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<document id="9352A4EDDD02222E4E1C55A9D3AD747D" ID-CLB-Dataset="30866" ID-DOI="10.11646/zootaxa.4377.2.3" ID-GBIF-Dataset="5cb64c13-76bb-45f3-9a12-c2413c64c537" ID-ISSN="1175-5326" ID-Zenodo-Dep="1163893" ID-ZooBank="B1EFF160-FD26-4A18-A274-4E605EEEEE2F" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1517471380876" checkinUser="plazi" docAuthor="Liu, Yubin, Hutchings, Pat &amp; Kupriyanova, Elena" docDate="2018" docId="BF203D7B315D385CDEE34CDB4788E74B" docLanguage="en" docName="zootaxa.4372.2.3.pdf" docOrigin="Zootaxa 4377 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Marphysa Quatrefages 1865" docType="treatment" docVersion="7" lastPageNumber="194" masterDocId="43194503315F385FDE744B724356E246" masterDocTitle="Two new species of Marphysa Quatrefages, 1865 (Polychaeta: Eunicida: Eunicidae) from northern coast of China and redescription for Marphysa orientalis Treadwell, 1936" masterLastPageNumber="215" masterPageNumber="191" pageNumber="193" updateTime="1698518448595" updateUser="ExternalLinkService">
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<mods:title id="292CCC42814FC76C934E8B1C55B6DDA3">Two new species of Marphysa Quatrefages, 1865 (Polychaeta: Eunicida: Eunicidae) from northern coast of China and redescription for Marphysa orientalis Treadwell, 1936</mods:title>
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<subSubSection id="7F93DFE6315D385DDEE34CDB418EE585" box="[151,728,1961,1987]" pageId="2" pageNumber="193" type="nomenclature">
<paragraph id="37368C6D315D385DDEE34CDB418EE585" blockId="2.[151,728,1961,1987]" box="[151,728,1961,1987]" pageId="2" pageNumber="193">
<heading id="6C7E3B01315D385DDEE34CDB418EE585" bold="true" box="[151,728,1961,1987]" fontSize="11" level="1" pageId="2" pageNumber="193" reason="1">
<emphasis id="05FD507F315D385DDEE34CDB418EE585" bold="true" box="[151,728,1961,1987]" pageId="2" pageNumber="193">
Genus
<taxonomicName id="F089F7EE315D385DDE9A4CDB4104E585" ID-CoL="84Q23" authority="Quatrefages, 1865" authorityName="Quatrefages" authorityYear="1865" box="[238,594,1961,1987]" class="Polychaeta" family="Eunicidae" genus="Marphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eunicida" pageId="2" pageNumber="193" phylum="Annelida" rank="genus">
<emphasis id="05FD507F315D385DDE9A4CDB423EE585" bold="true" box="[238,360,1961,1987]" italics="true" pageId="2" pageNumber="193">Marphysa</emphasis>
Quatrefages, 1865
</taxonomicName>
, restricted
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7F93DFE6315C385CDEE34BE54070E05E" pageId="3" pageNumber="194" type="diagnosis">
<paragraph id="37368C6D315C385CDEE34BE5407AE2F6" blockId="3.[151,1437,151,1293]" box="[151,812,151,176]" pageId="3" pageNumber="194">
<emphasis id="05FD507F315C385CDEE34BE5425CE2F6" bold="true" box="[151,266,151,176]" pageId="3" pageNumber="194">Diagnosis</emphasis>
. (after
<bibRefCitation id="5318F19C315C385CDF284BE54076E2F6" author="Molina-Acevedo, I. C. &amp; Carrera-Parra, L. F." box="[348,800,151,176]" journalOrPublisher="Zootaxa" pageId="3" pageNumber="194" pagination="1 - 62" part="4241" publicationUrl="https://doi.org/10.11646/zootaxa.4241.1.1" refString="Molina-Acevedo, I. C. &amp; Carrera-Parra, L. F. (2017) Revision of Marphysa de Quatrefages, 1865 and some species of Nicidion Kinberg, 1865 with the erection of a new genus (Polychaeta: Euncidae) from the Grand Caribbean. Zootaxa, 4241 (1), 1 - 62. https: // doi. org / 10.11646 / zootaxa. 4241.1.1" title="Revision of Marphysa de Quatrefages, 1865 and some species of Nicidion Kinberg, 1865 with the erection of a new genus (Polychaeta: Euncidae) from the Grand Caribbean" type="journal article" year="2017">Molina-Acevedo &amp; Carrera-Parra, 2017</bibRefCitation>
).
</paragraph>
<paragraph id="37368C6D315C385CDEB34BCE4070E05E" blockId="3.[151,1437,151,1293]" pageId="3" pageNumber="194">Prostomium slightly bilobed or completely bilobed; five prostomial appendages without articulations; eyes present or absent. Peristomium without peristomial cirri. Maxillary apparatus with four pairs of maxillae and an unpaired on left side; MI with falcal arch developed, extended, and with the outer edge of the base arched; MIII curved, forming part of distal arc, with attachment lamella of rectangular or irregular shape, situated at the centre of posterior edge of maxilla; MIV with circular or rectangular attachment lamella. Branchiae distributed throughout the body. Dorsal cirri without articulation; postchaetal lobe well developed in anterior region, ventral cirri with swollen base, oval or circular. Aciculae dark or translucent. Supracicular chaetae include limbate; pectinate isodont chaetae with slender teeth, and pectinate anodont chaetae with long teeth present or absent. Subacicular chaetae include compound falcigers or spinigers or both. Subacicular hook, dark or translucent, bidentate or unidentate. Pygidium with two pairs of anal cirri, without articulation.</paragraph>
</subSubSection>
<subSubSection id="7F93DFE6315C385CDEB349564788E74B" pageId="3" pageNumber="194" type="discussion">
<paragraph id="37368C6D315C385CDEB34956460BE153" blockId="3.[151,1437,151,1293]" pageId="3" pageNumber="194">
<emphasis id="05FD507F315C385CDEB349564262E07B" bold="true" box="[199,308,548,573]" pageId="3" pageNumber="194">Remarks</emphasis>
.
<taxonomicName id="F089F7EE315C385CDF32495642EEE07B" authorityName="Quatrefages" authorityYear="1865" box="[326,440,548,573]" class="Polychaeta" family="Eunicidae" genus="Marphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eunicida" pageId="3" pageNumber="194" phylum="Annelida" rank="genus">
<emphasis id="05FD507F315C385CDF32495642EEE07B" box="[326,440,548,573]" italics="true" pageId="3" pageNumber="194">Marphysa</emphasis>
</taxonomicName>
is a heterogeneous genus due to the vast variation of morphological features and its definition has been controversial.
<bibRefCitation id="5318F19C315C385CDC67493541B3E026" author="Zanol, J. &amp; Halanych, K. M. &amp; Fauchald, K." box="[531,741,583,608]" journalOrPublisher="Zoologica Scripta" pageId="3" pageNumber="194" pagination="79 - 100" part="43" publicationUrl="https://doi.org/10.1111/zsc.12034" refString="Zanol, J., Halanych, K. M. &amp; Fauchald, K. (2014) Reconciling taxonomy and phylogeny in the bristleworm family Eunicidae (polychaete, Annelida). Zoologica Scripta, 43, 79 - 100. https: // doi. org / 10.1111 / zsc. 12034" title="Reconciling taxonomy and phylogeny in the bristleworm family Eunicidae (polychaete, Annelida)" type="journal article" year="2014">
Zanol
<emphasis id="05FD507F315C385CDC2F493B41DAE026" box="[603,652,583,608]" italics="true" pageId="3" pageNumber="194">et al</emphasis>
. (2014)
</bibRefCitation>
proposed a phylogenetic hypothesis of
<taxonomicName id="F089F7EE315C385CDAD549354647E026" authorityName="Berthold" authorityYear="1827" box="[1185,1297,583,608]" class="Polychaeta" family="Eunicidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eunicida" pageId="3" pageNumber="194" phylum="Annelida" rank="family">Eunicidae</taxonomicName>
based on an analysis of morphological and molecular characters, and established two new genera which included species previously assigned to
<taxonomicName id="F089F7EE315C385CDFD149FD4141E0EE" authorityName="Quatrefages" authorityYear="1865" box="[421,535,655,680]" class="Polychaeta" family="Eunicidae" genus="Marphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eunicida" pageId="3" pageNumber="194" phylum="Annelida" rank="genus">
<emphasis id="05FD507F315C385CDFD149FD4141E0EE" box="[421,535,655,680]" italics="true" pageId="3" pageNumber="194">Marphysa</emphasis>
</taxonomicName>
. They suggest that characters previously used to delineate genera within the family, such as the number of prostomial appendages and the presence or absence of peristomial cirri are not informative. However, until a complete revision of the genus
<taxonomicName id="F089F7EE315C385CDD3F49A540EBE0B6" authorityName="Quatrefages" authorityYear="1865" box="[843,957,727,752]" class="Polychaeta" family="Eunicidae" genus="Marphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eunicida" pageId="3" pageNumber="194" phylum="Annelida" rank="genus">
<emphasis id="05FD507F315C385CDD3F49A540EBE0B6" box="[843,957,727,752]" italics="true" pageId="3" pageNumber="194">Marphysa</emphasis>
</taxonomicName>
is undertaken, we are following the above definition of
<bibRefCitation id="5318F19C315C385CDF5D498E41A1E152" author="Molina-Acevedo, I. C. &amp; Carrera-Parra, L. F." box="[297,759,764,789]" journalOrPublisher="Zootaxa" pageId="3" pageNumber="194" pagination="1 - 62" part="4241" publicationUrl="https://doi.org/10.11646/zootaxa.4241.1.1" refString="Molina-Acevedo, I. C. &amp; Carrera-Parra, L. F. (2017) Revision of Marphysa de Quatrefages, 1865 and some species of Nicidion Kinberg, 1865 with the erection of a new genus (Polychaeta: Euncidae) from the Grand Caribbean. Zootaxa, 4241 (1), 1 - 62. https: // doi. org / 10.11646 / zootaxa. 4241.1.1" title="Revision of Marphysa de Quatrefages, 1865 and some species of Nicidion Kinberg, 1865 with the erection of a new genus (Polychaeta: Euncidae) from the Grand Caribbean" type="journal article" year="2017">Molina-Acevedo &amp; Carrera-Parra (2017)</bibRefCitation>
, who acknowledge that the genus is not monophyletic.
</paragraph>
<paragraph id="37368C6D315C385CDEB3486D4788E74B" blockId="3.[151,1437,151,1293]" pageId="3" pageNumber="194">
<bibRefCitation id="5318F19C315C385CDEB3486D42D2E17E" author="Fauchald, K." box="[199,388,799,824]" journalOrPublisher="Allan Hancock Monograph in Marine Biology, 5" pageId="3" pageNumber="194" pagination="335" part="1" refString="Fauchald, K. (1970) Polychaetous annelids of the families Eunicidae, Lumbrineridae, Iphitimidae, Arabellidae, Lysaretidae and Dorvilleidae from western Mexico. Allan Hancock Monograph in Marine Biology, 5, 1 ¯ 335." title="Polychaetous annelids of the families Eunicidae, Lumbrineridae, Iphitimidae, Arabellidae, Lysaretidae and Dorvilleidae from western Mexico" type="journal article" year="1970">Fauchald (1970)</bibRefCitation>
reviewed the 48 species of
<taxonomicName id="F089F7EE315C385CDCAE486D401AE17E" authorityName="Quatrefages" authorityYear="1865" box="[730,844,799,824]" class="Polychaeta" family="Eunicidae" genus="Marphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eunicida" pageId="3" pageNumber="194" phylum="Annelida" rank="genus">
<emphasis id="05FD507F315C385CDCAE486D401AE17E" box="[730,844,799,824]" italics="true" pageId="3" pageNumber="194">Marphysa</emphasis>
</taxonomicName>
accepted at the time and arranged them into five artificial groups (referred to as groups
<emphasis id="05FD507F315C385CDC374836412EE11B" bold="true" box="[579,632,836,861]" pageId="3" pageNumber="194">AE</emphasis>
) based on the type of compound chaeta present, since then an additional 18 species have been described. According to the grouping, group
<emphasis id="05FD507F315C385CDDF9481540F7E1C6" bold="true" box="[909,929,871,896]" pageId="3" pageNumber="194">A</emphasis>
includes species lacking compound chaetae, species in
<emphasis id="05FD507F315C385CDF6648FE4273E1E3" bold="true" box="[274,293,908,933]" pageId="3" pageNumber="194">B</emphasis>
have compound spinigers only, species in
<emphasis id="05FD507F315C385CDD6848FE4066E1E3" bold="true" box="[796,816,908,933]" pageId="3" pageNumber="194">C</emphasis>
have compound falcigers only, those in
<emphasis id="05FD507F315C385CDB7848FE4676E1E3" bold="true" box="[1292,1312,908,933]" pageId="3" pageNumber="194">D</emphasis>
have both compound falcigers and spinigers, and
<emphasis id="05FD507F315C385CDC1548DD4122E18E" bold="true" box="[609,628,943,968]" pageId="3" pageNumber="194">E</emphasis>
unites species too poorly known with the respect of their compound chaetae to be categorised.
<bibRefCitation id="5318F19C315C385CDFBE48A641D0E1AA" author="Fauchald, K." box="[458,646,980,1005]" journalOrPublisher="Allan Hancock Monograph in Marine Biology, 5" pageId="3" pageNumber="194" pagination="335" part="1" refString="Fauchald, K. (1970) Polychaetous annelids of the families Eunicidae, Lumbrineridae, Iphitimidae, Arabellidae, Lysaretidae and Dorvilleidae from western Mexico. Allan Hancock Monograph in Marine Biology, 5, 1 ¯ 335." title="Polychaetous annelids of the families Eunicidae, Lumbrineridae, Iphitimidae, Arabellidae, Lysaretidae and Dorvilleidae from western Mexico" type="journal article" year="1970">Fauchald (1970)</bibRefCitation>
further subdivides species in groups
<emphasis id="05FD507F315C385CDA4D48A64738E1AB" bold="true" box="[1081,1134,980,1005]" pageId="3" pageNumber="194">AE</emphasis>
into those with branchiae restricted to a short anterior region only (1) and those with branchiae present throughout most of the body length (2). This classification was followed by
<bibRefCitation id="5318F19C315C385CDC1F4F6E40FAE672" author="Glasby, C. J. &amp; Hutchings, P. A." box="[619,940,1052,1077]" journalOrPublisher="Zootaxa" pageId="3" pageNumber="194" pagination="29 - 45" part="2352" refString="Glasby, C. J. &amp; Hutchings, P. A. (2010) A new species of Marphysa Quatrefages, 1865 (Polychaeta: Eunicida: Eunicidae) from northern Australia and a review of similar taxa from the Indo-West Pacific, including the genus Nauphanta Kinberg, 1865. Zootaxa, 2352, 29 - 45." title="A new species of Marphysa Quatrefages, 1865 (Polychaeta: Eunicida: Eunicidae) from northern Australia and a review of similar taxa from the Indo-West Pacific, including the genus Nauphanta Kinberg, 1865" type="journal article" year="2010">Glasby &amp; Hutchings (2010)</bibRefCitation>
and
<bibRefCitation id="5318F19C315C385CDD9C4F6E4790E673" author="Zanol, J. &amp; da Silva, T." box="[1000,1222,1052,1077]" journalOrPublisher="Invertebrate Biology, 135" pageId="3" pageNumber="194" pagination="328 - 344" part="4" publicationUrl="https://doi.org/10.1111/ivb.12146" refString="Zanol, J., da Silva, T. dos S. C. &amp; Hutchings, P. (2016) Integrative taxonomy of Marphysa (Eunicidae, Polychaeta, Annelida) species of the S anguinea-group from Australia. Invertebrate Biology, 135 (4), 328 - 344. https: // doi. org / 10.1111 / ivb. 12146" title="Integrative taxonomy of Marphysa (Eunicidae, Polychaeta, Annelida) species of the S anguinea-group from Australia" type="journal article" year="2016">
Zanol
<emphasis id="05FD507F315C385CDA474F6F4730E673" box="[1075,1126,1052,1077]" italics="true" pageId="3" pageNumber="194">et al</emphasis>
. (2016)
</bibRefCitation>
. Some support for these morphological groups was obtained during a molecular study of the
<taxonomicName id="F089F7EE315C385CDDAC4F4D471EE61E" authorityName="Berthold" authorityYear="1827" box="[984,1096,1087,1112]" class="Polychaeta" family="Eunicidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eunicida" pageId="3" pageNumber="194" phylum="Annelida" rank="family">Eunicidae</taxonomicName>
family by
<bibRefCitation id="5318F19C315C385CDAB64F4D46C1E61E" author="Zanol, J. &amp; Halanych, K. M. &amp; Fauchald, K." box="[1218,1431,1087,1112]" journalOrPublisher="Zoologica Scripta" pageId="3" pageNumber="194" pagination="79 - 100" part="43" publicationUrl="https://doi.org/10.1111/zsc.12034" refString="Zanol, J., Halanych, K. M. &amp; Fauchald, K. (2014) Reconciling taxonomy and phylogeny in the bristleworm family Eunicidae (polychaete, Annelida). Zoologica Scripta, 43, 79 - 100. https: // doi. org / 10.1111 / zsc. 12034" title="Reconciling taxonomy and phylogeny in the bristleworm family Eunicidae (polychaete, Annelida)" type="journal article" year="2014">
Zanol
<emphasis id="05FD507F315C385CDB7F4F334615E61E" box="[1291,1347,1087,1112]" italics="true" pageId="3" pageNumber="194">et al.</emphasis>
(2014)
</bibRefCitation>
, but they sequenced only 12 species of
<taxonomicName id="F089F7EE315C385CDC3C4F1641ECE63B" authorityName="Quatrefages" authorityYear="1865" box="[584,698,1124,1149]" class="Polychaeta" family="Eunicidae" genus="Marphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eunicida" pageId="3" pageNumber="194" phylum="Annelida" rank="genus">
<emphasis id="05FD507F315C385CDC3C4F1641ECE63B" box="[584,698,1124,1149]" italics="true" pageId="3" pageNumber="194">Marphysa</emphasis>
</taxonomicName>
and found two basal groups in
<emphasis id="05FD507F315C385CDA6F4F164799E63A" box="[1051,1231,1124,1149]" italics="true" pageId="3" pageNumber="194">
<taxonomicName id="F089F7EE315C385CDA6F4F1647DAE63B" authority="Liu &amp; Hutchings &amp; Kupriyanova, 2018" authorityName="Liu &amp; Hutchings &amp; Kupriyanova" authorityYear="2018" box="[1051,1164,1124,1149]" class="Polychaeta" family="Eunicidae" genus="Marphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eunicida" pageId="3" pageNumber="194" phylum="Annelida" rank="genus" sensu="stricto">Marphysa</taxonomicName>
<taxonomicNameLabel id="1ECEED04315C385CDAE04F174799E63A" box="[1172,1231,1125,1148]" pageId="3" pageNumber="194" sensu="stricto">s. str.</taxonomicNameLabel>
</emphasis>
which represent a “
<taxonomicName id="F089F7EE315C385CDED74FF542C3E6E6" baseAuthorityName="Montagu" baseAuthorityYear="1813" box="[163,405,1159,1184]" class="Polychaeta" family="Eunicidae" genus="Marphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eunicida" pageId="3" pageNumber="194" phylum="Annelida" rank="species" species="sanguinea">
<emphasis id="05FD507F315C385CDED74FF542C3E6E6" box="[163,405,1159,1184]" italics="true" pageId="3" pageNumber="194">Marphysa sanguinea</emphasis>
</taxonomicName>
-like” group
<emphasis id="05FD507F315C385CDC454FF54112E6E6" bold="true" box="[561,580,1159,1184]" pageId="3" pageNumber="194">B</emphasis>
and a “
<taxonomicName id="F089F7EE315C385CDCD94FF54008E6E6" box="[685,862,1159,1184]" class="Polychaeta" family="Eunicidae" genus="Marphysa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Eunicida" pageId="3" pageNumber="194" phylum="Annelida" rank="species" species="belli">
<emphasis id="05FD507F315C385CDCD94FF54008E6E6" box="[685,862,1159,1184]" italics="true" pageId="3" pageNumber="194">Marphysa belli</emphasis>
</taxonomicName>
-like”-group
<emphasis id="05FD507F315C385CDD814FF5475FE6E6" bold="true" box="[1013,1033,1159,1184]" pageId="3" pageNumber="194">D</emphasis>
.
<bibRefCitation id="5318F19C315C385CDA6A4FF54655E6E6" author="Zanol, J. &amp; Halanych, K. M. &amp; Fauchald, K." box="[1054,1283,1159,1184]" journalOrPublisher="Zoologica Scripta" pageId="3" pageNumber="194" pagination="79 - 100" part="43" publicationUrl="https://doi.org/10.1111/zsc.12034" refString="Zanol, J., Halanych, K. M. &amp; Fauchald, K. (2014) Reconciling taxonomy and phylogeny in the bristleworm family Eunicidae (polychaete, Annelida). Zoologica Scripta, 43, 79 - 100. https: // doi. org / 10.1111 / zsc. 12034" title="Reconciling taxonomy and phylogeny in the bristleworm family Eunicidae (polychaete, Annelida)" type="journal article" year="2014">
Zanol
<emphasis id="05FD507F315C385CDA194FFB47FAE6E6" box="[1133,1196,1159,1184]" italics="true" pageId="3" pageNumber="194">et al.</emphasis>
(2014)
</bibRefCitation>
provided an emended generic diagnosis, however
<bibRefCitation id="5318F19C315C385CDC494FDE475CE682" author="Molina-Acevedo, I. C. &amp; Carrera-Parra, L. F." box="[573,1034,1196,1221]" journalOrPublisher="Zootaxa" pageId="3" pageNumber="194" pagination="1 - 62" part="4241" publicationUrl="https://doi.org/10.11646/zootaxa.4241.1.1" refString="Molina-Acevedo, I. C. &amp; Carrera-Parra, L. F. (2017) Revision of Marphysa de Quatrefages, 1865 and some species of Nicidion Kinberg, 1865 with the erection of a new genus (Polychaeta: Euncidae) from the Grand Caribbean. Zootaxa, 4241 (1), 1 - 62. https: // doi. org / 10.11646 / zootaxa. 4241.1.1" title="Revision of Marphysa de Quatrefages, 1865 and some species of Nicidion Kinberg, 1865 with the erection of a new genus (Polychaeta: Euncidae) from the Grand Caribbean" type="journal article" year="2017">Molina-Acevedo &amp; Carrera-Parra (2017)</bibRefCitation>
added some further characters with regards to the maxillary apparatus and parapodial structures which we used here. We have followed Molina-
<bibRefCitation id="5318F19C315C385CDEE34F86415EE74B" author="Molina-Acevedo, I. C. &amp; Carrera-Parra, L. F." box="[151,520,1268,1293]" journalOrPublisher="Zootaxa" pageId="3" pageNumber="194" pagination="37 - 55" part="3925" publicationUrl="https://doi.org/10.11646/zootaxa.3925.1.3" refString="Molina-Acevedo, I. C. &amp; Carrera-Parra, L. F. (2015) Reinstatement of three Grand Caribbean species of the Marphysa sanguinea complex (Polychaeta: Eunicidae), Zootaxa, 3925 (1), 37 - 55. https: // doi. org / 10.11646 / zootaxa. 3925.1.3" title="Reinstatement of three Grand Caribbean species of the Marphysa sanguinea complex (Polychaeta: Eunicidae)" type="journal article" year="2015">Acevedo &amp; Carrera-Parra (2015)</bibRefCitation>
and
<bibRefCitation id="5318F19C315C385CDC4A4F864047E74B" author="Zanol, J. &amp; da Silva, T." box="[574,785,1268,1293]" journalOrPublisher="Invertebrate Biology, 135" pageId="3" pageNumber="194" pagination="328 - 344" part="4" publicationUrl="https://doi.org/10.1111/ivb.12146" refString="Zanol, J., da Silva, T. dos S. C. &amp; Hutchings, P. (2016) Integrative taxonomy of Marphysa (Eunicidae, Polychaeta, Annelida) species of the S anguinea-group from Australia. Invertebrate Biology, 135 (4), 328 - 344. https: // doi. org / 10.1111 / ivb. 12146" title="Integrative taxonomy of Marphysa (Eunicidae, Polychaeta, Annelida) species of the S anguinea-group from Australia" type="journal article" year="2016">
Zanol
<emphasis id="05FD507F315C385CDCF34F8741EEE74B" box="[647,696,1268,1293]" italics="true" pageId="3" pageNumber="194">et al</emphasis>
. (2016)
</bibRefCitation>
for terminology of the pectinate chaetae.
</paragraph>
</subSubSection>
</treatment>
</document>