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<document id="AAAA1E1FB893DAD6646152BA14EDDD91" ID-DOI="10.5852/ejt.2016.199" ID-GBIF-Dataset="d88184f4-ceef-4392-a444-0ac1552778d3" ID-ISSN="2118-9773" ID-Zenodo-Dep="3840484" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1590159499057" checkinUser="valdenar" docAuthor="Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak &amp; Backeljau, Thierry" docDate="2016" docId="0385FA5C9615FFA7FDE08E8ADE26C31C" docLanguage="en" docName="ejt-2016-199.pdf.imf" docOrigin="European Journal of Taxonomy 199" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Thyropygus sutchariti Pimvichai &amp; Enghoff &amp; Panha &amp; Backeljau 2016, sp. nov." docType="treatment" docVersion="15" lastPageNumber="27" masterDocId="FFBC8224960DFFBDFF848A31DC7FC003" masterDocTitle="A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae)" masterLastPageNumber="37" masterPageNumber="1" pageNumber="25" updateTime="1734450501215" updateUser="felipe" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="8056CD7C6665EABE479C746733BEB35D">
<mods:title id="3676912B968CA0844ADEA06A25DE27F0">A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae)</mods:title>
</mods:titleInfo>
<mods:name id="A44B599E4051EF5BC460EC161080555E" type="personal">
<mods:role id="4194FFCFAC92B985881AE846866D7618">
<mods:roleTerm id="97F16C4E057552C71848A8D429D80795">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FC35152923331414BBE5707512A66D6C">Pimvichai, Piyatida</mods:namePart>
<mods:nameIdentifier id="7B733944E9A01F708E874C52C50D0EC1" type="LSID">4D882A89-E0C1-4F78-93E7-FA1B5A2650F2</mods:nameIdentifier>
<mods:affiliation id="9134E6AEF64145A0C43EBBAC72781F56">Department of Biology, Faculty of Science, Mahasarakham University, Kantharawichai District, Maha Sarakham 44150, Thailand.</mods:affiliation>
<mods:nameIdentifier id="B89540C1D4E2D88B5C481AD8D8C6729F" type="email">piyatida_pimvichai@yahoo.com</mods:nameIdentifier>
</mods:name>
<mods:name id="AE2F8A0CEEF6F65E45A24C4BB64D4CAB" type="personal">
<mods:role id="45082D5EF04682CE88CE05A6733B21CA">
<mods:roleTerm id="04FC607269A49516626939D653479B34">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6545020E10B29A845A4F54CE3CBD0429">Enghoff, Henrik</mods:namePart>
<mods:nameIdentifier id="5A28049119FCCEBB4B71606AE45F7845" type="LSID">FB09A817-000D-43C3-BCC4-2BC1E5373635</mods:nameIdentifier>
<mods:affiliation id="18E70D53D0E0F3F400A4B57DF7A7C8F7">Natural History Museum of Denmark, University of Copenhagen, Universitetsparken 15, DK- 2100 Copenhagen Ø, Denmark.</mods:affiliation>
<mods:nameIdentifier id="6974F045BF0866E283028F21B164156D" type="email">henghoff@snm.ku.dk</mods:nameIdentifier>
</mods:name>
<mods:name id="999256AB6B87C31A76ACB1BD1EB67D53" type="personal">
<mods:role id="295A2EFEF7BC4E03EAFB5A6A3CEA2C36">
<mods:roleTerm id="95537B2DA6A347BCD37ED6A302C03874">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3691B1DAA931128FF745775B5A0C75FE">Panha, Somsak</mods:namePart>
<mods:nameIdentifier id="0D8C2E5960DDDAC3845D0DFD08CF6A63" type="LSID">AC935098-D901-4F35-A414-4B0D4FE44E79</mods:nameIdentifier>
<mods:affiliation id="709F1A6DA78209E69D84C4EB8B2304DB">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand.</mods:affiliation>
<mods:nameIdentifier id="0364D422922259FA286AF06AE9A99A51" type="email">somsakp@sc.chula.ac.th</mods:nameIdentifier>
</mods:name>
<mods:name id="106211419252CE0DDB12682EAA54EAA0" type="personal">
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<mods:roleTerm id="C1A97D1015EEED4AEA3CA07A1D07DAE8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9C812E20A90D6F165A9FA2D69022AFBF">Backeljau, Thierry</mods:namePart>
<mods:nameIdentifier id="88C82FA80219E88B7FC038871347B542" type="LSID">E6F42575-36AE-4AD7-98C6-D083EF052568</mods:nameIdentifier>
<mods:affiliation id="81F1CB8261D2CEEFE692AB760CDFA673">University of Antwerp, Evolutionary Ecology Group, Groenenborgerlaan 171, B- 2020 Antwerp, Belgium and Royal Belgian Institute of Natural Sciences, OD Taxonomy and Phylogeny, and JEMU, Vautierstraat 29, B- 1000 Brussels, Belgium. &amp; Thierry.</mods:affiliation>
<mods:nameIdentifier id="7D33DD85E779DDA1A995B97F93FB2464" type="email">Thierry.Backeljau@naturalsciences.be</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="70CBE255EEF56A9F95120FCAEB619746">text</mods:typeOfResource>
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<mods:title id="662E2B16CBF3154659943B27AE715AA4">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="6E9124BBD70C5AF09F05212B9DFB3802">
<mods:date id="9301CF922AA309AA145E1159FF0C7327">2016</mods:date>
<mods:detail id="31D2619B4D56EE68964786BEDE4F2651" type="pubDate">
<mods:number id="B5FA430ABB0AB2D5031FFF4A8A88E541">2016-05-24</mods:number>
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<mods:detail id="0AC2289FCEE8D9BF599C2C122CC1D1DD" type="volume">
<mods:number id="4AE01E355A20A49A61B066BF8B0457A0">199</mods:number>
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<mods:identifier id="F90317B4635FDFBC004E410FA554C0F8" type="DOI">10.5852/ejt.2016.199</mods:identifier>
<mods:identifier id="BFC787D3C915DE7F73D82A4ABD446242" type="GBIF-Dataset">d88184f4-ceef-4392-a444-0ac1552778d3</mods:identifier>
<mods:identifier id="ECF3F50407C4B937AF13F8BBC99A10F8" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="CD198E9C868CAAA3C7354EA7479378C8" type="Zenodo-Dep">3840484</mods:identifier>
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<treatment id="0385FA5C9615FFA7FDE08E8ADE26C31C" ID-DOI="http://doi.org/10.5281/zenodo.3853328" ID-GBIF-Taxon="164249487" ID-Zenodo-Dep="3853328" LSID="urn:lsid:zoobank.org:act:6804DF2C-7787-46FE-9185-2E5C40AF5ED3" httpUri="http://treatment.plazi.org/id/0385FA5C9615FFA7FDE08E8ADE26C31C" lastPageId="26" lastPageNumber="27" pageId="24" pageNumber="25" scope_class="Diplopoda" scope_family="Harpagophoridae" scope_order="Spirostreptida" scope_subFamily="Harpagophorinae">
<subSubSection id="C33618C19615FFA5FDE08E8ADFAFC4D5" box="[612,976,1211,1238]" pageId="24" pageNumber="25" type="nomenclature">
<paragraph id="8B934B4A9615FFA5FDE08E8ADFAFC4D5" blockId="24.[371,1216,1211,1308]" box="[612,976,1211,1238]" pageId="24" pageNumber="25">
<heading id="D0DBFC269615FFA5FDE08E8ADFAFC4D5" box="[612,976,1211,1238]" centered="true" fontSize="11" level="2" pageId="24" pageNumber="25" reason="2">
<taxonomicName id="4C2C30C99615FFA5FDE08E8ADF11C4D5" ID-CoL="56T4F" authority="Pimvichai &amp; Enghoff &amp; Panha &amp; Backeljau, 2016" authorityName="Pimvichai &amp; Enghoff &amp; Panha &amp; Backeljau" authorityYear="2016" box="[612,878,1211,1238]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="sutchariti" status="sp. nov.">
<emphasis id="B95897589615FFA5FDE08E8ADF11C4D5" bold="true" box="[612,878,1211,1238]" italics="true" pageId="24" pageNumber="25">Thyropygus sutchariti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26B2A239615FFA5FCF28E8DDFAFC4D5" box="[886,976,1212,1238]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33618C19615FFA5FEF78EEFDF27C51F" pageId="24" pageNumber="25" type="description">
<paragraph id="8B934B4A9615FFA5FEF78EEFD8BFC4FA" blockId="24.[371,1216,1211,1308]" box="[371,1216,1246,1273]" pageId="24" pageNumber="25">
<uri id="FFBD47489615FFA5FEF78EEFD8BFC4FA" box="[371,1216,1246,1273]" pageId="24" pageNumber="25">
urn:lsid:zoobank.org:act:
<uuid id="FF8A719F9615FFA5FD148EEFD8BFC4FA" box="[656,1216,1246,1273]" pageId="24" pageNumber="25">6804DF2C-7787-46FE-9185-2E5C40AF5ED3</uuid>
</uri>
</paragraph>
<paragraph id="8B934B4A9615FFA5FD5F8F33DF27C51F" blockId="24.[371,1216,1211,1308]" box="[731,856,1282,1308]" pageId="24" pageNumber="25">
<figureCitation id="131757CF9615FFA5FD5F8F33DF62C51F" box="[731,797,1282,1308]" captionStart="Fig" captionStartId="25.[189,232,1928,1954]" captionTargetBox="[206,1387,494,1893]" captionTargetId="figure@25.[191,1404,481,1908]" captionTargetPageId="25" captionText="Fig. 8. Thyropygus sutchariti sp. nov., from Kaeng Krachan, holotype (CUMZ-D00090), ♂, gonopods. A. Anterior view, left telopodite removed. B. Posterior view, left telopodite removed. C. Left telopodite, posterior-mesal view. D. Left telopodite, anterior-lateral view." figureDoi="http://doi.org/10.5281/zenodo.3840500" httpUri="https://zenodo.org/record/3840500/files/figure.png" pageId="24" pageNumber="25">Fig. 8</figureCitation>
A–D
</paragraph>
</subSubSection>
<subSubSection id="C33618C19615FFA5FF398F72DD9CC5F6" pageId="24" pageNumber="25" type="diagnosis">
<paragraph id="8B934B4A9615FFA5FF398F72DD49C55E" blockId="24.[189,310,1347,1373]" box="[189,310,1347,1373]" pageId="24" pageNumber="25">
<heading id="D0DBFC269615FFA5FF398F72DD49C55E" bold="true" box="[189,310,1347,1373]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="B95897589615FFA5FF398F72DD49C55E" bold="true" box="[189,310,1347,1373]" pageId="24" pageNumber="25">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B934B4A9615FFA5FF398F43DD9CC5F6" blockId="24.[189,1399,1394,1526]" pageId="24" pageNumber="25">
A species of the
<taxonomicName id="4C2C30C99615FFA5FEF28F42DDA4C58F" authorityName="Arw" baseAuthorityName="Karsch" baseAuthorityYear="1881" box="[374,475,1395,1420]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="opinatus">
<emphasis id="B95897589615FFA5FEF28F42DDA4C58F" box="[374,475,1395,1420]" italics="true" pageId="24" pageNumber="25">opinatus</emphasis>
</taxonomicName>
subgroup. Mesal process of anterior coxal fold (
<emphasis id="B95897589615FFA5FB818F42D847C58F" box="[1029,1080,1395,1420]" italics="true" pageId="24" pageNumber="25">amp</emphasis>
) directed slightly obliquely disto-mesad, tip overlapping tip of opposite
<emphasis id="B95897589615FFA5FD458FA7DE8BC5AC" box="[705,756,1430,1455]" italics="true" pageId="24" pageNumber="25">amp</emphasis>
. Similar in this respect to
<taxonomicName id="4C2C30C99615FFA5FBA28FA7D8DAC5AC" box="[1062,1189,1429,1455]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="chelatus">
<emphasis id="B95897589615FFA5FBA28FA7D8DAC5AC" box="[1062,1189,1429,1455]" italics="true" pageId="24" pageNumber="25">T. chelatus</emphasis>
</taxonomicName>
. Differs from this species by having
<emphasis id="B95897589615FFA5FE128F88DDB6C5D1" box="[406,457,1465,1490]" italics="true" pageId="24" pageNumber="25">amp</emphasis>
as long as
<emphasis id="B95897589615FFA5FDC88F89DE0EC5D1" box="[588,625,1464,1490]" italics="true" pageId="24" pageNumber="25">alp</emphasis>
and by having the lateral process of the posterior coxal fold (
<emphasis id="B95897589615FFA5FAC38F89D913C5D1" box="[1351,1388,1464,1490]" italics="true" pageId="24" pageNumber="25">plp</emphasis>
) as a massive, broad lobe.
</paragraph>
</subSubSection>
<subSubSection id="C33618C19615FFA5FF398C2CDD40C68B" pageId="24" pageNumber="25" type="etymology">
<paragraph id="8B934B4A9615FFA5FF398C2CDD3BC634" blockId="24.[189,324,1565,1591]" box="[189,324,1565,1591]" pageId="24" pageNumber="25">
<heading id="D0DBFC269615FFA5FF398C2CDD3BC634" bold="true" box="[189,324,1565,1591]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="B95897589615FFA5FF398C2CDD3BC634" bold="true" box="[189,324,1565,1591]" pageId="24" pageNumber="25">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8B934B4A9615FFA5FF398C7ADD40C68B" blockId="24.[189,1399,1611,1672]" pageId="24" pageNumber="25">The species is named in honour of Chirasak Sutcharit in recognition of his devotion to collecting millipedes.</paragraph>
</subSubSection>
<subSubSection id="C33618C19615FFA5FF398C81DDE4C7E9" pageId="24" pageNumber="25" type="materials_examined">
<paragraph id="8B934B4A9615FFA5FF398C81DDD5C6C9" blockId="24.[189,426,1712,1738]" box="[189,426,1712,1738]" pageId="24" pageNumber="25">
<heading id="D0DBFC269615FFA5FF398C81DDD5C6C9" bold="true" box="[189,426,1712,1738]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="B95897589615FFA5FF398C81DDD5C6C9" bold="true" box="[189,426,1712,1738]" pageId="24" pageNumber="25">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8B934B4A9615FFA5FF6E8CEEDDEAC761" blockId="24.[189,1399,1759,1890]" pageId="24" pageNumber="25">
<materialsCitation id="3B4441179615FFA5FF6E8CEEDDEAC761" ID-GBIF-Occurrence="2625222302" collectingDate="2012-05-14" collectorName="C. Sutcharit &amp; Animal Systematics Research Unit" country="Thailand" latitude="12.901944" location="Kaeng Krachan" longLatPrecision="21" longitude="99.59889" municipality="Kaeng Krachan District" pageId="24" pageNumber="25" specimenCode="CUMZ-D00090" specimenCount="1" specimenCount-male="1" stateProvince="Phetchaburi Province" typeStatus="holotype">
<emphasis id="B95897589615FFA5FF6E8CEEDD23C6FA" bold="true" box="[234,348,1759,1785]" pageId="24" pageNumber="25">
<typeStatus id="5497F5E89615FFA5FF6E8CEEDD23C6FA" box="[234,348,1759,1785]" pageId="24" pageNumber="25" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F33B0BDA9615FFA5FF398D33DD26C71F" box="[189,345,1794,1820]" name="Thailand" pageId="24" pageNumber="25">THAILAND</collectingCountry>
:
<specimenCount id="9D2A80C39615FFA5FEEA8D31DDFEC71F" box="[366,385,1792,1820]" pageId="24" pageNumber="25" type="male">♂</specimenCount>
,
<collectingRegion id="49E885A89615FFA5FE188D33DEDBC71F" box="[412,676,1794,1820]" country="Thailand" name="Phetchaburi" pageId="24" pageNumber="25">Phetchaburi Province</collectingRegion>
,
<collectingMunicipality id="6BF7D1309615FFA5FD3D8D33DF99C71F" box="[697,998,1794,1820]" pageId="24" pageNumber="25">Kaeng Krachan District</collectingMunicipality>
,
<location id="8EF31D919615FFA5FC7F8D33D8C1C71F" LSID="urn:lsid:plazi:treatment:0385FA5C9615FFA7FDE08E8ADE26C31C:8EF31D919615FFA5FC7F8D33D8C1C71F" box="[1019,1214,1794,1820]" country="Thailand" latitude="12.901944" longLatPrecision="21" longitude="99.59889" municipality="Kaeng Krachan District" name="Kaeng Krachan" pageId="24" pageNumber="25" stateProvince="Phetchaburi Province">Kaeng Krachan</location>
,
<geoCoordinate id="EE182D8D9615FFA5FB508D30D90DC71F" box="[1236,1394,1793,1820]" degrees="12" direction="north" minutes="54" orientation="latitude" pageId="24" pageNumber="25" precision="15" seconds="07" value="12.901944">12°54'07&quot; N</geoCoordinate>
,
<geoCoordinate id="EE182D8D9615FFA5FF398D14DD2FC73C" box="[189,336,1829,1856]" degrees="99" direction="east" minutes="35" orientation="longitude" pageId="24" pageNumber="25" precision="15" seconds="56" value="99.59889">99°35'56&quot; E</geoCoordinate>
,
<date id="FF926D8A9615FFA5FEDB8D14DE7CC73C" box="[351,515,1829,1855]" pageId="24" pageNumber="25" value="2012-05-14">
<collectingDate id="EFD694629615FFA5FEDB8D14DE7CC73C" box="[351,515,1829,1855]" pageId="24" pageNumber="25" value="2012-05-14">14 May 2012</collectingDate>
</date>
, leg.
<collectorName id="26D92E9C9615FFA5FDC28D14DEABC73C" box="[582,724,1829,1855]" pageId="24" pageNumber="25">C. Sutcharit</collectorName>
and members of the
<collectorName id="26D92E9C9615FFA5FC528D14D909C73C" box="[982,1398,1829,1855]" pageId="24" pageNumber="25">Animal Systematics Research Unit</collectorName>
(
<specimenCode id="DB8AE3319615FFA5FF418D79DDF6C761" box="[197,393,1864,1890]" collectionCode="CUMZ-D" pageId="24" pageNumber="25">CUMZ-D00090</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8B934B4A9615FFA5FF6E8DBBDDE4C7E9" blockId="24.[189,1398,1930,2026]" pageId="24" pageNumber="25">
<materialsCitation id="3B4441179615FFA5FF6E8DBBD8E5C7C4" ID-GBIF-Occurrence="2625222302" collectingDate="2012-05-14" collectorName="C. Sutcharit &amp; Animal Systematics Research Unit" country="Thailand" latitude="12.901944" location="THAILAND" longLatPrecision="21" longitude="99.59889" municipality="Kaeng Krachan District" pageId="24" pageNumber="25" specimenCode="CUMZ-D00098" specimenCount="5" specimenCount-female="2" specimenCount-male="2" specimenCount-subadult="1" stateProvince="Phetchaburi Province" typeStatus="paratype">
<emphasis id="B95897589615FFA5FF6E8DBBDD18C7A7" bold="true" box="[234,359,1930,1956]" pageId="24" pageNumber="25">
<typeStatus id="5497F5E89615FFA5FF6E8DBBDD18C7A7" box="[234,359,1930,1956]" pageId="24" pageNumber="25" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="F33B0BDA9615FFA5FF398D9CDD26C7C4" box="[189,345,1965,1991]" name="Thailand" pageId="24" pageNumber="25">THAILAND</collectingCountry>
:
<specimenCount id="9D2A80C39615FFA5FEE18D9CDDD7C7C4" box="[357,424,1963,1991]" count="2" pageId="24" pageNumber="25" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D2A80C39615FFA5FE3C8D9CDD84C7C4" box="[440,507,1965,1991]" count="2" pageId="24" pageNumber="25" type="female">2 ♀♀</specimenCount>
,
<specimenCount id="9D2A80C39615FFA5FD8F8D9CDEFAC7C4" box="[523,645,1965,1991]" count="1" pageId="24" pageNumber="25" type="subadult">1 subadult</specimenCount>
♀, same data as holotype (
<specimenCode id="DB8AE3319615FFA5FC4E8D9CD8F0C7C4" box="[970,1167,1965,1991]" pageId="24" pageNumber="25">CUMZ-D00098</specimenCode>
);
</materialsCitation>
<materialsCitation id="3B4441179615FFA5FB278D9CDDE4C7E9" ID-GBIF-Occurrence="2625222302" collectingDate="2012-05-14" collectionCode="ZMUC" collectorName="C. Sutcharit &amp; Animal Systematics Research Unit" country="Thailand" latitude="12.901944" location="Kaeng Krachan" longLatPrecision="21" longitude="99.59889" municipality="Kaeng Krachan District" pageId="24" pageNumber="25" specimenCount="1" specimenCount-male="1" stateProvince="Phetchaburi Province">
<specimenCount id="9D2A80C39615FFA5FB278D9CD8AFC7C4" box="[1187,1232,1963,1991]" count="1" pageId="24" pageNumber="25" type="male">1 ♂</specimenCount>
, same data as holotype (
<collectionCode id="ED3DD38F9615FFA5FEB78DE1DDF0C7E9" box="[307,399,2000,2026]" collectionName="Denmark, Kobenhavn [= Copenhagen], University of Copenhagen, Zoological Museum" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="24" pageNumber="25">ZMUC</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33618C19614FFA7FF398B3AD90FC21F" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="description">
<paragraph id="8B934B4A9614FFA4FF398B3ADD31C126" blockId="25.[189,334,267,293]" box="[189,334,267,293]" pageId="25" pageNumber="26">
<heading id="D0DBFC269614FFA4FF398B3ADD31C126" bold="true" box="[189,334,267,293]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="B95897589614FFA4FF398B3ADD31C126" bold="true" box="[189,334,267,293]" pageId="25" pageNumber="26">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B934B4A9614FFA4FF398B08DE3FC1BE" blockId="25.[189,1399,313,446]" pageId="25" pageNumber="26">
Adult males with 63–65 podous rings, no apodous rings. Length
<quantity id="4CD4E6AF9614FFA4FC428B0BD841C157" box="[966,1086,314,340]" metricMagnitude="-1" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="25" pageNumber="26" unit="cm" value="10.5" valueMax="11.0" valueMin="10.0">10–11 cm</quantity>
, width
<quantity id="4CD4E6AF9614FFA4FB128B08D957C157" box="[1174,1320,313,340]" metricMagnitude="-3" metricUnit="m" metricValue="5.55" metricValueMax="5.7" metricValueMin="5.4" pageId="25" pageNumber="26" unit="mm" value="5.55" valueMax="5.7" valueMin="5.4">5.4–5.7 mm</quantity>
. Adult females with 61–63 podous rings, no apodous rings. Length
<quantity id="4CD4E6AF9614FFA4FCFC8B6CDF91C174" box="[888,1006,349,375]" metricMagnitude="-1" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="25" pageNumber="26" unit="cm" value="12.5" valueMax="13.0" valueMin="12.0">12–13 cm</quantity>
, width
<quantity id="4CD4E6AF9614FFA4FBC68B6DD8AEC174" box="[1090,1233,348,375]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" metricValueMax="6.9" metricValueMin="6.7" pageId="25" pageNumber="26" unit="mm" value="6.8" valueMax="6.9" valueMin="6.7">6.7–6.9 mm</quantity>
. Colour in life unknown; preserved specimens with head, antennae, prozona, legs, epiproct, paraprocts and hypoproct brown; metazona reddish brown.
</paragraph>
<caption id="DF531BC29614FFA4FF398DB9DFEBC7EB" ID-DOI="http://doi.org/10.5281/zenodo.3840500" ID-Zenodo-Dep="3840500" httpUri="https://zenodo.org/record/3840500/files/figure.png" pageId="25" pageNumber="26" startId="25.[189,232,1928,1954]" targetBox="[206,1387,494,1893]" targetPageId="25">
<paragraph id="8B934B4A9614FFA4FF398DB9DFEBC7EB" blockId="25.[189,1399,1926,2025]" pageId="25" pageNumber="26">
<emphasis id="B95897589614FFA4FF398DB9DD75C7A1" bold="true" box="[189,266,1928,1954]" pageId="25" pageNumber="26">Fig. 8.</emphasis>
<taxonomicName id="4C2C30C99614FFA4FE968DB9DE6CC7A1" authority="Pimvichai &amp; Enghoff &amp; Panha &amp; Backeljau, 2016" authorityName="Pimvichai &amp; Enghoff &amp; Panha &amp; Backeljau" authorityYear="2016" box="[274,531,1928,1954]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="sutchariti" status="sp. nov.">
<emphasis id="B95897589614FFA4FE968DB9DE6CC7A1" box="[274,531,1928,1954]" italics="true" pageId="25" pageNumber="26">Thyropygus sutchariti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26B2A239614FFA4FD9F8DB9DE07C7A1" box="[539,632,1928,1954]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
, from Kaeng Krachan, holotype (CUMZ-D00090), ♂, gonopods.
<emphasis id="B95897589614FFA4FF398D9ADCADC7C6" bold="true" box="[189,210,1963,1989]" pageId="25" pageNumber="26">A</emphasis>
. Anterior view, left telopodite removed.
<emphasis id="B95897589614FFA4FD2F8D9ADEC0C7C6" bold="true" box="[683,703,1963,1989]" pageId="25" pageNumber="26">B</emphasis>
. Posterior view, left telopodite removed.
<emphasis id="B95897589614FFA4FB1B8D9AD8CBC7C6" bold="true" box="[1183,1204,1963,1989]" pageId="25" pageNumber="26">C</emphasis>
. Left telopodite, posterior-mesal view.
<emphasis id="B95897589614FFA4FE448DFFDDAAC7EB" bold="true" box="[448,469,1998,2024]" pageId="25" pageNumber="26">D</emphasis>
. Left telopodite, anterior-lateral view.
</paragraph>
</caption>
<paragraph id="8B934B4A9617FFA7FF398B3AD90FC21F" blockId="26.[189,1399,267,540]" pageId="26" pageNumber="27">
GONOPODS (
<figureCitation id="131757CF9617FFA7FEC78B3ADDFAC126" box="[323,389,267,293]" captionStart="Fig" captionStartId="25.[189,232,1928,1954]" captionTargetBox="[206,1387,494,1893]" captionTargetId="figure@25.[191,1404,481,1908]" captionTargetPageId="25" captionText="Fig. 8. Thyropygus sutchariti sp. nov., from Kaeng Krachan, holotype (CUMZ-D00090), ♂, gonopods. A. Anterior view, left telopodite removed. B. Posterior view, left telopodite removed. C. Left telopodite, posterior-mesal view. D. Left telopodite, anterior-lateral view." figureDoi="http://doi.org/10.5281/zenodo.3840500" httpUri="https://zenodo.org/record/3840500/files/figure.png" pageId="26" pageNumber="27">Fig. 8</figureCitation>
A–D). Anterior coxal fold (
<emphasis id="B95897589617FFA7FD418B3DDE9EC126" box="[709,737,268,293]" italics="true" pageId="26" pageNumber="27">ac</emphasis>
;
<figureCitation id="131757CF9617FFA7FD6B8B3ADF33C126" box="[751,844,267,293]" captionStart="Fig" captionStartId="25.[189,232,1928,1954]" captionTargetBox="[206,1387,494,1893]" captionTargetId="figure@25.[191,1404,481,1908]" captionTargetPageId="25" captionText="Fig. 8. Thyropygus sutchariti sp. nov., from Kaeng Krachan, holotype (CUMZ-D00090), ♂, gonopods. A. Anterior view, left telopodite removed. B. Posterior view, left telopodite removed. C. Left telopodite, posterior-mesal view. D. Left telopodite, anterior-lateral view." figureDoi="http://doi.org/10.5281/zenodo.3840500" httpUri="https://zenodo.org/record/3840500/files/figure.png" pageId="26" pageNumber="27">Fig. 8A</figureCitation>
): lateral process (
<emphasis id="B95897589617FFA7FB938B3AD843C126" box="[1047,1084,267,293]" italics="true" pageId="26" pageNumber="27">alp</emphasis>
) slender, regularly curved; mesal process (
<emphasis id="B95897589617FFA7FEFF8B1EDDD1C14B" box="[379,430,303,328]" italics="true" pageId="26" pageNumber="27">amp</emphasis>
) as long as
<emphasis id="B95897589617FFA7FDC98B1EDE0DC14A" box="[589,626,303,329]" italics="true" pageId="26" pageNumber="27">alp</emphasis>
, directed slightly obliquely disto-mesad, tip overlapping tip of opposite
<emphasis id="B95897589617FFA7FEAD8B62DD23C16F" box="[297,348,339,364]" italics="true" pageId="26" pageNumber="27">amp</emphasis>
. Posterior coxal fold (
<emphasis id="B95897589617FFA7FDE38B62DEFCC16F" box="[615,643,339,364]" italics="true" pageId="26" pageNumber="27">pc</emphasis>
;
<figureCitation id="131757CF9617FFA7FD178B63DE92C16F" box="[659,749,338,364]" captionStart="Fig" captionStartId="25.[189,232,1928,1954]" captionTargetBox="[206,1387,494,1893]" captionTargetId="figure@25.[191,1404,481,1908]" captionTargetPageId="25" captionText="Fig. 8. Thyropygus sutchariti sp. nov., from Kaeng Krachan, holotype (CUMZ-D00090), ♂, gonopods. A. Anterior view, left telopodite removed. B. Posterior view, left telopodite removed. C. Left telopodite, posterior-mesal view. D. Left telopodite, anterior-lateral view." figureDoi="http://doi.org/10.5281/zenodo.3840500" httpUri="https://zenodo.org/record/3840500/files/figure.png" pageId="26" pageNumber="27">Fig. 8B</figureCitation>
) basally with moderately high lateral paracoxites (
<emphasis id="B95897589617FFA7FACE8B62D919C16F" box="[1354,1382,339,364]" italics="true" pageId="26" pageNumber="27">px</emphasis>
), distally with two processes: mesal process (
<emphasis id="B95897589617FFA7FD3B8B47DE8DC18C" box="[703,754,374,399]" italics="true" pageId="26" pageNumber="27">pmp</emphasis>
) a rounded lobe; lateral process (
<emphasis id="B95897589617FFA7FBFF8B44D8DFC18C" box="[1147,1184,373,399]" italics="true" pageId="26" pageNumber="27">plp</emphasis>
) a massive, broad lobe, projecting laterad. Telopodite (
<figureCitation id="131757CF9617FFA7FDE28BA9DED6C1B1" box="[614,681,408,434]" captionStart="Fig" captionStartId="25.[189,232,1928,1954]" captionTargetBox="[206,1387,494,1893]" captionTargetId="figure@25.[191,1404,481,1908]" captionTargetPageId="25" captionText="Fig. 8. Thyropygus sutchariti sp. nov., from Kaeng Krachan, holotype (CUMZ-D00090), ♂, gonopods. A. Anterior view, left telopodite removed. B. Posterior view, left telopodite removed. C. Left telopodite, posterior-mesal view. D. Left telopodite, anterior-lateral view." figureDoi="http://doi.org/10.5281/zenodo.3840500" httpUri="https://zenodo.org/record/3840500/files/figure.png" pageId="26" pageNumber="27">Fig. 8</figureCitation>
C–D) leaving coxite between
<emphasis id="B95897589617FFA7FB838BA8D845C1B1" box="[1031,1082,409,434]" italics="true" pageId="26" pageNumber="27">pmp</emphasis>
and
<emphasis id="B95897589617FFA7FBF68BA9D8E8C1B1" box="[1138,1175,408,434]" italics="true" pageId="26" pageNumber="27">plp</emphasis>
; femoral spine (
<emphasis id="B95897589617FFA7FAD38BA9D913C1B1" box="[1367,1388,408,434]" italics="true" pageId="26" pageNumber="27">fe</emphasis>
) long, curved downward; telopodite distally to
<emphasis id="B95897589617FFA7FD6F8B8DDF7FC1D5" box="[747,768,444,470]" italics="true" pageId="26" pageNumber="27">fe</emphasis>
with large, round lobe (
<emphasis id="B95897589617FFA7FBAD8B8AD83FC1D6" box="[1065,1088,443,469]" italics="true" pageId="26" pageNumber="27">lo</emphasis>
) projecting distolaterally; tibial spine (
<emphasis id="B95897589617FFA7FED08BEEDD1BC1FB" box="[340,356,479,504]" italics="true" pageId="26" pageNumber="27">ti</emphasis>
) very long, curving in horizontal plane, tip close to basal part of
<emphasis id="B95897589617FFA7FBFE8BEED8EEC1FA" box="[1146,1169,479,505]" italics="true" pageId="26" pageNumber="27">lo</emphasis>
; spatulate lobe (
<emphasis id="B95897589617FFA7FADD8BEED912C1FA" box="[1369,1389,479,505]" italics="true" pageId="26" pageNumber="27">sl</emphasis>
) broad, rounded; palette (
<emphasis id="B95897589617FFA7FE598832DD85C21F" box="[477,506,515,540]" italics="true" pageId="26" pageNumber="27">pa</emphasis>
) simple, gutter-like, distally with about ten brownish blepharochaetae (
<emphasis id="B95897589617FFA7FAC68833D920C21F" box="[1346,1375,514,540]" italics="true" pageId="26" pageNumber="27">bp</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="C33618C19617FFA7FF398872DDF3C2AC" pageId="26" pageNumber="27" type="materials_examined">
<paragraph id="8B934B4A9617FFA7FF398872DD19C25E" blockId="26.[189,358,579,605]" box="[189,358,579,605]" pageId="26" pageNumber="27">
<heading id="D0DBFC269617FFA7FF398872DD19C25E" bold="true" box="[189,358,579,605]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="B95897589617FFA7FF398872DD19C25E" bold="true" box="[189,358,579,605]" pageId="26" pageNumber="27">DNA barcode</emphasis>
</heading>
</paragraph>
<paragraph id="8B934B4A9617FFA7FF398843DDF3C2AC" blockId="26.[189,1398,626,687]" pageId="26" pageNumber="27">
The GenBank accession number of the barcode of the
<typeStatus id="5497F5E89617FFA7FC118843DF83C28F" box="[917,1020,626,652]" pageId="26" pageNumber="27" type="holotype">holotype</typeStatus>
is
<accessionNumber id="947FD6A99617FFA7FBB68843D8CBC28E" box="[1074,1204,626,653]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KU306524" pageId="26" pageNumber="27">KU306524</accessionNumber>
(voucher code CUMZ-D00090).
</paragraph>
</subSubSection>
<subSubSection id="C33618C19617FFA7FF3988E7DE26C31C" pageId="26" pageNumber="27" type="distribution">
<paragraph id="8B934B4A9617FFA7FF3988E7DDBBC2F3" blockId="26.[189,452,726,752]" box="[189,452,726,752]" pageId="26" pageNumber="27">
<emphasis id="B95897589617FFA7FF3988E7DD29C2F3" bold="true" box="[189,342,726,752]" pageId="26" pageNumber="27">Distribution</emphasis>
(
<figureCitation id="131757CF9617FFA7FEE288E7DDC4C2F3" box="[358,443,726,752]" captionStart="Fig" captionStartId="30.[189,232,1998,2024]" captionTargetBox="[189,1398,829,1964]" captionTargetId="figure@30.[189,1398,829,1964]" captionTargetPageId="30" captionText="Fig. 12. Known distribution of the species of the T. opinatus subgroup." figureDoi="http://doi.org/10.5281/zenodo.3840510" httpUri="https://zenodo.org/record/3840510/files/figure.png" pageId="26" pageNumber="27">Fig. 12</figureCitation>
)
</paragraph>
<paragraph id="8B934B4A9617FFA7FF398934DE26C31C" blockId="26.[189,601,773,799]" box="[189,601,773,799]" pageId="26" pageNumber="27">
Known only from the
<typeStatus id="5497F5E89617FFA7FE468934DD8AC31C" box="[450,501,773,799]" pageId="26" pageNumber="27">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>
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