222 lines
24 KiB
XML
222 lines
24 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.75.884" ID-GBIF-Dataset="c11b212f-31f6-4c92-b218-c76d411c3941" ID-PMC="PMC3088044" ID-Pensoft-Pub="1313-2970-75-21" ID-PubMed="21594138" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-75-21" ModsDocOrigin="ZooKeys 75" ModsDocTitle="A new species of Amblyodus Westwood, 1878 (Coleoptera, Melolonthidae, Dynastinae) from South America" checkinTime="1451250841376" checkinUser="pensoft" docAuthor="Grossi, Paschoal Coelho & Grossi, Everardo Jose" docDate="2011" docId="9B788EC33C5935C5C8024C7C64ADA52D" docLanguage="en" docName="ZooKeys 75: 21-28" docOrigin="ZooKeys 75" docSource="http://dx.doi.org/10.3897/zookeys.75.884" docTitle="Amblyodus castroi Grossi & Grossi, sp. n." docType="treatment" docVersion="5" lastPageNumber="25" masterDocId="FFDBFFF8FFD7D943FF95B526FFFFAA79" masterDocTitle="A new species of Amblyodus Westwood, 1878 (Coleoptera, Melolonthidae, Dynastinae) from South America" masterLastPageNumber="28" masterPageNumber="21" pageNumber="23" updateTime="1668165716126" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A new species of Amblyodus Westwood, 1878 (Coleoptera, Melolonthidae, Dynastinae) from South America</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Grossi, Paschoal Coelho</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Grossi, Everardo Jose</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2011</mods:date>
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<mods:detail type="volume">
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<mods:number>75</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>21</mods:start>
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<mods:end>28</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.75.884</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.75.884</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-75-21</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="159361086" LSID="urn:lsid:zoobank.org:act:37121595-32A5-4339-93DD-BC7F1AE006B9" httpUri="http://treatment.plazi.org/id/9B788EC33C5935C5C8024C7C64ADA52D" lastPageId="4" lastPageNumber="25" pageId="2" pageNumber="23">
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<subSubSection pageId="2" pageNumber="23" type="nomenclature">
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<paragraph pageId="2" pageNumber="23">
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<taxonomicName LSID="urn:lsid:zoobank.org:act:37121595-32A5-4339-93DD-BC7F1AE006B9" authority="Grossi & Grossi" class="Insecta" family="Dynastidae" genus="Amblyodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblyodus castroi" order="Coleoptera" pageId="2" pageNumber="23" phylum="Arthropoda" rank="species" species="castroi">Amblyodus castroi Grossi & Grossi</taxonomicName>
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<taxonomicNameLabel pageId="2" pageNumber="23">sp. n.</taxonomicNameLabel>
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Figs 1-813, 14
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="3" lastPageNumber="24" pageId="2" pageNumber="23" type="material examined">
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<paragraph pageId="2" pageNumber="23">Material examined.</paragraph>
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<paragraph pageId="2" pageNumber="23">
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Holotype male, dissected, labeled: a) "male symbol label"; b) "Brasil, Amazonas,
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<taxonomicName lsidName="" pageId="2" pageNumber="23" rank="tribe" tribe="Uarini">Uarini</taxonomicName>
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,/ 03°02'57"S, 65°41'42"W,/ 22/VII-03/VIII/1995, P./
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<normalizedToken originalValue="Bührnhein">Buehrnhein</normalizedToken>
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, N.Aguiar & al."; c) handwritten label "NO ALBURNO DE/ TRONCO
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<normalizedToken originalValue="CAÍDO">CAIDO</normalizedToken>
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/ 01/VIII/1995" (UFAM).
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</paragraph>
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<paragraph lastPageId="3" lastPageNumber="24" pageId="2" pageNumber="23">
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Paratypes, 4 males and 7 females labeled: 1 female a) "female symbol label", b) "Brasil, Amazonas,
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<taxonomicName lsidName="" pageId="2" pageNumber="23" rank="tribe" tribe="Uarini">Uarini</taxonomicName>
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,/ 03°02'57"S, 65°41'42"W,/ 22/VII-03/VIII/1995, P./
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<normalizedToken originalValue="Bührnhein">Buehrnhein</normalizedToken>
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, N.Aguiar & al."; c) handwritten label "NO ALBURNO DE/ TRONCO
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<normalizedToken originalValue="CAÍDO">CAIDO</normalizedToken>
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/ 01/VIII/1995" (CEMT). 3 females labeled "Brasil, Amazonas, Coari,/ Duto
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<pageBreakToken pageId="3" pageNumber="24" start="start">Urucu</pageBreakToken>
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/Porto Terminal,/ 04°50'16"S, 65°20'36"W,/ 16/VI/1996, Buhmheim (sic),/ PF, Aguiar, NO, Arruda,/ AMR & Gualberto, TL col." (UFAM). 1 male labeled a) "Brasil, Amazonas, Coari,/ Duto Urucu/Porto Terminal,/ 04°50'16"S, 65°20'36"W,/ 16/VI/1996,
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<normalizedToken originalValue="Bühmheim">Buehmheim</normalizedToken>
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(sic),/ P.F. & Aguiar, N.O. col."; b) handwritten label "Em tronco/
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<normalizedToken originalValue="caído”">caido"</normalizedToken>
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(EPGC). 1 female labeled "PERU, Junin, Puerto/ Ocopa, I-2007, 600m/ 11°07'50"S, 74°17'46"W/ S.Castro col." (EPGC). 1 female a) "BRASIL:
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<normalizedToken originalValue="Rondônia">Rondonia</normalizedToken>
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: Porto/ Velho, margem direita/ do Rio Madeira, IX-2004" (EPCG). 1 male labeled "PERU, Ucayali, Via/ Puerto Ocopa-Ucayali,/ XII-2009, 1200m,/ 11°03'51"S, 74°15'39"W/ S. Castro col." (MIIS); 1 male labeled "PERU, Junin,Huerto/ Eden c.a, near Puerto/ Prado XII-2009/ 1800m, local collector" (JSC). 1 male labeled a) "BR 14, Km 92/
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<normalizedToken originalValue="Pará">Para</normalizedToken>
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, Brasil/ E. Lobato XII. 60" (IOC). 1 female labeled a) handwritten label "S. Paulo de/
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<normalizedToken originalValue="Olivença">Olivenca</normalizedToken>
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/
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<normalizedToken originalValue="Solimões">Solimoes</normalizedToken>
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/ VIII. - Fassl", b)
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<normalizedToken originalValue="“Coleção">"Colecao</normalizedToken>
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/ J. F. Zikan" (IOC). All paratypes additionally labeled, yellow label, "
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<taxonomicName class="Insecta" family="Dynastidae" genus="Amblyodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblyodus castroi" order="Coleoptera" pageId="3" pageNumber="24" phylum="Arthropoda" rank="species" species="castroi">Amblyodus castroi</taxonomicName>
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/ PARATYPE/ Grossi & Grossi.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="24" type="diagnosis">
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<paragraph pageId="3" pageNumber="24">Diagnosis.</paragraph>
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<paragraph pageId="3" pageNumber="24">
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Head with frontal horns less divergent, being almost parallel (Fig 3). Pronotal disc more declivous than in
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<taxonomicName class="Insecta" family="Dynastidae" genus="Amblyodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblyodus taurus" order="Coleoptera" pageId="3" pageNumber="24" phylum="Arthropoda" rank="species" species="taurus">Amblyodus taurus</taxonomicName>
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with lateral carinae distinctly higher in males majors (Fig 4), posterior furrow deeply excavated as opposed to a less excavated furrow in
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<taxonomicName class="Insecta" family="Dynastidae" genus="Amblyodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblyodus taurus" order="Coleoptera" pageId="3" pageNumber="24" phylum="Arthropoda" rank="species" species="taurus">Amblyodus taurus</taxonomicName>
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(Figs 9 and 10). Aedeagus with internal margin of parameres concave, not parallel; external margin less constricted than in
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<taxonomicName class="Insecta" family="Dynastidae" genus="Amblyodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblyodus taurus" order="Coleoptera" pageId="3" pageNumber="24" phylum="Arthropoda" rank="species" species="taurus">Amblyodus taurus</taxonomicName>
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where internal margin is almost parallel, and external margin abruptly constricted (Figs 11 and 12).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="4" lastPageNumber="25" pageId="3" pageNumber="24" type="holotype description">
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<paragraph pageId="3" pageNumber="24">Holotype description</paragraph>
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<paragraph lastPageId="4" lastPageNumber="25" pageId="3" pageNumber="24">
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(Figs 1 and 2). Total body length 20 mm. Total pronotal width 9.2 mm. Body elongated, about 2 times longer than wide, color dark brown dorsally and reddish brown ventrally; dorsal surface glabrous, with sterna being setose. Head: Surface smooth and glabrous with sparse micropunctures; in frontal view with two lateral, concave areas with scattered C-shaped puntures. Clypeus subtriangular, laterally bordered and slightly upturned, convex and with acute apex. Canthi intruding upon eyes just before first third. Frons with two erect, diverging, posteriorly recurving horns. Interocular width equals 3.3 mm. Mandibles tridentate with inner tooth more acute; teeth upturned. Antennae 10 segmented with scape as long as the 6 next segments; club with segments subequal in length. Pronotum: Shape trapezoidal, narrower than both elytra together; discal area with C-shaped punctures combined with reticulated areas, flat and declivous; anterior border incomplete. Lateral and posterior sides of pronotal depression strongly carinated, anterior carina more pronounced, higher, being convex and wider posteriorly; carina at base weakly emarginate; carina posteriorly interrupted at middle by a longitudinal furrow, internally weakly angulate; furrow about 4.5 times narrower than head width. Anterior angles acute, posterior angles rounded. Outer side and lateral posterior side of pronotal surface punctate; punctures ocellate, sometimes coalescent, posteriorly larger. Elytra: Discal area with 5 rows of large ocellate punctures, rows weakly impressed. Interstriae slightly convex, with scattered small punctures. Lateral edges with 5 similar rows of ocellate punctures, instead of by some ocellate punctures between third and fourth rows. Pygidium: Shape convex in lateral view; surface densely punctate; punctures ocellate, moderately sized, elongate near dorsal margin. Sixth ventrite sinuous at middle. Legs: Protibiae quadridentate, proximal tooth smaller, distal ones bigger. Metatibiae with 5 apical teeth. Parameres:
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<pageBreakToken pageId="4" pageNumber="25" start="start">Shape</pageBreakToken>
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symmetrical, narrower at apex; apex dilated, laterally roundly angulated, apically rounded, triangle shaped; internal side of each paramera concave, not parallel (Fig 5). In lateral view shaft constricted, with a post-anterior concavity; dorso-ventral carina incomplete, not reaching ventral margin; with a flat, truncate tooth near base (Fig 6).
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</paragraph>
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<caption pageId="4" pageNumber="25">
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<paragraph pageId="4" pageNumber="25">
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Figures 1-2. Habitus of
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<taxonomicName class="Insecta" family="Dynastidae" genus="Amblyodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblyodus castroi" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="species" species="castroi">Amblyodus castroi</taxonomicName>
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sp. n., male holotype. 1 Dorsal view 2 Lateral view.
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</paragraph>
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</caption>
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<paragraph pageId="4" pageNumber="25">Female paratypes. Similar to male in general aspects except for the following characters: Head: Frontal horns less developed and never reaching pronotal middle. Pronotum: Pronotal depression narrower, near anterior margin smooth, not punctate; lateral carina smaller with a weak acute projection before first third (Figs 7 and 8); pronotal furrow longer, ending just before posterior margin. Pygidium: Shape wider laterally and shorter, with punctures smaller. Sixth ventrite not sinuous at middle.</paragraph>
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<caption pageId="4" pageNumber="25">
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<paragraph pageId="4" pageNumber="25">
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Figures 3-14.
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<taxonomicName class="Insecta" family="Dynastidae" genus="Amblyodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblyodus" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="genus">Amblyodus</taxonomicName>
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spp. 3-4 Head and pronotum of a male major of
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<taxonomicName class="Insecta" family="Dynastidae" genus="Amblyodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblyodus castroi" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="species" species="castroi">Amblyodus castroi</taxonomicName>
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in dorsal and lateral views respectively 5-6 Parameres of
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<taxonomicName class="Insecta" family="Dynastidae" genus="Amblyodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblyodus castroi" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="species" species="castroi">Amblyodus castroi</taxonomicName>
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in caudal and lateral views respectively 7-8 Head and pronotum of a female of
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<taxonomicName class="Insecta" family="Dynastidae" genus="Amblyodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblyodus castroi" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="species" species="castroi">Amblyodus castroi</taxonomicName>
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in dorsal and lateral views respectively 9-10 Head and pronotum of
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<taxonomicName class="Insecta" family="Dynastidae" genus="Amblyodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblyodus taurus" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="species" species="taurus">Amblyodus taurus</taxonomicName>
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in dorsal and lateral views respectively 11-12 Parameres of
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<taxonomicName class="Insecta" family="Dynastidae" genus="Amblyodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblyodus taurus" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="species" species="taurus">Amblyodus taurus</taxonomicName>
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in caudal and lateral views respectively 13-14 Head and pronotum of a male minor of
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<taxonomicName class="Insecta" family="Dynastidae" genus="Amblyodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblyodus castroi" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="species" species="castroi">Amblyodus castroi</taxonomicName>
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in dorsal and lateral views respectively. Arrows indicate apical lateral carina, 6 (incomplete) and 12 (complete).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="25" type="variation of male paratypes">
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<paragraph pageId="4" pageNumber="25">Variation of male paratypes.</paragraph>
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<paragraph pageId="4" pageNumber="25">Smaller males possess less developed and less posteriorly recurving frontal horns. Pronotal carina very weakly pronounced (Figs 13 and 14).</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="25" type="etymology">
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<paragraph pageId="4" pageNumber="25">Etymology.</paragraph>
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<paragraph pageId="4" pageNumber="25">The species is named castroi in homage of Sergio Castro who provided us with part of the paratypes from Peru.</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="25" type="remarks">
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<paragraph pageId="4" pageNumber="25">Remarks.</paragraph>
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<paragraph pageId="4" pageNumber="25">
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When recording the new occurrence of
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<taxonomicName class="Insecta" family="Dynastidae" genus="Amblyodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblyodus taurus" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="species" species="taurus">Amblyodus taurus</taxonomicName>
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for South America,
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<bibRefCitation author="Gasca Alvarez, HJ" journalOrPublisher="Acta Zoologica Mexicana (nueva serie)" pageId="4" pageNumber="25" pagination="221 - 224" title="Primer Registro de Amblyodus taurus Westwood, 1878 (Scarabaeidae: Dynastinae: Phileurini) em Brasil." volume="24" year="2008">Gasca and Aguiar (2008)</bibRefCitation>
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misapplied the name
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<taxonomicName class="Insecta" family="Dynastidae" genus="Amblyodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblyodus taurus" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="species" species="taurus">Amblyodus taurus</taxonomicName>
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to the material they had on hand. Actually both species of
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<taxonomicName class="Insecta" family="Dynastidae" genus="Amblyodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblyodus" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="genus">Amblyodus</taxonomicName>
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are similar in appearance, which is common in
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<taxonomicName lsidName="" pageId="4" pageNumber="25" rank="tribe" tribe="Phileurini">Phileurini</taxonomicName>
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. The authors also stated that the only two genera of the tribe that possess elongate frontal horns were
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<taxonomicName class="Insecta" family="Dynastidae" genus="Amblyodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblyodus" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="genus">Amblyodus</taxonomicName>
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and
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<taxonomicName class="Insecta" family="Dynastidae" genus="Oryctophileurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oryctophileurus" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="genus">Oryctophileurus</taxonomicName>
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Kolbe, 1910. However,
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<taxonomicName class="Insecta" family="Dynastidae" genus="Ceratophileurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratophileurus" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="genus">Ceratophileurus</taxonomicName>
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Ohaus, 1911 also possess this character in both sexes (
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<bibRefCitation author="Gillett, CPDT" journalOrPublisher="ZooKeys" pageId="4" pageNumber="25" pagination="193 - 198" title="The rare rhinoceros beetle, Ceratophileurus lemouti Ohaus, 1911, in French Guiana and Suriname (Coleoptera, Scarabaeidae, Dynastinae, Phileurini)." volume="34" year="2010">Gillett et al. 2010</bibRefCitation>
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).
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</paragraph>
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<paragraph pageId="4" pageNumber="25">
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<taxonomicName class="Insecta" family="Dynastidae" genus="Amblyodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblyodus" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="genus">Amblyodus</taxonomicName>
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and
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<taxonomicName class="Insecta" family="Dynastidae" genus="Oryctophileurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oryctophileurus" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="genus">Oryctophileurus</taxonomicName>
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share the lateral pronotal carina and pronotal shape: elongate, widely flattened, depressed with discal depression totally wrinkled. Additionally, both genera have almost no external sexual dimorphism, as females also possess a well-developed frontal horn, a feature that makes the distinction between genders difficult, being the shape of the last sternite the better way to determine gender in these cases. Moreover parameres have almost the same shape, and the shape of prosternal process is also shared between
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<taxonomicName class="Insecta" family="Dynastidae" genus="Amblyodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblyodus" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="genus">Amblyodus</taxonomicName>
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and
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<taxonomicName class="Insecta" family="Dynastidae" genus="Oryctophileurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oryctophileurus" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="genus">Oryctophileurus</taxonomicName>
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, namely flat ventrally and with a spine-like process posteriorly. Based on several morphological characters, there is a possibility that
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<taxonomicName class="Insecta" family="Dynastidae" genus="Oryctophileurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oryctophileurus" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="genus">Oryctophileurus</taxonomicName>
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could be a synonym of
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<taxonomicName class="Insecta" family="Dynastidae" genus="Amblyodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblyodus" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="genus">Amblyodus</taxonomicName>
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because the unique feature to distinguishing both genera is the number of frontal horns, one or two, which is not a good character within
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<taxonomicName lsidName="" pageId="4" pageNumber="25" rank="subFamily" subFamily="Dynastinae">Dynastinae</taxonomicName>
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as some genera are usually extremely allometric. Only with the examination of the type species of
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<taxonomicName class="Insecta" family="Dynastidae" genus="Oryctophileurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oryctophileurus" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="genus">Oryctophileurus</taxonomicName>
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, will it be possible to affirm if they are congeneric or not.
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</paragraph>
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<paragraph pageId="4" pageNumber="25">
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The holotype male is an incomplete specimen and it was apparently damaged by an axe during collecting inside dead logs. It is the same male specimen illustrated by
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<bibRefCitation author="Gasca Alvarez, HJ" journalOrPublisher="Acta Zoologica Mexicana (nueva serie)" pageId="4" pageNumber="25" pagination="221 - 224" title="Primer Registro de Amblyodus taurus Westwood, 1878 (Scarabaeidae: Dynastinae: Phileurini) em Brasil." volume="24" year="2008">Gasca and Aguiar (2008, figure 1, male in lateral view)</bibRefCitation>
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.
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</paragraph>
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<paragraph pageId="4" pageNumber="25">
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The genus
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<taxonomicName class="Insecta" family="Dynastidae" genus="Amblyodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblyodus" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="genus">Amblyodus</taxonomicName>
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is recorded for the first time from Peru and from Brazil states of
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<normalizedToken originalValue="Pará">Para</normalizedToken>
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and
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<normalizedToken originalValue="Rondônia">Rondonia</normalizedToken>
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(Fig 15). It is possible that
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<taxonomicName class="Insecta" family="Dynastidae" genus="Amblyodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblyodus castroi" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="species" species="castroi">Amblyodus castroi</taxonomicName>
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occurs in all the Amazonian subregion.
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</paragraph>
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<caption pageId="4" pageNumber="25">
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<paragraph pageId="4" pageNumber="25">
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Figure 15. Known distribution map of
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<taxonomicName class="Insecta" family="Dynastidae" genus="Amblyodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblyodus" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="genus">Amblyodus</taxonomicName>
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species.
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<taxonomicName class="Insecta" family="Dynastidae" genus="Amblyodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblyodus taurus" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="species" species="taurus">Amblyodus taurus</taxonomicName>
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|
(circles) and
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<taxonomicName class="Insecta" family="Dynastidae" genus="Amblyodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblyodus castroi" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="species" species="castroi">Amblyodus castroi</taxonomicName>
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(squares).
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</paragraph>
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</caption>
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<paragraph pageId="4" pageNumber="25">
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Characters used to diagnose the genus by
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<bibRefCitation author="Ratcliffe, BC" journalOrPublisher="Bulletin of the University of the Nebraska State Museum 16:, 1 - 50" pageId="4" pageNumber="25" title="The Dynastine Scarab Beetles of Costa Rica and Panama (Coleoptera, Scarabaeidae, Dynastinae)." year="2003">Ratcliffe (2003)</bibRefCitation>
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and
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<bibRefCitation author="Ratcliffe, BC" journalOrPublisher="Bulletin of the University of the Nebraska State Museum 21:, 1 - 42" pageId="4" pageNumber="25" title="The Dynastine Scarab Beetles of Honduras, Nicaragua and El Salvador (Coleoptera: Scarabaeidae: Dynastinae)." year="2006">Ratcliffe and Cave (2006)</bibRefCitation>
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are confirmed with the new species described here, except for the presence of six teeth on the apex of the metatibiae. In
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<taxonomicName class="Insecta" family="Dynastidae" genus="Amblyodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblyodus castroi" order="Coleoptera" pageId="4" pageNumber="25" phylum="Arthropoda" rank="species" species="castroi">Amblyodus castroi</taxonomicName>
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|
, that is a variable character, as the number of teeth may be five or six.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |