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<document id="1BB334FF08D6689B003A33AFF35605A0" ID-DOI="10.5852/ejt.2021.761.1449" ID-GBIF-Dataset="bf24ff1f-569a-4222-afe1-8527c83d8003" ID-ISSN="2118-9773" ID-Zenodo-Dep="5156259" ID-ZooBank="519E2F56-40E1-4431-BC90-8484E308D16E" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="valdenar" IM.tables_approvedBy="valdenar" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628011133532" checkinUser="felipe" docAuthor="Tan, Ming Kai, Wahab, Rodzay bin Haji Abdul, Japir, Razy, Chung, Authur Y. C. &amp; Robillard, Tony" docDate="2021" docId="039487A4FFF7FFE2FDC5F533FEECFE3A" docLanguage="en" docName="ejt-761_1-75.pdf" docOrigin="European Journal of Taxonomy 761 (1)" docSource="http://dx.doi.org/10.5852/ejt.2021.761.1449" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Nisitrus crucius Robillard &amp; Tan 2021, sp. nov." docType="treatment" docUuid="3140F3A4-21FB-4BB4-ACC6-57BD749A6C3B" docUuidSource="ZooBank" docVersion="19" lastPageNumber="31" masterDocId="FFADFFDCFFECFFFCFFE0F365FFD5FFF9" masterDocTitle="Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species" masterLastPageNumber="75" masterPageNumber="1" pageNumber="28" updateTime="1701996970784" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="18998A2FD027C47CAFAE2758D18AB6FA">Revision of the cricket genus Nisitrus Saussure (Orthoptera: Gryllidae: Eneopterinae) and descriptions of five new species</mods:title>
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<mods:namePart id="ADBE2D01AF84CD2D3161DD60A440260F">Tan, Ming Kai</mods:namePart>
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<mods:affiliation id="A8017D8144FF49AECCFB4DB9F0314EE3">Institut de Systématique, Evolution et Biodiversité (ISYEB), Muséum national dʼhistoire naturelle, CNRS, SU, EPHE, UA, 57 rue Cuvier, CP 50, 75231 Paris Cedex 05, France.</mods:affiliation>
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<mods:namePart id="D9A0BBB3F578463ADA224BD5EB6106E4">Wahab, Rodzay bin Haji Abdul</mods:namePart>
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<mods:affiliation id="91CCE705BD5A9AD6A155190174D521AF">Institute for Biodiversity and Environmental Research, Universiti Brunei Darussalam, Jalan Universiti, BE 1410, Brunei Darussalam.</mods:affiliation>
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<mods:affiliation id="3B9658D1C848B7D775E89D307806FCAC">Forest Research Centre (Sepilok), Sabah Forestry Department, P. O. Box 1407, 90715 Sandakan, Sabah, Malaysia.</mods:affiliation>
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<mods:affiliation id="4B524F2714B624AF6F7A5BE222395D79">Forest Research Centre (Sepilok), Sabah Forestry Department, P. O. Box 1407, 90715 Sandakan, Sabah, Malaysia.</mods:affiliation>
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<taxonomicName id="4C3D4D31FFF7FFE7FDC5F533FC78F988" ID-CoL="8QKW6" authority="Robillard &amp; Tan" authorityName="Robillard &amp; Tan" authorityYear="2021" box="[549,941,1622,1649]" class="Insecta" family="Gryllidae" genus="Nisitrus" kingdom="Animalia" order="Orthoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="crucius" status="sp. nov.">
<emphasis id="B949EAA0FFF7FFE7FDC5F533FD32F989" bold="true" box="[549,743,1622,1648]" italics="true" pageId="27" pageNumber="28">Nisitrus crucius</emphasis>
Robillard &amp; Tan
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<figureCitation id="13062A37FFF7FFE7FE6DF5C4FE02F942" box="[397,471,1696,1723]" captionStart="Fig" captionStartId="9.[189,233,1963,1989]" captionTargetBox="[187,1396,991,1924]" captionTargetId="figure-297@9.[187,1396,990,1924]" captionTargetPageId="9" captionText="Fig. 5. Map of Southeast Asia showing the distribution of species of Nisitrus Saussure, 1878. Each coloured symbol represents one sampling or observation locality listed in the material in the text." figureDoi="http://doi.org/10.5281/zenodo.5156278" httpUri="https://zenodo.org/record/5156278/files/figure.png" pageId="27" pageNumber="28">Figs 5</figureCitation>
, 
<figureCitation id="13062A37FFF7FFE7FE05F5C4FDDCF942" box="[485,521,1697,1723]" captionStart="Fig" captionStartId="12.[189,232,1928,1954]" captionTargetBox="[220,1368,276,1865]" captionTargetId="figure-14@12.[220,1368,265,1889]" captionTargetPageId="12" captionText="Fig. 7. Habitus of ♂♂ in dorsal view. A. Nisitrus brunnerianus Saussure, 1878. B. N. crucius Robillard &amp; Tan sp. nov. C. N. hughtani Robillard &amp; Tan sp. nov. D. N. insignis Saussure, 1878. E. N. malaya Robillard &amp; Tan sp. nov. F. N. musicus Ingrisch, 1987. G. N. vittatus (Haan, 1844). Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5156284" httpUri="https://zenodo.org/record/5156284/files/figure.png" pageId="27" pageNumber="28">7B</figureCitation>
, 
<figureCitation id="13062A37FFF7FFE7FDF6F5C4FDEFF942" box="[534,570,1697,1723]" captionStart="Fig" captionStartId="13.[189,233,1926,1952]" captionTargetBox="[232,1318,284,1848]" captionTargetId="figure-11@13.[199,1388,265,1889]" captionTargetPageId="13" captionText="Fig. 8. Habitus of ♀♀ in dorsal view. A–B. Nisitrus brunnerianus Saussure, 1878. C. N. crucius Robillard &amp; Tan sp. nov. D. N. danum Robillard &amp; Tan sp. nov. E. N. hughtani Robillard &amp; Tan sp. nov. F. N. insignis Saussure, 1878. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5156286" httpUri="https://zenodo.org/record/5156286/files/figure.png" pageId="27" pageNumber="28">8C</figureCitation>
, 
<figureCitation id="13062A37FFF7FFE7FDA6F5C4FDACF942" box="[582,633,1697,1723]" captionStart="Fig" captionStartId="15.[189,232,1742,1768]" captionTargetBox="[216,1388,279,1679]" captionTargetId="figure-11@15.[189,1398,263,1705]" captionTargetPageId="15" captionText="Fig. 10. Face in anterior view. A. Nisitrus brunnerianus Saussure, 1878. B. N. crucius Robillard &amp; Tan sp. nov. C. N. danum Robillard &amp; Tan sp. nov. D. N. hughtani Robillard &amp; Tan sp. nov. E. N. insignis Saussure, 1878. F. N. malaya Robillard &amp; Tan sp. nov. G. N. musicus Ingrisch, 1987. H. N. rindu Robillard &amp; Tan sp. nov. I. N. vittatus (Haan, 1844). Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5156294" httpUri="https://zenodo.org/record/5156294/files/figure.png" pageId="27" pageNumber="28">10B</figureCitation>
, 
<figureCitation id="13062A37FFF7FFE7FD66F5C4FD6DF942" box="[646,696,1697,1723]" captionStart="Fig" captionStartId="16.[189,232,1892,1918]" captionTargetBox="[223,1354,324,1818]" captionTargetId="figure-15@16.[197,1389,265,1854]" captionTargetPageId="16" captionText="Fig. 11. Head and pronotum in dorsal view. A. Nisitrus brunnerianus Saussure, 1878. B. N. crucius Robillard &amp; Tan sp. nov. C. N. danum Robillard &amp; Tan sp. nov. D. N. hughtani Robillard &amp; Tan sp. nov. E. N. insignis Saussure, 1878. F. N. malaya Robillard &amp; Tan sp. nov. G. N. musicus Ingrisch, 1987. H. N. rindu Robillard &amp; Tan sp. nov. I. N. vittatus (Haan, 1844). Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5156298" httpUri="https://zenodo.org/record/5156298/files/figure.png" pageId="27" pageNumber="28">11B</figureCitation>
, 
<figureCitation id="13062A37FFF7FFE7FD25F5C4FD2DF942" box="[709,760,1697,1723]" captionStart="Fig" captionStartId="17.[189,232,1645,1671]" captionTargetBox="[210,1398,290,1606]" captionTargetId="figure-11@17.[189,1398,265,1606]" captionTargetPageId="17" captionText="Fig. 12. Body in lateral view. A. Nisitrus brunnerianus Saussure, 1878. B. N. crucius Robillard &amp; Tan sp. nov. C. N. danum Robillard &amp; Tan sp. nov. D. N. hughtani Robillard &amp; Tan sp. nov. E. N. insignis Saussure, 1878. F. N. malaya Robillard &amp; Tan sp. nov. G. N. musicus Ingrisch, 1987. H. N. rindu Robillard &amp; Tan sp. nov. I. N. vittatus (Haan, 1844). Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5156302" httpUri="https://zenodo.org/record/5156302/files/figure.png" pageId="27" pageNumber="28">12B</figureCitation>
, 
<figureCitation id="13062A37FFF7FFE7FCE4F5C4FCE2F942" box="[772,823,1697,1723]" captionStart="Fig" captionStartId="18.[189,231,1928,1954]" captionTargetBox="[243,1357,300,1857]" captionTargetId="figure-15@18.[219,1368,265,1891]" captionTargetPageId="18" captionText="Fig. 13. Male tegmina in dorsal view. A. Nisitrus brunnerianus Saussure, 1878.B. N. crucius Robillard &amp; Tan sp. nov. C. N. hughtani Robillard &amp; Tan sp. nov. D. N. insignis Saussure, 1878. E. N. malaya Robillard &amp; Tan sp. nov. F. N. musicus Ingrisch, 1987. G. N. vittatus (Haan, 1844). Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5156304" httpUri="https://zenodo.org/record/5156304/files/figure.png" pageId="27" pageNumber="28">13B</figureCitation>
, 
<figureCitation id="13062A37FFF7FFE7FCA4F5C4FCA2F942" box="[836,887,1697,1723]" captionStart="Fig" captionStartId="19.[189,233,1894,1920]" captionTargetBox="[216,1372,287,1817]" captionTargetId="figure-11@19.[193,1394,265,1855]" captionTargetPageId="19" captionText="Fig. 14. Female tegmina in dorsal view. A. Nisitrus brunnerianus Saussure, 1878. B. N. crucius Robillard &amp; Tan sp. nov. C. N. danum Robillard &amp; Tan sp. nov. D. N. hughtani Robillard &amp; Tan sp. nov. E. N. insignis Saussure, 1878. F. N. malaya Robillard &amp; Tan sp. nov. G. N. musicus Ingrisch, 1987. H. N. rindu Robillard &amp; Tan sp. nov. I. N. vittatus (Haan, 1844). Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5156308" httpUri="https://zenodo.org/record/5156308/files/figure.png" pageId="27" pageNumber="28">14B</figureCitation>
, 
<figureCitation id="13062A37FFF7FFE7FC63F5C4FC62F942" box="[899,951,1697,1723]" captionStart="Fig" captionStartId="21.[189,231,1640,1666]" captionTargetBox="[218,1372,288,1585]" captionTargetId="figure-11@21.[189,1398,265,1601]" captionTargetPageId="21" captionText="Fig. 16. Male genitalia in ventral view. A. Nisitrus brunnerianus Saussure, 1878. B. N. crucius Robillard &amp; Tan sp. nov. C. N. hughtani Robillard &amp; Tan sp. nov. D. N. insignis Saussure, 1878. E. N. malaya Robillard &amp; Tan sp. nov. F. N. musicus Ingrisch, 1987. G. N. vittatus (Haan, 1844). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5156316" httpUri="https://zenodo.org/record/5156316/files/figure.png" pageId="27" pageNumber="28">16B</figureCitation>
, 
<figureCitation id="13062A37FFF7FFE7FC23F5C4FBCEF942" box="[963,1051,1697,1723]" captionStart="Fig" captionStartId="22.[189,232,1809,1835]" captionTargetBox="[201,1393,282,1740]" captionTargetId="figure-14@22.[189,1398,265,1771]" captionText="Fig. 17. Female genitalia in lateral (A, C, E, F, H, J, L, N, P) and dorsal (B, D, G, I, K, M, O, Q) views. A–B. Nisitrus brunnerianus Saussure, 1878. C–D. N. crucius Robillard &amp; Tan sp. nov. E. N. danum Robillard &amp; Tan sp. nov. F–G. N. hughtani Robillard &amp; Tan sp. nov. H–I. N. insignis Saussure, 1878. J–K. N. malaya Robillard &amp; Tan sp. nov. L–M. N. musicus Ingrisch, 1987. N–O. N. rindu Robillard &amp; Tan sp. nov. P–Q. N. vittatus (Haan, 1844). Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5165004" httpUri="https://zenodo.org/record/5165004/files/figure.png" pageId="27" pageNumber="28">17C–D</figureCitation>
; 
<tableCitation id="C6BF0309FFF7FFE7FBC6F5C4FB5EF942" box="[1062,1163,1697,1723]" captionStart="Table 2" captionStartId="10.[189,258,265,291]" captionTargetPageId="10" captionText="Table 2. Status of original species epithets associated with Nisitrus Saussure, 1878." httpUri="http://table.plazi.org/id/DF42663AFFE6FFF6FF5DF26CFB5AFEDA" pageId="27" pageNumber="28" tableUuid="DF42663AFFE6FFF6FF5DF26CFB5AFEDA">Tables 2</tableCitation>
, 
<tableCitation id="C6BF0309FFF7FFE7FB77F5C4FB73F942" box="[1175,1190,1697,1723]" captionStart="Table 4" captionStartId="29.[189,258,265,291]" captionTargetBox="[202,1376,331,543]" captionTargetPageId="29" captionText="Table 4. Measurements of Nisitrus crucius Robillard &amp; Tan sp. nov. (in mm, means in brackets)." httpUri="http://table.plazi.org/id/DF42663AFFF1FFE1FF5DF26CFAFEFEDD" pageId="27" pageNumber="28" tableUuid="DF42663AFFF1FFE1FF5DF26CFAFEFEDD">4</tableCitation>
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</subSubSection>
<subSubSection id="C3276539FFF7FFE7FF5DF581FB4DF811" pageId="27" pageNumber="28" type="diagnosis">
<paragraph id="8B8236B2FFF7FFE7FF5DF581FEE3F907" blockId="27.[189,310,1764,1790]" box="[189,310,1764,1790]" pageId="27" pageNumber="28">
<heading id="D0CA81DEFFF7FFE7FF5DF581FEE3F907" bold="true" box="[189,310,1764,1790]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis id="B949EAA0FFF7FFE7FF5DF581FEE3F907" bold="true" box="[189,310,1764,1790]" pageId="27" pageNumber="28">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B8236B2FFF7FFE7FF5DF471FB4DF811" blockId="27.[189,1400,1812,2025]" pageId="27" pageNumber="28">
This species differs from all known congeners by the following combination of characters: frons yellow, with a cross-shaped black pattern above mouthparts; face part of fastigium between eyes black. Male FWs with a basal area generally paler; female ovipositor only about as long as FIII. 
<taxonomicName id="4C3D4D31FFF7FFE7FB5AF405FAADF880" authority="Tan &amp; Wahab &amp; Japir &amp; Chung &amp; Robillard, 2021" authorityName="Tan &amp; Wahab &amp; Japir &amp; Chung &amp; Robillard" authorityYear="2021" box="[1210,1400,1888,1913]" class="Insecta" family="Gryllidae" genus="Nisitrus" kingdom="Animalia" order="Orthoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="crucius" status="sp. nov.">
<emphasis id="B949EAA0FFF7FFE7FB5AF405FAADF880" box="[1210,1400,1888,1913]" italics="true" pageId="27" pageNumber="28">Nisitrus crucius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57DBFFF7FFE7FF5FF4E0FECFF866" box="[191,282,1925,1951]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
is similar to 
<taxonomicName id="4C3D4D31FFF7FFE7FE54F4E0FDA1F867" authorityName="Saussure" authorityYear="1878" box="[436,628,1924,1950]" class="Insecta" family="Gryllidae" genus="Nisitrus" kingdom="Animalia" order="Orthoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="brunnerianus">
<emphasis id="B949EAA0FFF7FFE7FE54F4E0FDA1F867" box="[436,628,1924,1950]" italics="true" pageId="27" pageNumber="28">N. brunnerianus</emphasis>
</taxonomicName>
, from which it differs by smaller size and shorter ovipositor. It is also similar in size to 
<taxonomicName id="4C3D4D31FFF7FFE7FE23F4CFFDE8F83A" baseAuthorityName="Haan" baseAuthorityYear="1844" box="[451,573,1962,1987]" class="Insecta" family="Gryllidae" genus="Nisitra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="vittatus">
<emphasis id="B949EAA0FFF7FFE7FE23F4CFFDE8F83A" box="[451,573,1962,1987]" italics="true" pageId="27" pageNumber="28">N. vittatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3D4D31FFF7FFE7FD97F4CFFD25F83A" authority="Tan &amp; Wahab &amp; Japir &amp; Chung &amp; Robillard, 2021" authorityName="Tan &amp; Wahab &amp; Japir &amp; Chung &amp; Robillard" authorityYear="2021" box="[631,752,1961,1987]" class="Insecta" family="Gryllidae" genus="Nisitrus" kingdom="Animalia" order="Orthoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="malaya" status="sp. nov.">
<emphasis id="B949EAA0FFF7FFE7FD97F4CFFD25F83A" box="[631,752,1961,1987]" italics="true" pageId="27" pageNumber="28">N. malaya</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57DBFFF7FFE7FD1AF4CFFC81F83D" box="[762,852,1962,1988]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
but differs by female FW with cells on dorsal field mostly infumated black, gradually becoming yellow near CuA, M and R veins.
</paragraph>
</subSubSection>
<subSubSection id="C3276539FFF0FFE0FF5DF26CFB21FEAD" box="[189,1268,265,340]" pageId="28" pageNumber="29" type="etymology">
<paragraph id="8B8236B2FFF0FFE0FF5DF26CFE91FEDA" blockId="28.[189,324,265,291]" box="[189,324,265,291]" pageId="28" pageNumber="29">
<heading id="D0CA81DEFFF0FFE0FF5DF26CFE91FEDA" bold="true" box="[189,324,265,291]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="B949EAA0FFF0FFE0FF5DF26CFE91FEDA" bold="true" box="[189,324,265,291]" pageId="28" pageNumber="29">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8B8236B2FFF0FFE0FF5DF25CFB21FEAD" blockId="28.[189,1268,313,340]" box="[189,1268,313,340]" pageId="28" pageNumber="29">
The species name refers to the dark cross marking on the frons (ʻ 
<emphasis id="B949EAA0FFF0FFE0FC57F25FFC2AFEAA" box="[951,1023,314,339]" italics="true" pageId="28" pageNumber="29">crucis</emphasis>
ʼ = ʻcrossʼ, in Latin).
</paragraph>
</subSubSection>
<subSubSection id="C3276539FFF0FFE0FF5DF219FBA5FBD1" pageId="28" pageNumber="29" type="materials_examined">
<paragraph id="8B8236B2FFF0FFE0FF5DF219FE7FFE6F" blockId="28.[189,426,380,406]" box="[189,426,380,406]" pageId="28" pageNumber="29">
<heading id="D0CA81DEFFF0FFE0FF5DF219FE7FFE6F" bold="true" box="[189,426,380,406]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="B949EAA0FFF0FFE0FF5DF219FE7FFE6F" bold="true" box="[189,426,380,406]" pageId="28" pageNumber="29">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8B8236B2FFF0FFE0FF0AF2C8FD1FFDE9" blockId="28.[189,1399,429,529]" pageId="28" pageNumber="29">
<materialsCitation id="3B553CEFFFF0FFE0FF0AF2C8FD1FFDE9" ID-GBIF-Occurrence="3333037407" collectingDate="1925-08-16" collectingDateMax="1925-08-20" collectingDateMin="1925-08-16" collectionCode="MNHN-EO-ENSIF" collectorName="H. C. Siebers" country="Indonesia" latitude="1.352" location="Biv. Long Hoet" longLatPrecision="1" longitude="116.51781" pageId="28" pageNumber="29" specimenCode="MNHN-EO-ENSIF11261" specimenCount="1" specimenCount-male="1" stateProvince="Kalimantan" typeStatus="holotype">
<emphasis id="B949EAA0FFF0FFE0FF0AF2C8FE89FE3E" bold="true" box="[234,348,429,455]" pageId="28" pageNumber="29">
<typeStatus id="54868810FFF0FFE0FF0AF2C8FE89FE3E" box="[234,348,429,455]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F32A7622FFF0FFE0FF5DF2B7FEBDFE15" box="[189,360,466,492]" name="Indonesia" pageId="28" pageNumber="29">INDONESIA</collectingCountry>
– 
<emphasis id="B949EAA0FFF0FFE0FE6BF2B4FDF7FE12" bold="true" box="[395,546,465,491]" pageId="28" pageNumber="29">
<collectingRegion id="49F9F850FFF0FFE0FE6BF2B4FDF7FE12" box="[395,546,465,491]" country="Indonesia" name="Kalimantan Utara" pageId="28" pageNumber="29">Kalimantan</collectingRegion>
</emphasis>
• 
<specimenCount id="9D3BFD3BFFF0FFE0FDDDF2B7FD83FE15" box="[573,598,466,492]" pageId="28" pageNumber="29" type="male">♂</specimenCount>
; “ 
<location id="8EE26069FFF0FFE0FD8FF2B4FCE5FE15" LSID="urn:lsid:plazi:treatment:039487A4FFF7FFE2FDC5F533FEECFE3A:8EE26069FFF0FFE0FD8FF2B4FCE5FE15" box="[623,816,465,492]" country="Indonesia" latitude="1.352" longLatPrecision="1" longitude="116.51781" name="Biv. Long Hoet" pageId="28" pageNumber="29" stateProvince="Kalimantan">Biv. Long Hoet</location>
”; 
<geoCoordinate id="EE095075FFF0FFE0FCA9F2B5FC0CFE15" box="[841,985,464,492]" degrees="1" direction="north" minutes="21" orientation="latitude" pageId="28" pageNumber="29" precision="1" seconds="7.2" value="1.352">1°21′7.2″ N</geoCoordinate>
, 
<geoCoordinate id="EE095075FFF0FFE0FC07F2B5FB44FE15" box="[999,1169,464,492]" degrees="116" direction="east" minutes="31" orientation="longitude" pageId="28" pageNumber="29" precision="1" seconds="4.1" value="116.51781">116°31′4.1″ E</geoCoordinate>
; 
<date id="FF831072FFF0FFE0FB7FF2B4FAA4FE15" box="[1183,1393,465,492]" pageId="28" pageNumber="29" value="1925-08-16" valueMax="1925-08-20" valueMin="1925-08-16">
<collectingDate id="EFC7E99AFFF0FFE0FB7FF2B4FAA4FE15" box="[1183,1393,465,492]" pageId="28" pageNumber="29" value="1925-08-16" valueMax="1925-08-20" valueMin="1925-08-16">16–20 Aug. 1925</collectingDate>
</date>
; 
<collectorName id="26C85364FFF0FFE0FF5DF292FE87FDE9" box="[189,338,502,529]" pageId="28" pageNumber="29">H.C. Siebers</collectorName>
leg.; 
<specimenCode id="DB9B9EC9FFF0FFE0FE74F293FD13FDE9" box="[404,710,502,528]" collectionCode="MNHN-EO-ENSIF" pageId="28" pageNumber="29">MNHN-EO-ENSIF11261</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8B8236B2FFF0FFE0FF0AF15FFE86FCA1" blockId="28.[189,1399,570,856]" pageId="28" pageNumber="29">
<materialsCitation id="3B553CEFFFF0FFE0FF0AF15FFE8FFD67" ID-GBIF-Occurrence="3333037525" collectorName="Long Nawang &amp; Mjoberg" country="Indonesia" location="Long Navang" municipality="Long Navang" pageId="28" pageNumber="29" specimenCode="MNHN-EO- ENSIF11262" specimenCount="1" specimenCount-female="1" stateProvince="Kalimantan" typeStatus="paratype">
<emphasis id="B949EAA0FFF0FFE0FF0AF15FFEB2FDAD" bold="true" box="[234,359,570,596]" pageId="28" pageNumber="29">
<typeStatus id="54868810FFF0FFE0FF0AF15FFEB2FDAD" box="[234,359,570,596]" pageId="28" pageNumber="29" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="F32A7622FFF0FFE0FF5DF13AFEBEFD80" box="[189,363,607,633]" name="Indonesia" pageId="28" pageNumber="29">INDONESIA</collectingCountry>
– 
<emphasis id="B949EAA0FFF0FFE0FE70F13AFDFFFD80" bold="true" box="[400,554,607,633]" pageId="28" pageNumber="29">
<collectingRegion id="49F9F850FFF0FFE0FE70F13AFDFFFD80" box="[400,554,607,633]" country="Indonesia" name="Kalimantan Utara" pageId="28" pageNumber="29">Kalimantan</collectingRegion>
</emphasis>
• 
<specimenCount id="9D3BFD3BFFF0FFE0FDA9F13AFDAFFD80" box="[585,634,607,633]" pageId="28" pageNumber="29" type="female">1 ♀</specimenCount>
; “ 
<collectingMunicipality id="6BE6ACC8FFF0FFE0FD76F13AFC91FD80" box="[662,836,607,633]" country="Indonesia" name="Long Navang" pageId="28" pageNumber="29" stateProvince="Kalimantan Utara">Long Navang</collectingMunicipality>
” [
<collectorName id="26C85364FFF0FFE0FCBFF13AFBCDFD80" box="[863,1048,606,633]" pageId="28" pageNumber="29">Long Nawang</collectorName>
]; 
<collectorName id="26C85364FFF0FFE0FBCEF13AFB40FD80" box="[1070,1173,607,633]" pageId="28" pageNumber="29">Mjöberg</collectorName>
leg.; 
<specimenCode id="DB9B9EC9FFF0FFE0FB3FF13AFE8FFD67" pageId="28" pageNumber="29">MNHN-EO- ENSIF11262</specimenCode>
</materialsCitation>
• 
<materialsCitation id="3B553CEFFFF0FFE0FE99F1E1FD16FD3A" ID-GBIF-Occurrence="3333037623" collectingDate="1925-11-10" collectingDateMax="1925-11-28" collectingDateMin="1925-11-10" collectionCode="MNHN-EO-ENSIF" collectorName="H. C. Siebers" country="Indonesia" latitude="0.9752777" location="Marah" longLatPrecision="15" longitude="116.554115" pageId="28" pageNumber="29" specimenCode="MNHN-EO-ENSIF11263" specimenCount="1" specimenCount-female="1" stateProvince="Kalimantan Utara" typeStatus="paratype">
<specimenCount id="9D3BFD3BFFF0FFE0FE99F1E1FE7CFD67" box="[377,425,644,670]" pageId="28" pageNumber="29" type="female">1 ♀</specimenCount>
; “ 
<locationDeviation id="9A302F90FFF0FFE0FE25F1E1FD61FD67" box="[453,692,644,670]" pageId="28" pageNumber="29">Midden O. Borneo</locationDeviation>
, 
<location id="8EE26069FFF0FFE0FD22F1E1FCC2FD67" LSID="urn:lsid:plazi:treatment:039487A4FFF7FFE2FDC5F533FEECFE3A:8EE26069FFF0FFE0FD22F1E1FCC2FD67" box="[706,791,644,670]" country="Indonesia" latitude="0.9752777" longLatPrecision="15" longitude="116.554115" name="Marah" pageId="28" pageNumber="29" stateProvince="Kalimantan Utara">Marah</location>
”; 
<geoCoordinate id="EE095075FFF0FFE0FCD0F1E7FC68FD67" box="[816,957,642,670]" degrees="0" direction="north" minutes="58" orientation="latitude" pageId="28" pageNumber="29" precision="15" seconds="31" value="0.9752777">0°58′31″ N</geoCoordinate>
, 
<geoCoordinate id="EE095075FFF0FFE0FC2CF1E7FB5CFD67" box="[972,1161,642,670]" degrees="116" direction="east" minutes="33" orientation="longitude" pageId="28" pageNumber="29" precision="1" seconds="14.8" value="116.554115">116°33′14.8″ E</geoCoordinate>
; 
<date id="FF831072FFF0FFE0FB79F1E1FAA7FD66" box="[1177,1394,644,671]" pageId="28" pageNumber="29" value="1925-11-10" valueMax="1925-11-28" valueMin="1925-11-10">
<collectingDate id="EFC7E99AFFF0FFE0FB79F1E1FAA7FD66" box="[1177,1394,644,671]" pageId="28" pageNumber="29" value="1925-11-10" valueMax="1925-11-28" valueMin="1925-11-10">10–28 Nov. 1925</collectingDate>
</date>
; 
<collectorName id="26C85364FFF0FFE0FF5DF1CCFE87FD3A" box="[189,338,681,707]" pageId="28" pageNumber="29">H.C. Siebers</collectorName>
leg.; 
<specimenCode id="DB9B9EC9FFF0FFE0FE76F1CCFD16FD3A" box="[406,707,681,707]" collectionCode="MNHN-EO-ENSIF" pageId="28" pageNumber="29">MNHN-EO-ENSIF11263</specimenCode>
</materialsCitation>
• 
<materialsCitation id="3B553CEFFFF0FFE0FD3DF1CCFC51FD11" ID-GBIF-Occurrence="3333037591" collectingDate="1925-08-16" collectingDateMax="1925-08-20" collectingDateMin="1925-08-16" collectionCode="MNHN-EO-ENSIF" collectorName="H. C. Siebers" country="Indonesia" latitude="1.352" location="Biv. Long Hoet" longLatPrecision="1" longitude="116.51781" pageId="28" pageNumber="29" specimenCode="MNHN-EO-ENSIF11264" specimenCount="1" specimenCount-female="1" stateProvince="Kalimantan Utara" typeStatus="paratype">
<specimenCount id="9D3BFD3BFFF0FFE0FD3DF1CCFCDEFD3A" box="[733,779,681,707]" pageId="28" pageNumber="29" type="female">1 ♀</specimenCount>
; “ 
<location id="8EE26069FFF0FFE0FCC5F1CCFC36FD3A" LSID="urn:lsid:plazi:treatment:039487A4FFF7FFE2FDC5F533FEECFE3A:8EE26069FFF0FFE0FCC5F1CCFC36FD3A" box="[805,995,681,707]" country="Indonesia" latitude="1.352" longLatPrecision="1" longitude="116.51781" name="Biv. Long Hoet" pageId="28" pageNumber="29" stateProvince="Kalimantan Utara">Biv. Long Hoet</location>
”; 
<geoCoordinate id="EE095075FFF0FFE0FC1AF1C2FB5CFD3A" box="[1018,1161,679,707]" degrees="1" direction="north" minutes="21" orientation="latitude" pageId="28" pageNumber="29" precision="1" seconds="7.2" value="1.352">1°21′7.2″ N</geoCoordinate>
, 
<geoCoordinate id="EE095075FFF0FFE0FB75F1C2FAEBFD3A" box="[1173,1342,679,707]" degrees="116" direction="east" minutes="31" orientation="longitude" pageId="28" pageNumber="29" precision="1" seconds="4.1" value="116.51781">116°31′4.1″ E</geoCoordinate>
; 
<date id="FF831072FFF0FFE0FAAAF1CCFEB0FD10" pageId="28" pageNumber="29" value="1925-08-16" valueMax="1925-08-20" valueMin="1925-08-16">
<collectingDate id="EFC7E99AFFF0FFE0FAAAF1CCFEB0FD10" pageId="28" pageNumber="29" value="1925-08-16" valueMax="1925-08-20" valueMin="1925-08-16">16– 20 Aug. 1925</collectingDate>
</date>
; 
<collectorName id="26C85364FFF0FFE0FE93F1ABFDD9FD11" box="[371,524,718,744]" pageId="28" pageNumber="29">H.C. Siebers</collectorName>
leg.; 
<specimenCode id="DB9B9EC9FFF0FFE0FDB2F1ABFC51FD11" box="[594,900,718,744]" collectionCode="MNHN-EO-ENSIF" pageId="28" pageNumber="29">MNHN-EO-ENSIF11264</specimenCode>
</materialsCitation>
• 
<materialsCitation id="3B553CEFFFF0FFE0FC41F1ABFBE9FCF4" ID-GBIF-Occurrence="3333037418" collectingDate="1924-11" collectingDateMax="1924-12-31" collectingDateMin="1924-11" collectionCode="MNHN-EO-ENSIF" collectorName="Blanchemanche" country="Indonesia" latitude="-0.5744444" location="Borneo" longLatPrecision="21" longitude="111.75139" municipality="Melawi" pageId="28" pageNumber="29" specimenCode="MNHN-EO-ENSIF1732" specimenCount="1" specimenCount-female="1" stateProvince="Kalimantan Utara" typeStatus="paratype">
<specimenCount id="9D3BFD3BFFF0FFE0FC41F1ABFC05FD10" box="[929,976,718,745]" pageId="28" pageNumber="29" type="female">1 ♀</specimenCount>
; “ 
<collectingMunicipality id="6BE6ACC8FFF0FFE0FC0BF1ABFB9CFD11" box="[1003,1097,718,744]" pageId="28" pageNumber="29">Melawi</collectingMunicipality>
W 
<location id="8EE26069FFF0FFE0FB96F1ABFB01FD11" LSID="urn:lsid:plazi:treatment:039487A4FFF7FFE2FDC5F533FEECFE3A:8EE26069FFF0FFE0FB96F1ABFB01FD11" box="[1142,1236,718,744]" country="Indonesia" latitude="-0.5744444" longLatPrecision="21" longitude="111.75139" municipality="Melawi" name="Borneo" pageId="28" pageNumber="29" stateProvince="Kalimantan Utara">Borneo</location>
”; 
<geoCoordinate id="EE095075FFF0FFE0FB0CF1A9FAA4FD11" box="[1260,1393,716,744]" degrees="0" direction="south" minutes="34" orientation="latitude" pageId="28" pageNumber="29" precision="15" seconds="28" value="-0.5744444">0°34′28″ S</geoCoordinate>
, 
<geoCoordinate id="EE095075FFF0FFE0FF5DF194FE99FCF7" box="[189,332,753,782]" degrees="111" direction="east" minutes="45" orientation="longitude" pageId="28" pageNumber="29" precision="15" seconds="5" value="111.75139">111°45′5″ E</geoCoordinate>
; 
<date id="FF831072FFF0FFE0FEB7F191FDCDFCF4" box="[343,536,755,782]" pageId="28" pageNumber="29" value="1924-11" valueMax="1924-12-31" valueMin="1924-11-00">
<collectingDate id="EFC7E99AFFF0FFE0FEB7F191FDCDFCF4" box="[343,536,755,782]" pageId="28" pageNumber="29" value="1924-11" valueMax="1924-12-31" valueMin="1924-11-00">Nov.–Dec. 1924</collectingDate>
</date>
; 
<collectorName id="26C85364FFF0FFE0FDC2F196FD08FCF4" box="[546,733,755,781]" pageId="28" pageNumber="29">Blanchemanche</collectorName>
leg.; 
<specimenCode id="DB9B9EC9FFF0FFE0FCFCF196FBE9FCF4" box="[796,1084,755,781]" collectionCode="MNHN-EO-ENSIF" pageId="28" pageNumber="29">MNHN-EO-ENSIF1732</specimenCode>
</materialsCitation>
• 
<materialsCitation id="3B553CEFFFF0FFE0FBB2F196FD6CFCCA" ID-GBIF-Occurrence="3333037425" collectingDate="1924-11" collectingDateMax="1924-12-31" collectingDateMin="1924-11" collectionCode="MNHN-EO-ENSIF" collectorName="Blanchemanche" country="Indonesia" latitude="-0.5744444" location="Borneo" longLatPrecision="21" longitude="111.75139" municipality="Melawi" pageId="28" pageNumber="29" specimenCode="MNHN-EO-ENSIF11265" specimenCount="1" specimenCount-male="1" stateProvince="Kalimantan Utara" typeStatus="paratype">
<specimenCount id="9D3BFD3BFFF0FFE0FBB2F196FBAAFCF7" box="[1106,1151,755,782]" pageId="28" pageNumber="29" type="male">1 ♂</specimenCount>
; same collection data as for preceding; 
<specimenCode id="DB9B9EC9FFF0FFE0FE6AF07DFD6CFCCA" box="[394,697,792,819]" collectionCode="MNHN-EO-ENSIF" pageId="28" pageNumber="29">MNHN-EO-ENSIF11265</specimenCode>
</materialsCitation>
• 
<materialsCitation id="3B553CEFFFF0FFE0FD32F07DFE9AFCA1" ID-GBIF-Occurrence="3333037781" collectionCode="ZMLU" collectorName="M. - E. Walsh" country="Indonesia" latitude="-0.49638888" location="Samarinda" longLatPrecision="21" longitude="117.13972" municipality="Samarinda" pageId="28" pageNumber="29" specimenCount="1" specimenCount-female="1" stateProvince="Kalimantan Utara" typeStatus="paratype">
<specimenCount id="9D3BFD3BFFF0FFE0FD32F07DFD2AFCCA" box="[722,767,792,819]" pageId="28" pageNumber="29" type="female">1 ♀</specimenCount>
; “ 
<collectingMunicipality id="6BE6ACC8FFF0FFE0FCF8F07DFC4AFCCB" box="[792,927,792,818]" country="Indonesia" latitude="-0.49638888" longLatPrecision="21" longitude="117.13972" name="Samarinda" pageId="28" pageNumber="29" stateProvince="Kalimantan Utara">Samarinda</collectingMunicipality>
”; 
<geoCoordinate id="EE095075FFF0FFE0FC55F073FBE3FCCB" box="[949,1078,790,818]" degrees="0" direction="south" minutes="29" orientation="latitude" pageId="28" pageNumber="29" precision="15" seconds="47" value="-0.49638888">0°29′47″ S</geoCoordinate>
, 
<geoCoordinate id="EE095075FFF0FFE0FBA4F073FB03FCCA" box="[1092,1238,790,819]" degrees="117" direction="east" minutes="8" orientation="longitude" pageId="28" pageNumber="29" precision="15" seconds="23" value="117.13972">117°8′23″ E</geoCoordinate>
; 
<collectorName id="26C85364FFF0FFE0FB02F07CFAA2FCCB" box="[1250,1399,792,819]" pageId="28" pageNumber="29">M.-E. Walsh</collectorName>
leg.; 
<collectionCode id="ED2CAE77FFF0FFE0FF18F05BFE9AFCA1" box="[248,335,830,856]" country="Sweden" httpUri="http://grbio.org/cool/nzgn-b6sw" name="Lunds Universitet, Zoologiska Institutionen" pageId="28" pageNumber="29">ZMLU</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8B8236B2FFF0FFE0FF5DF0E5FCAFFC1C" blockId="28.[189,1399,895,997]" pageId="28" pageNumber="29">
<materialsCitation id="3B553CEFFFF0FFE0FF5DF0E5FD80FC39" ID-GBIF-Occurrence="3333037345" collectingDate="1970-07-09" collectionCode="MNHN-EO-ENSIF" collectorName="Baebe &amp; Ind. Mus." country="Malaysia" latitude="1.7666667" location="Kapit" longLatPrecision="21" longitude="113.53667" municipality="Kapit" pageId="28" pageNumber="29" specimenCode="MNHN-EO-ENSIF1750" specimenCount="1" specimenCount-male="1" stateProvince="Sarawak">
<collectingCountry id="F32A7622FFF0FFE0FF5DF0E5FE8BFC63" box="[189,350,896,922]" name="Malaysia" pageId="28" pageNumber="29">MALAYSIA</collectingCountry>
– 
<emphasis id="B949EAA0FFF0FFE0FE9DF0E5FE39FC63" bold="true" box="[381,492,896,922]" pageId="28" pageNumber="29">
<collectingRegion id="49F9F850FFF0FFE0FE9DF0E5FE39FC63" box="[381,492,896,922]" country="Malaysia" name="Sarawak" pageId="28" pageNumber="29">Sarawak</collectingRegion>
</emphasis>
• 
<specimenCount id="9D3BFD3BFFF0FFE0FDE5F0E5FDE0FC62" box="[517,565,896,923]" pageId="28" pageNumber="29" type="male">1 ♂</specimenCount>
; “ 
<collectingMunicipality id="6BE6ACC8FFF0FFE0FDACF0E5FD40FC63" box="[588,661,896,922]" pageId="28" pageNumber="29">Kapit</collectingMunicipality>
”; 
<geoCoordinate id="EE095075FFF0FFE0FD4DF01AFCF0FC62" box="[685,805,895,923]" degrees="1" direction="north" minutes="46" orientation="latitude" pageId="28" pageNumber="29" precision="15" seconds="0" value="1.7666667">1°46′0″ N</geoCoordinate>
, 
<geoCoordinate id="EE095075FFF0FFE0FCD1F01AFC07FC62" box="[817,978,895,923]" degrees="113" direction="east" minutes="32" orientation="longitude" pageId="28" pageNumber="29" precision="15" seconds="12" value="113.53667">113°32′12″ E</geoCoordinate>
; 
<date id="FF831072FFF0FFE0FC3FF0E5FBB1FC63" box="[991,1124,896,922]" pageId="28" pageNumber="29" value="1970-07-09">
<collectingDate id="EFC7E99AFFF0FFE0FC3FF0E5FBB1FC63" box="[991,1124,896,922]" pageId="28" pageNumber="29" value="1970-07-09">9 Jul. 1970</collectingDate>
</date>
; 
<collectorName id="26C85364FFF0FFE0FB91F0E5FB6FFC63" box="[1137,1210,896,922]" pageId="28" pageNumber="29">Baebe</collectorName>
leg.; 
<collectorName id="26C85364FFF0FFE0FB1DF0E5FAA7FC62" box="[1277,1394,896,923]" pageId="28" pageNumber="29">Ind. Mus.</collectorName>
; PII: N56; 
<specimenCode id="DB9B9EC9FFF0FFE0FED0F0C0FD80FC39" box="[304,597,933,960]" collectionCode="MNHN-EO-ENSIF" pageId="28" pageNumber="29">MNHN-EO-ENSIF1750</specimenCode>
</materialsCitation>
. – 
<materialsCitation id="3B553CEFFFF0FFE0FD93F0C0FCA3FC1C" ID-GBIF-Occurrence="3333037827" collectingDate="1987-04-02" collectingDateMax="1987-04-13" collectingDateMin="1987-04-02" collectionCode="RMNH" collectorName="C. v. Achterberg" country="Malaysia" elevation="1200" latitude="4.4058332" location="West" longLatPrecision="21" longitude="115.72972" municipality="Long Pa Sia" pageId="28" pageNumber="29" specimenCount="1" specimenCount-male="1" stateProvince="Sabah">
<emphasis id="B949EAA0FFF0FFE0FD93F0C0FD14FC46" bold="true" box="[627,705,933,959]" pageId="28" pageNumber="29">
<collectingRegion id="49F9F850FFF0FFE0FD93F0C0FD14FC46" box="[627,705,933,959]" country="Malaysia" name="Sabah" pageId="28" pageNumber="29">Sabah</collectingRegion>
</emphasis>
• 
<specimenCount id="9D3BFD3BFFF0FFE0FD39F0C0FCD2FC39" box="[729,775,933,960]" pageId="28" pageNumber="29" type="male">1 ♂</specimenCount>
; “nr. 
<collectingMunicipality id="6BE6ACC8FFF0FFE0FCA4F0C3FC04FC46" box="[836,977,933,960]" pageId="28" pageNumber="29">Long Pa Sia</collectingMunicipality>
(
<location id="8EE26069FFF0FFE0FC00F0C0FBF6FC46" LSID="urn:lsid:plazi:treatment:039487A4FFF7FFE2FDC5F533FEECFE3A:8EE26069FFF0FFE0FC00F0C0FBF6FC46" box="[992,1059,933,959]" country="Malaysia" latitude="4.4058332" longLatPrecision="21" longitude="115.72972" municipality="Long Pa Sia" name="West" pageId="28" pageNumber="29" stateProvince="Sabah">West</location>
)”; 
<geoCoordinate id="EE095075FFF0FFE0FBA1F0C1FB12FC39" box="[1089,1223,932,960]" degrees="4" direction="north" minutes="24" orientation="latitude" pageId="28" pageNumber="29" precision="15" seconds="21" value="4.4058332">4°24′21″ N</geoCoordinate>
, 
<geoCoordinate id="EE095075FFF0FFE0FB32F0C1FAA4FC39" box="[1234,1393,932,960]" degrees="115" direction="east" minutes="43" orientation="longitude" pageId="28" pageNumber="29" precision="15" seconds="47" value="115.72972">115°43′47″ E</geoCoordinate>
; “c. 
<quantity id="4CC59B57FFF0FFE0FF06F0AEFEE8FC1C" box="[230,317,971,997]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="28" pageNumber="29" unit="m" value="1200.0">
<elevation id="0010D181FFF0FFE0FF06F0AEFEE8FC1C" box="[230,317,971,997]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="28" pageNumber="29" unit="m" value="1200.0">1200m</elevation>
</quantity>
”; 
<date id="FF831072FFF0FFE0FEB4F0AEFDD8FC1C" box="[340,525,971,997]" pageId="28" pageNumber="29" value="1987-04-02" valueMax="1987-04-13" valueMin="1987-04-02">
<collectingDate id="EFC7E99AFFF0FFE0FEB4F0AEFDD8FC1C" box="[340,525,971,997]" pageId="28" pageNumber="29" value="1987-04-02" valueMax="1987-04-13" valueMin="1987-04-02">2–13 Apr. 1987</collectingDate>
</date>
; 
<collectorName id="26C85364FFF0FFE0FDF9F0AEFD0DFC1D" box="[537,728,970,997]" pageId="28" pageNumber="29">C. v. Achterberg</collectorName>
leg.; 
<collectionCode id="ED2CAE77FFF0FFE0FCFAF0AEFCA3FC1C" box="[794,886,971,997]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="28" pageNumber="29" type="Museum">RMNH</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8B8236B2FFF0FFE0FF5DF76BFBA5FBD1" blockId="28.[189,1136,1038,1064]" box="[189,1136,1038,1064]" pageId="28" pageNumber="29">
<materialsCitation id="3B553CEFFFF0FFE0FF5DF76BFBA5FBD1" ID-GBIF-Occurrence="3333037426" box="[189,1136,1038,1064]" collectionCode="MNHN-EO-ENSIF" collectorName="Chaper" location="Borneo" municipality="Borneo" pageId="28" pageNumber="29" specimenCode="MNHN-EO-ENSIF11266" specimenCount="1" specimenCount-female="1">
NO COUNTRY • 
<specimenCount id="9D3BFD3BFFF0FFE0FE7EF76BFE1EFBD1" box="[414,459,1038,1064]" pageId="28" pageNumber="29" type="female">1 ♀</specimenCount>
; “ 
<collectingMunicipality id="6BE6ACC8FFF0FFE0FE04F76BFD96FBD1" box="[484,579,1038,1064]" pageId="28" pageNumber="29">Borneo</collectingMunicipality>
”; 1891; 
<collectorName id="26C85364FFF0FFE0FD44F76BFD2CFBD1" box="[676,761,1038,1064]" pageId="28" pageNumber="29">Chaper</collectorName>
leg.; 
<specimenCode id="DB9B9EC9FFF0FFE0FCDAF76BFBB9FBD1" box="[826,1132,1038,1064]" collectionCode="MNHN-EO-ENSIF" pageId="28" pageNumber="29">MNHN-EO-ENSIF11266</specimenCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3276539FFF0FFE1FF5DF734FE99F907" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29" type="description">
<paragraph id="8B8236B2FFF0FFE0FF5DF734FE9BFB92" blockId="28.[189,334,1105,1131]" box="[189,334,1105,1131]" pageId="28" pageNumber="29">
<heading id="D0CA81DEFFF0FFE0FF5DF734FE9BFB92" bold="true" box="[189,334,1105,1131]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="B949EAA0FFF0FFE0FF5DF734FE9BFB92" bold="true" box="[189,334,1105,1131]" pageId="28" pageNumber="29">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B8236B2FFF0FFE0FF5DF7E7FC2EF9CA" blockId="28.[188,1400,1153,1587]" pageId="28" pageNumber="29">
Average size (
<figureCitation id="13062A37FFF0FFE0FE82F7E4FE6BFB65" box="[354,446,1153,1180]" captionStart="Fig" captionStartId="12.[189,232,1928,1954]" captionTargetBox="[220,1368,276,1865]" captionTargetId="figure-14@12.[220,1368,265,1889]" captionTargetPageId="12" captionText="Fig. 7. Habitus of ♂♂ in dorsal view. A. Nisitrus brunnerianus Saussure, 1878. B. N. crucius Robillard &amp; Tan sp. nov. C. N. hughtani Robillard &amp; Tan sp. nov. D. N. insignis Saussure, 1878. E. N. malaya Robillard &amp; Tan sp. nov. F. N. musicus Ingrisch, 1987. G. N. vittatus (Haan, 1844). Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5156284" httpUri="https://zenodo.org/record/5156284/files/figure.png" pageId="28" pageNumber="29">Fig. 7B</figureCitation>
). Colour pattern somewhat intermediate between 
<taxonomicName id="4C3D4D31FFF0FFE0FBE0F7E7FB6AFB62" authorityName="Saussure" authorityYear="1878" box="[1024,1215,1153,1179]" class="Insecta" family="Gryllidae" genus="Nisitrus" kingdom="Animalia" order="Orthoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="brunnerianus">
<emphasis id="B949EAA0FFF0FFE0FBE0F7E7FB6AFB62" box="[1024,1215,1153,1179]" italics="true" pageId="28" pageNumber="29">N. brunnerianus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3D4D31FFF0FFE0FB16F7E7FABBFB62" baseAuthorityName="Haan" baseAuthorityYear="1844" box="[1270,1390,1154,1179]" class="Insecta" family="Gryllidae" genus="Nisitra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="vittatus">
<emphasis id="B949EAA0FFF0FFE0FB16F7E7FABBFB62" box="[1270,1390,1154,1179]" italics="true" pageId="28" pageNumber="29">N. vittatus</emphasis>
</taxonomicName>
. Vertex black with cream-coloured margin around eyes (
<figureCitation id="13062A37FFF0FFE0FCA9F7C3FC61FB39" box="[841,948,1190,1216]" captionStart="Fig" captionStartId="16.[189,232,1892,1918]" captionTargetBox="[223,1354,324,1818]" captionTargetId="figure-15@16.[197,1389,265,1854]" captionTargetPageId="16" captionText="Fig. 11. Head and pronotum in dorsal view. A. Nisitrus brunnerianus Saussure, 1878. B. N. crucius Robillard &amp; Tan sp. nov. C. N. danum Robillard &amp; Tan sp. nov. D. N. hughtani Robillard &amp; Tan sp. nov. E. N. insignis Saussure, 1878. F. N. malaya Robillard &amp; Tan sp. nov. G. N. musicus Ingrisch, 1987. H. N. rindu Robillard &amp; Tan sp. nov. I. N. vittatus (Haan, 1844). Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5156298" httpUri="https://zenodo.org/record/5156298/files/figure.png" pageId="28" pageNumber="29">Fig. 11B</figureCitation>
). Fastigium velvety black with creamcoloured margins. Scapes black dorsally, sometimes reddish ventrally or along dorso-inner margins. Antennae black. Frons yellow, face part of fastigium black between eyes, with a cross-shaped black pattern above mouthparts; clypeus sometimes darkened, mouthparts dark red brown to black (
<figureCitation id="13062A37FFF0FFE0FAE0F670FABEFAC9" box="[1280,1387,1301,1328]" captionStart="Fig" captionStartId="15.[189,232,1742,1768]" captionTargetBox="[216,1388,279,1679]" captionTargetId="figure-11@15.[189,1398,263,1705]" captionTargetPageId="15" captionText="Fig. 10. Face in anterior view. A. Nisitrus brunnerianus Saussure, 1878. B. N. crucius Robillard &amp; Tan sp. nov. C. N. danum Robillard &amp; Tan sp. nov. D. N. hughtani Robillard &amp; Tan sp. nov. E. N. insignis Saussure, 1878. F. N. malaya Robillard &amp; Tan sp. nov. G. N. musicus Ingrisch, 1987. H. N. rindu Robillard &amp; Tan sp. nov. I. N. vittatus (Haan, 1844). Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5156294" httpUri="https://zenodo.org/record/5156294/files/figure.png" pageId="28" pageNumber="29">Fig. 10B</figureCitation>
). Maxillary palpi cream-coloured. Head lateral side generally black. Pronotal disk black with white setae (
<figureCitation id="13062A37FFF0FFE0FF26F63AFEFAFA83" box="[198,303,1375,1402]" captionStart="Fig" captionStartId="16.[189,232,1892,1918]" captionTargetBox="[223,1354,324,1818]" captionTargetId="figure-15@16.[197,1389,265,1854]" captionTargetPageId="16" captionText="Fig. 11. Head and pronotum in dorsal view. A. Nisitrus brunnerianus Saussure, 1878. B. N. crucius Robillard &amp; Tan sp. nov. C. N. danum Robillard &amp; Tan sp. nov. D. N. hughtani Robillard &amp; Tan sp. nov. E. N. insignis Saussure, 1878. F. N. malaya Robillard &amp; Tan sp. nov. G. N. musicus Ingrisch, 1987. H. N. rindu Robillard &amp; Tan sp. nov. I. N. vittatus (Haan, 1844). Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5156298" httpUri="https://zenodo.org/record/5156298/files/figure.png" pageId="28" pageNumber="29">Fig. 11B</figureCitation>
). Lateral lobes of pronotum with dorsal third black, cream-coloured ventrally (
<figureCitation id="13062A37FFF0FFE0FB5FF63AFAFFFA83" box="[1215,1322,1375,1402]" captionStart="Fig" captionStartId="17.[189,232,1645,1671]" captionTargetBox="[210,1398,290,1606]" captionTargetId="figure-11@17.[189,1398,265,1606]" captionTargetPageId="17" captionText="Fig. 12. Body in lateral view. A. Nisitrus brunnerianus Saussure, 1878. B. N. crucius Robillard &amp; Tan sp. nov. C. N. danum Robillard &amp; Tan sp. nov. D. N. hughtani Robillard &amp; Tan sp. nov. E. N. insignis Saussure, 1878. F. N. malaya Robillard &amp; Tan sp. nov. G. N. musicus Ingrisch, 1987. H. N. rindu Robillard &amp; Tan sp. nov. I. N. vittatus (Haan, 1844). Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5156302" httpUri="https://zenodo.org/record/5156302/files/figure.png" pageId="28" pageNumber="29">Fig. 12B</figureCitation>
). Legs red brown, sometimes internal-basal area black. FIIIs dark red brown, knees dark brown to black; TIIIs brown with black spines and spurs, dark brown to black near distal end; tarsomeres dark brown to black. Hindwings dark hyaline, with a rounded transparent window near internal-basal area; longer than FWs, the dark brown tail exceeding the FWs about twice as long as pronotum. Tergites black, with a pale band laterally of variable thickness; sternites pale, with a black median area.
</paragraph>
<paragraph id="8B8236B2FFF0FFE0FF0AF539FEFFF98F" blockId="28.[189,1400,1628,2025]" box="[234,298,1628,1654]" pageId="28" pageNumber="29">
<emphasis id="B949EAA0FFF0FFE0FF0AF539FEFFF98F" bold="true" box="[234,298,1628,1654]" pageId="28" pageNumber="29">Male</emphasis>
</paragraph>
<paragraph id="8B8236B2FFF0FFE0FF5DF5E4FAF8F811" blockId="28.[189,1400,1628,2025]" pageId="28" pageNumber="29">
FOREWINGS. FW colouration (
<figureCitation id="13062A37FFF0FFE0FDCDF5E4FD48F962" box="[557,669,1665,1691]" captionStart="Fig" captionStartId="18.[189,231,1928,1954]" captionTargetBox="[243,1357,300,1857]" captionTargetId="figure-15@18.[219,1368,265,1891]" captionTargetPageId="18" captionText="Fig. 13. Male tegmina in dorsal view. A. Nisitrus brunnerianus Saussure, 1878.B. N. crucius Robillard &amp; Tan sp. nov. C. N. hughtani Robillard &amp; Tan sp. nov. D. N. insignis Saussure, 1878. E. N. malaya Robillard &amp; Tan sp. nov. F. N. musicus Ingrisch, 1987. G. N. vittatus (Haan, 1844). Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5156304" httpUri="https://zenodo.org/record/5156304/files/figure.png" pageId="28" pageNumber="29">Fig. 13B</figureCitation>
): dorsal field cells mostly transparent; veins mostly creamcoloured, sometimes black. Basal area brown, basally cream-coloured, brown towards distal part. R and Sc projections cream-coloured. Lateral field hyaline, with dorsal parts infumated cream-coloured. FW venation (
<figureCitation id="13062A37FFF0FFE0FE88F595FE00F8F3" box="[360,469,1776,1802]" captionStart="Fig" captionStartId="18.[189,231,1928,1954]" captionTargetBox="[243,1357,300,1857]" captionTargetId="figure-15@18.[219,1368,265,1891]" captionTargetPageId="18" captionText="Fig. 13. Male tegmina in dorsal view. A. Nisitrus brunnerianus Saussure, 1878.B. N. crucius Robillard &amp; Tan sp. nov. C. N. hughtani Robillard &amp; Tan sp. nov. D. N. insignis Saussure, 1878. E. N. malaya Robillard &amp; Tan sp. nov. F. N. musicus Ingrisch, 1987. G. N. vittatus (Haan, 1844). Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5156304" httpUri="https://zenodo.org/record/5156304/files/figure.png" pageId="28" pageNumber="29">Fig. 13B</figureCitation>
): 1A somewhat straight. Harp slightly longer than wide, with 2 oblique veins, distal one bifurcate basally. Cell c1 long and wide, c2 diamond-shaped; mirror (d1) longer than wide, not rounded, generally separated into two parts by a distinct straight transverse vein, the posterior part somewhat rectangular, much shorter than anterior part. Cell d2 narrower than d1, usually subdivided by accessory veins. Apical field short and rounded, with 3 wide cell alignments posterior to mirror and a narrow apical alignment; veins cream-coloured. Lateral field with 5 projections of Sc. Epiproct black. Subgenital plate pale cream-coloured with median area and anterior and posterior margins black.
</paragraph>
<caption id="DF42663AFFF1FFE1FF5DF26CFAFEFEDD" ID-Table-UUID="DF42663AFFF1FFE1FF5DF26CFAFEFEDD" box="[189,1323,265,292]" httpUri="http://table.plazi.org/id/DF42663AFFF1FFE1FF5DF26CFAFEFEDD" pageId="29" pageNumber="30" startId="29.[189,258,265,291]" targetBox="[202,1376,331,543]" targetIsTable="true" targetPageId="29">
<paragraph id="8B8236B2FFF1FFE1FF5DF26CFAFEFEDD" blockId="29.[189,1323,265,292]" box="[189,1323,265,292]" pageId="29" pageNumber="30">
<emphasis id="B949EAA0FFF1FFE1FF5DF26CFECAFEDA" bold="true" box="[189,287,265,291]" pageId="29" pageNumber="30">Table 4.</emphasis>
Measurements of 
<taxonomicName id="4C3D4D31FFF1FFE1FE19F26FFCAEFEDD" authority="Robillard &amp; Tan" authorityName="Robillard &amp; Tan" authorityYear="2021" box="[505,891,265,292]" class="Insecta" family="Gryllidae" genus="Nisitrus" kingdom="Animalia" order="Orthoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="crucius" status="sp. nov.">
<emphasis id="B949EAA0FFF1FFE1FE19F26FFD66FEDA" box="[505,691,266,291]" italics="true" pageId="29" pageNumber="30">Nisitrus crucius</emphasis>
Robillard &amp; Tan
</taxonomicName>
<taxonomicNameLabel id="A27A57DBFFF1FFE1FC61F26FFC09FEDD" box="[897,988,266,292]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
(in mm, means in brackets).
</paragraph>
</caption>
<paragraph id="8B8236B2FFF1FFE1FE78F22EFA9EFDE6" pageId="29" pageNumber="30">
<table id="F93DC412FFF10003FF2AF22EFAB5FDE6" box="[202,1376,331,543]" gridcols="9" gridrows="4" pageId="29" pageNumber="30" rowsContinueIn="29.[202,1175,559,813]">
<tr id="350D34F0FFF10003FF2AF22EFAB5FE9A" box="[202,1376,331,355]" gridrow="0" pageId="29" pageNumber="30" rowspan-0="1">
<th id="76DC5D8CFFF10003FE74F22EFE31FE9A" box="[404,484,331,355]" gridcol="1" gridrow="0" pageId="29" pageNumber="30">
<emphasis id="B949EAA0FFF1FFE1FE78F22EFE37FE9A" bold="true" box="[408,482,331,355]" pageId="29" pageNumber="30">PronL</emphasis>
</th>
<th id="76DC5D8CFFF10003FDE9F22EFD8EFE9A" box="[521,603,331,355]" gridcol="2" gridrow="0" pageId="29" pageNumber="30">
<emphasis id="B949EAA0FFF1FFE1FDE9F22EFD8EFE9A" bold="true" box="[521,603,331,355]" pageId="29" pageNumber="30">PronW</emphasis>
</th>
<th id="76DC5D8CFFF10003FD9DF22EFD33FE9A" box="[637,742,331,355]" gridcol="3" gridrow="0" pageId="29" pageNumber="30">
<emphasis id="B949EAA0FFF1FFE1FD74F22EFD04FE9A" bold="true" box="[660,721,331,355]" pageId="29" pageNumber="30">FWL</emphasis>
</th>
<th id="76DC5D8CFFF10003FCE7F22EFC82FE9A" box="[775,855,331,355]" gridcol="4" gridrow="0" pageId="29" pageNumber="30">
<emphasis id="B949EAA0FFF1FFE1FCECF22EFC87FE9A" bold="true" box="[780,850,331,355]" pageId="29" pageNumber="30">FWW</emphasis>
</th>
<th id="76DC5D8CFFF10003FC9DF22EFC18FE9A" box="[893,973,331,355]" gridcol="5" gridrow="0" pageId="29" pageNumber="30">
<emphasis id="B949EAA0FFF1FFE1FC64F22EFC10FE9A" bold="true" box="[900,965,331,355]" pageId="29" pageNumber="30">HWT</emphasis>
</th>
<th id="76DC5D8CFFF10003FC11F22EFB89FE9A" box="[1009,1116,331,355]" gridcol="6" gridrow="0" pageId="29" pageNumber="30">
<emphasis id="B949EAA0FFF1FFE1FBE6F22EFB92FE9A" bold="true" box="[1030,1095,331,355]" pageId="29" pageNumber="30">FIIIL</emphasis>
</th>
<th id="76DC5D8CFFF10003FB9FF22EFB1AFE9A" box="[1151,1231,331,355]" gridcol="7" gridrow="0" pageId="29" pageNumber="30">
<emphasis id="B949EAA0FFF1FFE1FB62F22EFB19FE9A" bold="true" box="[1154,1228,331,355]" pageId="29" pageNumber="30">FIIIW</emphasis>
</th>
<th id="76DC5D8CFFF10003FB16F22EFAB5FE9A" box="[1270,1376,331,355]" gridcol="8" gridrow="0" pageId="29" pageNumber="30">
<emphasis id="B949EAA0FFF1FFE1FAEAF22EFA98FE9A" bold="true" box="[1290,1357,331,355]" pageId="29" pageNumber="30">TIIIL</emphasis>
</th>
</tr>
<tr id="350D34F0FFF10003FF2AF211FAB5FE74" box="[202,1376,372,397]" gridrow="1" pageId="29" pageNumber="30">
<th id="76DC5D8CFFF10003FF2AF211FEA6FE74" box="[202,371,372,397]" gridcol="0" gridrow="1" pageId="29" pageNumber="30">Male holotype</th>
<td id="76DC5D8CFFF10003FE74F211FE31FE74" box="[404,484,372,397]" gridcol="1" gridrow="1" pageId="29" pageNumber="30">2.1</td>
<td id="76DC5D8CFFF10003FDE9F211FD8EFE74" box="[521,603,372,397]" gridcol="2" gridrow="1" pageId="29" pageNumber="30">2.9</td>
<td id="76DC5D8CFFF10003FD9DF211FD33FE74" box="[637,742,372,397]" gridcol="3" gridrow="1" pageId="29" pageNumber="30">10.8</td>
<td id="76DC5D8CFFF10003FCE7F211FC82FE74" box="[775,855,372,397]" gridcol="4" gridrow="1" pageId="29" pageNumber="30">3.7</td>
<td id="76DC5D8CFFF10003FC9DF211FC18FE74" box="[893,973,372,397]" gridcol="5" gridrow="1" pageId="29" pageNumber="30">6.9</td>
<td id="76DC5D8CFFF10003FC11F211FB89FE74" box="[1009,1116,372,397]" gridcol="6" gridrow="1" pageId="29" pageNumber="30">13.0</td>
<td id="76DC5D8CFFF10003FB9FF211FB1AFE74" box="[1151,1231,372,397]" gridcol="7" gridrow="1" pageId="29" pageNumber="30">2.3</td>
<td id="76DC5D8CFFF10003FB16F211FAB5FE74" box="[1270,1376,372,397]" gridcol="8" gridrow="1" pageId="29" pageNumber="30">12.0</td>
</tr>
<tr id="350D34F0FFF10003FF2AF2F8FAB5FE2F" box="[202,1376,413,470]" gridrow="2" pageId="29" pageNumber="30">
<th id="76DC5D8CFFF10003FF2AF2F8FEA6FE2F" box="[202,371,413,470]" gridcol="0" gridrow="2" pageId="29" pageNumber="30">Males (n = 3)</th>
<td id="76DC5D8CFFF10003FE74F2F8FE31FE2F" box="[404,484,413,470]" gridcol="1" gridrow="2" pageId="29" pageNumber="30">1.7–2.1 (1.9)</td>
<td id="76DC5D8CFFF10003FDE9F2F8FD8EFE2F" box="[521,603,413,470]" gridcol="2" gridrow="2" pageId="29" pageNumber="30">2.9–3.4 (3.1)</td>
<td id="76DC5D8CFFF10003FD9DF2F8FD33FE2F" box="[637,742,413,470]" gridcol="3" gridrow="2" pageId="29" pageNumber="30">9.9–10.8 (10.3)</td>
<td id="76DC5D8CFFF10003FCE7F2F8FC82FE2F" box="[775,855,413,470]" gridcol="4" gridrow="2" pageId="29" pageNumber="30">3.7–4.2 (4.0)</td>
<td id="76DC5D8CFFF10003FC9DF2F8FC18FE2F" box="[893,973,413,470]" gridcol="5" gridrow="2" pageId="29" pageNumber="30">5.3–7.0 (6.4)</td>
<td id="76DC5D8CFFF10003FC11F2F8FB89FE2F" box="[1009,1116,413,470]" gridcol="6" gridrow="2" pageId="29" pageNumber="30">12.5–13.0 (12.7)</td>
<td id="76DC5D8CFFF10003FB9FF2F8FB1AFE2F" box="[1151,1231,413,470]" gridcol="7" gridrow="2" pageId="29" pageNumber="30">2.3–2.8 (2.5)</td>
<td id="76DC5D8CFFF10003FB16F2F8FAB5FE2F" box="[1270,1376,413,470]" gridcol="8" gridrow="2" pageId="29" pageNumber="30">12.0–13.0 (12.5)</td>
</tr>
<tr id="350D34F0FFF10003FF2AF282FAB5FDE6" box="[202,1376,487,543]" gridrow="3" pageId="29" pageNumber="30">
<th id="76DC5D8CFFF10003FF2AF282FEA6FDE6" box="[202,371,487,543]" gridcol="0" gridrow="3" pageId="29" pageNumber="30">Females (n = 5)</th>
<td id="76DC5D8CFFF10003FE74F282FE31FDE6" box="[404,484,487,543]" gridcol="1" gridrow="3" pageId="29" pageNumber="30">1.9–2.1 (2.0)</td>
<td id="76DC5D8CFFF10003FDE9F282FD8EFDE6" box="[521,603,487,543]" gridcol="2" gridrow="3" pageId="29" pageNumber="30">2.8–3.5 (3.3)</td>
<td id="76DC5D8CFFF10003FD9DF282FD33FDE6" box="[637,742,487,543]" gridcol="3" gridrow="3" pageId="29" pageNumber="30">10.5–11.7 (11.3)</td>
<td id="76DC5D8CFFF10003FCE7F282FC82FDE6" box="[775,855,487,543]" gridcol="4" gridrow="3" pageId="29" pageNumber="30">3.0–3.3 (3.2)</td>
<td id="76DC5D8CFFF10003FC9DF282FC18FDE6" box="[893,973,487,543]" gridcol="5" gridrow="3" pageId="29" pageNumber="30">6.5–8.0 (7.1)</td>
<td id="76DC5D8CFFF10003FC11F282FB89FDE6" box="[1009,1116,487,543]" gridcol="6" gridrow="3" pageId="29" pageNumber="30">13.1–14.5 (13.9)</td>
<td id="76DC5D8CFFF10003FB9FF282FB1AFDE6" box="[1151,1231,487,543]" gridcol="7" gridrow="3" pageId="29" pageNumber="30">2.6–2.9 (2.7)</td>
<td id="76DC5D8CFFF10003FB16F282FAB5FDE6" box="[1270,1376,487,543]" gridcol="8" gridrow="3" pageId="29" pageNumber="30">13.0–15.0 (14.1)</td>
</tr>
</table>
</paragraph>
<paragraph id="8B8236B2FFF1FFE1FE4BF13CFB57FCD4" pageId="29" pageNumber="30">
<table id="F93DC412FFF10003FF2AF14AFB42FCD4" box="[202,1175,559,813]" gridcols="7" gridrows="4" pageId="29" pageNumber="30" rowsContinueFrom="29.[202,1376,331,543]">
<tr id="350D34F0FFF10003FF2AF14AFB42FD88" box="[202,1175,559,625]" gridrow="0" pageId="29" pageNumber="30" rowspan-0="1">
<th id="76DC5D8CFFF10003FE7AF14AFE08FD88" box="[410,477,559,625]" gridcol="1" gridrow="0" pageId="29" pageNumber="30">
<emphasis id="B949EAA0FFF1FFE1FE4BF13CFE18FD88" bold="true" box="[427,461,601,625]" pageId="29" pageNumber="30">Ias</emphasis>
</th>
<th id="76DC5D8CFFF10003FE1FF14AFD3EFD88" box="[511,747,559,625]" colspan="2" colspanRight="1" gridcol="2" gridrow="0" pageId="29" pageNumber="30">
<emphasis id="B949EAA0FFF1FFE1FE1FF155FD3EFDB1" bold="true" box="[511,747,559,584]" pageId="29" pageNumber="30">Dorsal spines of TIII</emphasis>
<emphasis id="B949EAA0FFF1FFE1FDC0F13CFD91FD88" bold="true" box="[544,580,601,625]" pageId="29" pageNumber="30">Ibs</emphasis>
<emphasis id="B949EAA0FFF1FFE1FD7BF13DFD12FD89" bold="true" box="[667,711,600,624]" pageId="29" pageNumber="30">Oas</emphasis>
</th>
<th id="76DC5D8CFFF10003FCF8F14AFC93FD88" box="[792,838,559,625]" gridcol="4" gridrow="0" pageId="29" pageNumber="30">
<emphasis id="B949EAA0FFF1FFE1FCF8F13DFC93FD89" bold="true" box="[792,838,600,624]" pageId="29" pageNumber="30">Obs</emphasis>
</th>
<th id="76DC5D8CFFF10003FC62F14AFC1DFD88" box="[898,968,559,625]" gridcol="5" gridrow="0" pageId="29" pageNumber="30">
<emphasis id="B949EAA0FFF1FFE1FC62F13CFC1DFD88" bold="true" box="[898,968,601,625]" pageId="29" pageNumber="30">TaIIIs</emphasis>
</th>
<th id="76DC5D8CFFF10003FBCCF14AFB42FD88" box="[1068,1175,559,625]" gridcol="6" gridrow="0" pageId="29" pageNumber="30">
<emphasis id="B949EAA0FFF1FFE1FBAFF13DFBA3FD89" bold="true" box="[1103,1142,600,624]" pageId="29" pageNumber="30">OL</emphasis>
</th>
</tr>
<tr id="350D34F0FFF10003FF2AF1E7FB42FD63" box="[202,1175,642,666]" gridrow="1" pageId="29" pageNumber="30">
<th id="76DC5D8CFFF10003FF2AF1E7FEA6FD63" box="[202,371,642,666]" gridcol="0" gridrow="1" pageId="29" pageNumber="30">Male holotype</th>
<td id="76DC5D8CFFF10003FE7AF1E7FE08FD63" box="[410,477,642,666]" gridcol="1" gridrow="1" pageId="29" pageNumber="30">16</td>
<td id="76DC5D8CFFF10003FE1FF1E7FD93FD63" box="[511,582,642,666]" gridcol="2" gridrow="1" pageId="29" pageNumber="30">3-5</td>
<td id="76DC5D8CFFF10003FD70F1E7FD3EFD63" box="[656,747,642,666]" gridcol="3" gridrow="1" pageId="29" pageNumber="30">23</td>
<td id="76DC5D8CFFF10003FCF8F1E7FC93FD63" box="[792,838,642,666]" gridcol="4" gridrow="1" pageId="29" pageNumber="30">3–5</td>
<td id="76DC5D8CFFF10003FC62F1E7FC1DFD63" box="[898,968,642,666]" gridcol="5" gridrow="1" pageId="29" pageNumber="30">4</td>
<td id="76DC5D8CFFF10003FBCCF1E7FB42FD63" box="[1068,1175,642,666]" gridcol="6" gridrow="1" pageId="29" pageNumber="30">–</td>
</tr>
<tr id="350D34F0FFF10003FF2AF1CEFB42FD1D" box="[202,1175,683,740]" gridrow="2" pageId="29" pageNumber="30">
<th id="76DC5D8CFFF10003FF2AF1CEFEA6FD1D" box="[202,371,683,740]" gridcol="0" gridrow="2" pageId="29" pageNumber="30">Males (n = 3)</th>
<td id="76DC5D8CFFF10003FE7AF1CEFE08FD1D" box="[410,477,683,740]" gridcol="1" gridrow="2" pageId="29" pageNumber="30">13–16 (15)</td>
<td id="76DC5D8CFFF10003FE1FF1CEFD93FD1D" box="[511,582,683,740]" gridcol="2" gridrow="2" pageId="29" pageNumber="30">3–5</td>
<td id="76DC5D8CFFF10003FD70F1CEFD3EFD1D" box="[656,747,683,740]" gridcol="3" gridrow="2" pageId="29" pageNumber="30">17–23 (20)</td>
<td id="76DC5D8CFFF10003FCF8F1CEFC93FD1D" box="[792,838,683,740]" gridcol="4" gridrow="2" pageId="29" pageNumber="30">3–7</td>
<td id="76DC5D8CFFF10003FC62F1CEFC1DFD1D" box="[898,968,683,740]" gridcol="5" gridrow="2" pageId="29" pageNumber="30">4</td>
<td id="76DC5D8CFFF10003FBCCF1CEFB42FD1D" box="[1068,1175,683,740]" gridcol="6" gridrow="2" pageId="29" pageNumber="30">–</td>
</tr>
<tr id="350D34F0FFF10003FF2AF191FB42FCD4" box="[202,1175,756,813]" gridrow="3" pageId="29" pageNumber="30">
<th id="76DC5D8CFFF10003FF2AF191FEA6FCD4" box="[202,371,756,813]" gridcol="0" gridrow="3" pageId="29" pageNumber="30">Females (n = 5)</th>
<td id="76DC5D8CFFF10003FE7AF191FE08FCD4" box="[410,477,756,813]" gridcol="1" gridrow="3" pageId="29" pageNumber="30">11–12 (12)</td>
<td id="76DC5D8CFFF10003FE1FF191FD93FCD4" box="[511,582,756,813]" gridcol="2" gridrow="3" pageId="29" pageNumber="30">3–4</td>
<td id="76DC5D8CFFF10003FD70F191FD3EFCD4" box="[656,747,756,813]" gridcol="3" gridrow="3" pageId="29" pageNumber="30">19–20 (20)</td>
<td id="76DC5D8CFFF10003FCF8F191FC93FCD4" box="[792,838,756,813]" gridcol="4" gridrow="3" pageId="29" pageNumber="30">3–8</td>
<td id="76DC5D8CFFF10003FC62F191FC1DFCD4" box="[898,968,756,813]" gridcol="5" gridrow="3" pageId="29" pageNumber="30">3–4</td>
<td id="76DC5D8CFFF10003FBCCF191FB42FCD4" box="[1068,1175,756,813]" gridcol="6" gridrow="3" pageId="29" pageNumber="30">13.8–15.0 (14.3)</td>
</tr>
</table>
</paragraph>
<paragraph id="8B8236B2FFF1FFE1FF5DF01CFC6BFB1E" blockId="29.[187,1400,888,1255]" pageId="29" pageNumber="30">
GENITALIA (
<figureCitation id="13062A37FFF1FFE1FEA4F01DFE7EFC6A" box="[324,427,888,915]" captionStart="Fig" captionStartId="21.[189,231,1640,1666]" captionTargetBox="[218,1372,288,1585]" captionTargetId="figure-11@21.[189,1398,265,1601]" captionTargetPageId="21" captionText="Fig. 16. Male genitalia in ventral view. A. Nisitrus brunnerianus Saussure, 1878. B. N. crucius Robillard &amp; Tan sp. nov. C. N. hughtani Robillard &amp; Tan sp. nov. D. N. insignis Saussure, 1878. E. N. malaya Robillard &amp; Tan sp. nov. F. N. musicus Ingrisch, 1987. G. N. vittatus (Haan, 1844). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5156316" httpUri="https://zenodo.org/record/5156316/files/figure.png" pageId="29" pageNumber="30">Fig. 16B</figureCitation>
). Pseudepiphallus sclerotized, stout (small but broad) compared to congeners, anterior margin somewhat broad and straight, posterior margin also straight, lateral margins faintly converging posteriorly. Posterior apex with paired lophi slightly longer than wide (at base), obtuse at apex, sclerotized laterally only and covered with short strong setae; narrowly spaced apart from one another. Rami swollen preapically, anterior apex somewhat truncated. Pseudepiphallic parameres narrow, divergent posteriorly, their basis membranous, with a sclerotized lobe on anterior apex. Ectophallic arc transverse, complete, with anterior and posterior margins straight. Ectophallic fold strong rounded lateral sclerites appearing bean-shaped. Ectophallic apodemes long, straight and slightly divergent basally. Endophallic sclerite large and sclerotized, its posterior apex with divergent lateral arms and with a short median expansion curved dorsally. Endophallic apodeme with lateral lamellae stout.
</paragraph>
<paragraph id="8B8236B2FFF1FFE1FF0AF675FE90FAD3" blockId="29.[189,1398,1296,1436]" box="[234,325,1296,1322]" pageId="29" pageNumber="30">
<emphasis id="B949EAA0FFF1FFE1FF0AF675FE90FAD3" bold="true" box="[234,325,1296,1322]" pageId="29" pageNumber="30">Female</emphasis>
</paragraph>
<paragraph id="8B8236B2FFF1FFE1FF5DF653FD2EFA62" blockId="29.[189,1398,1296,1436]" pageId="29" pageNumber="30">
FOREWINGS. FW with cells on dorsal field mostly infumated black, gradually becoming yellow near CuA, M and R veins; veins generally yellow (
<figureCitation id="13062A37FFF1FFE1FD07F639FC81FA8F" box="[743,852,1372,1398]" captionStart="Fig" captionStartId="19.[189,233,1894,1920]" captionTargetBox="[216,1372,287,1817]" captionTargetId="figure-11@19.[193,1394,265,1855]" captionTargetPageId="19" captionText="Fig. 14. Female tegmina in dorsal view. A. Nisitrus brunnerianus Saussure, 1878. B. N. crucius Robillard &amp; Tan sp. nov. C. N. danum Robillard &amp; Tan sp. nov. D. N. hughtani Robillard &amp; Tan sp. nov. E. N. insignis Saussure, 1878. F. N. malaya Robillard &amp; Tan sp. nov. G. N. musicus Ingrisch, 1987. H. N. rindu Robillard &amp; Tan sp. nov. I. N. vittatus (Haan, 1844). Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5156308" httpUri="https://zenodo.org/record/5156308/files/figure.png" pageId="29" pageNumber="30">Fig. 14B</figureCitation>
). Sc projections cream-coloured. Lateral field transparent, sometimes infumated black apically.
</paragraph>
<paragraph id="8B8236B2FFF1FFE1FF5DF6A0FDF5F9D2" blockId="29.[188,1399,1477,1579]" pageId="29" pageNumber="30">
GENITALIA. Ovipositor: only slightly longer than FIII. Copulatory papilla conical, smaller and stout; apex folded ventrally, short, pointed; dorsal face with a sclerotized area; ventro-anterior end forming an oval to pyriform rim (
<figureCitation id="13062A37FFF1FFE1FE63F574FDC1F9D2" box="[387,532,1553,1579]" captionStart="Fig" captionStartId="22.[189,232,1809,1835]" captionTargetBox="[201,1393,282,1740]" captionTargetId="figure-14@22.[189,1398,265,1771]" captionText="Fig. 17. Female genitalia in lateral (A, C, E, F, H, J, L, N, P) and dorsal (B, D, G, I, K, M, O, Q) views. A–B. Nisitrus brunnerianus Saussure, 1878. C–D. N. crucius Robillard &amp; Tan sp. nov. E. N. danum Robillard &amp; Tan sp. nov. F–G. N. hughtani Robillard &amp; Tan sp. nov. H–I. N. insignis Saussure, 1878. J–K. N. malaya Robillard &amp; Tan sp. nov. L–M. N. musicus Ingrisch, 1987. N–O. N. rindu Robillard &amp; Tan sp. nov. P–Q. N. vittatus (Haan, 1844). Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5165004" httpUri="https://zenodo.org/record/5165004/files/figure.png" pageId="29" pageNumber="30">Fig. 17C–D</figureCitation>
).
</paragraph>
<paragraph id="8B8236B2FFF1FFE1FF0AF531FE87F997" blockId="29.[189,338,1620,1684]" box="[234,338,1620,1646]" pageId="29" pageNumber="30">
<emphasis id="B949EAA0FFF1FFE1FF0AF531FE87F997" bold="true" box="[234,338,1620,1646]" pageId="29" pageNumber="30">Juvenile</emphasis>
</paragraph>
<paragraph id="8B8236B2FFF1FFE1FF5DF51FFEECF96D" blockId="29.[189,338,1620,1684]" box="[189,313,1658,1684]" pageId="29" pageNumber="30">Unknown.</paragraph>
<paragraph id="8B8236B2FFF1FFE1FF0AF5DAFE77F920" blockId="29.[189,418,1727,1790]" box="[234,418,1727,1753]" pageId="29" pageNumber="30">
<emphasis id="B949EAA0FFF1FFE1FF0AF5DAFE77F920" bold="true" box="[234,418,1727,1753]" pageId="29" pageNumber="30">Measurements</emphasis>
</paragraph>
<paragraph id="8B8236B2FFF1FFE1FF5DF581FE99F907" blockId="29.[189,418,1727,1790]" box="[189,332,1764,1790]" pageId="29" pageNumber="30">
See 
<tableCitation id="C6BF0309FFF1FFE1FF0EF581FE9DF907" box="[238,328,1764,1790]" captionStart="Table 4" captionStartId="29.[189,258,265,291]" captionTargetBox="[202,1376,331,543]" captionTargetPageId="29" captionText="Table 4. Measurements of Nisitrus crucius Robillard &amp; Tan sp. nov. (in mm, means in brackets)." httpUri="http://table.plazi.org/id/DF42663AFFF1FFE1FF5DF26CFAFEFEDD" pageId="29" pageNumber="30" tableUuid="DF42663AFFF1FFE1FF5DF26CFAFEFEDD">Table 4</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3276539FFF1FFE1FF5DF44DFEECF88A" box="[189,313,1832,1907]" pageId="29" pageNumber="30" type="biology_ecology">
<paragraph id="8B8236B2FFF1FFE1FF5DF44DFEF5F8BB" blockId="29.[189,288,1832,1858]" box="[189,288,1832,1858]" pageId="29" pageNumber="30">
<heading id="D0CA81DEFFF1FFE1FF5DF44DFEF5F8BB" bold="true" box="[189,288,1832,1858]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="B949EAA0FFF1FFE1FF5DF44DFEF5F8BB" bold="true" box="[189,288,1832,1858]" pageId="29" pageNumber="30">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="8B8236B2FFF1FFE1FF5DF43CFEECF88A" blockId="29.[189,313,1881,1907]" box="[189,313,1881,1907]" pageId="29" pageNumber="30">Unknown.</paragraph>
</subSubSection>
<subSubSection id="C3276539FFF1FFE1FF5DF4F8FD5CF810" box="[189,649,1949,2025]" pageId="29" pageNumber="30" type="distribution">
<paragraph id="8B8236B2FFF1FFE1FF5DF4F8FE83F84E" blockId="29.[189,342,1949,1975]" box="[189,342,1949,1975]" pageId="29" pageNumber="30">
<heading id="D0CA81DEFFF1FFE1FF5DF4F8FE83F84E" bold="true" box="[189,342,1949,1975]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="B949EAA0FFF1FFE1FF5DF4F8FE83F84E" bold="true" box="[189,342,1949,1975]" pageId="29" pageNumber="30">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8B8236B2FFF1FFE1FF5DF4AAFD5CF810" blockId="29.[189,649,1998,2025]" box="[189,649,1998,2025]" pageId="29" pageNumber="30">
Borneo (Kalimantan, 
<collectingRegion id="49F9F850FFF1FFE1FE5CF4ABFDD2F811" box="[444,519,1998,2024]" country="Malaysia" name="Sabah" pageId="29" pageNumber="30">Sabah</collectingRegion>
, 
<collectingCountry id="F32A7622FFF1FFE1FDF2F4ABFDA9F810" box="[530,636,1998,2025]" name="Malaysia" pageId="29" pageNumber="30">Sarawak</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C3276539FFF2FFE2FF5DF26CFE1BFEAB" pageId="30" pageNumber="31" type="materials_examined">
<paragraph id="8B8236B2FFF2FFE2FF5DF26CFE88FEDB" blockId="30.[189,349,264,291]" box="[189,349,264,291]" pageId="30" pageNumber="31">
<heading id="D0CA81DEFFF2FFE2FF5DF26CFE88FEDB" bold="true" box="[189,349,264,291]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="B949EAA0FFF2FFE2FF5DF26CFE88FEDB" bold="true" box="[189,349,264,291]" pageId="30" pageNumber="31">Type locality</emphasis>
</heading>
</paragraph>
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<collectingCountry id="F32A7622FFF2FFE2FF5DF25DFEE7FEAB" box="[189,306,312,338]" name="Indonesia" pageId="30" pageNumber="31">Indonesia</collectingCountry>
, 
<collectingRegion id="49F9F850FFF2FFE2FEDEF25DFE1EFEAB" box="[318,459,312,338]" country="Indonesia" name="Kalimantan Utara" pageId="30" pageNumber="31">Kalimantan</collectingRegion>
.
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<paragraph id="8B8236B2FFF2FFE2FF5DF21FFE8CFE6D" blockId="30.[189,345,378,404]" box="[189,345,378,404]" pageId="30" pageNumber="31">
<heading id="D0CA81DEFFF2FFE2FF5DF21FFE8CFE6D" bold="true" box="[189,345,378,404]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="B949EAA0FFF2FFE2FF5DF21FFE8CFE6D" bold="true" box="[189,345,378,404]" pageId="30" pageNumber="31">Calling song</emphasis>
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</paragraph>
<paragraph id="8B8236B2FFF2FFE2FF5DF2CCFEECFE3A" blockId="30.[189,313,425,451]" box="[189,313,425,451]" pageId="30" pageNumber="31">Unknown.</paragraph>
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