
|
||
<paragraph id="531936F79118F117A7D3FCA379CE79A2" blockId="9.[151,1437,1421,2021]" pageId="9" pageNumber="160">
|
||
<emphasis id="61D2EAE59118F117A7D3FCA37B717BE3" bold="true" box="[151,276,1421,1445]" pageId="9" pageNumber="160">FIGURE 2.</emphasis>
|
||
Male appendages of
|
||
<taxonomicName id="94A64D749118F117A6B7FCA378FB7BE2" box="[499,670,1421,1444]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="61D2EAE59118F117A6B7FCA378097BE2" box="[499,620,1421,1444]" italics="true" pageId="9" pageNumber="160">Coenagrion</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
of the
|
||
<taxonomicName id="94A64D749118F117A5ADFCA3794D7BE3" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[745,808,1421,1445]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="puella">
|
||
<emphasis id="61D2EAE59118F117A5ADFCA3794D7BE3" box="[745,808,1421,1445]" italics="true" pageId="9" pageNumber="160">puella</emphasis>
|
||
</taxonomicName>
|
||
-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—
|
||
<taxonomicName id="94A64D749118F117A63FFC9F78357B8F" authorityName="Dumont & Heidari" authorityYear="1996" box="[379,592,1457,1481]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="australocaspicum">
|
||
<emphasis id="61D2EAE59118F117A63FFC9F78357B8F" box="[379,592,1457,1481]" italics="true" pageId="9" pageNumber="160">C. australocaspicum</emphasis>
|
||
</taxonomicName>
|
||
(b—with the appendages expanded); d-k—
|
||
<taxonomicName id="94A64D749118F117A35AFC9F7E1B7B8F" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1054,1150,1457,1481]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="puella">
|
||
<emphasis id="61D2EAE59118F117A35AFC9F7E1B7B8F" box="[1054,1150,1457,1481]" italics="true" pageId="9" pageNumber="160">C. puella</emphasis>
|
||
</taxonomicName>
|
||
; l–m—
|
||
<taxonomicName id="94A64D749118F117A394FC9F7F2A7B8F" baseAuthorityName="Bartenef" baseAuthorityYear="1929" box="[1232,1359,1457,1481]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="ponticum">
|
||
<emphasis id="61D2EAE59118F117A394FC9F7F2A7B8F" box="[1232,1359,1457,1481]" italics="true" pageId="9" pageNumber="160">C. ponticum</emphasis>
|
||
</taxonomicName>
|
||
(males with a more (c) and less (d) expanded black abdominal maculation); n—
|
||
<taxonomicName id="94A64D749118F117A4E6FCFB7E457BAB" authorityName="Lohmann" authorityYear="1983" box="[930,1056,1493,1517]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="persicum">
|
||
<emphasis id="61D2EAE59118F117A4E6FCFB7E457BAB" box="[930,1056,1493,1517]" italics="true" pageId="9" pageNumber="160">C. persicum</emphasis>
|
||
</taxonomicName>
|
||
; o–p—
|
||
<taxonomicName id="94A64D749118F117A335FCFB7F667BAB" baseAuthorityName="Vander Linden" baseAuthorityYear="1825" box="[1137,1283,1493,1517]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="pulchellum">
|
||
<emphasis id="61D2EAE59118F117A335FCFB7F667BAB" box="[1137,1283,1493,1517]" italics="true" pageId="9" pageNumber="160">C. pulchellum</emphasis>
|
||
</taxonomicName>
|
||
; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="07D9667F911BF114A7D3FCB67BA779A9" ID-DOI="http://doi.org/10.5281/zenodo.12189912" ID-Zenodo-Dep="12189912" httpUri="https://zenodo.org/record/12189912/files/figure.png" pageId="10" pageNumber="161" startId="10.[151,250,1432,1456]" targetBox="[393,1180,188,1398]" targetPageId="10" targetType="figure">
|
||
<paragraph id="531936F7911BF114A7D3FCB67BA779A9" blockId="10.[151,1437,1432,2032]" pageId="10" pageNumber="161">
|
||
<emphasis id="61D2EAE5911BF114A7D3FCB67B757BF6" bold="true" box="[151,272,1432,1456]" pageId="10" pageNumber="161">FIGURE 3.</emphasis>
|
||
Male appendages of
|
||
<taxonomicName id="94A64D74911BF114A69AFCB678E07BE9" box="[478,645,1432,1455]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="61D2EAE5911BF114A69AFCB678327BE9" box="[478,599,1432,1455]" italics="true" pageId="10" pageNumber="161">Coenagrion</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
of the
|
||
<taxonomicName id="94A64D74911BF114A587FCB679677BF6" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[707,770,1432,1456]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="puella">
|
||
<emphasis id="61D2EAE5911BF114A587FCB679677BF6" box="[707,770,1432,1456]" italics="true" pageId="10" pageNumber="161">puella</emphasis>
|
||
</taxonomicName>
|
||
-group in the broad sense in lateral view: a–c—
|
||
<taxonomicName id="94A64D74911BF114A389FCB67FF97BF6" authorityName="Dumont & Heidari" authorityYear="1996" box="[1229,1436,1432,1456]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="australocaspicum">
|
||
<emphasis id="61D2EAE5911BF114A389FCB67FF97BF6" box="[1229,1436,1432,1456]" italics="true" pageId="10" pageNumber="161">C. australocaspicum</emphasis>
|
||
</taxonomicName>
|
||
(b—with the appendages expanded); d–k—
|
||
<taxonomicName id="94A64D74911BF114A513FC9278D07B92" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[599,693,1468,1492]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="puella">
|
||
<emphasis id="61D2EAE5911BF114A513FC9278D07B92" box="[599,693,1468,1492]" italics="true" pageId="10" pageNumber="161">C. puella</emphasis>
|
||
</taxonomicName>
|
||
; l–m—
|
||
<taxonomicName id="94A64D74911BF114A441FC9279E77B92" baseAuthorityName="Bartenef" baseAuthorityYear="1929" box="[773,898,1468,1492]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="ponticum">
|
||
<emphasis id="61D2EAE5911BF114A441FC9279E77B92" box="[773,898,1468,1492]" italics="true" pageId="10" pageNumber="161">C. ponticum</emphasis>
|
||
</taxonomicName>
|
||
(males with a more (c) and less (d) expanded black abdominal maculation); n—
|
||
<taxonomicName id="94A64D74911BF114A6FFFCCE785D7BBE" authorityName="Lohmann" authorityYear="1983" box="[443,568,1504,1528]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="persicum">
|
||
<emphasis id="61D2EAE5911BF114A6FFFCCE785D7BBE" box="[443,568,1504,1528]" italics="true" pageId="10" pageNumber="161">C. persicum</emphasis>
|
||
</taxonomicName>
|
||
; o–q—
|
||
<taxonomicName id="94A64D74911BF114A5CCFCCE797C7BBE" baseAuthorityName="Vander Linden" baseAuthorityYear="1825" box="[648,793,1504,1528]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="pulchellum">
|
||
<emphasis id="61D2EAE5911BF114A5CCFCCE797C7BBE" box="[648,793,1504,1528]" italics="true" pageId="10" pageNumber="161">C. pulchellum</emphasis>
|
||
</taxonomicName>
|
||
; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="07D9667F911AF115A7D3FD30789A7BA0" ID-DOI="http://doi.org/10.5281/zenodo.12189914" ID-Zenodo-Dep="12189914" httpUri="https://zenodo.org/record/12189914/files/figure.png" pageId="11" pageNumber="162" startId="11.[151,250,1054,1078]" targetBox="[189,1398,181,1021]" targetPageId="11" targetType="figure">
|
||
<paragraph id="531936F7911AF115A7D3FD30789A7BA0" blockId="11.[151,1437,1054,1511]" pageId="11" pageNumber="162">
|
||
<emphasis id="61D2EAE5911AF115A7D3FD307B7C7A70" bold="true" box="[151,281,1054,1078]" pageId="11" pageNumber="162">FIGURE 4.</emphasis>
|
||
Male appendages of
|
||
<taxonomicName id="94A64D74911AF115A548FD3178D97A70" box="[524,700,1055,1078]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="61D2EAE5911AF115A548FD3178E07A70" box="[524,645,1055,1078]" italics="true" pageId="11" pageNumber="162">Coenagrion</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
of the
|
||
<taxonomicName id="94A64D74911AF115A451FD3079317A70" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[789,852,1054,1078]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="puella">
|
||
<emphasis id="61D2EAE5911AF115A451FD3079317A70" box="[789,852,1054,1078]" italics="true" pageId="11" pageNumber="162">puella</emphasis>
|
||
</taxonomicName>
|
||
-group in the broad sense in posterior view: a–b—
|
||
<taxonomicName id="94A64D74911AF115A2C1FD317B2E7A1C" authorityName="Dumont & Heidari" authorityYear="1996" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="australocaspicum">
|
||
<emphasis id="61D2EAE5911AF115A2C1FD317B2E7A1C" italics="true" pageId="11" pageNumber="162">C. australocaspicum</emphasis>
|
||
</taxonomicName>
|
||
(b—with the appendages expanded); c–d—
|
||
<taxonomicName id="94A64D74911AF115A466FD6D79C77A1C" baseAuthorityName="Bartenef" baseAuthorityYear="1929" box="[802,930,1091,1114]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="ponticum">
|
||
<emphasis id="61D2EAE5911AF115A466FD6D79C77A1C" box="[802,930,1091,1114]" italics="true" pageId="11" pageNumber="162">C. ponticum</emphasis>
|
||
</taxonomicName>
|
||
(males with a more (c) and less (d) expanded black abdominal maculation); e-f—
|
||
<taxonomicName id="94A64D74911AF115A548FD4978FB7A38" baseAuthorityName="Vander Linden" baseAuthorityYear="1825" box="[524,670,1126,1150]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="pulchellum">
|
||
<emphasis id="61D2EAE5911AF115A548FD4978FB7A38" box="[524,670,1126,1150]" italics="true" pageId="11" pageNumber="162">C. pulchellum</emphasis>
|
||
</taxonomicName>
|
||
; g–k—
|
||
<taxonomicName id="94A64D74911AF115A5AAFD49792A7A38" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[750,847,1126,1150]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="puella">
|
||
<emphasis id="61D2EAE5911AF115A5AAFD49792A7A38" box="[750,847,1126,1150]" italics="true" pageId="11" pageNumber="162">C. puella</emphasis>
|
||
</taxonomicName>
|
||
; l—
|
||
<taxonomicName id="94A64D74911AF115A4C4FD49799B7A38" authorityName="Lohmann" authorityYear="1983" box="[896,1022,1127,1150]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="persicum">
|
||
<emphasis id="61D2EAE5911AF115A4C4FD49799B7A38" box="[896,1022,1127,1150]" italics="true" pageId="11" pageNumber="162">C. persicum</emphasis>
|
||
</taxonomicName>
|
||
; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c–d—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; e—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; f—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; g–h—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; i—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, a brook and pond, 25.06.2021; j—Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017. Scale bar 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="531936F7911DF112A783F9F079137CEF" blockId="12.[151,1437,150,2013]" pageId="12" pageNumber="163">
|
||
<emphasis id="61D2EAE5911DF112A783F9F0797D7EBE" bold="true" box="[199,792,222,248]" pageId="12" pageNumber="163">
|
||
Additional photographic registrations.
|
||
<materialsCitation id="E3CE3CAA911DF112A5D5F9F07BB27F23" ID-GBIF-Occurrence="4897992307" collectingDate="2021-06-24" county="Magaramkent District" elevation="17" latitude="48.5576" location="Samurskiy Zakaznik Stationary" longLatPrecision="6" longitude="48.5592" municipality="Samur Delta Forest" pageId="12" pageNumber="163" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Dagestan">
|
||
<collectingRegion id="9162F815911DF112A5D5F9F079EE7EBF" box="[657,907,222,249]" country="Russia" name="Dagestan" pageId="12" pageNumber="163">
|
||
Republic of
|
||
<emphasis id="61D2EAE5911DF112A459F9F179EE7EBF" bold="true" box="[797,907,223,249]" pageId="12" pageNumber="163">Dagestan</emphasis>
|
||
</collectingRegion>
|
||
:
|
||
<specimenCount id="45A0FD7E911DF112A4D3F9F179A67EBF" box="[919,963,223,249]" count="1" pageId="12" pageNumber="163" type="male">♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="45A0FD7E911DF112A489F9F179927EBF" box="[973,1015,223,249]" count="1" pageId="12" pageNumber="163" type="female">♀♀</specimenCount>
|
||
(196426531, 196426558, 196426592, 196426638),
|
||
<collectingCounty id="BA784E7B911DF112A66AF82D784C7F5B" box="[302,553,259,285]" pageId="12" pageNumber="163">Magaramkent District</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="B37DAC8D911DF112A571F82D79707F5B" box="[565,789,259,285]" pageId="12" pageNumber="163">Samur Delta Forest</collectingMunicipality>
|
||
,
|
||
<locationDeviation id="42AB2FD5911DF112A466F82D7E527F5B" box="[802,1079,259,285]" location="Primorskiy" pageId="12" pageNumber="163">2 km NW of Primorskiy</locationDeviation>
|
||
village, environs of
|
||
<location id="5679602C911DF112A261F82D7B1C7F07" LSID="urn:lsid:plazi:treatment:DB0F87E19119F10EA7D3FF1F7F207CEF:5679602C911DF112A261F82D7B1C7F07" county="Magaramkent District" latitude="48.5576" longLatPrecision="6" longitude="48.5592" municipality="Samur Delta Forest" name="Samurskiy Zakaznik Stationary" pageId="12" pageNumber="163" stateProvince="Dagestan">Samurskiy Zakaznik Stationary</location>
|
||
, a swampy overgrown estuary of a forest rivulet,
|
||
<geoCoordinate id="36925030911DF112A4D0F809798A7F07" box="[916,1007,295,321]" degrees="41.8619" direction="north" orientation="latitude" pageId="12" pageNumber="163" precision="5" value="41.8619">41.8619</geoCoordinate>
|
||
–41.8630 N, 48.5576–
|
||
<geoCoordinate id="36925030911DF112A3A9F8087F0D7F07" box="[1261,1384,294,321]" degrees="48.5592" direction="east" orientation="longitude" pageId="12" pageNumber="163" precision="5" value="48.5592">48.5592 E</geoCoordinate>
|
||
,
|
||
<quantity id="945E9B12911DF112A236F8097AB47F23" metricMagnitude="1" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="12" pageNumber="163" unit="m" value="17.0" valueMax="18.0" valueMin="16.0">
|
||
<elevation id="D88BD1C4911DF112A236F8097AB47F23" metricMagnitude="1" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="12" pageNumber="163" unit="m" value="17.0" valueMax="18.0" valueMin="16.0">16– 18 m</elevation>
|
||
</quantity>
|
||
below s.l.,
|
||
<date id="27181037911DF112A611F8657BB27F23" box="[341,471,331,357]" pageId="12" pageNumber="163" value="2021-06-24">
|
||
<collectingDate id="375CE9DF911DF112A611F8657BB27F23" box="[341,471,331,357]" pageId="12" pageNumber="163" value="2021-06-24">24.06.2021</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
</emphasis>
|
||
;
|
||
<specimenCount id="45A0FD7E911DF112A6A7F865786A7F23" box="[483,527,331,357]" count="1" pageId="12" pageNumber="163" type="male">♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="45A0FD7E911DF112A558F86578237F23" box="[540,582,331,357]" count="1" pageId="12" pageNumber="163" type="female">♀♀</specimenCount>
|
||
(196425929, 196425961, 196426001, 196426083, 196426117, 196426145, 196426182 (
|
||
<figureCitation id="CB9D2A72911DF112A669F8417BE47FCF" box="[301,385,367,393]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="12" pageNumber="163">Fig. 6j</figureCitation>
|
||
), 196426298, 196426322, 196432235, 196432236, 196432237, 196432239, 196432240, 196432241, 196432243, 196432244, 196432245), the same place,
|
||
<date id="27181037911DF112A4CDF8BD7E6E7FEB" box="[905,1035,403,429]" pageId="12" pageNumber="163" value="2021-06-26">26.06.2021</date>
|
||
;
|
||
<specimenCount id="45A0FD7E911DF112A353F8BD7E267FEB" box="[1047,1091,403,429]" count="1" pageId="12" pageNumber="163" type="male">♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="45A0FD7E911DF112A30BF8BD7E1C7FEB" box="[1103,1145,403,429]" count="1" pageId="12" pageNumber="163" type="female">♀♀</specimenCount>
|
||
(196431432, 196431467, 196431500, 196431519, 196431595, 196431732, 196431756, 196431799 (
|
||
<figureCitation id="CB9D2A72911DF112A4AFF8997E587F97" box="[1003,1085,439,465]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="12" pageNumber="163">Fig. 6e</figureCitation>
|
||
)), the same place,
|
||
<date id="27181037911DF112A252F8987FFD7F97" box="[1302,1432,438,465]" pageId="12" pageNumber="163" value="2023-05-28">28.05.2023</date>
|
||
;
|
||
<materialsCitation id="E3CE3CAA911DF112A7D3F8F57E6E7C5F" ID-GBIF-Occurrence="4897992317" collectingDate="2021-06-26" elevation="29" latitude="41.8668" location="Samur Delta Forest" longLatPrecision="41" longitude="48.558" municipality="Magaramkent District" pageId="12" pageNumber="163" specimenCount="1" stateProvince="Primorskiy">
|
||
copula (196432247,
|
||
<figureCitation id="CB9D2A72911DF112A6C0F8F57BBC7FB3" box="[388,473,475,501]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="12" pageNumber="163">Fig. 6k</figureCitation>
|
||
)),
|
||
<collectingMunicipality id="B37DAC8D911DF112A6BFF8F578927FB3" box="[507,759,475,501]" pageId="12" pageNumber="163">Magaramkent District</collectingMunicipality>
|
||
,
|
||
<location id="5679602C911DF112A441F8F5798D7FB3" LSID="urn:lsid:plazi:treatment:DB0F87E19119F10EA7D3FF1F7F207CEF:5679602C911DF112A441F8F5798D7FB3" box="[773,1000,475,501]" latitude="41.8668" longLatPrecision="41" longitude="48.558" municipality="Magaramkent District" name="Samur Delta Forest" pageId="12" pageNumber="163" stateProvince="Primorskiy">Samur Delta Forest</location>
|
||
,
|
||
<locationDeviation id="42AB2FD5911DF112A4B2F8F57F777FB3" box="[1014,1298,475,501]" location="Primorskiy" pageId="12" pageNumber="163">
|
||
<quantity id="945E9B12911DF112A4B2F8F57E567FB3" box="[1014,1075,475,501]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="12" pageNumber="163" unit="km" value="3.0">3 km</quantity>
|
||
NW of
|
||
<collectingRegion id="9162F815911DF112A3D2F8F57F777FB3" box="[1174,1298,475,501]" country="Russia" name="Primorskiy" pageId="12" pageNumber="163">Primorskiy</collectingRegion>
|
||
</locationDeviation>
|
||
village, the SE bank of a coastal lake,
|
||
<geoCoordinate id="36925030911DF112A680F8D178207C5F" box="[452,581,511,537]" degrees="41.8668" direction="north" orientation="latitude" pageId="12" pageNumber="163" precision="5" value="41.8668">41.8668 N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="36925030911DF112A515F8D078DA7C5F" box="[593,703,510,537]" degrees="48.558" direction="east" orientation="longitude" pageId="12" pageNumber="163" precision="55" value="48.558">48.558 E</geoCoordinate>
|
||
,
|
||
<quantity id="945E9B12911DF112A588F8D179637C5F" box="[716,774,511,537]" metricMagnitude="1" metricUnit="m" metricValue="2.9" pageId="12" pageNumber="163" unit="m" value="29.0">
|
||
<elevation id="D88BD1C4911DF112A588F8D179637C5F" box="[716,774,511,537]" metricMagnitude="1" metricUnit="m" metricValue="2.9" pageId="12" pageNumber="163" unit="m" value="29.0">29 m</elevation>
|
||
</quantity>
|
||
below s.l.,
|
||
<date id="27181037911DF112A4CDF8D17E6E7C5F" box="[905,1035,511,537]" pageId="12" pageNumber="163" value="2021-06-26">
|
||
<collectingDate id="375CE9DF911DF112A4CDF8D17E6E7C5F" box="[905,1035,511,537]" pageId="12" pageNumber="163" value="2021-06-26">26.06.2021</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="E3CE3CAA911DF112A353F8D179177CEF" ID-GBIF-Occurrence="4897992336" collectingDate="2021-06-26" elevation="20" location="Samur Delta Forest" municipality="Magaramkent District" pageId="12" pageNumber="163" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Primorskiy">
|
||
<specimenCount id="45A0FD7E911DF112A353F8D17E267C5F" box="[1047,1091,511,537]" count="1" pageId="12" pageNumber="163" type="male">♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="45A0FD7E911DF112A30BF8D17E1C7C5F" box="[1103,1145,511,537]" count="1" pageId="12" pageNumber="163" type="female">♀♀</specimenCount>
|
||
(196427626, 196428813, 196428891, 196428956, 196428980, 196429008, 196429029, 196429061, 196429094, 196429172, 196429235, 196429278, 196429336, 196429394, 196429463, 196429541, 196429578, 196429724, 196429748),
|
||
<collectingMunicipality id="B37DAC8D911DF112A245FB697A8B7CC3" pageId="12" pageNumber="163">Magaramkent District</collectingMunicipality>
|
||
,
|
||
<location id="5679602C911DF112A7BEFB457BB27CC3" LSID="urn:lsid:plazi:treatment:DB0F87E19119F10EA7D3FF1F7F207CEF:5679602C911DF112A7BEFB457BB27CC3" box="[250,471,619,645]" municipality="Magaramkent District" name="Samur Delta Forest" pageId="12" pageNumber="163" stateProvince="Primorskiy">Samur Delta Forest</location>
|
||
,
|
||
<locationDeviation id="42AB2FD5911DF112A6A7FB44796D7CC3" box="[483,776,618,645]" location="Primorskiy" pageId="12" pageNumber="163">
|
||
<quantity id="945E9B12911DF112A6A7FB4478547CC3" box="[483,561,618,645]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="12" pageNumber="163" unit="km" value="3.5">3.5 km</quantity>
|
||
NW of
|
||
<collectingRegion id="9162F815911DF112A5C8FB45796D7CC3" box="[652,776,619,645]" country="Russia" name="Primorskiy" pageId="12" pageNumber="163">Primorskiy</collectingRegion>
|
||
</locationDeviation>
|
||
village, bushy meadows at the NW bank of a coastal lake, 41.8709–8755 N, 48.5409–5459 E,
|
||
<quantity id="945E9B12911DF112A573FBA178157CEF" box="[567,624,655,681]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="163" unit="m" value="20.0">
|
||
<elevation id="D88BD1C4911DF112A573FBA178157CEF" box="[567,624,655,681]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="163" unit="m" value="20.0">20 m</elevation>
|
||
</quantity>
|
||
below s.l.,
|
||
<date id="27181037911DF112A5B4FBA179177CEF" box="[752,882,655,681]" pageId="12" pageNumber="163" value="2021-06-26">
|
||
<collectingDate id="375CE9DF911DF112A5B4FBA179177CEF" box="[752,882,655,681]" pageId="12" pageNumber="163" value="2021-06-26">26.06.2021</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="531936F7911DF112A783FB9C79017DAB" blockId="12.[151,1437,150,2013]" pageId="12" pageNumber="163">
|
||
<emphasis id="61D2EAE5911DF112A783FB9C7B317C8A" bold="true" box="[199,340,690,716]" pageId="12" pageNumber="163">Characters.</emphasis>
|
||
Male cercus trunk in dorsal view (
|
||
<figureCitation id="CB9D2A72911DF112A595FB9D795F7C8B" box="[721,826,691,717]" captionStart="FIGURE 2" captionStartId="9.[151,250,1421,1445]" captionTargetBox="[381,1200,187,1391]" captionTargetId="figure-19@9.[381,1206,181,1397]" captionTargetPageId="9" captionText="FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189910" httpUri="https://zenodo.org/record/12189910/files/figure.png" pageId="12" pageNumber="163">Fig. 2a–c</figureCitation>
|
||
) evenly thick, with a very gentle convection in place of bulge (so their bases do not touch each other), rather long. In lateral view (
|
||
<figureCitation id="CB9D2A72911DF112A48FFBF97E517CB7" box="[971,1076,727,753]" captionStart="FIGURE3" captionStartId="10.[151,250,1432,1456]" captionTargetBox="[393,1180,188,1398]" captionTargetId="figure-24@10.[393,1194,181,1408]" captionTargetPageId="10" captionText="FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189912" httpUri="https://zenodo.org/record/12189912/files/figure.png" pageId="12" pageNumber="163">Fig. 3a–c</figureCitation>
|
||
) it is smoothly rounded, without an expressed apex. Tubercle in dorsal view (
|
||
<figureCitation id="CB9D2A72911DF112A5CBFBD5789F7D53" box="[655,762,763,789]" captionStart="FIGURE 2" captionStartId="9.[151,250,1421,1445]" captionTargetBox="[381,1200,187,1391]" captionTargetId="figure-19@9.[381,1206,181,1397]" captionTargetPageId="9" captionText="FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189910" httpUri="https://zenodo.org/record/12189910/files/figure.png" pageId="12" pageNumber="163">Fig. 2a–c</figureCitation>
|
||
) of the same width as trunk and looks like its continuation, rather elongate, in lateral view (
|
||
<figureCitation id="CB9D2A72911DF112A6B1FA31783B7D7F" box="[501,606,799,825]" captionStart="FIGURE3" captionStartId="10.[151,250,1432,1456]" captionTargetBox="[393,1180,188,1398]" captionTargetId="figure-24@10.[393,1194,181,1408]" captionTargetPageId="10" captionText="FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189912" httpUri="https://zenodo.org/record/12189912/files/figure.png" pageId="12" pageNumber="163">Fig. 3a–c</figureCitation>
|
||
) always well seen, never hidden behind paraprocts. (It should be noted that the specimen shown in
|
||
<figureCitation id="CB9D2A72911DF112A6E4FA6D7B837D1B" box="[416,486,835,861]" captionStart="FIGURE 1" captionStartId="7.[151,250,790,814]" captionTargetBox="[201,1386,187,756]" captionTargetId="figure-19@7.[201,1386,179,764]" captionTargetPageId="7" captionText="FIGURE 1. Explanation of terminology (see the text) used for morphological details of male terminalia of Coenagrion spp. of the puella-group in the broad sense used in this paper, on examples of a male specimen of C. australocaspicum with an unusual expanded position of the cerci in lateral view (left, the same as Fig. 3a) and a male of C. persicum in dorsal view (right, the same as Fig. 2n)." figureDoi="http://doi.org/10.5281/zenodo.12189908" httpUri="https://zenodo.org/record/12189908/files/figure.png" pageId="12" pageNumber="163">Figs 1</figureCitation>
|
||
(left), 2b and 3b has dead with abnormally expanded cerci, while
|
||
<figureCitation id="CB9D2A72911DF112A395FA6D7F527D1B" box="[1233,1335,835,861]" captionStart="FIGURE 2" captionStartId="9.[151,250,1421,1445]" captionTargetBox="[381,1200,187,1391]" captionTargetId="figure-19@9.[381,1206,181,1397]" captionTargetPageId="9" captionText="FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189910" httpUri="https://zenodo.org/record/12189910/files/figure.png" pageId="12" pageNumber="163">Figs 2a,c</figureCitation>
|
||
and
|
||
<figureCitation id="CB9D2A72911DF112A22AFA6D7FF97D1B" box="[1390,1436,835,861]" captionStart="FIGURE3" captionStartId="10.[151,250,1432,1456]" captionTargetBox="[393,1180,188,1398]" captionTargetId="figure-24@10.[393,1194,181,1408]" captionTargetPageId="10" captionText="FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189912" httpUri="https://zenodo.org/record/12189912/files/figure.png" pageId="12" pageNumber="163">3a,c</figureCitation>
|
||
show their normal downward position. Nevertheless, the above described characters are well seen in
|
||
<figureCitation id="CB9D2A72911DF112A3BCFA497F2E7DC7" box="[1272,1355,871,897]" captionStart="FIGURE 2" captionStartId="9.[151,250,1421,1445]" captionTargetBox="[381,1200,187,1391]" captionTargetId="figure-19@9.[381,1206,181,1397]" captionTargetPageId="9" captionText="FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189910" httpUri="https://zenodo.org/record/12189910/files/figure.png" pageId="12" pageNumber="163">Figs 2b</figureCitation>
|
||
and
|
||
<figureCitation id="CB9D2A72911DF112A2C4FA497FF97DC7" box="[1408,1436,871,897]" captionStart="FIGURE3" captionStartId="10.[151,250,1432,1456]" captionTargetBox="[393,1180,188,1398]" captionTargetId="figure-24@10.[393,1194,181,1408]" captionTargetPageId="10" captionText="FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189912" httpUri="https://zenodo.org/record/12189912/files/figure.png" pageId="12" pageNumber="163">3b</figureCitation>
|
||
as well.) Tooth small, normally not seen in dorsal view unless cerci are unusually expanded as in
|
||
<figureCitation id="CB9D2A72911DF112A398FAA57F4A7DE3" box="[1244,1327,907,933]" captionStart="FIGURE 2" captionStartId="9.[151,250,1421,1445]" captionTargetBox="[381,1200,187,1391]" captionTargetId="figure-19@9.[381,1206,181,1397]" captionTargetPageId="9" captionText="FIGURE 2. Male appendages of Coenagrion spp. of the puella-group in the broad sense, from the Caucasus and elsewhere, in dorsal view: a–c—C. australocaspicum (b—with the appendages expanded); d-k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–p—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; j—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m— Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189910" httpUri="https://zenodo.org/record/12189910/files/figure.png" pageId="12" pageNumber="163">Fig. 2b</figureCitation>
|
||
, disposed at ca 3/4 of the trunk length. Process broadly triangular, somewhat hooked inside apically but rather blunt, reaching about the middle of the height of the paraproct base (
|
||
<figureCitation id="CB9D2A72911DF112A5ACFAFD793D7DAB" box="[744,856,979,1005]" captionStart="FIGURE 4" captionStartId="11.[151,250,1054,1078]" captionTargetBox="[189,1398,181,1021]" captionTargetId="figure-19@11.[189,1398,181,1030]" captionTargetPageId="11" captionText="FIGURE 4. Male appendages of Coenagrion spp. of the puella-group in the broad sense in posterior view: a–b—C. australocaspicum (b—with the appendages expanded); c–d—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); e-f—C. pulchellum; g–k—C. puella; l—C. persicum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c–d—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; e—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; f—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; g–h—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; i—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, a brook and pond, 25.06.2021; j—Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189914" httpUri="https://zenodo.org/record/12189914/files/figure.png" pageId="12" pageNumber="163">Fig. 4a–b</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="531936F7911DF112A783FAD97E237A3B" blockId="12.[151,1437,150,2013]" pageId="12" pageNumber="163">
|
||
Paraproct spine very long, directed strictly caudad in lateral view (
|
||
<figureCitation id="CB9D2A72911DF112A4E9FAD97E727A57" box="[941,1047,1015,1041]" captionStart="FIGURE3" captionStartId="10.[151,250,1432,1456]" captionTargetBox="[393,1180,188,1398]" captionTargetId="figure-24@10.[393,1194,181,1408]" captionTargetPageId="10" captionText="FIGURE3. Maleappendages of Coenagrion spp.ofthe puella-groupin thebroadsense inlateral view:a–c—C.australocaspicum (b—with the appendages expanded); d–k—C. puella; l–m—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); n—C. persicum; o–q—C. pulchellum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c—Russia, Dagestan, Kumtorkala District, the Shura-Ozen’ River at Kumtorkala Station, next to Sarykum Barkhan, 3.05.2021; d —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; e–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g–h—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; i—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; k—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l–m—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; n—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; o—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; p—Russia, N Caucasus, Karachay-Cherkes Republic, Zelenchuk District, Dausuz village W environs, S banks of a reservoir at the Bolzhoy Zelenchuk River, 1012 m a.s.l.; q—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189912" httpUri="https://zenodo.org/record/12189912/files/figure.png" pageId="12" pageNumber="163">Fig. 3a–c</figureCitation>
|
||
) where is ca 5/6 as long as S10 (as measured along the line of its dorsal side); spines diverging and slightly hooked in at tips dorsal view. Prominence weakly expressed as resembling a rounded right angle, incision between spine and prominense not expressed. S
|
||
<quantity id="945E9B12911DF112A2C4FD117AC87A3B" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="12" pageNumber="163" unit="in" value="10.0">10 in</quantity>
|
||
caudal view looks more flattened above (
|
||
<figureCitation id="CB9D2A72911DF112A5C5FD4D78887A3B" box="[641,749,1123,1149]" captionStart="FIGURE 4" captionStartId="11.[151,250,1054,1078]" captionTargetBox="[189,1398,181,1021]" captionTargetId="figure-19@11.[189,1398,181,1030]" captionTargetPageId="11" captionText="FIGURE 4. Male appendages of Coenagrion spp. of the puella-group in the broad sense in posterior view: a–b—C. australocaspicum (b—with the appendages expanded); c–d—C. ponticum (males with a more (c) and less (d) expanded black abdominal maculation); e-f—C. pulchellum; g–k—C. puella; l—C. persicum; a–b—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; c–d—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; e—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; f—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; g–h—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; i—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, a brook and pond, 25.06.2021; j—Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; k—Russia, Republic of Bashkortostan, Zinachurinskiy District, a small roadside oxbow lake 0.7 km E of Verkhniy Sarabil’ village, 5.07.2021; l—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189914" httpUri="https://zenodo.org/record/12189914/files/figure.png" pageId="12" pageNumber="163">Fig. 4a–b</figureCitation>
|
||
) as compared to other species.
|
||
</paragraph>
|
||
<paragraph id="531936F7911DF112A783FDA97BD07A83" blockId="12.[151,1437,150,2013]" pageId="12" pageNumber="163">
|
||
Male pronotum hind margin almost straight, with negligible, very shallow lateral depressions and a very slight median incision (
|
||
<figureCitation id="CB9D2A72911DF112A61DFD857BCD7A83" box="[345,424,1194,1221]" captionStart="FIGURE 5" captionStartId="13.[151,250,1570,1594]" captionTargetBox="[151,1436,181,1541]" captionTargetId="figure-19@13.[151,1436,181,1546]" captionTargetPageId="13" captionText="FIGURE 5. Male pronotum of Coenagrion spp. of the puella-group in the broad sense in dorsal view: a—C. persicum; b–c—C. pulchellum; d–e—C. ponticum (males with a more (d) and less (e) expanded black abdominal maculation); f—C. australocaspicum; g–l—C. puella; a—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; b—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; c—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; d–e—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; f—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; g —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; h–i—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021; j—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; k—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; l—Russia, Karachay- Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189916" httpUri="https://zenodo.org/record/12189916/files/figure.png" pageId="12" pageNumber="163">Fig. 5f</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="531936F7911DF112A783FDE178CE7B77" blockId="12.[151,1437,150,2013]" pageId="12" pageNumber="163">
|
||
Male abdomen is somewhat relatively longer than in
|
||
<taxonomicName id="94A64D74911DF112A456FDE179127AAF" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[786,887,1231,1257]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="puella">
|
||
<emphasis id="61D2EAE5911DF112A456FDE179127AAF" box="[786,887,1231,1257]" italics="true" pageId="12" pageNumber="163">C. puella</emphasis>
|
||
</taxonomicName>
|
||
, coloured as in that species but S9 blue with (
|
||
<figureCitation id="CB9D2A72911DF112A22BFDE17ABF7B4B" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="12" pageNumber="163">Fig. 6a, 6c</figureCitation>
|
||
) or without (
|
||
<figureCitation id="CB9D2A72911DF112A62DFDDD7BDC7B4B" box="[361,441,1267,1293]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="12" pageNumber="163">Fig. 6b</figureCitation>
|
||
) a pair of black dots or (rarely) black triangular marks (
|
||
<figureCitation id="CB9D2A72911DF112A359FDDD7E0B7B4B" box="[1053,1134,1267,1293]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="12" pageNumber="163">Fig. 6c</figureCitation>
|
||
), but never with a posterior black bar. S10 black dorsally and blue laterally.
|
||
</paragraph>
|
||
<paragraph id="531936F7911DF112A783FC15797B7BDB" blockId="12.[151,1437,150,2013]" pageId="12" pageNumber="163">
|
||
Female pronotum hind lobe margin almost straight when seen flat, perpendicularly to its surface, that is anteriodorsally with respect to the insect body axis, (
|
||
<figureCitation id="CB9D2A72911DF112A599FC7179487B3F" box="[733,813,1375,1401]" captionStart="FIGURE 7" captionStartId="15.[151,250,1565,1589]" captionTargetBox="[447,1140,184,1541]" captionTargetId="figure-19@15.[447,1140,181,1541]" captionTargetPageId="15" captionText="FIGURE 7. Female pronotum of Coenagrion spp. of the puella-group in the broad sense, a–h—in dorsal, with respect to the body main axis (above), and anteriodorsal, perpendicular to the posterior lobe surface (below), views; i–j—in dorsal view only; a—C. australocaspicum; b–g—C. puella; h—C. ponticum; i—C. persicum; j—C. pulchellum; a—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; b—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; c—Sergokala District, 3 km SW of Myurego village, the Inchkheozen’ River upper valley, 19.06.2021; d–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; h—Russia, Chelyabinsk Province, Nagaybakskiy District, small oxbow lakes at the Gumbeyka River left bank at Ostrolenskiy Settlement SE margin, 6.07.2021; i—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; j—Iran, Markazi Ostan, Shazand Shakhrestan, the Garehchay River at Gavigodar village, 34.1078-1111° N 49.3627-3657° E, 1803–1805 m a.s.l., 19.05.2017; k—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189924" httpUri="https://zenodo.org/record/12189924/files/figure.png" pageId="12" pageNumber="163">Fig. 7a</figureCitation>
|
||
, below), but is slightly wavy as can be seen in the dorsal view with respect to the insect body axis (
|
||
<figureCitation id="CB9D2A72911DF112A529FCAD78DA7BDB" box="[621,703,1411,1437]" captionStart="FIGURE 7" captionStartId="15.[151,250,1565,1589]" captionTargetBox="[447,1140,184,1541]" captionTargetId="figure-19@15.[447,1140,181,1541]" captionTargetPageId="15" captionText="FIGURE 7. Female pronotum of Coenagrion spp. of the puella-group in the broad sense, a–h—in dorsal, with respect to the body main axis (above), and anteriodorsal, perpendicular to the posterior lobe surface (below), views; i–j—in dorsal view only; a—C. australocaspicum; b–g—C. puella; h—C. ponticum; i—C. persicum; j—C. pulchellum; a—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; b—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; c—Sergokala District, 3 km SW of Myurego village, the Inchkheozen’ River upper valley, 19.06.2021; d–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; h—Russia, Chelyabinsk Province, Nagaybakskiy District, small oxbow lakes at the Gumbeyka River left bank at Ostrolenskiy Settlement SE margin, 6.07.2021; i—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; j—Iran, Markazi Ostan, Shazand Shakhrestan, the Garehchay River at Gavigodar village, 34.1078-1111° N 49.3627-3657° E, 1803–1805 m a.s.l., 19.05.2017; k—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12189924" httpUri="https://zenodo.org/record/12189924/files/figure.png" pageId="12" pageNumber="163">Fig. 7a</figureCitation>
|
||
, above).
|
||
</paragraph>
|
||
<paragraph id="531936F7911DF112A783FC897F3F7937" blockId="12.[151,1437,150,2013]" pageId="12" pageNumber="163">
|
||
As usual in the genus, females are polymorphic for the coloration pattern. Most frequent is the dark morph which resembles the corresponding morph in
|
||
<taxonomicName id="94A64D74911DF112A508FCE578D67BA3" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[588,691,1483,1509]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="puella">
|
||
<emphasis id="61D2EAE5911DF112A508FCE578D67BA3" box="[588,691,1483,1509]" italics="true" pageId="12" pageNumber="163">C. puella</emphasis>
|
||
</taxonomicName>
|
||
, with most of the abdomen dorsum black, with only expression of pale rings at S8–10 varying to complete disappearance (
|
||
<figureCitation id="CB9D2A72911DF112A44EFCC1791C784F" box="[778,889,1519,1545]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="12" pageNumber="163">Fig. 6d–e</figureCitation>
|
||
). Less frequent is a spotty morph, also paralleled by the analogous morph of
|
||
<taxonomicName id="94A64D74911DF112A68DFF3D784A786B" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[457,559,1555,1581]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="puella">
|
||
<emphasis id="61D2EAE5911DF112A68DFF3D784A786B" box="[457,559,1555,1581]" italics="true" pageId="12" pageNumber="163">C. puella</emphasis>
|
||
</taxonomicName>
|
||
, with anterior parts of most abdominal segments pale (
|
||
<figureCitation id="CB9D2A72911DF112A3D1FF3D7F64786B" box="[1173,1281,1555,1581]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="12" pageNumber="163">Fig. 6f–h</figureCitation>
|
||
). However, in these morphs, the pale ground colour occupies a larger area on S8 than in
|
||
<taxonomicName id="94A64D74911DF112A49FFF197E277817" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[987,1090,1591,1617]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="puella">
|
||
<emphasis id="61D2EAE5911DF112A49FFF197E277817" box="[987,1090,1591,1617]" italics="true" pageId="12" pageNumber="163">C. puella</emphasis>
|
||
</taxonomicName>
|
||
, varying from a large bipartite anterior pale spots (
|
||
<figureCitation id="CB9D2A72911DF112A6C4FF757B827833" box="[384,487,1627,1653]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="12" pageNumber="163">Fig. 6f–j</figureCitation>
|
||
) to, rather infrequently, S8 being pale with a black streak (
|
||
<figureCitation id="CB9D2A72911DF112A3E0FF757E9E7833" box="[1188,1275,1627,1653]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="12" pageNumber="163">Fig. 6h</figureCitation>
|
||
), often of the horseshoe shape—this version resembles females of rather
|
||
<taxonomicName id="94A64D74911DF112A47FFF5179BC78DF" baseAuthorityName="Vander Linden" baseAuthorityYear="1825" box="[827,985,1663,1689]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="pulchellum">
|
||
<emphasis id="61D2EAE5911DF112A47FFF5179BC78DF" box="[827,985,1663,1689]" italics="true" pageId="12" pageNumber="163">C. pulchellum</emphasis>
|
||
</taxonomicName>
|
||
than
|
||
<taxonomicName id="94A64D74911DF112A35FFF517EE678DF" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1051,1155,1663,1689]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="163" phylum="Arthropoda" rank="species" species="puella">
|
||
<emphasis id="61D2EAE5911DF112A35FFF517EE678DF" box="[1051,1155,1663,1689]" italics="true" pageId="12" pageNumber="163">C. puella</emphasis>
|
||
</taxonomicName>
|
||
. The extreme version of the spotty morph are extra-pale females the maculation of which rather resembles males of
|
||
<emphasis id="61D2EAE5911DF112A3CBFF8D7FF978FB" box="[1167,1436,1699,1725]" italics="true" pageId="12" pageNumber="163">Enallahma cyathigerum</emphasis>
|
||
(
|
||
<bibRefCitation id="37374B06911DF112A7DBFFE97B0878A7" author="Charpentier, T." box="[159,365,1735,1761]" pageId="12" pageNumber="163" refId="ref27675" refString="Charpentier, T. (de) (1840) Libellulinae europaeae descripta ac depictae. L. Voss, Lipsiae, 181 pp. https: // doi. org / 10.5962 / t. 173037" type="book" year="1840">Charpentier, 1840</bibRefCitation>
|
||
): with a black arrowhead mark on S2, black spots in the posterior parts of S2–7, producing one to three processes in the anterior direction, a paired black mark on S8 and S9–10 black at dorsum (
|
||
<figureCitation id="CB9D2A72911DF112A3F8FFC57F6D7943" box="[1212,1288,1771,1797]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="12" pageNumber="163">Fig. 6i</figureCitation>
|
||
). The ground pale colour varies (
|
||
<figureCitation id="CB9D2A72911DF112A634FE217BBE796F" box="[368,475,1807,1833]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="12" pageNumber="163">Fig. 6d–i</figureCitation>
|
||
), at least in part with age, from pale greyish (
|
||
<figureCitation id="CB9D2A72911DF112A49BFE217E57796F" box="[991,1074,1807,1833]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="12" pageNumber="163">Fig. 6h</figureCitation>
|
||
), sometime pale orange, to blue (
|
||
<figureCitation id="CB9D2A72911DF112A7DBFE1D7B69790B" box="[159,268,1843,1869]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="12" pageNumber="163">Fig. 6e, g</figureCitation>
|
||
), often this is grey but with the anteriodorsal part of the abdominal segments blue (
|
||
<figureCitation id="CB9D2A72911DF112A3F5FE1D7E9A790B" box="[1201,1279,1843,1869]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="12" pageNumber="163">Fig. 6i</figureCitation>
|
||
). However, in mature blue females, the antehumeral stripes are usually green, as well as the eye lower part (
|
||
<figureCitation id="CB9D2A72911DF112A3E9FE797F2B7937" box="[1197,1358,1879,1905]" captionStart="FIGURE 6" captionStartId="14.[151,250,1745,1769]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-24@14.[151,1436,181,1721]" captionTargetPageId="14" captionText="FIGURE 6. Males (a–c), females (d–i) and copulae (j–k) of Coenagrion australocaspicum in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="12" pageNumber="163">Fig. 6e, g, j–k</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1BBC657C911DF10EA783FE547F207CEF" lastPageId="16" lastPageNumber="167" pageId="12" pageNumber="163" type="distribution">
|
||
<paragraph id="531936F7911DF10EA783FE547EC27F23" blockId="12.[151,1437,150,2013]" lastBlockId="16.[151,1437,151,681]" lastPageId="16" lastPageNumber="167" pageId="12" pageNumber="163">
|
||
<emphasis id="61D2EAE5911DF112A783FE547B9C79D2" bold="true" box="[199,505,1914,1941]" pageId="12" pageNumber="163">Distribution and habitat.</emphasis>
|
||
Found in the Russian Caucasus only in
|
||
<collectingRegion id="9162F815911DF112A49EFE557E2079D3" box="[986,1093,1915,1941]" country="Russia" name="Dagestan" pageId="12" pageNumber="163">Dagestan</collectingRegion>
|
||
, in the coastal plains of both southern (Samur Delta Forest) and northern (Khamamatyurt)
|
||
<collectingRegion id="9162F815911DF112A47BFEB179C279FF" box="[831,935,1951,1977]" country="Russia" name="Dagestan" pageId="12" pageNumber="163">Dagestan</collectingRegion>
|
||
and in the foothills near the famous Sarykum Barkhan (the Shura-Ozen’ River). However, the Khamamatyurt locality is situated at the right bank of the Terek River which serves there as the border with Chechen Republic, where the species certainly occurs as well. The latter is the northermost known locality of this species in the world, of
|
||
<geoCoordinate id="369250309101F10EA43BF99579AB7E93" box="[895,974,187,213]" degrees="43.6" direction="north" orientation="latitude" pageId="16" pageNumber="167" precision="5555" value="43.6">43.6 N</geoCoordinate>
|
||
. Beyond
|
||
<collectingCountry id="2BB176679101F10EA37EF9957EED7E93" box="[1082,1160,187,213]" name="Russia" pageId="16" pageNumber="167">Russia</collectingCountry>
|
||
, the species is known to occur along the southern coast of the Caspian Sea in
|
||
<collectingCountry id="2BB176679101F10EA5B8F9F1794C7EBF" box="[764,809,223,249]" name="Iran" pageId="16" pageNumber="167">Iran</collectingCountry>
|
||
and in the coastal Lankoran and Kachmaz Districts of
|
||
<collectingCountry id="2BB176679101F10EA7D3F82D7B777F5B" box="[151,274,259,285]" name="Azerbaijan" pageId="16" pageNumber="167">Azerbaijan</collectingCountry>
|
||
(
|
||
<bibRefCitation id="37374B069101F10EA665F82D78597F5B" author="Dumont, H. J. & Heidari, H." box="[289,572,259,285]" pageId="16" pageNumber="167" pagination="63 - 78" refId="ref27800" refString="Dumont, H. J. & Heidari, H. (1996) On a collection of spring Odonata from Iran, with the description of Coenagrion australocaspicum n. sp. Bulletin et Annales de la Societe Royale Belge d'Entomologie, 132, 63 - 78." type="journal article" year="1996">Dumont & Heidari 1996</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="37374B069101F10EA50DF82D788C7F5B" author="Dumont, H. J." box="[585,745,259,285]" pageId="16" pageNumber="167" pagination="87 - 92" refId="ref27765" refString="Dumont, H. J. (2004) Dragonflies from Azerbaijan. Zoology in the Middle East, 31, 87 - 92. https: // doi. org / 10.1080 / 09397140.2004.10638027" type="journal article" year="2004">Dumont 2004</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="37374B069101F10EA5B1F82D79AE7F5B" author="Boudot, J. - P. & Borisov, S. & De Knijf, G. & Van Grunsven, R. & Schroter, A. & Kalkman, V. J." box="[757,971,259,285]" pageId="16" pageNumber="167" pagination="3 - 248" refId="ref27573" refString="Boudot, J. - P., Borisov, S., De Knijf, G., Van Grunsven, R., Schroter, A. & Kalkman, V. J. (2021) Atlas of the dragonflies and damselflies of West and Central Asia. Brachytron, 22, Supplement, 3 - 248." type="journal article" year="2021">
|
||
Boudot
|
||
<emphasis id="61D2EAE59101F10EA40BF82D79EC7F5B" box="[847,905,259,285]" italics="true" pageId="16" pageNumber="167">et al.</emphasis>
|
||
2021
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="37374B069101F10EA49CF82D7EFD7F5B" author="Snegovaya, N. Y." box="[984,1176,259,285]" pageId="16" pageNumber="167" pagination="1 - 33" refId="ref29587" refString="Snegovaya, N. Y. (2024) Odonata collected in 2023 in Azerbaijan, including the confirmation of Cordulia aenea Linnaeus, 1758 for Azerbaijan. International Dragonfly Fund Report, 186, 1 - 33." type="journal article" year="2024">Snegovaya 2024</bibRefCitation>
|
||
). In the latter district it was near Şollar (
|
||
<bibRefCitation id="37374B069101F10EA612F8097B937F07" author="Dumont, H. J." box="[342,502,295,321]" pageId="16" pageNumber="167" pagination="87 - 92" refId="ref27765" refString="Dumont, H. J. (2004) Dragonflies from Azerbaijan. Zoology in the Middle East, 31, 87 - 92. https: // doi. org / 10.1080 / 09397140.2004.10638027" type="journal article" year="2004">Dumont 2004</bibRefCitation>
|
||
) and Narban (
|
||
<bibRefCitation id="37374B069101F10EA5DDF809793D7F07" author="Snegovaya, N. Y." box="[665,856,295,321]" pageId="16" pageNumber="167" pagination="1 - 33" refId="ref29587" refString="Snegovaya, N. Y. (2024) Odonata collected in 2023 in Azerbaijan, including the confirmation of Cordulia aenea Linnaeus, 1758 for Azerbaijan. International Dragonfly Fund Report, 186, 1 - 33." type="journal article" year="2024">Snegovaya 2024</bibRefCitation>
|
||
) villages, the latter being situated near the Russian border, just
|
||
<quantity id="945E9B129101F10EA659F8657B017F23" box="[285,356,331,357]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="16" pageNumber="167" unit="km" value="12.0">12 km</quantity>
|
||
south-east from our closest findings of this species in Samur Delta Forest.
|
||
</paragraph>
|
||
<caption id="07D9667F911CF113A7D3FF0C7B9B79AF" ID-DOI="http://doi.org/10.5281/zenodo.12189916" ID-Zenodo-Dep="12189916" httpUri="https://zenodo.org/record/12189916/files/figure.png" pageId="13" pageNumber="164" startId="13.[151,250,1570,1594]" targetBox="[151,1436,181,1541]" targetPageId="13" targetType="figure">
|
||
<paragraph id="531936F7911CF113A7D3FF0C7B9B79AF" blockId="13.[151,1437,1570,2026]" pageId="13" pageNumber="164">
|
||
<emphasis id="61D2EAE5911CF113A7D3FF0C7B72787C" bold="true" box="[151,279,1570,1594]" pageId="13" pageNumber="164">FIGURE 5.</emphasis>
|
||
Male pronotum of
|
||
<taxonomicName id="94A64D74911CF113A6A9FF0C78FE787F" box="[493,667,1570,1593]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="61D2EAE5911CF113A6A9FF0C7803787F" box="[493,614,1570,1593]" italics="true" pageId="13" pageNumber="164">Coenagrion</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
of the
|
||
<taxonomicName id="94A64D74911CF113A5AAFF0C7948787C" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[750,813,1570,1594]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="puella">
|
||
<emphasis id="61D2EAE5911CF113A5AAFF0C7948787C" box="[750,813,1570,1594]" italics="true" pageId="13" pageNumber="164">puella</emphasis>
|
||
</taxonomicName>
|
||
-group in the broad sense in dorsal view: a—
|
||
<taxonomicName id="94A64D74911CF113A253FF0C7FF0787C" authorityName="Lohmann" authorityYear="1983" box="[1303,1429,1570,1594]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="persicum">
|
||
<emphasis id="61D2EAE5911CF113A253FF0C7FF0787C" box="[1303,1429,1570,1594]" italics="true" pageId="13" pageNumber="164">C. persicum</emphasis>
|
||
</taxonomicName>
|
||
; b–c—
|
||
<taxonomicName id="94A64D74911CF113A791FF687B027818" baseAuthorityName="Vander Linden" baseAuthorityYear="1825" box="[213,359,1606,1630]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="pulchellum">
|
||
<emphasis id="61D2EAE5911CF113A791FF687B027818" box="[213,359,1606,1630]" italics="true" pageId="13" pageNumber="164">C. pulchellum</emphasis>
|
||
</taxonomicName>
|
||
; d–e—
|
||
<taxonomicName id="94A64D74911CF113A6F0FF6878567818" baseAuthorityName="Bartenef" baseAuthorityYear="1929" box="[436,563,1606,1630]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="ponticum">
|
||
<emphasis id="61D2EAE5911CF113A6F0FF6878567818" box="[436,563,1606,1630]" italics="true" pageId="13" pageNumber="164">C. ponticum</emphasis>
|
||
</taxonomicName>
|
||
(males with a more (d) and less (e) expanded black abdominal maculation); f—
|
||
<taxonomicName id="94A64D74911CF113A2C1FF687B2E78C4" authorityName="Dumont & Heidari" authorityYear="1996" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="australocaspicum">
|
||
<emphasis id="61D2EAE5911CF113A2C1FF687B2E78C4" italics="true" pageId="13" pageNumber="164">C. australocaspicum</emphasis>
|
||
</taxonomicName>
|
||
; g–l—
|
||
<taxonomicName id="94A64D74911CF113A6D0FF447B9778C4" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[404,498,1642,1666]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="puella">
|
||
<emphasis id="61D2EAE5911CF113A6D0FF447B9778C4" box="[404,498,1642,1666]" italics="true" pageId="13" pageNumber="164">C. puella</emphasis>
|
||
</taxonomicName>
|
||
; a—Iran, Markazi Ostan, Shazand Shakhrestan, Emarat village env., a spring and swampy reservoir, 20.05.2017; b—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021; c—Russia, Dagestan, Makhachkala N suburbs, pools between dunes at the Krivaya Balka River left bank in its lowermost reaches, 23.05.2023; d–e—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; f—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; g —Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; h–i—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021; j—Russia, Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021; k—Russia, Dagestan, Buynaksk District, Buynaksk S environs, Nizhnee Kazanishche village environs, a water reservoir, 2020; l—Russia, KarachayCherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019. Scale bar 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="07D9667F911FF110A7D3FFFF7ECA793F" ID-DOI="http://doi.org/10.5281/zenodo.12189920" ID-Zenodo-Dep="12189920" httpUri="https://zenodo.org/record/12189920/files/figure.png" pageId="14" pageNumber="165" startId="14.[151,250,1745,1769]" targetBox="[151,1436,181,1721]" targetPageId="14" targetType="figure">
|
||
<paragraph id="531936F7911FF110A7D3FFFF7ECA793F" blockId="14.[151,1437,1745,1913]" pageId="14" pageNumber="165">
|
||
<emphasis id="61D2EAE5911FF110A7D3FFFF7B7778AF" bold="true" box="[151,274,1745,1769]" pageId="14" pageNumber="165">FIGURE 6.</emphasis>
|
||
Males (a–c), females (d–i) and copulae (j–k) of
|
||
<taxonomicName id="94A64D74911FF110A5BFFFFC7E4B78AF" authorityName="Dumont & Heidari" authorityYear="1996" box="[763,1070,1745,1769]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="165" phylum="Arthropoda" rank="species" species="australocaspicum">
|
||
<emphasis id="61D2EAE5911FF110A5BFFFFC7E4B78AF" box="[763,1070,1745,1769]" italics="true" pageId="14" pageNumber="165">Coenagrion australocaspicum</emphasis>
|
||
</taxonomicName>
|
||
in nature in the Samur Delta Forest, 2-4 km NW of Primorskiy village, Magaramkent District of Dagestan, Russia; a—27.05.2023 (196433323); b—26.05.2023 (196432917); c—26.05.2023 (196503424); d—27.05.2023 (196430622); e—28.05.2023 (196431799); f—26.05.2023 (196432921); g—25.05.2021 (196425479); h—26.05.2023 (196427547); i—27.05.2023 (196430322); j—26.06.2021 (196426182); k—26.06.2021 (196432247); a–c, f, k—photo by OEK, d–e, h–j—by VVO. Not to scale.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="07D9667F911EF111A7D3FF33794979A3" ID-DOI="http://doi.org/10.5281/zenodo.12189924" ID-Zenodo-Dep="12189924" httpUri="https://zenodo.org/record/12189924/files/figure.png" pageId="15" pageNumber="166" startId="15.[151,250,1565,1589]" targetBox="[447,1140,184,1541]" targetPageId="15" targetType="figure">
|
||
<paragraph id="531936F7911EF111A7D3FF33794979A3" blockId="15.[151,1437,1565,2022]" pageId="15" pageNumber="166">
|
||
<emphasis id="61D2EAE5911EF111A7D3FF337B717873" bold="true" box="[151,276,1565,1589]" pageId="15" pageNumber="166">FIGURE 7.</emphasis>
|
||
Female pronotum of
|
||
<taxonomicName id="94A64D74911EF111A6B0FF3378FA7873" box="[500,671,1565,1589]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="61D2EAE5911EF111A6B0FF3378087872" box="[500,621,1565,1588]" italics="true" pageId="15" pageNumber="166">Coenagrion</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
of the
|
||
<taxonomicName id="94A64D74911EF111A5ACFF3379427873" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[744,807,1565,1589]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="puella">
|
||
<emphasis id="61D2EAE5911EF111A5ACFF3379427873" box="[744,807,1565,1589]" italics="true" pageId="15" pageNumber="166">puella</emphasis>
|
||
</taxonomicName>
|
||
-group in the broad sense, a–h—in dorsal, with respect to the body main axis (above), and anteriodorsal, perpendicular to the posterior lobe surface (below), views; i–j—in dorsal view only; a—
|
||
<taxonomicName id="94A64D74911EF111A7BEFF4B7BB5783B" authorityName="Dumont & Heidari" authorityYear="1996" box="[250,464,1637,1661]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="australocaspicum">
|
||
<emphasis id="61D2EAE5911EF111A7BEFF4B7BB5783B" box="[250,464,1637,1661]" italics="true" pageId="15" pageNumber="166">C. australocaspicum</emphasis>
|
||
</taxonomicName>
|
||
; b–g—
|
||
<taxonomicName id="94A64D74911EF111A565FF4B78E4783B" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[545,641,1637,1661]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="puella">
|
||
<emphasis id="61D2EAE5911EF111A565FF4B78E4783B" box="[545,641,1637,1661]" italics="true" pageId="15" pageNumber="166">C. puella</emphasis>
|
||
</taxonomicName>
|
||
; h—
|
||
<taxonomicName id="94A64D74911EF111A5FDFF4B795C783B" baseAuthorityName="Bartenef" baseAuthorityYear="1929" box="[697,825,1637,1661]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="ponticum">
|
||
<emphasis id="61D2EAE5911EF111A5FDFF4B795C783B" box="[697,825,1637,1661]" italics="true" pageId="15" pageNumber="166">C. ponticum</emphasis>
|
||
</taxonomicName>
|
||
; i—
|
||
<taxonomicName id="94A64D74911EF111A42FFF4B798C783B" authorityName="Lohmann" authorityYear="1983" box="[875,1001,1637,1661]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="persicum">
|
||
<emphasis id="61D2EAE5911EF111A42FFF4B798C783B" box="[875,1001,1637,1661]" italics="true" pageId="15" pageNumber="166">C. persicum</emphasis>
|
||
</taxonomicName>
|
||
; j—
|
||
<taxonomicName id="94A64D74911EF111A35FFF4B7ECB783B" baseAuthorityName="Vander Linden" baseAuthorityYear="1825" box="[1051,1198,1637,1661]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="pulchellum">
|
||
<emphasis id="61D2EAE5911EF111A35FFF4B7ECB783B" box="[1051,1198,1637,1661]" italics="true" pageId="15" pageNumber="166">C. pulchellum</emphasis>
|
||
</taxonomicName>
|
||
; a—Russia, Dagestan, Magaramkent District, Samur Delta Forest, 2 km NW of Primorskiy village, a swampy estuary of a forest rivulet, 24.06.2021; b—Georgia, Dzau District, 4 km NNE Kvaisa, Lake Koz (Kvedi), 28−30.06.2021; c—Sergokala District, 3 km SW of Myurego village, the Inchkheozen’ River upper valley, 19.06.2021; d–f—Russia, Dagestan, Rutul District; the Samur River left arm between Kufa and Kala villages, 26.06.2021; g—Russia, Karachay-Cherkes Republic, Zelenchuk District, 10 km WSW of Arkhyz, Lunnaya Polyana terrain, pools in the Dukka River valley left side, 2.07.2019; h—Russia, Chelyabinsk Province, Nagaybakskiy District, small oxbow lakes at the Gumbeyka River left bank at Ostrolenskiy Settlement SE margin, 6.07.2021; i—Russia, Krasnodarskiy Kray, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 27.04.2019; j—Iran, Markazi Ostan, Shazand Shakhrestan, the Garehchay River at Gavigodar village, 34.1078-1111° N 49.3627-3657° E, 1803–1805 m a.s.l., 19.05.2017; k—Russia, Dagestan, Makhachkala N env., Muraveynik Protected Area at the Sulak River mouth, pools among coastal dunes, 1.06.2021. Scale bar 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="531936F79101F10EA783F8417F207CEF" blockId="16.[151,1437,151,681]" pageId="16" pageNumber="167">
|
||
These damselflies were invariably found near slow flowing overgrown brooks. They often concentrate at forest roads and margins (and also around an old plane tree surrounded by a meadow in dozen metres apart from water). A very abundant population was found in the Samur Delta Forest, composed of tall oak and hornbeam trees woven with many lianas, which grow, below the conventional sea level at the Caspian Sea coast, in the semidesert zone because the delta of the large Samur River forms a number of fast and cold arms locally called ‘Karasu’. In that forest, this species seems to nearly segregate with the visually very similar
|
||
<taxonomicName id="94A64D749101F10EA4BFFB0D7E017C7B" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1019,1124,547,573]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="167" phylum="Arthropoda" rank="species" species="puella">
|
||
<emphasis id="61D2EAE59101F10EA4BFFB0D7E017C7B" box="[1019,1124,547,573]" italics="true" pageId="16" pageNumber="167">C. puella</emphasis>
|
||
</taxonomicName>
|
||
, with just
|
||
<specimenCount id="45A0FD7E9101F10EA3A4FB0D7F197C7B" box="[1248,1404,547,573]" count="1" pageId="16" pageNumber="167" type="generic">one specimen</specimenCount>
|
||
of
|
||
<taxonomicName id="94A64D749101F10EA7D3FB697A9B7C27" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[151,254,583,609]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="167" phylum="Arthropoda" rank="species" species="puella">
|
||
<emphasis id="61D2EAE59101F10EA7D3FB697A9B7C27" box="[151,254,583,609]" italics="true" pageId="16" pageNumber="167">C. puella</emphasis>
|
||
</taxonomicName>
|
||
found (on
|
||
<date id="271810379101F10EA63FFB687B9E7C27" box="[379,507,582,609]" pageId="16" pageNumber="167" value="2023-05-26">26.05.2023</date>
|
||
) among dozens of
|
||
<taxonomicName id="94A64D749101F10EA595FB6979DC7C27" authorityName="Dumont & Heidari" authorityYear="1996" box="[721,953,583,609]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="167" phylum="Arthropoda" rank="species" species="australocaspicum">
|
||
<emphasis id="61D2EAE59101F10EA595FB6979DC7C27" box="[721,953,583,609]" italics="true" pageId="16" pageNumber="167">C. australocaspicum</emphasis>
|
||
</taxonomicName>
|
||
at the Samurskiy Zakaznik Stationary, and none among numerous representatives of
|
||
<taxonomicName id="94A64D749101F10EA537FB4579397CC3" authorityName="Dumont & Heidari" authorityYear="1996" box="[627,860,619,645]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="167" phylum="Arthropoda" rank="species" species="australocaspicum">
|
||
<emphasis id="61D2EAE59101F10EA537FB4579397CC3" box="[627,860,619,645]" italics="true" pageId="16" pageNumber="167">C. australocaspicum</emphasis>
|
||
</taxonomicName>
|
||
at the famous old plane tree, while only
|
||
<taxonomicName id="94A64D749101F10EA270FB457FF97CC3" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1332,1436,619,645]" class="Insecta" family="Coenagrionidae" genus="Coenagrion" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="167" phylum="Arthropoda" rank="species" species="puella">
|
||
<emphasis id="61D2EAE59101F10EA270FB457FF97CC3" box="[1332,1436,619,645]" italics="true" pageId="16" pageNumber="167">C. puella</emphasis>
|
||
</taxonomicName>
|
||
was present at a brook in Bil’-Bil’ Kazmalyar village just
|
||
<quantity id="945E9B129101F10EA45BFBA0793D7CEF" box="[799,856,654,681]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="16" pageNumber="167" unit="km" value="5.0">5 km</quantity>
|
||
SSE of the Samurskiy Zakaznik Stationary.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |