treatments-xml/data/B9/70/A0/B970A055FFC3FFE6FE1E2605FDC1FBF7.xml
2024-06-21 12:49:17 +02:00

181 lines
18 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="6D98397F4BA5D91A821261B965ED61CB" ID-DOI="10.5281/zenodo.5330604" ID-ISSN="0374-1036" ID-Zenodo-Dep="5330604" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630168017471" checkinUser="felipe" docAuthor="Fikáček, Martin, Prokin, Alexander, Angus, Robert B., Pono, Alexander, Marenko, Yue, Yanli, Ren, Dong &amp; Prokop, Jakub" docDate="2012" docId="B970A055FFC3FFE6FE1E2605FDC1FBF7" docLanguage="en" docName="ActaEntMusNatPra.2012.52.1.89-127.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 52 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Hydrophilopsia Ponomarenko 1987" docType="treatment" docVersion="4" lastPageNumber="92" masterDocId="4549D82DFFC0FFE5FF9B2756FFB9FFFB" masterDocTitle="Revision of Mesozoic fossils of the helophorid lineage of the superfamily Hydrophiloidea (Coleoptera: Polyphaga)" masterLastPageNumber="127" masterPageNumber="89" pageNumber="92" updateTime="1699171640213" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="A49CD5909FB3D404AE2779099479FEF9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9C74881384C5D8DACA302F204B008441">
<mods:title id="25E021D29F6B22AB536DECB53D7D7F3C">Revision of Mesozoic fossils of the helophorid lineage of the superfamily Hydrophiloidea (Coleoptera: Polyphaga)</mods:title>
</mods:titleInfo>
<mods:name id="136356CDD510CE376164DB8258A15F69" type="personal">
<mods:role id="55E3C86D6FB0A381679496138B67574A">
<mods:roleTerm id="7405A2023C0202D755FE04D509C157D0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="70CB1B8CCC2FD79EBAA4C97688FBAA79">Fikáček, Martin</mods:namePart>
<mods:affiliation id="8B3202ED8043B949EF61E9F6A1CF18BD">Department of Entomology, National Museum, Kunratice 1, CZ- 148 00 Praha 4, Czech Republic; e-mail: mfikacek @ gmail. com &amp; Department of Zoology, Faculty of Science, Charles University in Prague, Viničná 7, CZ- 128 44 Praha 2, Czech Republic; e-mail: jprokop @ natur. cuni. cz</mods:affiliation>
</mods:name>
<mods:name id="FDBB800915053159B09D84DB573F01BB" type="personal">
<mods:role id="0A2DA47A40DD7354B2CE3882BCBB392F">
<mods:roleTerm id="2AF4FBD255EFB202EDCE8F7ED91B0D2E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="38EC3D94F7A0A3675E7334692F765A38">Prokin, Alexander</mods:namePart>
<mods:affiliation id="B186765DB123B346EFEFE7B6C9EE278A">Papanin Institute for Inland Water Biology, Russian Academy of Science, Borok, Russia; e-mail: prokina @ mail. ru &amp; Voronezh State University, Research-Educational Centre “ Venevitinovo ”, Voronezh, Russia</mods:affiliation>
</mods:name>
<mods:name id="0E7C79F3A1BEDA2A17DD324C3DDEDFCA" type="personal">
<mods:role id="4D9E341A82ED0D347119980FE41ADC7D">
<mods:roleTerm id="5C6A7A770398C13B4AB09D966B22A4D9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="185D55E8D2F109EBB18E4CABFC7EF2E3">Angus, Robert B.</mods:namePart>
<mods:affiliation id="FA5155FEBCC07406F4BF0364699B4BEE">Department of Entomology, The Natural History Museum, London, UK; e-mail: r. angus @ rhul. ac. uk</mods:affiliation>
</mods:name>
<mods:name id="50476598E829D7E3055741E737E9BBE8" type="personal">
<mods:role id="1E6EC3C3A13307C95C6FDB29EF130D13">
<mods:roleTerm id="5D4B386214996BE042B36542BF80A807">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5051583D0CBD53B36E770619728A222A">Pono, Alexander</mods:namePart>
</mods:name>
<mods:name id="1D57B0730DAD038CEE5CC18C991462F5" type="personal">
<mods:role id="A2A0FD88259E4A2F3FE83E57186532AA">
<mods:roleTerm id="C686266E6656B9AC42A28F558875F769">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="20E6904D435874527196EEE852185FB4">Marenko</mods:namePart>
<mods:affiliation id="EBDF83F7FFD4DD026E5482A11EADBBCF">Borissiak Paleontological Institute, Russian Academy of Sciences, Moscow, Russia; e-mail: aponom @ paleo. ru</mods:affiliation>
</mods:name>
<mods:name id="9E86CA30EEB4606671C92D6BF63ED19C" type="personal">
<mods:role id="71A9584A8638FACCA8852EE35C3A08D6">
<mods:roleTerm id="53B3E2C0300314A4D94FCE91D62EE7C8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7E536FC54D96CDEDF61F9A368B34FAEA">Yue, Yanli</mods:namePart>
<mods:affiliation id="F5818FEF2084D70660D4CE5D15F66BFD">School of Life Science, Ningxia University, Ningxia, China; e-mail: yueyanli 0407 @ yahoo. cn</mods:affiliation>
</mods:name>
<mods:name id="B4EFB78524F083B3BE4958E2366FBF90" type="personal">
<mods:role id="36114F0AF40EDB482B988527CBDC6A0A">
<mods:roleTerm id="46447603F3D0074B8AAACA1C62862FA2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D9C31F35EFE1A80664756260AC428CDB">Ren, Dong</mods:namePart>
</mods:name>
<mods:name id="A56D21AF379B44CD90744F126DE5B776" type="personal">
<mods:role id="CE8A167203B043FF6A3C2622C70621EC">
<mods:roleTerm id="ED175E96B59294F6DF4B95363CC0CDE1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F24A7A21F9E67C7E0860D2769B5A2422">Prokop, Jakub</mods:namePart>
<mods:affiliation id="4249EC09F68DC27F52FF4CCD96037B23">Department of Zoology, Faculty of Science, Charles University in Prague, Viničná 7, CZ- 128 44 Praha 2, Czech Republic; e-mail: jprokop @ natur. cuni. cz</mods:affiliation>
</mods:name>
<mods:typeOfResource id="7551440A740ED5E978FBCD710707EF0B">text</mods:typeOfResource>
<mods:relatedItem id="C1C58B4961655A20A518E4C9D630B0BB" type="host">
<mods:titleInfo id="05DE3CF02884F76C2D27ED01D62A7B7A">
<mods:title id="C4F928390917362C3B3524DB5D357312">Acta Entomologica Musei Nationalis Pragae</mods:title>
</mods:titleInfo>
<mods:part id="31BE159B1648A2D098A3658FF8634692">
<mods:date id="8B8DCD0A839676DC8F43BED35C381617">2012</mods:date>
<mods:detail id="AB032774A40D89737DD7D457A077442C" type="pubDate">
<mods:number id="967929FC5E5558354E863042E8520DBB">2012-06-30</mods:number>
</mods:detail>
<mods:detail id="5CC5C59ADF808F627727DE90D36A7DA2" type="volume">
<mods:number id="41B6BA13058296998C758D597740134D">52</mods:number>
</mods:detail>
<mods:detail id="7CFD478251B94D9DBA931E502C093E03" type="issue">
<mods:number id="6985CDDA32F752F9A6690E192B09A0BD">1</mods:number>
</mods:detail>
<mods:extent id="087336191DE9E47754BC5637D7E7ECA9" unit="page">
<mods:start id="2843052239DDA7AAB52476B8919802C6">89</mods:start>
<mods:end id="55C144665F356F1F582A24DD12D7F72E">127</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="F9D700D2C4085EDEA92CB45ABD4982F8">journal article</mods:classification>
<mods:identifier id="AC67FCB9FACBA950BDF3B106A5C96BCE" type="DOI">10.5281/zenodo.5330604</mods:identifier>
<mods:identifier id="D1B155A8D5FE7BC46EF346FDDB696AA5" type="ISSN">0374-1036</mods:identifier>
<mods:identifier id="0CF64B535B29981AF4B207962E302524" type="Zenodo-Dep">5330604</mods:identifier>
</mods:mods>
<treatment id="B970A055FFC3FFE6FE1E2605FDC1FBF7" LSID="urn:lsid:plazi:treatment:B970A055FFC3FFE6FE1E2605FDC1FBF7" httpUri="http://treatment.plazi.org/id/B970A055FFC3FFE6FE1E2605FDC1FBF7" lastPageNumber="92" pageId="3" pageNumber="92">
<subSubSection id="79C342C8FFC3FFE6FE1E2605FC95FE90" box="[389,812,339,364]" pageId="3" pageNumber="92" type="nomenclature">
<paragraph id="31661143FFC3FFE6FE1E2605FC95FE90" blockId="3.[389,812,339,364]" box="[389,812,339,364]" pageId="3" pageNumber="92">
<heading id="6A2EA62FFFC3FFE6FE1E2605FC95FE90" bold="true" box="[389,812,339,364]" centered="true" fontSize="8" level="2" pageId="3" pageNumber="92" reason="3">
<emphasis id="03ADCD51FFC3FFE6FE1E2605FC95FE90" bold="true" box="[389,812,339,364]" pageId="3" pageNumber="92">
<taxonomicName id="F6D96AC0FFC3FFE6FE022605FC95FE90" ID-CoL="8XLWT" authority="Ponomarenko, 1987" authorityName="Ponomarenko" authorityYear="1987" box="[409,812,339,364]" class="Insecta" family="Helophoridae" genus="Hydrophilopsia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="03ADCD51FFC3FFE6FE022605FDFAFE97" bold="true" box="[409,579,339,364]" italics="true" pageId="3" pageNumber="92">Hydrophilopsia</emphasis>
<bibRefCitation id="55486CB2FFC3FFE6FDD12602FC95FE90" author="PONOMARENKO A. G." box="[586,812,339,364]" pageId="3" pageNumber="92" pagination="83 - 97" refId="ref15732" refString="PONOMARENKO A. G. 1987: Novye mezozoiskye vodnye zhestkokrylye (Insecta, Coleoptera) iz Azii. [New Mesozoic aquatic beetles (Insecta, Coleoptera) from Asia]. Paleontologischeskiy Zhurnal 1987: 83 - 97 (in Russian)." type="journal article" year="1987">Ponomarenko, 1987</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="79C342C8FFC3FFE6FFEA262CFE76FE76" box="[113,463,377,397]" pageId="3" pageNumber="92" type="reference_group">
<paragraph id="31661143FFC3FFE6FFEA262CFE76FE76" blockId="3.[113,463,377,397]" box="[113,463,377,397]" pageId="3" pageNumber="92">
<treatmentCitationGroup id="11C9366DFFC3FFE6FFEA262CFE76FE76" box="[113,463,377,397]" pageId="3" pageNumber="92">
<treatmentCitation id="B0783752FFC3FFE6FFEA262CFE72FE76" author="PONOMARENKO A. G." box="[113,459,377,397]" page="93" pageId="3" pageNumber="92" year="1987">
<taxonomicName id="F6D96AC0FFC3FFE6FFEA262CFE72FE76" ID-CoL="8XLWT" authority="Ponomarenko, 1987: 93" authorityName="Ponomarenko" authorityPageNumber="93" authorityYear="1987" box="[113,459,377,397]" class="Insecta" family="Helophoridae" genus="Hydrophilopsia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="03ADCD51FFC3FFE6FFEA262CFF4FFE77" box="[113,246,378,396]" italics="true" pageId="3" pageNumber="92">Hydrophilopsia</emphasis>
<bibRefCitation id="55486CB2FFC3FFE6FF67262FFE72FE76" author="PONOMARENKO A. G." box="[252,459,377,397]" pageId="3" pageNumber="92" pagination="83 - 97" refId="ref15732" refString="PONOMARENKO A. G. 1987: Novye mezozoiskye vodnye zhestkokrylye (Insecta, Coleoptera) iz Azii. [New Mesozoic aquatic beetles (Insecta, Coleoptera) from Asia]. Paleontologischeskiy Zhurnal 1987: 83 - 97 (in Russian)." type="journal article" year="1987">Ponomarenko, 1987: 93</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="79C342C8FFC3FFE6FFEA26F3FD64FE26" pageId="3" pageNumber="92" type="type_taxon">
<paragraph id="31661143FFC3FFE6FFEA26F3FD64FE26" blockId="3.[113,1075,421,477]" pageId="3" pageNumber="92">
<emphasis id="03ADCD51FFC3FFE6FFEA26F3FEBCFE46" bold="true" box="[113,261,421,445]" pageId="3" pageNumber="92">
<typeStatus id="EE62AFE1FFC3FFE6FFEA26F3FF10FE46" box="[113,169,421,445]" pageId="3" pageNumber="92">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="F6D96AC0FFC3FFE6FE9726F0FCA4FE46" ID-CoL="3N7WD" authority="Ponomarenko, 1987" authorityName="Ponomarenko" authorityYear="1987" box="[268,797,421,445]" class="Insecta" family="Helophoridae" genus="Hydrophilopsia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="92" phylum="Arthropoda" rank="species" species="longitarsalis">
<emphasis id="03ADCD51FFC3FFE6FE9726F0FDF9FE46" box="[268,576,422,445]" italics="true" pageId="3" pageNumber="92">Hydrophilopsia longitarsalis</emphasis>
<bibRefCitation id="55486CB2FFC3FFE6FDDD26F3FCA4FE46" author="PONOMARENKO A. G." box="[582,797,421,445]" pageId="3" pageNumber="92" pagination="83 - 97" refId="ref15732" refString="PONOMARENKO A. G. 1987: Novye mezozoiskye vodnye zhestkokrylye (Insecta, Coleoptera) iz Azii. [New Mesozoic aquatic beetles (Insecta, Coleoptera) from Asia]. Paleontologischeskiy Zhurnal 1987: 83 - 97 (in Russian)." type="journal article" year="1987">Ponomarenko, 1987</bibRefCitation>
</taxonomicName>
(by original designation).
<emphasis id="03ADCD51FFC3FFE6FFEA2693FF42FE26" bold="true" box="[113,251,453,477]" pageId="3" pageNumber="92">Time range.</emphasis>
Latest Jurassic, Tithonian, ca. 151146 mya.
</paragraph>
</subSubSection>
<subSubSection id="79C342C8FFC3FFE6FFEA26A2FDC1FBF7" pageId="3" pageNumber="92" type="diagnosis">
<paragraph id="31661143FFC3FFE6FFEA26A2FE41FCF7" blockId="3.[113,1089,499,1036]" pageId="3" pageNumber="92">
<emphasis id="03ADCD51FFC3FFE6FFEA26A2FE89FDF7" bold="true" box="[113,304,499,524]" pageId="3" pageNumber="92">
Diagnosis.
<emphasis id="03ADCD51FFC3FFE6FF7126A5FE9EFDF7" bold="true" box="[234,295,499,524]" italics="true" pageId="3" pageNumber="92">Adult</emphasis>
:
</emphasis>
Head and pronotum without setiferous granules, gula wide, pronotal flanks extremely wide, elytra without elevated costae, legs with dense series of swimming hairs on tibiae and tarsi; it is also probable that the pronotum lacks longitudinal grooves and that the median portion of frontoclypeal suture is not grooved (both these characters are incompletely preserved in the single specimen available). In the helophorid lineage, the extremely wide pronotal flanks delimited by a non-sinuate inner ridge are only developed in the modern
<taxonomicName id="F6D96AC0FFC3FFE6FFEA25E3FF56FD37" authorityName="Fabricius" authorityYear="1775" box="[113,239,693,716]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="03ADCD51FFC3FFE6FFEA25E3FF56FD37" box="[113,239,693,716]" italics="true" pageId="3" pageNumber="92">Helophorus</emphasis>
</taxonomicName>
subgenera
<taxonomicName id="F6D96AC0FFC3FFE6FEF025E3FDDAFD37" authority="Hope, 1838" authorityName="Hope" authorityYear="1838" box="[363,611,692,716]" class="Insecta" family="Curculionidae" genus="Empleurus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="03ADCD51FFC3FFE6FEF025E3FE66FD37" box="[363,479,693,716]" italics="true" pageId="3" pageNumber="92">Empleurus</emphasis>
Hope, 1838
</taxonomicName>
and
<taxonomicName id="F6D96AC0FFC3FFE6FD0125E3FBBCFD37" authority="Orchymont, 1927" authorityName="Orchymont" authorityYear="1927" box="[666,1029,692,716]" class="Insecta" family="Helophoridae" genus="Orphelophorus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="03ADCD51FFC3FFE6FD0125E3FC84FD37" box="[666,829,693,716]" italics="true" pageId="3" pageNumber="92">Orphelophorus</emphasis>
Orchymont, 1927
</taxonomicName>
; both of them bear very distinct granulation on the head and pronotum, lack swimming hairs on legs and bear strongly costate elytra.
</paragraph>
<paragraph id="31661143FFC3FFE6FFEA2442FDC1FBF7" blockId="3.[113,1089,499,1036]" pageId="3" pageNumber="92">
<emphasis id="03ADCD51FFC3FFE6FFEA2442FE89FCD7" bold="true" box="[113,304,788,812]" pageId="3" pageNumber="92">Taxonomic note.</emphasis>
The genus is largely defined by plesiomorphies, but bears at least one autapomorphy, i.e. extremely wide pronotal flanks, hence it may represent an extinct late Jurassic clade of the helophorid lineage. All characters shared between
<taxonomicName id="F6D96AC0FFC3FFE6FCC52403FBBDFC97" authorityName="Ponomarenko" authorityYear="1987" box="[862,1028,853,876]" class="Insecta" family="Helophoridae" genus="Hydrophilopsia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="92" phylum="Arthropoda" rank="species" species="longitarsalis">
<emphasis id="03ADCD51FFC3FFE6FCC52403FBBDFC97" box="[862,1028,853,876]" italics="true" pageId="3" pageNumber="92">H. longitarsalis</emphasis>
</taxonomicName>
(
<typeStatus id="EE62AFE1FFC3FFE6FB8A2402FBF9FC97" box="[1041,1088,852,876]" pageId="3" pageNumber="92">type</typeStatus>
species) and the remaining species assigned previously to this genus are plesiomorphic and
<taxonomicName id="F6D96AC0FFC3FFE6FFEA24C3FEAEFC57" authorityName="Ponomarenko" authorityYear="1987" box="[113,279,917,940]" class="Insecta" family="Helophoridae" genus="Hydrophilopsia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="03ADCD51FFC3FFE6FFEA24C3FEAEFC57" box="[113,279,917,940]" italics="true" pageId="3" pageNumber="92">Hydrophilopsia</emphasis>
</taxonomicName>
in its original concept seems to be paraphyletic or polyphyletic for that reason. In order to keep
<taxonomicName id="F6D96AC0FFC3FFE6FED524E3FE4DFC37" authorityName="Ponomarenko" authorityYear="1987" box="[334,500,949,972]" class="Insecta" family="Helophoridae" genus="Hydrophilopsia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="03ADCD51FFC3FFE6FED524E3FE4DFC37" box="[334,500,949,972]" italics="true" pageId="3" pageNumber="92">Hydrophilopsia</emphasis>
</taxonomicName>
monophyletic and easily diagnosable, we are excluding all other species into a separate genus
<taxonomicName id="F6D96AC0FFC3FFE6FD8A2483FDC3FC17" authority="Fikáček &amp; Prokin &amp; Angus &amp; Pono &amp; Marenko &amp; Yue &amp; Ren &amp; Prokop, 2012" authorityName="Fikáček &amp; Prokin &amp; Angus &amp; Pono &amp; Marenko &amp; Yue &amp; Ren &amp; Prokop" authorityYear="2012" box="[529,634,981,1004]" class="Insecta" family="Helophoridae" genus="Laetopsia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="92" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="03ADCD51FFC3FFE6FD8A2483FDC3FC17" box="[529,634,981,1004]" italics="true" pageId="3" pageNumber="92">Laetopsia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="189E702AFFC3FFE6FD192482FD5FFC10" box="[642,742,980,1004]" pageId="3" pageNumber="92" rank="genus">gen. nov.</taxonomicNameLabel>
, which makes
<taxonomicName id="F6D96AC0FFC3FFE6FC1C2483FB94FC17" authorityName="Ponomarenko" authorityYear="1987" box="[903,1069,981,1004]" class="Insecta" family="Helophoridae" genus="Hydrophilopsia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="03ADCD51FFC3FFE6FC1C2483FB94FC17" box="[903,1069,981,1004]" italics="true" pageId="3" pageNumber="92">Hydrophilopsia</emphasis>
</taxonomicName>
a monotypic genus confined to the Latest Jurassic.
</paragraph>
</subSubSection>
</treatment>
</document>