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<document id="FA92C14ECF0F91E84960CE2AA928D80C" ID-ISSN="1175­5334" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629612136760" checkinUser="felipe" docAuthor="Ballantyne, Lesley A. &amp; Lambkin, Christine" docDate="2009" docId="0394D665BE37FFA1FF3C53AC22D4ED9A" docLanguage="en" docName="zt01997p188.pdf" docOrigin="Zootaxa 1997" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Aquilonia Ballantyne &amp; Lambkin 2009, gen. n." docType="treatment" docVersion="3" lastPageNumber="30" masterDocId="FFADAE1DBE2CFFBCFFAB57492227EE30" masterDocTitle="Systematics of Indo-Pacific fireflies with a redefinition of Australasian Atyphella Olliff, Madagascan Photuroluciola Pic, and description of seven new genera from the Luciolinae (Coleoptera: Lampyridae)" masterLastPageNumber="188" masterPageNumber="1" pageNumber="28" updateTime="1699083011144" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="2F9C32031FBD8A664937590EC4E51E03">Systematics of Indo-Pacific fireflies with a redefinition of Australasian Atyphella Olliff, Madagascan Photuroluciola Pic, and description of seven new genera from the Luciolinae (Coleoptera: Lampyridae)</mods:title>
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<mods:namePart id="1D18B7AC9CB03E2874F55D4859CB4209">Ballantyne, Lesley A.</mods:namePart>
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<mods:namePart id="77BB776B7220A26F7779BBBC18CB2D91">Lambkin, Christine</mods:namePart>
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<subSubSection id="C32734F8BE37FFA7FF3C53AC234EEACF" box="[151,361,1253,1279]" pageId="27" pageNumber="28" type="nomenclature">
<paragraph id="8B826773BE37FFA7FF3C53AC234EEACF" blockId="27.[151,361,1253,1314]" box="[151,361,1253,1279]" pageId="27" pageNumber="28">
<heading id="D0CAD01FBE37FFA7FF3C53AC234EEACF" bold="true" box="[151,361,1253,1279]" fontSize="11" level="1" pageId="27" pageNumber="28" reason="1">
<emphasis id="B949BB61BE37FFA7FF3C53AC234EEACF" bold="true" box="[151,361,1253,1279]" pageId="27" pageNumber="28">
<taxonomicName id="4C3D1CF0BE37FFA7FF3C53AC2328EACF" ID-CoL="ZHV" authority="Ballantyne &amp; Lambkin, 2009" authorityName="Ballantyne &amp; Lambkin" authorityYear="2009" box="[151,271,1253,1279]" class="Insecta" family="Lampyridae" genus="Aquilonia" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B949BB61BE37FFA7FF3C53AC2328EACF" bold="true" box="[151,271,1253,1279]" italics="true" pageId="27" pageNumber="28">Aquilonia</emphasis>
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</emphasis>
</heading>
</paragraph>
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<subSubSection id="C32734F8BE37FFA7FF3C524122C8EB12" box="[151,239,1288,1314]" pageId="27" pageNumber="28" type="description">
<paragraph id="8B826773BE37FFA7FF3C524122C8EB12" blockId="27.[151,361,1253,1314]" box="[151,239,1288,1314]" pageId="27" pageNumber="28">
(
<figureCitation id="13067BF6BE37FFA7FF34524122C2EB12" box="[159,229,1288,1314]" captionStart="FIGURES 414" captionStartId="146.[151,269,1910,1934]" captionTargetBox="[211,1379,194,1877]" captionTargetId="figure-81@146.[211,1380,194,1877]" captionTargetPageId="146" captionText="FIGURES 414. Habitus Genera 1. Aquilonia costata male dorsal (Katherine NT, ANIC). 5, 6, 8, Asymmetricata circumdata, 5, 6 males dorsal ventral, 8 female ventral (Thailand, ANIC). 9, 10 Atyphella atra males dorsal, ventral (9 paratype National Park Q; 10 Springbrook Q, ANIC). Atyphella lychnus male dorsal (Lansdowne State Forest, ANIC). Convexa wolfi males 12, 13 dorsal, ventral, 14 female ventral (Kukugai, BPBM)." pageId="27" pageNumber="28">Fig. 4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32734F8BE37FFA7FF3C521E20D8EB42" box="[151,767,1367,1394]" pageId="27" pageNumber="28" type="type_taxon">
<paragraph id="8B826773BE37FFA7FF3C521E20D8EB42" blockId="27.[151,1436,1367,2034]" box="[151,767,1367,1394]" pageId="27" pageNumber="28">
<emphasis id="B949BB61BE37FFA7FF3C521E231CEB41" bold="true" box="[151,315,1367,1393]" pageId="27" pageNumber="28">
<typeStatus id="5486D9D1BE37FFA7FF3C521E22F3EB41" box="[151,212,1367,1393]" pageId="27" pageNumber="28">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="4C3D1CF0BE37FFA7FEE852112072EB42" authority="(Lea)" baseAuthorityName="Lea" box="[323,597,1368,1394]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="costata">
<emphasis id="B949BB61BE37FFA7FEE852112028EB41" box="[323,527,1368,1393]" italics="true" pageId="27" pageNumber="28">Atyphella costata</emphasis>
(Lea)
</taxonomicName>
by monotypy.
</paragraph>
</subSubSection>
<subSubSection id="C32734F8BE37FFA1FF6C523623EFEC5A" lastPageId="29" lastPageNumber="30" pageId="27" pageNumber="28" type="diagnosis">
<paragraph id="8B826773BE37FFA7FF6C523626D0E882" blockId="27.[151,1436,1367,2034]" pageId="27" pageNumber="28">
<emphasis id="B949BB61BE37FFA7FF6C52362318EBA9" bold="true" box="[199,319,1407,1433]" pageId="27" pageNumber="28">Diagnosis</emphasis>
.
<taxonomicName id="4C3D1CF0BE37FFA7FEE552C923E6EBA9" authorityName="Ballantyne &amp; Lambkin" authorityYear="2009" box="[334,449,1408,1433]" class="Insecta" family="Lampyridae" genus="Aquilonia" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B949BB61BE37FFA7FEE552C923E6EBA9" box="[334,449,1408,1433]" italics="true" pageId="27" pageNumber="28">Aquilonia</emphasis>
</taxonomicName>
is distinguished most obviously by its dorsal colour pattern, which is approached in the study area by two species of
<taxonomicName id="4C3D1CF0BE37FFA7FDBF52E120BBEBF1" authority="Ballantyne &amp; Lambkin, 2009" authorityName="Ballantyne &amp; Lambkin" authorityYear="2009" box="[532,668,1448,1473]" class="Insecta" family="Lampyridae" genus="Gilvainsula" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B949BB61BE37FFA7FDBF52E120BBEBF1" box="[532,668,1448,1473]" italics="true" pageId="27" pageNumber="28">Gilvainsula</emphasis>
</taxonomicName>
<emphasis id="B949BB61BE37FFA7FD0F52EE20D1EBF1" bold="true" box="[676,758,1447,1473]" pageId="27" pageNumber="28">
<taxonomicNameLabel id="A27A061ABE37FFA7FD0F52EE20D1EBF1" box="[676,758,1447,1473]" pageId="27" pageNumber="28" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="4C3D1CF0BE37FFA7FC9D52E1261AEBF1" authority="Ballantyne &amp; Lambkin, 2009" authorityName="Ballantyne &amp; Lambkin" authorityYear="2009" box="[822,1085,1448,1473]" class="Insecta" family="Lampyridae" genus="Pygatyphella" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="russellia" status="sp. nov.">
<emphasis id="B949BB61BE37FFA7FC9D52E1261AEBF1" box="[822,1085,1448,1473]" italics="true" pageId="27" pageNumber="28">Pygatyphella russellia</emphasis>
</taxonomicName>
<emphasis id="B949BB61BE37FFA7FBEF52EE26A0EBF1" bold="true" box="[1092,1159,1447,1473]" pageId="27" pageNumber="28">
<taxonomicNameLabel id="A27A061ABE37FFA7FBEF52EE26A0EBF1" box="[1092,1159,1447,1473]" pageId="27" pageNumber="28" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
It differs from
<taxonomicName id="4C3D1CF0BE37FFA7FA9152E122EFEBD9" authority="Ballantyne &amp; Lambkin, 2009" authorityName="Ballantyne &amp; Lambkin" authorityYear="2009" class="Insecta" family="Lampyridae" genus="Gilvainsula" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B949BB61BE37FFA7FA9152E122EFEBD9" italics="true" pageId="27" pageNumber="28">Gilvainsula</emphasis>
</taxonomicName>
<emphasis id="B949BB61BE37FFA7FF7952862300EBD9" bold="true" box="[210,295,1487,1513]" pageId="27" pageNumber="28">
<taxonomicNameLabel id="A27A061ABE37FFA7FF7952862300EBD9" box="[210,295,1487,1513]" pageId="27" pageNumber="28" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
by the parallel-sided elytra and pronotum, and the toothed apical segment of labial palpi, and from
<emphasis id="B949BB61BE37FFA7FF7252B123B5E821" box="[217,402,1528,1553]" italics="true" pageId="27" pageNumber="28">
Pygat.
<taxonomicName id="4C3D1CF0BE37FFA7FE8652B123B5E821" authorityName="Ballantyne &amp; Lambkin" authorityYear="2009" box="[301,402,1528,1553]" class="Insecta" family="Lampyridae" genus="Pygatyphella" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="russellia">russellia</taxonomicName>
</emphasis>
by the absence of an arched V7 posterior to the LO, and tergite 8 lacking any median ridges. It differs from most
<taxonomicName id="4C3D1CF0BE37FFA7FE725169206EE809" authorityName="Olliff, Madagascan" authorityYear="1890" box="[473,585,1568,1593]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B949BB61BE37FFA7FE725169206EE809" box="[473,585,1568,1593]" italics="true" pageId="27" pageNumber="28">Atyphella</emphasis>
</taxonomicName>
by its distinctive dorsal colour, the subparallel-sided pronotum, pronotal width subequal to that of elytral humerus, [two features it shares with
<taxonomicName id="4C3D1CF0BE37FFA7FC6251012651E851" authorityName="Ballantyne &amp; Lambkin" authorityYear="2009" baseAuthorityName="Ballantyne &amp; Lambkin" baseAuthorityYear="2009" box="[969,1142,1608,1633]" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="inconspicua">
<emphasis id="B949BB61BE37FFA7FC6251012651E851" box="[969,1142,1608,1633]" italics="true" pageId="27" pageNumber="28">A. inconspicua</emphasis>
</taxonomicName>
], and it shares with most species of
<taxonomicName id="4C3D1CF0BE37FFA7FEBD513923AAE8B9" authority="Ballantyne &amp; Lambkin, 2009" authorityName="Ballantyne &amp; Lambkin" authorityYear="2009" box="[278,397,1648,1673]" class="Insecta" family="Lampyridae" genus="Lloydiella" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B949BB61BE37FFA7FEBD513923AAE8B9" box="[278,397,1648,1673]" italics="true" pageId="27" pageNumber="28">Lloydiella</emphasis>
</taxonomicName>
<emphasis id="B949BB61BE37FFA7FE3D512623CDE8B9" bold="true" box="[406,490,1647,1673]" pageId="27" pageNumber="28">
<taxonomicNameLabel id="A27A061ABE37FFA7FE3D512623CDE8B9" box="[406,490,1647,1673]" pageId="27" pageNumber="28" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
an aedeagal complex pattern with lateral lobes not divergent along their dorsal length and their inner preapical margins obliquely truncate, while differing in the colour pattern
</paragraph>
<paragraph id="8B826773BE37FFA7FF6C51F62004E9C2" blockId="27.[151,1436,1367,2034]" pageId="27" pageNumber="28">
<emphasis id="B949BB61BE37FFA7FF6C51F6232AE8E9" bold="true" box="[199,269,1727,1753]" pageId="27" pageNumber="28">Male.</emphasis>
Pronotum (
<bibRefCitation id="EFAC1A82BE37FFA7FE32518920D5E8EA" author="Ballantyne, L. A. &amp; Lambkin, C." box="[409,754,1728,1754]" pageId="27" pageNumber="28" pagination="15 - 93" refId="ref103088" refString="Ballantyne, L. A. &amp; Lambkin, C. (2000) Lampyridae of Australia (Coleoptera: Lampyridae: Luciolinae: Luciolini). Memoirs of the Queensland Museum 46 (1), 15 - 93." type="journal article" year="2000">Ballantyne &amp; Lambkin 2000</bibRefCitation>
:
<figureCitation id="13067BF6BE37FFA7FD5E51892160E8EA" box="[757,839,1728,1754]" captionStart="FIGURE 1" captionStartId="20.[151,255,1948,1972]" captionTargetBox="[159,1426,193,1918]" captionTargetId="figure-43@20.[151,1436,183,1922]" captionTargetPageId="20" captionText="FIGURE 1. Strict consensus of 739 most parsimonious trees of length 3387 from phylogenetic analysis of 112 taxa and 343 morphological characters in Table 1." pageId="27" pageNumber="28">Fig. 1h</figureCitation>
) dorsal surface lacking irregularities in posterolateral areas and longitudinal groove in lateral areas; punctation dense. Anterior margin not explanate. Pronotum subparallel-sided; pronotal width subequal to humeral width. Anterolateral corners rounded obtuse; lateral margins in anterior half subparallel-sided; lateral margins in posterior half subparallel-sided; lateral margins lacking indentation at mid-point, or sinuousity in either horizontal or vertical plane; subparallel-sided margins straight; lacking indentation in lateral margin near posterolateral corner, and irregularities at corner; posterolateral corners rounded obtuse; posterolateral corners project as far as median posterior margin and separated from it by shallow emarginations.
</paragraph>
<paragraph id="8B826773BE30FFA0FF6C57D1271AEEEA" blockId="28.[151,1437,152,2018]" pageId="28" pageNumber="29">Hypomera closed. Median area of hypomeron not elevated in vertical direction; anterior area of hypomeron not flat, posterior area flat, closely adpressed or not; pronotal width/ GHW index 1.41.5.</paragraph>
<paragraph id="8B826773BE30FFA0FF6C57A12779EFFA" blockId="28.[151,1437,152,2018]" pageId="28" pageNumber="29">Elytron punctation dense, not linear, not as large as that of pronotum, nor widely and evenly spaced; apices not deflexed; epipleuron and suture extend beyond mid-point, almost to apex but not as ridge around apex, neither thickened in apical half; 2 interstitial lines (1, 2) not exceeding suture; elytral carina absent; in horizontal specimen viewed from below epipleuron at elytral base wide, covering humerus, viewed from above epipleuron arises anterior to posterior margin of MS; epipleuron developed as a lateral ridge along most of length; sutural margins approximate along most of length in closed elytra; lateral margins parallel-sided.</paragraph>
<paragraph id="8B826773BE30FFA0FF6C569122E7ED6A" blockId="28.[151,1437,152,2018]" pageId="28" pageNumber="29">Head moderately depressed between eyes; well exposed in front of pronotum, not capable of complete retraction within prothoracic cavity; eyes moderately separated beneath at level of posterior margin of mouthpart complex; eyes above labrum moderately separated; frons-vertex junction rounded, lacking median elevation; posterolateral eye excavation not strongly developed, not visible in resting head position; antennal sockets on head between eyes, not contiguous, separated by up to ASW; clypeolabral suture present, flexible, not in front of anterior eye margin when head viewed with labrum horizontal; outer edges of labrum reach inner edges of closed mandibles. Mouthparts functional; apical segment of labial palpi non-lunate, strongly flattened, of form of wide triangle, with inner edge dentate (2 teeth). Antennae 11 segmented; length&gt;GHW to twice GHW; no segments flattened, shortened, or expanded; pedicel not produced; FS1 not shorter than pedicel.</paragraph>
<paragraph id="8B826773BE30FFA0FF6C5421238CED9A" blockId="28.[151,1437,152,2018]" pageId="28" pageNumber="29">Legs with inner tarsal claw not split; lacking MFC; no femora or tibiae swollen or curved; no basitarsi expanded or excavated.</paragraph>
<paragraph id="8B826773BE30FFA0FF6C54F12037E862" blockId="28.[151,1437,152,2018]" pageId="28" pageNumber="29">
Abdomen (
<bibRefCitation id="EFAC1A82BE30FFA0FEE154F120BAEDE2" author="Ballantyne, L. A. &amp; Lambkin, C." box="[330,669,952,978]" pageId="28" pageNumber="29" pagination="15 - 93" refId="ref103088" refString="Ballantyne, L. A. &amp; Lambkin, C. (2000) Lampyridae of Australia (Coleoptera: Lampyridae: Luciolinae: Luciolini). Memoirs of the Queensland Museum 46 (1), 15 - 93." type="journal article" year="2000">Ballantyne &amp; Lambkin 2000</bibRefCitation>
<figureCitation id="13067BF6BE30FFA0FD0E54F1213AEDE2" box="[677,797,952,978]" captionStart="FIGURE 3" captionStartId="22.[151,255,1622,1646]" captionText="FIGURE 3 Preferred MPT equivalent to majority rule consensus of 739 most parsimonious trees of length 3387 from phylogenetic analysis of 112 taxa and 343 morphological characters in Table 1. % of MPTs with the node, BS (bold), and node numbers along the branches. Where TreeRot v.2 was unable to find the shortest MPT, the BS is not bold. Part 2." pageId="28" pageNumber="29">Fig. 3 d, e</figureCitation>
) lacking cuticular remnants in association with aedeagal sheath; no ventrites with curved posterior margins nor extending anteriorly into emarginated posterior margin of anterior segment; LO in V7 entire occupying more than half V7, reaching to sides but not posterior margin [
<bibRefCitation id="EFAC1A82BE30FFA0FF5C53792076EA7A" author="Ballantyne, L. A. &amp; Lambkin, C." box="[247,593,1072,1098]" pageId="28" pageNumber="29" pagination="15 - 93" refId="ref103088" refString="Ballantyne, L. A. &amp; Lambkin, C. (2000) Lampyridae of Australia (Coleoptera: Lampyridae: Luciolinae: Luciolini). Memoirs of the Queensland Museum 46 (1), 15 - 93." type="journal article" year="2000">Ballantyne &amp; Lambkin (2000</bibRefCitation>
<figureCitation id="13067BF6BE30FFA0FDF253792083EA7A" box="[601,676,1072,1098]" captionStart="FIGURE 3" captionStartId="22.[151,255,1622,1646]" captionText="FIGURE 3 Preferred MPT equivalent to majority rule consensus of 739 most parsimonious trees of length 3387 from phylogenetic analysis of 112 taxa and 343 morphological characters in Table 1. % of MPTs with the node, BS (bold), and node numbers along the branches. Where TreeRot v.2 was unable to find the shortest MPT, the BS is not bold. Part 2." pageId="28" pageNumber="29">
Fig
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) illustrated V 7 withdrawn beneath V 6 thus the full extent of V7 and lateral margins of LO not fully visible]; posterior half of V7 not arched or swollen, muscle impressions not visible in this area; neither anterior nor posterior margin of LO emarginate; LO present in V6, occupying almost all V6. MPP present, symmetrical, apex rounded, entire, not laterally compressed, L=W, not inclined dorsally nor engulfed by T8 apex, lacking dorsal ridge, median longitudinal trough. V7 lacking median carina, median longitudinal trough, anteromedian depression on face of LO, PLP, incurving lobes or pointed projections, median dimple, or reflexed lobes. T7 lacking prolonged posterolateral corners. T8 not strongly sclerotised, symmetrical, W=L, visible posterior area not narrowing abruptly, lacking prolonged posterolateral corners, median posterior emargination, median posterior projections, not inclined ventrally nor engulfing posterior margin of V7 nor MPP, not extending conspicuously beyond posterior margin of V7; T8 ventral surface lacking flanges, lateral depressed troughs, median longitudinal trough, asymmetrical projections, median posterior ridge; concealed anterolateral arms of T8 very short, not laterally emarginated before their origins, not expanded dorsoventrally, lacking bifurcation of inner margin and ventrally directed pieces; lateral margins of T8 not enfolding sides of V7.
</paragraph>
<paragraph id="8B826773BE30FFA0FF6C51292048E972" blockId="28.[151,1437,152,2018]" pageId="28" pageNumber="29">Aedeagal sheath never&gt; 4 times as long as wide; lacking paraprocts; asymmetrical in posterior area where sheath sternite emarginated on right from point of attachment of tergite; sternite not angulate on L or R sides, not subparallel-sided, posterior margin entire, rounded, not emarginated on either side preapically, apex rounded; anterior half of sternite broad, apically rounded; tergite lacking lateral arms extending anteriorly at sides of sheath sternite; tergite not subdivided, lacking projecting pieces along posterior margin of tergite 9, anterior margin lacking transverse band.</paragraph>
<paragraph id="8B826773BE30FFA1FF6C5019265EEF4A" blockId="28.[151,1437,152,2018]" lastBlockId="29.[151,1436,152,938]" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">Aedeagus L/W 3/1 or shorter; subparallel-sided; LL lack lateral appendages; LL visible from beneath at sides of ML, LL/ML wide to moderate; LL of equal length, slightly shorter than ML, not diverging along inner dorsal margins, separated there by&gt; half their length; LL base width not = LL apex width which is narrower than that of ML; LL apices not expanded in horizontal plane; dorsal base of LL symmetrical, not excavated, median margin prolonged, broadly rounded; LL lacking lateral hairy appendages along their outer ventral margins, not produced preapically nor narrowly on inner apical margin, obliquely truncate along preapical inner margins; apices of LL not inturned, nor outturned; lacking projection on left LL; inner margins lacking slender leaf-like projection; ML symmetrical, lacking paired lateral teeth and tooth to left side, not strongly arched, apex not shaped like arrowhead, not bulbous, not inclined ventrally; BP not very narrow, not strongly sclerotised, not hooded, not strongly emarginated along anterior margin.</paragraph>
<paragraph id="8B826773BE31FFA1FF6C56CE260EEC72" blockId="29.[151,1436,152,938]" pageId="29" pageNumber="30">
<emphasis id="B949BB61BE31FFA1FF6C56CE230EEF91" bold="true" box="[199,297,391,417]" pageId="29" pageNumber="30">Female.</emphasis>
Macropterous and observed in flight. Pronotum lacking irregularities in posterolateral areas; punctation moderate to dense; pronotum subequal to humeral width; lacking indentation of lateral margin, irregularities at posterolateral corner; outline similar to that of male. Elytral punctation not as large as that of pronotum, nor evenly spaced; 2 interstitial lines; elytral carina absent. No legs or parts thereof swollen and /or curved. LO in V6 only, lacking any elevations or depressions or ridges on V7.
</paragraph>
<paragraph id="8B826773BE31FFA1FF6C550623EFEC5A" blockId="29.[151,1436,152,938]" box="[199,456,591,618]" pageId="29" pageNumber="30">
<emphasis id="B949BB61BE31FFA1FF6C55062334EC59" bold="true" box="[199,275,591,617]" pageId="29" pageNumber="30">Larva</emphasis>
not associated.
</paragraph>
</subSubSection>
<subSubSection id="C32734F8BE31FFA1FF6C553E2072EC8A" pageId="29" pageNumber="30" type="etymology">
<paragraph id="8B826773BE31FFA1FF6C553E2072EC8A" blockId="29.[151,1436,152,938]" pageId="29" pageNumber="30">
<emphasis id="B949BB61BE31FFA1FF6C553E236BECA1" bold="true" box="[199,332,631,657]" pageId="29" pageNumber="30">Etymology</emphasis>
.
<taxonomicName id="4C3D1CF0BE31FFA1FEF7553123E8ECA1" authorityName="Ballantyne &amp; Lambkin" authorityYear="2009" box="[348,463,632,657]" class="Insecta" family="Lampyridae" genus="Aquilonia" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="B949BB61BE31FFA1FEF7553123E8ECA1" box="[348,463,632,657]" italics="true" pageId="29" pageNumber="30">Aquilonia</emphasis>
</taxonomicName>
(feminine, from aquilonius Latin, northern) describes the northerly distribution in
<collectingCountry id="F32A27E3BE31FFA1FF3C55E92325EC8A" box="[151,258,672,698]" name="Australia" pageId="29" pageNumber="30">Australia</collectingCountry>
of the single known species.
</paragraph>
</subSubSection>
<subSubSection id="C32734F8BE31FFA1FF6C558E22D4ED9A" pageId="29" pageNumber="30" type="discussion">
<paragraph id="8B826773BE31FFA1FF6C558E22D4ED9A" blockId="29.[151,1436,152,938]" pageId="29" pageNumber="30">
<emphasis id="B949BB61BE31FFA1FF6C558E231EECD1" bold="true" box="[199,313,711,737]" pageId="29" pageNumber="30">Remarks</emphasis>
.
<taxonomicName id="4C3D1CF0BE31FFA1FEE155812399ECD1" authority="Ballantyne &amp; Lambkin, 2009" authorityName="Ballantyne &amp; Lambkin" authorityYear="2009" box="[330,446,712,737]" class="Insecta" family="Lampyridae" genus="Aquilonia" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B949BB61BE31FFA1FEE155812399ECD1" box="[330,446,712,737]" italics="true" pageId="29" pageNumber="30">Aquilonia</emphasis>
</taxonomicName>
<emphasis id="B949BB61BE31FFA1FE6C558E203CECD1" bold="true" box="[455,539,711,737]" pageId="29" pageNumber="30">
<taxonomicNameLabel id="A27A061ABE31FFA1FE6C558E203CECD1" box="[455,539,711,737]" pageId="29" pageNumber="30" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
is known from one Australian species, which is unusual in its distribution (restricted to the northern tropical areas in open forest), and its colour. While the dorsal colour pattern of light brown to yellow/orange, often with elytral apices black, is widespread in Asia, it is rare in
<collectingCountry id="F32A27E3BE31FFA1FB6B5451270BED02" box="[1216,1324,792,818]" name="Australia" pageId="29" pageNumber="30">Australia</collectingCountry>
and New
<collectingCountry id="F32A27E3BE31FFA1FF3C540922CBED6A" box="[151,236,832,858]" name="Guinea-Bissau" pageId="29" pageNumber="30">Guinea</collectingCountry>
and seen only in this species, two species of
<taxonomicName id="4C3D1CF0BE31FFA1FD56540921A2ED69" authority="Ballantyne &amp; Lambkin, 2009" authorityName="Ballantyne &amp; Lambkin" authorityYear="2009" box="[765,901,832,857]" class="Insecta" family="Lampyridae" genus="Gilvainsula" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B949BB61BE31FFA1FD56540921A2ED69" box="[765,901,832,857]" italics="true" pageId="29" pageNumber="30">Gilvainsula</emphasis>
</taxonomicName>
<emphasis id="B949BB61BE31FFA1FC25547621C7ED69" bold="true" box="[910,992,831,857]" pageId="29" pageNumber="30">
<taxonomicNameLabel id="A27A061ABE31FFA1FC25547621C7ED69" box="[910,992,831,857]" pageId="29" pageNumber="30" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="4C3D1CF0BE31FFA1FB8B54092700ED69" authority="Ballantyne &amp; Lambkin, 2009" authorityName="Ballantyne &amp; Lambkin" authorityYear="2009" box="[1056,1319,832,857]" class="Insecta" family="Lampyridae" genus="Pygatyphella" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="russellia" status="sp. nov.">
<emphasis id="B949BB61BE31FFA1FB8B54092700ED69" box="[1056,1319,832,857]" italics="true" pageId="29" pageNumber="30">Pygatyphella russellia</emphasis>
</taxonomicName>
<emphasis id="B949BB61BE31FFA1FA8454762756ED69" bold="true" box="[1327,1393,831,857]" pageId="29" pageNumber="30">
<taxonomicNameLabel id="A27A061ABE31FFA1FA8454762756ED69" box="[1327,1393,831,857]" pageId="29" pageNumber="30" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
No Asian species with this colouration investigated thus far (Ballantyne work in progress) conforms with
<taxonomicName id="4C3D1CF0BE31FFA1FA29542122CBED99" class="Insecta" family="Lampyridae" genus="Atyphella" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="costata">
<emphasis id="B949BB61BE31FFA1FA29542122CBED99" italics="true" pageId="29" pageNumber="30">A. costata</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>