343 lines
43 KiB
XML
343 lines
43 KiB
XML
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<mods:title id="3CE3B328A099FDE37A24E1F200EE2676">Chinchaysuyoa, a new genus of the fish family Ariidae (Siluriformes), with a redescription of Chinchaysuyoa labiata from Ecuador and a new species description from Peru</mods:title>
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<mods:namePart id="7763E07984452FE3EBF611329DDAC750">Marceniuk, Alexandre Pires</mods:namePart>
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<mods:namePart id="DFAC2D54F59AE05624F7FC843418111C">Marchena, Jose</mods:namePart>
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<mods:namePart id="3ED81E68236FDB5F764B7D211E140922">Oliveira, Claudio</mods:namePart>
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<mods:namePart id="5C5F32CBB4DB28D89B938377BFCECF77">Betancur-R, Ricardo</mods:namePart>
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<mods:date id="BD2A414E098EAB98AD0F48A4610B3C23">2019</mods:date>
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<treatment id="03D48796EC74FFA0F8909D4CFE04FEA5" ID-DOI="http://doi.org/10.5281/zenodo.5935997" ID-GBIF-Taxon="154533746" ID-Zenodo-Dep="5935997" LSID="urn:lsid:plazi:treatment:03D48796EC74FFA0F8909D4CFE04FEA5" httpUri="http://treatment.plazi.org/id/03D48796EC74FFA0F8909D4CFE04FEA5" lastPageId="11" lastPageNumber="372" pageId="7" pageNumber="368">
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<heading id="D08A81ECEC74FFACF8909D4CFDEAFA71" bold="true" box="[151,581,1401,1428]" fontSize="11" level="1" pageId="7" pageNumber="368" reason="1">
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<emphasis id="B909EA92EC74FFACF8909D4CFDEAFA71" bold="true" box="[151,581,1401,1428]" pageId="7" pageNumber="368">
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<taxonomicName id="4C7D4D03EC74FFACF8909D4CFE01FA76" authority="Marceniuk & Marchena & Oliveira & Betancur-R, 2019" authorityName="Marceniuk & Marchena & Oliveira & Betancur-R" authorityYear="2019" box="[151,430,1401,1427]" class="Actinopterygii" family="Ariidae" genus="Chinchaysuyoa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="368" phylum="Chordata" rank="species" species="ortegai" status="sp. nov.">
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<emphasis id="B909EA92EC74FFACF8909D4CFE01FA76" bold="true" box="[151,430,1401,1427]" italics="true" pageId="7" pageNumber="368">Chinchaysuyoa ortegai</emphasis>
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<taxonomicNameLabel id="A23A57E9EC74FFACF9B29D4FFDEAFA71" box="[437,581,1402,1428]" pageId="7" pageNumber="368" rank="species">new species</taxonomicNameLabel>
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<paragraph id="8BC23680EC74FFACF8909DA9FE28FA50" blockId="7.[151,581,1401,1461]" box="[151,391,1436,1461]" pageId="7" pageNumber="368">
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<figureCitation id="13462A05EC74FFACF8909DA9FF57FA50" box="[151,248,1436,1461]" captionStart="FIGURE 4" captionStartId="8.[151,250,1735,1757]" captionTargetBox="[150,1436,193,1714]" captionTargetId="figure@8.[150,1436,193,1714]" captionTargetPageId="8" captionText="FIGURE 4. Body in lateral view. Chinchaysuyoa labiata, (A) holotype 475 mm SL (MZUT 1540), (B) Hexanematichthys henni, holotype 139 mm SL (CAS 60620), (C) Chinchaysuyoa ortegai, holotype 256 mm SL (MUSM 63800), and (D) fresh non type specimen, not cataloged, 315 mm SL." httpUri="https://zenodo.org/record/2622976/files/figure.png" pageId="7" pageNumber="368">Figure 4</figureCitation>
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,
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<figureCitation id="13462A05EC74FFACF8FA9DA9FEA3FA50" box="[253,268,1436,1461]" captionStart="FIGURE 5" captionStartId="9.[151,250,1410,1432]" captionTargetBox="[150,1436,409,1389]" captionTargetId="figure@9.[150,1436,409,1389]" captionTargetPageId="9" captionText="FIGURE 5. Head in dorsal view. Chinchaysuyoa labiata, (A) holotype 475 mm SL (MZUT 1540), and Chinchaysuyoa ortegai, (B) holotype 256 mm SL (MUSM 63800)." httpUri="https://zenodo.org/record/2622978/files/figure.png" pageId="7" pageNumber="368">5</figureCitation>
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,
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<figureCitation id="13462A05EC74FFACF9169DA9FE8EFA50" box="[273,289,1436,1461]" captionStart="FIGURE 6" captionStartId="10.[151,250,1977,1999]" captionTargetBox="[150,1436,589,1955]" captionTargetId="figure@10.[150,1436,589,1955]" captionTargetPageId="10" captionText="FIGURE 6. Dorsal view of the neurocranium and the nuchal plate of Chinchaysuyoa labiata, (A) paratype 113 mm SL (CAS 60621), and Chinchaysuyoa ortegai, (B) paratype 155 mm SL (MPEG 35976)." httpUri="https://zenodo.org/record/2622980/files/figure.png" pageId="7" pageNumber="368">6</figureCitation>
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,
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<tableCitation id="C6FF033BEC74FFACF92A9DA9FE2EFA50" box="[301,385,1436,1461]" captionStart="TABLE 1" captionStartId="13.[151,239,332,355]" captionText="TABLE 1. Morphometric data for Chinchaysuyoa labiata and C. ortegai. Standard length is expressed in millimeters and all other measurements expressed in percents of standard length." pageId="7" pageNumber="368">Table 1</tableCitation>
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.
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</paragraph>
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<taxonomicName id="4C7D4D03EC74FFACF8909DD1FE39FA1F" ID-CoL="3LGKM" authorityName="Fisher & Eigenmann" authorityYear="1922" box="[151,406,1508,1530]" class="Actinopterygii" family="Ariidae" genus="Hexanematichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="368" phylum="Chordata" rank="species" species="henni">
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<emphasis id="B909EA92EC74FFACF8909DD1FE39FA1F" box="[151,406,1508,1530]" italics="true" pageId="7" pageNumber="368">Hexanematichthys henni</emphasis>
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</taxonomicName>
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(non Fisher & Eigenmann).—
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<treatmentCitation id="0ADC1091EC74FFACFADB9DD1FC19FA1F" author="Chirichigno, F. N." box="[732,950,1508,1530]" page="27" pageId="7" pageNumber="368" year="1963">
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<bibRefCitation id="EFEC4B71EC74FFACFADB9DD1FC37FA1F" author="Chirichigno, F. N." box="[732,920,1508,1530]" pageId="7" pageNumber="368" pagination="1 - 87" refId="ref7250" refString="Chirichigno, F. N. (1963) Estudio de la fauna ictiologica de los esteros y parte baja de los rios del Departamento de Tumbes [Peru]. Serrie de divulgacion cientifica / Servicio de Pesqueria, Ministerio de Agriculture, Lima, 22, 1 - 87." type="journal article" year="1963">Chirichigno 1963</bibRefCitation>
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:27
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(
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<collectingRegion id="49B9F862EC74FFACFBCE9DD1FBB2FA1F" box="[969,1053,1508,1530]" country="Peru" name="Tumbes" pageId="7" pageNumber="368">Tumbes</collectingRegion>
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).—
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<treatmentCitation id="0ADC1091EC74FFACFC459DD1FA91FA1F" author="Ortega, H. & Vari, R. P." box="[1090,1342,1508,1530]" page="13" pageId="7" pageNumber="368" year="1986">
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<bibRefCitation id="EFEC4B71EC74FFACFC459DD1FA8DFA1F" author="Ortega, H. & Vari, R. P." box="[1090,1314,1508,1530]" pageId="7" pageNumber="368" pagination="25" refId="ref9057" refString="Ortega, H. & Vari, R. P. (1986) Annotated checklist of the freshwater fishes of Peru. Smithsonian Contributions to Zoology, 437, iii- 25." type="journal article" year="1986">Ortega & Vari 1986</bibRefCitation>
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:13
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||
</treatmentCitation>
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||
(species list).—
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<treatmentCitation id="0ADC1091EC74FFACF90C9E36FE4BF9FC" author="Ortega, H. & Hidalgo, M. & Trevejo, G. & Correa, E. & Cortijo, A. M. & Meza, V. & Espino, J." box="[267,484,1539,1561]" page="42" pageId="7" pageNumber="368" year="2012">
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<bibRefCitation id="EFEC4B71EC74FFACF90C9E36FE68F9FC" author="Ortega, H. & Hidalgo, M. & Trevejo, G. & Correa, E. & Cortijo, A. M. & Meza, V. & Espino, J." box="[267,455,1539,1561]" pageId="7" pageNumber="368" refId="ref9090" refString="Ortega, H., Hidalgo, M., Trevejo, G., Correa, E., Cortijo, A. M., Meza, V. & Espino, J. (2012) Lista anotada de los peces de aguas continentales de Peru. Estado actual Del" type="book" year="2012">
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Ortega
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||
<emphasis id="B909EA92EC74FFACF95E9E31FE29F9FC" box="[345,390,1539,1561]" italics="true" pageId="7" pageNumber="368">et al</emphasis>
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. 2012
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</bibRefCitation>
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:42
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||
</treatmentCitation>
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(species list).
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<typeStatus id="54C68822EC74FFACF8909E73FEABF9BA" box="[151,260,1606,1631]" pageId="7" pageNumber="368" type="holotype">Holotype</typeStatus>
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:
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<specimenCode id="DBDB9EFBEC74FFACF9149E72FE17F9BA" box="[275,440,1606,1631]" collectionCode="MUSM" pageId="7" pageNumber="368">
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MUSM
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,
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mm SL, reservoir
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, middle
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,
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basin,
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<collectingCountry id="F36A7610EC74FFACFCBA9E72FB5BF9BB" box="[1213,1268,1607,1630]" name="Peru" pageId="7" pageNumber="368">Peru</collectingCountry>
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<typeStatus id="54C68822EC74FFACF8C09E5FFE9BF966" box="[199,308,1642,1667]" pageId="7" pageNumber="368" type="paratype">Paratype</typeStatus>
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</emphasis>
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:
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<specimenCode id="DBDB9EFBEC74FFACF9439E5FFE47F966" box="[324,488,1642,1667]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="7" pageNumber="368">ANSP 282226</specimenCode>
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(
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SL), the same locality of the
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<typeStatus id="54C68822EC74FFACFBCE9E5FFB81F966" box="[969,1070,1642,1667]" pageId="7" pageNumber="368" type="holotype">holotype</typeStatus>
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;
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<specimenCode id="DBDB9EFBEC74FFACFC3C9E5FFB78F966" box="[1083,1239,1642,1667]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="7" pageNumber="368">MPEG 35976</specimenCode>
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(2, 209–
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SL), the same locality of the
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|
||
<typeStatus id="54C68822EC74FFACF9E09EBBFDE3F942" box="[487,588,1678,1703]" pageId="7" pageNumber="368" type="holotype">holotype</typeStatus>
|
||
;
|
||
<specimenCode id="DBDB9EFBEC74FFACFA5E9EBAFCBCF942" box="[601,787,1679,1703]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="7" pageNumber="368">MZUSP 123828</specimenCode>
|
||
(2, 252–
|
||
<quantity id="4C859B65EC74FFACFB7B9EBBFC75F943" box="[892,986,1678,1703]" metricMagnitude="-1" metricUnit="m" metricValue="2.92" pageId="7" pageNumber="368" unit="mm" value="292.0">292 mm</quantity>
|
||
SL). the same locality of the
|
||
</materialsCitation>
|
||
<materialsCitation id="3B153CDDEC74FFACFD349EBBFC97F92E" ID-GBIF-Occurrence="2234225257" collectionCode="MUSM" pageId="7" pageNumber="368" specimenCode="MUSM 48796" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus id="54C68822EC74FFACFD349EBBFA37F942" box="[1331,1432,1678,1703]" pageId="7" pageNumber="368" type="holotype">holotype</typeStatus>
|
||
;
|
||
<specimenCode id="DBDB9EFBEC74FFACF8909E87FE96F92E" box="[151,313,1714,1739]" collectionCode="MUSM" pageId="7" pageNumber="368">MUSM 48796</specimenCode>
|
||
(5, 155–
|
||
<quantity id="4C859B65EC74FFACF99D9E87FE57F92F" box="[410,504,1714,1739]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" pageId="7" pageNumber="368" unit="mm" value="250.0">250 mm</quantity>
|
||
SL), the same locality of the
|
||
</materialsCitation>
|
||
<materialsCitation id="3B153CDDEC74FFACFB399E87FE30F90A" ID-GBIF-Occurrence="2234225261" collectingDate="2018-07-02" collectionCode="USNM" pageId="7" pageNumber="368" specimenCount="2" typeStatus="holotype">
|
||
<typeStatus id="54C68822EC74FFACFB399E87FC0CF92E" box="[830,931,1714,1739]" pageId="7" pageNumber="368" type="holotype">holotype</typeStatus>
|
||
;
|
||
<collectionCode id="ED6CAE45EC74FFACFBA99E87FC50F92F" box="[942,1023,1714,1738]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="368" type="Museum">USNM</collectionCode>
|
||
xxxx (
|
||
<date id="FFC31040EC74FFACFC4A9E87FB3EF92E" box="[1101,1169,1714,1739]" pageId="7" pageNumber="368" value="2018-07-02">
|
||
<collectingDate id="EF87E9A8EC74FFACFC4A9E87FB3EF92E" box="[1101,1169,1714,1739]" pageId="7" pageNumber="368" value="2018-07-02">
|
||
2, 18
|
||
<quantity id="4C859B65EC74FFACFC839E87FB3EF92E" box="[1156,1169,1714,1739]" metricMagnitude="-1" metricUnit="m" metricValue="1.065" metricValueMax="2.06" metricValueMin="0.06999999999999999" pageId="7" pageNumber="368" unit="mm" value="106.5" valueMax="206.0" valueMin="7.0">7</quantity>
|
||
</collectingDate>
|
||
</date>
|
||
–206 mm SL), the same locality of the
|
||
<typeStatus id="54C68822EC74FFACF93E9EE3FE30F90A" box="[313,415,1750,1775]" pageId="7" pageNumber="368" type="holotype">holotype</typeStatus>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C367650BEC74FFACF8C09ECFFDD6F822" pageId="7" pageNumber="368" type="diagnosis">
|
||
<paragraph id="8BC23680EC74FFACF8C09ECFFDD6F822" blockId="7.[151,1437,1606,1991]" pageId="7" pageNumber="368">
|
||
<emphasis id="B909EA92EC74FFACF8C09ECFFE95F8F6" bold="true" box="[199,314,1786,1811]" pageId="7" pageNumber="368">Diagnosis</emphasis>
|
||
.
|
||
<taxonomicName id="4C7D4D03EC74FFACF9489ECFFDF4F8F7" authorityName="Marceniuk & Marchena & Oliveira & Betancur-R" authorityYear="2019" box="[335,603,1786,1810]" class="Actinopterygii" family="Ariidae" genus="Chinchaysuyoa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="368" phylum="Chordata" rank="species" species="ortegai">
|
||
<emphasis id="B909EA92EC74FFACF9489ECFFDF4F8F7" box="[335,603,1786,1810]" italics="true" pageId="7" pageNumber="368">Chinchaysuyoa ortegai</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C7D4D03EC74FFACFB029ECFFCD3F8F7" baseAuthorityName="Boulenger" baseAuthorityYear="1898" box="[773,892,1786,1810]" class="Actinopterygii" family="Ariidae" genus="Chinchaysuyoa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="368" phylum="Chordata" rank="species" species="labiata">
|
||
<emphasis id="B909EA92EC74FFACFB029ECFFCD3F8F7" box="[773,892,1786,1810]" italics="true" pageId="7" pageNumber="368">C. labiata</emphasis>
|
||
</taxonomicName>
|
||
by thicker granulation irregularly distributed granulation in the cephalic shield (
|
||
<emphasis id="B909EA92EC74FFACFA1A9F2AFD9BF8D3" box="[541,564,1823,1846]" italics="true" pageId="7" pageNumber="368">vs</emphasis>
|
||
. finer and homogeneously distributed in the cephalic shield, especially in the sphenotic, pterotic and parieto-supraoccipital,
|
||
<figureCitation id="13462A05EC74FFACFA999F77FD44F8BE" box="[670,747,1858,1883]" captionStart="FIGURE 5" captionStartId="9.[151,250,1410,1432]" captionTargetBox="[150,1436,409,1389]" captionTargetId="figure@9.[150,1436,409,1389]" captionTargetPageId="9" captionText="FIGURE 5. Head in dorsal view. Chinchaysuyoa labiata, (A) holotype 475 mm SL (MZUT 1540), and Chinchaysuyoa ortegai, (B) holotype 256 mm SL (MUSM 63800)." httpUri="https://zenodo.org/record/2622978/files/figure.png" pageId="7" pageNumber="368">Figs. 5</figureCitation>
|
||
and
|
||
<figureCitation id="13462A05EC74FFACFB269F77FC9EF8BE" box="[801,817,1858,1883]" captionStart="FIGURE 6" captionStartId="10.[151,250,1977,1999]" captionTargetBox="[150,1436,589,1955]" captionTargetId="figure@10.[150,1436,589,1955]" captionTargetPageId="10" captionText="FIGURE 6. Dorsal view of the neurocranium and the nuchal plate of Chinchaysuyoa labiata, (A) paratype 113 mm SL (CAS 60621), and Chinchaysuyoa ortegai, (B) paratype 155 mm SL (MPEG 35976)." httpUri="https://zenodo.org/record/2622980/files/figure.png" pageId="7" pageNumber="368">6</figureCitation>
|
||
), snout truncated anteriorly (
|
||
<emphasis id="B909EA92EC74FFACFC719F76FB22F8BF" box="[1142,1165,1859,1882]" italics="true" pageId="7" pageNumber="368">vs</emphasis>
|
||
. pointed anteriorly,
|
||
<figureCitation id="13462A05EC74FFACFD779F77FF09F89A" captionStart="FIGURE 5" captionStartId="9.[151,250,1410,1432]" captionTargetBox="[150,1436,409,1389]" captionTargetId="figure@9.[150,1436,409,1389]" captionTargetPageId="9" captionText="FIGURE 5. Head in dorsal view. Chinchaysuyoa labiata, (A) holotype 475 mm SL (MZUT 1540), and Chinchaysuyoa ortegai, (B) holotype 256 mm SL (MUSM 63800)." httpUri="https://zenodo.org/record/2622978/files/figure.png" pageId="7" pageNumber="368">Fig. 5</figureCitation>
|
||
), distance between anterior nostrils 6.3–7.6% SL (
|
||
<emphasis id="B909EA92EC74FFACFAE59F52FD56F89B" box="[738,761,1895,1918]" italics="true" pageId="7" pageNumber="368">vs</emphasis>
|
||
. 5.1–5.9% SL,
|
||
<tableCitation id="C6FF033BEC74FFACFBAC9F53FC52F89A" box="[939,1021,1894,1919]" captionStart="TABLE 1" captionStartId="13.[151,239,332,355]" captionText="TABLE 1. Morphometric data for Chinchaysuyoa labiata and C. ortegai. Standard length is expressed in millimeters and all other measurements expressed in percents of standard length." pageId="7" pageNumber="368">Table 1</tableCitation>
|
||
), distance between posterior nostrils 6.4–7.5% SL, rarely less than 6.4 (
|
||
<emphasis id="B909EA92EC74FFACFA259FBEFD96F847" box="[546,569,1931,1954]" italics="true" pageId="7" pageNumber="368">vs</emphasis>
|
||
. 5.2–6.2% SL,
|
||
<tableCitation id="C6FF033BEC74FFACFAEC9FBFFC91F847" box="[747,830,1930,1955]" captionStart="TABLE 1" captionStartId="13.[151,239,332,355]" captionText="TABLE 1. Morphometric data for Chinchaysuyoa labiata and C. ortegai. Standard length is expressed in millimeters and all other measurements expressed in percents of standard length." pageId="7" pageNumber="368">Table 1</tableCitation>
|
||
), caudal–fin lobes relatively, rounded posteriorly (
|
||
<emphasis id="B909EA92EC74FFACFD7A9FBEFA3BF847" box="[1405,1428,1931,1954]" italics="true" pageId="7" pageNumber="368">vs</emphasis>
|
||
. relatively long, pointed posteriorly,
|
||
<figureCitation id="13462A05EC74FFACFA2F9F9BFDC4F822" box="[552,619,1966,1991]" captionStart="FIGURE 5" captionStartId="9.[151,250,1410,1432]" captionTargetBox="[150,1436,409,1389]" captionTargetId="figure@9.[150,1436,409,1389]" captionTargetPageId="9" captionText="FIGURE 5. Head in dorsal view. Chinchaysuyoa labiata, (A) holotype 475 mm SL (MZUT 1540), and Chinchaysuyoa ortegai, (B) holotype 256 mm SL (MUSM 63800)." httpUri="https://zenodo.org/record/2622978/files/figure.png" pageId="7" pageNumber="368">Fig. 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF026608EC7BFFA3F8909EF2FDDBF8FE" httpUri="https://zenodo.org/record/2622976/files/figure.png" pageId="8" pageNumber="369" startId="8.[151,250,1735,1757]" targetBox="[150,1436,193,1714]" targetPageId="8">
|
||
<paragraph id="8BC23680EC7BFFA3F8909EF2FDDBF8FE" blockId="8.[151,1436,1735,1819]" pageId="8" pageNumber="369">
|
||
<emphasis id="B909EA92EC7BFFA3F8909EF2FEBAF939" bold="true" box="[151,277,1735,1757]" pageId="8" pageNumber="369">FIGURE 4.</emphasis>
|
||
Body in lateral view.
|
||
<taxonomicName id="4C7D4D03EC7BFFA3FA059EF2FD40F938" baseAuthorityName="Boulenger" baseAuthorityYear="1898" box="[514,751,1735,1757]" class="Actinopterygii" family="Ariidae" genus="Chinchaysuyoa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="369" phylum="Chordata" rank="species" species="labiata">
|
||
<emphasis id="B909EA92EC7BFFA3FA059EF2FD40F938" box="[514,751,1735,1757]" italics="true" pageId="8" pageNumber="369">Chinchaysuyoa labiata</emphasis>
|
||
</taxonomicName>
|
||
, (A) holotype 475 mm SL (MZUT 1540), (B)
|
||
<taxonomicName id="4C7D4D03EC7BFFA3FCE69EF2FF60F919" authorityName="Fisher & Eigenmann" authorityYear="1922" class="Actinopterygii" family="Ariidae" genus="Hexanematichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="369" phylum="Chordata" rank="species" species="henni">
|
||
<emphasis id="B909EA92EC7BFFA3FCE69EF2FF60F919" italics="true" pageId="8" pageNumber="369">Hexanematichthys henni</emphasis>
|
||
</taxonomicName>
|
||
, holotype 139 mm SL (CAS 60620), (C)
|
||
<taxonomicName id="4C7D4D03EC7BFFA3FA869ED3FCC2F919" authorityName="Marceniuk & Marchena & Oliveira & Betancur-R" authorityYear="2019" box="[641,877,1766,1788]" class="Actinopterygii" family="Ariidae" genus="Chinchaysuyoa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="369" phylum="Chordata" rank="species" species="ortegai">
|
||
<emphasis id="B909EA92EC7BFFA3FA869ED3FCC2F919" box="[641,877,1766,1788]" italics="true" pageId="8" pageNumber="369">Chinchaysuyoa ortegai</emphasis>
|
||
</taxonomicName>
|
||
, holotype 256 mm SL (MUSM 63800), and (D) fresh non type specimen, not cataloged, 315 mm SL.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C367650BEC7BFFA1F8C09F72FE25FE6D" lastPageId="10" lastPageNumber="371" pageId="8" pageNumber="369" type="description">
|
||
<paragraph id="8BC23680EC7BFFA2F8C09F72FD6FFE80" blockId="8.[151,1437,1863,2032]" lastBlockId="9.[151,1436,151,357]" lastPageId="9" lastPageNumber="370" pageId="8" pageNumber="369">
|
||
<emphasis id="B909EA92EC7BFFA3F8C09F72FEFEF885" bold="true" box="[199,337,1863,1888]" pageId="8" pageNumber="369">Description</emphasis>
|
||
. Morphometric data in
|
||
<tableCitation id="C6FF033BEC7BFFA3FA5F9F72FD04F885" box="[600,683,1863,1888]" captionStart="TABLE 1" captionStartId="13.[151,239,332,355]" captionText="TABLE 1. Morphometric data for Chinchaysuyoa labiata and C. ortegai. Standard length is expressed in millimeters and all other measurements expressed in percents of standard length." pageId="8" pageNumber="369">Table 1</tableCitation>
|
||
. Head depressed, profile elevated posteriorly, straight at frontal and parieto-supraoccipital area. Snout long and previously truncated. Anterior nostril round, with fleshy edge, posterior nostril covered by flap of skin; nostrils distant to one another and distant to orbit, not connected by fleshy furrow. Eye lateral and large; eyes distant to one another. Three pairs of relatively long teretiform barbels; maxillary barbel usually reaching or past reaching gill membrane edge, mental barbels not reaching gill membrane edge. Osseous bridge formed by lateral ethmoid and frontal relatively long and slender, little evident under skin. Cephalic shield exposed, very rough and granulated; long and large on lateral ethmoid, frontal, and supracleithral areas. Anterior portion of dorsomedial groove of neurocranium fleshy, conspicuous and continuous to level of posterior nares; posterior portion of osseous groove, deep and conspicuous, with straight margins, narrowing posteriorly. Parietosupraoccipital process funnel-shaped, long and moderately wide on posterior portion, and profile straight. Nuchal plate crescent-shaped, long and moderately wide.
|
||
</paragraph>
|
||
<caption id="DF026608EC7AFFA2F8909DB7FD33FA52" httpUri="https://zenodo.org/record/2622978/files/figure.png" pageId="9" pageNumber="370" startId="9.[151,250,1410,1432]" targetBox="[150,1436,409,1389]" targetPageId="9">
|
||
<paragraph id="8BC23680EC7AFFA2F8909DB7FD33FA52" blockId="9.[151,1436,1410,1463]" pageId="9" pageNumber="370">
|
||
<emphasis id="B909EA92EC7AFFA2F8909DB7FEB8FA72" bold="true" box="[151,279,1410,1432]" pageId="9" pageNumber="370">FIGURE 5.</emphasis>
|
||
Head in dorsal view.
|
||
<taxonomicName id="4C7D4D03EC7AFFA2FA0C9DB7FD55FA7D" baseAuthorityName="Boulenger" baseAuthorityYear="1898" box="[523,762,1410,1432]" class="Actinopterygii" family="Ariidae" genus="Chinchaysuyoa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="370" phylum="Chordata" rank="species" species="labiata">
|
||
<emphasis id="B909EA92EC7AFFA2FA0C9DB7FD55FA7D" box="[523,762,1410,1432]" italics="true" pageId="9" pageNumber="370">Chinchaysuyoa labiata</emphasis>
|
||
</taxonomicName>
|
||
, (A) holotype 475 mm SL (MZUT 1540), and
|
||
<taxonomicName id="4C7D4D03EC7AFFA2FD079DB7FF4FFA52" authorityName="Marceniuk & Marchena & Oliveira & Betancur-R" authorityYear="2019" class="Actinopterygii" family="Ariidae" genus="Chinchaysuyoa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="370" phylum="Chordata" rank="species" species="ortegai">
|
||
<emphasis id="B909EA92EC7AFFA2FD079DB7FF4FFA52" italics="true" pageId="9" pageNumber="370">Chinchaysuyoa ortegai</emphasis>
|
||
</taxonomicName>
|
||
, (B) holotype 256 mm SL (MUSM 63800).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC23680EC7AFFA2F8C09DEBFEE7F92A" blockId="9.[151,1437,1502,2031]" pageId="9" pageNumber="370">Mouth subterminal to terminal, moderately large; lower jaw arched. Lips relatively thick, upper lip more thick than lower lip. Vomerine tooth plates absent. One pair of oval shaped accessory tooth plates, very small and transversely elongate and narrow, distant one another, with sharp teeth. Premaxilla with sharp teeth. Dentary separated at midline, with sharp teeth. Gill membranes fused, attached to isthmus. Eighteen to twenty acicular gill rakers on first arch, 20–23 spike-shaped gill rakers on second arch. Mesial surfaces of all gill arches with developed gill rakers, lateral and mesial surfaces of first and second gill arches lacking fleshy papillae intercalated with gill rakers.</paragraph>
|
||
<paragraph id="8BC23680EC7AFFA1F8C09EEFFE8CFEF8" blockId="9.[151,1437,1502,2031]" lastBlockId="10.[151,1437,151,536]" lastPageId="10" lastPageNumber="371" pageId="9" pageNumber="370">Body width wider than depth in pectoral-girdle area, progressively more compressed from pectoral fin to caudal peduncle. Lateral line sloping ventrally on anterior one-third, extending posteriorly to caudal peduncle, bending abruptly onto dorsal lobe of caudal fin. Dorsal-fin spine moderately long and thick; anterior margin with granules on basal two-thirds and distal one-third with serrations; posterior margin smooth on basal half and distal half with serrations. Seven soft dorsal-fin rays. Pectoral-fin spine moderately long, shorter than dorsal-fin spine; anterior margin with granules on basal two-thirds and distal one-third with short serrations; posterior margin with granules on basal, distal half with serrations. Ten to eleven soft pectoral-fin rays. Posterior cleithral process exposed, rough and triangular shaped, very long and pointed posteriorly. Pelvic fin deep and moderately long at base, with six rays. Adipose-fin base relatively long, its base about half as long as the anal-fin base. Anal fin deep and relatively long at base, with 19–20 rays and distal margin slightly concave. Caudal peduncle relatively deep. Caudal fin forked, dorsal and ventral lobes moderately long, wide and rounded posteriorly; dorsal lobe longer than ventral lobe.</paragraph>
|
||
<paragraph id="8BC23680EC79FFA1F8C09912FE25FE6D" blockId="10.[151,1437,151,536]" pageId="10" pageNumber="371">
|
||
<emphasis id="B909EA92EC79FFA1F8C09912FEE7FEA5" bold="true" box="[199,328,295,320]" pageId="10" pageNumber="371">Coloration</emphasis>
|
||
. Grayish to dark brown on dorsum, silvery on flanks and white on abdomen. In alcohol, dark brown on dorsum, flanks brownish to silvery and white on venter; fins black. Maxillary barbel dark, mental barbel lighter (
|
||
<figureCitation id="13462A05EC79FFA1F8E8995AFECFFE6D" box="[239,352,367,392]" captionStart="FIGURE 4" captionStartId="8.[151,250,1735,1757]" captionTargetBox="[150,1436,193,1714]" captionTargetId="figure@8.[150,1436,193,1714]" captionTargetPageId="8" captionText="FIGURE 4. Body in lateral view. Chinchaysuyoa labiata, (A) holotype 475 mm SL (MZUT 1540), (B) Hexanematichthys henni, holotype 139 mm SL (CAS 60620), (C) Chinchaysuyoa ortegai, holotype 256 mm SL (MUSM 63800), and (D) fresh non type specimen, not cataloged, 315 mm SL." httpUri="https://zenodo.org/record/2622976/files/figure.png" pageId="10" pageNumber="371">Figs. 4a,b</figureCitation>
|
||
,
|
||
<figureCitation id="13462A05EC79FFA1F96B995AFED3FE6D" box="[364,380,367,392]" captionStart="FIGURE 5" captionStartId="9.[151,250,1410,1432]" captionTargetBox="[150,1436,409,1389]" captionTargetId="figure@9.[150,1436,409,1389]" captionTargetPageId="9" captionText="FIGURE 5. Head in dorsal view. Chinchaysuyoa labiata, (A) holotype 475 mm SL (MZUT 1540), and Chinchaysuyoa ortegai, (B) holotype 256 mm SL (MUSM 63800)." httpUri="https://zenodo.org/record/2622978/files/figure.png" pageId="10" pageNumber="371">5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C367650BEC79FFA1F8C099A1FBDCFE35" pageId="10" pageNumber="371" type="distribution">
|
||
<paragraph id="8BC23680EC79FFA1F8C099A1FBDCFE35" blockId="10.[151,1437,151,536]" pageId="10" pageNumber="371">
|
||
<emphasis id="B909EA92EC79FFA1F8C099A1FE47FE48" bold="true" box="[199,488,404,429]" pageId="10" pageNumber="371">Distribution and habitat</emphasis>
|
||
.
|
||
<taxonomicName id="4C7D4D03EC79FFA1F9F199A1FD55FE49" authorityName="Marceniuk & Marchena & Oliveira & Betancur-R" authorityYear="2019" box="[502,762,404,428]" class="Actinopterygii" family="Ariidae" genus="Chinchaysuyoa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="10" pageNumber="371" phylum="Chordata" rank="species" species="ortegai">
|
||
<emphasis id="B909EA92EC79FFA1F9F199A1FD55FE49" box="[502,762,404,428]" italics="true" pageId="10" pageNumber="371">Chinchaysuyoa ortegai</emphasis>
|
||
</taxonomicName>
|
||
is described to Poechos reservoir, in the middle Rio Chira in
|
||
<collectingCountry id="F36A7610EC79FFA1F890998DFF62FE2A" box="[151,205,440,463]" name="Peru" pageId="10" pageNumber="371">Peru</collectingCountry>
|
||
, Catamayo Basin, that born in the Andes Occidental Cordillera in
|
||
<collectingCountry id="F36A7610EC79FFA1FBB39982FBBEFE35" box="[948,1041,439,464]" name="Ecuador" pageId="10" pageNumber="371">Ecuador</collectingCountry>
|
||
(
|
||
<figureCitation id="13462A05EC79FFA1FC269982FBCAFE35" box="[1057,1125,439,464]" captionStart="FIGURE 8" captionStartId="12.[151,250,1381,1403]" captionTargetBox="[151,1435,193,1360]" captionTargetId="figure@12.[151,1435,193,1360]" captionTargetPageId="12" captionText="FIGURE 8. Sampling localities of the material examined in the present study, showing the geographic distribution of Chinchaysuyoa labiata, holotype (light blue circle), holotype and paratype of Hexanematichthys henni (red and pink circles, respectively), non–type specimen (yellow circle), and Chinchaysuyoa ortegai, type-specimens (green circle)." httpUri="https://zenodo.org/record/2622984/files/figure.png" pageId="10" pageNumber="371">Fig. 8</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C367650BEC79FFA1F8D699E9FE43FDF2" pageId="10" pageNumber="371" type="etymology">
|
||
<paragraph id="8BC23680EC79FFA1F8D699E9FE43FDF2" blockId="10.[151,1437,151,536]" pageId="10" pageNumber="371">
|
||
<emphasis id="B909EA92EC79FFA1F8D699E9FEFFFE10" bold="true" box="[209,336,476,501]" pageId="10" pageNumber="371">Etymology</emphasis>
|
||
. The name of the species is in honor of Dr. Hernán Ortega Torres Teofilo the most important ichthyologist working in
|
||
<collectingCountry id="F36A7610EC79FFA1F9B69A35FE47FDF2" box="[433,488,512,535]" name="Peru" pageId="10" pageNumber="371">Peru</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF026608EC79FFA1F8909F8CFC1AF808" httpUri="https://zenodo.org/record/2622980/files/figure.png" pageId="10" pageNumber="371" startId="10.[151,250,1977,1999]" targetBox="[150,1436,589,1955]" targetPageId="10">
|
||
<paragraph id="8BC23680EC79FFA1F8909F8CFC1AF808" blockId="10.[151,1436,1977,2030]" pageId="10" pageNumber="371">
|
||
<emphasis id="B909EA92EC79FFA1F8909F8CFEA2F82B" bold="true" box="[151,269,1977,1999]" pageId="10" pageNumber="371">FIGURE 6</emphasis>
|
||
. Dorsal view of the neurocranium and the nuchal plate of
|
||
<taxonomicName id="4C7D4D03EC79FFA1FB649F8CFBE1F82A" baseAuthorityName="Boulenger" baseAuthorityYear="1898" box="[867,1102,1977,1999]" class="Actinopterygii" family="Ariidae" genus="Chinchaysuyoa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="10" pageNumber="371" phylum="Chordata" rank="species" species="labiata">
|
||
<emphasis id="B909EA92EC79FFA1FB649F8CFBE1F82A" box="[867,1102,1977,1999]" italics="true" pageId="10" pageNumber="371">Chinchaysuyoa labiata</emphasis>
|
||
</taxonomicName>
|
||
, (A) paratype 113 mm SL (CAS 60621), and
|
||
<taxonomicName id="4C7D4D03EC79FFA1F9119FE2FDAEF80B" authorityName="Marceniuk & Marchena & Oliveira & Betancur-R" authorityYear="2019" box="[278,513,2007,2030]" class="Actinopterygii" family="Ariidae" genus="Chinchaysuyoa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="10" pageNumber="371" phylum="Chordata" rank="species" species="ortegai">
|
||
<emphasis id="B909EA92EC79FFA1F9119FE2FDAEF80B" box="[278,513,2007,2030]" italics="true" pageId="10" pageNumber="371">Chinchaysuyoa ortegai</emphasis>
|
||
</taxonomicName>
|
||
, (B) paratype 155 mm SL (MPEG 35976).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C367650BEC78FFA0F8C998A2FE04FEA5" pageId="11" pageNumber="372" type="discussion">
|
||
<paragraph id="8BC23680EC78FFA0F8C998A2FE04FEA5" blockId="11.[151,1436,151,320]" pageId="11" pageNumber="372">
|
||
<emphasis id="B909EA92EC78FFA0F8C998A2FE94FF55" bold="true" box="[206,315,151,176]" pageId="11" pageNumber="372">Remarks</emphasis>
|
||
.
|
||
<taxonomicName id="4C7D4D03EC78FFA0F94E98ADFDE2FF4A" authorityName="Marceniuk & Marchena & Oliveira & Betancur-R" authorityYear="2019" box="[329,589,152,176]" class="Actinopterygii" family="Ariidae" genus="Chinchaysuyoa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="11" pageNumber="372" phylum="Chordata" rank="species" species="ortegai">
|
||
<emphasis id="B909EA92EC78FFA0F94E98ADFDE2FF4A" box="[329,589,152,176]" italics="true" pageId="11" pageNumber="372">Chinchaysuyoa ortegai</emphasis>
|
||
</taxonomicName>
|
||
can be differentiated from
|
||
<taxonomicName id="4C7D4D03EC78FFA0FB7998ADFC41FF55" baseAuthorityName="Boulenger" baseAuthorityYear="1898" box="[894,1006,152,176]" class="Actinopterygii" family="Ariidae" genus="Chinchaysuyoa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="11" pageNumber="372" phylum="Chordata" rank="species" species="labiata">
|
||
<emphasis id="B909EA92EC78FFA0FB7998ADFC41FF55" box="[894,1006,152,176]" italics="true" pageId="11" pageNumber="372">C. labiata</emphasis>
|
||
</taxonomicName>
|
||
based on its morphological characters (see Remarks on
|
||
<taxonomicName id="4C7D4D03EC78FFA0F9649889FE79FF31" baseAuthorityName="Boulenger" baseAuthorityYear="1898" box="[355,470,188,212]" class="Actinopterygii" family="Ariidae" genus="Chinchaysuyoa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="11" pageNumber="372" phylum="Chordata" rank="species" species="labiata">
|
||
<emphasis id="B909EA92EC78FFA0F9649889FE79FF31" box="[355,470,188,212]" italics="true" pageId="11" pageNumber="372">C. labiata</emphasis>
|
||
</taxonomicName>
|
||
), which is further reinforced by the disjunct geographic ranges of the two species, which are restricted to the drainage basins of
|
||
<collectingCountry id="F36A7610EC78FFA0FAAD98EAFCA8FF1D" box="[682,775,223,248]" name="Ecuador" pageId="11" pageNumber="372">Ecuador</collectingCountry>
|
||
and
|
||
<collectingCountry id="F36A7610EC78FFA0FB4498D5FCD6FF12" box="[835,889,224,247]" name="Peru" pageId="11" pageNumber="372">Peru</collectingCountry>
|
||
, respectively (
|
||
<figureCitation id="13462A05EC78FFA0FC2798EAFBC9FF1D" box="[1056,1126,223,248]" captionStart="FIGURE 8" captionStartId="12.[151,250,1381,1403]" captionTargetBox="[151,1435,193,1360]" captionTargetId="figure@12.[151,1435,193,1360]" captionTargetPageId="12" captionText="FIGURE 8. Sampling localities of the material examined in the present study, showing the geographic distribution of Chinchaysuyoa labiata, holotype (light blue circle), holotype and paratype of Hexanematichthys henni (red and pink circles, respectively), non–type specimen (yellow circle), and Chinchaysuyoa ortegai, type-specimens (green circle)." httpUri="https://zenodo.org/record/2622984/files/figure.png" pageId="11" pageNumber="372">Fig. 8</figureCitation>
|
||
), while the molecular data support the recognition of the species as distinct from the other genera of the family recorded on the Pacific coast of the Americas (
|
||
<figureCitation id="13462A05EC78FFA0F95E9912FE35FEA5" box="[345,410,295,320]" captionStart="FIGURE 1" captionStartId="5.[151,250,1948,1970]" captionTargetBox="[208,1344,214,1925]" captionTargetId="figure@5.[197,1397,195,1935]" captionTargetPageId="5" captionText="FIGURE 1. Phylogenetic tree inferred from combined evidence from molecular and morphological data based on Parsimony Analysis (A), Bayesian inference (B) and Maximum Likelihood (C), with bootstrap values greater than 50%. In red, neotropical freshwater species, included in Potamarius sensu Marceniuk & Menezes (2007)." httpUri="https://zenodo.org/record/2622969/files/figure.png" pageId="11" pageNumber="372">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |