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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.340.5470" ID-GBIF-Dataset="35e1f7dd-6c8a-457f-a648-4cb716ffe363" ID-PMC="PMC3800796" ID-Pensoft-Pub="1313-2970-340-1" ID-PubMed="24146589" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-340-1" ModsDocOrigin="ZooKeys 340" ModsDocTitle="Taxonomic and nomenclatural notes on the genera Themus Motschulsky and Lycocerus Gorham (Coleoptera, Cantharidae)" checkinTime="1451246833308" checkinUser="pensoft" docAuthor="Yang, Yuxia, Kopetz, Andreas &amp; Yang, Xingke" docDate="2013" docId="9B042F861CE013EF2E4E170F1D47016D" docLanguage="en" docName="ZooKeys 340: 1-19" docOrigin="ZooKeys 340" docSource="http://dx.doi.org/10.3897/zookeys.340.5470" docTitle="Themus (Telephorops) uniformis Wittmer 1983, stat. n." docType="treatment" docVersion="4" lastPageNumber="3" masterDocId="FFE3C7763B12FFC4A9220B69FFF7FFC8" masterDocTitle="Taxonomic and nomenclatural notes on the genera Themus Motschulsky and Lycocerus Gorham (Coleoptera, Cantharidae)" masterLastPageNumber="19" masterPageNumber="1" pageNumber="3" updateTime="1668156592509" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Taxonomic and nomenclatural notes on the genera Themus Motschulsky and Lycocerus Gorham (Coleoptera, Cantharidae)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Yang, Yuxia</mods:namePart>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Kopetz, Andreas</mods:namePart>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Yang, Xingke</mods:namePart>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2013</mods:date>
<mods:detail type="volume">
<mods:number>340</mods:number>
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<mods:start>1</mods:start>
<mods:end>19</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.340.5470</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.340.5470</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-340-1</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3499694" ID-GBIF-Taxon="152048630" ID-Zenodo-Dep="3499694" LSID="urn:lsid:plazi:treatment:9B042F861CE013EF2E4E170F1D47016D" httpUri="http://treatment.plazi.org/id/9B042F861CE013EF2E4E170F1D47016D" lastPageNumber="3" pageId="2" pageNumber="3">
<subSubSection pageId="2" pageNumber="3" type="nomenclature">
<paragraph pageId="2" pageNumber="3">
<taxonomicName authority="Wittmer, 1983" authorityName="Wittmer" authorityYear="1983" class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Themus (Telephorops) uniformis" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="uniformis" subGenus="Telephorops">Themus (Telephorops) uniformis Wittmer, 1983</taxonomicName>
<taxonomicNameLabel pageId="2" pageNumber="3">stat. n.</taxonomicNameLabel>
Figs 1-2
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="3" type="reference_group">
<paragraph pageId="2" pageNumber="3">
<taxonomicName class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Themus" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">Themus</taxonomicName>
(s.str.) bitinctus uniformis
<bibRefCitation author="Wittmer, W" journalOrPublisher="Entomologischen Arbeiten aus dem Museum G. Frey" pageId="13" pageNumber="14" pagination="189 - 239" title="Beitrag zur einer Revision der Gattung Themus Motsch. Coleoptera: Cantharidae." volume="31 - 32" year="1983 a">Wittmer 1983a</bibRefCitation>
: 218, fig. 30.
</paragraph>
<paragraph pageId="2" pageNumber="3">
<taxonomicName class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Themus" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">Themus</taxonomicName>
(s.str.) cribripennis
<bibRefCitation author="Wittmer, W" journalOrPublisher="Entomological Review of Japan" pageId="13" pageNumber="14" pagination="147 - 172" title="Die Familie Cantharidae (Col.) auf Taiwan (2. Teil)." volume="38" year="1983 b">Wittmer 1983b</bibRefCitation>
: 151, figs. 46, 49.
</paragraph>
<paragraph pageId="2" pageNumber="3">
<taxonomicName class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Themus (Telephorops) bitinctus subsp. uniformis" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="subSpecies" species="bitinctus" subGenus="Telephorops" subSpecies="uniformis">Themus (Telephorops) bitinctus uniformis</taxonomicName>
:
<bibRefCitation author="Svihla, V" journalOrPublisher="Veroeffentlichungen des Naturkundemuseums Erfurt" pageId="12" pageNumber="13" pagination="183 - 190" title="Redescription of the subgenera of the genus Themus Motschulsky, 1858, with description of five new species (Coleoptera: Cantharidae)." volume="27" year="2008">
<normalizedToken originalValue="Švihla">Svihla</normalizedToken>
2008
</bibRefCitation>
: 187.
</paragraph>
<paragraph pageId="2" pageNumber="3">
<taxonomicName class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Themus (Telephorops) cribripennis" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="cribripennis" subGenus="Telephorops">Themus (Telephorops) cribripennis</taxonomicName>
:
<bibRefCitation author="Svihla, V" journalOrPublisher="Veroeffentlichungen des Naturkundemuseums Erfurt" pageId="12" pageNumber="13" pagination="183 - 190" title="Redescription of the subgenera of the genus Themus Motschulsky, 1858, with description of five new species (Coleoptera: Cantharidae)." volume="27" year="2008">
<normalizedToken originalValue="Švihla">Svihla</normalizedToken>
2008
</bibRefCitation>
: 187. syn. n.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="3" type="type material examined">
<paragraph pageId="2" pageNumber="3">Type material examined.</paragraph>
<paragraph pageId="2" pageNumber="3">
<taxonomicName class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Themus" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">Themus</taxonomicName>
(s.str.) bitinctus uniformis: Holotype ♂ (NHMB): [p]&quot;Yen-ping, China \ VII.21. 1917 \ Ac. 5148&quot;, [h]&quot;bitinctus \ uniformis&quot;, [p]
<normalizedToken originalValue="“HOLOTYPUS”">&quot;HOLOTYPUS&quot;</normalizedToken>
, [p]&quot;Naturhist. \ Museum Basel \ coll. W. Wittmer&quot;, [p]&quot;CANTHARIDAE \ CANTH00000449&quot;.
</paragraph>
<paragraph pageId="2" pageNumber="3">
<taxonomicName class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Themus" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">Themus</taxonomicName>
(s.str.) cribripennis: Holotype ♂ (MNHN): [p]&quot;Taihorinsho \ Formosa \ Sauter, VIII_ 7_09&quot;, [h]&quot;Cantharis \ davidis Fairm.&quot;, [h]&quot;Themus (s.str.) \ cribripennis \ Wittm. \ det. W. Wittmer&quot;, [p]
<normalizedToken originalValue="“HOLOTYPUS”">&quot;HOLOTYPUS&quot;</normalizedToken>
, [h]
<normalizedToken originalValue="“136”">&quot;136&quot;</normalizedToken>
. Paratype: 1♀ (NHMB): [p]&quot;Suisharyo \ Formosa \ H. Sauter, X.1911&quot;, [h]&quot;Themus s.str. \ cribripennis \ Wittm. \ det. W. Wittmer&quot;, [p]
<normalizedToken originalValue="“PARATYPUS”">&quot;PARATYPUS&quot;</normalizedToken>
, [p]&quot;CANTHARIDAE \ CANTH00002654&quot;.
</paragraph>
<caption pageId="2" pageNumber="3">
<paragraph pageId="2" pageNumber="3">
Figures 1-9. Habitus, dorsal view 1 Holotype of
<taxonomicName class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Themus" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">Themus</taxonomicName>
(s.str.) bitinctus uniformis Wittmer, 1983 2 Holotype of
<taxonomicName class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Themus" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">Themus</taxonomicName>
(s.str.) cribripennis Wittmer, 1983 3 Holotype of
<taxonomicName class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Themus licenti" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="licenti">Themus licenti</taxonomicName>
Pic, 1938 4 Holotype of
<taxonomicName class="Insecta" family="Cantharidae" genus="Telephorus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Telephorus coriaceipennis" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="coriaceipennis">Telephorus coriaceipennis</taxonomicName>
Fairmaire, 1889 5 Lectotype of
<taxonomicName class="Insecta" family="Lampyridae" genus="Podabrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Podabrus aenescens" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="aenescens">Podabrus aenescens</taxonomicName>
Fairmaire, 1889 6 Holotype of
<taxonomicName class="Insecta" family="Lampyridae" genus="Cantharis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cantharis tcheonana" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="tcheonana">Cantharis tcheonana</taxonomicName>
Pic, 1922 7 Holotype of
<taxonomicName class="Insecta" family="Cantharidae" genus="Athemus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Athemus (Isathemus) bilineatus" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="bilineatus" subGenus="Isathemus">Athemus (Isathemus) bilineatus</taxonomicName>
Wittmer, 19958 Holotype of
<taxonomicName class="Insecta" family="Cantharidae" genus="Athemus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Athemus" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">Athemus</taxonomicName>
(s.str.) amplus Wittmer, 1995 9 Holotype of
<taxonomicName class="Insecta" family="Cantharidae" genus="Telephorus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Telephorus dimidiaticrus" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="dimidiaticrus">Telephorus dimidiaticrus</taxonomicName>
Fairmaire, 1889. 1-2, 5, 7-8, 9 male 3-4, 6 female.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="2" pageNumber="3" type="distribution">
<paragraph pageId="2" pageNumber="3">Distribution.</paragraph>
<paragraph pageId="2" pageNumber="3">China (Fujian, Taiwan).</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="3" type="remarks">
<paragraph pageId="2" pageNumber="3">Remarks.</paragraph>
<paragraph pageId="2" pageNumber="3">
Having examined the holotypes of
<taxonomicName class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Themus cribripennis" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="cribripennis">Themus cribripennis</taxonomicName>
Wittmer, 1983b and
<taxonomicName class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Themus bitinctus subsp. uniformis" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="subSpecies" species="bitinctus" subSpecies="uniformis">Themus bitinctus uniformis</taxonomicName>
Wittmer, 1983a, we were unable to find differences justifying their separation, which has led us to consider all the examined specimens of both nominal species to be conspecific. Therefore, we synonymized
<taxonomicName class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Themus cribripennis" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="cribripennis">Themus cribripennis</taxonomicName>
under
<taxonomicName class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Themus bitinctus subsp. uniformis" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="subSpecies" species="bitinctus" subSpecies="uniformis">Themus bitinctus uniformis</taxonomicName>
. Furthermore,
<taxonomicName class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Themus bitinctus subsp. uniformis" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="subSpecies" species="bitinctus" subSpecies="uniformis">Themus bitinctus uniformis</taxonomicName>
should be upgraded to the specific rank, because it is obviously different from
<taxonomicName class="Insecta" family="Cantharidae" genus="Themus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Themus bitinctus" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="bitinctus">Themus bitinctus</taxonomicName>
Wittmer, 1982 (located in Vietnam) in the aedeagus, except the difference in the elytra coloration from the latter.
</paragraph>
</subSubSection>
</treatment>
</document>
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